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<document id="46E2231F696C57911BE23A0B66E33ED7" ID-CLB-Dataset="263342" ID-DOI="10.11646/zootaxa.5286.1.1" ID-GBIF-Dataset="67e7a105-1f62-414a-83d9-f758039eb726" ID-ISSN="1175-5326" ID-Zenodo-Dep="7958937" ID-ZooBank="F9834684-24D3-4795-B5EB-77B451DF856D" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="diego" checkinTime="1684793492880" checkinUser="plazi" docAuthor="Shi, Hongliang &amp; Liang, Hongbin" docDate="2023" docId="03877623627FFFDE2DEFB62EFDAD5E50" docLanguage="en" docName="zootaxa.5286.1.1.pdf" docOrigin="Zootaxa 5286 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5286.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Parena (Bothynoptera) tesari" docType="treatment" docVersion="8" lastPageNumber="62" masterDocId="FFBE0E5B6246FFE32D78B27BFFE75D5A" masterDocTitle="Taxonomic revision of the genus Parena Motschulsky, 1860 (Coleoptera, Carabidae, Lebiini, Metallicina)" masterLastPageNumber="144" masterPageNumber="1" pageNumber="58" updateTime="1699506634024" updateUser="plazi">
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<mods:title id="2D048F1551630F20A6CC6054B053CB8E">Taxonomic revision of the genus Parena Motschulsky, 1860 (Coleoptera, Carabidae, Lebiini, Metallicina)</mods:title>
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<mods:namePart id="DDB397F1470ECE13A94D9E9D8ABE2145">Shi, Hongliang</mods:namePart>
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<mods:namePart id="4DACAA6C37B5518968260217569A67AC">Liang, Hongbin</mods:namePart>
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<mods:title id="EC83A046F5A9D423812099E7979A58DB">Zootaxa</mods:title>
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<mods:date id="F624FD1EC328DE6CB7FBBB334ED39B81">2023</mods:date>
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<mods:number id="0708CC830974CCD9210A67A4B8558CE9">2023-05-18</mods:number>
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<mods:number id="55E5C5A3B092B4F5375EE99421C3A861">5286</mods:number>
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<treatment id="03877623627FFFDE2DEFB62EFDAD5E50" ID-DOI="http://doi.org/10.5281/zenodo.7963982" ID-GBIF-Taxon="209628989" ID-Zenodo-Dep="7963982" LSID="urn:lsid:plazi:treatment:03877623627FFFDE2DEFB62EFDAD5E50" httpUri="http://treatment.plazi.org/id/03877623627FFFDE2DEFB62EFDAD5E50" lastPageId="61" lastPageNumber="62" pageId="57" pageNumber="58">
<subSubSection id="C33494BE627FFFDA2DEFB62EFD1A5935" box="[151,765,1108,1135]" pageId="57" pageNumber="58" type="nomenclature">
<paragraph id="8B91C735627FFFDA2DEFB62EFD1A5935" blockId="57.[151,1133,1108,1171]" box="[151,765,1108,1135]" pageId="57" pageNumber="58">
<emphasis id="B95A1B27627FFFDA2DEFB62EFD1A5935" bold="true" box="[151,765,1108,1135]" pageId="57" pageNumber="58">
[13]
<taxonomicName id="4C2EBCB6627FFFDA2DB7B62EFD1A5935" ID-CoL="4DKQQ" authority="(Jedlicka, 1951)" baseAuthorityName="Jedlicka" baseAuthorityYear="1951" box="[207,765,1108,1135]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="tesari" subGenus="Bothynoptera">
<emphasis id="B95A1B27627FFFDA2DB7B62EFEC05935" bold="true" box="[207,295,1109,1135]" italics="true" pageId="57" pageNumber="58">Parena</emphasis>
(
<emphasis id="B95A1B27627FFFDA2C40B62FFE3A5935" bold="true" box="[312,477,1108,1135]" italics="true" pageId="57" pageNumber="58">Bothynoptera</emphasis>
)
<emphasis id="B95A1B27627FFFDA2C96B62EFDD65935" bold="true" box="[494,561,1109,1135]" italics="true" pageId="57" pageNumber="58">tesari</emphasis>
(
<bibRefCitation id="EFBFBAC4627FFFDA2F38B62EFD125935" author="Jedlicka, A." box="[576,757,1109,1135]" pageId="57" pageNumber="58" pagination="59 - 60" refId="ref111384" refString="Jedlicka, A. (1951) Stukie O. Carabidech Palaearcticke Fauny. Etudes sur les Carabides de la faune palaearctique. Acta Musei Silesiae, (A) 1, 59 - 60." type="journal article" year="1951">Jedlička, 1951</bibRefCitation>
)
</taxonomicName>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C33494BE627FFFDA2DEFB602FB8A59C9" box="[151,1133,1145,1171]" pageId="57" pageNumber="58" type="description">
<paragraph id="8B91C735627FFFDA2DEFB602FB8A59C9" blockId="57.[151,1133,1108,1171]" box="[151,1133,1145,1171]" pageId="57" pageNumber="58">
Habitus:
<figureCitation id="1315DBB0627FFFDA2D85B602FE6C59C9" box="[253,395,1145,1171]" captionStart="FIGURE 32" captionStartId="59.[151,250,1549,1573]" captionTargetBox="[156,1430,184,1511]" captionTargetId="figure-19@59.[151,1436,181,1525]" captionTargetPageId="59" captionText="FIGURE 32. Habitus of Parena (Bothynoptera) tesari species group. A Holotype of Parena tesari (Jedlička). B Holotype of Parena kataevi Kirschenhofer [=Parena tesari (Jedlička)]. C P. tesari (Jedlička), male from Taiwan. D P. tesari (Jedlička), fe- male from Yunnan. E Holotype of Parena obscura Mateu. F P. obscura Mateu, male from Nepal. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7959003" httpUri="https://zenodo.org/record/7959003/files/figure.png" pageId="57" pageNumber="58">Figs 32A–D</figureCitation>
. Male genitalia:
<figureCitation id="1315DBB0627FFFDA2F3DB602FD7D59C9" box="[581,666,1145,1171]" captionStart="FIGURE 33" captionStartId="60.[151,250,1277,1301]" captionTargetBox="[162,1428,460,1246]" captionTargetId="figure-174@60.[152,1436,453,1253]" captionTargetPageId="60" captionText="FIGURE 33. Median lobe of male genitalia for Parena (Bothynoptera) tesari (Jedlička), right-lateral (r), ventral (v), left-lateral (l), and dorsal (d) views, Taiwan. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7959005" httpUri="https://zenodo.org/record/7959005/files/figure.png" pageId="57" pageNumber="58">Figs 33</figureCitation>
,
<figureCitation id="1315DBB0627FFFDA2FDEB602FD2259C9" box="[678,709,1145,1171]" captionStart="FIGURE 34" captionStartId="60.[151,250,1977,2001]" captionTargetBox="[160,1428,1372,1949]" captionTargetId="figure-208@60.[151,1437,1361,1954]" captionTargetPageId="60" captionText="FIGURE 34. Median lobe of male genitalia for Parena (Bothynoptera) tesari (Jedlička) from different localities, left-lateral (l) and dorsal (d) views: A Yunnan. B Taiwan. C Zhejiang. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7959007" httpUri="https://zenodo.org/record/7959007/files/figure.png" pageId="57" pageNumber="58">34</figureCitation>
. Gonocoxites of ovipositor:
<figureCitation id="1315DBB0627FFFDA297BB602FB8F59C9" box="[1027,1128,1145,1171]" captionStart="FIGURE 11" captionStartId="18.[151,250,150,174]" captionTargetId="figure-19@17.[155,1432,181,1971]" captionTargetPageId="17" captionText="FIGURE 11. Gonocoxites (left side) of ovipositor for Parena spp., ventral side (dorsal side for fig. Q). A–J subgenus Bothynoptera: A P. (Bothynoptera) heteronycha sp.n., Laos, paratype. B P. (Bothynoptera) dorsigera (Schaum), Shaanxi. C P. (Bothynoptera) kurosai Habu, Guangdong. D P. (Bothynoptera) tesari (Jedlička), Taiwan. E P. (Bothynoptera) taiwana Hua, Taiwan. F P. (Bothynoptera) tripunctata (Bates), Shaanxi. G P. (Bothynoptera) malaisei (Andrewes), Taiwan. H P. (Bothynoptera) quadrisignata Mateu, Yunnan. I P. (Bothynoptera) emarginata sp. n., holotype. J P. (Bothynoptera) triguttata sp. n., Sichuan, paratype. K–X subgenus Parena: K P. (Parena) bicolor Motschulsky, Sulawesi. L P. (Parena) rubripicta Andrewes, Yunnan. M P. (Parena) fasciata (Chaudoir), Taiwan. N P. (Parena) nigrolineata (Chaudoir), Taiwan. O P. (Parena) picipes sp. n., Borneo, paratype. P P. (Parena) andrewesi Jedlička, Philippine. Q P. (Parena) politissima (Chaudoir), Vanuatu. R P. (Parena) latecincta (Bates), Guizhou. S P. (Parena) circumdata Shibata, Taiwan. T P. (Parena) pendleburyi Andrewes, Borneo. U P. (Parena) fulva sp. n., Zimbabwe, paratype. V P. (Parena) madagascariensis (Alluaud), Madagascar. W P. (Parena) ruficornis sp.n., CentralAfrica, paratype. X P. (Parena) africana (Alluaud), Uganda. Y–γ subgenus Crossoglossa: Y P. (Crossoglossa) cavipennis (Bates), Taiwan. Z P. (Crossoglossa) laesipennis (Bates), Yunnan. α P. (Crossoglossa) testacea (Chaudoir), Zhejiang. β P. (Crossoglossa) sulawesiensis Kirschenhofer, Sulawesi. γ P. (Crossoglossa) sciakyi sp. n., holotype. Scale bar = 0.2mm." figureDoi="http://doi.org/10.5281/zenodo.7963954" httpUri="https://zenodo.org/record/7963954/files/figure.png" pageId="57" pageNumber="58">Fig. 11D</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33494BE627FFFDA2DEFB6C3FB5058D2" pageId="57" pageNumber="58" type="reference_group">
<paragraph id="8B91C735627FFFDA2DEFB6C3FC8A59B5" blockId="57.[151,1437,1208,1416]" pageId="57" pageNumber="58">
<bibRefCitation id="EFBFBAC4627FFFDA2DEFB6C3FEB6598A" author="Jedlicka, A." box="[151,337,1208,1232]" pageId="57" pageNumber="58" pagination="59 - 60" refId="ref111384" refString="Jedlicka, A. (1951) Stukie O. Carabidech Palaearcticke Fauny. Etudes sur les Carabides de la faune palaearctique. Acta Musei Silesiae, (A) 1, 59 - 60." type="journal article" year="1951">Jedlička, 1951: 60</bibRefCitation>
(original:
<taxonomicName id="4C2EBCB6627FFFDA2CC5B6C3FDA0598A" ID-CoL="8LR5S" authority="Schaum, 1863" box="[445,583,1208,1232]" class="Insecta" family="Carabidae" genus="Bothynoptera" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="B95A1B27627FFFDA2CC5B6C3FDA0598A" box="[445,583,1208,1232]" italics="true" pageId="57" pageNumber="58">Bothynoptera</emphasis>
</taxonomicName>
; type locality:
<collectingCountry id="F33987A5627FFFDA2F99B6C3FCC8598A" box="[737,815,1208,1232]" name="Taiwan" pageId="57" pageNumber="58">Taiwan</collectingCountry>
: Karenko;
<typeStatus id="54957997627FFFDA2EE6B6C3FC10598A" box="[926,1015,1208,1232]" pageId="57" pageNumber="58" type="holotype">holotype</typeStatus>
in NMPC);
<bibRefCitation id="EFBFBAC4627FFFDA290EB6C3FADA598A" author="Jedlicka, A." box="[1142,1341,1208,1232]" pageId="57" pageNumber="58" pagination="269 - 579" refId="ref111456" refString="Jedlicka, A. (1963) Monographie der Truncatipennen aus Ostasien, Lebiinae- Odacanthinae- Braehyninae (Coleptera, Carabidae). Entomologische Abhandlungen und Berichte aus dem Staatlichen Museum fuer Tierkunde in Dresden, 28, 269 - 579." type="journal article" year="1963">Jedlička, 1963: 446</bibRefCitation>
(
<taxonomicName id="4C2EBCB6627FFFDA2834B6C3FEEE59B4" ID-CoL="8LR5S" authority="Schaum, 1863" class="Insecta" family="Carabidae" genus="Bothynoptera" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="B95A1B27627FFFDA2834B6C3FEEE59B4" italics="true" pageId="57" pageNumber="58">Bothynoptera</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFBFBAC4627FFFDA2C67B6ACFE1559B5" author="Xie, W. P. &amp; Yu, P. Y." box="[287,498,1239,1263]" pageId="57" pageNumber="58" pagination="185 - 195" refId="ref113326" refString="Xie, W. P. &amp; Yu, P. Y. (1993) On the Chinese species of Parena Motschulsky (Coleoptera: Carabidae). Sinozoologia, 10, 185 - 195." type="journal article" year="1993">Xie &amp; Yu, 1993: 192</bibRefCitation>
(Zhejiang);
<bibRefCitation id="EFBFBAC4627FFFDA2F09B6ACFC8E59B5" author="Kirschenhofer, E." box="[625,873,1239,1263]" pageId="57" pageNumber="58" pagination="87 - 103" refId="ref111612" refString="Kirschenhofer, E. (2006) Neue Arten der Gattung Parena Motschulsky aus Japan, China Und Indonesien (Sulawesi) (Coleoptera: Carabidae). Mitteilungen des Internationalen Entomologischen Vereins E. V. Frankfurt, 31 (3 - 4), 87 - 103." type="journal article" year="2006">Kirschenhofer, 2006: 89</bibRefCitation>
.
