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<document id="F63241B0E3650B534524BB2F73D9E8EA" ID-CLB-Dataset="263342" ID-DOI="10.11646/zootaxa.5286.1.1" ID-GBIF-Dataset="67e7a105-1f62-414a-83d9-f758039eb726" ID-ISSN="1175-5326" ID-Zenodo-Dep="7958937" ID-ZooBank="F9834684-24D3-4795-B5EB-77B451DF856D" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="diego" checkinTime="1684793492880" checkinUser="plazi" docAuthor="Shi, Hongliang &amp; Liang, Hongbin" docDate="2023" docId="038776236234FF972DEFB127FAEF5CF7" docLanguage="en" docName="zootaxa.5286.1.1.pdf" docOrigin="Zootaxa 5286 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5286.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Parena (Parena) circumdata Shibata 1987" docType="treatment" docVersion="8" lastPageNumber="117" masterDocId="FFBE0E5B6246FFE32D78B27BFFE75D5A" masterDocTitle="Taxonomic revision of the genus Parena Motschulsky, 1860 (Coleoptera, Carabidae, Lebiini, Metallicina)" masterLastPageNumber="144" masterPageNumber="1" pageNumber="115" updateTime="1699506634024" updateUser="plazi">
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<mods:title id="049AA10D26A376EE5038A819AF254424">Taxonomic revision of the genus Parena Motschulsky, 1860 (Coleoptera, Carabidae, Lebiini, Metallicina)</mods:title>
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<mods:namePart id="29270BBE3EA4C3B2D6F8506E5267BEBD">Shi, Hongliang</mods:namePart>
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<mods:namePart id="9ACF39A9A1253901CD22DEE26569807F">Liang, Hongbin</mods:namePart>
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<mods:title id="93BAB88EF8C4AF499E69B1F27C3AFFBA">Zootaxa</mods:title>
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<mods:date id="66CB9ADE87E6B6270F32BC1CBA5E34A6">2023</mods:date>
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<mods:number id="E3BEBB497DFAC1E87CD35C027003C9AC">2023-05-18</mods:number>
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<treatment id="038776236234FF972DEFB127FAEF5CF7" ID-DOI="http://doi.org/10.5281/zenodo.7963793" ID-GBIF-Taxon="209629024" ID-Zenodo-Dep="7963793" LSID="urn:lsid:plazi:treatment:038776236234FF972DEFB127FAEF5CF7" httpUri="http://treatment.plazi.org/id/038776236234FF972DEFB127FAEF5CF7" lastPageId="116" lastPageNumber="117" pageId="114" pageNumber="115">
<subSubSection id="C33494BE6234FF912DEFB127FD3B5E2C" box="[151,732,859,886]" pageId="114" pageNumber="115" type="nomenclature">
<paragraph id="8B91C7356234FF912DEFB127FD3B5E2C" blockId="114.[151,1108,859,922]" box="[151,732,859,886]" pageId="114" pageNumber="115">
<emphasis id="B95A1B276234FF912DEFB127FD3B5E2C" bold="true" box="[151,732,859,886]" pageId="114" pageNumber="115">
[34]
<taxonomicName id="4C2EBCB66234FF912DB7B127FD3B5E2C" ID-CoL="4DKP7" authority="Shibata, 1987" authorityName="Shibata" authorityYear="1987" box="[207,732,859,886]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="114" pageNumber="115" phylum="Arthropoda" rank="species" species="circumdata" subGenus="Parena">
<emphasis id="B95A1B276234FF912DB7B127FEC05E2C" bold="true" box="[207,295,860,886]" italics="true" pageId="114" pageNumber="115">Parena</emphasis>
(
<emphasis id="B95A1B276234FF912C40B127FE775E2C" bold="true" box="[312,400,860,886]" italics="true" pageId="114" pageNumber="115">Parena</emphasis>
)
<emphasis id="B95A1B276234FF912CDAB120FDCC5E2C" bold="true" box="[418,555,859,886]" italics="true" pageId="114" pageNumber="115">circumdata</emphasis>
<bibRefCitation id="EFBFBAC46234FF912F4AB127FD3B5E2C" author="Shibata, T." box="[562,732,860,886]" pageId="114" pageNumber="115" pagination="63 - 67" refId="ref112910" refString="Shibata, T. (1987) Three new species and a new record of Carabidae from Taiwan (Truncatipennes) (Coleoptera). Entomological Review of Japan, 42 (1), 63 - 67." type="journal article" year="1987">Shibata, 1987</bibRefCitation>
</taxonomicName>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C33494BE6234FF912DEFB1FBFBB35EC0" box="[151,1108,896,922]" pageId="114" pageNumber="115" type="description">
<paragraph id="8B91C7356234FF912DEFB1FBFBB35EC0" blockId="114.[151,1108,859,922]" box="[151,1108,896,922]" pageId="114" pageNumber="115">
Habitus:
<figureCitation id="1315DBB06234FF912D85B1FBFE435EC0" box="[253,420,896,922]" captionStart="FIGURE 69" captionStartId="115.[151,250,1672,1696]" captionTargetBox="[165,1413,170,1629]" captionTargetId="figure-19@115.[164,1423,166,1639]" captionTargetPageId="115" captionText="FIGURE 69. Habitus of Parena (Parena) latecincta species group. A P. circumdata Shibata, male from Taiwan. B P. circumdata Shibata, female from India. C P. monticola Shibata, male from Jiangxi. D P. monticola Shibata, male from Sichuan. E Holotype of Parena pendleburyi Andrewes. F P. pendleburyi Andrewes, female from Borneo. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7959082" httpUri="https://zenodo.org/record/7959082/files/figure.png" pageId="114" pageNumber="115">Figs 69A, 69B</figureCitation>
. Male genitalia:
<figureCitation id="1315DBB06234FF912F27B1FBFD565EC0" box="[607,689,896,922]" captionStart="FIGURE 70" captionStartId="116.[151,250,1154,1178]" captionTargetBox="[164,1422,448,1123]" captionTargetId="figure-167@116.[152,1436,438,1131]" captionTargetPageId="116" captionText="FIGURE 70. Median lobe of male genitalia for Parena (Parena) circumdata Shibata, right-lateral (r), ventral (v), left-lateral (l), and dorsal (d) views, Guangdong. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7959084" httpUri="https://zenodo.org/record/7959084/files/figure.png" pageId="114" pageNumber="115">Fig. 70</figureCitation>
