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<mods:title id="87326020A46058FEE70EA298845D4681">A revision of the extant species of Theodoxus (Gastropoda, Neritidae) in Asia, with the description of three new species</mods:title>
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<treatment id="F1893B31F4125F84A45984E956D005F6" ID-GBIF-Taxon="161939040" LSID="urn:lsid:plazi:treatment:F1893B31F4125F84A45984E956D005F6" httpUri="http://treatment.plazi.org/id/F1893B31F4125F84A45984E956D005F6" lastPageNumber="25" pageId="0" pageNumber="25" scope_class="Gastropoda" scope_family="Neritidae" scope_subFamily="Neritininae" scope_superFamily="Neritoidea">
<subSubSection id="FBEEB63AB30DC7013971B301EB2E1A0F" pageId="0" pageNumber="25" type="nomenclature">
<paragraph id="B75CF21DE706B2CE0B253401514B3C53" pageId="0" pageNumber="25">
<taxonomicName id="4E9AA4DA15FCC2EC78F36E2715F8A568" ID-CoL="56C5W" LSID="F1893B31-F412-5F84-A459-84E956D005F6" authority="(Linnaeus, 1758)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus fluviatilis" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="fluviatilis">Theodoxus fluviatilis (Linnaeus, 1758)</taxonomicName>
<figureCitation id="8A94825D1A93E40A612BC689ABBD7481" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Theodoxus fluviatilis (Linnaeus, 1758). A - D. Specimen collected close to Oued Laabid, Morocco (UGSB 18106) (Fig. 2); E - H. Specimen from the Danube River, Woerth an der Donau, Germany (UGSB 24173); I - L. Specimen (T. euxinus - morphotype) collected in the Papuc Creek, Kiyikoey, Turkey (UGSB 24172) (Fig. 2); M - P. Specimen (T. euxinus - morphotype) from Ovidiopol, Ukraine (UGSB 18124); Q - S. Specimen (T. heldreichi - morphotype) collected in Lake Egirdir, Turkey (UGSB 20361); T, U. Specimen (T. danasteri - morphotype) collected in Krasna Kosa, Ukraine (UGSB 18417) (Fig. 2); V. Opercula of a T. danasteri - morphotype from Lake Skadar, Montenegro (UGSB 24171). Specimens A-D, I-L and T-U were used in the phylogeny (Fig. 2). Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure9" httpUri="https://binary.pensoft.net/fig/376458" pageId="0" pageNumber="25">Figures 9A-S</figureCitation>
<figureCitation id="788DEB6172A132A8764EA23FB0C37B2E" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Theodoxus fluviatilis (Linnaeus, 1758). A - D. Specimen (conforming to T. sarmaticus) from the Dnieper-Bug estuary, Ukraine (UGSB 18328); E - H. Specimen (T. subthermalis - morphotype) from Novy Afon, Abkhazia, Georgia (UGSB 18402); I - K. Syntype of N. saulcyi from near the Penteli Monastery, Greece (MHNG-MOLL- 111736); L. Specimen (T. saulcyi - morphotype) from Stilos, Crete, Greece (UGSB 20519) incorporated into the phylogeny (Fig. 2). Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure10" httpUri="https://binary.pensoft.net/fig/376459" pageId="0" pageNumber="25">, 10A-L</figureCitation>
</paragraph>
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<taxonomicName id="45475FEDC2B4154E75D68986938639DB" authorityName="Linnaeus" authorityYear="1758" class="Gastropoda" family="Neritidae" genus="Nerita" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nerita fluviatilis" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="fluviatilis">Nerita fluviatilis</taxonomicName>
<bibRefCitation id="C3130712430BC28CF7C2F580E1EF4700" author="Linnaeus, C" journalOrPublisher="Laurentius Salvius, Holmiae" pageId="0" pageNumber="25" publicationUrl="https://biodiversitylibrary.org/page/726886" refId="B93" refString="Linnaeus, C, 1758. Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Editio decima, reformata. Laurentius Salvius, Holmiae, https://biodiversitylibrary.org/page/726886" title="Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Editio decima, reformata." url="https://biodiversitylibrary.org/page/726886" year="1758">Linnaeus 1758</bibRefCitation>
: 777.
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<paragraph id="ACFCD1067252C4151075E7073F288057" pageId="0" pageNumber="25">
<taxonomicName id="317B07F4D47EB9F198C664D36CE1A3F3" authorityName="Montfort" authorityYear="1810" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus lutetianus" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="lutetianus">Theodoxus lutetianus</taxonomicName>
<bibRefCitation id="B178966275E25E24526850690C526C45" author="Montfort, PDd" journalOrPublisher="Schoell, Paris" pageId="0" pageNumber="25" publicationUrl="https://biodiversitylibrary.org/page/11065017" refId="B107" refString="Montfort, PDd, 1810. Conchyliologie systematique et classification methodique de coquilles; offrant leurs figures, leur arrangement generique, leurs descriptions caracteristiques, leurs noms; ainsi que leur synonymie en plusieurs langues. Ouvrage destine a faciliter l'etude des coquilles, ainsi que leur disposition dans les cabinets d'histoire naturelle. Coquilles univalves, non cloisonnees. Tome second. Schoell, Paris, https://biodiversitylibrary.org/page/11065017" title="Conchyliologie systematique et classification methodique de coquilles; offrant leurs figures, leur arrangement generique, leurs descriptions caracteristiques, leurs noms; ainsi que leur synonymie en plusieurs langues. Ouvrage destine a faciliter l'etude des coquilles, ainsi que leur disposition dans les cabinets d'histoire naturelle. Coquilles univalves, non cloisonnees. Tome second." url="https://biodiversitylibrary.org/page/11065017" year="1810">Montfort 1810</bibRefCitation>
: 351-352.
</paragraph>
<paragraph id="91F6D4A9047129384CB4D3F5B3045E34" pageId="0" pageNumber="25">
<taxonomicName id="061E47BA548AC691DA18A26A6A24937B" class="Gastropoda" family="Neritidae" genus="Neritina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neritina" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="genus">Neritina</taxonomicName>
Saulcyi
<bibRefCitation id="B6BF50459C04E992F2C4604447B0F54D" author="Bourguignat, JR" journalOrPublisher="Bailliere, Lutetia" pageId="0" pageNumber="25" publicationUrl="https://gallica.bnf.fr/ark:/12148/bpt6k99037b.image" refId="B30" refString="Bourguignat, JR, 1852. Testacea novissima quae Cl. de Saulcy in itinere per Orientem annis 1850 et 1851, collegit. J.-B. Bailliere, Lutetia, https://gallica.bnf.fr/ark:/12148/bpt6k99037b.image" title="Testacea novissima quae Cl. de Saulcy in itinere per Orientem annis 1850 et 1851, collegit. J. - B." url="https://gallica.bnf.fr/ark:/12148/bpt6k99037b.image" year="1852">Bourguignat 1852</bibRefCitation>
: 25-26.
</paragraph>
<paragraph id="9757A9F0E8A8DC64188D4161B49F0D0E" pageId="0" pageNumber="25">
<taxonomicName id="30B1182B3C523DDD954B4D4C235C4D06" authorityName="Issel" authorityYear="1865" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus fluviatilis var. subthermalis" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="variety" species="fluviatilis" variety="subthermalis">Theodoxus fluviatilis var. subthermalis</taxonomicName>
<bibRefCitation id="8003350852011F13DF1FF5F04C557C53" author="Issel, A" journalOrPublisher="Stamperia Reale, Torino" pageId="0" pageNumber="25" publicationUrl="http://magteca-fi-ese.inera.it/unifi/opac/unifi/scheda.jsp?pid=mag:13023" refId="B72" refString="Issel, A, 1865. Catalogo dei molluschi raccolti dalla missione italiana in Persia aggiuntavi la descrizione delle specie nuove o poco note. Stamperia Reale, Torino, http://magteca-fi-ese.inera.it/unifi/opac/unifi/scheda.jsp?pid=mag:13023" title="Catalogo dei molluschi raccolti dalla missione italiana in Persia aggiuntavi la descrizione delle specie nuove o poco note." url="http://magteca-fi-ese.inera.it/unifi/opac/unifi/scheda.jsp?pid=mag:13023" year="1865">Issel 1865</bibRefCitation>
: 22-23;
<bibRefCitation id="89064B224F6665995E74E67F357EB2DD" author="Issel, A" journalOrPublisher="serie" pageId="0" pageNumber="25" pagination="387 - 439" publicationUrl="https://biodiversitylibrary.org/page/31197934" refId="B73" refString="Issel, A, 1866. Catalogo dei molluschi raccolti dalla missione italiana in Persia aggiuntavi la descrizione delle specie nuove o poco note. Memorie della Reale accademia delle scienze di Torino, 2. serie 23: 387 - 439, https://biodiversitylibrary.org/page/31197934" title="Catalogo dei molluschi raccolti dalla missione italiana in Persia aggiuntavi la descrizione delle specie nuove o poco note. Memorie della Reale accademia delle scienze di Torino, 2." url="https://biodiversitylibrary.org/page/31197934" volume="23" year="1866">Issel 1866</bibRefCitation>
: 406-407.
</paragraph>
<paragraph id="57BF62A7F4677217CAF172C46E1DCE29" pageId="0" pageNumber="25">
<taxonomicName id="4C770541C2BBFAAB80722B99EE1B5373" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Gastropoda" family="Neritidae" genus="Neritina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neritina fluviatilis" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="fluviatilis">Neritina fluviatilis</taxonomicName>
:
<bibRefCitation id="8787034C32FF722873985B6CF89F40BB" author="Martens, Ev" journalOrPublisher="Bauer &amp; Raspe, Nuernberg" pageId="0" pageNumber="25" publicationUrl="https://biodiversitylibrary.org/page/52484536" refId="B99" refString="Martens, Ev, 1879. Die Gattung Neritina. - Systematisches Conchylien-Cabinet von Martini und Chemnitz 2 (10). Bauer &amp; Raspe, Nuernberg, https://biodiversitylibrary.org/page/52484536" title="Die Gattung Neritina. - Systematisches Conchylien-Cabinet von Martini und Chemnitz 2 (10)." url="https://biodiversitylibrary.org/page/52484536" year="1879">Martens 1879</bibRefCitation>
: 204-221, pl. 21, figs 1-6;
<bibRefCitation id="E3786978E5DA8B5056F4179CA82D6324" author="Kobelt, W" journalOrPublisher="Aquila" pageId="0" pageNumber="25" publicationUrl="https://biodiversitylibrary.org/page/16299729" refId="B83" refString="Kobelt, W, 1899. Iconographie der Land- &amp; Suesswasser-Mollusken mit vorzueglicher Beruecksichtigung der europaeischen noch nicht abgebildeten Arten von E. A. Rossmaessler. VIII. Band. C. W. Kreidel, Wiesbaden, 1-11. [pls 211-240] https://biodiversitylibrary.org/page/16299729" title="Iconographie der Land- &amp; Suesswasser-Mollusken mit vorzueglicher Beruecksichtigung der europaeischen noch nicht abgebildeten Arten von E. A. Rossmaessler. VIII. Band. C. W. Kreidel, Wiesbaden, 1 - 11. [pls 211 - 240]" url="https://biodiversitylibrary.org/page/16299729" year="1899">Kobelt 1899</bibRefCitation>
: 16-18, pl. 215, figs 1359-1365.
