plazi/treatments-xml
2
0
Fork 0
Code Issues Pull requests Releases Activity
treatments-xml/data/03/9D/09/039D09066501FFECFF38FEACFC2EF830.xml
maintenance bc2564723d added 03
2024-06-21 12:22:17 +02:00

311 lines
54 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="BEA2CE193BDB9F51F647BB8065F14BE1" ID-CLB-Dataset="28353" ID-DOI="10.11646/zootaxa.4567.2.3" ID-GBIF-Dataset="7bd398c4-9441-4286-b150-0c421f91ec7e" ID-ISSN="1175-5326" ID-Zenodo-Dep="2594934" ID-ZooBank="76FF8984-7ABC-4751-8FA9-943501F9D63A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1552656074612" checkinUser="plazi" docAuthor="Rafael, José Albertino &amp; Marques, Dayse Willkenia Almeida" docDate="2019" docId="039D09066501FFECFF38FEACFC2EF830" docLanguage="en" docName="zootaxa.4567.2.3.pdf" docOrigin="Zootaxa 4567 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Macrostomus trifidus Rafael &amp; Marques 2019, sp. nov." docType="treatment" docUuid="809DA5CB-62C5-4153-873D-8097436197E1" docUuidSource="ZooBank" docVersion="9" lastPageNumber="269" masterDocId="FFA4717E6511FFFEFFAFFFB1FFCEFFE6" masterDocTitle="Five new species of Macrostomus Wiedemann and a checklist of Empididae (s. str.) (Diptera) from Bolivia" masterLastPageNumber="275" masterPageNumber="251" pageNumber="267" updateTime="1698701900396" updateUser="ExternalLinkService">
<mods:mods id="C664C7BF23513EAF73DE0CC63667EA0B" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="FB5A14007E916A3300201EB7565D9607">
<mods:title id="75A3B156AAC255BEB4A5FCAABB1CFD56">Five new species of Macrostomus Wiedemann and a checklist of Empididae (s. str.) (Diptera) from Bolivia</mods:title>
</mods:titleInfo>
<mods:name id="1316B64303DAB2DCD50BFF3E6BF1E7CA" type="personal">
<mods:role id="809938CF1499E494D53A968A64F69E4E">
<mods:roleTerm id="24F9017C2D421462C9297B5D2FD75818">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5124D04C8008F4DCB6BB962507518FCD">Rafael, José Albertino</mods:namePart>
</mods:name>
<mods:name id="ECF86CC107665AAA097C40A7932A026B" type="personal">
<mods:role id="D31133EA07E778D0D74365A96BC9C219">
<mods:roleTerm id="313759F7F403796EBC0B68299DCA3719">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8E5F9EDED627606ED288A9D3571C217F">Marques, Dayse Willkenia Almeida</mods:namePart>
</mods:name>
<mods:typeOfResource id="8A64C8032A0BA1D5B63FFEBFC08EAE74">text</mods:typeOfResource>
<mods:relatedItem id="06E7CAFD3BD899A78B335AB8C7E17C28" type="host">
<mods:titleInfo id="2BFA37E0B8DE1F88BE6C8A8B47BAF7A6">
<mods:title id="A6841BA51AC7C49B91E4DA2A8C0ED841">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="9184459CFB8FE6468D9C1E92AADA9C51">
<mods:date id="B6A695072DADA4D09EF3B8CC8A55D35C">2019</mods:date>
<mods:detail id="35C9B636A939F909067BCD7AB6E0B1FD" type="pubDate">
<mods:number id="2E6480DCFBF2987533DC1ABF5C59E41A">2019-03-15</mods:number>
</mods:detail>
<mods:detail id="B8F7FB62EF7313A1568A8C73CFA9D23A" type="volume">
<mods:number id="59F2E17786059EA2B965ECFB16AD9714">4567</mods:number>
</mods:detail>
<mods:detail id="1AB76E2D43CD3FDFF679D737C5C3B296" type="issue">
<mods:number id="A09F1442F74AB5A5AF795C495BAF2011">2</mods:number>
</mods:detail>
<mods:extent id="ED3DC18DD9683F39BEB9A63FFD94D548" unit="page">
<mods:start id="EE334496EA979A49C3AE3EDF1637D877">251</mods:start>
<mods:end id="A64B6DCDE7A2F1FF8544880EE4D10BAE">275</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="84CE8FFF0E493125E38C61A579168405">journal article</mods:classification>
<mods:identifier id="732315E02ECE2503F998EEA4F347C383" type="CLB-Dataset">28353</mods:identifier>
<mods:identifier id="3C3BFCE034ED22F49F7EEA17206974CD" type="DOI">10.11646/zootaxa.4567.2.3</mods:identifier>
<mods:identifier id="34702798680A6ED9B3E2021AB3FB57E0" type="GBIF-Dataset">7bd398c4-9441-4286-b150-0c421f91ec7e</mods:identifier>
<mods:identifier id="73ACA47E1F7874EA73793DD471C5E779" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="872637C28D260587AC64895B939DDEFA" type="Zenodo-Dep">2594934</mods:identifier>
<mods:identifier id="3AF8B16C8E1D260DCA5C20C9983330CD" type="ZooBank">76FF8984-7ABC-4751-8FA9-943501F9D63A</mods:identifier>
</mods:mods>
<treatment id="039D09066501FFECFF38FEACFC2EF830" ID-DOI="http://doi.org/10.5281/zenodo.5925358" ID-GBIF-Taxon="154402319" ID-Zenodo-Dep="5925358" LSID="urn:lsid:zoobank.org:act:809DA5CB-62C5-4153-873D-8097436197E1" httpUri="http://treatment.plazi.org/id/039D09066501FFECFF38FEACFC2EF830" lastPageId="18" lastPageNumber="269" pageId="16" pageNumber="267">
<subSubSection id="C32EEB9B6501FFEEFF38FEACFE31FED1" box="[151,511,285,311]" pageId="16" pageNumber="267" type="nomenclature">
<paragraph id="8B8BB8106501FFEEFF38FEACFE31FED1" blockId="16.[151,952,285,381]" box="[151,511,285,311]" pageId="16" pageNumber="267">
<heading id="D0C30F7C6501FFEEFF38FEACFE31FED1" bold="true" box="[151,511,285,311]" fontSize="11" level="1" pageId="16" pageNumber="267" reason="1">
<emphasis id="B94064026501FFEEFF38FEACFE31FED1" bold="true" box="[151,511,285,311]" pageId="16" pageNumber="267">
<taxonomicName id="4C34C3936501FFEEFF38FEACFE55FED1" authority="Rafael &amp; Marques, 2019" authorityName="Rafael &amp; Marques" authorityYear="2019" box="[151,411,285,311]" class="Insecta" family="Empididae" genus="Macrostomus" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="267" phylum="Arthropoda" rank="species" species="trifidus" status="sp. nov.">
<emphasis id="B94064026501FFEEFF38FEACFE55FED1" bold="true" box="[151,411,285,311]" italics="true" pageId="16" pageNumber="267">Macrostomus trifidus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A273D9796501FFEEFE0DFEACFE31FED1" box="[418,511,285,311]" pageId="16" pageNumber="267" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32EEB9B6501FFEEFF38FE8EFC76FE9B" pageId="16" pageNumber="267" type="description">
<paragraph id="8B8BB8106501FFEEFF38FE8EFE5EFEBE" blockId="16.[151,952,285,381]" box="[151,400,319,344]" pageId="16" pageNumber="267">
(