. Gonocoxites of ovipositor:
<figureCitation id="1315DBB06234FF912E88B1FBFBB65EC0" box="[1008,1105,896,922]" captionStart="FIGURE 11" captionStartId="18.[151,250,150,174]" captionTargetId="figure-19@17.[155,1432,181,1971]" captionTargetPageId="17" captionText="FIGURE 11. Gonocoxites (left side) of ovipositor for Parena spp., ventral side (dorsal side for fig. Q). A–J subgenus Bothynoptera: A P. (Bothynoptera) heteronycha sp.n., Laos, paratype. B P. (Bothynoptera) dorsigera (Schaum), Shaanxi. C P. (Bothynoptera) kurosai Habu, Guangdong. D P. (Bothynoptera) tesari (Jedlička), Taiwan. E P. (Bothynoptera) taiwana Hua, Taiwan. F P. (Bothynoptera) tripunctata (Bates), Shaanxi. G P. (Bothynoptera) malaisei (Andrewes), Taiwan. H P. (Bothynoptera) quadrisignata Mateu, Yunnan. I P. (Bothynoptera) emarginata sp. n., holotype. J P. (Bothynoptera) triguttata sp. n., Sichuan, paratype. K–X subgenus Parena: K P. (Parena) bicolor Motschulsky, Sulawesi. L P. (Parena) rubripicta Andrewes, Yunnan. M P. (Parena) fasciata (Chaudoir), Taiwan. N P. (Parena) nigrolineata (Chaudoir), Taiwan. O P. (Parena) picipes sp. n., Borneo, paratype. P P. (Parena) andrewesi Jedlička, Philippine. Q P. (Parena) politissima (Chaudoir), Vanuatu. R P. (Parena) latecincta (Bates), Guizhou. S P. (Parena) circumdata Shibata, Taiwan. T P. (Parena) pendleburyi Andrewes, Borneo. U P. (Parena) fulva sp. n., Zimbabwe, paratype. V P. (Parena) madagascariensis (Alluaud), Madagascar. W P. (Parena) ruficornis sp.n., CentralAfrica, paratype. X P. (Parena) africana (Alluaud), Uganda. Y–γ subgenus Crossoglossa: Y P. (Crossoglossa) cavipennis (Bates), Taiwan. Z P. (Crossoglossa) laesipennis (Bates), Yunnan. α P. (Crossoglossa) testacea (Chaudoir), Zhejiang. β P. (Crossoglossa) sulawesiensis Kirschenhofer, Sulawesi. γ P. (Crossoglossa) sciakyi sp. n., holotype. Scale bar = 0.2mm." figureDoi="http://doi.org/10.5281/zenodo.7963954" httpUri="https://zenodo.org/record/7963954/files/figure.png" pageId="114" pageNumber="115">Fig. 11S</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33494BE6234FF912DEFB1C4FD435EAC" pageId="114" pageNumber="115" type="reference_group">
<paragraph id="8B91C7356234FF912DEFB1C4FD435EAC" blockId="114.[151,1436,959,1014]" pageId="114" pageNumber="115">
<bibRefCitation id="EFBFBAC46234FF912DEFB1C4FEAC5E8D" author="Shibata, T." box="[151,331,959,983]" pageId="114" pageNumber="115" pagination="63 - 67" refId="ref112910" refString="Shibata, T. (1987) Three new species and a new record of Carabidae from Taiwan (Truncatipennes) (Coleoptera). Entomological Review of Japan, 42 (1), 63 - 67." type="journal article" year="1987">Shibata, 1987: 64</bibRefCitation>
(type locality:
<materialsCitation id="3B46CD686234FF912C9FB1C4FA7B5E8D" ID-GBIF-Occurrence="4115871593" box="[487,1436,959,984]" country="Taiwan" location="Kashihara City Museum of Insects" pageId="114" pageNumber="115" specimenCount="1" stateProvince="Taoyuan" typeStatus="holotype">
Taiwan,
<collectingRegion id="49EA09D76234FF912F46B1BBFD7F5E82" box="[574,664,960,984]" country="Taiwan" name="Taoyuan" pageId="114" pageNumber="115">Taoyuan</collectingRegion>
;
<typeStatus id="549579976234FF912FDBB1C4FD1B5E8D" box="[675,764,959,983]" pageId="114" pageNumber="115" type="holotype">holotype</typeStatus>
in KCMI:
<location id="8EF191EE6234FF912E16B1C4FB335E8D" LSID="urn:lsid:plazi:treatment:038776236234FF972DEFB127FAEF5CF7:8EF191EE6234FF912E16B1C4FB335E8D" box="[878,1236,959,983]" country="Taiwan" name="Kashihara City Museum of Insects" pageId="114" pageNumber="115" stateProvince="Taoyuan">Kashihara City Museum of Insects</location>
, Kashihara,
<collectingCountry id="F33987A56234FF91282BB1C4FA765E8D" box="[1363,1425,959,983]" name="Japan" pageId="114" pageNumber="115">Japan</collectingCountry>
);
</materialsCitation>
<bibRefCitation id="EFBFBAC46234FF912DBFB1A5FE7A5EAC" author="Xie, W. P. &amp; Yu, P. Y." box="[199,413,990,1014]" pageId="114" pageNumber="115" pagination="185 - 195" refId="ref113326" refString="Xie, W. P. &amp; Yu, P. Y. (1993) On the Chinese species of Parena Motschulsky (Coleoptera: Carabidae). Sinozoologia, 10, 185 - 195." type="journal article" year="1993">Xie &amp; Yu, 1993: 189</bibRefCitation>
;
<bibRefCitation id="EFBFBAC46234FF912CDFB1A5FD475EAC" author="Kirschenhofer, E." box="[423,672,990,1014]" pageId="114" pageNumber="115" pagination="87 - 103" refId="ref111612" refString="Kirschenhofer, E. (2006) Neue Arten der Gattung Parena Motschulsky aus Japan, China Und Indonesien (Sulawesi) (Coleoptera: Carabidae). Mitteilungen des Internationalen Entomologischen Vereins E. V. Frankfurt, 31 (3 - 4), 87 - 103." type="journal article" year="2006">Kirschenhofer, 2006: 88</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33494BE6234FF912DEFB65FFD3A58D8" pageId="114" pageNumber="115" type="materials_examined">
<paragraph id="8B91C7356234FF912DEFB65FFD3A58D8" blockId="114.[151,1436,1060,1951]" pageId="114" pageNumber="115">
<emphasis id="B95A1B276234FF912DEFB65FFE095964" bold="true" box="[151,494,1060,1086]" pageId="114" pageNumber="115">Non-type material examined</emphasis>
.
