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<document id="8173D181FA5B7213C3452EF09DE136B1" ID-CLB-Dataset="263342" ID-DOI="10.11646/zootaxa.5286.1.1" ID-GBIF-Dataset="67e7a105-1f62-414a-83d9-f758039eb726" ID-ISSN="1175-5326" ID-Zenodo-Dep="7958937" ID-ZooBank="F9834684-24D3-4795-B5EB-77B451DF856D" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="diego" checkinTime="1684793492880" checkinUser="plazi" docAuthor="Shi, Hongliang &amp; Liang, Hongbin" docDate="2023" docId="03877623621EFFBA2DEFB3E9FA655E4F" docLanguage="en" docName="zootaxa.5286.1.1.pdf" docOrigin="Zootaxa 5286 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5286.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Parena (Parena) rubripicta Andrewes. A 1928" docType="treatment" docVersion="8" lastPageNumber="90" masterDocId="FFBE0E5B6246FFE32D78B27BFFE75D5A" masterDocTitle="Taxonomic revision of the genus Parena Motschulsky, 1860 (Coleoptera, Carabidae, Lebiini, Metallicina)" masterLastPageNumber="144" masterPageNumber="1" pageNumber="89" updateTime="1699506634024" updateUser="plazi">
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<mods:title id="EE61B7C06BE6DCEF712E0855D2EBBC47">Taxonomic revision of the genus Parena Motschulsky, 1860 (Coleoptera, Carabidae, Lebiini, Metallicina)</mods:title>
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<mods:namePart id="1C8A7721A32D408CFDFE5FA4E2350D2A">Shi, Hongliang</mods:namePart>
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<treatment id="03877623621EFFBA2DEFB3E9FA655E4F" ID-DOI="http://doi.org/10.5281/zenodo.7963771" ID-GBIF-Taxon="209628970" ID-Zenodo-Dep="7963771" LSID="urn:lsid:plazi:treatment:03877623621EFFBA2DEFB3E9FA655E4F" httpUri="http://treatment.plazi.org/id/03877623621EFFBA2DEFB3E9FA655E4F" lastPageId="89" lastPageNumber="90" pageId="88" pageNumber="89">
<subSubSection id="C33494BE621EFFBB2DEFB3E9FD0F5CF6" box="[151,744,402,429]" pageId="88" pageNumber="89" type="nomenclature">
<paragraph id="8B91C735621EFFBB2DEFB3E9FD0F5CF6" blockId="88.[151,1164,402,465]" box="[151,744,402,429]" pageId="88" pageNumber="89">
<emphasis id="B95A1B27621EFFBB2DEFB3E9FD0F5CF6" bold="true" box="[151,744,402,429]" pageId="88" pageNumber="89">
[25]
<taxonomicName id="4C2EBCB6621EFFBB2DB7B3E8FD0F5CF6" ID-CoL="4DKQC" authority="Andrewes, 1928" authorityName="Andrewes. A" authorityYear="1928" box="[207,744,402,429]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="rubripicta" subGenus="Parena">
<emphasis id="B95A1B27621EFFBB2DB7B3E8FEC05CF7" bold="true" box="[207,295,403,429]" italics="true" pageId="88" pageNumber="89">Parena</emphasis>
(
<emphasis id="B95A1B27621EFFBB2C40B3E8FE775CF7" bold="true" box="[312,400,403,429]" italics="true" pageId="88" pageNumber="89">Parena</emphasis>
)
<emphasis id="B95A1B27621EFFBB2CDAB3E9FDFC5CF7" bold="true" box="[418,539,402,429]" italics="true" pageId="88" pageNumber="89">rubripicta</emphasis>
<bibRefCitation id="EFBFBAC4621EFFBB2F5AB3E9FD0F5CF6" author="Andrewes, H. E." box="[546,744,402,428]" pageId="88" pageNumber="89" pagination="1 - 24" refId="ref109566" refString="Andrewes, H. E. (1928) On the types of Oriental Carabidae described by V. de Motschulsky. The Transactions of the Entomological Society of London, 76, 1 - 24. https: // doi. org / 10.1111 / j. 1365 - 2311.1928. tb 01185. x" type="journal article" year="1928">Andrewes, 1928</bibRefCitation>
</taxonomicName>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C33494BE621EFFBB2DEFB3CCFB6B5C8B" box="[151,1164,438,465]" pageId="88" pageNumber="89" type="description">
<paragraph id="8B91C735621EFFBB2DEFB3CCFB6B5C8B" blockId="88.[151,1164,402,465]" box="[151,1164,438,465]" pageId="88" pageNumber="89">
Habitus:
<figureCitation id="1315DBB0621EFFBB2D85B3CCFE3D5C8B" box="[253,474,438,465]" captionStart="FIGURE 50" captionStartId="85.[151,250,1856,1880]" captionTargetBox="[176,1411,484,1822]" captionTargetId="figure-128@85.[171,1416,478,1832]" captionTargetPageId="85" captionText="FIGURE 50. Habitus of Parena (Parena) bicolor Motschulsky and Parena (Parena) rubripicta Andrewes. A P. bicolor Motschulsky, male from Java. B P. bicolor Motschulsky, male from Sumatra. C P. bicolor Motschulsky, male from Philippine. D Holotype of Parena rubripicta Andrewes. E P. rubripicta Andrewes, female from Yunnan. F P. rubripicta Andrewes, female from Guangxi. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7959041" httpUri="https://zenodo.org/record/7959041/files/figure.png" pageId="88" pageNumber="89">
Figs 50D, 50E,
<specimenCount id="9D280CBC621EFFBB2CD6B3CDFE3D5C8B" box="[430,474,438,465]" pageId="88" pageNumber="89" type="generic">50F</specimenCount>
</figureCitation>
. Male genitalia:
<figureCitation id="1315DBB0621EFFBB2FEDB3CCFD005C8B" box="[661,743,438,465]" captionStart="FIGURE 52" captionStartId="89.[151,250,1641,1665]" captionTargetBox="[173,1413,825,1608]" captionTargetId="figure-331@89.[165,1421,813,1618]" captionTargetPageId="89" captionText="FIGURE 52. Median lobe of male genitalia for Parena (Parena) rubripicta Andrewes, right-lateral (r), ventral (v), left-lateral (l), and dorsal (d) views, Xizang (Medog). Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7959045" httpUri="https://zenodo.org/record/7959045/files/figure.png" pageId="88" pageNumber="89">Fig. 52</figureCitation>
. Gonocoxites of ovipositor:
<figureCitation id="1315DBB0621EFFBB295DB3CCFB625C8B" box="[1061,1157,439,465]" captionStart="FIGURE 11" captionStartId="18.[151,250,150,174]" captionTargetId="figure-19@17.[155,1432,181,1971]" captionTargetPageId="17" captionText="FIGURE 11. Gonocoxites (left side) of ovipositor for Parena spp., ventral side (dorsal side for fig. Q). AJ subgenus Bothynoptera: A P. (Bothynoptera) heteronycha sp.n., Laos, paratype. B P. (Bothynoptera) dorsigera (Schaum), Shaanxi. C P. (Bothynoptera) kurosai Habu, Guangdong. D P. (Bothynoptera) tesari (Jedlička), Taiwan. E P. (Bothynoptera) taiwana Hua, Taiwan. F P. (Bothynoptera) tripunctata (Bates), Shaanxi. G P. (Bothynoptera) malaisei (Andrewes), Taiwan. H P. (Bothynoptera) quadrisignata Mateu, Yunnan. I P. (Bothynoptera) emarginata sp. n., holotype. J P. (Bothynoptera) triguttata sp. n., Sichuan, paratype. KX subgenus Parena: K P. (Parena) bicolor Motschulsky, Sulawesi. L P. (Parena) rubripicta Andrewes, Yunnan. M P. (Parena) fasciata (Chaudoir), Taiwan. N P. (Parena) nigrolineata (Chaudoir), Taiwan. O P. (Parena) picipes sp. n., Borneo, paratype. P P. (Parena) andrewesi Jedlička, Philippine. Q P. (Parena) politissima (Chaudoir), Vanuatu. R P. (Parena) latecincta (Bates), Guizhou. S P. (Parena) circumdata Shibata, Taiwan. T P. (Parena) pendleburyi Andrewes, Borneo. U P. (Parena) fulva sp. n., Zimbabwe, paratype. V P. (Parena) madagascariensis (Alluaud), Madagascar. W P. (Parena) ruficornis sp.n., CentralAfrica, paratype. X P. (Parena) africana (Alluaud), Uganda. Yγ subgenus Crossoglossa: Y P. (Crossoglossa) cavipennis (Bates), Taiwan. Z P. (Crossoglossa) laesipennis (Bates), Yunnan. α P. (Crossoglossa) testacea (Chaudoir), Zhejiang. β P. (Crossoglossa) sulawesiensis Kirschenhofer, Sulawesi. γ P. (Crossoglossa) sciakyi sp. n., holotype. Scale bar = 0.2mm." figureDoi="http://doi.org/10.5281/zenodo.7963954" httpUri="https://zenodo.org/record/7963954/files/figure.png" pageId="88" pageNumber="89">
Fig.