<figureCitation id="130FA4956501FFEEFF30FE8EFF3EFEBE" box="[159,240,319,344]" captionStart="FIGURE 11" captionStartId="17.[151,250,1795,1817]" captionTargetBox="[222,1364,200,1778]" captionTargetId="figure@17.[221,1364,193,1780]" captionTargetPageId="17" captionText="FIGURE 11. Macrostomus trifidus sp. nov., holotype ♂. A. Habitus, lateral view; B. Apex of abdomen after macerated, lateral view (pieces with white outline); C. Tergite 8, dorsal view; D. Tergite and sternite 8, lateral view; E. Sternite 8, ventral view; F. Epandrium detached, lateral view; G. Epandrium detached, posterior view; H. Terminalia, posterior view; I. Terminalia (epandrium, hypandrium and phallus removed), right lateral view; J. Branchs of posterior cerci, posterolateral view (black arrows indicating the three branchs); K. Detail of cercal bridge of anterior cercus, ventral projection of cercus and dorsal lobe of hypoproct, anterior view; L. Posterior cercus, posterior view; M. Hypandrium, ejaculatory apodeme and phallus, lateral view; N. Hypandrium, posterior view; O. Holotype labels. Scale bars, figures A = 1 mm; B–N = 0.2 mm. Abbreviations: cbrd, cercal bridge of anterior cercus; vpc, ventral projection of cercus; DHyL, Dorsal hypoproctal lobe;" httpUri="https://zenodo.org/record/2594957/files/figure.png" pageId="16" pageNumber="267">Figs 11</figureCitation>
A–O, 12A–E)
</paragraph>
<paragraph id="8B8BB8106501FFEEFF38FED5FC76FE9B" blockId="16.[151,952,285,381]" box="[151,952,356,381]" pageId="16" pageNumber="267">
<uri id="FFA5B4126501FFEEFF38FED5FC76FE9B" box="[151,952,356,381]" pageId="16" pageNumber="267">
urn:lsid:zoobank.org:act:
<uuid id="FF9282C56501FFEEFE06FED5FC76FE9B" box="[425,952,356,381]" pageId="16" pageNumber="267">809DA5CB-62C5-4153-873D-8097436197E1</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C32EEB9B6501FFEEFF38FE17FEE0FD06" pageId="16" pageNumber="267" type="diagnosis">
<paragraph id="8B8BB8106501FFEEFF38FE17FEE0FD06" blockId="16.[151,1437,422,2028]" pageId="16" pageNumber="267">
<emphasis id="B94064026501FFEEFF38FE17FEDFFE59" bold="true" box="[151,273,422,447]" pageId="16" pageNumber="267">Diagnosis.</emphasis>
Thorax shiny yellow with scutellum and mediotergite brown, gray-yellow pruinosity denser on disc of scutum and on scutellum. Wing brown, infuscation darker along costal and r
<subScript id="17B0BA556501FFEEFC53FE68FBCAFE01" attach="left" box="[1020,1028,473,487]" fontSize="6" pageId="16" pageNumber="267">1</subScript>
cells and apex of cell r
<subScript id="17B0BA556501FFEEFABFFE68FAE6FE01" attach="right" box="[1296,1320,473,487]" fontSize="6" pageId="16" pageNumber="267">2+3</subScript>
. Tergite 8 elongate distally, basally more sclerotized with a sclerotized protuberance at base of the projection connecting with mesial cercal bridge of anterior cercus. Sternite 8 with shallow basal medial sinus, with saddle-shaped posterodorsal protuberance. Epandrium with longer setae posteriorly and mainly on posterior lobe. Anterior cercus multilobate, with few longer setae posterodorsally; posterior cercus trifid, all branch yellowish, inwardly curved; ventral projection of cercus downwardly directed, horizontally placed distally when viewed anteriorly or posteriorly, glabrous and not touching each other. Hypandrium well sclerotized, same width throughout, with a hook distally.
</paragraph>
</subSubSection>
<subSubSection id="C32EEB9B6501FFECFF68FD5AFD59F98B" lastPageId="18" lastPageNumber="269" pageId="16" pageNumber="267" type="description">
<paragraph id="8B8BB8106501FFEEFF68FD5AFCB7F8DF" blockId="16.[151,1437,422,2028]" pageId="16" pageNumber="267">
<emphasis id="B94064026501FFEEFF68FD5AFE97FCE2" bold="true" box="[199,345,747,772]" pageId="16" pageNumber="267">Description.</emphasis>
<typeStatus id="548F06B26501FFEEFECDFD5AFE04FCE2" box="[354,458,747,772]" pageId="16" pageNumber="267" type="holotype">Holotype</typeStatus>
<specimenCount id="9D3273996501FFEEFE7BFD5DFE27FCE5" box="[468,489,748,771]" pageId="16" pageNumber="267" type="male">♂</specimenCount>
(
<figureCitation id="130FA4956501FFEEFE54FD5AFDA8FCE5" box="[507,614,747,772]" captionStart="FIGURE 11" captionStartId="17.[151,250,1795,1817]" captionTargetBox="[222,1364,200,1778]" captionTargetId="figure@17.[221,1364,193,1780]" captionTargetPageId="17" captionText="FIGURE 11. Macrostomus trifidus sp. nov., holotype ♂. A. Habitus, lateral view; B. Apex of abdomen after macerated, lateral view (pieces with white outline); C. Tergite 8, dorsal view; D. Tergite and sternite 8, lateral view; E. Sternite 8, ventral view; F. Epandrium detached, lateral view; G. Epandrium detached, posterior view; H. Terminalia, posterior view; I. Terminalia (epandrium, hypandrium and phallus removed), right lateral view; J. Branchs of posterior cerci, posterolateral view (black arrows indicating the three branchs); K. Detail of cercal bridge of anterior cercus, ventral projection of cercus and dorsal lobe of hypoproct, anterior view; L. Posterior cercus, posterior view; M. Hypandrium, ejaculatory apodeme and phallus, lateral view; N. Hypandrium, posterior view; O. Holotype labels. Scale bars, figures A = 1 mm; B–N = 0.2 mm. Abbreviations: cbrd, cercal bridge of anterior cercus; vpc, ventral projection of cercus; DHyL, Dorsal hypoproctal lobe;" httpUri="https://zenodo.org/record/2594957/files/figure.png" pageId="16" pageNumber="267">Fig. 11A</figureCitation>
). Body length
<quantity id="4CCC15F56501FFEEFCBCFD5AFCA1FCE5" box="[787,879,747,771]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="16" pageNumber="267" unit="mm" value="3.3">3.3 mm</quantity>
; wing length
<quantity id="4CCC15F56501FFEEFBA0FD5AFBA5FCE5" box="[1039,1131,747,771]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="16" pageNumber="267" unit="mm" value="3.8">3.8 mm</quantity>
. Head narrowly dichoptic. Frons shiny black, slightly narrower than anterior ocellus width. One pair of proclinate ocellar setae. Face slightly narrower than frons width. Postcranium shiny black, entirely and sparsely gray pruinose. Inner vertical seta stouter than outer vertical seta, latter subequal in length to uniseriate postocular row of setae; 2 pairs of dorsal occipital setae. Gena with 2 setae and postgena with 1 weak seta. Antenna velvety black with postpedicel short pubescent. Proboscis dark brown with median third yellow and apical 2/3 of labellum yellow. Palpus black. Thorax shiny yellow with scutellum and mediotergite brown; scutum gray-yellow pruinose posteriorly and on scutellum. Thoracic chaetotaxy: 3 antepronotals; 1 postpronotal stout and 2 minutes; 3–4 dorsocentral setae; 1 presutural supra-alar; 0 postsutural supra-alar; 1 postalar; 4 proepisternals weak; 1 notopleural stout, 1 weak; 2 pairs of scutellars, the outer pair weaker; 4 laterotergitals. Legs shiny yellow with apex of fore and mid tarsi, distal 1/3 of hind femur, entire hind tibia and tarsus brown to black. Hind femur slightly clavate. Legs longer setae: hind femur with 1 slightly stouter anteroventral seta; hind tibia with 5 longer dorsal setae being with one at basal fourth, one near middle and three along distal third; hind tarsomere 1 with 1 longer median dorsal seta. Wing (