<materialsCitation id="3B46CD686234FF912C86B65FFB905938" ID-GBIF-Occurrence="4115871410" collectingDate="2008-07-25" country="China" elevation="354" latitude="24.7232" location="Yang Zhuo" longLatPrecision="1" longitude="114.2564" municipality="Shixing county" pageId="114" pageNumber="115" specimenCount="1" specimenCount-male="1" stateProvince="Guangdong">
<emphasis id="B95A1B276234FF912C86B65FFDB75964" bold="true" box="[510,592,1060,1086]" pageId="114" pageNumber="115">
<collectingCountry id="F33987A56234FF912C86B65FFDAC5964" box="[510,587,1060,1086]" name="China" pageId="114" pageNumber="115">China</collectingCountry>
:
</emphasis>
<specimenCount id="9D280CBC6234FF912F21B65FFD425964" box="[601,677,1060,1086]" pageId="114" pageNumber="115" type="male">1 male</specimenCount>
(IZAS), &quot;
<collectingRegion id="49EA09D76234FF912E61B65FFC785964" box="[793,927,1060,1086]" country="China" name="Guangdong" pageId="114" pageNumber="115">Guangdong</collectingRegion>
,
<collectingMunicipality id="6BF55D4F6234FF912ED3B65FFBBF5965" box="[939,1112,1060,1087]" pageId="114" pageNumber="115">Shixing county</collectingMunicipality>
, Chebaling, Zhangdongshui,
<geoCoordinate id="EE1AA1F26234FF912DEFB633FEC45938" box="[151,291,1096,1122]" degrees="24.72320" direction="north" orientation="latitude" pageId="114" pageNumber="115" precision="1" value="24.7232">N24.72320°</geoCoordinate>
,
<geoCoordinate id="EE1AA1F26234FF912C55B633FE245939" box="[301,451,1096,1123]" degrees="114.25640" direction="east" orientation="longitude" pageId="114" pageNumber="115" precision="1" value="114.2564">E114.25640°</geoCoordinate>
,
<quantity id="4CD66AD06234FF912CB5B633FDF15939" box="[461,534,1096,1123]" metricMagnitude="2" metricUnit="m" metricValue="3.54" pageId="114" pageNumber="115" unit="m" value="354.0">
<elevation id="000320066234FF912CB5B633FDF15939" box="[461,534,1096,1123]" metricMagnitude="2" metricUnit="m" metricValue="3.54" pageId="114" pageNumber="115" unit="m" value="354.0">354 m</elevation>
</quantity>
,
<date id="FF90E1F56234FF912F59B633FD4B5939" box="[545,684,1096,1123]" pageId="114" pageNumber="115" value="2008-07-25">
<collectingDate id="EFD4181D6234FF912F59B633FD4B5939" box="[545,684,1096,1123]" pageId="114" pageNumber="115" value="2008-07-25">2008.VII.25</collectingDate>
</date>
, on vegetation,
<location id="8EF191EE6234FF912E24B632FC3F5938" LSID="urn:lsid:plazi:treatment:038776236234FF972DEFB127FAEF5CF7:8EF191EE6234FF912E24B632FC3F5938" box="[860,984,1096,1123]" country="China" latitude="24.7232" longLatPrecision="1" longitude="114.2564" municipality="Shixing county" name="Yang Zhuo" pageId="114" pageNumber="115" stateProvince="Guangdong">Yang Zhuo</location>
lgt.&quot; &lt;
<figureCitation id="1315DBB06234FF91295DB633FB905938" box="[1061,1143,1096,1122]" captionStart="FIGURE 70" captionStartId="116.[151,250,1154,1178]" captionTargetBox="[164,1422,448,1123]" captionTargetId="figure-167@116.[152,1436,438,1131]" captionTargetPageId="116" captionText="FIGURE 70. Median lobe of male genitalia for Parena (Parena) circumdata Shibata, right-lateral (r), ventral (v), left-lateral (l), and dorsal (d) views, Guangdong. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7959084" httpUri="https://zenodo.org/record/7959084/files/figure.png" pageId="114" pageNumber="115">Fig. 70</figureCitation>
</materialsCitation>
&gt;.
<materialsCitation id="3B46CD686234FF9129E9B633FCDA59DC" ID-GBIF-Occurrence="4115871408" collectingDate="1994-07-09" country="China" county="Hsinchu county" location="Dalu forest" municipality="Wufeng" pageId="114" pageNumber="115" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D280CBC6234FF9129E9B633FB155938" box="[1169,1266,1096,1122]" pageId="114" pageNumber="115" type="female">1 female</specimenCount>
(CCCC), &quot;Taiwan,
<collectingCounty id="62F0BFB96234FF912DABB617FE6459DD" box="[211,387,1132,1159]" pageId="114" pageNumber="115">Hsinchu county</collectingCounty>
,
<collectingMunicipality id="6BF55D4F6234FF912CF5B617FE0E59DC" box="[397,489,1132,1158]" pageId="114" pageNumber="115">Wufeng</collectingMunicipality>
,
<location id="8EF191EE6234FF912C8BB617FD8859DC" LSID="urn:lsid:plazi:treatment:038776236234FF972DEFB127FAEF5CF7:8EF191EE6234FF912C8BB617FD8859DC" box="[499,623,1132,1158]" country="China" county="Hsinchu county" municipality="Wufeng" name="Dalu forest" pageId="114" pageNumber="115">Dalu forest</location>
road,
<date id="FF90E1F56234FF912FCCB617FCD359DC" box="[692,820,1132,1158]" pageId="114" pageNumber="115" value="1994-07-09">
<collectingDate id="EFD4181D6234FF912FCCB617FCD359DC" box="[692,820,1132,1158]" pageId="114" pageNumber="115" value="1994-07-09">1994.VII.9</collectingDate>
</date>
&quot;
</materialsCitation>
.
