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<document id="D9A293F75A191D10209779494BF14BD3" ID-DOI="10.11646/zootaxa.5399.1.6" ID-ISSN="1175-5326" ID-Zenodo-Dep="10494478" ID-ZooBank="8B269581-73B6-472C-8379-C1367767D24F" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1705013715613" checkinUser="plazi" docAuthor="Nguyen, Hieu Van, Nguyen, Manh Thanh &amp; Nguyen, Lien Thi Phuong" docDate="2024" docId="03870712D2122E0BFF34FED33A7532A4" docLanguage="en" docName="zootaxa.5399.1.6.pdf" docOrigin="Zootaxa 5399 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5399.1.6" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Okinawepipona yty , Nguyen 2018" docType="treatment" docVersion="2" lastPageNumber="80" masterDocId="FFBE7F6AD2132E0AFFA3FFB538343557" masterDocTitle="Discovery of the male of Okinawepipona yty Nguyen, 2018 (Hymenoptera: Vespidae: Eumeninae) from Vietnam" masterLastPageNumber="84" masterPageNumber="79" pageNumber="80" updateTime="1705014392585" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="A8CD3EAE2C8CAD3AE9638DA053750D24" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="2FAF0CF7659D736AAF402944A82B68E2">Discovery of the male of Okinawepipona yty Nguyen, 2018 (Hymenoptera: Vespidae: Eumeninae) from Vietnam</mods:title>
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<mods:namePart id="11BEFEE2392AEC4994B309E8185A09CF">Nguyen, Hieu Van</mods:namePart>
<mods:affiliation id="994FEF9C4EF5378B9B1EC96AF087ACED">Faculty of Biology and Agricultural Engineering, Hanoi Pedagogical University 2, 32 Nguyen Van Linh Street, Phuc Yen city, Vinh Phuc province, Vietnam</mods:affiliation>
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<mods:namePart id="1A6222C1578E6E5DC5AFDE40058D453A">Nguyen, Manh Thanh</mods:namePart>
<mods:affiliation id="9278DF3261968F3ED0910A9E93E7C85D">Institute of Ecology and Biological Resources (IEBR), Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet Road, Nghia Do, Cau Giay, Hanoi, Vietnam</mods:affiliation>
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<mods:namePart id="1C2ACE5EDE3002D266DC90CE220306B9">Nguyen, Lien Thi Phuong</mods:namePart>
<mods:affiliation id="5D66B040D40C5E8C9A7E3C5ECB487C2D">Institute of Ecology and Biological Resources (IEBR), Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet Road, Nghia Do, Cau Giay, Hanoi, Vietnam &amp; Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet Road, Nghia Do, Cau Giay, Hanoi, Vietnam</mods:affiliation>
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<mods:title id="CCBD28FC29E27D67A68789F7B836530B">Zootaxa</mods:title>
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<mods:date id="F20D30D3FD9F3503FA71D9EDF02EC2C9">2024</mods:date>
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<mods:number id="79C3A3365BAF758F01818A469A39C53E">5399</mods:number>
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<treatment id="03870712D2122E0BFF34FED33A7532A4" LSID="urn:lsid:plazi:treatment:03870712D2122E0BFF34FED33A7532A4" httpUri="http://treatment.plazi.org/id/03870712D2122E0BFF34FED33A7532A4" lastPageNumber="80" pageId="1" pageNumber="80">
<subSubSection id="C334E58FD2122E0BFF34FED33A0534D7" box="[151,561,358,385]" pageId="1" pageNumber="80" type="nomenclature">
<paragraph id="8B91B604D2122E0BFF34FED33A0534D7" blockId="1.[151,561,358,421]" box="[151,561,358,385]" pageId="1" pageNumber="80">
<heading id="D0D90168D2122E0BFF34FED33A0534D7" bold="true" box="[151,561,358,385]" fontSize="11" level="1" pageId="1" pageNumber="80" reason="1">