<specimenCount id="9D280CBC621EFFBB2921B3CCFB625C8B" box="[1113,1157,439,465]" pageId="88" pageNumber="89" type="generic">11L</specimenCount>
</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33494BE621EFFBB2DEFB380FCD35F68" pageId="88" pageNumber="89" type="reference_group">
<paragraph id="8B91C735621EFFBB2DEFB380FCD35F68" blockId="88.[151,1436,507,562]" pageId="88" pageNumber="89">
<bibRefCitation id="EFBFBAC4621EFFBB2DEFB380FE8F5F49" author="Andrewes, H. E." box="[151,360,507,531]" pageId="88" pageNumber="89" pagination="1 - 24" refId="ref109566" refString="Andrewes, H. E. (1928) On the types of Oriental Carabidae described by V. de Motschulsky. The Transactions of the Entomological Society of London, 76, 1 - 24. https: // doi. org / 10.1111 / j. 1365 - 2311.1928. tb 01185. x" type="journal article" year="1928">Andrewes, 1928: 17</bibRefCitation>
(type locality: Nilgiri Hills;
<typeStatus id="54957997621EFFBB2FEFB380FD175F49" box="[663,752,507,531]" pageId="88" pageNumber="89" type="holotype">holotype</typeStatus>
in NHML);
<bibRefCitation id="EFBFBAC4621EFFBB2E00B380FBA25F49" author="Jedlicka, A." box="[888,1093,507,531]" pageId="88" pageNumber="89" pagination="269 - 579" refId="ref111456" refString="Jedlicka, A. (1963) Monographie der Truncatipennen aus Ostasien, Lebiinae- Odacanthinae- Braehyninae (Coleptera, Carabidae). Entomologische Abhandlungen und Berichte aus dem Staatlichen Museum fuer Tierkunde in Dresden, 28, 269 - 579." type="journal article" year="1963">Jedlička, 1963: 441</bibRefCitation>
;
<bibRefCitation id="EFBFBAC4621EFFBB292AB380FAC95F49" author="Xie, W. P. &amp; Yu, P. Y." box="[1106,1326,507,531]" pageId="88" pageNumber="89" pagination="185 - 195" refId="ref113326" refString="Xie, W. P. &amp; Yu, P. Y. (1993) On the Chinese species of Parena Motschulsky (Coleoptera: Carabidae). Sinozoologia, 10, 185 - 195." type="journal article" year="1993">Xie &amp; Yu, 1993: 188</bibRefCitation>
(Yunnan);
<bibRefCitation id="EFBFBAC4621EFFBB2DBFB061FE305F68" author="Kirschenhofer, E." box="[199,471,538,562]" pageId="88" pageNumber="89" pagination="999 - 1067" refId="ref111570" refString="Kirschenhofer, E. (1994) Neue und wenig bekannte Carabidae aus der palaarktischen und orientalischen Region (Col. Carabidae, Lebiinae, Odacanthinae, Brachininae, Panagaeinae). Linzer Biologische Beitrage, 26, 999 - 1067." type="journal article" year="1994">Kirschenhofer, 1994: 1047</bibRefCitation>
(
<collectingCountry id="F33987A5621EFFBB2C9CB061FDC15F68" box="[484,550,538,562]" name="Nepal" pageId="88" pageNumber="89">Nepal</collectingCountry>
);
<bibRefCitation id="EFBFBAC4621EFFBB2F40B061FCD75F68" author="Kirschenhofer, E." box="[568,816,538,562]" pageId="88" pageNumber="89" pagination="87 - 103" refId="ref111612" refString="Kirschenhofer, E. (2006) Neue Arten der Gattung Parena Motschulsky aus Japan, China Und Indonesien (Sulawesi) (Coleoptera: Carabidae). Mitteilungen des Internationalen Entomologischen Vereins E. V. Frankfurt, 31 (3 - 4), 87 - 103." type="journal article" year="2006">Kirschenhofer, 2006: 89</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33494BE621EFFBB2DEFB01BFAF75B8C" pageId="88" pageNumber="89" type="materials_examined">
<paragraph id="8B91C735621EFFBB2DEFB01BFB645E09" blockId="88.[151,1437,607,2039]" pageId="88" pageNumber="89">
<emphasis id="B95A1B27621EFFBB2DEFB01BFE5F5F20" bold="true" box="[151,440,608,634]" pageId="88" pageNumber="89">Type material examined</emphasis>
.