<materialsCitation id="3B46CD686234FF912E3FB617FD8B59F0" ID-GBIF-Occurrence="4115871455" collectingDate="2003-06-15" country="China" location="Dalu forest" municipality="Hsinchu county" pageId="114" pageNumber="115" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D280CBC6234FF912E3FB617FC4059DC" box="[839,935,1132,1158]" pageId="114" pageNumber="115" type="female">1 female</specimenCount>
(CCCC), &quot;Taiwan,
<collectingMunicipality id="6BF55D4F6234FF9129FAB617FAD559DD" box="[1154,1330,1132,1159]" pageId="114" pageNumber="115">Hsinchu county</collectingMunicipality>
, Wufeng,
<location id="8EF191EE6234FF912DEFB6EBFEF159F0" LSID="urn:lsid:plazi:treatment:038776236234FF972DEFB127FAEF5CF7:8EF191EE6234FF912DEFB6EBFEF159F0" box="[151,278,1168,1194]" country="China" municipality="Hsinchu county" name="Dalu forest" pageId="114" pageNumber="115">Dalu forest</location>
road,
<date id="FF90E1F56234FF912C18B6EBFE0459F1" box="[352,483,1168,1195]" pageId="114" pageNumber="115" value="2003-06-15">
<collectingDate id="EFD4181D6234FF912C18B6EBFE0459F1" box="[352,483,1168,1195]" pageId="114" pageNumber="115" value="2003-06-15">2003.VI.15</collectingDate>
</date>
&quot; &lt;
<figureCitation id="1315DBB06234FF912F7CB6EBFD8B59F0" box="[516,620,1168,1194]" captionStart="FIGURE 69" captionStartId="115.[151,250,1672,1696]" captionTargetBox="[165,1413,170,1629]" captionTargetId="figure-19@115.[164,1423,166,1639]" captionTargetPageId="115" captionText="FIGURE 69. Habitus of Parena (Parena) latecincta species group. A P. circumdata Shibata, male from Taiwan. B P. circumdata Shibata, female from India. C P. monticola Shibata, male from Jiangxi. D P. monticola Shibata, male from Sichuan. E Holotype of Parena pendleburyi Andrewes. F P. pendleburyi Andrewes, female from Borneo. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7959082" httpUri="https://zenodo.org/record/7959082/files/figure.png" pageId="114" pageNumber="115">Fig. 69A</figureCitation>
</materialsCitation>
&gt;.
<materialsCitation id="3B46CD686234FF912FF0B6EBFEAA5994" ID-GBIF-Occurrence="4115871495" collectingDate="1994-04-20" country="China" county="Taipei county" location="Chen Changchin" municipality="Fushan" pageId="114" pageNumber="115" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D280CBC6234FF912FF0B6EBFD0C59F0" box="[648,747,1168,1194]" pageId="114" pageNumber="115" type="female">1 female</specimenCount>
(CCCC), &quot;Taiwan,
<collectingCounty id="62F0BFB96234FF912EB5B6EBFB8F59F1" box="[973,1128,1168,1195]" pageId="114" pageNumber="115">Taipei county</collectingCounty>
,
<collectingMunicipality id="6BF55D4F6234FF91290DB6EBFB2E59F0" box="[1141,1225,1168,1194]" pageId="114" pageNumber="115">Fushan</collectingMunicipality>
,
<date id="FF90E1F56234FF9129AEB6EBFAB459F0" box="[1238,1363,1168,1194]" pageId="114" pageNumber="115" value="1994-04-20">
<collectingDate id="EFD4181D6234FF9129AEB6EBFAB459F0" box="[1238,1363,1168,1194]" pageId="114" pageNumber="115" value="1994-04-20">1994.IV.20</collectingDate>
</date>
,
<location id="8EF191EE6234FF912819B6EBFEF75994" LSID="urn:lsid:plazi:treatment:038776236234FF972DEFB127FAEF5CF7:8EF191EE6234FF912819B6EBFEF75994" country="China" county="Taipei county" municipality="Fushan" name="Chen Changchin" pageId="114" pageNumber="115">Chen Changchin</location>
lgt.&quot;
</materialsCitation>
.
<materialsCitation id="3B46CD686234FF912C23B6CFFAC85994" ID-GBIF-Occurrence="4115871405" box="[347,1327,1204,1231]" country="China" county="Nantou county" location="Chinching" municipality="Ren'ai" pageId="114" pageNumber="115" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D280CBC6234FF912C23B6CFFE585994" box="[347,447,1204,1230]" pageId="114" pageNumber="115" type="female">1 female</specimenCount>
(CCCC), &quot;Taiwan,
<collectingCounty id="62F0BFB96234FF912FDDB6CEFCA95995" box="[677,846,1205,1231]" pageId="114" pageNumber="115">Nantou county</collectingCounty>
,
<collectingMunicipality id="6BF55D4F6234FF912E24B6CFFC425994" box="[860,933,1204,1230]" pageId="114" pageNumber="115">Ren'ai</collectingMunicipality>
,
<location id="8EF191EE6234FF912ECBB6CFFBCC5994" LSID="urn:lsid:plazi:treatment:038776236234FF972DEFB127FAEF5CF7:8EF191EE6234FF912ECBB6CFFBCC5994" box="[947,1067,1204,1230]" country="China" county="Nantou county" municipality="Ren'ai" name="Chinching" pageId="114" pageNumber="115">Chinching</location>
, 1995.VIII&quot; &lt;
<figureCitation id="1315DBB06234FF9129B5B6CFFAC85994" box="[1229,1327,1204,1230]" captionStart="FIGURE 11" captionStartId="18.[151,250,150,174]" captionTargetId="figure-19@17.[155,1432,181,1971]" captionTargetPageId="17" captionText="FIGURE 11. Gonocoxites (left side) of ovipositor for Parena spp., ventral side (dorsal side for fig. Q). A–J subgenus Bothynoptera: A P. (Bothynoptera) heteronycha sp.n., Laos, paratype. B P. (Bothynoptera) dorsigera (Schaum), Shaanxi. C P. (Bothynoptera) kurosai Habu, Guangdong. D P. (Bothynoptera) tesari (Jedlička), Taiwan. E P. (Bothynoptera) taiwana Hua, Taiwan. F P. (Bothynoptera) tripunctata (Bates), Shaanxi. G P. (Bothynoptera) malaisei (Andrewes), Taiwan. H P. (Bothynoptera) quadrisignata Mateu, Yunnan. I P. (Bothynoptera) emarginata sp. n., holotype. J P. (Bothynoptera) triguttata sp. n., Sichuan, paratype. K–X subgenus Parena: K P. (Parena) bicolor Motschulsky, Sulawesi. L P. (Parena) rubripicta Andrewes, Yunnan. M P. (Parena) fasciata (Chaudoir), Taiwan. N P. (Parena) nigrolineata (Chaudoir), Taiwan. O P. (Parena) picipes sp. n., Borneo, paratype. P P. (Parena) andrewesi Jedlička, Philippine. Q P. (Parena) politissima (Chaudoir), Vanuatu. R P. (Parena) latecincta (Bates), Guizhou. S P. (Parena) circumdata Shibata, Taiwan. T P. (Parena) pendleburyi Andrewes, Borneo. U P. (Parena) fulva sp. n., Zimbabwe, paratype. V P. (Parena) madagascariensis (Alluaud), Madagascar. W P. (Parena) ruficornis sp.n., CentralAfrica, paratype. X P. (Parena) africana (Alluaud), Uganda. Y–γ subgenus Crossoglossa: Y P. (Crossoglossa) cavipennis (Bates), Taiwan. Z P. (Crossoglossa) laesipennis (Bates), Yunnan. α P. (Crossoglossa) testacea (Chaudoir), Zhejiang. β P. (Crossoglossa) sulawesiensis Kirschenhofer, Sulawesi. γ P. (Crossoglossa) sciakyi sp. n., holotype. Scale bar = 0.2mm." figureDoi="http://doi.org/10.5281/zenodo.7963954" httpUri="https://zenodo.org/record/7963954/files/figure.png" pageId="114" pageNumber="115">Fig. 11S</figureCitation>
</materialsCitation>
&gt;.