<figureCitation id="130FA4956501FFEEFB4DFB26FA88FB56" box="[1250,1350,1175,1200]" captionStart="FIGURE 11" captionStartId="17.[151,250,1795,1817]" captionTargetBox="[222,1364,200,1778]" captionTargetId="figure@17.[221,1364,193,1780]" captionTargetPageId="17" captionText="FIGURE 11. Macrostomus trifidus sp. nov., holotype ♂. A. Habitus, lateral view; B. Apex of abdomen after macerated, lateral view (pieces with white outline); C. Tergite 8, dorsal view; D. Tergite and sternite 8, lateral view; E. Sternite 8, ventral view; F. Epandrium detached, lateral view; G. Epandrium detached, posterior view; H. Terminalia, posterior view; I. Terminalia (epandrium, hypandrium and phallus removed), right lateral view; J. Branchs of posterior cerci, posterolateral view (black arrows indicating the three branchs); K. Detail of cercal bridge of anterior cercus, ventral projection of cercus and dorsal lobe of hypoproct, anterior view; L. Posterior cercus, posterior view; M. Hypandrium, ejaculatory apodeme and phallus, lateral view; N. Hypandrium, posterior view; O. Holotype labels. Scale bars, figures A = 1 mm; B–N = 0.2 mm. Abbreviations: cbrd, cercal bridge of anterior cercus; vpc, ventral projection of cercus; DHyL, Dorsal hypoproctal lobe;" httpUri="https://zenodo.org/record/2594957/files/figure.png" pageId="16" pageNumber="267">Fig. 11A</figureCitation>
) brown infuscated, darker along costal and r
<subScript id="17B0BA556501FFEEFD98FB7BFDF1FB3E" attach="left" box="[567,575,1226,1240]" fontSize="6" pageId="16" pageNumber="267">1</subScript>
cells and apex of cell r
<subScript id="17B0BA556501FFEEFCE1FB7BFCA8FB3E" attach="left" box="[846,870,1226,1240]" fontSize="6" pageId="16" pageNumber="267">2+3</subScript>
. Veins M
<subScript id="17B0BA556501FFEEFC79FB7BFC10FB3E" attach="left" box="[982,990,1226,1240]" fontSize="6" pageId="16" pageNumber="267">1</subScript>
, M
<subScript id="17B0BA556501FFEEFBA8FB7BFBC1FB3E" attach="left" box="[1031,1039,1226,1240]" fontSize="6" pageId="16" pageNumber="267">2</subScript>
and CuA+CuP weakly evanescent distally. Last section of vein M
<subScript id="17B0BA556501FFEEFE41FB40FE38FB19" attach="left" box="[494,502,1265,1279]" fontSize="6" pageId="16" pageNumber="267">4</subScript>
slightly longer than posterior length of elongate and blunt cell dm. Anal lobe very narrow. Halter light yellow. Abdomen shiny black. Tergite 1–2 with longer setae posteriorly. Tergite
<quantity id="4CCC15F56501FFEEFA8FFAB8FA83FAC7" box="[1312,1357,1288,1313]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="16" pageNumber="267" unit="in" value="8.0">8 in</quantity>
lateral view (
<figureCitation id="130FA4956501FFEEFF4EFA9DFEBDFAA3" box="[225,371,1324,1349]" captionStart="FIGURE 11" captionStartId="17.[151,250,1795,1817]" captionTargetBox="[222,1364,200,1778]" captionTargetId="figure@17.[221,1364,193,1780]" captionTargetPageId="17" captionText="FIGURE 11. Macrostomus trifidus sp. nov., holotype ♂. A. Habitus, lateral view; B. Apex of abdomen after macerated, lateral view (pieces with white outline); C. Tergite 8, dorsal view; D. Tergite and sternite 8, lateral view; E. Sternite 8, ventral view; F. Epandrium detached, lateral view; G. Epandrium detached, posterior view; H. Terminalia, posterior view; I. Terminalia (epandrium, hypandrium and phallus removed), right lateral view; J. Branchs of posterior cerci, posterolateral view (black arrows indicating the three branchs); K. Detail of cercal bridge of anterior cercus, ventral projection of cercus and dorsal lobe of hypoproct, anterior view; L. Posterior cercus, posterior view; M. Hypandrium, ejaculatory apodeme and phallus, lateral view; N. Hypandrium, posterior view; O. Holotype labels. Scale bars, figures A = 1 mm; B–N = 0.2 mm. Abbreviations: cbrd, cercal bridge of anterior cercus; vpc, ventral projection of cercus; DHyL, Dorsal hypoproctal lobe;" httpUri="https://zenodo.org/record/2594957/files/figure.png" pageId="16" pageNumber="267">Figs 11B, D</figureCitation>
) curved, with sclerotized protuberance at base of projection connecting with mesial cercal bridge of anterior cercus; in dorsal view (
<figureCitation id="130FA4956501FFEEFDC2FAE1FD1FFA8E" box="[621,721,1360,1385]" captionStart="FIGURE 11" captionStartId="17.[151,250,1795,1817]" captionTargetBox="[222,1364,200,1778]" captionTargetId="figure@17.[221,1364,193,1780]" captionTargetPageId="17" captionText="FIGURE 11. Macrostomus trifidus sp. nov., holotype ♂. A. Habitus, lateral view; B. Apex of abdomen after macerated, lateral view (pieces with white outline); C. Tergite 8, dorsal view; D. Tergite and sternite 8, lateral view; E. Sternite 8, ventral view; F. Epandrium detached, lateral view; G. Epandrium detached, posterior view; H. Terminalia, posterior view; I. Terminalia (epandrium, hypandrium and phallus removed), right lateral view; J. Branchs of posterior cerci, posterolateral view (black arrows indicating the three branchs); K. Detail of cercal bridge of anterior cercus, ventral projection of cercus and dorsal lobe of hypoproct, anterior view; L. Posterior cercus, posterior view; M. Hypandrium, ejaculatory apodeme and phallus, lateral view; N. Hypandrium, posterior view; O. Holotype labels. Scale bars, figures A = 1 mm; B–N = 0.2 mm. Abbreviations: cbrd, cercal bridge of anterior cercus; vpc, ventral projection of cercus; DHyL, Dorsal hypoproctal lobe;" httpUri="https://zenodo.org/record/2594957/files/figure.png" pageId="16" pageNumber="267">Fig. 11C</figureCitation>
) distally with elongate projection (setae broken on projection); basally more sclerotized with deep basal cleft. Sternite 8 (
<figureCitation id="130FA4956501FFEEFCEEFAC5FC1FFA6A" box="[833,977,1396,1421]" captionStart="FIGURE 11" captionStartId="17.[151,250,1795,1817]" captionTargetBox="[222,1364,200,1778]" captionTargetId="figure@17.[221,1364,193,1780]" captionTargetPageId="17" captionText="FIGURE 11. Macrostomus trifidus sp. nov., holotype ♂. A. Habitus, lateral view; B. Apex of abdomen after macerated, lateral view (pieces with white outline); C. Tergite 8, dorsal view; D. Tergite and sternite 8, lateral view; E. Sternite 8, ventral view; F. Epandrium detached, lateral view; G. Epandrium detached, posterior view; H. Terminalia, posterior view; I. Terminalia (epandrium, hypandrium and phallus removed), right lateral view; J. Branchs of posterior cerci, posterolateral view (black arrows indicating the three branchs); K. Detail of cercal bridge of anterior cercus, ventral projection of cercus and dorsal lobe of hypoproct, anterior view; L. Posterior cercus, posterior view; M. Hypandrium, ejaculatory apodeme and phallus, lateral view; N. Hypandrium, posterior view; O. Holotype labels. Scale bars, figures A = 1 mm; B–N = 0.2 mm. Abbreviations: cbrd, cercal bridge of anterior cercus; vpc, ventral projection of cercus; DHyL, Dorsal hypoproctal lobe;" httpUri="https://zenodo.org/record/2594957/files/figure.png" pageId="16" pageNumber="267">Figs 11D, E</figureCitation>