<emphasis id="B95A1B27621EFFBB2CBEB01BFC685F20" bold="true" box="[454,911,607,634]" pageId="88" pageNumber="89">
<taxonomicName id="4C2EBCB6621EFFBB2CBEB01BFCF15F20" authority="Andrewes" authorityName="Andrewes. A" authorityYear="1928" box="[454,790,607,634]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="rubripicta">
<emphasis id="B95A1B27621EFFBB2CBEB01BFD735F20" bold="true" box="[454,660,607,634]" italics="true" pageId="88" pageNumber="89">Parena rubripicta</emphasis>
Andrewes
</taxonomicName>
:
<typeStatus id="54957997621EFFBB2E5AB01BFC685F20" box="[802,911,608,634]" pageId="88" pageNumber="89" type="holotype">Holotype</typeStatus>
</emphasis>
(NHML,
<figureCitation id="1315DBB0621EFFBB2978B01BFB8F5F21" box="[1024,1128,608,635]" captionStart="FIGURE 50" captionStartId="85.[151,250,1856,1880]" captionTargetBox="[176,1411,484,1822]" captionTargetId="figure-128@85.[171,1416,478,1832]" captionTargetPageId="85" captionText="FIGURE 50. Habitus of Parena (Parena) bicolor Motschulsky and Parena (Parena) rubripicta Andrewes. A P. bicolor Motschulsky, male from Java. B P. bicolor Motschulsky, male from Sumatra. C P. bicolor Motschulsky, male from Philippine. D Holotype of Parena rubripicta Andrewes. E P. rubripicta Andrewes, female from Yunnan. F P. rubripicta Andrewes, female from Guangxi. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7959041" httpUri="https://zenodo.org/record/7959041/files/figure.png" pageId="88" pageNumber="89">Fig. 50D</figureCitation>
): female, body length =
<quantity id="4CD66AD0621EFFBB2801B01BFF225FC5" metricMagnitude="-3" metricUnit="m" metricValue="8.1" pageId="88" pageNumber="89" unit="mm" value="8.1">8.1 mm</quantity>
, pin mounted, &quot;H.L. Andrewes. / Nilgiri Hills.&quot;, &quot;
<emphasis id="B95A1B27621EFFBB2F81B0FFFCC45FC4" box="[761,803,644,670]" italics="true" pageId="88" pageNumber="89">788</emphasis>
&quot;, &quot;Type&quot; [red label]; &quot;
<emphasis id="B95A1B27621EFFBB295BB0FEFA6A5FC4" box="[1059,1421,644,670]" italics="true" pageId="88" pageNumber="89">
<taxonomicName id="4C2EBCB6621EFFBB295BB0FEFB935FC4" box="[1059,1140,645,670]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="genus">Parena</taxonomicName>
/
<taxonomicName id="4C2EBCB6621EFFBB29F4B0FFFB195FC4" authorityName="Andrewes. A" authorityYear="1928" box="[1164,1278,644,670]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="rubripicta">rubripicta</taxonomicName>
/ Type Andr.
</emphasis>
/ H.E. Andrewes det.&quot;, &quot;H.E. Andrewes Coll. / B.M. 1945-97.&quot;.
<emphasis id="B95A1B27621EFFBB2E28B0D3FC365F98" bold="true" box="[848,977,680,706]" pageId="88" pageNumber="89">
<typeStatus id="54957997621EFFBB2E28B0D3FC2B5F98" box="[848,972,680,706]" pageId="88" pageNumber="89" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<specimenCount id="9D280CBC621EFFBB2EA0B0D3FBC45F98" box="[984,1059,680,706]" pageId="88" pageNumber="89" type="male">1 male</specimenCount>
(NHML), &quot;Co-type&quot; [round label with yellow circle], &quot;
<emphasis id="B95A1B27621EFFBB2CFCB0B7FE075FBC" box="[388,480,716,742]" italics="true" pageId="88" pageNumber="89">Doherty</emphasis>
&quot;, &quot;
<emphasis id="B95A1B27621EFFBB2F7DB0B7FDAC5FBC" box="[517,587,716,742]" italics="true" pageId="88" pageNumber="89">22809</emphasis>
&quot;, &quot;
<emphasis id="B95A1B27621EFFBB2F08B0B7FCD75FBC" box="[624,816,716,742]" italics="true" pageId="88" pageNumber="89">Birmah / Momeit</emphasis>
&quot;, &quot;Fry Coll. / 1905.100.&quot;, &quot;
<emphasis id="B95A1B27621EFFBB291DB0B6FF045E50" italics="true" pageId="88" pageNumber="89">
<taxonomicName id="4C2EBCB6621EFFBB291DB0B6FB515FBC" box="[1125,1206,717,742]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="genus">Parena</taxonomicName>
/
<taxonomicName id="4C2EBCB6621EFFBB29B4B0B7FADF5FBC" authorityName="Andrewes. A" authorityYear="1928" box="[1228,1336,716,742]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="rubripicta">rubripicta</taxonomicName>
/ Cotype Andr. /
</emphasis>
H.E. Andrewes det.&quot;.
<specimenCount id="9D280CBC621EFFBB2C9CB08BFDA15E50" box="[484,582,752,778]" pageId="88" pageNumber="89" type="female">1 female</specimenCount>
(NHML), &quot;Niambur, Madras. / S.N. Chatterjee.,
<emphasis id="B95A1B27621EFFBB2900B08BFB485E50" box="[1144,1199,752,778]" italics="true" pageId="88" pageNumber="89">7.XI.</emphasis>
192
<emphasis id="B95A1B27621EFFBB29A0B08BFB015E50" box="[1240,1254,752,778]" italics="true" pageId="88" pageNumber="89">5</emphasis>
&quot;, &quot;Predaceous / on Hybloea / puera larva&quot;, &quot;
<emphasis id="B95A1B27621EFFBB2CAAB16FFE1B5E74" box="[466,508,788,814]" italics="true" pageId="88" pageNumber="89">301</emphasis>
&quot;, &quot;For. Res. Ins. / Dehra
<collectingRegion id="49EA09D7621EFFBB2E61B16FFCA85E74" box="[793,847,788,814]" country="United Kingdom" name="Down" pageId="88" pageNumber="89">Dun</collectingRegion>
&quot;, &quot;Co-type&quot; [round label with green circle], &quot;
<emphasis id="B95A1B27621EFFBB2833B16EFE205E08" italics="true" pageId="88" pageNumber="89">
<taxonomicName id="4C2EBCB6621EFFBB2833B16EFA7B5E74" box="[1355,1436,789,814]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="genus">Parena</taxonomicName>
/
<taxonomicName id="4C2EBCB6621EFFBB2DDEB143FEF55E08" authorityName="Andrewes. A" authorityYear="1928" box="[166,274,824,850]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="rubripicta">rubripicta</taxonomicName>
/ Cotype Andr. /
</emphasis>
H.E. Andrewes det.&quot;, &quot;H.E. Andrewes Coll. / B.M. 1945-97.&quot;.
</paragraph>
<paragraph id="8B91C735621EFFBB2DBFB127FAF75B8C" blockId="88.[151,1437,607,2039]" pageId="88" pageNumber="89">
<emphasis id="B95A1B27621EFFBB2DBFB127FDFB5E2C" bold="true" box="[199,540,860,886]" pageId="88" pageNumber="89">Non-type material examined</emphasis>
.
<emphasis id="B95A1B27621EFFBB2F52B127FD8D5E2C" bold="true" box="[554,618,860,886]" pageId="88" pageNumber="89">
<collectingCountry id="F33987A5621EFFBB2F52B127FD8D5E2C" box="[554,618,860,886]" name="India" pageId="88" pageNumber="89">India</collectingCountry>
</emphasis>
: 1 ex (CMB), &quot;India, West Bengalen, Distr. Darjeeling, Kurseong, Kunti village,
<quantity id="4CD66AD0621EFFBB2D88B1FBFEDF5EC1" box="[240,312,896,923]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="88" pageNumber="89" unit="m" value="900.0">900 m</quantity>
,
<date id="FF90E1F5621EFFBB2C3BB1FBFE315EC0" box="[323,470,896,922]" pageId="88" pageNumber="89" value="1996-08-11">11.VIII.1996</date>
, leg. N. Dangal&quot;. 3 ex (NHMB), &quot;NE India, Arunachal Pr., Etalin vicinity, 800+-
<quantity id="4CD66AD0621EFFBB2837B1FBFA705EC1" box="[1359,1431,896,923]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="88" pageNumber="89" unit="m" value="100.0">100 m</quantity>
, 28°35'
<geoCoordinate id="EE1AA1F2621EFFBB2DA5B1DFFEDF5EE5" box="[221,312,932,959]" degrees="95" direction="north" minutes="52" orientation="latitude" pageId="88" pageNumber="89" precision="925" value="95.86667">N95°52</geoCoordinate>
E, Pacholatko leg.