<materialsCitation id="3B46CD686234FF912836B6CFFBFC59A8" ID-GBIF-Occurrence="4115871461" collectingDate="1995-03-18" country="China" county="Taipei county" location="Shizitou" municipality="Sanhsia" pageId="114" pageNumber="115" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D280CBC6234FF912836B6CFFA7B5994" box="[1358,1436,1204,1230]" pageId="114" pageNumber="115" type="male">1 male</specimenCount>
(CCCC), &quot;Taiwan,
<collectingCounty id="62F0BFB96234FF912C13B6A3FDE459A9" box="[363,515,1240,1267]" pageId="114" pageNumber="115">Taipei county</collectingCounty>
,
<collectingMunicipality id="6BF55D4F6234FF912F75B6A3FD8059A8" box="[525,615,1240,1266]" pageId="114" pageNumber="115">Sanhsia</collectingMunicipality>
,
<location id="8EF191EE6234FF912F09B6A3FD3659A8" LSID="urn:lsid:plazi:treatment:038776236234FF972DEFB127FAEF5CF7:8EF191EE6234FF912F09B6A3FD3659A8" box="[625,721,1240,1266]" country="China" county="Taipei county" municipality="Sanhsia" name="Shizitou" pageId="114" pageNumber="115">Shizitou</location>
,
<date id="FF90E1F56234FF912FA3B6A3FCBB59A9" box="[731,860,1240,1267]" pageId="114" pageNumber="115" value="1995-03-18">
<collectingDate id="EFD4181D6234FF912FA3B6A3FCBB59A9" box="[731,860,1240,1267]" pageId="114" pageNumber="115" value="1995-03-18">1995.III.18</collectingDate>
</date>
,
<location id="8EF191EE6234FF912E1EB6A2FC0359A8" LSID="urn:lsid:plazi:treatment:038776236234FF972DEFB127FAEF5CF7:8EF191EE6234FF912E1EB6A2FC0359A8" box="[870,996,1240,1267]" country="China" county="Taipei county" municipality="Sanhsia" name="Lo Chinchi" pageId="114" pageNumber="115">Lo Chinchi</location>
lgt.&quot;
</materialsCitation>
.
<materialsCitation id="3B46CD686234FF91295EB6A3FC6C584C" ID-GBIF-Occurrence="4115871559" collectingDate="1999-05-01" country="India" county="Nantou county" location="Chen Changchin" municipality="Guandaoshan" pageId="114" pageNumber="115" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D280CBC6234FF91295EB6A3FB8859A8" box="[1062,1135,1240,1266]" pageId="114" pageNumber="115" type="male">1 male</specimenCount>
(CCCC), &quot;Taiwan,
<collectingCounty id="62F0BFB96234FF912832B6A2FF03584D" pageId="114" pageNumber="115">Nantou county</collectingCounty>
, Puli,
<collectingMunicipality id="6BF55D4F6234FF912C55B687FE2F584C" box="[301,456,1276,1302]" pageId="114" pageNumber="115">Guandaoshan</collectingMunicipality>
,
<date id="FF90E1F56234FF912CACB687FDDE584C" box="[468,569,1276,1303]" pageId="114" pageNumber="115" value="1999-05-01">
<collectingDate id="EFD4181D6234FF912CACB687FDDE584C" box="[468,569,1276,1303]" pageId="114" pageNumber="115" value="1999-05-01">1999.V.1</collectingDate>
</date>
,
<location id="8EF191EE6234FF912F30B687FCE3584C" LSID="urn:lsid:plazi:treatment:038776236234FF972DEFB127FAEF5CF7:8EF191EE6234FF912F30B687FCE3584C" box="[584,772,1276,1302]" country="India" county="Nantou county" municipality="Guandaoshan" name="Chen Changchin" pageId="114" pageNumber="115">Chen Changchin</location>
lgt.&quot;.