</paragraph>
<paragraph id="D1DCB81650635F7B9EF1FFA8D2D26A01" pageId="0" pageNumber="25">
<taxonomicName id="06F3868BEF2B519D6766171334D4452A" class="Gastropoda" family="Neritidae" genus="Neritina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neritina" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="genus">Neritina</taxonomicName>
Heldreichi
<bibRefCitation id="30707459164915C87A9A6A87FEAE2464" author="Martens, Ev" journalOrPublisher="Bauer &amp; Raspe, Nuernberg" pageId="0" pageNumber="25" publicationUrl="https://biodiversitylibrary.org/page/52484536" refId="B99" refString="Martens, Ev, 1879. Die Gattung Neritina. - Systematisches Conchylien-Cabinet von Martini und Chemnitz 2 (10). Bauer &amp; Raspe, Nuernberg, https://biodiversitylibrary.org/page/52484536" title="Die Gattung Neritina. - Systematisches Conchylien-Cabinet von Martini und Chemnitz 2 (10)." url="https://biodiversitylibrary.org/page/52484536" year="1879">Martens 1879</bibRefCitation>
: 225, pl. 22, figs 9, 10;
<bibRefCitation id="00A73251F9FCA374E70603414599F0E2" author="Kobelt, W" journalOrPublisher="Aquila" pageId="0" pageNumber="25" publicationUrl="https://biodiversitylibrary.org/page/16299729" refId="B83" refString="Kobelt, W, 1899. Iconographie der Land- &amp; Suesswasser-Mollusken mit vorzueglicher Beruecksichtigung der europaeischen noch nicht abgebildeten Arten von E. A. Rossmaessler. VIII. Band. C. W. Kreidel, Wiesbaden, 1-11. [pls 211-240] https://biodiversitylibrary.org/page/16299729" title="Iconographie der Land- &amp; Suesswasser-Mollusken mit vorzueglicher Beruecksichtigung der europaeischen noch nicht abgebildeten Arten von E. A. Rossmaessler. VIII. Band. C. W. Kreidel, Wiesbaden, 1 - 11. [pls 211 - 240]" url="https://biodiversitylibrary.org/page/16299729" year="1899">Kobelt 1899</bibRefCitation>
: 9, pl. 213, fig. 1337.
</paragraph>
<paragraph id="C418433C5309BAFF140329BBF852ADCC" pageId="0" pageNumber="25">
<taxonomicName id="2117F73AB62A5C78FC064EB6DB8D422A" authorityName="Clessin" authorityYear="1885" class="Gastropoda" family="Neritidae" genus="Neritina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neritina euxina" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="euxina">Neritina euxina</taxonomicName>
<bibRefCitation id="0BE79A98093CEAAAF93725B5724F7448" author="Clessin, S" journalOrPublisher="Malakozoologische Blaetter (Neue Folge)" pageId="0" pageNumber="25" pagination="49 - 56" publicationUrl="https://biodiversitylibrary.org/page/35594868" refId="B36" refString="Clessin, S, 1886. Binnenmollusken aus Rumaenien. Malakozoologische Blaetter (Neue Folge) 8: 49 - 56, https://biodiversitylibrary.org/page/35594868" title="Binnenmollusken aus Rumaenien." url="https://biodiversitylibrary.org/page/35594868" volume="8" year="1886">Clessin 1886</bibRefCitation>
: 55.
</paragraph>
<paragraph id="FF4894A3797B6621E972F2F95CC37FA7" pageId="0" pageNumber="25">
<taxonomicName id="940391C75F3EEEEB28E907F5FFF30AC8" class="Gastropoda" family="Neritidae" genus="Neritina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neritina heldreichi var. graeca" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="variety" species="heldreichi" variety="graeca">Neritina (Theodoxus) heldreichi var. graeca</taxonomicName>
<bibRefCitation id="A1BD8BC716F9ECA33185C9B79F271376" author="Westerlund, CA" journalOrPublisher="ZooKeys" pageId="0" pageNumber="25" publicationUrl="https://biodiversitylibrary.org/page/10887359" refId="B160" refString="Westerlund, CA, 1886. Fauna der in der palaearctischen Region (Europa, Kaukasien, Sibirien, Turan, Persien, Kurdistan, Armenien, Mesopotamien, Kleinasien, Syrien, Arabien, Egypten, Tripolis, Tunesien, Algerien und Marocco) lebenden Binnenconchylien. VI. Fam. Ampullariidae, Paludinidae, Hydrobiidae, Melaniidae, Valvatidae &amp; Neritidae. Hakan Ohlsson, Lund, 156 + 13 pp. https://biodiversitylibrary.org/page/10887359" title="Fauna der in der palaearctischen Region (Europa, Kaukasien, Sibirien, Turan, Persien, Kurdistan, Armenien, Mesopotamien, Kleinasien, Syrien, Arabien, Egypten, Tripolis, Tunesien, Algerien und Marocco) lebenden Binnenconchylien. VI. Fam. Ampullariidae, Paludinidae, Hydrobiidae, Melaniidae, Valvatidae &amp; Neritidae. Hakan Ohlsson, Lund, 156 + 13 pp." url="https://biodiversitylibrary.org/page/10887359" year="1886">Westerlund 1886</bibRefCitation>
: 152.
</paragraph>
<paragraph id="5CFB6B5C07D88CEA73966ACFD0D84DC5" pageId="0" pageNumber="25">
<taxonomicName id="F4A7DE947B61CD6FE31AA2AD7C72828D" class="Gastropoda" family="Neritidae" genus="Neritina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neritina brauneri" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="brauneri">Neritina brauneri</taxonomicName>
<bibRefCitation id="0D918E6B67F47F2472C056E6B7907869" author="Lindholm, WA" journalOrPublisher="Zapiski Novorossijskago Obshchestva Estestvoispytatelej" pageId="0" pageNumber="25" pagination="199 - 232" refId="B91" refString="Lindholm, WA, 1908. Materialien zur Molluskenfauena [sic] von Suedwestrussland, Polen und der Krim. Zapiski Novorossijskago Obshchestva Estestvoispytatelej 31: 199 - 232" title="Materialien zur Molluskenfauena [sic] von Suedwestrussland, Polen und der Krim." volume="31" year="1908">Lindholm 1908</bibRefCitation>
: 217.
</paragraph>
<paragraph id="7AF460D22C45E5B0E87A89E7934646F4" pageId="0" pageNumber="25">
<taxonomicName id="2E876706AB6750E0F9B6ADF269B483E1" baseAuthorityName="Pfeiffer" baseAuthorityYear="1828" class="Gastropoda" family="Neritidae" genus="Neritina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neritina danubialis" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="danubialis">Neritina danubialis</taxonomicName>
var.
<taxonomicName id="1C1BD9C18E40CB9B5DB577A3329D97DE" authorityName="Lindholm" authorityYear="1908" class="Gastropoda" family="Neritidae" genus="Neritina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="var. Danasteri" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="variety" species="danubialis" variety="Danasteri">Danasteri</taxonomicName>
<bibRefCitation id="C2A78391450B9AF89BFEE1DD8766486F" author="Lindholm, WA" journalOrPublisher="Zapiski Novorossijskago Obshchestva Estestvoispytatelej" pageId="0" pageNumber="25" pagination="199 - 232" refId="B91" refString="Lindholm, WA, 1908. Materialien zur Molluskenfauena [sic] von Suedwestrussland, Polen und der Krim. Zapiski Novorossijskago Obshchestva Estestvoispytatelej 31: 199 - 232" title="Materialien zur Molluskenfauena [sic] von Suedwestrussland, Polen und der Krim." volume="31" year="1908">Lindholm 1908</bibRefCitation>
: 215-217.
</paragraph>
<paragraph id="606B601FA2252186097C9FE9CC071A03" pageId="0" pageNumber="25">
<taxonomicName id="B9C73534D4D65886764B604526B55379" class="Gastropoda" family="Neritidae" genus="Neritina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neritina fluviatilis var. abrauensis" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="variety" species="fluviatilis" variety="abrauensis">Neritina fluviatilis var. abrauensis</taxonomicName>
<bibRefCitation id="71C890878180C3101DAD30458A5F0C29" author="Milaschewitsch, KO" journalOrPublisher="Ezhegodnik Zoologicheskogo Muzeya Imperatorskoi Akademii Nauk" pageId="0" pageNumber="25" pagination="420 - 430" refId="B103" refString="Milaschewitsch, KO, 1914. Liste des mollusques, collectionnes par Mr. S.A. Zernov pres des cotes du Caucase en mai 1910. Ezhegodnik Zoologicheskogo Muzeya Imperatorskoi Akademii Nauk 18: 420 - 430" title="Liste des mollusques, collectionnes par Mr. S. A. Zernov pres des cotes du Caucase en mai 1910." volume="18" year="1914">Milaschewitsch 1914</bibRefCitation>
: 429-430.
</paragraph>
<paragraph id="E7292F391BB348E0155CD58BBA453332" pageId="0" pageNumber="25">
<taxonomicName id="F875A1FF131C2298F8441241E8A85A7A" class="Gastropoda" family="Neritidae" genus="Theodoxia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxia" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="genus">Theodoxia</taxonomicName>
Ghigii
<bibRefCitation id="65085C020E1F00707FA5515F99C73690" author="Gambetta, L" journalOrPublisher="Archivio zoologico italiano" pageId="0" pageNumber="25" pagination="45 - 117" refId="B56" refString="Gambetta, L, 1929. Molluschi. Archivio zoologico italiano 13: 45 - 117" title="Molluschi." volume="13" year="1929">Gambetta 1929</bibRefCitation>
: 107, fig. 23.
</paragraph>
<paragraph id="57A9A7584525AA063EDA8B7E4C4C79C8" pageId="0" pageNumber="25">
<taxonomicName id="ADA114A8898563259E61149271323B7C" authorityName="Put'" authorityYear="1972" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus dniestroviensis" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="dniestroviensis">Theodoxus dniestroviensis</taxonomicName>
<bibRefCitation id="F9DD7175BE35492F4322F56763DC1D15" author="Put', AL" journalOrPublisher="Dopovidi Akademii Nauk Ukrainskoi RSR, Seriya B, Geologiya, Khimiya, Biologiya" pageId="0" pageNumber="25" pagination="78 - 83" refId="B125" refString="Put', AL, 1972. Do vyvchennya lunkovykh (Neritidae) Ukrayini. Dopovidi Akademii Nauk Ukrainskoi RSR, Seriya B, Geologiya, Khimiya, Biologiya 1: 78 - 83" title="Do vyvchennya lunkovykh (Neritidae) Ukrayini." volume="1" year="1972">
<normalizedToken id="A46B0BD9F48F72AEB2C7160D15CAC62D" originalValue="Put">Put'</normalizedToken>
1972
</bibRefCitation>
: 80-82, text fig. 5.