<taxonomicName id="4C2ECD87D2122E0BFF34FED33A0534D7" authority="Nguyen, 2018" authorityName=", Nguyen" authorityYear="2018" box="[151,561,358,385]" class="Insecta" family="Eumenidae" genus="Okinawepipona" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="80" phylum="Arthropoda" rank="species" species="yty">
<emphasis id="B95A6A16D2122E0BFF34FED33A0534D7" bold="true" box="[151,561,358,385]" pageId="1" pageNumber="80">
<emphasis id="B95A6A16D2122E0BFF34FED3394B34D6" bold="true" box="[151,383,358,385]" italics="true" pageId="1" pageNumber="80">Okinawepipona yty</emphasis>
<bibRefCitation id="EFBFCBF5D2122E0BFE24FED23A0534D7" author="Nguyen, L. T. P. &amp; Nguyen, H. T. T. &amp; Bozdogan, H." box="[391,561,358,385]" pageId="1" pageNumber="80" pagination="592 - 596" refId="ref2638" refString="Nguyen, L. T. P., Nguyen, H. T. T. &amp; Bozdogan, H. (2018) Contribution to the genus Okinawepipona Yamane (Hymenoptera: Vespidae: Eumeninae) from Vietnam, with description of a new species. Zootaxa, 4462 (4), 592 - 596. https: // doi. org / 10.11646 / zootaxa. 4462.4.10" type="journal article" year="2018">Nguyen, 2018</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C334E58FD2122E0BFF34FE3E393434F2" box="[151,256,395,421]" pageId="1" pageNumber="80" type="description">
<paragraph id="8B91B604D2122E0BFF34FE3E393434F2" blockId="1.[151,561,358,421]" box="[151,256,395,421]" pageId="1" pageNumber="80">
<figureCitation id="1315AA81D2122E0BFF34FE3E393434F2" box="[151,256,395,421]" captionStart-0="FIGURES 1–4" captionStart-1="FIGURES 5–10" captionStartId-0="2.[152,270,1670,1694]" captionStartId-1="3.[152,270,1938,1962]" captionTargetBox-0="[158,1430,181,1641]" captionTargetBox-1="[423,1163,181,1919]" captionTargetId-0="figure-21@2.[158,1430,181,1652]" captionTargetId-1="figure-19@3.[423,1163,181,1919]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURES 1–4. Okinawepipona yty Nguyen, 2018, male. 1 Head, frontal view 2 Head, dorsal view 3 Antenna 4 Habitus. Scale bars = 1 mm." captionText-1="FIGURES 5–10. Okinawepipona yty Nguyen, 2018, male (5–7) and Okinawepipona curcipunctura Nguyen &amp; Xu, 2014, male (8–10): 5, 8 Genitalia, inner aspect of paramere with volsella and digitus 6, 9 Aedeagus, ventral view 7, 10 Aedeagus, lateral view. Scale bars = 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.10494480" figureDoi-1="http://doi.org/10.5281/zenodo.10494482" httpUri-0="https://zenodo.org/record/10494480/files/figure.png" httpUri-1="https://zenodo.org/record/10494482/files/figure.png" pageId="1" pageNumber="80">Figs. 1–7</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C334E58FD2122E0BFF34FE7A3DA63704" pageId="1" pageNumber="80" type="reference_group">
<paragraph id="8B91B604D2122E0BFF34FE7A3C6134B0" blockId="1.[151,1109,463,488]" box="[151,1109,463,488]" pageId="1" pageNumber="80">
<treatmentCitationGroup id="AB3E912AD2122E0BFF34FE7A3C6134B0" box="[151,1109,463,488]" pageId="1" pageNumber="80">
<treatmentCitation id="0A8F9015D2122E0BFF34FE7A3A1334B0" ID-DOI="http://doi.org/10.5281/zenodo.5988705" ID-Zenodo-Dep="5988705" author="Nguyen, L. T. P. &amp; Nguyen, H. T. T. &amp; Bozdogan, H." box="[151,551,463,488]" httpUri="http://treatment.plazi.org/id/7C783A27FFE90C44FF78F8C2ED418615" page="592" pageId="1" pageNumber="80" year="2018">
<taxonomicName id="4C2ECD87D2122E0BFF34FE7A3A1334B0" authority="Nguyen, 2018: 592" authorityName=", Nguyen" authorityPageNumber="592" authorityYear="2018" box="[151,551,463,488]" class="Insecta" family="Eumenidae" genus="Okinawepipona" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="80" phylum="Arthropoda" rank="species" species="yty">