<emphasis id="B95A1B276234FF912E33B687FC6C584C" bold="true" box="[843,907,1276,1302]" pageId="114" pageNumber="115">
<collectingCountry id="F33987A56234FF912E33B687FC6C584C" box="[843,907,1276,1302]" name="India" pageId="114" pageNumber="115">India</collectingCountry>
</emphasis>
</materialsCitation>
:
<materialsCitation id="3B46CD686234FF912EE3B687FD5D5860" ID-GBIF-Occurrence="4115871467" country="India" elevation="800" location="Chairy Cow." pageId="114" pageNumber="115" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D280CBC6234FF912EE3B687FC1A584C" box="[923,1021,1276,1302]" pageId="114" pageNumber="115" type="female">1 female</specimenCount>
(NHMB), &quot;India Darjeeling D., C.J. Rai&quot;, &quot;
<location id="8EF191EE6234FF912DA6B75BFE895860" LSID="urn:lsid:plazi:treatment:038776236234FF972DEFB127FAEF5CF7:8EF191EE6234FF912DA6B75BFE895860" box="[222,366,1312,1338]" country="India" name="Chairy Cow." pageId="114" pageNumber="115">Chairy Cow.</location>
26.VIII.85
<quantity id="4CD66AD06234FF912C88B75BFDDF5860" box="[496,568,1312,1338]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="114" pageNumber="115" unit="m" value="800.0">
<elevation id="000320066234FF912C88B75BFDDF5860" box="[496,568,1312,1338]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="114" pageNumber="115" unit="m" value="800.0">800 m</elevation>
</quantity>
&quot; &lt;
<figureCitation id="1315DBB06234FF912F2EB75BFD5D5860" box="[598,698,1312,1338]" captionStart="FIGURE 69" captionStartId="115.[151,250,1672,1696]" captionTargetBox="[165,1413,170,1629]" captionTargetId="figure-19@115.[164,1423,166,1639]" captionTargetPageId="115" captionText="FIGURE 69. Habitus of Parena (Parena) latecincta species group. A P. circumdata Shibata, male from Taiwan. B P. circumdata Shibata, female from India. C P. monticola Shibata, male from Jiangxi. D P. monticola Shibata, male from Sichuan. E Holotype of Parena pendleburyi Andrewes. F P. pendleburyi Andrewes, female from Borneo. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7959082" httpUri="https://zenodo.org/record/7959082/files/figure.png" pageId="114" pageNumber="115">Fig. 69B</figureCitation>
</materialsCitation>
&gt;.
<materialsCitation id="3B46CD686234FF912FABB75BFE3F5805" ID-GBIF-Occurrence="4115871536" collectingDate="1985-06-29" country="India" location="Lower Tanake" municipality="Kalimpong" pageId="114" pageNumber="115" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D280CBC6234FF912FABB75BFCD45860" box="[723,819,1312,1338]" pageId="114" pageNumber="115" type="female">1 female</specimenCount>
(NHMB), &quot;India Darjeeling D., C.J. Rai&quot;, &quot;
<collectingMunicipality id="6BF55D4F6234FF91286BB75BFA7F5860" box="[1299,1432,1312,1338]" pageId="114" pageNumber="115">Kalimpong</collectingMunicipality>
,
<location id="8EF191EE6234FF912DEFB73EFEDB5804" LSID="urn:lsid:plazi:treatment:038776236234FF972DEFB127FAEF5CF7:8EF191EE6234FF912DEFB73EFEDB5804" box="[151,316,1348,1375]" country="India" municipality="Kalimpong" name="Lower Tanake" pageId="114" pageNumber="115">Lower Tanake</location>
,
<date id="FF90E1F56234FF912C31B73FFE295805" box="[329,462,1348,1375]" pageId="114" pageNumber="115" value="1985-06-29">
<collectingDate id="EFD4181D6234FF912C31B73FFE295805" box="[329,462,1348,1375]" pageId="114" pageNumber="115" value="1985-06-29">29.VI.1985</collectingDate>
</date>
&quot;
</materialsCitation>
.
<materialsCitation id="3B46CD686234FF912C9CB73FFD3E58D8" ID-GBIF-Occurrence="4115871515" country="Vietnam" county="Darjeeling Distr." elevation="2200" location="Tonkin" municipality="Hoah" pageId="114" pageNumber="115" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D280CBC6234FF912C9CB73FFDA05804" box="[484,583,1348,1374]" pageId="114" pageNumber="115" type="female">1 female</specimenCount>
(NHMB), &quot;
<collectingCounty id="62F0BFB96234FF912FB6B73FFC685804" box="[718,911,1348,1374]" pageId="114" pageNumber="115">Darjeeling Distr.</collectingCounty>
India, Bhakta B.&quot;, &quot;Laua
<quantity id="4CD66AD06234FF9129C3B73FFAF35805" box="[1211,1300,1348,1375]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="114" pageNumber="115" unit="m" value="2200.0">
<elevation id="000320066234FF9129C3B73FFAF35805" box="[1211,1300,1348,1375]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="114" pageNumber="115" unit="m" value="2200.0">2200 m</elevation>
</quantity>
, 7.VIII.78&quot;.
<emphasis id="B95A1B276234FF912DEFB713FF1A58D8" bold="true" box="[151,253,1384,1410]" pageId="114" pageNumber="115">
<collectingCountry id="F33987A56234FF912DEFB713FF1A58D8" box="[151,253,1384,1410]" name="Vietnam" pageId="114" pageNumber="115">Vietnam</collectingCountry>
</emphasis>
: 1 ex (MNHN), &quot;env.
<collectingMunicipality id="6BF55D4F6234FF912C80B713FDD258D8" box="[504,565,1384,1410]" pageId="114" pageNumber="115">Hoah</collectingMunicipality>
binh,
<location id="8EF191EE6234FF912F03B713FD3658D8" LSID="urn:lsid:plazi:treatment:038776236234FF972DEFB127FAEF5CF7:8EF191EE6234FF912F03B713FD3658D8" box="[635,721,1384,1410]" country="Vietnam" county="Darjeeling Distr." municipality="Hoah" name="Tonkin" pageId="114" pageNumber="115">Tonkin</location>
&quot;
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33494BE6234FF912DBFB7F7FC9E5B00" pageId="114" pageNumber="115" type="diagnosis">
<paragraph id="8B91C7356234FF912DBFB7F7FC9E5B00" blockId="114.[151,1436,1060,1951]" pageId="114" pageNumber="115">
<emphasis id="B95A1B276234FF912DBFB7F7FE8158FC" bold="true" box="[199,358,1420,1446]" pageId="114" pageNumber="115">Comparisons</emphasis>
. This species differs from other species of the genus by the combination of: elytra with metallic green lateral stripes, elytra disc reddish brown with metallic hue, postgenae without suborbital setae, and males with rudimentary adhesive setae on mesotarsomere 1. It is most similar to
<taxonomicName id="4C2EBCB66234FF912ED8B7AEFBCE58B4" baseAuthorityName="Bates" baseAuthorityYear="1873" box="[928,1065,1492,1518]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="114" pageNumber="115" phylum="Arthropoda" rank="species" species="latecincta">
<emphasis id="B95A1B276234FF912ED8B7AEFBCE58B4" box="[928,1065,1492,1518]" italics="true" pageId="114" pageNumber="115">P. latecincta</emphasis>
</taxonomicName>
, with both having metallic green lateral stripes on the elytra. However,
<taxonomicName id="4C2EBCB66234FF912F34B782FD0F5B48" authorityName="Shibata" authorityYear="1987" box="[588,744,1528,1554]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="114" pageNumber="115" phylum="Arthropoda" rank="species" species="circumdata">
<emphasis id="B95A1B276234FF912F34B782FD0F5B48" box="[588,744,1528,1554]" italics="true" pageId="114" pageNumber="115">P. circumdata</emphasis>
</taxonomicName>
is readily differentiated from the latter species in having the elytra lateral stripes narrower, only extended medially to interval
<quantity id="4CD66AD06234FF912E01B466FC425B6C" box="[889,933,1564,1591]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="114" pageNumber="115" unit="in" value="7.0">7 in</quantity>
the middle, elytra disc with distinct metallic hue, and males with adhesive setae ventrally on mesotarsomere 1.