) with shallow basal medial sinus, with saddle-shaped posterodorsal protuberance. Tergite 8 and sternite 8 strongly connected anteriorly. Epandrium (
<figureCitation id="130FA4956501FFEEFAC5FA26FF2AFA32" captionStart="FIGURE 11" captionStartId="17.[151,250,1795,1817]" captionTargetBox="[222,1364,200,1778]" captionTargetId="figure@17.[221,1364,193,1780]" captionTargetPageId="17" captionText="FIGURE 11. Macrostomus trifidus sp. nov., holotype ♂. A. Habitus, lateral view; B. Apex of abdomen after macerated, lateral view (pieces with white outline); C. Tergite 8, dorsal view; D. Tergite and sternite 8, lateral view; E. Sternite 8, ventral view; F. Epandrium detached, lateral view; G. Epandrium detached, posterior view; H. Terminalia, posterior view; I. Terminalia (epandrium, hypandrium and phallus removed), right lateral view; J. Branchs of posterior cerci, posterolateral view (black arrows indicating the three branchs); K. Detail of cercal bridge of anterior cercus, ventral projection of cercus and dorsal lobe of hypoproct, anterior view; L. Posterior cercus, posterior view; M. Hypandrium, ejaculatory apodeme and phallus, lateral view; N. Hypandrium, posterior view; O. Holotype labels. Scale bars, figures A = 1 mm; B–N = 0.2 mm. Abbreviations: cbrd, cercal bridge of anterior cercus; vpc, ventral projection of cercus; DHyL, Dorsal hypoproctal lobe;" httpUri="https://zenodo.org/record/2594957/files/figure.png" pageId="16" pageNumber="267">Figs 11F, G</figureCitation>
) with very small anterodorsal excision and very small anteroventral excision; longer setae posteriorly and mainly on posterior lobe; anterior and posterior lobes of same length; epandrium in posterior view (
<figureCitation id="130FA4956501FFEEFB59FA6EFA94FA11" box="[1270,1370,1503,1528]" captionStart="FIGURE 11" captionStartId="17.[151,250,1795,1817]" captionTargetBox="[222,1364,200,1778]" captionTargetId="figure@17.[221,1364,193,1780]" captionTargetPageId="17" captionText="FIGURE 11. Macrostomus trifidus sp. nov., holotype ♂. A. Habitus, lateral view; B. Apex of abdomen after macerated, lateral view (pieces with white outline); C. Tergite 8, dorsal view; D. Tergite and sternite 8, lateral view; E. Sternite 8, ventral view; F. Epandrium detached, lateral view; G. Epandrium detached, posterior view; H. Terminalia, posterior view; I. Terminalia (epandrium, hypandrium and phallus removed), right lateral view; J. Branchs of posterior cerci, posterolateral view (black arrows indicating the three branchs); K. Detail of cercal bridge of anterior cercus, ventral projection of cercus and dorsal lobe of hypoproct, anterior view; L. Posterior cercus, posterior view; M. Hypandrium, ejaculatory apodeme and phallus, lateral view; N. Hypandrium, posterior view; O. Holotype labels. Scale bars, figures A = 1 mm; B–N = 0.2 mm. Abbreviations: cbrd, cercal bridge of anterior cercus; vpc, ventral projection of cercus; DHyL, Dorsal hypoproctal lobe;" httpUri="https://zenodo.org/record/2594957/files/figure.png" pageId="16" pageNumber="267">Fig. 11G</figureCitation>
) with stout protuberance mesially and several yellow distal short setae posterodorsally. Anterior cercus well sclerotized, upwardly directed, multilobate, with few longer setae posterodorsally; when viewed anteriorly (
<figureCitation id="130FA4956501FFEEFB6EF997FA84F9D9" box="[1217,1354,1574,1599]" captionStart="FIGURE 11" captionStartId="17.[151,250,1795,1817]" captionTargetBox="[222,1364,200,1778]" captionTargetId="figure@17.[221,1364,193,1780]" captionTargetPageId="17" captionText="FIGURE 11. Macrostomus trifidus sp. nov., holotype ♂. A. Habitus, lateral view; B. Apex of abdomen after macerated, lateral view (pieces with white outline); C. Tergite 8, dorsal view; D. Tergite and sternite 8, lateral view; E. Sternite 8, ventral view; F. Epandrium detached, lateral view; G. Epandrium detached, posterior view; H. Terminalia, posterior view; I. Terminalia (epandrium, hypandrium and phallus removed), right lateral view; J. Branchs of posterior cerci, posterolateral view (black arrows indicating the three branchs); K. Detail of cercal bridge of anterior cercus, ventral projection of cercus and dorsal lobe of hypoproct, anterior view; L. Posterior cercus, posterior view; M. Hypandrium, ejaculatory apodeme and phallus, lateral view; N. Hypandrium, posterior view; O. Holotype labels. Scale bars, figures A = 1 mm; B–N = 0.2 mm. Abbreviations: cbrd, cercal bridge of anterior cercus; vpc, ventral projection of cercus; DHyL, Dorsal hypoproctal lobe;" httpUri="https://zenodo.org/record/2594957/files/figure.png" pageId="16" pageNumber="267">Figs 11H, K</figureCitation>
) cercal bridge V-shaped, fused medially and ventrally. Median cercus (
<figureCitation id="130FA4956501FFEEFCC2F9FBFC07F985" box="[877,969,1610,1635]" captionStart="FIGURE 11" captionStartId="17.[151,250,1795,1817]" captionTargetBox="[222,1364,200,1778]" captionTargetId="figure@17.[221,1364,193,1780]" captionTargetPageId="17" captionText="FIGURE 11. Macrostomus trifidus sp. nov., holotype ♂. A. Habitus, lateral view; B. Apex of abdomen after macerated, lateral view (pieces with white outline); C. Tergite 8, dorsal view; D. Tergite and sternite 8, lateral view; E. Sternite 8, ventral view; F. Epandrium detached, lateral view; G. Epandrium detached, posterior view; H. Terminalia, posterior view; I. Terminalia (epandrium, hypandrium and phallus removed), right lateral view; J. Branchs of posterior cerci, posterolateral view (black arrows indicating the three branchs); K. Detail of cercal bridge of anterior cercus, ventral projection of cercus and dorsal lobe of hypoproct, anterior view; L. Posterior cercus, posterior view; M. Hypandrium, ejaculatory apodeme and phallus, lateral view; N. Hypandrium, posterior view; O. Holotype labels. Scale bars, figures A = 1 mm; B–N = 0.2 mm. Abbreviations: cbrd, cercal bridge of anterior cercus; vpc, ventral projection of cercus; DHyL, Dorsal hypoproctal lobe;" httpUri="https://zenodo.org/record/2594957/files/figure.png" pageId="16" pageNumber="267">
Fig.