</paragraph>
<paragraph id="8B91C735627FFFDA2DEFB68DFB2D5876" blockId="57.[151,1437,1208,1416]" pageId="57" pageNumber="58">
<taxonomicName id="4C2EBCB6627FFFDA2DEFB68DFE3A5857" ID-CoL="8KW9P" authority="Habu, 1969: 115" authorityName="Habu" authorityPageNumber="115" authorityYear="1969" box="[151,477,1269,1293]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="esakii">
<emphasis id="B95A1B27627FFFDA2DEFB68DFECE5857" bold="true" box="[151,297,1269,1293]" italics="true" pageId="57" pageNumber="58">Parena esakii</emphasis>
<materialsCitation id="3B46CD68627FFFDA2C48B68EFE3A5857" ID-GBIF-Occurrence="4115871399" box="[304,477,1269,1293]" country="Taiwan" location="Entomological Laboratory of Kyushu University" pageId="57" pageNumber="58" specimenCount="1" typeStatus="holotype">
<bibRefCitation id="EFBFBAC4627FFFDA2C48B68EFE3A5857" author="Habu, A." box="[304,477,1269,1293]" pageId="57" pageNumber="58" pagination="115 - 119" refId="ref110804" refString="Habu, A. (1969) A new species of Parena (Lebiini: Calleidina) from Formosa, with notes on the generic name Calleida (Coleoptera, Carabidae). Kontyu, 37, 115 - 119." type="journal article" year="1969">Habu, 1969: 115</bibRefCitation>
</materialsCitation>
</taxonomicName>
(type locality:
<collectingCountry id="F33987A5627FFFDA2F05B68EFD2C5857" box="[637,715,1269,1293]" name="Taiwan" pageId="57" pageNumber="58">Taiwan</collectingCountry>
: Taichu-shu;
<typeStatus id="54957997627FFFDA2E2EB68EFC485857" box="[854,943,1269,1293]" pageId="57" pageNumber="58" type="holotype">holotype</typeStatus>
in KUK:
<location id="8EF191EE627FFFDA2961B68EFED45876" LSID="urn:lsid:plazi:treatment:03877623627FFFDE2DEFB62EFDAD5E50:8EF191EE627FFFDA2961B68EFED45876" country="Taiwan" name="Entomological Laboratory of Kyushu University" pageId="57" pageNumber="58">Entomological Laboratory of Kyushu University</location>
,
<collectingCountry id="F33987A5627FFFDA2C45B76FFE9C5876" box="[317,379,1300,1324]" name="Japan" pageId="57" pageNumber="58">Japan</collectingCountry>
);
<bibRefCitation id="EFBFBAC4627FFFDA2CF5B76FFD845876" author="Xie, W. P. &amp; Yu, P. Y." box="[397,611,1300,1324]" pageId="57" pageNumber="58" pagination="185 - 195" refId="ref113326" refString="Xie, W. P. &amp; Yu, P. Y. (1993) On the Chinese species of Parena Motschulsky (Coleoptera: Carabidae). Sinozoologia, 10, 185 - 195." type="journal article" year="1993">Xie &amp; Yu, 1993: 194</bibRefCitation>
;
<bibRefCitation id="EFBFBAC4627FFFDA2F15B76FFC855876" author="Kirschenhofer, E." box="[621,866,1300,1324]" pageId="57" pageNumber="58" pagination="87 - 103" refId="ref111612" refString="Kirschenhofer, E. (2006) Neue Arten der Gattung Parena Motschulsky aus Japan, China Und Indonesien (Sulawesi) (Coleoptera: Carabidae). Mitteilungen des Internationalen Entomologischen Vereins E. V. Frankfurt, 31 (3 - 4), 87 - 103." type="journal article" year="2006">Kirschenhofer, 2006: 90</bibRefCitation>
(synonymized with
<taxonomicName id="4C2EBCB6627FFFDA294AB76EFB5B5876" ID-CoL="4DKQQ" baseAuthorityName="Jedlicka" baseAuthorityYear="1951" box="[1074,1212,1301,1324]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="tesari">
<emphasis id="B95A1B27627FFFDA294AB76EFB5B5876" box="[1074,1212,1301,1324]" italics="true" pageId="57" pageNumber="58">Parena tesari</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8B91C735627FFFDA2DEFB748FDFA5830" blockId="57.[151,1437,1208,1416]" pageId="57" pageNumber="58">
<taxonomicName id="4C2EBCB6627FFFDA2DEFB748FD925811" ID-CoL="4DKPW" authority="Kirschenhofer, 1996: 769" authorityName="Kirschenhofer" authorityPageNumber="769" authorityYear="1996" box="[151,629,1331,1355]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="nantouensis">
<emphasis id="B95A1B27627FFFDA2DEFB748FE8E5811" bold="true" box="[151,361,1331,1355]" italics="true" pageId="57" pageNumber="58">Parena nantouensis</emphasis>
<materialsCitation id="3B46CD68627FFFDA2C08B748FD925811" ID-GBIF-Occurrence="4115871433" box="[368,629,1331,1355]" country="Taiwan" location="Naturhistorisches Museum" pageId="57" pageNumber="58" specimenCount="1" stateProvince="Wien" typeStatus="holotype">Kirschenhofer, 1996: 769</materialsCitation>
</taxonomicName>
(type locality:
<collectingCountry id="F33987A5627FFFDA2E69B748FCB85811" box="[785,863,1331,1355]" name="Taiwan" pageId="57" pageNumber="58">Taiwan</collectingCountry>
: Nantou;
<typeStatus id="54957997627FFFDA2EBAB748FBFC5811" box="[962,1051,1331,1355]" pageId="57" pageNumber="58" type="holotype">holotype</typeStatus>
in NMW:
<location id="8EF191EE627FFFDA29F3B748FA7A5810" LSID="urn:lsid:plazi:treatment:03877623627FFFDE2DEFB62EFDAD5E50:8EF191EE627FFFDA29F3B748FA7A5810" box="[1163,1437,1331,1355]" country="Taiwan" name="Naturhistorisches Museum" pageId="57" pageNumber="58" stateProvince="Wien">Naturhistorisches Museum</location>
<collectingRegion id="49EA09D7627FFFDA2DBFB72AFEE75833" box="[199,256,1361,1385]" country="Austria" name="Wien" pageId="57" pageNumber="58">Wien</collectingRegion>
,
<collectingRegion id="49EA09D7627FFFDA2C71B72AFEB45833" box="[265,339,1361,1385]" country="Austria" name="Wien" pageId="57" pageNumber="58">Vienne</collectingRegion>
,
<collectingCountry id="F33987A5627FFFDA2C24B72AFE4C5833" box="[348,427,1361,1385]" name="Austria" pageId="57" pageNumber="58">Austria</collectingCountry>
).
<emphasis id="B95A1B27627FFFDA2CC4B72AFDFA5830" bold="true" box="[444,541,1361,1386]" pageId="57" pageNumber="58">Syn. nov.</emphasis>
</paragraph>
<paragraph id="8B91C735627FFFDA2DEFB70BFB5058D2" blockId="57.[151,1437,1208,1416]" box="[151,1207,1392,1416]" pageId="57" pageNumber="58">
<taxonomicName id="4C2EBCB6627FFFDA2DEFB70BFDD758D2" ID-CoL="4DKPJ" authority="Kirschenhofer, 2006: 97" authorityName="Kirschenhofer" authorityPageNumber="97" authorityYear="2006" box="[151,560,1392,1416]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="kataevi">
<emphasis id="B95A1B27627FFFDA2DEFB70BFED258D2" bold="true" box="[151,309,1392,1416]" italics="true" pageId="57" pageNumber="58">Parena kataevi</emphasis>
<materialsCitation id="3B46CD68627FFFDA2C42B70BFDD758D2" ID-GBIF-Occurrence="4115871375" box="[314,560,1392,1416]" location="Muyuping" pageId="57" pageNumber="58" specimenCount="1" stateProvince="Hubei" typeStatus="holotype">
<bibRefCitation id="EFBFBAC4627FFFDA2C42B70BFDD758D2" author="Kirschenhofer, E." box="[314,560,1392,1416]" pageId="57" pageNumber="58" pagination="87 - 103" refId="ref111612" refString="Kirschenhofer, E. (2006) Neue Arten der Gattung Parena Motschulsky aus Japan, China Und Indonesien (Sulawesi) (Coleoptera: Carabidae). Mitteilungen des Internationalen Entomologischen Vereins E. V. Frankfurt, 31 (3 - 4), 87 - 103." type="journal article" year="2006">Kirschenhofer, 2006: 97</bibRefCitation>
</materialsCitation>
</taxonomicName>
(type locality:
<collectingRegion id="49EA09D7627FFFDA2FB2B70BFCED58D2" box="[714,778,1392,1416]" country="China" name="Hubei" pageId="57" pageNumber="58">Hubei</collectingRegion>
,
<location id="8EF191EE627FFFDA2E6DB70BFC6558D2" LSID="urn:lsid:plazi:treatment:03877623627FFFDE2DEFB62EFDAD5E50:8EF191EE627FFFDA2E6DB70BFC6558D2" box="[789,898,1392,1416]" name="Muyuping" pageId="57" pageNumber="58" stateProvince="Hubei">Muyuping</location>
;
<typeStatus id="54957997627FFFDA2EF4B70BFC0258D2" box="[908,997,1392,1416]" pageId="57" pageNumber="58" type="holotype">holotype</typeStatus>
in CDW).
<emphasis id="B95A1B27627FFFDA292DB70BFB5058D2" bold="true" box="[1109,1207,1392,1416]" pageId="57" pageNumber="58">Syn. nov.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C33494BE627FFFDA2DEFB7CAFB035B25" pageId="57" pageNumber="58" type="materials_examined">
<paragraph id="8B91C735627FFFDA2DEFB7CAFD215B49" blockId="57.[151,1437,1456,2023]" pageId="57" pageNumber="58">
<emphasis id="B95A1B27627FFFDA2DEFB7CAFE515891" bold="true" box="[151,438,1457,1483]" pageId="57" pageNumber="58">Type material examined</emphasis>
.
<taxonomicName id="4C2EBCB6627FFFDA2CBCB7CBFCF65891" authority="Jedlicka" authorityName="Jedlicka" authorityYear="1951" box="[452,785,1456,1483]" class="Insecta" family="Carabidae" genus="Bothynoptera" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="tesari">
<emphasis id="B95A1B27627FFFDA2CBCB7CBFCF65891" bold="true" box="[452,785,1456,1483]" pageId="57" pageNumber="58">
<emphasis id="B95A1B27627FFFDA2CBCB7CBFD405891" bold="true" box="[452,679,1456,1483]" italics="true" pageId="57" pageNumber="58">Bothynoptera tesari</emphasis>
Jedlička
</emphasis>
</taxonomicName>
:
<emphasis id="B95A1B27627FFFDA2E66B7CAFC6C5891" bold="true" box="[798,907,1457,1483]" pageId="57" pageNumber="58">
<typeStatus id="54957997627FFFDA2E66B7CAFC6C5891" box="[798,907,1457,1483]" pageId="57" pageNumber="58" type="holotype">Holotype</typeStatus>
</emphasis>
(NMPC,
<figureCitation id="1315DBB0627FFFDA2E8FB7CAFB875891" box="[1015,1120,1457,1483]" captionStart="FIGURE 32" captionStartId="59.[151,250,1549,1573]" captionTargetBox="[156,1430,184,1511]" captionTargetId="figure-19@59.[151,1436,181,1525]" captionTargetPageId="59" captionText="FIGURE 32. Habitus of Parena (Bothynoptera) tesari species group. A Holotype of Parena tesari (Jedlička). B Holotype of Parena kataevi Kirschenhofer [=Parena tesari (Jedlička)]. C P. tesari (Jedlička), male from Taiwan. D P. tesari (Jedlička), fe- male from Yunnan. E Holotype of Parena obscura Mateu. F P. obscura Mateu, male from Nepal. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7959003" httpUri="https://zenodo.org/record/7959003/files/figure.png" pageId="57" pageNumber="58">Fig. 32A</figureCitation>
): female, body length =
<quantity id="4CD66AD0627FFFDA2813B7CAFF2258AA" metricMagnitude="-2" metricUnit="m" metricValue="1.05" pageId="57" pageNumber="58" unit="mm" value="10.5">10.5 mm</quantity>
, pin mounted, &quot;
<collectingCountry id="F33987A5627FFFDA2C00B7AEFE3958B5" box="[376,478,1493,1519]" name="Taiwan" pageId="57" pageNumber="58">Formosa</collectingCountry>
/ Karenko, -19. / VII 20-VIII 4. / T. Okuni&quot;, &quot;TYPUS&quot; [red label]; &quot;
<emphasis id="B95A1B27627FFFDA298BB7AEFEC95B49" italics="true" pageId="57" pageNumber="58">
<taxonomicName id="4C2EBCB6627FFFDA298BB7AEFA6C58B5" authority="Schaum, 1863" box="[1267,1419,1493,1519]" class="Insecta" family="Carabidae" genus="Bothynoptera" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="genus">Bothynoptera</taxonomicName>
/
<taxonomicName id="4C2EBCB6627FFFDA2DEFB781FF3B5B49" authority="Shi &amp; Liang, 2023" authorityName="Shi &amp; Liang" authorityYear="2023" box="[151,220,1530,1555]" class="Insecta" family="Carabidae" genus="Bothynoptera" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="tesari" status="sp. nov.">Tesari</taxonomicName>
<taxonomicNameLabel id="A269A65C627FFFDA2D9BB781FEF85B49" box="[227,287,1530,1555]" pageId="57" pageNumber="58" rank="species">sp. n.</taxonomicNameLabel>
/
</emphasis>
det. ING. JEDLIČKA&quot; [pink label].