<date id="FF90E1F5621EFFBB2F72B1DFFD7F5EE4" box="[522,664,932,958]" pageId="88" pageNumber="89" value="2007-06-01" valueMax="2007-06-03" valueMin="2007-06-01">1.-3.vi.2007</date>
&quot;. 1 ex (NHMB), &quot;India-S.
<collectingRegion id="49EA09D7621EFFBB2EBBB1DFFBF75EE4" box="[963,1040,932,958]" country="India" name="Kerala" pageId="88" pageNumber="89">Kerala</collectingRegion>
, Periyar Lk.
<quantity id="4CD66AD0621EFFBB29D9B1DFFB0D5EE5" box="[1185,1258,932,959]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="88" pageNumber="89" unit="m" value="900.0">900 m</quantity>
,
<date id="FF90E1F5621EFFBB298EB1DFFA705EE4" box="[1270,1431,932,959]" pageId="88" pageNumber="89" value="1991-05-13" valueMax="1991-05-20" valueMin="1991-05-13">13.-20.V.1991</date>
, leg. Jiri Kolibac&quot;. 4 ex (NHMB), &quot;Kalimpong, 6th mi
<quantity id="4CD66AD0621EFFBB2F84B1B3FCA25EB9" box="[764,837,968,995]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="88" pageNumber="89" unit="m" value="800.0">800 m</quantity>
,
<date id="FF90E1F5621EFFBB2E2AB1B3FC345EB9" box="[850,979,968,995]" pageId="88" pageNumber="89" value="1985-04-15">15.IV.1985</date>
&quot;, &quot;Indien, Darjeeling D. Ch.J. Rai&quot;. 1 ex (NHMB), &quot;Kalimpong, Chibo
<quantity id="4CD66AD0621EFFBB2C80B197FDA3595D" box="[504,580,1004,1031]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="88" pageNumber="89" unit="m" value="900.0">900 m</quantity>
,
<date id="FF90E1F5621EFFBB2F2BB197FD33595C" box="[595,724,1004,1030]" pageId="88" pageNumber="89" value="1987-04-27">27.IV.1987</date>
&quot;, &quot;Indien, Darjeeling D. Ch.J. Rai&quot;. 1 ex (NHMB), &quot;S. India,
<collectingRegion id="49EA09D7621EFFBB2DEFB66BFF035970" box="[151,228,1040,1066]" country="India" name="Kerala" pageId="88" pageNumber="89">Kerala</collectingRegion>
, Thekkady; Periyar Lake; 9,34N, 77,10E,
<quantity id="4CD66AD0621EFFBB2FBDB66BFCB75971" box="[709,848,1040,1067]" metricMagnitude="2" metricUnit="m" metricValue="9.5" metricValueMax="10.0" metricValueMin="9.0" pageId="88" pageNumber="89" unit="m" value="950.0" valueMax="1000.0" valueMin="900.0">900-1000 m</quantity>
,
<date id="FF90E1F5621EFFBB2E25B66BFBE45970" box="[861,1027,1040,1066]" pageId="88" pageNumber="89" value="1997-04-19" valueMax="1997-04-27" valueMin="1997-04-19">19.-27.iv.1997</date>
; Dembicky &amp; Pacholatko leg&quot;. 1 ex (NHMB), &quot;S. India,
<collectingRegion id="49EA09D7621EFFBB2C06B64FFE135914" box="[382,500,1076,1102]" country="India" name="Karnataka" pageId="88" pageNumber="89">Karnataka</collectingRegion>
, W. Ghats,
<quantity id="4CD66AD0621EFFBB2F0DB64FFD515914" box="[629,694,1076,1102]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="88" pageNumber="89" unit="km" value="20.0">20km</quantity>
W Talguppa, JOG Falls, 14°14'
<geoCoordinate id="EE1AA1F2621EFFBB2961B64FFB9B5914" box="[1049,1148,1076,1102]" degrees="74" direction="north" minutes="44" orientation="latitude" pageId="88" pageNumber="89" precision="925" value="74.73333">N 74°44</geoCoordinate>
E, 500+-
<quantity id="4CD66AD0621EFFBB29A4B64FFAC05915" box="[1244,1319,1076,1103]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="88" pageNumber="89" unit="m" value="200.0">200 m</quantity>
, P. Pacholatko leg.
<date id="FF90E1F5621EFFBB2C73B623FE4B5928" box="[267,428,1112,1138]" pageId="88" pageNumber="89" value="2002-05-22" valueMax="2002-05-28" valueMin="2002-05-22">22.-28.v.2002</date>
&quot;. 1 ex (NHMB), &quot;Pudung
<quantity id="4CD66AD0621EFFBB2FA5B623FCCE5929" box="[733,809,1112,1139]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="88" pageNumber="89" unit="m" value="900.0">900 m</quantity>
,
<date id="FF90E1F5621EFFBB2E4FB623FC475929" box="[823,928,1112,1139]" pageId="88" pageNumber="89" value="1985-05-03">3.V.1985</date>
&quot;, &quot;Darjeeling D. India Bhakta B.&quot;.
<emphasis id="B95A1B27621EFFBB284FB623FA9A5928" bold="true" box="[1335,1405,1112,1138]" pageId="88" pageNumber="89">
<collectingCountry id="F33987A5621EFFBB284FB623FA9A5928" box="[1335,1405,1112,1138]" name="Nepal" pageId="88" pageNumber="89">Nepal</collectingCountry>
</emphasis>
: 1 ex (NHMB), &quot;Kali-G. Khola, Tatopani
<quantity id="4CD66AD0621EFFBB2F2FB607FD1759CD" box="[599,752,1148,1175]" metricMagnitude="3" metricUnit="m" metricValue="1.25" metricValueMax="1.4" metricValueMin="1.1" pageId="88" pageNumber="89" unit="m" value="1250.0" valueMax="1400.0" valueMin="1100.0">1100-1400 m</quantity>
,
<date id="FF90E1F5621EFFBB2F86B607FC4F59CC" box="[766,936,1148,1174]" pageId="88" pageNumber="89" value="1986-06-27" valueMax="1986-06-28" valueMin="1986-06-27">27-28.VI.1986</date>
&quot;, &quot;Nepal,
<collectingRegion id="49EA09D7621EFFBB2964B607FB4E59CC" box="[1052,1193,1148,1174]" country="Nepal" name="Dhawalagiri" pageId="88" pageNumber="89">Dhawalagiri</collectingRegion>
, C. Holzschuh&quot;. 1 ex (NHMB), &quot;Nepal, Kosi-#14, Barabishe 27°26'N/ 87°18'E to Deorali 27°24'N/ 87°20'E,
<quantity id="4CD66AD0621EFFBB29E8B6DBFAF959E1" box="[1168,1310,1184,1211]" metricMagnitude="3" metricUnit="m" metricValue="1.005" metricValueMax="1.45" metricValueMin="0.5599999999999999" pageId="88" pageNumber="89" unit="m" value="1005.0" valueMax="1450.0" valueMin="560.0">560-1450 m</quantity>
, 13.vi.01&quot;, &quot;NHMB Basel, expedition to Nepal 2001&quot;.