<emphasis id="B95A6A16D2122E0BFF34FE7A396834B0" box="[151,348,463,487]" italics="true" pageId="1" pageNumber="80">Okinawepipona yty</emphasis>
<bibRefCitation id="EFBFCBF5D2122E0BFEC1FE653A1334B0" author="Nguyen, L. T. P. &amp; Nguyen, H. T. T. &amp; Bozdogan, H." box="[354,551,463,488]" pageId="1" pageNumber="80" pagination="592 - 596" refId="ref2638" refString="Nguyen, L. T. P., Nguyen, H. T. T. &amp; Bozdogan, H. (2018) Contribution to the genus Okinawepipona Yamane (Hymenoptera: Vespidae: Eumeninae) from Vietnam, with description of a new species. Zootaxa, 4462 (4), 592 - 596. https: // doi. org / 10.11646 / zootaxa. 4462.4.10" type="journal article" year="2018">Nguyen, 2018: 592</bibRefCitation>
</taxonomicName>
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, 596 (key), female – “Y Ty, Bat Xat,
<collectingRegion id="49EA78E6D2122E0BFC02FE653BC734B0" box="[929,1011,463,487]" country="Vietnam" name="Lao Cai" pageId="1" pageNumber="80">Lao Cai</collectingRegion>
” (IEBR).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B91B604D2122E0BFF34FDA03DA63704" blockId="1.[151,1437,533,2035]" pageId="1" pageNumber="80">
The male specimens we examined are from the
<typeStatus id="549508A6D2122E0BFD31FDA33AF53767" box="[658,705,534,560]" pageId="1" pageNumber="80">type</typeStatus>
locality and agree well with the description by
<bibRefCitation id="EFBFCBF5D2122E0BFB1BFDA33DAD3778" author="Nguyen, L. T. P. &amp; Nguyen, H. T. T. &amp; Bozdogan, H." box="[1208,1433,533,560]" pageId="1" pageNumber="80" pagination="592 - 596" refId="ref2638" refString="Nguyen, L. T. P., Nguyen, H. T. T. &amp; Bozdogan, H. (2018) Contribution to the genus Okinawepipona Yamane (Hymenoptera: Vespidae: Eumeninae) from Vietnam, with description of a new species. Zootaxa, 4462 (4), 592 - 596. https: // doi. org / 10.11646 / zootaxa. 4462.4.10" type="journal article" year="2018">
Nguyen
<emphasis id="B95A6A16D2122E0BFAB0FDA33D753778" box="[1299,1345,533,559]" italics="true" pageId="1" pageNumber="80">et al</emphasis>
. (2018)
</bibRefCitation>
. As described below, the male is similar to the female both in structure and coloration except for some few characters.
</paragraph>
</subSubSection>
<subSubSection id="C334E58FD2122E0BFF64FDE83BBA3797" pageId="1" pageNumber="80" type="materials_examined">
<paragraph id="8B91B604D2122E0BFF64FDE83BBA3797" blockId="1.[151,1437,533,2035]" pageId="1" pageNumber="80">
<emphasis id="B95A6A16D2122E0BFF64FDE839843720" bold="true" box="[199,432,605,631]" pageId="1" pageNumber="80">Material examined.</emphasis>
<materialsCitation id="3B46BC59D2122E0BFE14FDEB3AB437CC" collectingDate="2019-05-06" collectorName="Y Ty &amp; Bat Xat &amp; Nguyen Quang Cuong" country="Vietnam" elevation="1869" latitude="22.608194" location="Bat Xat" longLatPrecision="1" longitude="103.62489" pageId="1" pageNumber="80" specimenCount="12" specimenCount-female="12" stateProvince="Lao Cai">
<collectingCountry id="F339F694D2122E0BFE14FDEB3A09372F" box="[439,573,606,632]" name="Vietnam" pageId="1" pageNumber="80">VIETNAM</collectingCountry>
:
<emphasis id="B95A6A16D2122E0BFDE4FDE83A9B3720" bold="true" box="[583,687,605,631]" pageId="1" pageNumber="80">
<collectingRegion id="49EA78E6D2122E0BFDE4FDE83A9E3720" box="[583,682,605,631]" country="Vietnam" name="Lao Cai" pageId="1" pageNumber="80">Lao Cai</collectingRegion>