</paragraph>
<paragraph id="8B91C735627FFFDA2DBFB466FB035B25" blockId="57.[151,1437,1456,2023]" pageId="57" pageNumber="58">
<emphasis id="B95A1B27627FFFDA2DBFB466FDD55B6D" bold="true" box="[199,562,1564,1591]" pageId="57" pageNumber="58">
<taxonomicName id="4C2EBCB6627FFFDA2DBFB466FDCA5B6D" authority="Kirschenhofer" authorityName="Kirschenhofer" authorityYear="2006" box="[199,557,1564,1591]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="kataevi">
<emphasis id="B95A1B27627FFFDA2DBFB466FE935B6D" bold="true" box="[199,372,1564,1591]" italics="true" pageId="57" pageNumber="58">Parena kataevi</emphasis>
Kirschenhofer
</taxonomicName>
:
</emphasis>
<materialsCitation id="3B46CD68627FFFDA2F41B466FB035B25" ID-GBIF-Occurrence="4115871349" collectorName="Jaroslav Turna" country="China" determinerName="Kirschenhofer" elevation="1300" latitude="31.45" location="Hubei" longitude="110.4" pageId="57" pageNumber="58" specimenCount="2" stateProvince="Hubei" typeStatus="holotype">
<emphasis id="B95A1B27627FFFDA2F41B466FD415B6D" bold="true" box="[569,678,1565,1591]" pageId="57" pageNumber="58">
<typeStatus id="54957997627FFFDA2F41B466FD415B6D" box="[569,678,1565,1591]" pageId="57" pageNumber="58" type="holotype">Holotype</typeStatus>
</emphasis>
(CDW,
<figureCitation id="1315DBB0627FFFDA2E7BB466FC8D5B6D" box="[771,874,1565,1591]" captionStart="FIGURE 32" captionStartId="59.[151,250,1549,1573]" captionTargetBox="[156,1430,184,1511]" captionTargetId="figure-19@59.[151,1436,181,1525]" captionTargetPageId="59" captionText="FIGURE 32. Habitus of Parena (Bothynoptera) tesari species group. A Holotype of Parena tesari (Jedlička). B Holotype of Parena kataevi Kirschenhofer [=Parena tesari (Jedlička)]. C P. tesari (Jedlička), male from Taiwan. D P. tesari (Jedlička), fe- male from Yunnan. E Holotype of Parena obscura Mateu. F P. obscura Mateu, male from Nepal. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7959003" httpUri="https://zenodo.org/record/7959003/files/figure.png" pageId="57" pageNumber="58">Fig. 32B</figureCitation>
): female, body length =
<quantity id="4CD66AD0627FFFDA2903B466FB065B6D" box="[1147,1249,1565,1591]" metricMagnitude="-2" metricUnit="m" metricValue="1.02" pageId="57" pageNumber="58" unit="mm" value="10.2">10.2 mm</quantity>
, board mounted, &quot;
<collectingCountry id="F33987A5627FFFDA2DE7B43AFF0F5B01" box="[159,232,1601,1627]" name="China" pageId="57" pageNumber="58">China</collectingCountry>
, W
<collectingRegion id="49EA09D7627FFFDA2C6CB43AFEBC5B01" box="[276,347,1601,1627]" country="China" name="Hubei" pageId="57" pageNumber="58">Hubei</collectingRegion>
, 20.-21.VI. / MUYUPING S. env. /
<geoCoordinate id="EE1AA1F2627FFFDA2F88B43AFC7A5B01" box="[752,925,1601,1628]" degrees="110.4" direction="north" orientation="latitude" pageId="57" pageNumber="58" precision="5555" value="110.4">31.45N 110.4E</geoCoordinate>
, ~
<quantity id="4CD66AD0627FFFDA2ECEB43AFBEB5B06" box="[950,1036,1601,1628]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="57" pageNumber="58" unit="m" value="1300.0">
<elevation id="00032006627FFFDA2ECEB43AFBEB5B06" box="[950,1036,1601,1628]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="57" pageNumber="58" unit="m" value="1300.0">1300 m</elevation>
</quantity>
/
<collectorName id="26DBA2E3627FFFDA295AB43AFB205B06" box="[1058,1223,1601,1628]" pageId="57" pageNumber="58">Jaroslav Turna</collectorName>
leg. 2003&quot;, &quot;Holotypus /
<taxonomicName id="4C2EBCB6627FFFDA2D92B41DFED15BDA" box="[234,310,1638,1664]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="genus">Parena</taxonomicName>
/ kataevi
<taxonomicNameLabel id="A269A65C627FFFDA2CDBB41DFE385BDA" box="[419,479,1638,1664]" pageId="57" pageNumber="58" rank="species">sp. n.</taxonomicNameLabel>
/ det.
<determinerName id="7E4D2BF1627FFFDA2F5DB41EFD225B25" box="[549,709,1637,1663]" pageId="57" pageNumber="58">Kirschenhofer</determinerName>
2004&quot; [red label], &quot;COLL.WRASE / BERLIN&quot;.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33494BE627FFFD92DBFB4F2FB8E5DA2" lastPageId="58" lastPageNumber="59" pageId="57" pageNumber="58" type="discussion">
<paragraph id="8B91C735627FFFDA2DBFB4F2FDBD5BB1" blockId="57.[151,1437,1456,2023]" pageId="57" pageNumber="58">
<emphasis id="B95A1B27627FFFDA2DBFB4F2FDCB5BF9" bold="true" box="[199,556,1673,1699]" pageId="57" pageNumber="58">Notes on types and synonyms</emphasis>
.
<taxonomicName id="4C2EBCB6627FFFDA2F44B4F3FC685BF9" authority="Jedlicka" authorityName="Jedlicka" authorityYear="1951" box="[572,911,1672,1699]" class="Insecta" family="Carabidae" genus="Bothynoptera" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="tesari">
<emphasis id="B95A1B27627FFFDA2F44B4F3FC685BF9" bold="true" box="[572,911,1672,1699]" pageId="57" pageNumber="58">
<emphasis id="B95A1B27627FFFDA2F44B4F3FCC55BF9" bold="true" box="[572,802,1672,1699]" italics="true" pageId="57" pageNumber="58">Bothynoptera tesari</emphasis>
Jedlička
</emphasis>
</taxonomicName>
: This species was originally described from a single specimen from Karenko. The female we examined in the collection of NMPC in accord with the original description is undoubtedly the
<typeStatus id="54957997627FFFDA2C89B4AAFDB05BB1" box="[497,599,1745,1771]" pageId="57" pageNumber="58" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="8B91C735627FFFDA2DBFB48EFC875A69" blockId="57.[151,1437,1456,2023]" pageId="57" pageNumber="58">
<taxonomicName id="4C2EBCB6627FFFDA2DBFB48EFE545A55" authority="Habu" authorityName="Habu" authorityYear="1969" box="[199,435,1780,1807]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="esakii">
<emphasis id="B95A1B27627FFFDA2DBFB48EFE545A55" bold="true" box="[199,435,1780,1807]" pageId="57" pageNumber="58">
<emphasis id="B95A1B27627FFFDA2DBFB48EFE805A55" bold="true" box="[199,359,1780,1807]" italics="true" pageId="57" pageNumber="58">Parena esakii</emphasis>
Habu
</emphasis>
</taxonomicName>
:
<bibRefCitation id="EFBFBAC4627FFFDA2CBCB48EFD5E5A55" author="Kirschenhofer, E." box="[452,697,1781,1807]" pageId="57" pageNumber="58" pagination="87 - 103" refId="ref111612" refString="Kirschenhofer, E. (2006) Neue Arten der Gattung Parena Motschulsky aus Japan, China Und Indonesien (Sulawesi) (Coleoptera: Carabidae). Mitteilungen des Internationalen Entomologischen Vereins E. V. Frankfurt, 31 (3 - 4), 87 - 103." type="journal article" year="2006">Kirschenhofer (2006)</bibRefCitation>
firstly proposed it is a synonym of
<taxonomicName id="4C2EBCB6627FFFDA2920B48DFB515A55" authorityName="Jedlicka" authorityYear="1951" box="[1112,1206,1782,1807]" class="Insecta" family="Carabidae" genus="Bothynoptera" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="tesari">
<emphasis id="B95A1B27627FFFDA2920B48DFB515A55" box="[1112,1206,1782,1807]" italics="true" pageId="57" pageNumber="58">P. tesari</emphasis>
</taxonomicName>
. We agree with this treatment according to the original description and illustrations.
</paragraph>
<paragraph id="8B91C735627FFFDA2DBFB546FB0C5ABD" blockId="57.[151,1437,1456,2023]" pageId="57" pageNumber="58">
<taxonomicName id="4C2EBCB6627FFFDA2DBFB546FD845A0D" authority="Kirschenhofer" authorityName="Kirschenhofer" authorityYear="1996" box="[199,611,1853,1879]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="nantouensis">
<emphasis id="B95A1B27627FFFDA2DBFB546FD845A0D" bold="true" box="[199,611,1853,1879]" pageId="57" pageNumber="58">
<emphasis id="B95A1B27627FFFDA2DBFB546FE495A0D" bold="true" box="[199,430,1853,1879]" italics="true" pageId="57" pageNumber="58">Parena nantouensis</emphasis>
Kirschenhofer
</emphasis>
</taxonomicName>
: We did not examine the type of this species. However, from the description and illustrations in the original description (Kirschenhofer, 1996), this species is almost identical to
<taxonomicName id="4C2EBCB6627FFFDA2847B519FA7B5A21" authorityName="Jedlicka" authorityYear="1951" box="[1343,1436,1890,1915]" class="Insecta" family="Carabidae" genus="Bothynoptera" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="tesari">
<emphasis id="B95A1B27627FFFDA2847B519FA7B5A21" box="[1343,1436,1890,1915]" italics="true" pageId="57" pageNumber="58">P. tesari</emphasis>
</taxonomicName>
except for its size (slightly smaller) and interval 3 with only two setigerous pores. We examined
<specimenCount id="9D280CBC627FFFDA29B2B5FEFA845AC5" box="[1226,1379,1925,1951]" pageId="57" pageNumber="58" type="generic">one specimen</specimenCount>
of
<taxonomicName id="4C2EBCB6627FFFDA28F0B5FDFF305A99" authorityName="Jedlicka" authorityYear="1951" class="Insecta" family="Carabidae" genus="Bothynoptera" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="tesari">
<emphasis id="B95A1B27627FFFDA28F0B5FDFF305A99" italics="true" pageId="57" pageNumber="58">P. tesari</emphasis>
</taxonomicName>
from
<collectingCountry id="F33987A5627FFFDA2C65B5D2FE885A99" box="[285,367,1961,1987]" name="Taiwan" pageId="57" pageNumber="58">Taiwan</collectingCountry>
with two setigerous pores (the middle one missing) on the right elytron. Thus, we believe that the
<typeStatus id="54957997627FFFDA2DB8B5B6FEC55ABD" box="[192,290,1997,2023]" pageId="57" pageNumber="58" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C2EBCB6627FFFDA2C30B5B5FE0E5ABD" authorityName="Kirschenhofer" authorityYear="1996" box="[328,489,1998,2023]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="nantouensis">
<emphasis id="B95A1B27627FFFDA2C30B5B5FE0E5ABD" box="[328,489,1998,2023]" italics="true" pageId="57" pageNumber="58">P. nantouensis</emphasis>
</taxonomicName>
is just an aberrant individual but not representing a separate species.
</paragraph>
<paragraph id="8B91C735627CFFD92DBFB2ECFB8E5DA2" blockId="58.[151,1437,150,2041]" pageId="58" pageNumber="59">
<taxonomicName id="4C2EBCB6627CFFD92DBFB2ECFDC15DEA" authority="Kirschenhofer" authorityName="Kirschenhofer" authorityYear="2006" box="[199,550,150,177]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="kataevi">
<emphasis id="B95A1B27627CFFD92DBFB2ECFDC15DEA" bold="true" box="[199,550,150,177]" pageId="58" pageNumber="59">
<emphasis id="B95A1B27627CFFD92DBFB2ECFE945DEB" bold="true" box="[199,371,150,177]" italics="true" pageId="58" pageNumber="59">Parena kataevi</emphasis>
Kirschenhofer
</emphasis>
</taxonomicName>
: This species was described from
<collectingRegion id="49EA09D7627CFFD92ED9B2ECFC0F5DEB" box="[929,1000,151,177]" country="China" name="Hubei" pageId="58" pageNumber="59">Hubei</collectingRegion>
, Shennongjia. We examined specimens from
<collectingRegion id="49EA09D7627CFFD92DACB2C0FEFF5D8F" box="[212,280,187,213]" country="China" name="Hubei" pageId="58" pageNumber="59">Hubei</collectingRegion>
and many other localities in central
<collectingCountry id="F33987A5627CFFD92FD4B2C0FD085D8F" box="[684,751,187,213]" name="China" pageId="58" pageNumber="59">China</collectingCountry>
that are in accord with it. Their male genitalia are identical to that of
<taxonomicName id="4C2EBCB6627CFFD92D9EB2A4FEA55DA2" authorityName="Jedlicka" authorityYear="1951" box="[230,322,223,248]" class="Insecta" family="Carabidae" genus="Bothynoptera" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="tesari">
<emphasis id="B95A1B27627CFFD92D9EB2A4FEA55DA2" box="[230,322,223,248]" italics="true" pageId="58" pageNumber="59">P. tesari</emphasis>
</taxonomicName>
from
<collectingCountry id="F33987A5627CFFD92CFEB2A4FE3B5DA3" box="[390,476,223,249]" name="Taiwan" pageId="58" pageNumber="59">Taiwan</collectingCountry>
. Thus, we treat
<taxonomicName id="4C2EBCB6627CFFD92FF4B2A4FD1F5DA3" authorityName="Kirschenhofer" authorityYear="2006" box="[652,760,223,249]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="kataevi">
<emphasis id="B95A1B27627CFFD92FF4B2A4FD1F5DA3" box="[652,760,223,249]" italics="true" pageId="58" pageNumber="59">P. kataevi</emphasis>
</taxonomicName>
as a junior synonym of
<emphasis id="B95A1B27627CFFD9297FB2A4FB8E5DA2" box="[1031,1129,223,248]" italics="true" pageId="58" pageNumber="59">
<taxonomicName id="4C2EBCB6627CFFD9297FB2A4FB825DA2" authorityName="Jedlicka" authorityYear="1951" box="[1031,1125,223,248]" class="Insecta" family="Carabidae" genus="Bothynoptera" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="tesari">P. tesari</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C33494BE627CFFD92DBFB378FE105A30" pageId="58" pageNumber="59" type="materials_examined">
<paragraph id="8B91C735627CFFD92DBFB378FE105A30" blockId="58.[151,1437,150,2041]" pageId="58" pageNumber="59">
<emphasis id="B95A1B27627CFFD92DBFB378FDFB5C46" bold="true" box="[199,540,258,285]" pageId="58" pageNumber="59">Non-type material examined</emphasis>
.