</paragraph>
</subSubSection>
<subSubSection id="C33494BE6234FF972DBFB41FFABE5CD3" lastPageId="116" lastPageNumber="117" pageId="114" pageNumber="115" type="description">
<paragraph id="8B91C7356234FF972DBFB41FFABE5CD3" blockId="114.[151,1436,1060,1951]" lastBlockId="116.[151,1437,151,429]" lastPageId="116" lastPageNumber="117" pageId="114" pageNumber="115">
<emphasis id="B95A1B276234FF912DBFB41FFEB65B24" bold="true" box="[199,337,1636,1662]" pageId="114" pageNumber="115">Description</emphasis>
. Body length 10.0–11.0 mm. Head and pronotum dark brown, with faint metallic green hue; mouthparts yellowish brown, apices of mandibles dark brown; basal four antennomeres reddish brown, apical ones distinctly darker; elytra disc reddish brown, with distinct metallic purple hue, with narrow metallic green lateral stripes, extended medially over intervals 8 and 9 and half of interval 7 at middle, stripes narrowly continuous at base, nearly extended to sutural angles at apex, interval 1 completely brown; elytral lateral margins and epipleura brown with strong metallic hue; legs dark brown, apices of femora piceous with metallic green hue; venter dark brown. Vertex finely and densely punctate; postgenae without suborbital setae; antennae barely extended to pronotal base; labrum quadrate, apex nearly straight; mentum with a pair of minute median setae, barely visible in some specimens. Pro notum sub-cordate, PW/PL = 1.35–1.43, usually slightly wider than head, PW/HW = 0.99–1.04; widest at anterior third, strongly narrowed to base, more or less sinuate before posterior angles; posterior angles widely obtuse; disc usually with sparse punctures. Elytra slightly dilated to apex, with shallow isodiametric microsculpture; striae well incised, with fine puncture rows; intervals sparsely punctate; disc with large shallow depressions near basal third of intervals 3 to 6. Males with uniseriate adhesive setae near apex of mesotarsomere 1, with biseriate setae on apical half of mesotarsomere 2 and almost full length of mesotarsomere 3. Apex of abdominal sternite VII nearly straight in males. Median lobe of aedeagus stout (AL/AW = 3.9–4.1), right margin weakly pointed before apical lamella; apical lamella thick, ovate in dorsal view, LL subequal to LW, not declined to left; subuliform in lateral view, apex not bent upward. Endophallus with very large flared basal expansion of primary sclerite; flagellum extending almost to the base of apical orifice; apical sclerite narrowly V-shaped, weakly defined, basal core indistinct; basal sheath coarsely scaled, apical sheath finely scaled, similar size as basal sheath; squamate sac well divided, near middle of median lobe, dorsal-right to squamate sheath; distal sac smaller and closer to apex than proximal sac. Gonocoxite II of ovipositor nearly trapezoid, slightly longer than width, apex curved, angulate near inner apical angle, with two ensiform setae on outer apical angle, three on inner apical angle, one of them slightly distant from other two.
</paragraph>
</subSubSection>
<caption id="DF5197BD6235FF902DEFB4F3FA7A5BB2" ID-DOI="http://doi.org/10.5281/zenodo.7959082" ID-Zenodo-Dep="7959082" httpUri="https://zenodo.org/record/7959082/files/figure.png" pageId="115" pageNumber="116" startId="115.[151,250,1672,1696]" targetBox="[165,1413,170,1629]" targetPageId="115" targetType="figure">
<paragraph id="8B91C7356235FF902DEFB4F3FA7A5BB2" blockId="115.[151,1437,1672,1768]" pageId="115" pageNumber="116">
<emphasis id="B95A1B276235FF902DEFB4F3FEFB5BFA" bold="true" box="[151,284,1672,1696]" pageId="115" pageNumber="116">FIGURE 69</emphasis>
. Habitus of
<taxonomicName id="4C2EBCB66235FF902CDBB4F2FD595BFA" baseAuthorityName="Bates" baseAuthorityYear="1873" box="[419,702,1672,1696]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="115" pageNumber="116" phylum="Arthropoda" rank="species" species="latecincta" subGenus="Parena">
<emphasis id="B95A1B276235FF902CDBB4F2FE0A5BFA" box="[419,493,1673,1696]" italics="true" pageId="115" pageNumber="116">Parena</emphasis>
(
<emphasis id="B95A1B276235FF902C86B4F2FDAF5BFA" box="[510,584,1673,1696]" italics="true" pageId="115" pageNumber="116">Parena</emphasis>
)
<emphasis id="B95A1B276235FF902F22B4F3FD595BFA" box="[602,702,1672,1696]" italics="true" pageId="115" pageNumber="116">latecincta</emphasis>
</taxonomicName>
species group.
<emphasis id="B95A1B276235FF902E1CB4F3FC915BFA" bold="true" box="[868,886,1672,1696]" pageId="115" pageNumber="116">A</emphasis>
<taxonomicName id="4C2EBCB66235FF902EF8B4F2FB805BFA" authority="Shibata" authorityName="Shibata" authorityYear="1987" box="[896,1127,1672,1696]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="115" pageNumber="116" phylum="Arthropoda" rank="species" species="circumdata">
<emphasis id="B95A1B276235FF902EF8B4F2FBE95BFA" box="[896,1038,1672,1696]" italics="true" pageId="115" pageNumber="116">P. circumdata</emphasis>
Shibata
</taxonomicName>
, male from Taiwan.