:
</emphasis>
<specimenCount id="9D287D8DD2122E0BFD15FDE83ADA372F" box="[694,750,605,632]" count="12" pageId="1" pageNumber="80" type="female">12 ♀</specimenCount>
, 16 ♁,
<collectorName id="26DBD3D2D2122E0BFCE3FDEB3B4C372F" box="[832,888,606,632]" pageId="1" pageNumber="80">Y Ty</collectorName>
,
<collectorName id="26DBD3D2D2122E0BFC20FDEB3BEF372F" box="[899,987,606,632]" pageId="1" pageNumber="80">
<location id="8EF1E0DFD2122E0BFC20FDEB3BEF372F" LSID="urn:lsid:plazi:treatment:03870712D2122E0BFF34FED33A7532A4:8EF1E0DFD2122E0BFC20FDEB3BEF372F" box="[899,987,606,632]" country="Vietnam" latitude="22.608194" longLatPrecision="1" longitude="103.62489" name="Bat Xat" pageId="1" pageNumber="80" stateProvince="Lao Cai">Bat Xat</location>
</collectorName>
,
<geoCoordinate id="EE1AD0C3D2122E0BFC44FDEA3CB5372D" box="[999,1153,607,634]" degrees="22" direction="north" minutes="36" orientation="latitude" pageId="1" pageNumber="80" precision="1" seconds="29.5" value="22.608194">22°36′29.5″N</geoCoordinate>
<geoCoordinate id="EE1AD0C3D2122E0BFB2BFDEA3D1B372D" box="[1160,1327,607,634]" degrees="103" direction="east" minutes="37" orientation="longitude" pageId="1" pageNumber="80" precision="1" seconds="29.6" value="103.62489">103°37′29.6″E</geoCoordinate>
, alt.
<quantity id="4CD61BE1D2122E0BFAC7FDE8389B37CB" metricMagnitude="3" metricUnit="m" metricValue="1.869" pageId="1" pageNumber="80" unit="m" value="1869.0">
<elevation id="00035137D2122E0BFAC7FDE8389B37CB" metricMagnitude="3" metricUnit="m" metricValue="1.869" pageId="1" pageNumber="80" unit="m" value="1869.0">1869 m</elevation>
</quantity>
,
<date id="FF9090C4D2122E0BFF18FD34397137CC" box="[187,325,641,668]" pageId="1" pageNumber="80" value="2019-05-06">
<collectingDate id="EFD4692CD2122E0BFF18FD34397137CC" box="[187,325,641,668]" pageId="1" pageNumber="80" value="2019-05-06">6 May 2019</collectingDate>
</date>
,
<collectorName id="26DBD3D2D2122E0BFEF3FD373A7937CC" box="[336,589,641,668]" pageId="1" pageNumber="80">Nguyen Quang Cuong</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B46BC59D2122E0BFD28FD343BBA3797" collectingDate="2023-07-15" collectionCode="IEBR" collectorName="Y Ty &amp; Bat Xat &amp; Nguyen Quang Cuong" country="Vietnam" elevation="1850" latitude="22.620695" location="Bat Xat" longLatPrecision="1" longitude="103.62375" pageId="1" pageNumber="80" specimenCount="1" stateProvince="Lao Cai">
3 ♁,
<collectorName id="26DBD3D2D2122E0BFD61FD373ACE37CB" box="[706,762,642,668]" pageId="1" pageNumber="80">Y Ty</collectorName>
,
<collectorName id="26DBD3D2D2122E0BFCA6FD373B6A37CB" box="[773,862,642,668]" pageId="1" pageNumber="80">
<location id="8EF1E0DFD2122E0BFCA6FD373B6A37CB" LSID="urn:lsid:plazi:treatment:03870712D2122E0BFF34FED33A7532A4:8EF1E0DFD2122E0BFCA6FD373B6A37CB" box="[773,862,642,668]" country="Vietnam" latitude="22.620695" longLatPrecision="1" longitude="103.62375" name="Bat Xat" pageId="1" pageNumber="80" stateProvince="Lao Cai">Bat Xat</location>
</collectorName>
,
<geoCoordinate id="EE1AD0C3D2122E0BFCC9FD363C3737C9" box="[874,1027,643,670]" degrees="22" direction="north" minutes="37" orientation="latitude" pageId="1" pageNumber="80" precision="1" seconds="14.5" value="22.620695">22°37′14.5″N</geoCoordinate>
<geoCoordinate id="EE1AD0C3D2122E0BFBA8FD363C8637C9" box="[1035,1202,643,670]" degrees="103" direction="east" minutes="37" orientation="longitude" pageId="1" pageNumber="80" precision="1" seconds="25.5" value="103.62375">103°37′25.5″E</geoCoordinate>
, alt.