<emphasis id="B95A1B27627CFFD92F52B379FD945C46" bold="true" box="[554,627,258,284]" pageId="58" pageNumber="59">
<collectingCountry id="F33987A5627CFFD92F52B379FD945C46" box="[554,627,258,284]" name="China" pageId="58" pageNumber="59">China</collectingCountry>
</emphasis>
:
<specimenCount id="9D280CBC627CFFD92FFAB378FD3F5C47" box="[642,728,259,285]" pageId="58" pageNumber="59" type="male">2 males</specimenCount>
(HBUM), &quot;
<collectingRegion id="49EA09D7627CFFD92E24B378FC4F5C47" box="[860,936,259,285]" country="China" name="Anhui" pageId="58" pageNumber="59">Anhui</collectingRegion>
, Yuexi, Yaoluoping,
<date id="FF90E1F5627CFFD929E9B378FA955C47" box="[1169,1394,259,285]" pageId="58" pageNumber="59" value="2007-07-30" valueMax="2007-08-04" valueMin="2007-07-30">2007.VII.30- VIII.4</date>
, Ba Yibin, Lang Juntong, Wang Fengyan lgt.&quot;.
<specimenCount id="9D280CBC627CFFD92F06B35CFD075C1B" box="[638,736,295,321]" pageId="58" pageNumber="59" type="female">1 female</specimenCount>
(HBUM), &quot;
<collectingRegion id="49EA09D7627CFFD92E1EB35CFC555C1B" box="[870,946,295,321]" country="China" name="Anhui" pageId="58" pageNumber="59">Anhui</collectingRegion>
, Yuexi, Yaoluoping nat. resv.,
<date id="FF90E1F5627CFFD92872B35CFF525C3F" pageId="58" pageNumber="59" value="2007-07-17" valueMax="2007-07-21" valueMin="2007-07-17">2007.VII.17- 21</date>
, Ba Yibin, Lang Juntong, Wang Fengyan lgt.&quot;. 1 ex (IZAS), &quot;Tiemushan,
<date id="FF90E1F5627CFFD92E9CB330FB8E5C3F" box="[996,1129,331,357]" pageId="58" pageNumber="59" value="1927-06-20">1927.VI.20</date>
&quot;. 1 ex (IZAS), &quot;Tiemushan,
<date id="FF90E1F5627CFFD92DEFB314FEFB5CD3" box="[151,284,367,393]" pageId="58" pageNumber="59" value="1936-06-19">1936.VI.19</date>
&quot;. 1 ex (IZAS), &quot;Tiemushan,
<date id="FF90E1F5627CFFD92F1BB315FD1B5CD3" box="[611,764,366,393]" pageId="58" pageNumber="59" value="1935-08-18">1935.VIII.18</date>
&quot;. 1 ex (IZAS), &quot;T’ienmu Shan,
<date id="FF90E1F5627CFFD92910B314FB145CD3" box="[1128,1267,367,393]" pageId="58" pageNumber="59" value="1936-07-24">1936.VII.24</date>
, O. Piel coll.&quot;. 1 ex (IZAS), &quot;T’ienmu Shan,
<date id="FF90E1F5627CFFD92C91B3E8FD8C5CF7" box="[489,619,403,429]" pageId="58" pageNumber="59" value="1936-06-16">1936.VI.16</date>
, O. Piel coll.&quot;. 1 ex (IZAS), &quot;T’ienmu Shan,
<date id="FF90E1F5627CFFD92913B3E8FB115CF7" box="[1131,1270,403,429]" pageId="58" pageNumber="59" value="1936-07-20">1936.VII.20</date>
, O. Piel coll.&quot;. 1 ex (IZAS), &quot;Tianmushan,
<date id="FF90E1F5627CFFD92CAAB3CCFDAD5C8B" box="[466,586,439,465]" pageId="58" pageNumber="59" value="1947-09-07">1947.IX.7</date>
&quot;. 1 ex (IZAS), &quot;
<collectingRegion id="49EA09D7627CFFD92F85B3CCFC805C8B" box="[765,871,439,465]" country="China" name="Zhejiang" pageId="58" pageNumber="59">Zhejiang</collectingRegion>
, Tianmushan,
<date id="FF90E1F5627CFFD92973B3CDFB985C8B" box="[1035,1151,438,465]" pageId="58" pageNumber="59" value="1981-09-05">1981.IX.5</date>
, Zhang Baolin lgt.&quot;. 5 ex (IZAS), &quot;
<collectingRegion id="49EA09D7627CFFD92D87B3A0FE8E5CAF" box="[255,361,475,501]" country="China" name="Zhejiang" pageId="58" pageNumber="59">Zhejiang</collectingRegion>
, West Tianmushan. Traps.
<date id="FF90E1F5627CFFD92FE2B3A0FCEA5CAF" box="[666,781,475,501]" pageId="58" pageNumber="59" value="2018-07-16">2018.7.16</date>
. Li Chao,
<quantity id="4CD66AD0627CFFD92EFDB3A0FC075CAF" box="[901,992,475,501]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="58" pageNumber="59" unit="m" value="1400.0">1400 m</quantity>
&quot;. 1 ex (IZAS), &quot;
<collectingRegion id="49EA09D7627CFFD929EEB3A0FAE75CAF" box="[1174,1280,475,501]" country="China" name="Zhejiang" pageId="58" pageNumber="59">Zhejiang</collectingRegion>
, Tianmushan, 1993.VI, Ma Jufa lgt.&quot;. 2 ex (IZAS), &quot;Zehjiang, Tianmushan, 1999.VII&quot;. 1 ex (IZAS), &quot;
<collectingRegion id="49EA09D7627CFFD929EBB384FB1A5F43" box="[1171,1277,511,537]" country="China" name="Zhejiang" pageId="58" pageNumber="59">Zhejiang</collectingRegion>
, Tianmushan,
<date id="FF90E1F5627CFFD92DEFB058FEC55F67" box="[151,290,547,573]" pageId="58" pageNumber="59" value="1973-07-22">1973.VII.22</date>
, Yu Peiyu lgt.&quot;.
<specimenCount id="9D280CBC627CFFD92CA6B058FDD15F67" box="[478,566,547,573]" pageId="58" pageNumber="59" type="male">2 males</specimenCount>
(IZAS), &quot;
<collectingRegion id="49EA09D7627CFFD92FDFB058FCF65F67" box="[679,785,547,573]" country="China" name="Zhejiang" pageId="58" pageNumber="59">Zhejiang</collectingRegion>
, Lin'an W. Tianmushan, 2018.VI, Huang Hao leg.&quot; &lt;
<figureCitation id="1315DBB0627CFFD92808B058FF2E5F3B" captionStart="FIGURE 34" captionStartId="60.[151,250,1977,2001]" captionTargetBox="[160,1428,1372,1949]" captionTargetId="figure-208@60.[151,1437,1361,1954]" captionTargetPageId="60" captionText="FIGURE 34. Median lobe of male genitalia for Parena (Bothynoptera) tesari (Jedlička) from different localities, left-lateral (l) and dorsal (d) views: A Yunnan. B Taiwan. C Zhejiang. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7959007" httpUri="https://zenodo.org/record/7959007/files/figure.png" pageId="58" pageNumber="59">Fig. 34C</figureCitation>
&gt;. 1 ex (IZAS), &quot;
<collectingRegion id="49EA09D7627CFFD92C07B03CFE315F3B" box="[383,470,583,609]" country="China" name="Jiangxi" pageId="58" pageNumber="59">Jiangxi</collectingRegion>
, Wugongshan, forestry station, on vegetation,
<geoCoordinate id="EE1AA1F2627CFFD92EA4B03DFBBC5F3B" box="[988,1115,582,609]" degrees="27.44591" direction="north" orientation="latitude" pageId="58" pageNumber="59" precision="1" value="27.44591">N27.44591</geoCoordinate>
,
<geoCoordinate id="EE1AA1F2627CFFD9291FB03CFB165F3B" box="[1127,1265,583,609]" degrees="114.18827" direction="east" orientation="longitude" pageId="58" pageNumber="59" precision="1" value="114.18827">E114.18827</geoCoordinate>
,
<quantity id="4CD66AD0627CFFD92983B03CFAB55F3B" box="[1275,1362,583,609]" metricMagnitude="3" metricUnit="m" metricValue="1.22" pageId="58" pageNumber="59" unit="m" value="1220.0">1220 m</quantity>
,
<date id="FF90E1F5627CFFD92825B03CFF3D5FDE" pageId="58" pageNumber="59" value="2006-06-29">2006. VI.29</date>
, Liu Ye, Liang Hongbin, Sota T. Lgt.&quot;. 1 ex (IZAS), &quot;
<collectingRegion id="49EA09D7627CFFD92E51B011FC935FDE" box="[809,884,618,644]" country="China" name="Hubei" pageId="58" pageNumber="59">Hubei</collectingRegion>
, Xingshan, Longmenhe, light trap,
<quantity id="4CD66AD0627CFFD92984B011FAB55FDF" box="[1276,1362,618,645]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="58" pageNumber="59" unit="m" value="1200.0">1200 m</quantity>
,
<date id="FF90E1F5627CFFD92825B011FF3D5FF2" pageId="58" pageNumber="59" value="1993-06-14">1993. VI.14</date>
, Li Hongxing lgt.&quot;. 1 ex (HBUM), &quot;
<collectingRegion id="49EA09D7627CFFD92F0FB0F5FD255FF2" box="[631,706,654,680]" country="China" name="Hubei" pageId="58" pageNumber="59">Hubei</collectingRegion>
, Shenongjia, Muyu,
<date id="FF90E1F5627CFFD92ED5B0F5FBC55FF2" box="[941,1058,654,680]" pageId="58" pageNumber="59" value="2004-09-03">2004.IX.3</date>
, Zhang Zhisheng, Chen Huiming lgt.&quot;. 1 ex (IZAS), &quot;
<collectingRegion id="49EA09D7627CFFD92C05B0C9FE235F96" box="[381,452,690,716]" country="China" name="Hubei" pageId="58" pageNumber="59">Hubei</collectingRegion>
, Zhuxi, Quanxi,
<geoCoordinate id="EE1AA1F2627CFFD92FFEB0C9FD125F96" box="[646,757,690,716]" degrees="32.0686" direction="north" orientation="latitude" pageId="58" pageNumber="59" precision="5" value="32.0686">N32.0686</geoCoordinate>
<geoCoordinate id="EE1AA1F2627CFFD92F86B0C9FC9C5F96" box="[766,891,690,716]" degrees="109.6614" direction="east" orientation="longitude" pageId="58" pageNumber="59" precision="5" value="109.6614">E109.6614</geoCoordinate>
,
<quantity id="4CD66AD0627CFFD92EFFB0C9FC355F97" box="[903,978,690,717]" metricMagnitude="2" metricUnit="m" metricValue="8.559999999999999" pageId="58" pageNumber="59" unit="m" value="856.0">856 m</quantity>
, Guo PL. Lgt.
<date id="FF90E1F5627CFFD92907B0C9FB175F96" box="[1151,1264,690,716]" pageId="58" pageNumber="59" value="2017-07-13">2017.7.13</date>
&quot;. 1 ex (IZAS), &quot;
<collectingRegion id="49EA09D7627CFFD92DDBB0ADFF0D5FAA" box="[163,234,726,752]" country="China" name="Hubei" pageId="58" pageNumber="59">Hubei</collectingRegion>
, Zhuxi, Quanxi,
<geoCoordinate id="EE1AA1F2627CFFD92CD7B0ADFDFA5FAA" box="[431,541,726,752]" degrees="32.0911" direction="north" orientation="latitude" pageId="58" pageNumber="59" precision="5" value="32.0911">N32.0911</geoCoordinate>
<geoCoordinate id="EE1AA1F2627CFFD92F5FB0ADFD435FAA" box="[551,676,726,752]" degrees="109.6733" direction="east" orientation="longitude" pageId="58" pageNumber="59" precision="5" value="109.6733">E109.6733</geoCoordinate>
,
<quantity id="4CD66AD0627CFFD92FCAB0ADFD195FAB" box="[690,766,726,753]" metricMagnitude="2" metricUnit="m" metricValue="7.9" pageId="58" pageNumber="59" unit="m" value="790.0">790 m</quantity>
, Xia ZZ. Lgt.