</paragraph>
<paragraph id="2E4110DBA250FF2E56F4FCD4C7E90E18" pageId="0" pageNumber="25">
<taxonomicName id="264E90C2CEDC62DF30D65C8F69D5EC4B" authority="fluvicola" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus heldreichi" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="heldreichi">Theodoxus (Theodoxus) heldreichi fluvicola</taxonomicName>
<bibRefCitation id="653AE4D2FBED266639109F38AE02B0D1" DOI="https://doi.org/10.1080/09397140.1992.10637614" author="Schuett, H" journalOrPublisher="Zoology in the Middle East" pageId="0" pageNumber="25" pagination="63 - 68" refId="B146" refString="Schuett, H, Sesen, R, 1992. The genus Theodoxus in South-western Anatolia, Turkey (Gastropoda, Prosobranchia, Neritidae). Zoology in the Middle East 6 (1): 63 - 68, DOI: https://doi.org/10.1080/09397140.1992.10637614" title="The genus Theodoxus in South-western Anatolia, Turkey (Gastropoda, Prosobranchia, Neritidae)." url="https://doi.org/10.1080/09397140.1992.10637614" volume="6" year="1992">
<normalizedToken id="17E9EAC5C7A39694FE0957A8FAFA36C9" originalValue="Schütt">Schuett</normalizedToken>
and
<normalizedToken id="7FA3DA18D8C7D714E38D7B807F4AAA1A" originalValue="Şeşen">Sesen</normalizedToken>
1992
</bibRefCitation>
: 64-66.
</paragraph>
<paragraph id="2925C30BE89EDFD554C724F502B1F45C" pageId="0" pageNumber="25">
<taxonomicName id="3446413080538383695009EB3903655D" authority="euxinus" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus fluviatilis" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="fluviatilis">Theodoxus fluviatilis euxinus</taxonomicName>
:
<bibRefCitation id="2ADA1A283166E4F7B78C2BA821C500A5" author="Yildirim, MZ" journalOrPublisher="Turkish Journal of Zoology" pageId="0" pageNumber="25" pagination="877 - 900" refId="B161" refString="Yildirim, MZ, 1999. Tuerkiye Prosobranchia (Gastropoda: Mollusca) tuerleri ve zoocografik yayilislari. 1. Tatli ve aci sular. Turkish Journal of Zoology 23 (3): 877 - 900" title="Tuerkiye Prosobranchia (Gastropoda: Mollusca) tuerleri ve zoocografik yayilislari. 1. Tatli ve aci sular." volume="23" year="1999">
<normalizedToken id="EE078245E567D5D14AB444C242A09617" originalValue="Yıldırım">Yildirim</normalizedToken>
1999
</bibRefCitation>
: 885.
</paragraph>
<paragraph id="0D9D41175C385A37D571DB02B3F39641" pageId="0" pageNumber="25">
<taxonomicName id="5A9D9616FDE5AFA0B9CF9C4085B18C6D" authority="fluviatilis" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus fluviatilis" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="fluviatilis">Theodoxus fluviatilis fluviatilis</taxonomicName>
:
<bibRefCitation id="2D09A1C7523910E5D9FF32F51A85694B" author="Yildirim, MZ" journalOrPublisher="Turkish Journal of Zoology" pageId="0" pageNumber="25" pagination="877 - 900" refId="B161" refString="Yildirim, MZ, 1999. Tuerkiye Prosobranchia (Gastropoda: Mollusca) tuerleri ve zoocografik yayilislari. 1. Tatli ve aci sular. Turkish Journal of Zoology 23 (3): 877 - 900" title="Tuerkiye Prosobranchia (Gastropoda: Mollusca) tuerleri ve zoocografik yayilislari. 1. Tatli ve aci sular." volume="23" year="1999">
<normalizedToken id="F6C3B4E58EC18C7FF636D3FBBF58BC15" originalValue="Yıldırım">Yildirim</normalizedToken>
1999
</bibRefCitation>
: 884.
</paragraph>
<paragraph id="F789080996AB6FDD9986CDC105CF3920" pageId="0" pageNumber="25">
<taxonomicName id="0AA9770219F3E8754E517D86D03F797E" authority="fluvicola" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus heldreichi" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="heldreichi">Theodoxus heldreichi fluvicola</taxonomicName>
:
<bibRefCitation id="B1CEF62A0A29D4AA612A7D40BEF5D962" author="Yildirim, MZ" journalOrPublisher="Turkish Journal of Zoology" pageId="0" pageNumber="25" pagination="877 - 900" refId="B161" refString="Yildirim, MZ, 1999. Tuerkiye Prosobranchia (Gastropoda: Mollusca) tuerleri ve zoocografik yayilislari. 1. Tatli ve aci sular. Turkish Journal of Zoology 23 (3): 877 - 900" title="Tuerkiye Prosobranchia (Gastropoda: Mollusca) tuerleri ve zoocografik yayilislari. 1. Tatli ve aci sular." volume="23" year="1999">
<normalizedToken id="880476FE984F21B882369EF313000F4E" originalValue="Yıldırım">Yildirim</normalizedToken>
1999
</bibRefCitation>
: 885 (as &quot;
<emphasis id="92074380659ABD6F62EF1EC040EE5EF2" italics="true" pageId="0" pageNumber="25">fluviocola</emphasis>
&quot;);
<bibRefCitation id="135CDFF9E2024DB5CBED10C2C801655A" author="Kebapci, Ue" journalOrPublisher="Oltenia Journal for Studies in Natural Sciences" pageId="0" pageNumber="25" pagination="75 - 83" refId="B80" refString="Kebapci, Ue, Yildirim, MZ, 2010. Freshwater snails of the Lakes Region (Goeller Boelgesi), Turkey. Oltenia Journal for Studies in Natural Sciences 26 (2): 75 - 83" title="Freshwater snails of the Lakes Region (Goeller Boelgesi), Turkey." volume="26" year="2010">
<normalizedToken id="2EE976543C430B377CF71AF4E95932AB" originalValue="Kebapçı">Kebapci</normalizedToken>
and
<normalizedToken id="A9C014592B52C9E04C8E12E48F9FC0FD" originalValue="Yıldırım">Yildirim</normalizedToken>
2010
</bibRefCitation>
: 77;
<bibRefCitation id="D8DE4B4E0BE3D71C8D36E1B09F573116" author="Guerlek, ME" journalOrPublisher="Fresenius Enviromental Bulletin" pageId="0" pageNumber="25" pagination="2992 - 3013" refId="B69" refString="Guerlek, ME, Sahin, SK, Doekuemcue, N, Yildirim, MZ, 2019. Checklist of the freshwater mollusca of Turkey (Mollusca: Gastropoda, Bivalvia). Fresenius Enviromental Bulletin 28 (4): 2992 - 3013" title="Checklist of the freshwater mollusca of Turkey (Mollusca: Gastropoda, Bivalvia)." volume="28" year="2019">
<normalizedToken id="0C3BEF750044B4B98787C8FD1E529B82" originalValue="Gürlek">Guerlek</normalizedToken>
et al. 2019
</bibRefCitation>
: 2993.
</paragraph>
<paragraph id="A626B5E488476459528AF066E55892BA" pageId="0" pageNumber="25">
<taxonomicName id="AEC6ED9BDA65FA8D4981367BD7FDEE9C" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus heldreichi" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="heldreichi">Theodoxus (Theodoxus) heldreichi</taxonomicName>
:
<bibRefCitation id="CDC6D613C2C4741D90739A61832FAB57" author="Yildirim, MZ" journalOrPublisher="Turkish Journal of Zoology" pageId="0" pageNumber="25" pagination="877 - 900" refId="B161" refString="Yildirim, MZ, 1999. Tuerkiye Prosobranchia (Gastropoda: Mollusca) tuerleri ve zoocografik yayilislari. 1. Tatli ve aci sular. Turkish Journal of Zoology 23 (3): 877 - 900" title="Tuerkiye Prosobranchia (Gastropoda: Mollusca) tuerleri ve zoocografik yayilislari. 1. Tatli ve aci sular." volume="23" year="1999">
<normalizedToken id="B4B11F71232C28F7EAF76D7C3042D756" originalValue="Yıldırım">Yildirim</normalizedToken>
1999
</bibRefCitation>
: 885.
</paragraph>
<paragraph id="506C5564183F1EAE004AE0F418549076" pageId="0" pageNumber="25">
<taxonomicName id="116C242AFA65A2FAEA627D5DD5A66B78" authorityName="Bourguignat in Issel" authorityYear="1865" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus subthermalis" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="subthermalis">Theodoxus subthermalis</taxonomicName>
:
<bibRefCitation id="BE318EE0CB9677B8F1874AF5889D3841" author="Yildirim, MZ" journalOrPublisher="Turkish Journal of Zoology" pageId="0" pageNumber="25" pagination="877 - 900" refId="B161" refString="Yildirim, MZ, 1999. Tuerkiye Prosobranchia (Gastropoda: Mollusca) tuerleri ve zoocografik yayilislari. 1. Tatli ve aci sular. Turkish Journal of Zoology 23 (3): 877 - 900" title="Tuerkiye Prosobranchia (Gastropoda: Mollusca) tuerleri ve zoocografik yayilislari. 1. Tatli ve aci sular." volume="23" year="1999">
<normalizedToken id="720DF484F9E0478DC1BB44ED60ECCB39" originalValue="Yıldırım">Yildirim</normalizedToken>
1999
</bibRefCitation>
: 885-886;
<bibRefCitation id="852F5B72A0B641FC99D8498369567BB2" author="Sitnikova, TYa" journalOrPublisher="Zoology in the Middle East" pageId="0" pageNumber="25" refId="B147" refString="Sitnikova, TYa, Kijashko, PV, Sysoev, AV, 2012. Species names of J.-R. Bourguignat and their application in current taxonomy of fresh-water gastropods of the Russian fauna. The Bulletin of the Russian Far East Malacological Society 15/16: 87-116." title="Species names of J. - R. Bourguignat and their application in current taxonomy of fresh-water gastropods of the Russian fauna. The Bulletin of the Russian Far East Malacological Society 15 / 16: 87 - 116." year="2012">Sitnikova et al. 2012</bibRefCitation>
: 88-89, fig. 1A-F.
</paragraph>
<paragraph id="C81AEE10A516A6E6819809CD64F3A230" pageId="0" pageNumber="25">
<taxonomicName id="85D9B9E27B19FF590CEF5CFE13BBB7F3" authority="heldreichi" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus heldreichi" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="heldreichi">Theodoxus (Theodoxus) heldreichi heldreichi</taxonomicName>
:
<bibRefCitation id="2FAFD48758D0819808B48ED15CB3B09C" author="Yildirim, MZ" journalOrPublisher="Turkish Journal of Zoology" pageId="0" pageNumber="25" pagination="97 - 102" refId="B162" refString="Yildirim, MZ, 2004. The Gastropods of Lake Egirdir. Turkish Journal of Zoology 28 (1): 97 - 102" title="The Gastropods of Lake Egirdir." volume="28" year="2004">
<normalizedToken id="D2AA168256F3B0DD4931855E7210678B" originalValue="Yıldırım">Yildirim</normalizedToken>
2004
</bibRefCitation>
: 99, pl. 1, fig. 1.