<quantity id="4CD61BE1D2122E0BFB4BFD343D0B37CB" box="[1256,1343,641,668]" metricMagnitude="3" metricUnit="m" metricValue="1.85" pageId="1" pageNumber="80" unit="m" value="1850.0">
<elevation id="00035137D2122E0BFB4BFD343D0B37CB" box="[1256,1343,641,668]" metricMagnitude="3" metricUnit="m" metricValue="1.85" pageId="1" pageNumber="80" unit="m" value="1850.0">1850 m</elevation>
</quantity>
,
<date id="FF9090C4D2122E0BFAE8FD3438E637E8" pageId="1" pageNumber="80" value="2023-07-15">
<collectingDate id="EFD4692CD2122E0BFAE8FD3438E637E8" pageId="1" pageNumber="80" value="2023-07-15">15 July 2023</collectingDate>
</date>
, Nguyen Thi Phuong Lien,
<collectorName id="26DBD3D2D2122E0BFDABFD133B3037E8" box="[520,772,677,704]" pageId="1" pageNumber="80">Nguyen Quang Cuong</collectorName>
leg. [
<collectionCode id="ED3F2EC1D2122E0BFCE1FD133BB33797" box="[834,903,678,704]" pageId="1" pageNumber="80">IEBR</collectionCode>
]
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C334E58FD2122E0BFF64FD7C39CA3298" pageId="1" pageNumber="80" type="description">
<paragraph id="8B91B604D2122E0BFF64FD7C39B630AC" blockId="1.[151,1437,533,2035]" pageId="1" pageNumber="80">
<emphasis id="B95A6A16D2122E0BFF64FD7C396D37B4" bold="true" box="[199,345,713,739]" pageId="1" pageNumber="80">Description.</emphasis>
<emphasis id="B95A6A16D2122E0BFEC1FD7C39A837B4" box="[354,412,713,739]" italics="true" pageId="1" pageNumber="80">Male</emphasis>
(
<figureCitation id="1315AA81D2122E0BFE0EFD7C39C737B4" box="[429,499,713,739]" captionStart="FIGURES 1–4" captionStartId="2.[152,270,1670,1694]" captionTargetBox="[158,1430,181,1641]" captionTargetId="figure-21@2.[158,1430,181,1652]" captionTargetPageId="2" captionText="FIGURES 1–4. Okinawepipona yty Nguyen, 2018, male. 1 Head, frontal view 2 Head, dorsal view 3 Antenna 4 Habitus. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10494480" httpUri="https://zenodo.org/record/10494480/files/figure.png" pageId="1" pageNumber="80">Fig. 4</figureCitation>
) [female characters in square brackets]. Body length
<quantity id="4CD61BE1D2122E0BFBFEFD7C3CEE37B3" box="[1117,1242,713,740]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.2" metricValueMin="1.1" pageId="1" pageNumber="80" unit="mm" value="11.5" valueMax="12.0" valueMin="11.0">11–12 mm</quantity>
; forewing length
<quantity id="4CD61BE1D2122E0BFF34FD583920365F" box="[151,276,749,776]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.2" metricValueMin="1.1" pageId="1" pageNumber="80" unit="mm" value="11.5" valueMax="12.0" valueMin="11.0">11–12 mm</quantity>
. Head in frontal view subcircular, wider than high, about 1.2× as wide as high [1.1× as wide as high] (
<figureCitation id="1315AA81D2122E0BFF3CFCA438D5367C" box="[159,225,785,811]" captionStart="FIGURES 1–4" captionStartId="2.[152,270,1670,1694]" captionTargetBox="[158,1430,181,1641]" captionTargetId="figure-21@2.[158,1430,181,1652]" captionTargetPageId="2" captionText="FIGURES 1–4. Okinawepipona yty Nguyen, 2018, male. 1 Head, frontal view 2 Head, dorsal view 3 Antenna 4 Habitus. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10494480" httpUri="https://zenodo.org/record/10494480/files/figure.png" pageId="1" pageNumber="80">Fig. 1</figureCitation>
). Vertex without cephalic foveae [with cephalic foveae small, bearing dense pubescence, situated very close to each other, almost touching each other] (
<figureCitation id="1315AA81D2122E0BFDD5FC803A8F3618" box="[630,699,821,847]" captionStart="FIGURES 1–4" captionStartId="2.[152,270,1670,1694]" captionTargetBox="[158,1430,181,1641]" captionTargetId="figure-21@2.[158,1430,181,1652]" captionTargetPageId="2" captionText="FIGURES 1–4. Okinawepipona yty Nguyen, 2018, male. 1 Head, frontal view 2 Head, dorsal view 3 Antenna 4 Habitus. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10494480" httpUri="https://zenodo.org/record/10494480/files/figure.png" pageId="1" pageNumber="80">Fig. 2</figureCitation>
). Distance from posterior ocellus to apical margin of vertex nearly 1.6× distance from posterior ocellus to inner compound eye margin [more than 1.7× distance from posterior ocellus to inner compound eye margin] (