<emphasis id="B95A1B27621EFFBB2F01B6BFFD385984" bold="true" box="[633,735,1220,1246]" pageId="88" pageNumber="89">
<collectingCountry id="F33987A5621EFFBB2F01B6BFFD385984" box="[633,735,1220,1246]" name="Vietnam" pageId="88" pageNumber="89">Vietnam</collectingCountry>
</emphasis>
: 1 ex (MNHN), &quot;Hoa-Binh, Tonkin, A. de Cooman&quot;, &quot;Museum Paris, Coll. Ch. Alluaud&quot;. 1 ex (MNHN), &quot;Tonkin, Region de, Hoa-Binh&quot;, &quot;Museum Paris, 1931, A. de Cooman&quot;, &quot;Museum Paris ex coll. R. Oberthur, 1952&quot;. 1 ex (MNHN), &quot;Chapa, Tonkin, (ex. Jeanvoine), Coll. Clermont&quot;, &quot;Museum Paris, 1988, Coll. J. Negre&quot;. 1 ex (MNHN), &quot;Hoa-binh (Tonkin), (A. de cooman), Coll. J. Clermont&quot;, &quot;Museum Paris, 1988, Coll. J. Negre&quot;. 1 ex (MNHN), &quot;env. Hoah binh, Tonkin&quot;, &quot;Museum Paris, 1988, Coll. J. Negre&quot;.
<specimenCount id="9D280CBC621EFFBB2D8DB703FEA758C8" box="[245,320,1400,1426]" pageId="88" pageNumber="89" type="male">1 male</specimenCount>
(NHML), &quot;Cochinchine, Tayninh, le
<date id="FF90E1F5621EFFBB2F9DB703FC8458C8" box="[741,867,1400,1426]" pageId="88" pageNumber="89" value="1924-12-01">1.XII.1924</date>
, R. Vitalis de Salvaza&quot;.
<emphasis id="B95A1B27621EFFBB2908B703FB0F58C8" bold="true" box="[1136,1256,1400,1426]" pageId="88" pageNumber="89">
<collectingCountry id="F33987A5621EFFBB2908B703FB0F58C8" box="[1136,1256,1400,1426]" name="Myanmar" pageId="88" pageNumber="89">Myanmar</collectingCountry>
</emphasis>
: 1 ex (MNHN), &quot;Hte Birmanie, Etat de Momeit,
<quantity id="4CD66AD0621EFFBB2F71B7E7FDB458ED" box="[521,595,1436,1463]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="88" pageNumber="89" unit="m" value="600.0">600 m</quantity>
, Doherty, 1890&quot;, &quot;Museum Paris, 1952, Coll. R. OBERthur&quot;.
<emphasis id="B95A1B27621EFFBB286AB7E7FA9858EC" bold="true" box="[1298,1407,1436,1462]" pageId="88" pageNumber="89">
<collectingCountry id="F33987A5621EFFBB286AB7E7FA9858EC" box="[1298,1407,1436,1462]" name="Thailand" pageId="88" pageNumber="89">Thailand</collectingCountry>
</emphasis>
: 1 ex (MNHN), &quot;Museum Paris, Isth, de KRA, harmand, 1885&quot;, &quot;Museum Paris, ex Coll. R. Oberthur, 1952&quot;.
<emphasis id="B95A1B27621EFFBB2833B7BBFA735880" bold="true" box="[1355,1428,1472,1498]" pageId="88" pageNumber="89">
<collectingCountry id="F33987A5621EFFBB2833B7BBFA735880" box="[1355,1428,1472,1498]" name="China" pageId="88" pageNumber="89">China</collectingCountry>
</emphasis>
:
<specimenCount id="9D280CBC621EFFBB2DEFB79FFF1E58A4" box="[151,249,1508,1534]" pageId="88" pageNumber="89" type="female">1 female</specimenCount>
(IZAS), &quot;
<collectingRegion id="49EA09D7621EFFBB2C15B79FFE3758A4" box="[365,464,1508,1534]" country="China" name="Guangxi" pageId="88" pageNumber="89">Guangxi</collectingRegion>
, Longzhou, Daqingshan,
<quantity id="4CD66AD0621EFFBB2F8AB79FFCDA58A5" box="[754,829,1508,1535]" metricMagnitude="2" metricUnit="m" metricValue="3.6" pageId="88" pageNumber="89" unit="m" value="360.0">360 m</quantity>
,
<date id="FF90E1F5621EFFBB2E32B79FFC2058A4" box="[842,967,1508,1534]" pageId="88" pageNumber="89" value="1963-04-19">1963.IV.19</date>
, Wang Shuyong lgt.&quot; &lt;
<figureCitation id="1315DBB0621EFFBB29A9B79FFAD358A5" box="[1233,1332,1508,1535]" captionStart="FIGURE 50" captionStartId="85.[151,250,1856,1880]" captionTargetBox="[176,1411,484,1822]" captionTargetId="figure-128@85.[171,1416,478,1832]" captionTargetPageId="85" captionText="FIGURE 50. Habitus of Parena (Parena) bicolor Motschulsky and Parena (Parena) rubripicta Andrewes. A P. bicolor Motschulsky, male from Java. B P. bicolor Motschulsky, male from Sumatra. C P. bicolor Motschulsky, male from Philippine. D Holotype of Parena rubripicta Andrewes. E P. rubripicta Andrewes, female from Yunnan. F P. rubripicta Andrewes, female from Guangxi. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7959041" httpUri="https://zenodo.org/record/7959041/files/figure.png" pageId="88" pageNumber="89">Fig. 50F</figureCitation>
&gt;.
<specimenCount id="9D280CBC621EFFBB2828B79FFA7B58A4" box="[1360,1436,1508,1534]" pageId="88" pageNumber="89" type="male">1 male</specimenCount>
(IZAS), &quot;
<collectingRegion id="49EA09D7621EFFBB2D86B473FEBD5B78" box="[254,346,1544,1570]" country="China" name="Yunnan" pageId="88" pageNumber="89">Yunnan</collectingRegion>
, Xishuangbanna, Menga,
<quantity id="4CD66AD0621EFFBB2FF0B473FCC75B79" box="[648,800,1544,1571]" metricMagnitude="3" metricUnit="m" metricValue="1.065" metricValueMax="1.08" metricValueMin="1.05" pageId="88" pageNumber="89" unit="m" value="1065.0" valueMax="1080.0" valueMin="1050.0">1050-1080 m</quantity>
,
<date id="FF90E1F5621EFFBB2E54B473FC465B78" box="[812,929,1544,1571]" pageId="88" pageNumber="89" value="1958-05-29">1958.V.29</date>
, Wang Shuyong lgt.&quot;. 1 ex (IZAS), &quot;
<collectingRegion id="49EA09D7621EFFBB2844B473FA7F5B78" box="[1340,1432,1544,1570]" country="China" name="Yunnan" pageId="88" pageNumber="89">Yunnan</collectingRegion>
, Xishuangbanna, Menglun Botany Garden,
<date id="FF90E1F5621EFFBB2F0EB457FD145B1C" box="[630,755,1580,1606]" pageId="88" pageNumber="89" value="2009-12-01">
2009.XII.
<quantity id="4CD66AD0621EFFBB2F92B457FD145B1C" box="[746,755,1580,1606]" metricMagnitude="3" metricUnit="m" metricValue="1.56" pageId="88" pageNumber="89" unit="m" value="1560.0">1</quantity>
</date>
, 560 m, Tang Guo lgt.&quot;.