<date id="FF8A9ED06501FFEEFC0BF9FAFC07F985" box="[932,969,1611,1635]" pageId="16" pageNumber="267" value="2011-01">11I</date>
</figureCitation>
) inverted C-shaped, with elongate setae, less sclerotized distally. Posterior cercus (
<figureCitation id="130FA4956501FFEEFDC0F9DCFD05F960" box="[623,715,1645,1670]" captionStart="FIGURE 11" captionStartId="17.[151,250,1795,1817]" captionTargetBox="[222,1364,200,1778]" captionTargetId="figure@17.[221,1364,193,1780]" captionTargetPageId="17" captionText="FIGURE 11. Macrostomus trifidus sp. nov., holotype ♂. A. Habitus, lateral view; B. Apex of abdomen after macerated, lateral view (pieces with white outline); C. Tergite 8, dorsal view; D. Tergite and sternite 8, lateral view; E. Sternite 8, ventral view; F. Epandrium detached, lateral view; G. Epandrium detached, posterior view; H. Terminalia, posterior view; I. Terminalia (epandrium, hypandrium and phallus removed), right lateral view; J. Branchs of posterior cerci, posterolateral view (black arrows indicating the three branchs); K. Detail of cercal bridge of anterior cercus, ventral projection of cercus and dorsal lobe of hypoproct, anterior view; L. Posterior cercus, posterior view; M. Hypandrium, ejaculatory apodeme and phallus, lateral view; N. Hypandrium, posterior view; O. Holotype labels. Scale bars, figures A = 1 mm; B–N = 0.2 mm. Abbreviations: cbrd, cercal bridge of anterior cercus; vpc, ventral projection of cercus; DHyL, Dorsal hypoproctal lobe;" httpUri="https://zenodo.org/record/2594957/files/figure.png" pageId="16" pageNumber="267">Fig. 11J</figureCitation>
) trifid, all branches yellow, inwardly curved, apical two setose. Ventral projection of cercus (
<figureCitation id="130FA4956501FFEEFE4EF920FD89F94C" box="[481,583,1681,1706]" captionStart="FIGURE 11" captionStartId="17.[151,250,1795,1817]" captionTargetBox="[222,1364,200,1778]" captionTargetId="figure@17.[221,1364,193,1780]" captionTargetPageId="17" captionText="FIGURE 11. Macrostomus trifidus sp. nov., holotype ♂. A. Habitus, lateral view; B. Apex of abdomen after macerated, lateral view (pieces with white outline); C. Tergite 8, dorsal view; D. Tergite and sternite 8, lateral view; E. Sternite 8, ventral view; F. Epandrium detached, lateral view; G. Epandrium detached, posterior view; H. Terminalia, posterior view; I. Terminalia (epandrium, hypandrium and phallus removed), right lateral view; J. Branchs of posterior cerci, posterolateral view (black arrows indicating the three branchs); K. Detail of cercal bridge of anterior cercus, ventral projection of cercus and dorsal lobe of hypoproct, anterior view; L. Posterior cercus, posterior view; M. Hypandrium, ejaculatory apodeme and phallus, lateral view; N. Hypandrium, posterior view; O. Holotype labels. Scale bars, figures A = 1 mm; B–N = 0.2 mm. Abbreviations: cbrd, cercal bridge of anterior cercus; vpc, ventral projection of cercus; DHyL, Dorsal hypoproctal lobe;" httpUri="https://zenodo.org/record/2594957/files/figure.png" pageId="16" pageNumber="267">Fig. 11K</figureCitation>
) downwardly directed, horizontally placed distally when viewed anteriorly or posteriorly, glabrous and not touching each other. Dorsal lobe of hypoproct (
<figureCitation id="130FA4956501FFEEFBB4F904FB4FF928" box="[1051,1153,1717,1742]" captionStart="FIGURE 11" captionStartId="17.[151,250,1795,1817]" captionTargetBox="[222,1364,200,1778]" captionTargetId="figure@17.[221,1364,193,1780]" captionTargetPageId="17" captionText="FIGURE 11. Macrostomus trifidus sp. nov., holotype ♂. A. Habitus, lateral view; B. Apex of abdomen after macerated, lateral view (pieces with white outline); C. Tergite 8, dorsal view; D. Tergite and sternite 8, lateral view; E. Sternite 8, ventral view; F. Epandrium detached, lateral view; G. Epandrium detached, posterior view; H. Terminalia, posterior view; I. Terminalia (epandrium, hypandrium and phallus removed), right lateral view; J. Branchs of posterior cerci, posterolateral view (black arrows indicating the three branchs); K. Detail of cercal bridge of anterior cercus, ventral projection of cercus and dorsal lobe of hypoproct, anterior view; L. Posterior cercus, posterior view; M. Hypandrium, ejaculatory apodeme and phallus, lateral view; N. Hypandrium, posterior view; O. Holotype labels. Scale bars, figures A = 1 mm; B–N = 0.2 mm. Abbreviations: cbrd, cercal bridge of anterior cercus; vpc, ventral projection of cercus; DHyL, Dorsal hypoproctal lobe;" httpUri="https://zenodo.org/record/2594957/files/figure.png" pageId="16" pageNumber="267">Fig. 11K</figureCitation>
) larger with longer setae than narrower ventral lobe. Hypandrium (
<figureCitation id="130FA4956501FFEEFD28F968FCEAF917" box="[647,804,1753,1778]" captionStart="FIGURE 11" captionStartId="17.[151,250,1795,1817]" captionTargetBox="[222,1364,200,1778]" captionTargetId="figure@17.[221,1364,193,1780]" captionTargetPageId="17" captionText="FIGURE 11. Macrostomus trifidus sp. nov., holotype ♂. A. Habitus, lateral view; B. Apex of abdomen after macerated, lateral view (pieces with white outline); C. Tergite 8, dorsal view; D. Tergite and sternite 8, lateral view; E. Sternite 8, ventral view; F. Epandrium detached, lateral view; G. Epandrium detached, posterior view; H. Terminalia, posterior view; I. Terminalia (epandrium, hypandrium and phallus removed), right lateral view; J. Branchs of posterior cerci, posterolateral view (black arrows indicating the three branchs); K. Detail of cercal bridge of anterior cercus, ventral projection of cercus and dorsal lobe of hypoproct, anterior view; L. Posterior cercus, posterior view; M. Hypandrium, ejaculatory apodeme and phallus, lateral view; N. Hypandrium, posterior view; O. Holotype labels. Scale bars, figures A = 1 mm; B–N = 0.2 mm. Abbreviations: cbrd, cercal bridge of anterior cercus; vpc, ventral projection of cercus; DHyL, Dorsal hypoproctal lobe;" httpUri="https://zenodo.org/record/2594957/files/figure.png" pageId="16" pageNumber="267">Figs 11M, N</figureCitation>
) well sclerotized, same width throughout, with hook distally, longer setae restricted to posterior median third. Ejaculatory apodeme (
<figureCitation id="130FA4956501FFEEFBBCF94CFBB3F8F3" box="[1043,1149,1789,1814]" captionStart="FIGURE 11" captionStartId="17.[151,250,1795,1817]" captionTargetBox="[222,1364,200,1778]" captionTargetId="figure@17.[221,1364,193,1780]" captionTargetPageId="17" captionText="FIGURE 11. Macrostomus trifidus sp. nov., holotype ♂. A. Habitus, lateral view; B. Apex of abdomen after macerated, lateral view (pieces with white outline); C. Tergite 8, dorsal view; D. Tergite and sternite 8, lateral view; E. Sternite 8, ventral view; F. Epandrium detached, lateral view; G. Epandrium detached, posterior view; H. Terminalia, posterior view; I. Terminalia (epandrium, hypandrium and phallus removed), right lateral view; J. Branchs of posterior cerci, posterolateral view (black arrows indicating the three branchs); K. Detail of cercal bridge of anterior cercus, ventral projection of cercus and dorsal lobe of hypoproct, anterior view; L. Posterior cercus, posterior view; M. Hypandrium, ejaculatory apodeme and phallus, lateral view; N. Hypandrium, posterior view; O. Holotype labels. Scale bars, figures A = 1 mm; B–N = 0.2 mm. Abbreviations: cbrd, cercal bridge of anterior cercus; vpc, ventral projection of cercus; DHyL, Dorsal hypoproctal lobe;" httpUri="https://zenodo.org/record/2594957/files/figure.png" pageId="16" pageNumber="267">Fig. 11M</figureCitation>
) tetralamellar, all lamella subequal. Phallus (
<figureCitation id="130FA4956501FFEEFEC6F891FE37F8DF" box="[361,505,1824,1849]" captionStart="FIGURE 11" captionStartId="17.[151,250,1795,1817]" captionTargetBox="[222,1364,200,1778]" captionTargetId="figure@17.[221,1364,193,1780]" captionTargetPageId="17" captionText="FIGURE 11. Macrostomus trifidus sp. nov., holotype ♂. A. Habitus, lateral view; B. Apex of abdomen after macerated, lateral view (pieces with white outline); C. Tergite 8, dorsal view; D. Tergite and sternite 8, lateral view; E. Sternite 8, ventral view; F. Epandrium detached, lateral view; G. Epandrium detached, posterior view; H. Terminalia, posterior view; I. Terminalia (epandrium, hypandrium and phallus removed), right lateral view; J. Branchs of posterior cerci, posterolateral view (black arrows indicating the three branchs); K. Detail of cercal bridge of anterior cercus, ventral projection of cercus and dorsal lobe of hypoproct, anterior view; L. Posterior cercus, posterior view; M. Hypandrium, ejaculatory apodeme and phallus, lateral view; N. Hypandrium, posterior view; O. Holotype labels. Scale bars, figures A = 1 mm; B–N = 0.2 mm. Abbreviations: cbrd, cercal bridge of anterior cercus; vpc, ventral projection of cercus; DHyL, Dorsal hypoproctal lobe;" httpUri="https://zenodo.org/record/2594957/files/figure.png" pageId="16" pageNumber="267">Figs 11M, N</figureCitation>
) slightly longer than hypandrium.