<figureCitation id="1315AA81D2122E0BFDB0FCC83A6F36C0" box="[531,603,893,919]" captionStart="FIGURES 1–4" captionStartId="2.[152,270,1670,1694]" captionTargetBox="[158,1430,181,1641]" captionTargetId="figure-21@2.[158,1430,181,1652]" captionTargetPageId="2" captionText="FIGURES 1–4. Okinawepipona yty Nguyen, 2018, male. 1 Head, frontal view 2 Head, dorsal view 3 Antenna 4 Habitus. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10494480" httpUri="https://zenodo.org/record/10494480/files/figure.png" pageId="1" pageNumber="80">Fig. 2</figureCitation>
). Gena slightly narrower than compound eye, in lateral view about 0.9× as wide as compound eye [much narrower than compound eye, in lateral view about 0.7× as wide as compound eye]. Occipital carina complete, present along entire length of gena, but dorsally somewhat weak. Inner compound eye margins strongly converging ventrally; in frontal view about 1.8× further apart from each other at vertex than at clypeus [1.3× further apart from each other at vertex than at clypeus]. Disc of clypeus in lateral view weakly convex at basal half, then straight to near apical margin; in frontal view 1.3× higher than wide [about as wide as high] (
<figureCitation id="1315AA81D2122E0BFF41FBE039103138" box="[226,292,1109,1135]" captionStart="FIGURES 1–4" captionStartId="2.[152,270,1670,1694]" captionTargetBox="[158,1430,181,1641]" captionTargetId="figure-21@2.[158,1430,181,1652]" captionTargetPageId="2" captionText="FIGURES 1–4. Okinawepipona yty Nguyen, 2018, male. 1 Head, frontal view 2 Head, dorsal view 3 Antenna 4 Habitus. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10494480" httpUri="https://zenodo.org/record/10494480/files/figure.png" pageId="1" pageNumber="80">Fig. 1</figureCitation>
), with basal margin almost straight [slightly convex medially] and distinctly separated from antennal sockets; apical margin deeply emarginate medially, forming very sharp tooth on each lateral side [forming sharp tooth on each lateral side] (
<figureCitation id="1315AA81D2122E0BFE64FB283A3C31E0" box="[455,520,1181,1207]" captionStart="FIGURES 1–4" captionStartId="2.[152,270,1670,1694]" captionTargetBox="[158,1430,181,1641]" captionTargetId="figure-21@2.[158,1430,181,1652]" captionTargetPageId="2" captionText="FIGURES 1–4. Okinawepipona yty Nguyen, 2018, male. 1 Head, frontal view 2 Head, dorsal view 3 Antenna 4 Habitus. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10494480" httpUri="https://zenodo.org/record/10494480/files/figure.png" pageId="1" pageNumber="80">Fig. 1</figureCitation>
), without carina [with two faint carinae running from tooth at apical point to base direction]; width of emargination slightly greater than 1/3 width of clypeus between inner compound eye margins [width of emargination slightly less than 1/3 width of clypeus between inner compound eye margins]. Mandible with prominent teeth, second and third teeth with inner side almost straight [second and third teeth with inner side produced with round margin], fourth tooth pointed apically. Antennal scape about 3.3× as long as its maximum width [about 3.7× as long as its maximum width]; flagellomere I about 1.8× as long as wide [about 1.5× as long as wide], flagellomeres II and III slightly longer than wide [flagellomere III as wide as long], flagellomere IV as wide as long, flagellomeres V–IX wider than long, terminal flagellomere small, slightly curved, 3.5× as long as its basal width, reaching to near base of flagellomere IX when folded [terminal flagellomere bullet-shaped, as long as its basal width] (
<figureCitation id="1315AA81D2122E0BFE93FA54394130AC" box="[304,373,1505,1531]" captionStart="FIGURES 1–4" captionStartId="2.[152,270,1670,1694]" captionTargetBox="[158,1430,181,1641]" captionTargetId="figure-21@2.[158,1430,181,1652]" captionTargetPageId="2" captionText="FIGURES 1–4. Okinawepipona yty Nguyen, 2018, male. 1 Head, frontal view 2 Head, dorsal view 3 Antenna 4 Habitus. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10494480" httpUri="https://zenodo.org/record/10494480/files/figure.png" pageId="1" pageNumber="80">Fig. 3</figureCitation>
).