<specimenCount id="9D280CBC621EFFBB297DB457FB815B1C" box="[1029,1126,1580,1606]" pageId="88" pageNumber="89" type="female">1 female</specimenCount>
(IZAS), &quot;
<collectingRegion id="49EA09D7621EFFBB29ABB457FAC85B1C" box="[1235,1327,1580,1606]" country="China" name="Yunnan" pageId="88" pageNumber="89">Yunnan</collectingRegion>
, Lincang, Zhenkang county,
<geoCoordinate id="EE1AA1F2621EFFBB2C1FB42BFE105B31" box="[359,503,1616,1643]" degrees="23" direction="north" minutes="54" orientation="latitude" pageId="88" pageNumber="89" precision="15" seconds="18" value="23.904999">N23°5418</geoCoordinate>
,
<geoCoordinate id="EE1AA1F2621EFFBB2F7BB42BFD775B31" box="[515,656,1616,1643]" degrees="98" direction="east" minutes="55" orientation="longitude" pageId="88" pageNumber="89" precision="15" seconds="15" value="98.92083">E98°5515</geoCoordinate>
,
<quantity id="4CD66AD0621EFFBB2FE4B42BFD025B31" box="[668,741,1616,1643]" metricMagnitude="2" metricUnit="m" metricValue="8.55" pageId="88" pageNumber="89" unit="m" value="855.0">855 m</quantity>
,
<date id="FF90E1F5621EFFBB2F89B42BFC935B30" box="[753,884,1616,1642]" pageId="88" pageNumber="89" value="2009-11-14">2009.XI.14</date>
, beating, Zhang Xiaoning lgt.&quot; &lt;
<figureCitation id="1315DBB0621EFFBB299EB42BFAAD5B31" box="[1254,1354,1616,1643]" captionStart="FIGURE 11" captionStartId="18.[151,250,150,174]" captionTargetId="figure-19@17.[155,1432,181,1971]" captionTargetPageId="17" captionText="FIGURE 11. Gonocoxites (left side) of ovipositor for Parena spp., ventral side (dorsal side for fig. Q). AJ subgenus Bothynoptera: A P. (Bothynoptera) heteronycha sp.n., Laos, paratype. B P. (Bothynoptera) dorsigera (Schaum), Shaanxi. C P. (Bothynoptera) kurosai Habu, Guangdong. D P. (Bothynoptera) tesari (Jedlička), Taiwan. E P. (Bothynoptera) taiwana Hua, Taiwan. F P. (Bothynoptera) tripunctata (Bates), Shaanxi. G P. (Bothynoptera) malaisei (Andrewes), Taiwan. H P. (Bothynoptera) quadrisignata Mateu, Yunnan. I P. (Bothynoptera) emarginata sp. n., holotype. J P. (Bothynoptera) triguttata sp. n., Sichuan, paratype. KX subgenus Parena: K P. (Parena) bicolor Motschulsky, Sulawesi. L P. (Parena) rubripicta Andrewes, Yunnan. M P. (Parena) fasciata (Chaudoir), Taiwan. N P. (Parena) nigrolineata (Chaudoir), Taiwan. O P. (Parena) picipes sp. n., Borneo, paratype. P P. (Parena) andrewesi Jedlička, Philippine. Q P. (Parena) politissima (Chaudoir), Vanuatu. R P. (Parena) latecincta (Bates), Guizhou. S P. (Parena) circumdata Shibata, Taiwan. T P. (Parena) pendleburyi Andrewes, Borneo. U P. (Parena) fulva sp. n., Zimbabwe, paratype. V P. (Parena) madagascariensis (Alluaud), Madagascar. W P. (Parena) ruficornis sp.n., CentralAfrica, paratype. X P. (Parena) africana (Alluaud), Uganda. Yγ subgenus Crossoglossa: Y P. (Crossoglossa) cavipennis (Bates), Taiwan. Z P. (Crossoglossa) laesipennis (Bates), Yunnan. α P. (Crossoglossa) testacea (Chaudoir), Zhejiang. β P. (Crossoglossa) sulawesiensis Kirschenhofer, Sulawesi. γ P. (Crossoglossa) sciakyi sp. n., holotype. Scale bar = 0.2mm." figureDoi="http://doi.org/10.5281/zenodo.7963954" httpUri="https://zenodo.org/record/7963954/files/figure.png" pageId="88" pageNumber="89">Figs 11L</figureCitation>
,
<figureCitation id="1315DBB0621EFFBB2820B42BFA6F5B31" box="[1368,1416,1616,1643]" captionStart="FIGURE 50" captionStartId="85.[151,250,1856,1880]" captionTargetBox="[176,1411,484,1822]" captionTargetId="figure-128@85.[171,1416,478,1832]" captionTargetPageId="85" captionText="FIGURE 50. Habitus of Parena (Parena) bicolor Motschulsky and Parena (Parena) rubripicta Andrewes. A P. bicolor Motschulsky, male from Java. B P. bicolor Motschulsky, male from Sumatra. C P. bicolor Motschulsky, male from Philippine. D Holotype of Parena rubripicta Andrewes. E P. rubripicta Andrewes, female from Yunnan. F P. rubripicta Andrewes, female from Guangxi. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7959041" httpUri="https://zenodo.org/record/7959041/files/figure.png" pageId="88" pageNumber="89">50E</figureCitation>
&gt;.
<specimenCount id="9D280CBC621EFFBB2DEFB40FFF1C5BD4" box="[151,251,1652,1678]" pageId="88" pageNumber="89" type="female">1 female</specimenCount>
(HBUM), &quot;
<date id="FF90E1F5621EFFBB2CFCB40FFE175BD4" box="[388,496,1652,1678]" pageId="88" pageNumber="89" value="2007-08-08">2007-8-8</date>
,
<collectingRegion id="49EA09D7621EFFBB2C85B40EFDB15BD5" box="[509,598,1653,1679]" country="China" name="Yunnan" pageId="88" pageNumber="89">Yunnan</collectingRegion>
, Mengla, Menglun, Shi FM &amp; Mao SL leg.&quot;.
<specimenCount id="9D280CBC621EFFBB2912B40FFB5F5BD4" box="[1130,1208,1652,1678]" pageId="88" pageNumber="89" type="male">1 male</specimenCount>
(HBUM), &quot;
<collectingRegion id="49EA09D7621EFFBB2839B40FFA7F5BD4" box="[1345,1432,1652,1678]" country="China" name="Xizang" pageId="88" pageNumber="89">Xizang</collectingRegion>
, Mêdog, between Baibung-Mêdog,
<quantity id="4CD66AD0621EFFBB2F59B4E3FD4B5BE9" box="[545,684,1688,1715]" metricMagnitude="3" metricUnit="m" metricValue="1.05" metricValueMax="1.3" metricValueMin="0.8" pageId="88" pageNumber="89" unit="m" value="1050.0" valueMax="1300.0" valueMin="800.0">800-1300 m</quantity>
,
<date id="FF90E1F5621EFFBB2FC0B4E3FCAA5BE8" box="[696,845,1688,1714]" pageId="88" pageNumber="89" value="2003-08-13">2003.VIII.13</date>
, Ren Guodong lgt.&quot; &lt;
<figureCitation id="1315DBB0621EFFBB293DB4E3FB4D5BE8" box="[1093,1194,1688,1714]" captionStart="FIGURE 10" captionStartId="16.[151,250,1909,1933]" captionTargetBox="[163,1429,1250,1879]" captionTargetId="figure-182@16.[152,1436,1241,1885]" captionTargetPageId="16" captionText="FIGURE 10. Parameres of male genitalia for Parena spp. A P. (Crossoglossa) cavipennis (Bates), Beijing. B P. (Crossoglossa) testacea (Chaudoir), Shanghai. C P. (Bothynoptera) dorsigera (Schaum), Zhejiang. D P. (Bothynoptera) taiwana Hua, Taiwan. E P. (Bothynoptera) tripunctata (Bates), Shaanxi. F P. (Parena) rubripicta Andrewes, Xizang. G P. (Parena) fasciata (Chaudoir), Taiwan. H P. (Parena) nigrolineata (Chaudoir), Taiwan. I P. (Parena) latecincta (Bates), Yunnan. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7958957" httpUri="https://zenodo.org/record/7958957/files/figure.png" pageId="88" pageNumber="89">Figs 10F</figureCitation>
,
<figureCitation id="1315DBB0621EFFBB29CEB4E3FB325BE9" box="[1206,1237,1688,1715]" captionStart="FIGURE 52" captionStartId="89.[151,250,1641,1665]" captionTargetBox="[173,1413,825,1608]" captionTargetId="figure-331@89.[165,1421,813,1618]" captionTargetPageId="89" captionText="FIGURE 52. Median lobe of male genitalia for Parena (Parena) rubripicta Andrewes, right-lateral (r), ventral (v), left-lateral (l), and dorsal (d) views, Xizang (Medog). Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7959045" httpUri="https://zenodo.org/record/7959045/files/figure.png" pageId="88" pageNumber="89">52</figureCitation>
&gt;.