</paragraph>
<paragraph id="1409F7EE495BDA5B78C658DB2C29C86F" pageId="0" pageNumber="25">
<taxonomicName id="E3ECAD9D666567012D15CBDFC3B579F7" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus fluviatilis" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="fluviatilis">Theodoxus fluviatilis</taxonomicName>
:
<bibRefCitation id="6B19D10CA062270D4D350422EF44238B" author="Anistratenko, VV" journalOrPublisher="Vestnik Zoologii" pageId="0" pageNumber="25" pagination="3 - 10" refId="B15" refString="Anistratenko, VV, 2005. Lectotypes for Tricolia pullus, Gibbula divaricata and Theodoxus fluviatilis revisited. Vestnik Zoologii 39 (6): 3 - 10" title="Lectotypes for Tricolia pullus, Gibbula divaricata and Theodoxus fluviatilis revisited." volume="39" year="2005">Anistratenko 2005</bibRefCitation>
: 7-8, figs 3, 4;
<bibRefCitation id="D80DF6D26BF79022A7EF624EAF7D675C" author="Guerlek, ME" journalOrPublisher="Fresenius Enviromental Bulletin" pageId="0" pageNumber="25" pagination="2992 - 3013" refId="B69" refString="Guerlek, ME, Sahin, SK, Doekuemcue, N, Yildirim, MZ, 2019. Checklist of the freshwater mollusca of Turkey (Mollusca: Gastropoda, Bivalvia). Fresenius Enviromental Bulletin 28 (4): 2992 - 3013" title="Checklist of the freshwater mollusca of Turkey (Mollusca: Gastropoda, Bivalvia)." volume="28" year="2019">
<normalizedToken id="681DF65EE969ED27AC112DAF7912B6C1" originalValue="Gürlek">Guerlek</normalizedToken>
et al. 2019
</bibRefCitation>
: 2993;
<bibRefCitation id="AEA2E2BB5D5E7AA5FA863DBC2C9861B7" author="Gloeer, P" journalOrPublisher="Peter Gloeer, Hetlingen, Germany" pageId="0" pageNumber="25" refId="B62" refString="Gloeer, P, 2019. The freshwater gastropods of the West-Palaearctis. Vol. 1. Fresh- and brackish waters except spring and subterranean snails. Peter Gloeer, Hetlingen, Germany" title="The freshwater gastropods of the West-Palaearctis. Vol. 1. Fresh- and brackish waters except spring and subterranean snails." year="2019">
<normalizedToken id="8465627873E56E67AD04E6374514627F" originalValue="Glöer">Gloeer</normalizedToken>
2019
</bibRefCitation>
: 41-43, figs 26, 27.
</paragraph>
<paragraph id="DBA4FFBE4F4B0595C62E5A2247BDC474" pageId="0" pageNumber="25">
<taxonomicName id="029B8D0670800598F909AAD833A131FC" authority="heldreichi" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus heldreichi" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="heldreichi">Theodoxus heldreichi heldreichi</taxonomicName>
:
<bibRefCitation id="946281C5F98E601DD05ECCDA17E8A7B1" author="Kebapci, Ue" journalOrPublisher="Oltenia Journal for Studies in Natural Sciences" pageId="0" pageNumber="25" pagination="75 - 83" refId="B80" refString="Kebapci, Ue, Yildirim, MZ, 2010. Freshwater snails of the Lakes Region (Goeller Boelgesi), Turkey. Oltenia Journal for Studies in Natural Sciences 26 (2): 75 - 83" title="Freshwater snails of the Lakes Region (Goeller Boelgesi), Turkey." volume="26" year="2010">
<normalizedToken id="5136D593E8565179FABDF525A8EDA6E8" originalValue="Kebapçı">Kebapci</normalizedToken>
and
<normalizedToken id="C64C876E63662595CFB37447D64C12C7" originalValue="Yıldırım">Yildirim</normalizedToken>
2010
</bibRefCitation>
: 77;
<bibRefCitation id="56B5F57B37A91E5AA924B07ED6A015EB" author="Guerlek, ME" journalOrPublisher="Fresenius Enviromental Bulletin" pageId="0" pageNumber="25" pagination="2992 - 3013" refId="B69" refString="Guerlek, ME, Sahin, SK, Doekuemcue, N, Yildirim, MZ, 2019. Checklist of the freshwater mollusca of Turkey (Mollusca: Gastropoda, Bivalvia). Fresenius Enviromental Bulletin 28 (4): 2992 - 3013" title="Checklist of the freshwater mollusca of Turkey (Mollusca: Gastropoda, Bivalvia)." volume="28" year="2019">
<normalizedToken id="EFFB5D4E692D0A87EFC2412081C8D093" originalValue="Gürlek">Guerlek</normalizedToken>
et al. 2019
</bibRefCitation>
: 2993.
</paragraph>
<paragraph id="F9066D1E101826A8866D30BD7A0CB7F9" pageId="0" pageNumber="25">
<taxonomicName id="EB06AB17EC6FF6390F734761AEFD0751" authorityName="Lindholm" authorityYear="1908" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus brauneri" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="brauneri">Theodoxus brauneri</taxonomicName>
:
<bibRefCitation id="36F5CB07876FE0E80E502FD661FA530A" author="Welter-Schultes, FW" journalOrPublisher="Planet Poster Editions, Goettingen" pageId="0" pageNumber="25" refId="B158" refString="Welter-Schultes, FW, 2012. European non-marine molluscs, a guide for species identification. Planet Poster Editions, Goettingen" title="European non-marine molluscs, a guide for species identification." year="2012">Welter-Schultes 2012</bibRefCitation>
: 27.
</paragraph>
<paragraph id="9BC4166F00FE1332DCEC388C7DCF4441" pageId="0" pageNumber="25">
<taxonomicName id="51B673FE4FEB76042C89730C307FD9B0" baseAuthorityName="Clessin" baseAuthorityYear="1886" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus euxinus" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="euxinus">Theodoxus euxinus</taxonomicName>
:
<bibRefCitation id="1520BC7E18B5E8A1815A4B1B9D69BF21" author="Welter-Schultes, FW" journalOrPublisher="Planet Poster Editions, Goettingen" pageId="0" pageNumber="25" refId="B158" refString="Welter-Schultes, FW, 2012. European non-marine molluscs, a guide for species identification. Planet Poster Editions, Goettingen" title="European non-marine molluscs, a guide for species identification." year="2012">Welter-Schultes 2012</bibRefCitation>
: 27, unnumbered text figures.
</paragraph>
<paragraph id="CA2418F26D8165234D0E8C8370B23CCF" pageId="0" pageNumber="25">
<taxonomicName id="CA8479E8117C586393E7ABCC0FA0A963" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus heldreichi" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="heldreichi">Theodoxus heldreichi</taxonomicName>
:
<bibRefCitation id="120067332899C56DDE1872AB68D689D2" author="Welter-Schultes, FW" journalOrPublisher="Planet Poster Editions, Goettingen" pageId="0" pageNumber="25" refId="B158" refString="Welter-Schultes, FW, 2012. European non-marine molluscs, a guide for species identification. Planet Poster Editions, Goettingen" title="European non-marine molluscs, a guide for species identification." year="2012">Welter-Schultes 2012</bibRefCitation>
: 28, unnumbered text figures.
</paragraph>
<paragraph id="C47FB8708C34926FAB9F6329403EEC79" pageId="0" pageNumber="25">
<taxonomicName id="DE7751FC792D404D367F4F866D472F00" baseAuthorityName="Bourguignat" baseAuthorityYear="1852" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus saulcyi" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="saulcyi">Theodoxus saulcyi</taxonomicName>
:
<bibRefCitation id="EF55A05E72E3BAA5C74ABC39FF5F6B04" author="Welter-Schultes, FW" journalOrPublisher="Planet Poster Editions, Goettingen" pageId="0" pageNumber="25" refId="B158" refString="Welter-Schultes, FW, 2012. European non-marine molluscs, a guide for species identification. Planet Poster Editions, Goettingen" title="European non-marine molluscs, a guide for species identification." year="2012">Welter-Schultes 2012</bibRefCitation>
: 29, unnumbered text figures.
</paragraph>
<paragraph id="8AFE1597206055DCBAFB914EFE4FFE87" pageId="0" pageNumber="25">
<taxonomicName id="9A953E4D4DCDB0132AA3933F4520EEF2" baseAuthorityName="Clessin" baseAuthorityYear="1886" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus euxinus" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="euxinus">Theodoxus (Theodoxus) euxinus</taxonomicName>
:
<bibRefCitation id="F3C4C98F694B077D54A0CA1461908CD1" author="Vinarski, MV" journalOrPublisher="Severtsov Institute of Ecology and Evolution of RAS, Moscow" pageId="0" pageNumber="25" refId="B157" refString="Vinarski, MV, Kantor, YI, 2016. Analytical catalogue of fresh and brackish water molluscs of Russia and adjacent countries. A.N. Severtsov Institute of Ecology and Evolution of RAS, Moscow" title="Analytical catalogue of fresh and brackish water molluscs of Russia and adjacent countries. A. N." year="2016">Vinarski and Kantor 2016</bibRefCitation>
: 155.
</paragraph>
<paragraph id="75E546D88181D7C407B715E0ABAB0C4C" pageId="0" pageNumber="25">
<taxonomicName id="6C10FEE11557D7B871EBBB5DD5EF3FB9" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus fluviatilis" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="fluviatilis">Theodoxus (Theodoxus) fluviatilis</taxonomicName>
:
<bibRefCitation id="316C9FD197D447024DA9822144B22AD7" author="Vinarski, MV" journalOrPublisher="Severtsov Institute of Ecology and Evolution of RAS, Moscow" pageId="0" pageNumber="25" refId="B157" refString="Vinarski, MV, Kantor, YI, 2016. Analytical catalogue of fresh and brackish water molluscs of Russia and adjacent countries. A.N. Severtsov Institute of Ecology and Evolution of RAS, Moscow" title="Analytical catalogue of fresh and brackish water molluscs of Russia and adjacent countries. A. N." year="2016">Vinarski and Kantor 2016</bibRefCitation>
: 154-155.