<date id="FF90E1F5627CFFD92ED2B0ADFBFD5FAA" box="[938,1050,726,752]" pageId="58" pageNumber="59" value="2017-07-13">2017.7.13</date>
&quot;. 4 ex (IZAS), &quot;
<collectingRegion id="49EA09D7627CFFD92998B0ADFAC05FAA" box="[1248,1319,726,752]" country="China" name="Hubei" pageId="58" pageNumber="59">Hubei</collectingRegion>
, Xuan-en, Shadaogou, Longtan station,
<geoCoordinate id="EE1AA1F2627CFFD92C91B081FDA35E4E" box="[489,580,762,788]" degrees="29.6998" direction="north" orientation="latitude" pageId="58" pageNumber="59" precision="5" value="29.6998">29.6998</geoCoordinate>
<geoCoordinate id="EE1AA1F2627CFFD92F36B081FD5E5E4F" box="[590,697,762,789]" degrees="109.6511" direction="east" orientation="longitude" pageId="58" pageNumber="59" precision="5" value="109.6511">109.6511</geoCoordinate>
,
<quantity id="4CD66AD0627CFFD92FB0B081FCF35E4F" box="[712,788,762,789]" metricMagnitude="2" metricUnit="m" metricValue="6.31" pageId="58" pageNumber="59" unit="m" value="631.0">631 m</quantity>
,
<date id="FF90E1F5627CFFD92E5BB081FC735E4E" box="[803,916,762,788]" pageId="58" pageNumber="59" value="2016-04-30">2016.4.30</date>
D Liang HB., Zhao KD. lgt.&quot;. 1 ex (IZAS), &quot;
<collectingRegion id="49EA09D7627CFFD92DDBB164FF175E63" box="[163,240,799,825]" country="China" name="Hunan" pageId="58" pageNumber="59">Hunan</collectingRegion>
, Sangzhi, Tianpingshan,
<quantity id="4CD66AD0627CFFD92F7FB165FD785E63" box="[519,671,798,825]" metricMagnitude="3" metricUnit="m" metricValue="1.47" metricValueMax="1.5699999999999998" metricValueMin="1.3699999999999999" pageId="58" pageNumber="59" unit="m" value="1470.0" valueMax="1570.0" valueMin="1370.0">1370-1570 m</quantity>
,
<date id="FF90E1F5627CFFD92FD2B165FCD95E62" box="[682,830,798,824]" pageId="58" pageNumber="59" value="1988-08-13">1988.VIII.13</date>
, Wang Shuyong lgt.&quot;. 1 ex (IZAS), &quot;
<collectingRegion id="49EA09D7627CFFD929AFB164FAC35E63" box="[1239,1316,799,825]" country="China" name="Hunan" pageId="58" pageNumber="59">Hunan</collectingRegion>
, Tianpingshan,
<date id="FF90E1F5627CFFD92DA1B139FE835E06" box="[217,356,834,860]" pageId="58" pageNumber="59" value="1988-07-20">1988.VII.20</date>
, Lin Su lgt.&quot;.
<specimenCount id="9D280CBC627CFFD92F7BB139FD835E06" box="[515,612,834,860]" pageId="58" pageNumber="59" type="female">1 female</specimenCount>
(HBUM), &quot;
<collectingRegion id="49EA09D7627CFFD92F95B139FCAD5E06" box="[749,842,834,860]" country="China" name="Sichuan" pageId="58" pageNumber="59">Sichuan</collectingRegion>
, Tianquan, Labahe,
<date id="FF90E1F5627CFFD92955B139FB5F5E06" box="[1069,1208,834,860]" pageId="58" pageNumber="59" value="2004-07-29">2004.VII.29</date>
, Yang Xiujuan, Hua Huiran lgt.&quot;.
<specimenCount id="9D280CBC627CFFD92C50B11DFE6F5EDA" box="[296,392,870,896]" pageId="58" pageNumber="59" type="female">1 female</specimenCount>
(IZAS), &quot;
<collectingRegion id="49EA09D7627CFFD92C80B11DFDB25EDA" box="[504,597,870,896]" country="China" name="Sichuan" pageId="58" pageNumber="59">Sichuan</collectingRegion>
, Emeishan, Xixinsuo,
<quantity id="4CD66AD0627CFFD92E37B11DFC425EDB" box="[847,933,870,897]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="58" pageNumber="59" unit="m" value="1300.0">1300 m</quantity>
,
<date id="FF90E1F5627CFFD92EC8B11DFB8E5EDB" box="[944,1129,870,897]" pageId="58" pageNumber="59" value="2012-08-10" valueMax="2012-08-15" valueMin="2012-08-10">2012.VIII.10-15</date>
, Huang Hao leg.&quot;.
<specimenCount id="9D280CBC627CFFD92844B11DFA7B5EDA" box="[1340,1436,870,896]" pageId="58" pageNumber="59" type="female">1 female</specimenCount>
(CCCC), &quot;
<collectingRegion id="49EA09D7627CFFD92C76B1F1FE8C5EFE" box="[270,363,906,932]" country="China" name="Sichuan" pageId="58" pageNumber="59">Sichuan</collectingRegion>
, Emeishan, Xixinsuo,
<quantity id="4CD66AD0627CFFD92F11B1F1FD275EFF" box="[617,704,906,933]" metricMagnitude="3" metricUnit="m" metricValue="1.35" pageId="58" pageNumber="59" unit="m" value="1350.0">1350 m</quantity>
,
<date id="FF90E1F5627CFFD92FB5B1F1FCA65EFE" box="[717,833,906,933]" pageId="58" pageNumber="59" value="2017-05-24">2017.V.24</date>
, Yang Xiaodong leg.&quot;.
<specimenCount id="9D280CBC627CFFD9293FB1F1FB755EFE" box="[1095,1170,906,932]" pageId="58" pageNumber="59" type="male">1 male</specimenCount>
(IZAS), &quot;
<collectingRegion id="49EA09D7627CFFD9287DB1F1FA8F5EFE" box="[1285,1384,906,932]" country="China" name="Guizhou" pageId="58" pageNumber="59">Guizhou</collectingRegion>
, Zunyi, Kuankuoshui,
<date id="FF90E1F5627CFFD92C09B1D5FE025E92" box="[369,485,942,968]" pageId="58" pageNumber="59" value="2010-06-03">2010.VI.3</date>
, Wang Zhiliang lgt.&quot;. 1 ex (IZAS), &quot;
<collectingRegion id="49EA09D7627CFFD92EF5B1D5FC175E92" box="[909,1008,942,968]" country="China" name="Guizhou" pageId="58" pageNumber="59">Guizhou</collectingRegion>
, Leishan, Lanhuaping,
<quantity id="4CD66AD0627CFFD92985B1D5FA705E92" box="[1277,1431,942,969]" metricMagnitude="3" metricUnit="m" metricValue="1.8" metricValueMax="2.1" metricValueMin="1.5" pageId="58" pageNumber="59" unit="m" value="1800.0" valueMax="2100.0" valueMin="1500.0">1500-2100 m</quantity>
,
<date id="FF90E1F5627CFFD92DEFB1AAFEFE5EB6" box="[151,281,977,1004]" pageId="58" pageNumber="59" value="2005-09-16">2005.IX.16</date>
, Liu Ye lgt.&quot;. 1 ex (IZAS), &quot;
<collectingRegion id="49EA09D7627CFFD92F35B1AAFD575EB1" box="[589,688,977,1003]" country="China" name="Guizhou" pageId="58" pageNumber="59">Guizhou</collectingRegion>
, Leishan, Leigongshan,
<quantity id="4CD66AD0627CFFD92EC6B1A9FBF45EB6" box="[958,1043,978,1004]" metricMagnitude="3" metricUnit="m" metricValue="1.17" pageId="58" pageNumber="59" unit="m" value="1170.0">1170 m</quantity>
,
<date id="FF90E1F5627CFFD92965B1A9FB7D5EB6" box="[1053,1178,978,1004]" pageId="58" pageNumber="59" value="1988-07-03">1988.VII.3</date>
, Zhang Xiaochun lgt.&quot;.
<specimenCount id="9D280CBC627CFFD92DEFB18EFF035955" box="[151,228,1013,1039]" pageId="58" pageNumber="59" type="male">1 male</specimenCount>
(HBUM), &quot;
<collectingRegion id="49EA09D7627CFFD92C16B18EFE365955" box="[366,465,1013,1039]" country="China" name="Guizhou" pageId="58" pageNumber="59">Guizhou</collectingRegion>
, Leigongshan,
<date id="FF90E1F5627CFFD92F06B18EFCC2594A" box="[638,805,1013,1040]" pageId="58" pageNumber="59" value="2005-09-13" valueMax="2005-09-14" valueMin="2005-09-13">2005.IX.13-14</date>
, Wang Jiliang, Gao Chao lgt.&quot;. 1 ex (IZAS), &quot;
<collectingRegion id="49EA09D7627CFFD9284DB18EFA7F5955" box="[1333,1432,1013,1039]" country="China" name="Guizhou" pageId="58" pageNumber="59">Guizhou</collectingRegion>
, Daozhen, Dashahe,
<quantity id="4CD66AD0627CFFD92C04B662FE32596E" box="[380,469,1049,1076]" metricMagnitude="3" metricUnit="m" metricValue="1.42" pageId="58" pageNumber="59" unit="m" value="1420.0">1420 m</quantity>
,
<date id="FF90E1F5627CFFD92C9BB662FD9A5969" box="[483,637,1049,1076]" pageId="58" pageNumber="59" value="2006-05-13" valueMax="2006-05-16" valueMin="2006-05-13">2006.V.13-16</date>
, Tang Yi lgt.&quot;. 1 ex (HBUM), &quot;
<collectingRegion id="49EA09D7627CFFD92E95B662FBB75969" box="[1005,1104,1049,1075]" country="China" name="Guizhou" pageId="58" pageNumber="59">Guizhou</collectingRegion>
, Daozhen, Sanqiao, Yu Yang lgt.,
<date id="FF90E1F5627CFFD92DB2B646FEA6590D" box="[202,321,1085,1111]" pageId="58" pageNumber="59" value="2004-05-27">2004.V.27</date>
&quot;.
<specimenCount id="9D280CBC627CFFD92C2EB646FE46590D" box="[342,417,1085,1111]" pageId="58" pageNumber="59" type="male">1 male</specimenCount>
(HBUM), &quot;
<collectingRegion id="49EA09D7627CFFD92F51B646FD6B590D" box="[553,652,1085,1111]" country="China" name="Guizhou" pageId="58" pageNumber="59">Guizhou</collectingRegion>
, Daozhen, Yangsui,
<date id="FF90E1F5627CFFD92E08B646FC03590D" box="[880,996,1085,1111]" pageId="58" pageNumber="59" value="2004-05-31">2004.V.31</date>
, Yu Yang lgt.&quot;. 1 ex (IZAS), &quot;
<collectingRegion id="49EA09D7627CFFD9284DB646FA7F590D" box="[1333,1432,1085,1111]" country="China" name="Guizhou" pageId="58" pageNumber="59">Guizhou</collectingRegion>
, Jiangkou, Fanjingshan, Huixiangping,
<quantity id="4CD66AD0627CFFD92F34B61BFD435921" box="[588,676,1120,1147]" metricMagnitude="3" metricUnit="m" metricValue="1.775" pageId="58" pageNumber="59" unit="m" value="1775.0">1775 m</quantity>
,
<date id="FF90E1F5627CFFD92FC9B61AFCD45921" box="[689,819,1121,1147]" pageId="58" pageNumber="59" value="2009-06-22">2009.VI.22</date>
, Lin Wensin lgt.&quot;.
<specimenCount id="9D280CBC627CFFD92971B61AFB8C5920" box="[1033,1131,1120,1147]" pageId="58" pageNumber="59" type="female">1 female</specimenCount>
(IZAS), &quot;
<collectingRegion id="49EA09D7627CFFD929A2B61AFAD15921" box="[1242,1334,1121,1147]" country="China" name="Yunnan" pageId="58" pageNumber="59">Yunnan</collectingRegion>
, Jinping, Fenshuiling,
<quantity id="4CD66AD0627CFFD92C54B6FFFE6159C5" box="[300,390,1156,1183]" metricMagnitude="3" metricUnit="m" metricValue="1.8699999999999999" pageId="58" pageNumber="59" unit="m" value="1870.0">1870 m</quantity>
,
<date id="FF90E1F5627CFFD92CECB6FFFDEE59C4" box="[404,521,1156,1183]" pageId="58" pageNumber="59" value="2009-05-27">2009.V.27</date>
, Bi Wenxuan leg.&quot; &lt;
<figureCitation id="1315DBB0627CFFD92F84B6FFFC8259C4" box="[764,869,1156,1182]" captionStart="FIGURE 32" captionStartId="59.[151,250,1549,1573]" captionTargetBox="[156,1430,184,1511]" captionTargetId="figure-19@59.[151,1436,181,1525]" captionTargetPageId="59" captionText="FIGURE 32. Habitus of Parena (Bothynoptera) tesari species group. A Holotype of Parena tesari (Jedlička). B Holotype of Parena kataevi Kirschenhofer [=Parena tesari (Jedlička)]. C P. tesari (Jedlička), male from Taiwan. D P. tesari (Jedlička), fe- male from Yunnan. E Holotype of Parena obscura Mateu. F P. obscura Mateu, male from Nepal. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7959003" httpUri="https://zenodo.org/record/7959003/files/figure.png" pageId="58" pageNumber="59">Fig. 32D</figureCitation>
&gt;.