<specimenCount id="9D280CBC621EFFBB2988B4E3FADB5BE8" box="[1264,1340,1688,1714]" pageId="88" pageNumber="89" type="male">1 male</specimenCount>
(IZAS), &quot;China,
<collectingRegion id="49EA09D7621EFFBB2D8BB4C7FED75B8C" box="[243,304,1724,1750]" country="China" name="Xizang" pageId="88" pageNumber="89">Tibet</collectingRegion>
, Mêdog, road to Gelin village,
<geoCoordinate id="EE1AA1F2621EFFBB2FF5B4C7FD085B8C" box="[653,751,1724,1750]" degrees="29.481" direction="north" orientation="latitude" pageId="88" pageNumber="89" precision="55" value="29.481">N29.481</geoCoordinate>
<geoCoordinate id="EE1AA1F2621EFFBB2F8EB4C7FC835B8D" box="[758,868,1724,1751]" degrees="95.1881" direction="east" orientation="longitude" pageId="88" pageNumber="89" precision="5" value="95.1881">E95.1881</geoCoordinate>
,
<quantity id="4CD66AD0621EFFBB2E08B4C7FC5E5B8D" box="[880,953,1724,1751]" metricMagnitude="2" metricUnit="m" metricValue="9.6" pageId="88" pageNumber="89" unit="m" value="960.0">960 m</quantity>
,
<date id="FF90E1F5621EFFBB2EBDB4C7FBDE5B8D" box="[965,1081,1724,1751]" pageId="88" pageNumber="89" value="2015-08-24">2015.8.24</date>
D, Liang HB coll.&quot;.
</paragraph>
</subSubSection>
<subSubSection id="C33494BE621EFFBB2DBFB49BFD795AF5" pageId="88" pageNumber="89" type="diagnosis">
<paragraph id="8B91C735621EFFBB2DBFB49BFD795AF5" blockId="88.[151,1437,607,2039]" pageId="88" pageNumber="89">
<emphasis id="B95A1B27621EFFBB2DBFB49BFE815BA0" bold="true" box="[199,358,1760,1786]" pageId="88" pageNumber="89">Comparisons</emphasis>
. This species is most similar to
<taxonomicName id="4C2EBCB6621EFFBB2FB5B49AFCDE5BA0" authorityName="Motschulsky" authorityYear="1860" box="[717,825,1760,1786]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="B95A1B27621EFFBB2FB5B49AFCDE5BA0" box="[717,825,1760,1786]" italics="true" pageId="88" pageNumber="89">P. bicolor</emphasis>
</taxonomicName>
, distinguished as follows: (1) elytra with a median red patch of one to four intervals width, versus elytra entirely metallic in
<taxonomicName id="4C2EBCB6621EFFBB2ECCB57EFBC55A44" authorityName="Motschulsky" authorityYear="1860" box="[948,1058,1796,1822]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="B95A1B27621EFFBB2ECCB57EFBC55A44" box="[948,1058,1796,1822]" italics="true" pageId="88" pageNumber="89">P. bicolor</emphasis>
</taxonomicName>
; (2) right margin of median lobe distinctly sinuate before the apex (
<figureCitation id="1315DBB0621EFFBB2F60B553FD8C5A19" box="[536,619,1832,1859]" captionStart="FIGURE 52" captionStartId="89.[151,250,1641,1665]" captionTargetBox="[173,1413,825,1608]" captionTargetId="figure-331@89.[165,1421,813,1618]" captionTargetPageId="89" captionText="FIGURE 52. Median lobe of male genitalia for Parena (Parena) rubripicta Andrewes, right-lateral (r), ventral (v), left-lateral (l), and dorsal (d) views, Xizang (Medog). Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7959045" httpUri="https://zenodo.org/record/7959045/files/figure.png" pageId="88" pageNumber="89">Fig. 52</figureCitation>
), versus only very weakly sinuate in
<taxonomicName id="4C2EBCB6621EFFBB2972B552FB905A18" authorityName="Motschulsky" authorityYear="1860" box="[1034,1143,1832,1858]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="B95A1B27621EFFBB2972B552FB905A18" box="[1034,1143,1832,1858]" italics="true" pageId="88" pageNumber="89">P. bicolor</emphasis>
</taxonomicName>
(
<figureCitation id="1315DBB0621EFFBB29FEB553FB3E5A19" box="[1158,1241,1832,1859]" captionStart="FIGURE 51" captionStartId="86.[151,250,1801,1825]" captionTargetBox="[181,1411,211,1767]" captionTargetId="figure-28@86.[151,1436,181,1777]" captionTargetPageId="86" captionText="FIGURE 51. Median lobe of male genitalia for Parena (Parena) bicolor Motschulsky, right-lateral (r), ventral (v), left-lateral (l), and dorsal (d) views. A Sumatra. B Java. C Philippine. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7959043" httpUri="https://zenodo.org/record/7959043/files/figure.png" pageId="88" pageNumber="89">Fig. 51</figureCitation>
); (3)
<taxonomicName id="4C2EBCB6621EFFBB286DB552FA7B5A18" authorityName="Andrewes" authorityYear="1928" box="[1301,1436,1832,1858]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="ribripicta">
<emphasis id="B95A1B27621EFFBB286DB552FA7B5A18" box="[1301,1436,1832,1858]" italics="true" pageId="88" pageNumber="89">P. ribripicta</emphasis>
</taxonomicName>
occurs on the Asian mainland and
<taxonomicName id="4C2EBCB6621EFFBB2F62B536FD605A3C" authorityName="Motschulsky" authorityYear="1860" box="[538,647,1868,1894]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="B95A1B27621EFFBB2F62B536FD605A3C" box="[538,647,1868,1894]" italics="true" pageId="88" pageNumber="89">P. bicolor</emphasis>
</taxonomicName>
is known only from the Malay Archipelago.
<taxonomicName id="4C2EBCB6621EFFBB29F9B536FAE85A3C" authorityName="Andrewes. A" authorityYear="1928" box="[1153,1295,1868,1894]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="rubripicta">
<emphasis id="B95A1B27621EFFBB29F9B536FAE85A3C" box="[1153,1295,1868,1894]" italics="true" pageId="88" pageNumber="89">P. rubripicta</emphasis>
</taxonomicName>
may be also confused with some species in the
<taxonomicName id="4C2EBCB6621EFFBB2F65B50AFD405AD0" baseAuthorityName="Bates" baseAuthorityYear="1873" box="[541,679,1904,1930]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="latecincta">
<emphasis id="B95A1B27621EFFBB2F65B50AFD405AD0" box="[541,679,1904,1930]" italics="true" pageId="88" pageNumber="89">P. latecincta</emphasis>
</taxonomicName>
group based on a similar appearance, but it is readily distinguished from them by the presence of suborbital setae.