</paragraph>
<paragraph id="A99D9A981C1A7DE5D77BC941CDDBDCA6" pageId="0" pageNumber="25">
<taxonomicName id="9EC965301012783F08A4C00D47EF2BC8" authorityName="Bourguignat in Issel" authorityYear="1865" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus subthermalis" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="subthermalis">Theodoxus (Theodoxus) subthermalis</taxonomicName>
:
<bibRefCitation id="6C2C434607C50943601171F2D67E2AC3" author="Vinarski, MV" journalOrPublisher="Severtsov Institute of Ecology and Evolution of RAS, Moscow" pageId="0" pageNumber="25" refId="B157" refString="Vinarski, MV, Kantor, YI, 2016. Analytical catalogue of fresh and brackish water molluscs of Russia and adjacent countries. A.N. Severtsov Institute of Ecology and Evolution of RAS, Moscow" title="Analytical catalogue of fresh and brackish water molluscs of Russia and adjacent countries. A. N." year="2016">Vinarski and Kantor 2016</bibRefCitation>
: 157-158;
<bibRefCitation id="B91DE9A075C4983962F2C5B6895FB7B5" author="Guerlek, ME" journalOrPublisher="Fresenius Enviromental Bulletin" pageId="0" pageNumber="25" pagination="2992 - 3013" refId="B69" refString="Guerlek, ME, Sahin, SK, Doekuemcue, N, Yildirim, MZ, 2019. Checklist of the freshwater mollusca of Turkey (Mollusca: Gastropoda, Bivalvia). Fresenius Enviromental Bulletin 28 (4): 2992 - 3013" title="Checklist of the freshwater mollusca of Turkey (Mollusca: Gastropoda, Bivalvia)." volume="28" year="2019">
<normalizedToken id="2B6A980EB2E267354E549378199DF991" originalValue="Gürlek">Guerlek</normalizedToken>
et al. 2019
</bibRefCitation>
: 2993.
</paragraph>
</subSubSection>
<subSubSection id="892F040CE5697D7D1D8F391D1B900A6B" pageId="0" pageNumber="25" type="type locality">
<paragraph id="6964FFA9E1EADC60C6B1DEB31DCD8CB5" pageId="0" pageNumber="25">
<typeStatus id="E4920495E5C6C5F815C3C0CC16D6CA77">Type</typeStatus>
locality.
</paragraph>
<paragraph id="ADF709523B67D9E02F16CC754F43A060" pageId="0" pageNumber="25">Near Uppsala, Sweden.</paragraph>
</subSubSection>
<subSubSection id="31B986CE07ACF43504D729FE13561569" pageId="0" pageNumber="25" type="type material">
<paragraph id="A13AF8D29F08E84CE0CC28684F8CAAB8" pageId="0" pageNumber="25">
<typeStatus id="CDD99AF9127BC47CEE896B090D46887D">Type</typeStatus>
material.
</paragraph>
<paragraph id="E027587275E623A9E16F55139C7FA78E" pageId="0" pageNumber="25">
<materialsCitation id="4B7322929EA7C718A8C41B7C5FD466C4" collectionCode="LSL" country="Turkey" typeStatus="Lectotype">
<typeStatus id="3C33667D8035B94C129851777FF1DBB0">Lectotype</typeStatus>
of
<taxonomicName id="BFE52C6F39FF3C29EDA96F6FA07EF5AB" lsidName="T. fluviatilis" pageId="0" pageNumber="25" rank="species" species="fluviatilis">
<emphasis id="B38909B64A393AA8B717F3B96EF46938" italics="true" pageId="0" pageNumber="25">T. fluviatilis</emphasis>
</taxonomicName>
are stored in
<collectionCode id="FB3C96923DAD69AACE8DFFA5CB5FF0FC" collectionName="LSL">LSL</collectionCode>
(coll. no.
<collectionCode id="2CB42A8EF9EEF2A98AFF377346720BA5" collectionName="LSL">LSL</collectionCode>
.566;
<typeStatus id="ACF0F4385EE8E5FCC67578776A1306D6">lectotype</typeStatus>
designated by
<bibRefCitation id="75FD977F56C90F106B4494F9E6BB1942" author="Anistratenko, OY" journalOrPublisher="Vestnik Zoologii" pageId="0" pageNumber="25" pagination="11 - 19" refId="B14" refString="Anistratenko, OY, Starobogatov, YI, Anistratenko, VV, 1999. Mollyuski roda Theodoxus (Gastropoda, Pectinibranchia, Neritidae) Azovo-Chernomorskogo basseyna. Vestnik Zoologii 33 (3): 11 - 19" title="Mollyuski roda Theodoxus (Gastropoda, Pectinibranchia, Neritidae) Azovo-Chernomorskogo basseyna." volume="33" year="1999">Anistratenko et al. 1999</bibRefCitation>
; also see
<bibRefCitation id="AC72787E3F091C7100BBB2B4AAC3D58A" author="Anistratenko, VV" journalOrPublisher="Vestnik Zoologii" pageId="0" pageNumber="25" pagination="3 - 10" refId="B15" refString="Anistratenko, VV, 2005. Lectotypes for Tricolia pullus, Gibbula divaricata and Theodoxus fluviatilis revisited. Vestnik Zoologii 39 (6): 3 - 10" title="Lectotypes for Tricolia pullus, Gibbula divaricata and Theodoxus fluviatilis revisited." volume="39" year="2005">Anistratenko 2005</bibRefCitation>
)
</materialsCitation>
.
<materialsCitation id="7729E1E08B1D51264344B246D56C7BA8" collectionCode="MHNG" country="Turkey" typeStatus="Syntypes">
<typeStatus id="C410057AD24BCA7F01DACC3613F22651">Syntypes</typeStatus>
of
<taxonomicName id="F6C51A041209E8632BF334A3AC3A3A95" lsidName="N. saulcyi" pageId="0" pageNumber="25" rank="species" species="saulcyi">
<emphasis id="7E23809BDDA039FC32837ED14E43BC75" italics="true" pageId="0" pageNumber="25">N. saulcyi</emphasis>
</taxonomicName>
are stored in
<collectionCode id="F06E9D496841C07BA166EDD8085B4F19" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" type="Museum">MHNG</collectionCode>
(coll. no. MHNG-MOLL-111736)
</materialsCitation>
.
<materialsCitation id="C7E08794BD8E15E5F528E542FE3FE8D5" collectionCode="MHNG" country="Turkey" typeStatus="syntypes">
Seven
<typeStatus id="93EB198A9C44E4B7ABF75CB4E9DCB34C">syntypes</typeStatus>
of
<taxonomicName id="4DD4DCEDCD954F8231A22B59D4531BB2" lsidName="T. fluviatilis var. subthermalis" pageId="0" pageNumber="25" rank="variety" species="fluviatilis" variety="subthermalis">T. fluviatilis var. subthermalis</taxonomicName>
are stored in
<collectionCode id="2F2675EE2F341EECDA9F386614B3D45C" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" type="Museum">MHNG</collectionCode>
(coll. no. MHNG-MOLL-11737)
</materialsCitation>
.
<materialsCitation id="BC76A31903E2F2FEC634C7EA92EF6AFA" collectionCode="NSMF" country="Turkey" typeStatus="type">
The
<typeStatus id="98840766CC10D701C131F377BE359178">type</typeStatus>
material of
<taxonomicName id="49BA2FD6B00726E67144C2698FF491FB" authority="fluvicola" lsidName="T. heldreichi" pageId="0" pageNumber="25" rank="species" species="heldreichi">
<emphasis id="634B617D7D5D686A72DAB6C8C679D525" italics="true" pageId="0" pageNumber="25">T. heldreichi fluvicola</emphasis>
</taxonomicName>
is stored in
<collectionCode id="FDFFD595FDC3F9A8369DA73671DAA231" collectionName="NSMF">NSMF</collectionCode>
(coll. no. 309.439 and 309.440)
</materialsCitation>
.
<materialsCitation id="80300C40128D92CB5C0E19C9D336018A" collectionCode="ZIN" country="Turkey" typeStatus="syntypes">
Most taxa described by Lindholm are deposited in
<collectionCode id="FFE3262B29DECFFD6109924D3310A315" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" type="Museum">ZIN</collectionCode>
, including five
<typeStatus id="3A7B84F027A03D8C3BE9057ABABA6959">syntypes</typeStatus>
of
<taxonomicName id="44B547C052B6407D1C90362AEB116F40" lsidName="N. fluviatilis var. cereoflava" pageId="0" pageNumber="25" rank="variety" species="fluviatilis" variety="cereoflava">N. fluviatilis var. cereoflava</taxonomicName>
(coll. no. 6055/1)
</materialsCitation>
,
<materialsCitation id="0EFC114418CCF7435863D9E7C91CD1E4" country="Turkey" typeStatus="syntypes">
three
<typeStatus id="72CAF5366C01F06F733C0431E06CAB1F">syntypes</typeStatus>
of
<taxonomicName id="76EDB1D80FFF58FC4E89DD30F0352263" lsidName="N. brauneri" pageId="0" pageNumber="25" rank="species" species="brauneri">
<emphasis id="9C0EC0A84A18789B32430F3ADFCDD0D6" italics="true" pageId="0" pageNumber="25">N. brauneri</emphasis>
</taxonomicName>
(coll. no. 6046/1)
</materialsCitation>
,
<materialsCitation id="46AA2E4D2AB3CD0A2553A460C40F9D71" country="Turkey" typeStatus="syntype">
one
<typeStatus id="55DB9BD290D46ED063BED8AEB8CB388F">syntype</typeStatus>
of
<taxonomicName id="11E1B7873E28935CF7E1193886123078" form="alboguttata" lsidName="N. brauneri" pageId="0" pageNumber="25" rank="form" species="brauneri">N. brauneri f. alboguttata</taxonomicName>
(coll. no. 6051/1)
</materialsCitation>
,
<materialsCitation id="40A43877CDDFAB35773904DC98C3055E" country="Turkey" typeStatus="syntypes">
two
<typeStatus id="E1AB5F8BF220B58BFF94E43EC63F5E86">syntypes</typeStatus>
of
<taxonomicName id="B5A3A099061A295E02E88389C6D9BFEB" form="lacrymans" lsidName="N. brauneri" pageId="0" pageNumber="25" rank="form" species="brauneri">N. brauneri f. lacrymans</taxonomicName>
(coll. no. 6052/1)
</materialsCitation>
,
<materialsCitation id="523A359821FA2C5403A3A351C3D1FBF3" country="Turkey" typeStatus="syntypes">
one
<typeStatus id="67E79AAFA309D23434903A0654F1790C">syntypes</typeStatus>
of
<taxonomicName id="9F8052ACC6B60C8193D8C33BB5293E74" form="pulcherrima" lsidName="N. brauneri" pageId="0" pageNumber="25" rank="form" species="brauneri">N. brauneri f. pulcherrima</taxonomicName>
(coll. no. 6053/1)
</materialsCitation>
,
<materialsCitation id="FEAA9E19ED8A8B936EA1B8FF107025B3" country="Turkey" typeStatus="syntypes">
and three
<typeStatus id="524D13D033361958D019E58388625701">syntypes</typeStatus>
of
<taxonomicName id="B0C719FCFFB43B774341901ED775D9BE" lsidName="N. danubialis var. danasteri" pageId="0" pageNumber="25" rank="variety" species="danubialis" variety="danasteri">N. danubialis var. danasteri</taxonomicName>
(coll. no. 5910/3)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="5D2931C772C3EB57533F5272E9327503" pageId="0" pageNumber="25" type="remarks">
<paragraph id="430EF287A26BC1FA578C1F6B0122CB45" pageId="0" pageNumber="25">Remarks.</paragraph>
<paragraph id="C08F31AAEF355A1F7AA4ED59B16D0A7C" pageId="0" pageNumber="25">
<taxonomicName id="710BE4E30E6901004FF8300C28B5A252" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus fluviatilis" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="fluviatilis">
<emphasis id="7C2D7D1CED6A15E33431A033FB2FC5D7" italics="true" pageId="0" pageNumber="25">Theodoxus fluviatilis</emphasis>
</taxonomicName>
exhibits considerable variation in periostracum colouration (
<bibRefCitation id="CF14039577FCD2AAA9319BF9F70D0B3D" DOI="https://doi.org/10.1007/BF00408578" author="Neumann, D" journalOrPublisher="Zeitschrift fuer Morphologie und Oekologie der Tiere" pageId="0" pageNumber="25" pagination="349 - 411" refId="B114" refString="Neumann, D, 1959. Morphologische und experimentelle Untersuchungen ueber die Variabilitaet der Farbmuster auf der Schale von Theodoxus fluviatilis L. Zeitschrift fuer Morphologie und Oekologie der Tiere 48 (4): 349 - 411, DOI: https://doi.org/10.1007/BF00408578" title="Morphologische und experimentelle Untersuchungen ueber die Variabilitaet der Farbmuster auf der Schale von Theodoxus fluviatilis L." url="https://doi.org/10.1007/BF00408578" volume="48" year="1959">Neumann 1959</bibRefCitation>
;