<specimenCount id="9D280CBC627CFFD92EFBB6FFFC3759C4" box="[899,976,1156,1182]" pageId="58" pageNumber="59" type="male">1 male</specimenCount>
(IZAS), &quot;
<collectingRegion id="49EA09D7627CFFD9293CB6FFFB4759C4" box="[1092,1184,1156,1182]" country="China" name="Yunnan" pageId="58" pageNumber="59">Yunnan</collectingRegion>
, Jinping, Fenshuiling,
<quantity id="4CD66AD0627CFFD92DEFB6D3FF0A5999" box="[151,237,1192,1219]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="58" pageNumber="59" unit="m" value="1900.0">1900 m</quantity>
,
<date id="FF90E1F5627CFFD92D80B6D3FE915999" box="[248,374,1192,1219]" pageId="58" pageNumber="59" value="2010-04-15">2010.IV.15</date>
, Zhu Xiaoyu leg.&quot; &lt;
<figureCitation id="1315DBB0627CFFD92F20B6D3FD595998" box="[600,702,1192,1218]" captionStart="FIGURE 34" captionStartId="60.[151,250,1977,2001]" captionTargetBox="[160,1428,1372,1949]" captionTargetId="figure-208@60.[151,1437,1361,1954]" captionTargetPageId="60" captionText="FIGURE 34. Median lobe of male genitalia for Parena (Bothynoptera) tesari (Jedlička) from different localities, left-lateral (l) and dorsal (d) views: A Yunnan. B Taiwan. C Zhejiang. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7959007" httpUri="https://zenodo.org/record/7959007/files/figure.png" pageId="58" pageNumber="59">Fig. 34A</figureCitation>
&gt;. 1 ex (CCCC), &quot;Nantou county, Ren'ai, Songgang,
<date id="FF90E1F5627CFFD9287BB6D3FA7F5998" box="[1283,1432,1192,1218]" pageId="58" pageNumber="59" value="1994-08-24">1994.VIII.24</date>
, Chen Changchin lgt.&quot;. 1 ex (CCCC), &quot; Nantou county, Ren'ai, Nanshanhsi,
<date id="FF90E1F5627CFFD92E9BB6B7FB8759BC" box="[995,1120,1228,1254]" pageId="58" pageNumber="59" value="1993-07-01">1993.VII.1</date>
, Chen Changchin lgt.&quot;. 2 ex (CCCC), &quot;Nantou, Puli, Guandaoshan,
<date id="FF90E1F5627CFFD92F2BB68BFD335850" box="[595,724,1263,1290]" pageId="58" pageNumber="59" value="1993-06-15">1993.VI.15</date>
, Chen Changchin lgt.&quot;. 1 ex (CCCC), &quot; Nantou county, Ren'ai, Songgang,
<quantity id="4CD66AD0627CFFD92C6DB768FE8B5874" box="[277,364,1299,1326]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="58" pageNumber="59" unit="m" value="2000.0">2000 m</quantity>
,
<date id="FF90E1F5627CFFD92C00B768FE125877" box="[376,501,1299,1326]" pageId="58" pageNumber="59" value="1995-04-17">1995.IV.17</date>
, Lo Chinchi lgt.&quot;. 5 ex (CCCC), &quot; Nantou county, Ren'ai, Songgang,
<date id="FF90E1F5627CFFD92987B768FA735877" box="[1279,1428,1299,1325]" pageId="58" pageNumber="59" value="1994-08-16">1994.VIII.16</date>
&quot; &lt;
<figureCitation id="1315DBB0627CFFD92DDFB74CFEEB580B" box="[167,268,1335,1361]" captionStart="FIGURE 11" captionStartId="18.[151,250,150,174]" captionTargetId="figure-19@17.[155,1432,181,1971]" captionTargetPageId="17" captionText="FIGURE 11. Gonocoxites (left side) of ovipositor for Parena spp., ventral side (dorsal side for fig. Q). A–J subgenus Bothynoptera: A P. (Bothynoptera) heteronycha sp.n., Laos, paratype. B P. (Bothynoptera) dorsigera (Schaum), Shaanxi. C P. (Bothynoptera) kurosai Habu, Guangdong. D P. (Bothynoptera) tesari (Jedlička), Taiwan. E P. (Bothynoptera) taiwana Hua, Taiwan. F P. (Bothynoptera) tripunctata (Bates), Shaanxi. G P. (Bothynoptera) malaisei (Andrewes), Taiwan. H P. (Bothynoptera) quadrisignata Mateu, Yunnan. I P. (Bothynoptera) emarginata sp. n., holotype. J P. (Bothynoptera) triguttata sp. n., Sichuan, paratype. K–X subgenus Parena: K P. (Parena) bicolor Motschulsky, Sulawesi. L P. (Parena) rubripicta Andrewes, Yunnan. M P. (Parena) fasciata (Chaudoir), Taiwan. N P. (Parena) nigrolineata (Chaudoir), Taiwan. O P. (Parena) picipes sp. n., Borneo, paratype. P P. (Parena) andrewesi Jedlička, Philippine. Q P. (Parena) politissima (Chaudoir), Vanuatu. R P. (Parena) latecincta (Bates), Guizhou. S P. (Parena) circumdata Shibata, Taiwan. T P. (Parena) pendleburyi Andrewes, Borneo. U P. (Parena) fulva sp. n., Zimbabwe, paratype. V P. (Parena) madagascariensis (Alluaud), Madagascar. W P. (Parena) ruficornis sp.n., CentralAfrica, paratype. X P. (Parena) africana (Alluaud), Uganda. Y–γ subgenus Crossoglossa: Y P. (Crossoglossa) cavipennis (Bates), Taiwan. Z P. (Crossoglossa) laesipennis (Bates), Yunnan. α P. (Crossoglossa) testacea (Chaudoir), Zhejiang. β P. (Crossoglossa) sulawesiensis Kirschenhofer, Sulawesi. γ P. (Crossoglossa) sciakyi sp. n., holotype. Scale bar = 0.2mm." figureDoi="http://doi.org/10.5281/zenodo.7963954" httpUri="https://zenodo.org/record/7963954/files/figure.png" pageId="58" pageNumber="59">Fig. 11D</figureCitation>
&gt;.
<specimenCount id="9D280CBC627CFFD92C50B74CFE94580B" box="[296,371,1335,1361]" pageId="58" pageNumber="59" type="male">1 male</specimenCount>
(CCCC), &quot;Taiwan, Nantou, Renai, N. Dongyanshan,
<quantity id="4CD66AD0627CFFD92EB6B74CFBC2580B" box="[974,1061,1335,1361]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="58" pageNumber="59" unit="m" value="1800.0">1800 m</quantity>
,
<date id="FF90E1F5627CFFD92949B74CFB48580B" box="[1073,1199,1335,1361]" pageId="58" pageNumber="59" value="2010-04-22">2010.IV.22</date>
, Chou Wen-I leg.&quot;. 1 ex (SYUM), &quot;Formose Musha (Wuse), Teichung Distr.,
<quantity id="4CD66AD0627CFFD92E77B720FC80582F" box="[783,871,1371,1397]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="58" pageNumber="59" unit="m" value="1000.0">1000 m</quantity>
,
<date id="FF90E1F5627CFFD92E0CB720FBEE582F" box="[884,1033,1371,1397]" pageId="58" pageNumber="59" value="1947-08-24">1947.VIII.24</date>
, J. Linsley, Gressitt&quot;. 1 ex (CCCC), &quot;Hsinchu county, Wufeng, Talu Forest Road,
<date id="FF90E1F5627CFFD92FF4B705FD1458C2" box="[652,755,1406,1433]" pageId="58" pageNumber="59" value="1994-05-08">1994.V.8</date>
&quot;. 1 ex (CCCC), &quot; Hsinchu county, Wufeng, Talu Forest Road,
<date id="FF90E1F5627CFFD92DEFB7D9FEC558E6" box="[151,290,1442,1468]" pageId="58" pageNumber="59" value="1993-07-17">1993.VII.17</date>
, Chen Changchin lgt.&quot;. 1 ex (CCCC), &quot; Hsinchu county, Wufeng, Taping,
<date id="FF90E1F5627CFFD9292DB7D9FB3058E6" box="[1109,1239,1442,1468]" pageId="58" pageNumber="59" value="1994-06-12">1994.VI.12</date>
, Chen Changchin lgt.&quot;. 1 ex (CCCC), &quot;Taiwan, Hsinchu county, Wufeng, Meikang,
<date id="FF90E1F5627CFFD92E0DB7BDFC1058BA" box="[885,1015,1478,1504]" pageId="58" pageNumber="59" value="1994-09-10">1994.IX.10</date>
, Chen Changchin lgt.&quot;. 2 ex (CCCC), &quot;Taiwan, Hsinchu county, Jianshi, Ninglao vill.,
<date id="FF90E1F5627CFFD92FC7B791FCD45B5E" box="[703,819,1514,1540]" pageId="58" pageNumber="59" value="1997-06-01">1997.VI.1</date>
, Chen Changchin lgt.&quot;. 1 ex (CCCC), &quot;Hsinchu county, Jianshi,
<date id="FF90E1F5627CFFD92C65B475FE7C5B72" box="[285,411,1550,1577]" pageId="58" pageNumber="59" value="2005-04-21">2005.IV.21</date>
, Chen Changchin lgt.&quot;. 1 ex (CCCC), &quot; Hsinchu county, Skaru,
<date id="FF90E1F5627CFFD929EAB475FACC5B72" box="[1170,1323,1550,1577]" pageId="58" pageNumber="59" value="2005-08-29">2005.VIII.29</date>
&quot;.
<specimenCount id="9D280CBC627CFFD9283AB475FA7B5B72" box="[1346,1436,1550,1576]" pageId="58" pageNumber="59" type="male">2 males</specimenCount>
(CCCC), &quot;Taiwan, Taitung county, Haiduan,
<date id="FF90E1F5627CFFD92FF6B449FCE55B16" box="[654,770,1586,1612]" pageId="58" pageNumber="59" value="1996-05-18">1996.V.18</date>
, Chen Changchin lgt.&quot; &lt;
<figureCitation id="1315DBB0627CFFD9296FB449FB9B5B16" box="[1047,1148,1586,1612]" captionStart="FIGURE 34" captionStartId="60.[151,250,1977,2001]" captionTargetBox="[160,1428,1372,1949]" captionTargetId="figure-208@60.[151,1437,1361,1954]" captionTargetPageId="60" captionText="FIGURE 34. Median lobe of male genitalia for Parena (Bothynoptera) tesari (Jedlička) from different localities, left-lateral (l) and dorsal (d) views: A Yunnan. B Taiwan. C Zhejiang. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7959007" httpUri="https://zenodo.org/record/7959007/files/figure.png" pageId="58" pageNumber="59">Fig. 34B</figureCitation>
&gt;. 1 ex (CCCC), &quot;Taiwan, Taoyuan county, Fuxing county, Siling,
<date id="FF90E1F5627CFFD92F21B42EFD2A5B2A" box="[601,717,1621,1648]" pageId="58" pageNumber="59" value="1996-05-20">1996.V.20</date>
, Chen Changchin lgt.&quot;. 1 ex (CCCC), &quot;Taiwan, Hualien county, Ruisui Forest Road,
<date id="FF90E1F5627CFFD92CF8B402FE1A5BCE" box="[384,509,1657,1684]" pageId="58" pageNumber="59" value="1996-07-05">1996.VII.5</date>
, Chou Wen-I lgt.&quot;. 1 ex (CCCC), &quot;Taipei county, Sanxia, Shizaitou,
<date id="FF90E1F5627CFFD9287AB402FA9D5BCE" box="[1282,1402,1657,1684]" pageId="58" pageNumber="59" value="1994-09-05">1994.IX.5</date>
&quot;.
<specimenCount id="9D280CBC627CFFD928F6B402FF2A5BED" pageId="58" pageNumber="59" type="male">1 male</specimenCount>
(CCCC), &quot;Taiwan, Miaoli county, Sanyi, Guandaoshan,
<date id="FF90E1F5627CFFD92E36B4E6FC2B5BED" box="[846,972,1693,1720]" pageId="58" pageNumber="59" value="1995-07-02">1995.VII.2</date>
&quot; &lt;
<figureCitation id="1315DBB0627CFFD92E93B4E6FBB35BED" box="[1003,1108,1693,1719]" captionStart="FIGURE 32" captionStartId="59.[151,250,1549,1573]" captionTargetBox="[156,1430,184,1511]" captionTargetId="figure-19@59.[151,1436,181,1525]" captionTargetPageId="59" captionText="FIGURE 32. Habitus of Parena (Bothynoptera) tesari species group. A Holotype of Parena tesari (Jedlička). B Holotype of Parena kataevi Kirschenhofer [=Parena tesari (Jedlička)]. C P. tesari (Jedlička), male from Taiwan. D P. tesari (Jedlička), fe- male from Yunnan. E Holotype of Parena obscura Mateu. F P. obscura Mateu, male from Nepal. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7959003" httpUri="https://zenodo.org/record/7959003/files/figure.png" pageId="58" pageNumber="59">Figs 32C</figureCitation>
,
<figureCitation id="1315DBB0627CFFD92919B4E6FB985BED" box="[1121,1151,1693,1719]" captionStart="FIGURE 33" captionStartId="60.[151,250,1277,1301]" captionTargetBox="[162,1428,460,1246]" captionTargetId="figure-174@60.[152,1436,453,1253]" captionTargetPageId="60" captionText="FIGURE 33. Median lobe of male genitalia for Parena (Bothynoptera) tesari (Jedlička), right-lateral (r), ventral (v), left-lateral (l), and dorsal (d) views, Taiwan. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7959005" httpUri="https://zenodo.org/record/7959005/files/figure.png" pageId="58" pageNumber="59">33</figureCitation>
&gt;.
<specimenCount id="9D280CBC627CFFD929E3B4E6FB1A5BED" box="[1179,1277,1693,1719]" pageId="58" pageNumber="59" type="female">1 female</specimenCount>
(IZAS), &quot;Taiwan, Hsinpei, Beichatianshan,
<quantity id="4CD66AD0627CFFD92C8EB4BAFDA85B81" box="[502,591,1729,1755]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="58" pageNumber="59" unit="m" value="1700.0">1700 m</quantity>
,
<date id="FF90E1F5627CFFD92F24B4BAFD395B81" box="[604,734,1729,1755]" pageId="58" pageNumber="59" value="2020-06-12">2020.VI.12</date>
, Huang Jialong lgt.&quot; &lt;
<figureCitation id="1315DBB0627CFFD92E98B4BAFBD15B86" box="[992,1078,1729,1756]" captionStart="FIGURE 5" captionStartId="12.[151,250,1858,1882]" captionTargetBox="[185,1402,181,1834]" captionTargetId="figure-28@12.[185,1402,181,1834]" captionTargetPageId="12" captionText="FIGURE 5. Elytral apex (left) of Parena spp. A P. (Parena) africana (Alluaud), Cameroon. B P. (Parena) ruficornis sp. n., Central Africa, paratype. C P. (Parena) valeriae Facchini, holotype. D P. (Parena) madagascariensis (Alluaud), Madagascar. E P. (Bothynoptera) tesari (Jedlička), Taiwan. F P. (Bothynoptera) heteronycha sp. n., Laos, paratype. G P. (Bothynoptera) kurosai Habu, Japan. H P. (Bothynoptera) dorsigera (Schaum), Yunnan. I P. (Bothynoptera) dorsigera (Schaum), Beijing. J P. (Bothynoptera) dorsigera (Schaum), Japan. Scale bars = 0.5mm" figureDoi="http://doi.org/10.5281/zenodo.7958947" httpUri="https://zenodo.org/record/7958947/files/figure.png" pageId="58" pageNumber="59">Fig. 5E</figureCitation>
&gt;. 3 ex (MTMB), &quot;Taiwan, Ilan county, Mingchyh Forest, Recreation area,
<quantity id="4CD66AD0627CFFD92FF9B49EFD3E5BA5" box="[641,729,1765,1791]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="58" pageNumber="59" unit="m" value="1200.0">1200 m</quantity>
, at light,
<date id="FF90E1F5627CFFD92E3DB49FFC515BA5" box="[837,950,1764,1791]" pageId="58" pageNumber="59" value="2001-04-05">5.IV.2001</date>
, leg. Gy. Fabian &amp; O. Merkl&quot;.