</paragraph>
<paragraph id="8B91B604D2122E0BFF64F9B33C4133DB" blockId="1.[151,1437,533,2035]" pageId="1" pageNumber="80">Mesosoma, metasoma (except tergum VII and sternum VII), and body sculpture as same as in female except clypeus with dense, small, deep punctures [clypeus with dense, large, flat-bottomed punctures, each bearing silver bristle, punctures at center larger than at sides, space between punctures larger than puncture diameter]. Tergum VII and sternum VII with some small and sparse punctures come between minute punctures.</paragraph>
<paragraph id="8B91B604D2122E0BFF64F9203BD833A0" blockId="1.[151,1437,533,2035]" pageId="1" pageNumber="80">
<emphasis id="B95A6A16D2122E0BFF64F920393C33F8" box="[199,264,1685,1711]" italics="true" pageId="1" pageNumber="80">Color</emphasis>
. Black; similar to female except clypeus almost entirely yellow [large spots on upper lateral corner and a large spot on lower middle of clypeus]; head black [narrow band along inner compound eye margin extending from bottom of frons nearly to ocular sinus, large spot between antennal sockets].
</paragraph>
<paragraph id="8B91B604D2122E0BFF64F8B439CA3298" blockId="1.[151,1437,533,2035]" pageId="1" pageNumber="80">
<emphasis id="B95A6A16D2122E0BFF64F8B43905324C" box="[199,305,1793,1819]" italics="true" pageId="1" pageNumber="80">Genitalia</emphasis>
. As in
<figureCitation id="1315AA81D2122E0BFEDEF8B439D3324B" box="[381,487,1793,1820]" captionStart="FIGURES 5–10" captionStartId="3.[152,270,1938,1962]" captionTargetBox="[423,1163,181,1919]" captionTargetId="figure-19@3.[423,1163,181,1919]" captionTargetPageId="3" captionText="FIGURES 5–10. Okinawepipona yty Nguyen, 2018, male (5–7) and Okinawepipona curcipunctura Nguyen &amp; Xu, 2014, male (8–10): 5, 8 Genitalia, inner aspect of paramere with volsella and digitus 6, 9 Aedeagus, ventral view 7, 10 Aedeagus, lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10494482" httpUri="https://zenodo.org/record/10494482/files/figure.png" pageId="1" pageNumber="80">Figs. 5–7</figureCitation>
. Parameral spine lacking setae. Volsella flattened, spatulate, wide on inner aspect, and without setae at top (
<figureCitation id="1315AA81D2122E0BFE23F89039F03217" box="[384,452,1829,1856]" captionStart="FIGURES 5–10" captionStartId="3.[152,270,1938,1962]" captionTargetBox="[423,1163,181,1919]" captionTargetId="figure-19@3.[423,1163,181,1919]" captionTargetPageId="3" captionText="FIGURES 5–10. Okinawepipona yty Nguyen, 2018, male (5–7) and Okinawepipona curcipunctura Nguyen &amp; Xu, 2014, male (8–10): 5, 8 Genitalia, inner aspect of paramere with volsella and digitus 6, 9 Aedeagus, ventral view 7, 10 Aedeagus, lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10494482" httpUri="https://zenodo.org/record/10494482/files/figure.png" pageId="1" pageNumber="80">Fig. 5</figureCitation>
). Digitus knife-shaped, almost parallel at one-third from base, then gradually narrowing to top, with medium long setae on outer margin (
<figureCitation id="1315AA81D2122E0BFD1AF8FC3AC93233" box="[697,765,1865,1892]" captionStart="FIGURES 5–10" captionStartId="3.[152,270,1938,1962]" captionTargetBox="[423,1163,181,1919]" captionTargetId="figure-19@3.[423,1163,181,1919]" captionTargetPageId="3" captionText="FIGURES 5–10. Okinawepipona yty Nguyen, 2018, male (5–7) and Okinawepipona curcipunctura Nguyen &amp; Xu, 2014, male (8–10): 5, 8 Genitalia, inner aspect of paramere with volsella and digitus 6, 9 Aedeagus, ventral view 7, 10 Aedeagus, lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10494482" httpUri="https://zenodo.org/record/10494482/files/figure.png" pageId="1" pageNumber="80">Fig. 5</figureCitation>