</paragraph>
<paragraph id="8B8BB8106501FFEEFF68F8F5FE3EF80A" blockId="16.[151,1437,422,2028]" pageId="16" pageNumber="267">
<emphasis id="B94064026501FFEEFF68F8F5FEEBF8BB" bold="true" box="[199,293,1860,1885]" pageId="16" pageNumber="267">Female.</emphasis>
Body length
<quantity id="4CCC15F56501FFEEFE69F8F4FDEDF8BA" box="[454,547,1861,1885]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="16" pageNumber="267" unit="mm" value="4.1">4.1 mm</quantity>
; wing length
<quantity id="4CCC15F56501FFEEFD6BF8F4FCEEF8BA" box="[708,800,1861,1885]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="16" pageNumber="267" unit="mm" value="3.8">3.8 mm</quantity>
. Frons slightly wider than anterior ocellus width. Face narrower than frons width. Wing slightly darker and wider (
<figureCitation id="130FA4956501FFEEFC92F8D9FC68F866" box="[829,934,1896,1921]" captionStart="FIGURE 12" captionStartId="18.[151,250,1493,1515]" captionTargetBox="[150,1436,230,1418]" captionTargetId="figure@18.[150,1436,230,1418]" captionTargetPageId="18" captionText="FIGURE 12. Macrostomus trifidus sp. nov., paratype ♀. A. Habitus, lateral view; B. apex of abdomen, lateral view; C. Tergite 8, 9+10 and cerci, dorsal view; D. Sternite 8, ventral view; E. genital fork and genital chamber, ventral view. Scale bars, figures A = 1 mm; B–E = 0.2 mm." httpUri="https://zenodo.org/record/2594959/files/figure.png" pageId="16" pageNumber="267">Fig. 12A</figureCitation>
). Tergite 8 (
<figureCitation id="130FA4956501FFEEFB80F8D9FB74F866" box="[1071,1210,1896,1921]" captionStart="FIGURE 12" captionStartId="18.[151,250,1493,1515]" captionTargetBox="[150,1436,230,1418]" captionTargetId="figure@18.[150,1436,230,1418]" captionTargetPageId="18" captionText="FIGURE 12. Macrostomus trifidus sp. nov., paratype ♀. A. Habitus, lateral view; B. apex of abdomen, lateral view; C. Tergite 8, 9+10 and cerci, dorsal view; D. Sternite 8, ventral view; E. genital fork and genital chamber, ventral view. Scale bars, figures A = 1 mm; B–E = 0.2 mm." httpUri="https://zenodo.org/record/2594959/files/figure.png" pageId="16" pageNumber="267">Figs 12B, C</figureCitation>
) with shallow basal and deeper distal sinus. Sternite 8 (
<figureCitation id="130FA4956501FFEEFD9FF83DFD0FF842" box="[560,705,1932,1957]" captionStart="FIGURE 12" captionStartId="18.[151,250,1493,1515]" captionTargetBox="[150,1436,230,1418]" captionTargetId="figure@18.[150,1436,230,1418]" captionTargetPageId="18" captionText="FIGURE 12. Macrostomus trifidus sp. nov., paratype ♀. A. Habitus, lateral view; B. apex of abdomen, lateral view; C. Tergite 8, 9+10 and cerci, dorsal view; D. Sternite 8, ventral view; E. genital fork and genital chamber, ventral view. Scale bars, figures A = 1 mm; B–E = 0.2 mm." httpUri="https://zenodo.org/record/2594959/files/figure.png" pageId="16" pageNumber="267">Figs 12B, D</figureCitation>
) with distinct protuberance ventrally; in ventral view with less sclerotized mid-longitudinal stripe from base to middle. Genital fork (
<figureCitation id="130FA4956501FFEEFC64F81EFBFCF82E" box="[971,1074,1967,1992]" captionStart="FIGURE 12" captionStartId="18.[151,250,1493,1515]" captionTargetBox="[150,1436,230,1418]" captionTargetId="figure@18.[150,1436,230,1418]" captionTargetPageId="18" captionText="FIGURE 12. Macrostomus trifidus sp. nov., paratype ♀. A. Habitus, lateral view; B. apex of abdomen, lateral view; C. Tergite 8, 9+10 and cerci, dorsal view; D. Sternite 8, ventral view; E. genital fork and genital chamber, ventral view. Scale bars, figures A = 1 mm; B–E = 0.2 mm." httpUri="https://zenodo.org/record/2594959/files/figure.png" pageId="16" pageNumber="267">Fig. 12E</figureCitation>
) thin, elongated, bifid at apex. Genital chamber membranous.
</paragraph>
<caption id="DF4BE8986500FFEFFF38F8B2FBB2F816" httpUri="https://zenodo.org/record/2594957/files/figure.png" pageId="17" pageNumber="268" startId="17.[151,250,1795,1817]" targetBox="[222,1364,200,1778]" targetPageId="17">
<paragraph id="8B8BB8106500FFEFFF38F8B2FBB2F816" blockId="17.[151,1436,1795,2032]" pageId="17" pageNumber="268">
<emphasis id="B94064026500FFEFFF38F8B2FED0F8FE" bold="true" box="[151,286,1795,1817]" pageId="17" pageNumber="268">FIGURE 11.</emphasis>
<taxonomicName id="4C34C3936500FFEFFE8BF8B5FE32F8FF" authority="Rafael &amp; Marques, 2019" authorityName="Rafael &amp; Marques" authorityYear="2019" box="[292,508,1795,1817]" class="Insecta" family="Empididae" genus="Macrostomus" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="268" phylum="Arthropoda" rank="species" species="trifidus" status="sp. nov.">
<emphasis id="B94064026500FFEFFE8BF8B5FE32F8FF" box="[292,508,1795,1817]" italics="true" pageId="17" pageNumber="268">Macrostomus trifidus</emphasis>
</taxonomicName>
<emphasis id="B94064026500FFEFFDADF8B5FD97F8FF" bold="true" box="[514,601,1796,1817]" pageId="17" pageNumber="268">
<taxonomicNameLabel id="A273D9796500FFEFFDADF8B5FD9BF8FF" box="[514,597,1796,1817]" pageId="17" pageNumber="268" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
holotype ♂.