<emphasis id="B95A1B27627CFFD92861B49FFA985BA4" bold="true" box="[1305,1407,1764,1790]" pageId="58" pageNumber="59">
<collectingCountry id="F33987A5627CFFD92861B49FFA985BA4" box="[1305,1407,1764,1790]" name="Vietnam" pageId="58" pageNumber="59">Vietnam</collectingCountry>
</emphasis>
:
<specimenCount id="9D280CBC627CFFD928F6B49EFF2A5A78" pageId="58" pageNumber="59" type="male">1 male</specimenCount>
(CRS), &quot;N. Vietnam Mt. Fan-sipan N-Seite Cha-pa (=Sapa)
<quantity id="4CD66AD0627CFFD92EFEB573FC3A5A79" box="[902,989,1800,1827]" metricMagnitude="3" metricUnit="m" metricValue="1.525" pageId="58" pageNumber="59" unit="m" value="1525.0">1525 m</quantity>
22.17'
<geoCoordinate id="EE1AA1F2627CFFD92951B573FB705A78" box="[1065,1175,1800,1826]" degrees="103.44" direction="north" orientation="latitude" pageId="58" pageNumber="59" precision="555" value="103.44">N 103.44</geoCoordinate>
'E, prim. Urwald
<date id="FF90E1F5627CFFD92823B573FEE05A1C" pageId="58" pageNumber="59" value="1994-10-28" valueMax="1994-11-03" valueMin="1994-10-28">28.x.- 3.xi.1994</date>
&quot;. 1 ex (MNHN), &quot;Chapa, Tonkin&quot;.
<specimenCount id="9D280CBC627CFFD92FEEB557FD055A1C" box="[662,738,1836,1862]" pageId="58" pageNumber="59" type="male">1 male</specimenCount>
(IZAS), &quot;Vietnam, LaoCai Prov., SaPa, Hoang Lien National Park,
<date id="FF90E1F5627CFFD92DA0B52BFE8A5A30" box="[216,365,1871,1898]" pageId="58" pageNumber="59" value="2009-07-05" valueMax="2009-07-09" valueMin="2009-07-05">2009.VII.5-9</date>
, Li Hu leg.&quot;
</paragraph>
</subSubSection>
<subSubSection id="C33494BE627CFFD92DBFB508FE595AA3" pageId="58" pageNumber="59" type="diagnosis">
<paragraph id="8B91C735627CFFD92DBFB508FE595AA3" blockId="58.[151,1437,150,2041]" pageId="58" pageNumber="59">
<emphasis id="B95A1B27627CFFD92DBFB508FE815AD7" bold="true" box="[199,358,1907,1933]" pageId="58" pageNumber="59">Comparisons</emphasis>
.
<taxonomicName id="4C2EBCB6627CFFD92C0BB50FFE2A5AD7" authorityName="Jedlicka" authorityYear="1951" box="[371,461,1908,1933]" class="Insecta" family="Carabidae" genus="Bothynoptera" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="tesari">
<emphasis id="B95A1B27627CFFD92C0BB50FFE2A5AD7" box="[371,461,1908,1933]" italics="true" pageId="58" pageNumber="59">P. tesari</emphasis>
</taxonomicName>
is different from other species of subgenus
<taxonomicName id="4C2EBCB6627CFFD92ED6B508FBA15AD7" authority="Schaum, 1863" box="[942,1094,1907,1933]" class="Insecta" family="Carabidae" genus="Bothynoptera" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis id="B95A1B27627CFFD92ED6B508FBA15AD7" box="[942,1094,1907,1933]" italics="true" pageId="58" pageNumber="59">Bothynoptera</emphasis>
</taxonomicName>
(except for
<taxonomicName id="4C2EBCB6627CFFD929B3B50FFAA75AD7" authorityName="Mateu. F" authorityYear="1977" box="[1227,1344,1907,1933]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="obscura">
<emphasis id="B95A1B27627CFFD929B3B50FFAA75AD7" box="[1227,1344,1907,1933]" italics="true" pageId="58" pageNumber="59">P. obscura</emphasis>
</taxonomicName>
) in having the elongate tempora, which are gradually narrowed behind eyes. In other species, the tempora are very short and abruptly narrowed behind the eyes with the length of tempora plus neck-constriction approximately one-third of the diameter of the eye.
</paragraph>
</subSubSection>
<caption id="DF5197BD627DFFD82DEFB476FADB5B37" ID-DOI="http://doi.org/10.5281/zenodo.7959003" ID-Zenodo-Dep="7959003" httpUri="https://zenodo.org/record/7959003/files/figure.png" pageId="59" pageNumber="60" startId="59.[151,250,1549,1573]" targetBox="[156,1430,184,1511]" targetPageId="59" targetType="figure">
<paragraph id="8B91C735627DFFD82DEFB476FADB5B37" blockId="59.[151,1436,1549,1645]" pageId="59" pageNumber="60">
<emphasis id="B95A1B27627DFFD82DEFB476FEFD5B7F" bold="true" box="[151,282,1549,1573]" pageId="59" pageNumber="60">FIGURE 32</emphasis>
. Habitus of
<taxonomicName id="4C2EBCB6627DFFD82CE3B475FD2F5B7F" baseAuthorityName="Jedlicka" baseAuthorityYear="1951" box="[411,712,1549,1573]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="tesari" subGenus="Bothynoptera">
<emphasis id="B95A1B27627DFFD82CE3B475FE025B7F" box="[411,485,1550,1573]" italics="true" pageId="59" pageNumber="60">Parena</emphasis>
(
<emphasis id="B95A1B27627DFFD82C8DB476FD985B7F" box="[501,639,1549,1573]" italics="true" pageId="59" pageNumber="60">Bothynoptera</emphasis>
)
<emphasis id="B95A1B27627DFFD82FF6B475FD2F5B7F" box="[654,712,1550,1573]" italics="true" pageId="59" pageNumber="60">tesari</emphasis>
</taxonomicName>
species group.
<emphasis id="B95A1B27627DFFD82E11B476FC9C5B7F" bold="true" box="[873,891,1549,1573]" pageId="59" pageNumber="60">A</emphasis>
Holotype of
<taxonomicName id="4C2EBCB6627DFFD8297DB475FB195B7F" authority="(Jedlicka)" baseAuthorityName="Jedlicka" baseAuthorityYear="1951" box="[1029,1278,1549,1573]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="tesari">
<emphasis id="B95A1B27627DFFD8297DB475FB775B7F" box="[1029,1168,1550,1573]" italics="true" pageId="59" pageNumber="60">Parena tesari</emphasis>
(Jedlička)
</taxonomicName>
.
<emphasis id="B95A1B27627DFFD82871B476FAFD5B7F" bold="true" box="[1289,1306,1549,1573]" pageId="59" pageNumber="60">B</emphasis>
Holotype of
<taxonomicName id="4C2EBCB6627DFFD82DEFB449FE2E5B13" authority="Kirschenhofer" authorityName="Kirschenhofer" authorityYear="2006" box="[151,457,1585,1609]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="kataevi">
<emphasis id="B95A1B27627DFFD82DEFB449FED65B13" box="[151,305,1585,1609]" italics="true" pageId="59" pageNumber="60">Parena kataevi</emphasis>
Kirschenhofer
</taxonomicName>
[=
<taxonomicName id="4C2EBCB6627DFFD82C9EB449FD065B13" authority="(Jedlicka)" baseAuthorityName="Jedlicka" baseAuthorityYear="1951" box="[486,737,1585,1609]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="tesari">
<emphasis id="B95A1B27627DFFD82C9EB449FD965B13" box="[486,625,1586,1609]" italics="true" pageId="59" pageNumber="60">Parena tesari</emphasis>
(Jedlička)
</taxonomicName>
].
<emphasis id="B95A1B27627DFFD82F89B44AFCE45B13" bold="true" box="[753,771,1585,1609]" pageId="59" pageNumber="60">C</emphasis>
<taxonomicName id="4C2EBCB6627DFFD82E72B449FC2D5B13" authority="(Jedlicka)" baseAuthorityName="Jedlicka" baseAuthorityYear="1951" box="[778,970,1585,1609]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="tesari">
<emphasis id="B95A1B27627DFFD82E72B449FCBA5B13" box="[778,861,1586,1609]" italics="true" pageId="59" pageNumber="60">P. tesari</emphasis>
(Jedlička)
</taxonomicName>
, male from Taiwan.
<emphasis id="B95A1B27627DFFD829E4B44AFB495B13" bold="true" box="[1180,1198,1585,1609]" pageId="59" pageNumber="60">D</emphasis>
<taxonomicName id="4C2EBCB6627DFFD829CDB449FA925B13" authority="(Jedlicka)" baseAuthorityName="Jedlicka" baseAuthorityYear="1951" box="[1205,1397,1585,1609]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="tesari">
<emphasis id="B95A1B27627DFFD829CDB449FAEF5B13" box="[1205,1288,1586,1609]" italics="true" pageId="59" pageNumber="60">P. tesari</emphasis>
(Jedlička)
</taxonomicName>
, female from Yunnan.
<emphasis id="B95A1B27627DFFD82C27B42EFE975B37" bold="true" box="[351,368,1621,1645]" pageId="59" pageNumber="60">E</emphasis>
Holotype of
<taxonomicName id="4C2EBCB6627DFFD82C8FB42DFD1C5B37" authority="Mateu. F" authorityName="Mateu. F" authorityYear="1977" box="[503,763,1621,1645]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="obscura">
<emphasis id="B95A1B27627DFFD82C8FB42DFD7E5B37" box="[503,665,1621,1645]" italics="true" pageId="59" pageNumber="60">Parena obscura</emphasis>
Mateu.
<emphasis id="B95A1B27627DFFD82F94B42EFD1C5B37" bold="true" box="[748,763,1621,1645]" pageId="59" pageNumber="60">F</emphasis>
</taxonomicName>
<taxonomicName id="4C2EBCB6627DFFD82E7AB42DFC515B37" authority="Mateu" authorityName="Mateu. F" authorityYear="1977" box="[770,950,1621,1645]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="obscura">
<emphasis id="B95A1B27627DFFD82E7AB42DFC8A5B37" box="[770,877,1621,1645]" italics="true" pageId="59" pageNumber="60">P. obscura</emphasis>
Mateu
</taxonomicName>
, male from Nepal. Scale bars = 2 mm.
</paragraph>
</caption>
<subSubSection id="C33494BE627DFFDF2DBFB4E0FAC65CF7" lastPageId="60" lastPageNumber="61" pageId="59" pageNumber="60" type="description">
<paragraph id="8B91C735627DFFDF2DBFB4E0FAC65CF7" blockId="59.[151,1437,1691,2042]" lastBlockId="60.[151,1437,151,429]" lastPageId="60" lastPageNumber="61" pageId="59" pageNumber="60">
<emphasis id="B95A1B27627DFFD82DBFB4E0FEB65BEF" bold="true" box="[199,337,1691,1717]" pageId="59" pageNumber="60">Description</emphasis>
. Body length 8.8–11.0 mm. Dorsum reddish brown to piceous; in dark color specimens, legs, antennae, and mouthparts distinctly lighter than body. Tempora gradually narrowed behind eyes, tempora plus neck-constriction two-thirds as long as eyes; vertex often with a pair of additional short setae postermedial to the posterior supraorbital seta; mentum with a pair of short setae. Pronotum nearly quadrate, PW/PL = 1.20–1.32, much narrower than head in most specimens, PW/HW = 0.85–0.95; lateral explanations narrow; widest at anterior third, slightly rounded at anterior half, and then clearly sinuate before posterior angles in most specimens; posterior angles nearly rectangular, more or less pointed outward. Elytra wide, slightly dilated to apex, surface without microsculpture; striae not incised or shallowly incised, with fine puncture rows in striae; intervals flat or slightly convex, with fine and sparse punctures; interval 3 with three setigerous pores, only two in some specimens; apical truncation indistinct, evenly rounded, sutural angles indistinct. Males with uniseriate adhesive setae on apical half of mesotarsomere 2 and biseriate adhesive setae on nearly full length of mesotarsomere 3. Apex of abdominal sternite VII with three setae on each side in both sexes, four pairs present in a few specimens. Median lobe of aedeagus slender (AL/AW = 4.5–4.8), apical lamella laminar, LL subequal to LW, slightly narrowed to apex, apex rounded. Endophallus with relatively small primary sclerite, flared basal expansion straight, apical flagellum very short, ending at basal third of median lobe; apical sclerite small and well defined, basal core elongate; basal sheath well scaled, apical sheath composed of three heavily scaled regions; squamate sac not divided, large and elongated, near middle of median lobe, right to squamate sheath. Gonocoxite II of ovipositor nearly quadrate, length slightly greater than basal width, apex slightly concave, with three or four ensiform setae on outer angle, and two or three on inner angle.