</paragraph>
</subSubSection>
<subSubSection id="C33494BE621EFFBA2DBFB5C3FD995FF3" lastPageId="89" lastPageNumber="90" pageId="88" pageNumber="89" type="description">
<paragraph id="8B91C735621EFFBA2DBFB5C3FD995FF3" blockId="88.[151,1437,607,2039]" lastBlockId="89.[151,1437,151,789]" lastPageId="89" lastPageNumber="90" pageId="88" pageNumber="89">
<emphasis id="B95A1B27621EFFBB2DBFB5C3FEB65A88" bold="true" box="[199,337,1976,2002]" pageId="88" pageNumber="89">Description</emphasis>
. Body length
<quantity id="4CD66AD0621EFFBB2C95B5C3FD915A89" box="[493,630,1976,2003]" metricMagnitude="-3" metricUnit="m" metricValue="7.95" metricValueMax="8.8" metricValueMin="7.1" pageId="88" pageNumber="89" unit="mm" value="7.95" valueMax="8.8" valueMin="7.1">7.18.8 mm</quantity>
. Head and pronotum reddish yellow to dark brown; antennae completely reddish brown or apical seven antennomeres slightly darker than basal ones; elytra metallic green, copperish green in some specimens, with a median red patch of one to four intervals width, with the patch edge not distinctly defined; epipleura reddish brown; venter of head and prothorax same as dorsum, venter of mesothorax, metathorax and abdomen red to dark reddish brown; legs reddish brown, femoral apices darker in some specimens. Postgenae with a pair of suborbital setae; antennae barely extended pronotum base; labrum quadrate, apex very weakly protruded at middle; mentum with a pair of short median setae. Pronotum nearly cordate, PW/PL = 1.281.40, nearly same width as head, PW/HW = 0.991.04; widest at anterior third, slightly rounded at anterior half, and then distinctly sinuate before posterior angles; posterior angles widely obtuse. Elytra slightly dilated to apex, surface without or with very faint isodiametric microsculpture; striae very shallowly incised, with fine puncture rows; intervals weakly convex, very sparsely punctate; disc with large shallow depressions near basal third of intervals 3 to 6. Males with biseriate adhesive setae on apical half of mesotarsomere 1, and most of length of mesotarsomeres 2 and 3. Apex of abdominal sternite VII weakly emarginate in males. Median lobe of aedeagus slightly slender, gradually narrowed to apex, ventral margin nearly straight; right margin distinctly sinuate before apex in dorsal view; apical lamella wide and short, apex rounded, weakly bent upward in lateral view; endophallus identical to that of
<taxonomicName id="4C2EBCB6621FFFBA29FCB03CFB165F3B" authorityName="Motschulsky" authorityYear="1860" box="[1156,1265,583,609]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="B95A1B27621FFFBA29FCB03CFB165F3B" box="[1156,1265,583,609]" italics="true" pageId="89" pageNumber="90">P. bicolor</emphasis>
</taxonomicName>
. Gonocoxite II of ovipositor nearly quadrate, same length as basal width, apex strongly concave, with three ensiform setae on inner apical angle, and two on outer apical angle.
</paragraph>
</subSubSection>
<subSubSection id="C33494BE621FFFBA2DBFB0C9FA6C5F97" box="[199,1419,690,717]" pageId="89" pageNumber="90" type="distribution">
<paragraph id="8B91C735621FFFBA2DBFB0C9FA6C5F97" blockId="89.[151,1437,151,789]" box="[199,1419,690,717]" pageId="89" pageNumber="90">
<emphasis id="B95A1B27621FFFBA2DBFB0C9FEBE5F96" bold="true" box="[199,345,690,716]" pageId="89" pageNumber="90">Distribution</emphasis>
(
<figureCitation id="1315DBB0621FFFBA2C10B0C8FE535F97" box="[360,436,691,717]" captionStart="MAP 9" captionStartId="84.[152,211,1891,1915]" captionTargetBox="[151,1436,1041,1868]" captionTargetId="figure-773@84.[151,1436,1041,1868]" captionText="MAP 9. Distributions for species in the Parena (Parena) bicolor group." figureDoi="http://doi.org/10.5281/zenodo.7963679" httpUri="https://zenodo.org/record/7963679/files/figure.png" pageId="89" pageNumber="90">Map 9</figureCitation>
, green).
<collectingCountry id="F33987A5621FFFBA2F6EB0C8FDB55F97" box="[534,594,691,717]" name="India" pageId="89" pageNumber="90">India</collectingCountry>
,
<collectingCountry id="F33987A5621FFFBA2F25B0C8FD445F97" box="[605,675,691,717]" name="Nepal" pageId="89" pageNumber="90">Nepal</collectingCountry>
,
<collectingCountry id="F33987A5621FFFBA2FD5B0C8FCE85F97" box="[685,783,691,717]" name="Vietnam" pageId="89" pageNumber="90">Vietnam</collectingCountry>
,
<collectingCountry id="F33987A5621FFFBA2E62B0C8FC6E5F97" box="[794,905,691,717]" name="Myanmar" pageId="89" pageNumber="90">Myanmar</collectingCountry>
,
<collectingCountry id="F33987A5621FFFBA2EEBB0C8FC1E5F97" box="[915,1017,691,717]" name="Thailand" pageId="89" pageNumber="90">Thailand</collectingCountry>
,
<collectingCountry id="F33987A5621FFFBA297DB0C8FBAF5F97" box="[1029,1096,691,717]" name="China" pageId="89" pageNumber="90">China</collectingCountry>
(
<collectingRegion id="49EA09D7621FFFBA292FB0C8FB4B5F97" box="[1111,1196,691,717]" country="China" name="Xizang" pageId="89" pageNumber="90">Xizang</collectingRegion>
,
<collectingRegion id="49EA09D7621FFFBA29CEB0C8FAE85F97" box="[1206,1295,691,717]" country="China" name="Yunnan" pageId="89" pageNumber="90">Yunnan</collectingRegion>
,
<collectingRegion id="49EA09D7621FFFBA2861B0C8FA995F97" box="[1305,1406,691,717]" country="China" name="Guangxi" pageId="89" pageNumber="90">Guangxi</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C33494BE621FFFBA2DBFB0ACFA655E4F" pageId="89" pageNumber="90" type="discussion">
<paragraph id="8B91C735621FFFBA2DBFB0ACFA655E4F" blockId="89.[151,1437,151,789]" pageId="89" pageNumber="90">
<emphasis id="B95A1B27621FFFBA2DBFB0ACFED35FAB" bold="true" box="[199,308,727,753]" pageId="89" pageNumber="90">Remarks</emphasis>
. This species differs from
<taxonomicName id="4C2EBCB6621FFFBA2F1CB0ACFD365FAB" authorityName="Motschulsky" authorityYear="1860" box="[612,721,727,753]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="B95A1B27621FFFBA2F1CB0ACFD365FAB" box="[612,721,727,753]" italics="true" pageId="89" pageNumber="90">P. bicolor</emphasis>
</taxonomicName>
only in the color of elytra and subtle differences in the apex of male genitalia. Considering their allopatric distributions and stable differences, we treat them as distinct species.
</paragraph>
</subSubSection>
</treatment>
</document>