<emphasis id="B94064026500FFEFFD72F8B2FD38F8FE" bold="true" box="[733,758,1795,1816]" pageId="17" pageNumber="268">A.</emphasis>
Habitus, lateral view;
<emphasis id="B94064026500FFEFFC74F8B5FC3CF8FF" bold="true" box="[987,1010,1796,1817]" pageId="17" pageNumber="268">B.</emphasis>
Apex of abdomen after macerated, lateral view (pieces with white outline);
<emphasis id="B94064026500FFEFFE46F893FDCFF8DE" bold="true" box="[489,513,1826,1848]" pageId="17" pageNumber="268">C.</emphasis>
Tergite 8, dorsal view;
<emphasis id="B94064026500FFEFFD5FF893FCC6F8D1" bold="true" box="[752,776,1826,1847]" pageId="17" pageNumber="268">D.</emphasis>
Tergite and sternite 8, lateral view;
<emphasis id="B94064026500FFEFFBDCF893FB44F8D1" bold="true" box="[1139,1162,1826,1847]" pageId="17" pageNumber="268">E.</emphasis>
Sternite 8, ventral view;
<emphasis id="B94064026500FFEFFA27F893FA54F8D1" bold="true" box="[1416,1434,1826,1847]" pageId="17" pageNumber="268">F.</emphasis>
Epandrium detached, lateral view;
<emphasis id="B94064026500FFEFFDA1F8F0FDEDF8B1" bold="true" box="[526,547,1857,1879]" pageId="17" pageNumber="268">G.</emphasis>
Epandrium detached, posterior view;
<emphasis id="B94064026500FFEFFC6BF8F0FC10F8B0" bold="true" box="[964,990,1857,1878]" pageId="17" pageNumber="268">H.</emphasis>
Terminalia, posterior view;
<emphasis id="B94064026500FFEFFABCF8F0FAEDF8B0" bold="true" box="[1299,1315,1857,1878]" pageId="17" pageNumber="268">I.</emphasis>
Terminalia (epandrium, hypandrium and phallus removed), right lateral view;
<emphasis id="B94064026500FFEFFCFBF8D1FCA8F893" bold="true" box="[852,870,1888,1909]" pageId="17" pageNumber="268">J.</emphasis>
Branchs of posterior cerci, posterolateral view (black arrows indicating the three branchs);
<emphasis id="B94064026500FFEFFDBBF8CEFDE0F872" bold="true" box="[532,558,1919,1940]" pageId="17" pageNumber="268">K.</emphasis>
Detail of cercal bridge of anterior cercus, ventral projection of cercus and dorsal lobe of hypoproct, anterior view;
<emphasis id="B94064026500FFEFFE6AF82CFE12F854" bold="true" box="[453,476,1949,1970]" pageId="17" pageNumber="268">L.</emphasis>
Posterior cercus, posterior view;
<emphasis id="B94064026500FFEFFCEFF82CFC91F854" bold="true" box="[832,863,1949,1970]" pageId="17" pageNumber="268">M.</emphasis>
Hypandrium, ejaculatory apodeme and phallus, lateral view;
<emphasis id="B94064026500FFEFFF7AF80DFF20F837" bold="true" box="[213,238,1980,2001]" pageId="17" pageNumber="268">N.</emphasis>
Hypandrium, posterior view;
<emphasis id="B94064026500FFEFFD8EF80AFDF5F837" bold="true" box="[545,571,1979,2001]" pageId="17" pageNumber="268">O.</emphasis>
Holotype labels. Scale bars, figures A = 1 mm; B–N = 0.2 mm. Abbreviations: cbrd, cercal bridge of anterior cercus; vpc, ventral projection of cercus; DHyL, Dorsal hypoproctal lobe;
</paragraph>
</caption>
<caption id="DF4BE8986503FFECFF38FA64FE66F9CE" httpUri="https://zenodo.org/record/2594959/files/figure.png" pageId="18" pageNumber="269" startId="18.[151,250,1493,1515]" targetBox="[150,1436,230,1418]" targetPageId="18">
<paragraph id="8B8BB8106503FFECFF38FA64FE66F9CE" blockId="18.[151,1436,1493,1576]" pageId="18" pageNumber="269">
<emphasis id="B94064026503FFECFF38FA64FED1FA0C" bold="true" box="[151,287,1493,1515]" pageId="18" pageNumber="269">FIGURE 12.</emphasis>
<taxonomicName id="4C34C3936503FFECFE8AFA67FE33FA0D" authority="Rafael &amp; Marques, 2019" authorityName="Rafael &amp; Marques" authorityYear="2019" box="[293,509,1493,1515]" class="Insecta" family="Empididae" genus="Macrostomus" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="269" phylum="Arthropoda" rank="species" species="trifidus" status="sp. nov.">
<emphasis id="B94064026503FFECFE8AFA67FE33FA0D" box="[293,509,1493,1515]" italics="true" pageId="18" pageNumber="269">Macrostomus trifidus</emphasis>
</taxonomicName>
<emphasis id="B94064026503FFECFDACFA64FD9DFA0C" bold="true" box="[515,595,1493,1514]" pageId="18" pageNumber="269">
<taxonomicNameLabel id="A273D9796503FFECFDACFA64FD9DFA0C" box="[515,595,1493,1514]" pageId="18" pageNumber="269" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, paratype ♀.
<emphasis id="B94064026503FFECFD72FA64FD38FA0C" bold="true" box="[733,758,1493,1514]" pageId="18" pageNumber="269">A.</emphasis>
Habitus, lateral view;
<emphasis id="B94064026503FFECFC74FA64FC3CFA0C" bold="true" box="[987,1010,1493,1514]" pageId="18" pageNumber="269">B.</emphasis>
apex of abdomen, lateral view;
<emphasis id="B94064026503FFECFA99FA64FA81FA0D" bold="true" box="[1334,1359,1493,1515]" pageId="18" pageNumber="269">C.</emphasis>
Tergite 8, 9+10 and cerci, dorsal view;
<emphasis id="B94064026503FFECFE7BFA45FE23F9EF" bold="true" box="[468,493,1524,1545]" pageId="18" pageNumber="269">D.</emphasis>
Sternite 8, ventral view;
<emphasis id="B94064026503FFECFD45FA45FCCFF9EF" bold="true" box="[746,769,1524,1545]" pageId="18" pageNumber="269">E.</emphasis>
genital fork and genital chamber, ventral view. Scale bars, figures A = 1 mm; B–E = 0.2 mm.
</paragraph>
</caption>
<paragraph id="8B8BB8106503FFECFF68F9E5FD59F98B" blockId="18.[151,1437,1620,2006]" box="[199,663,1620,1645]" pageId="18" pageNumber="269">
<emphasis id="B94064026503FFECFF68F9E5FE1EF98B" bold="true" box="[199,464,1620,1645]" pageId="18" pageNumber="269">Geographical records.</emphasis>
<collectingCountry id="F323F8806503FFECFE78F9E5FDE7F98B" box="[471,553,1620,1645]" name="Bolivia" pageId="18" pageNumber="269">Bolivia</collectingCountry>
(
<collectingRegion id="49F076F26503FFECFD97F9E4FD45F98B" box="[568,651,1621,1645]" country="Bolivia" name="La Paz" pageId="18" pageNumber="269">La Paz</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C32EEB9B6503FFECFF68F9C9FAFEF918" pageId="18" pageNumber="269" type="materials_examined">
<paragraph id="8B8BB8106503FFECFF68F9C9FBDEF93F" blockId="18.[151,1437,1620,2006]" pageId="18" pageNumber="269">
<emphasis id="B94064026503FFECFF68F9C9FD51F977" bold="true" box="[199,671,1656,1681]" pageId="18" pageNumber="269">
Type material examined.
<materialsCitation id="3B5CB24D6503FFECFDAEF9C9FD51F977" ID-GBIF-Occurrence="2013156518" box="[513,671,1656,1681]" collectingDate="22. iv." collectionCode="NHMUK" country="Bolivia" county="Rio" elevation="270" location="San Buenaventura" municipality="San Buenaventura" pageId="18" pageNumber="269" specimenCode="NHMUK 010864590" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="La Paz" typeStatus="holotype">
<typeStatus id="548F06B26503FFECFDAEF9C9FD51F977" box="[513,671,1656,1681]" pageId="18" pageNumber="269" type="holotype">HOLOTYPE</typeStatus>
</materialsCitation>
</emphasis>
<specimenCount id="9D3273996503FFECFD05F9C8FD0FF976" box="[682,705,1657,1680]" pageId="18" pageNumber="269" type="male">♂</specimenCount>
,
<typeStatus id="548F06B26503FFECFD7EF9C8FCFFF977" box="[721,817,1657,1681]" pageId="18" pageNumber="269" type="paratype">paratype</typeStatus>
<specimenCount id="9D3273996503FFECFC93F9C8FC9FF976" box="[828,849,1657,1680]" pageId="18" pageNumber="269" type="female">♀</specimenCount>
, on same pin: “
<collectingCountry id="F323F8806503FFECFBA1F9C8FB48F977" box="[1038,1158,1657,1681]" name="Bolivia" pageId="18" pageNumber="269">BOLIVIA</collectingCountry>
,
<collectingRegion id="49F076F26503FFECFB39F9C8FB27F976" box="[1174,1257,1657,1680]" country="Bolivia" name="La Paz" pageId="18" pageNumber="269">La Paz</collectingRegion>
,
<collectingCounty id="62EAC09C6503FFECFB57F9C9FAEEF977" box="[1272,1312,1656,1681]" pageId="18" pageNumber="269">Rio</collectingCounty>