</paragraph>
</subSubSection>
<caption id="DF5197BD627AFFDF2DEFB686FD5D5863" ID-DOI="http://doi.org/10.5281/zenodo.7959005" ID-Zenodo-Dep="7959005" httpUri="https://zenodo.org/record/7959005/files/figure.png" pageId="60" pageNumber="61" startId="60.[151,250,1277,1301]" targetBox="[162,1428,460,1246]" targetPageId="60" targetType="figure">
<paragraph id="8B91C735627AFFDF2DEFB686FD5D5863" blockId="60.[151,1437,1277,1337]" pageId="60" pageNumber="61">
<emphasis id="B95A1B27627AFFDF2DEFB686FEFF584F" bold="true" box="[151,280,1277,1301]" pageId="60" pageNumber="61">FIGURE 33</emphasis>
. Median lobe of male genitalia for
<taxonomicName id="4C2EBCB6627AFFDF2F03B685FBF6584F" authority="(Jedlicka)" baseAuthorityName="Jedlicka" baseAuthorityYear="1951" box="[635,1041,1277,1301]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="tesari" subGenus="Bothynoptera">
<emphasis id="B95A1B27627AFFDF2F03B685FD22584F" box="[635,709,1278,1301]" italics="true" pageId="60" pageNumber="61">Parena</emphasis>
(
<emphasis id="B95A1B27627AFFDF2FABB686FCBA584F" box="[723,861,1277,1301]" italics="true" pageId="60" pageNumber="61">Bothynoptera</emphasis>
)
<emphasis id="B95A1B27627AFFDF2E13B685FC42584F" box="[875,933,1278,1301]" italics="true" pageId="60" pageNumber="61">tesari</emphasis>
(Jedlička)
</taxonomicName>
, right-lateral (r), ventral (v), left-lateral (l), and dorsal (d) views, Taiwan. Scale bar = 0.5 mm.
</paragraph>
</caption>
<caption id="DF5197BD627AFFDF2DEFB5C2FC785AAF" ID-DOI="http://doi.org/10.5281/zenodo.7959007" ID-Zenodo-Dep="7959007" httpUri="https://zenodo.org/record/7959007/files/figure.png" pageId="60" pageNumber="61" startId="60.[151,250,1977,2001]" targetBox="[160,1428,1372,1949]" targetPageId="60" targetType="figure">
<paragraph id="8B91C735627AFFDF2DEFB5C2FC785AAF" blockId="60.[151,1436,1977,2038]" pageId="60" pageNumber="61">
<emphasis id="B95A1B27627AFFDF2DEFB5C2FEFE5A8B" bold="true" box="[151,281,1977,2001]" pageId="60" pageNumber="61">FIGURE 34</emphasis>
. Median lobe of male genitalia for
<taxonomicName id="4C2EBCB6627AFFDF2F06B5C1FBF45A88" authority="(Jedlicka)" baseAuthorityName="Jedlicka" baseAuthorityYear="1951" box="[638,1043,1978,2002]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="tesari" subGenus="Bothynoptera">
<emphasis id="B95A1B27627AFFDF2F06B5C1FD2F5A8B" box="[638,712,1978,2001]" italics="true" pageId="60" pageNumber="61">Parena</emphasis>
(
<emphasis id="B95A1B27627AFFDF2FAEB5C1FC875A88" box="[726,864,1978,2002]" italics="true" pageId="60" pageNumber="61">Bothynoptera</emphasis>
)
<emphasis id="B95A1B27627AFFDF2E17B5C1FC4E5A8B" box="[879,937,1978,2001]" italics="true" pageId="60" pageNumber="61">tesari</emphasis>
(Jedlička)
</taxonomicName>
from different localities, left-lateral (l) and dorsal (d) views:
<emphasis id="B95A1B27627AFFDF2C0BB5A6FE625AAF" bold="true" box="[371,389,2013,2037]" pageId="60" pageNumber="61">A</emphasis>
Yunnan.
<emphasis id="B95A1B27627AFFDF2C9CB5A5FE125AAC" bold="true" box="[484,501,2014,2038]" pageId="60" pageNumber="61">B</emphasis>
Taiwan.
<emphasis id="B95A1B27627AFFDF2F2AB5A6FD835AAF" bold="true" box="[594,612,2013,2037]" pageId="60" pageNumber="61">C</emphasis>
Zhejiang. Scale bar = 0.5 mm.
</paragraph>
</caption>
<subSubSection id="C33494BE627BFFDE2DBFB2EDFDAD5E50" pageId="61" pageNumber="62" type="distribution">
<paragraph id="8B91C735627BFFDE2DBFB2EDFEA45D8F" blockId="61.[151,1436,150,285]" pageId="61" pageNumber="62">
<emphasis id="B95A1B27627BFFDE2DBFB2EDFEBE5DEA" bold="true" box="[199,345,150,176]" pageId="61" pageNumber="62">Distribution</emphasis>
(
<figureCitation id="1315DBB0627BFFDE2C11B2ECFE515DEB" box="[361,438,151,177]" captionStart="MAP 7" captionStartId="57.[152,211,1002,1026]" captionTargetBox="[200,1388,181,979]" captionTargetId="figure-19@57.[200,1388,181,979]" captionTargetPageId="57" captionText="MAP 7. Distributions for species in the Parena (Bothynoptera) tesari group" figureDoi="http://doi.org/10.5281/zenodo.7963671" httpUri="https://zenodo.org/record/7963671/files/figure.png" pageId="61" pageNumber="62">Map 7</figureCitation>
, blue).
<collectingCountry id="F33987A5627BFFDE2F73B2ECFDA95DEB" box="[523,590,151,177]" name="China" pageId="61" pageNumber="62">China</collectingCountry>
(
<collectingRegion id="49EA09D7627BFFDE2F26B2ECFD4F5DEB" box="[606,680,151,177]" country="China" name="Anhui" pageId="61" pageNumber="62">Anhui</collectingRegion>
,
<collectingRegion id="49EA09D7627BFFDE2FCDB2ECFCFB5DEB" box="[693,796,151,177]" country="China" name="Zhejiang" pageId="61" pageNumber="62">Zhejiang</collectingRegion>
,
<collectingRegion id="49EA09D7627BFFDE2E5FB2ECFC9C5DEB" box="[807,891,151,177]" country="China" name="Jiangxi" pageId="61" pageNumber="62">Jiangxi</collectingRegion>
,
<collectingRegion id="49EA09D7627BFFDE2EFFB2ECFC295DEB" box="[903,974,151,177]" country="China" name="Hubei" pageId="61" pageNumber="62">Hubei</collectingRegion>
,
<collectingRegion id="49EA09D7627BFFDE2EA2B2ECFBC05DEB" box="[986,1063,151,177]" country="China" name="Hunan" pageId="61" pageNumber="62">Hunan</collectingRegion>
,
<collectingRegion id="49EA09D7627BFFDE294CB2ECFB765DEB" box="[1076,1169,151,177]" country="China" name="Sichuan" pageId="61" pageNumber="62">Sichuan</collectingRegion>
,
<collectingRegion id="49EA09D7627BFFDE29E5B2ECFAE75DEB" box="[1181,1280,151,177]" country="China" name="Guizhou" pageId="61" pageNumber="62">Guizhou</collectingRegion>
,
<collectingRegion id="49EA09D7627BFFDE2874B2ECFA825DEB" box="[1292,1381,151,177]" country="China" name="Yunnan" pageId="61" pageNumber="62">Yunnan</collectingRegion>
,
<collectingRegion id="49EA09D7627BFFDE2817B2ECFF355D8F" country="China" name="Taiwan)" pageId="61" pageNumber="62">Taiwan)</collectingRegion>
,
<collectingCountry id="F33987A5627BFFDE2DA5B2C0FEA75D8F" box="[221,320,187,213]" name="Vietnam" pageId="61" pageNumber="62">Vietnam</collectingCountry>
.
</paragraph>
<paragraph id="8B91C735627BFFDE2DBFB2A5FDBF5C47" blockId="61.[151,1436,150,285]" pageId="61" pageNumber="62">
<emphasis id="B95A1B27627BFFDE2DBFB2A5FE045DA2" bold="true" box="[199,483,222,248]" pageId="61" pageNumber="62">Geographical variation.</emphasis>
This species ranges throughout southern
<collectingCountry id="F33987A5627BFFDE2EC8B2A4FC105DA3" box="[944,1015,223,249]" name="China" pageId="61" pageNumber="62">China</collectingCountry>
. Its external features vary slightly and geographically in the following aspects:
</paragraph>
<paragraph id="8B91C735627BFFDE2DEFB33EFD955CFD" blockId="61.[151,1436,325,640]" pageId="61" pageNumber="62">
(1) Coloration: Dorsum is gradually darker from north to south. In most parts of
<collectingCountry id="F33987A5627BFFDE2952B33EFB975C05" box="[1066,1136,325,351]" name="China" pageId="61" pageNumber="62">China</collectingCountry>
, the dorsum is always reddish brown. In southern provinces such as
<collectingCountry id="F33987A5627BFFDE2FC9B312FCE45CD9" box="[689,771,361,387]" name="Taiwan" pageId="61" pageNumber="62">Taiwan</collectingCountry>
and Yunnan, the dorsum is often nearly piceous, but dark brown individuals occur there as well.
</paragraph>
<paragraph id="8B91C735627BFFDE2DEFB3CAFD085F49" blockId="61.[151,1436,325,640]" pageId="61" pageNumber="62">
(2) Elytra striae: For specimens from most localities of
<collectingCountry id="F33987A5627BFFDE2E6DB3CAFCBC5C91" box="[789,859,433,459]" name="China" pageId="61" pageNumber="62">China</collectingCountry>
, elytra striae are well incised and intervals slightly convex. However, for specimens from
<collectingRegion id="49EA09D7627BFFDE2FFDB3ADFD3C5CAA" box="[645,731,470,496]" country="China" name="Yunnan" pageId="61" pageNumber="62">Yunnan</collectingRegion>
and
<collectingCountry id="F33987A5627BFFDE2E6EB3AEFC925CB5" box="[790,885,469,495]" name="Vietnam" pageId="61" pageNumber="62">Vietnam</collectingCountry>
(
<figureCitation id="1315DBB0627BFFDE2EFEB3AEFC155CB5" box="[902,1010,469,495]" captionStart="FIGURE 32" captionStartId="59.[151,250,1549,1573]" captionTargetBox="[156,1430,184,1511]" captionTargetId="figure-19@59.[151,1436,181,1525]" captionTargetPageId="59" captionText="FIGURE 32. Habitus of Parena (Bothynoptera) tesari species group. A Holotype of Parena tesari (Jedlička). B Holotype of Parena kataevi Kirschenhofer [=Parena tesari (Jedlička)]. C P. tesari (Jedlička), male from Taiwan. D P. tesari (Jedlička), fe- male from Yunnan. E Holotype of Parena obscura Mateu. F P. obscura Mateu, male from Nepal. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7959003" httpUri="https://zenodo.org/record/7959003/files/figure.png" pageId="61" pageNumber="62">Fig. 32D</figureCitation>
), the elytra striae are so shallow that striae 7 and 8 are hardly visible in the apical half.
</paragraph>
<paragraph id="8B91C735627BFFDE2DEFB066FBF35F25" blockId="61.[151,1436,325,640]" pageId="61" pageNumber="62">
(3) Vertex setae: For specimens from mainland
<collectingCountry id="F33987A5627BFFDE2FC7B066FCE55F6D" box="[703,770,541,567]" name="China" pageId="61" pageNumber="62">China</collectingCountry>
and
<collectingCountry id="F33987A5627BFFDE2E43B066FC7A5F6D" box="[827,925,541,567]" name="Vietnam" pageId="61" pageNumber="62">Vietnam</collectingCountry>
, there is a pair of additional setae on vertex, which are shorter than the primary posterior supraorbital setae and placed posteromedial to them. However, such additional setae are absent from all specimens examined from
<collectingCountry id="F33987A5627BFFDE2EC3B01EFBF65F25" box="[955,1041,613,639]" name="Taiwan" pageId="61" pageNumber="62">Taiwan</collectingCountry>
.
</paragraph>
<paragraph id="8B91C735627BFFDE2DBFB0D3FDAD5E50" blockId="61.[151,1436,680,779]" pageId="61" pageNumber="62">
Based on the above external differences,
<taxonomicName id="4C2EBCB6627BFFDE2FE2B0D2FD115F98" baseAuthorityName="Jedlicka" baseAuthorityYear="1951" box="[666,758,681,706]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="tesari">
<emphasis id="B95A1B27627BFFDE2FE2B0D2FD115F98" box="[666,758,681,706]" italics="true" pageId="61" pageNumber="62">P. tesari</emphasis>
</taxonomicName>
can be divided into three populations:
<collectingCountry id="F33987A5627BFFDE29C8B0D3FAE25F98" box="[1200,1285,680,706]" name="Taiwan" pageId="61" pageNumber="62">Taiwan</collectingCountry>
, S. Yunnan + N.
<collectingCountry id="F33987A5627BFFDE2DC1B0B7FEFC5FBC" box="[185,283,716,742]" name="Vietnam" pageId="61" pageNumber="62">Vietnam</collectingCountry>
, and other localities of
<collectingCountry id="F33987A5627BFFDE2F5AB0B7FD8E5FBC" box="[546,617,716,742]" name="China" pageId="61" pageNumber="62">China</collectingCountry>
. We do not think they should be treated as subspecies because their male genitalia are almost identical (
<figureCitation id="1315DBB0627BFFDE2C91B08BFDD95E50" box="[489,574,752,778]" captionStart="FIGURE 34" captionStartId="60.[151,250,1977,2001]" captionTargetBox="[160,1428,1372,1949]" captionTargetId="figure-208@60.[151,1437,1361,1954]" captionTargetPageId="60" captionText="FIGURE 34. Median lobe of male genitalia for Parena (Bothynoptera) tesari (Jedlička) from different localities, left-lateral (l) and dorsal (d) views: A Yunnan. B Taiwan. C Zhejiang. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7959007" httpUri="https://zenodo.org/record/7959007/files/figure.png" pageId="61" pageNumber="62">Fig. 34</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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