<bibRefCitation id="4B351BF1A14DDF6DBF6004B64D919DA2" author="Gloeer, P" journalOrPublisher="Ecologica Montenegrina" pageId="0" pageNumber="25" pagination="88 - 92" refId="B64" refString="Gloeer, P, Pesic, V, 2015. The morphological plasticity of Theodoxus fluviatilis (Linnaeus, 1758) (Mollusca: Gastropoda: Neritidae). Ecologica Montenegrina 2 (2): 88 - 92" title="The morphological plasticity of Theodoxus fluviatilis (Linnaeus, 1758) (Mollusca: Gastropoda: Neritidae)." volume="2" year="2015">
<normalizedToken id="7A36846B539A5353A7CEC1D1F355A17F" originalValue="Glöer">Gloeer</normalizedToken>
and
<normalizedToken id="D35D81A7F84E9058216D7E68F6ABDE8E" originalValue="Pešić">Pesic</normalizedToken>
2015
</bibRefCitation>
). It is not surprising that molecular studies already support the synonymy of a number taxa such as
<taxonomicName id="4395ED85DC30272B9700AA58E71E3ED8" lsidName="T. euxinus" pageId="0" pageNumber="25" rank="species" species="euxinus">
<emphasis id="ED2366042D5546E3FB63C293421F92CD" italics="true" pageId="0" pageNumber="25">T. euxinus</emphasis>
</taxonomicName>
(Clessin, 1886),
<taxonomicName id="BAA18B2CCA433F8F8E86F0F84C33531B" authority="abrauensis" lsidName="T. fluviatilis" pageId="0" pageNumber="25" rank="species" species="fluviatilis">
<emphasis id="57A60C8B9AD54AF535DA03B8346CE507" italics="true" pageId="0" pageNumber="25">T. fluviatilis abrauensis</emphasis>
</taxonomicName>
(Milaschewitsch, 1914)
<taxonomicName id="CDD5609EAA03EBE896D7E77A183B78C6" lsidName="T. danasteri" pageId="0" pageNumber="25" rank="species" species="danasteri">
<emphasis id="DAE24F98637E75CCBEA9454031AF1E80" italics="true" pageId="0" pageNumber="25">T. danasteri</emphasis>
</taxonomicName>
(Lindholm, 1908), and
<taxonomicName id="6DB8FFAE3799D2816EAA851C7D991308" lsidName="T. subthermalis" pageId="0" pageNumber="25" rank="species" species="subthermalis">
<emphasis id="80B64A13BF08C803142AA8A3E6DF8CFF" italics="true" pageId="0" pageNumber="25">T. subthermalis</emphasis>
</taxonomicName>
Issel, 1865, and further suggest the inclusion of
<taxonomicName id="6E8F94E8A556B409D187CCCE0243E927" lsidName="T. saulcyi" pageId="0" pageNumber="25" rank="species" species="saulcyi">
<emphasis id="72EAC24BDA4AE14E19C02F8096B4F7DB" italics="true" pageId="0" pageNumber="25">T. saulcyi</emphasis>
</taxonomicName>
(Bourguignat, 1852) (including
<taxonomicName id="941FC7AF452CDDF414396AD980AA1723" lsidName="N. graeca" pageId="0" pageNumber="25" rank="species" species="graeca">
<emphasis id="8C68E524A6F6DC93CAE6708AF3EAFC1C" italics="true" pageId="0" pageNumber="25">N. graeca</emphasis>
</taxonomicName>
Westerlund, 1886,
<taxonomicName id="78D2CEDD677A90A1D0C547E4CD59C8E8" lsidName="T. ghigii" pageId="0" pageNumber="25" rank="species" species="ghigii">
<emphasis id="A6EC9E7B42047C6737B2F1329E28160A" italics="true" pageId="0" pageNumber="25">T. ghigii</emphasis>
</taxonomicName>
Gambetta, 1929; both listed as synonyms of
<taxonomicName id="BDA946CB79572D70E37475A2375AEFFF" lsidName="T. saulcyi" pageId="0" pageNumber="25" rank="species" species="saulcyi">
<emphasis id="C6B41FF3D3B857E279A70F13C01F81A9" italics="true" pageId="0" pageNumber="25">T. saulcyi</emphasis>
</taxonomicName>
by
<bibRefCitation id="073494BD98A17C797981E65B66B2E02F" author="Bank, RA" journalOrPublisher="Heldia" pageId="0" pageNumber="25" pagination="51 - 86" refId="B21" refString="Bank, RA, 2006. Towards a catalogue and bibliography of the freshwater Mollusca of Greece. Heldia 6 (1): 51 - 86" title="Towards a catalogue and bibliography of the freshwater Mollusca of Greece." volume="6" year="2006">Bank (2006)</bibRefCitation>
) and
<taxonomicName id="20FE4FD42B0C70CC8BAF7246D2525C0F" lsidName="T. heldreichi" pageId="0" pageNumber="25" rank="species" species="heldreichi">
<emphasis id="1B2ED45E6BCB79E058EBB87CE1965E4C" italics="true" pageId="0" pageNumber="25">T. heldreichi</emphasis>
</taxonomicName>
(Martens, 1879) (
<bibRefCitation id="D365150DE41777A6CD3A2E397535C5C9" DOI="https://doi.org/10.1016/j.ympev.2006.06.026" author="Bunje, PME" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="0" pageNumber="25" pagination="373 - 387" refId="B35" refString="Bunje, PME, Lindberg, DR, 2007. Lineage divergence of a freshwater snail clade associated with post-Tethys marine basin development. Molecular Phylogenetics and Evolution 42 (2): 373 - 387, DOI: https://doi.org/10.1016/j.ympev.2006.06.026" title="Lineage divergence of a freshwater snail clade associated with post-Tethys marine basin development." url="https://doi.org/10.1016/j.ympev.2006.06.026" volume="42" year="2007">Bunje and Lindberg 2007</bibRefCitation>
;
<bibRefCitation id="7920E4A6AB43272359148E26C054EFF9" DOI="https://doi.org/10.1111/jbi.13590" author="Sands, AF" journalOrPublisher="Journal of Biogeography" pageId="0" pageNumber="25" pagination="1274 - 1286" refId="B138" refString="Sands, AF, Sereda, SV, Stelbrink, B, Neubauer, TA, Lazarev, S, Wilke, T, Albrecht, C, 2019a. Contributions of biogeographical functions to species accumulation may change over time in refugial regions. Journal of Biogeography 46 (6): 1274 - 1286, DOI: https://doi.org/10.1111/jbi.13590" title="Contributions of biogeographical functions to species accumulation may change over time in refugial regions." url="https://doi.org/10.1111/jbi.13590" volume="46" year="2019 a">Sands et al. 2019a</bibRefCitation>
). The above species share near identical operculum structures, with a strong rib-shield, rib-pouch and the absence of a pseudo-apophysis (Figs
<figureCitation id="D33A29BE84606CA49D222B0AD6E6FD43" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Theodoxus fluviatilis (Linnaeus, 1758). A - D. Specimen collected close to Oued Laabid, Morocco (UGSB 18106) (Fig. 2); E - H. Specimen from the Danube River, Woerth an der Donau, Germany (UGSB 24173); I - L. Specimen (T. euxinus - morphotype) collected in the Papuc Creek, Kiyikoey, Turkey (UGSB 24172) (Fig. 2); M - P. Specimen (T. euxinus - morphotype) from Ovidiopol, Ukraine (UGSB 18124); Q - S. Specimen (T. heldreichi - morphotype) collected in Lake Egirdir, Turkey (UGSB 20361); T, U. Specimen (T. danasteri - morphotype) collected in Krasna Kosa, Ukraine (UGSB 18417) (Fig. 2); V. Opercula of a T. danasteri - morphotype from Lake Skadar, Montenegro (UGSB 24171). Specimens A-D, I-L and T-U were used in the phylogeny (Fig. 2). Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure9" httpUri="https://binary.pensoft.net/fig/376458" pageId="0" pageNumber="25">9</figureCitation>
,
<figureCitation id="9F5A569D68DF24CDBBFD3C2AF9E338A6" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Theodoxus fluviatilis (Linnaeus, 1758). A - D. Specimen (conforming to T. sarmaticus) from the Dnieper-Bug estuary, Ukraine (UGSB 18328); E - H. Specimen (T. subthermalis - morphotype) from Novy Afon, Abkhazia, Georgia (UGSB 18402); I - K. Syntype of N. saulcyi from near the Penteli Monastery, Greece (MHNG-MOLL- 111736); L. Specimen (T. saulcyi - morphotype) from Stilos, Crete, Greece (UGSB 20519) incorporated into the phylogeny (Fig. 2). Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure10" httpUri="https://binary.pensoft.net/fig/376459" pageId="0" pageNumber="25">10</figureCitation>
), which were often disregarded in early
<taxonomicName id="BA760809BD78F1814E7C7E24E156B5A3" authorityName="Montfort" authorityYear="1810" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="genus">
<emphasis id="04C35BEEB4F351C7BBED91ABAD041D3B" italics="true" pageId="0" pageNumber="25">Theodoxus</emphasis>
</taxonomicName>
species descriptions. Moreover, a number of taxa including
<taxonomicName id="5E18BBBE20D3DEB641219B5439D5E9F3" lsidName="T. brauneri" pageId="0" pageNumber="25" rank="species" species="brauneri">
<emphasis id="8231A1FEE1E329F83EAC9BA499600651" italics="true" pageId="0" pageNumber="25">T. brauneri</emphasis>
</taxonomicName>
(Lindholm, 1908) (along with subspecies
<taxonomicName id="B389DDEC65293768CB95FF177FE1EE81" authority="alboguttatus" lsidName="T. b." pageId="0" pageNumber="25" rank="species" species="b.">
<emphasis id="BC837FDD9A81F1D442042E3CE0771C4B" italics="true" pageId="0" pageNumber="25">T. b. alboguttatus</emphasis>
</taxonomicName>
(Lindholm, 1908),
<taxonomicName id="CC473E1F126DE83E50A2921EE787E60F" authority="lacrymans" lsidName="T. b." pageId="0" pageNumber="25" rank="species" species="b.">
<emphasis id="F47A26A50EA3E3733FB986DBAD2ABE75" italics="true" pageId="0" pageNumber="25">T. b. lacrymans</emphasis>
</taxonomicName>
(Lindholm, 1908) and
<taxonomicName id="B0EE3EF6FD82AD2613FE2B112C2ACA7A" authority="pulcherrimus" lsidName="T. b." pageId="0" pageNumber="25" rank="species" species="b.">
<emphasis id="2259FFE7D30CC57B313D7A1333819B2D" italics="true" pageId="0" pageNumber="25">T. b. pulcherrimus</emphasis>
</taxonomicName>
(
<emphasis id="56EEC8898865A2DBCA1793EB9BE61D96" italics="true" pageId="0" pageNumber="25">Lindholm</emphasis>
,
<emphasis id="67AD42243397CD15600C4D2C80087878" italics="true" pageId="0" pageNumber="25">1908</emphasis>
)),
<taxonomicName id="891D80ABEE92D5AABB0E72E638B544C5" authority="cereoflava" lsidName="T. fluviatilis" pageId="0" pageNumber="25" rank="species" species="fluviatilis">
<emphasis id="F823239572E88CF624BA7FA5D1571F41" italics="true" pageId="0" pageNumber="25">T. fluviatilis cereoflava</emphasis>
</taxonomicName>
(Lindholm, 1913), and
<taxonomicName id="97D6D09F2D264144FD5C8124FAB97153" lsidName="T. dniestroviensis" pageId="0" pageNumber="25" rank="species" species="dniestroviensis">
<emphasis id="9EACEC99DB6C21C9A7D28DCC91365411" italics="true" pageId="0" pageNumber="25">T. dniestroviensis</emphasis>
</taxonomicName>
<bibRefCitation id="52EC285C8E65F7F5E9DED1C4A17E5FE8" author="Put', AL" journalOrPublisher="Dopovidi Akademii Nauk Ukrainskoi RSR, Seriya B, Geologiya, Khimiya, Biologiya" pageId="0" pageNumber="25" pagination="78 - 83" refId="B125" refString="Put', AL, 1972. Do vyvchennya lunkovykh (Neritidae) Ukrayini. Dopovidi Akademii Nauk Ukrainskoi RSR, Seriya B, Geologiya, Khimiya, Biologiya 1: 78 - 83" title="Do vyvchennya lunkovykh (Neritidae) Ukrayini." volume="1" year="1972">
<normalizedToken id="63B469C78DA7971992D2C2AEF6550EA1" originalValue="Put">Put'</normalizedToken>
, 1972
</bibRefCitation>
are well within the variability of
<taxonomicName id="77F913737C6B4AAE155F9E48AEF0E7C5" lsidName="T. fluviatilis" pageId="0" pageNumber="25" rank="species" species="fluviatilis">
<emphasis id="57849C80CF24C4EB3ECF0BB346433327" italics="true" pageId="0" pageNumber="25">T. fluviatilis</emphasis>
</taxonomicName>
and are considered junior synonyms herein as well.