<collectingRegion id="49F076F26503FFECFA84F9C9FAADF977" box="[1323,1379,1656,1681]" country="Bolivia" name="Beni" pageId="18" pageNumber="269">Beni</collectingRegion>
,
<collectingMunicipality id="6BEF226A6503FFECFADDF9C8FEF8F953" pageId="18" pageNumber="269">San Buenaventura</collectingMunicipality>
,
<quantity id="4CCC15F56503FFECFEEFF92CFE45F953" box="[320,395,1693,1717]" metricMagnitude="2" metricUnit="m" metricValue="2.7" pageId="18" pageNumber="269" unit="m" value="270.0">
<elevation id="00195F236503FFECFEEFF92CFE45F953" box="[320,395,1693,1717]" metricMagnitude="2" metricUnit="m" metricValue="2.7" pageId="18" pageNumber="269" unit="m" value="270.0">270 m</elevation>
</quantity>
”; “in copula,
<location id="8EEBEECB6503FFECFD8CF92CFD3DF953" LSID="urn:lsid:plazi:treatment:039D09066501FFECFF38FEACFC2EF830:8EEBEECB6503FFECFD8CF92CFD3DF953" box="[547,755,1693,1717]" country="Bolivia" county="Rio" municipality="San Buenaventura" name="San Buenaventura" pageId="18" pageNumber="269" stateProvince="La Paz">San Buenaventura</location>
,
<date id="FF8A9ED06503FFECFD52F92CFCF0F950" box="[765,830,1693,1718]" pageId="18" pageNumber="269">
<collectingDate id="EFCE67386503FFECFD52F92CFCF0F950" box="[765,830,1693,1718]" pageId="18" pageNumber="269">22.iv.</collectingDate>
</date>
[19]79”; “QR code:
<specimenCode id="DB92106B6503FFECFBB1F92FFADDF950" box="[1054,1299,1693,1718]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="18" pageNumber="269" type="Museum">NHMUK 010864590</specimenCode>
”; “
<typeStatus id="548F06B26503FFECFA9DF92CFA53F950" box="[1330,1437,1693,1718]" pageId="18" pageNumber="269" type="holotype">Holotype</typeStatus>
<specimenCount id="9D3273996503FFECFF38F970FF62F93E" box="[151,172,1729,1752]" pageId="18" pageNumber="269" type="male">♂</specimenCount>
<typeStatus id="548F06B26503FFECFF1CF970FEDBF93F" box="[179,277,1729,1753]" pageId="18" pageNumber="269" type="paratype">Paratype</typeStatus>
<specimenCount id="9D3273996503FFECFEB3F970FEFFF93E" box="[284,305,1729,1752]" pageId="18" pageNumber="269" type="female">♀</specimenCount>
<taxonomicName id="4C34C3936503FFECFE97F970FD31F93F" authority="Rafael &amp; Marques" authorityName="Rafael &amp; Marques" authorityYear="2019" box="[312,767,1728,1753]" class="Insecta" family="Empididae" genus="Macrostomus" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="269" phylum="Arthropoda" rank="species" species="trifidus">
<emphasis id="B94064026503FFECFE97F970FDE9F93F" box="[312,551,1729,1753]" italics="true" pageId="18" pageNumber="269">Macrostomus trifidus</emphasis>
Rafael &amp; Marques
</taxonomicName>
” (
<figureCitation id="130FA4956503FFECFCB6F971FCB3F93F" box="[793,893,1728,1753]" captionStart="FIGURE 11" captionStartId="17.[151,250,1795,1817]" captionTargetBox="[222,1364,200,1778]" captionTargetId="figure@17.[221,1364,193,1780]" captionTargetPageId="17" captionText="FIGURE 11. Macrostomus trifidus sp. nov., holotype ♂. A. Habitus, lateral view; B. Apex of abdomen after macerated, lateral view (pieces with white outline); C. Tergite 8, dorsal view; D. Tergite and sternite 8, lateral view; E. Sternite 8, ventral view; F. Epandrium detached, lateral view; G. Epandrium detached, posterior view; H. Terminalia, posterior view; I. Terminalia (epandrium, hypandrium and phallus removed), right lateral view; J. Branchs of posterior cerci, posterolateral view (black arrows indicating the three branchs); K. Detail of cercal bridge of anterior cercus, ventral projection of cercus and dorsal lobe of hypoproct, anterior view; L. Posterior cercus, posterior view; M. Hypandrium, ejaculatory apodeme and phallus, lateral view; N. Hypandrium, posterior view; O. Holotype labels. Scale bars, figures A = 1 mm; B–N = 0.2 mm. Abbreviations: cbrd, cercal bridge of anterior cercus; vpc, ventral projection of cercus; DHyL, Dorsal hypoproctal lobe;" httpUri="https://zenodo.org/record/2594957/files/figure.png" pageId="18" pageNumber="269">Fig. 11O</figureCitation>
) (NHMUK).
</paragraph>
<paragraph id="8B8BB8106503FFECFF68F954FAFEF918" blockId="18.[151,1437,1620,2006]" box="[199,1328,1765,1790]" pageId="18" pageNumber="269">
<emphasis id="B94064026503FFECFF68F954FE7CF918" bold="true" box="[199,434,1765,1790]" pageId="18" pageNumber="269">
<typeStatus id="548F06B26503FFECFF68F954FEFAF918" box="[199,308,1765,1790]" pageId="18" pageNumber="269" type="holotype">Holotype</typeStatus>
condition.
</emphasis>
Good, pinned, double mounted, abdomen dissected, in microvial with glycerin.
</paragraph>
</subSubSection>
<subSubSection id="C32EEB9B6503FFECFF68F8B9FE0CF8A3" pageId="18" pageNumber="269" type="etymology">
<paragraph id="8B8BB8106503FFECFF68F8B9FE0CF8A3" blockId="18.[151,1437,1620,2006]" pageId="18" pageNumber="269">
<emphasis id="B94064026503FFECFF68F8B9FE80F8C7" bold="true" box="[199,334,1800,1825]" pageId="18" pageNumber="269">Etymology.</emphasis>
From the Latin
<emphasis id="B94064026503FFECFDA9F8B8FDEFF8C6" box="[518,545,1801,1824]" italics="true" pageId="18" pageNumber="269">tri</emphasis>
= three and -
<emphasis id="B94064026503FFECFD17F8B8FD18F8C7" box="[696,726,1801,1825]" italics="true" pageId="18" pageNumber="269">fid</emphasis>
= suffix denoting division, which refers to the three digitiform branch of posterior cercus.
</paragraph>
</subSubSection>
<subSubSection id="C32EEB9B6503FFECFF68F8E1FC2EF830" pageId="18" pageNumber="269" type="discussion">
<paragraph id="8B8BB8106503FFECFF68F8E1FC2EF830" blockId="18.[151,1437,1620,2006]" pageId="18" pageNumber="269">
<emphasis id="B94064026503FFECFF68F8E1FEF5F88F" bold="true" box="[199,315,1872,1897]" pageId="18" pageNumber="269">Remarks.</emphasis>
Characters allowing a well-founded placement are presently not available for
<taxonomicName id="4C34C3936503FFECFB03F8E0FAEDF88F" authority="Rafael &amp; Marques, 2019" authorityName="Rafael &amp; Marques" authorityYear="2019" box="[1196,1315,1873,1897]" class="Insecta" family="Empididae" genus="Macrostomus" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="269" phylum="Arthropoda" rank="species" species="trifidus" status="sp. nov.">
<emphasis id="B94064026503FFECFB03F8E0FAEDF88F" box="[1196,1315,1873,1897]" italics="true" pageId="18" pageNumber="269">M. trifidus</emphasis>
</taxonomicName>
<emphasis id="B94064026503FFECFA85F8E1FA4AF88F" bold="true" box="[1322,1412,1872,1897]" pageId="18" pageNumber="269">
<taxonomicNameLabel id="A273D9796503FFECFA85F8E1FA4AF88F" box="[1322,1412,1872,1897]" pageId="18" pageNumber="269" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
It differs remarkably from other known
<taxonomicName id="4C34C3936503FFECFD94F8C7FD1CF86B" authorityName="in Rafael &amp; Cumming" authorityYear="2004" box="[571,722,1910,1933]" class="Insecta" family="Empididae" genus="Macrostomus" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="269" phylum="Arthropoda" rank="genus">
<emphasis id="B94064026503FFECFD94F8C7FD1CF86B" box="[571,722,1910,1933]" italics="true" pageId="18" pageNumber="269">Macrostomus</emphasis>
</taxonomicName>
species by the epandrium emitting mesially and porterodorsally a stout protuberance with several distal short setae; by the posterior cercus trifid with each branch digitiform and by the cercal bridge of anterior cercus V-shaped, fused medially and ventrally.
</paragraph>
</subSubSection>
</treatment>
</document>
Reference in a new issue View git blame Copy permalink