). Penis valves of aedeagus long, about 1.6× as long as basal apodeme, in ventral view proximal part produced laterally into a round lobe laterally with blunt apex in inner margin apically (
<figureCitation id="1315AA81D2122E0BFF5CF824397632FC" box="[255,322,1937,1963]" captionStart="FIGURES 5–10" captionStartId="3.[152,270,1938,1962]" captionTargetBox="[423,1163,181,1919]" captionTargetId="figure-19@3.[423,1163,181,1919]" captionTargetPageId="3" captionText="FIGURES 5–10. Okinawepipona yty Nguyen, 2018, male (5–7) and Okinawepipona curcipunctura Nguyen &amp; Xu, 2014, male (8–10): 5, 8 Genitalia, inner aspect of paramere with volsella and digitus 6, 9 Aedeagus, ventral view 7, 10 Aedeagus, lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10494482" httpUri="https://zenodo.org/record/10494482/files/figure.png" pageId="1" pageNumber="80">Fig. 6</figureCitation>
); in profile apical part produced into a round projection (
<figureCitation id="1315AA81D2122E0BFC1EF8243C3532FC" box="[957,1025,1937,1963]" captionStart="FIGURES 5–10" captionStartId="3.[152,270,1938,1962]" captionTargetBox="[423,1163,181,1919]" captionTargetId="figure-19@3.[423,1163,181,1919]" captionTargetPageId="3" captionText="FIGURES 5–10. Okinawepipona yty Nguyen, 2018, male (5–7) and Okinawepipona curcipunctura Nguyen &amp; Xu, 2014, male (8–10): 5, 8 Genitalia, inner aspect of paramere with volsella and digitus 6, 9 Aedeagus, ventral view 7, 10 Aedeagus, lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10494482" httpUri="https://zenodo.org/record/10494482/files/figure.png" pageId="1" pageNumber="80">Fig. 7</figureCitation>
); dorsal rod of aedeagus shorter than basal apodeme apically (
<figureCitation id="1315AA81D2122E0BFE0FF80039C43298" box="[428,496,1973,1999]" captionStart="FIGURES 5–10" captionStartId="3.[152,270,1938,1962]" captionTargetBox="[423,1163,181,1919]" captionTargetId="figure-19@3.[423,1163,181,1919]" captionTargetPageId="3" captionText="FIGURES 5–10. Okinawepipona yty Nguyen, 2018, male (5–7) and Okinawepipona curcipunctura Nguyen &amp; Xu, 2014, male (8–10): 5, 8 Genitalia, inner aspect of paramere with volsella and digitus 6, 9 Aedeagus, ventral view 7, 10 Aedeagus, lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10494482" httpUri="https://zenodo.org/record/10494482/files/figure.png" pageId="1" pageNumber="80">Fig. 7</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C334E58FD2122E0BFF64F86C3A7532A4" box="[199,577,2009,2035]" pageId="1" pageNumber="80" type="distribution">
<paragraph id="8B91B604D2122E0BFF64F86C3A7532A4" blockId="1.[151,1437,533,2035]" box="[199,577,2009,2035]" pageId="1" pageNumber="80">
<emphasis id="B95A6A16D2122E0BFF64F86C395532A4" bold="true" box="[199,353,2009,2035]" pageId="1" pageNumber="80">Distribution.</emphasis>
<collectingCountry id="F339F694D2122E0BFEC4F86C39F232A4" box="[359,454,2009,2035]" name="Vietnam" pageId="1" pageNumber="80">Vietnam</collectingCountry>
(
<collectingRegion id="49EA78E6D2122E0BFE76F86C3A0032A4" box="[469,564,2009,2035]" country="Vietnam" name="Lao Cai" pageId="1" pageNumber="80">Lao Cai</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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