</paragraph>
<caption id="BE48657C8FF8948B695F17E73C0B50D6" doi="10.3897/zse.96.48312.figure9" httpUri="https://binary.pensoft.net/fig/376458" pageId="0" pageNumber="25" start="Figure 9" startId="F9">
<paragraph id="AEB513EAC99FD85ECDFE8545CC03CEBD" pageId="0" pageNumber="25">
<emphasis id="5EC75D4F776853F5A5174A63754DAC5C" bold="true" pageId="0" pageNumber="25">Figure 9.</emphasis>
<taxonomicName id="5AB9E37C84C65D49D0EBC5127F44B57F" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus fluviatilis" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="fluviatilis">
<emphasis id="36B981DA0C6CDBC03F2C2462DC759829" italics="true" pageId="0" pageNumber="25">Theodoxus fluviatilis</emphasis>
</taxonomicName>
(Linnaeus, 1758).
<emphasis id="79C0169A9589A8DABC5E70C02517D9DE" bold="true" pageId="0" pageNumber="25">A</emphasis>
-
<emphasis id="B30CD1C1DF97C2B1C7C4B5E18CBB5762" bold="true" pageId="0" pageNumber="25">D</emphasis>
. Specimen collected close to Oued Laabid, Morocco (
<collectionCode id="B6E9D7B2E62CE0C62D1A0F299998826D" collectionName="UGSB">UGSB</collectionCode>
18106) (Fig.
<figureCitation id="A54D0123664E87EDAE02CD8832CC67CC" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dated phylogeny of the genus Theodoxus constructed with BEAST, based on existing GenBank and newly incorporated data for COI, 16 S and ATPα (Suppl. material 1: Table S 1). Morphospecies where associations are queried are highlighted by a question mark after the species name (see the &quot; Systematic checklist &quot; for more information). Small green (PP ≥ 0.95) and red (PP &lt;0.95) dots on nodes indicate the relative posterior probability support of divergence events. Node labels (1 - 32) correspond to divergence dates and HPD intervals in millions of years ago (Mya) for selected nodes (see Suppl. material 1: Table S 3). Dashed dark blue and light blue lines represent the minimum interspecific and maximum intraspecific variation determined by Sands et al. (2019 a) to aid Theodoxus spp. delimitation (see see Methods, Phylogenetics and species delimitation and that study for more information on the delimitation methods)." figureDoi="10.3897/zse.96.48312.figure2" httpUri="https://binary.pensoft.net/fig/376451" pageId="0" pageNumber="25">2</figureCitation>
);
<emphasis id="75730A299E24ED2396EF1773C747742F" bold="true" pageId="0" pageNumber="25">E</emphasis>
-
<emphasis id="97D25CCD0EE52EA8EA597A726BACB9A6" bold="true" pageId="0" pageNumber="25">H</emphasis>
. Specimen from the Danube River,
<normalizedToken id="044FC5F3A45AD09874D3B26599920000" originalValue="Wörth">Woerth</normalizedToken>
an der Donau, Germany (
<collectionCode id="CC62E09879D8B849AEAC52DB54A0C668" collectionName="UGSB">UGSB</collectionCode>
24173);
<emphasis id="28CB6EC1E9A468955177342CAE67E61D" bold="true" pageId="0" pageNumber="25">I</emphasis>
-
<emphasis id="FE56E2EE71A0ADB01F9DD3AE02607738" bold="true" pageId="0" pageNumber="25">L</emphasis>
. Specimen (
<taxonomicName id="106F308C71FF47D0CE4198E9AE75165C" lsidName="T. euxinus" pageId="0" pageNumber="25" rank="species" species="euxinus">
<emphasis id="C32B25134534F3188BF119374DED74E5" italics="true" pageId="0" pageNumber="25">T. euxinus</emphasis>
</taxonomicName>
-morphotype) collected in the
<normalizedToken id="C99740C6831689E7916CFAC673A27115" originalValue="Papuç">Papuc</normalizedToken>
Creek,
<normalizedToken id="71F27796F33A943A78D2955EA8868FCB" originalValue="Kıyıköy">Kiyikoey</normalizedToken>
, Turkey (
<collectionCode id="09E9A405759351AE64C60F887F35BA4A" collectionName="UGSB">UGSB</collectionCode>
24172) (Fig.
<figureCitation id="DA7FB1075DE906E21AEB48C03E3D6FF4" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dated phylogeny of the genus Theodoxus constructed with BEAST, based on existing GenBank and newly incorporated data for COI, 16 S and ATPα (Suppl. material 1: Table S 1). Morphospecies where associations are queried are highlighted by a question mark after the species name (see the &quot; Systematic checklist &quot; for more information). Small green (PP ≥ 0.95) and red (PP &lt;0.95) dots on nodes indicate the relative posterior probability support of divergence events. Node labels (1 - 32) correspond to divergence dates and HPD intervals in millions of years ago (Mya) for selected nodes (see Suppl. material 1: Table S 3). Dashed dark blue and light blue lines represent the minimum interspecific and maximum intraspecific variation determined by Sands et al. (2019 a) to aid Theodoxus spp. delimitation (see see Methods, Phylogenetics and species delimitation and that study for more information on the delimitation methods)." figureDoi="10.3897/zse.96.48312.figure2" httpUri="https://binary.pensoft.net/fig/376451" pageId="0" pageNumber="25">2</figureCitation>
);
<emphasis id="DC2206E468A497C342E3F2DF88F2204A" bold="true" pageId="0" pageNumber="25">M</emphasis>
-
<emphasis id="94BA5CB556195332253F43B47F2D702B" bold="true" pageId="0" pageNumber="25">P</emphasis>
. Specimen (
<taxonomicName id="4DB9CA0A2402AB768833541B1983E368" lsidName="T. euxinus" pageId="0" pageNumber="25" rank="species" species="euxinus">
<emphasis id="2BB3949FEE9314227CD1BE593B5BDABA" italics="true" pageId="0" pageNumber="25">T. euxinus</emphasis>
</taxonomicName>
-morphotype) from Ovidiopol, Ukraine (
<collectionCode id="2E1DA87B1099FC7F380F69D8F0911403" collectionName="UGSB">UGSB</collectionCode>
18124);
<emphasis id="F16D1E2DCB4E5794BA04D06F06ECA6BB" bold="true" pageId="0" pageNumber="25">Q</emphasis>
-
<emphasis id="42AFFA635CD586BC00BF28FA70CD85BE" bold="true" pageId="0" pageNumber="25">S</emphasis>
. Specimen (
<taxonomicName id="31A82CAE5BDFA41FE73916FD66189A44" lsidName="T. heldreichi" pageId="0" pageNumber="25" rank="species" species="heldreichi">
<emphasis id="0271C5133953F2E182AC890F0A0D670E" italics="true" pageId="0" pageNumber="25">T. heldreichi</emphasis>
</taxonomicName>
-morphotype) collected in Lake
<normalizedToken id="A73CA3037BE5CB7F700A618D48A8BEA2" originalValue="Eğirdir">Egirdir</normalizedToken>
, Turkey (
<collectionCode id="CF16F896D116E84D6D53F4DEABD1DAA1" collectionName="UGSB">UGSB</collectionCode>
20361);
<emphasis id="B02BE665600C16117E89C15E289B6977" bold="true" pageId="0" pageNumber="25">T</emphasis>
,
<emphasis id="72E6955798923009B101D55C6210C891" bold="true" pageId="0" pageNumber="25">U</emphasis>
. Specimen (
<taxonomicName id="A39CDA992292294F29F9E560B9C54A82" lsidName="T. danasteri" pageId="0" pageNumber="25" rank="species" species="danasteri">
<emphasis id="5807E055ACDB56289218BD33793B25C0" italics="true" pageId="0" pageNumber="25">T. danasteri</emphasis>
</taxonomicName>
-morphotype) collected in Krasna Kosa, Ukraine (
<collectionCode id="58040C07534E902DD21EB4F565D50D26" collectionName="UGSB">UGSB</collectionCode>
18417) (Fig.