<emphasis id="B95A1B276235FF90283AB4F3FAB45BFA" bold="true" box="[1346,1363,1672,1696]" pageId="115" pageNumber="116">B</emphasis>
<taxonomicName id="4C2EBCB66235FF902824B4F2FEA05B9E" authority="Shibata" authorityName="Shibata" authorityYear="1987" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="115" pageNumber="116" phylum="Arthropoda" rank="species" species="circumdata">
<emphasis id="B95A1B276235FF902824B4F2FF095B9E" italics="true" pageId="115" pageNumber="116">P. circumdata</emphasis>
Shibata
</taxonomicName>
, female from India.
<emphasis id="B95A1B276235FF902F58B4D7FDD55B9E" bold="true" box="[544,562,1708,1732]" pageId="115" pageNumber="116">C</emphasis>
<taxonomicName id="4C2EBCB66235FF902F44B4D6FCF25B9E" authority="Shibata" authorityName="Shibata" authorityYear="1987" box="[572,789,1708,1732]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="115" pageNumber="116" phylum="Arthropoda" rank="species" species="monticola">
<emphasis id="B95A1B276235FF902F44B4D6FD5B5B9E" box="[572,700,1708,1732]" italics="true" pageId="115" pageNumber="116">P. monticola</emphasis>
Shibata
</taxonomicName>
, male from Jiangxi.
<emphasis id="B95A1B276235FF902E88B4D7FBE55B9E" bold="true" box="[1008,1026,1708,1732]" pageId="115" pageNumber="116">D</emphasis>
<taxonomicName id="4C2EBCB66235FF902973B4D6FB025B9E" authority="Shibata" authorityName="Shibata" authorityYear="1987" box="[1035,1253,1708,1732]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="115" pageNumber="116" phylum="Arthropoda" rank="species" species="monticola">
<emphasis id="B95A1B276235FF902973B4D6FB6B5B9E" box="[1035,1164,1708,1732]" italics="true" pageId="115" pageNumber="116">P. monticola</emphasis>
Shibata
</taxonomicName>
, male from Sichuan.
<emphasis id="B95A1B276235FF902DB4B4ABFF3A5BB2" bold="true" box="[204,221,1744,1768]" pageId="115" pageNumber="116">E</emphasis>
Holotype of
<taxonomicName id="4C2EBCB66235FF902C15B4AAFD205BB2" authority="Andrewes. F" authorityName="Andrewes. F" box="[365,711,1744,1768]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="115" pageNumber="116" phylum="Arthropoda" rank="species" species="pendleburyi">
<emphasis id="B95A1B276235FF902C15B4AAFDDE5BB2" box="[365,569,1744,1768]" italics="true" pageId="115" pageNumber="116">Parena pendleburyi</emphasis>
Andrewes.
<emphasis id="B95A1B276235FF902FC0B4ABFD205BB2" bold="true" box="[696,711,1744,1768]" pageId="115" pageNumber="116">F</emphasis>
</taxonomicName>
<taxonomicName id="4C2EBCB66235FF902FA9B4AAFC3C5BB2" authority="Andrewes" authorityName="Andrewes, Borneo. U" authorityYear="1931" box="[721,987,1744,1768]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="115" pageNumber="116" phylum="Arthropoda" rank="species" species="pendleburyi">
<emphasis id="B95A1B276235FF902FA9B4AAFD075BB2" box="[721,736,1745,1768]" italics="true" pageId="115" pageNumber="116">P</emphasis>
.
<emphasis id="B95A1B276235FF902F88B4ABFC8E5BB2" box="[752,873,1744,1768]" italics="true" pageId="115" pageNumber="116">pendleburyi</emphasis>
Andrewes
</taxonomicName>
, female from Borneo. Scale bars = 2 mm.
</paragraph>
</caption>
<subSubSection id="C33494BE6232FF972DBFB3E9FAEF5CF7" box="[199,1288,402,429]" pageId="116" pageNumber="117" type="distribution">
<paragraph id="8B91C7356232FF972DBFB3E9FAEF5CF7" blockId="116.[151,1437,151,429]" box="[199,1288,402,429]" pageId="116" pageNumber="117">
<emphasis id="B95A1B276232FF972DBFB3E9FEBE5CF6" bold="true" box="[199,345,402,428]" pageId="116" pageNumber="117">Distribution</emphasis>
(
<figureCitation id="1315DBB06232FF972C10B3E8FE265CF7" box="[360,449,403,429]" captionStart="MAP 11" captionStartId="109.[152,211,1274,1298]" captionTargetBox="[151,1436,381,1251]" captionTargetId="figure-118@109.[151,1436,381,1251]" captionText="MAP 11. Distributions for species in the Parena (Parena) latecincta group" figureDoi="http://doi.org/10.5281/zenodo.7963693" httpUri="https://zenodo.org/record/7963693/files/figure.png" pageId="116" pageNumber="117">Map 11</figureCitation>
, blue).
<collectingCountry id="F33987A56232FF972F6DB3E8FDBF5CF7" box="[533,600,403,429]" name="China" pageId="116" pageNumber="117">China</collectingCountry>
(
<collectingCountry id="F33987A56232FF972F1EB3E8FD5A5CF7" box="[614,701,403,429]" name="Taiwan" pageId="116" pageNumber="117">Taiwan</collectingCountry>
, Guangdong),
<collectingCountry id="F33987A56232FF972E19B3E8FC275CF7" box="[865,960,403,429]" name="Vietnam" pageId="116" pageNumber="117">Vietnam</collectingCountry>
(
<collectingCountry id="F33987A56232FF972EB7B3E8FBC35CF7" box="[975,1060,403,429]" name="Vietnam" pageId="116" pageNumber="117">Tonkin</collectingCountry>
),
<collectingCountry id="F33987A56232FF97294FB3E8FB965CF7" box="[1079,1137,403,429]" name="India" pageId="116" pageNumber="117">India</collectingCountry>
(Darjeeling).
</paragraph>
</subSubSection>
</treatment>
</document>
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