<figureCitation id="54E4A147B67CACE794FCF555C3C5CAA6" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dated phylogeny of the genus Theodoxus constructed with BEAST, based on existing GenBank and newly incorporated data for COI, 16 S and ATPα (Suppl. material 1: Table S 1). Morphospecies where associations are queried are highlighted by a question mark after the species name (see the &quot; Systematic checklist &quot; for more information). Small green (PP ≥ 0.95) and red (PP &lt;0.95) dots on nodes indicate the relative posterior probability support of divergence events. Node labels (1 - 32) correspond to divergence dates and HPD intervals in millions of years ago (Mya) for selected nodes (see Suppl. material 1: Table S 3). Dashed dark blue and light blue lines represent the minimum interspecific and maximum intraspecific variation determined by Sands et al. (2019 a) to aid Theodoxus spp. delimitation (see see Methods, Phylogenetics and species delimitation and that study for more information on the delimitation methods)." figureDoi="10.3897/zse.96.48312.figure2" httpUri="https://binary.pensoft.net/fig/376451" pageId="0" pageNumber="25">2</figureCitation>
);
<emphasis id="0585F1903B7837094E472D98F0D2D49B" bold="true" pageId="0" pageNumber="25">V</emphasis>
. Opercula of a
<taxonomicName id="D53816B505D9B4511E905DB85E85D62B" lsidName="T. danasteri" pageId="0" pageNumber="25" rank="species" species="danasteri">
<emphasis id="1A90AB9773340BBD987F5F3F1E76DE72" italics="true" pageId="0" pageNumber="25">T. danasteri</emphasis>
</taxonomicName>
-morphotype from Lake Skadar, Montenegro (
<collectionCode id="57FAD05F171297C37EED8630B75EB5AC" collectionName="UGSB">UGSB</collectionCode>
24171). Specimens A-D, I-L and T-U were used in the phylogeny (Fig.
<figureCitation id="31F3FD79BFD8198228D8DA41B446179A" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dated phylogeny of the genus Theodoxus constructed with BEAST, based on existing GenBank and newly incorporated data for COI, 16 S and ATPα (Suppl. material 1: Table S 1). Morphospecies where associations are queried are highlighted by a question mark after the species name (see the &quot; Systematic checklist &quot; for more information). Small green (PP ≥ 0.95) and red (PP &lt;0.95) dots on nodes indicate the relative posterior probability support of divergence events. Node labels (1 - 32) correspond to divergence dates and HPD intervals in millions of years ago (Mya) for selected nodes (see Suppl. material 1: Table S 3). Dashed dark blue and light blue lines represent the minimum interspecific and maximum intraspecific variation determined by Sands et al. (2019 a) to aid Theodoxus spp. delimitation (see see Methods, Phylogenetics and species delimitation and that study for more information on the delimitation methods)." figureDoi="10.3897/zse.96.48312.figure2" httpUri="https://binary.pensoft.net/fig/376451" pageId="0" pageNumber="25">2</figureCitation>
). Scale bars:
<quantity id="C9BBDA90B50F2326EBB6C9AA31C8A99A" metricMagnitude="-3" metricUnit="m" metricValue="1.0" unit="mm" value="1.0">1 mm</quantity>
.
</paragraph>
</caption>
<caption id="CAEE0677E5F73C5477E8F437B93331E4" doi="10.3897/zse.96.48312.figure10" httpUri="https://binary.pensoft.net/fig/376459" pageId="0" pageNumber="25" start="Figure 10" startId="F10">
<paragraph id="81485C2ECA9B2D96FDDAB0D9FEFABDD8" pageId="0" pageNumber="25">
<emphasis id="6FADEB633D208CEABFF37FAA0A1F9F53" bold="true" pageId="0" pageNumber="25">Figure 10.</emphasis>
<taxonomicName id="390492377561AFC35EE8D2E8FBB6C08D" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus fluviatilis" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="fluviatilis">
<emphasis id="CC2D0A545DC293F1C88C72748864F068" italics="true" pageId="0" pageNumber="25">Theodoxus fluviatilis</emphasis>
</taxonomicName>
(Linnaeus, 1758).
<emphasis id="FD429940D77239827B1D91B71B169BD6" bold="true" pageId="0" pageNumber="25">A</emphasis>
-
<emphasis id="AB348A4C201BF688EB7538859542DF17" bold="true" pageId="0" pageNumber="25">D</emphasis>
. Specimen (conforming to
<taxonomicName id="47321B984835ABC7563B6645921AF5AF" lsidName="T. sarmaticus" pageId="0" pageNumber="25" rank="species" species="sarmaticus">
<emphasis id="872F2BC82A11F5D628AD937BCEEEF766" italics="true" pageId="0" pageNumber="25">T. sarmaticus</emphasis>
</taxonomicName>
) from the Dnieper-Bug estuary, Ukraine (
<collectionCode id="5D27253CE24A61E6A3E383306CE4EEFB" collectionName="UGSB">UGSB</collectionCode>
18328);
<emphasis id="54C8E4B55FD11966176C54CE507AE9D0" bold="true" pageId="0" pageNumber="25">E</emphasis>
-
<emphasis id="6A732C371DE3ADB2F472DD1037699C04" bold="true" pageId="0" pageNumber="25">H</emphasis>
. Specimen (
<taxonomicName id="F118BA1C7880CD1D3DA530A822D55E6A" lsidName="T. subthermalis" pageId="0" pageNumber="25" rank="species" species="subthermalis">
<emphasis id="1B473EC02A032AFFC0E4BA5EE9A15399" italics="true" pageId="0" pageNumber="25">T. subthermalis</emphasis>
</taxonomicName>
-morphotype) from Novy Afon, Abkhazia, Georgia (
<collectionCode id="04EA23DD96A1E722D60C116168CE44EE" collectionName="UGSB">UGSB</collectionCode>
18402);
<emphasis id="9E131A7609495F0ACF0F16B1944455F0" bold="true" pageId="0" pageNumber="25">I</emphasis>
-
<emphasis id="5CB9B8026F9DD499491F47A9327E4EFF" bold="true" pageId="0" pageNumber="25">K</emphasis>
.
<typeStatus id="CCCB1DE0DFF20559C9FEF9ECE58334E0">Syntype</typeStatus>
of
<taxonomicName id="099C7C91BB6368567E6619FAD6C75601" lsidName="N. saulcyi" pageId="0" pageNumber="25" rank="species" species="saulcyi">
<emphasis id="75D314D669956CA14EBADA131213288B" italics="true" pageId="0" pageNumber="25">N. saulcyi</emphasis>
</taxonomicName>
from near the Penteli Monastery, Greece (MHNG-MOLL-111736);
<emphasis id="EAAD64FB764E2E73473E7644D9F7B49C" bold="true" pageId="0" pageNumber="25">L</emphasis>
. Specimen (
<taxonomicName id="9595325EF984C91BD319DC27521B6B49" lsidName="T. saulcyi" pageId="0" pageNumber="25" rank="species" species="saulcyi">
<emphasis id="6A2AB7A4019FADF09B7B91BB19478599" italics="true" pageId="0" pageNumber="25">T. saulcyi</emphasis>
</taxonomicName>
-morphotype) from Stilos, Crete, Greece (
<collectionCode id="BE7DD64E4F9D18883C960809B7EAC24D" collectionName="UGSB">UGSB</collectionCode>
20519) incorporated into the phylogeny (Fig.
<figureCitation id="6CABB75F4E1B70815DEB73B86E4B24FA" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dated phylogeny of the genus Theodoxus constructed with BEAST, based on existing GenBank and newly incorporated data for COI, 16 S and ATPα (Suppl. material 1: Table S 1). Morphospecies where associations are queried are highlighted by a question mark after the species name (see the &quot; Systematic checklist &quot; for more information). Small green (PP ≥ 0.95) and red (PP &lt;0.95) dots on nodes indicate the relative posterior probability support of divergence events. Node labels (1 - 32) correspond to divergence dates and HPD intervals in millions of years ago (Mya) for selected nodes (see Suppl. material 1: Table S 3). Dashed dark blue and light blue lines represent the minimum interspecific and maximum intraspecific variation determined by Sands et al. (2019 a) to aid Theodoxus spp. delimitation (see see Methods, Phylogenetics and species delimitation and that study for more information on the delimitation methods)." figureDoi="10.3897/zse.96.48312.figure2" httpUri="https://binary.pensoft.net/fig/376451" pageId="0" pageNumber="25">2</figureCitation>
). Scale bars:
<quantity id="B27030FA1800F3A072C7CE814AF0C3EB" metricMagnitude="-3" metricUnit="m" metricValue="1.0" unit="mm" value="1.0">1 mm</quantity>
.
</paragraph>
</caption>
<paragraph id="D11F861CDCC4CB62E4D88D75F90D642F" pageId="0" pageNumber="25">
<bibRefCitation id="ECCC62474E19C4EB41679F861B9410DC" DOI="https://doi.org/10.1016/j.ympev.2006.06.026" author="Bunje, PME" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="0" pageNumber="25" pagination="373 - 387" refId="B35" refString="Bunje, PME, Lindberg, DR, 2007. Lineage divergence of a freshwater snail clade associated with post-Tethys marine basin development. Molecular Phylogenetics and Evolution 42 (2): 373 - 387, DOI: https://doi.org/10.1016/j.ympev.2006.06.026" title="Lineage divergence of a freshwater snail clade associated with post-Tethys marine basin development." url="https://doi.org/10.1016/j.ympev.2006.06.026" volume="42" year="2007">Bunje and Lindberg (2007)</bibRefCitation>
also suggested
<taxonomicName id="C8271CBA6C67276E4010F1A99E9F895A" lsidName="T. velox" pageId="0" pageNumber="25" rank="species" species="velox">
<emphasis id="9780E7D0622A9EF37C8022B5F597D60E" italics="true" pageId="0" pageNumber="25">T. velox</emphasis>
</taxonomicName>
V. Anistratenko in O. Anistratenko et al., 1999 as conspecific with
<taxonomicName id="7972C278244A6178BEE0F53F03308F86" lsidName="T. fluviatilis" pageId="0" pageNumber="25" rank="species" species="fluviatilis">
<emphasis id="345CF333F13A61D582BFE96AC48FED9D" italics="true" pageId="0" pageNumber="25">T. fluviatilis</emphasis>
</taxonomicName>
. However,
<bibRefCitation id="223ADB2CCA9572D2E309BBDEA6A6E7BF" DOI="https://doi.org/10.1111/jbi.13590" author="Sands, AF" journalOrPublisher="Journal of Biogeography" pageId="0" pageNumber="25" pagination="1274 - 1286" refId="B138" refString="Sands, AF, Sereda, SV, Stelbrink, B, Neubauer, TA, Lazarev, S, Wilke, T, Albrecht, C, 2019a. Contributions of biogeographical functions to species accumulation may change over time in refugial regions. Journal of Biogeography 46 (6): 1274 - 1286, DOI: https://doi.org/10.1111/jbi.13590" title="Contributions of biogeographical functions to species accumulation may change over time in refugial regions." url="https://doi.org/10.1111/jbi.13590" volume="46" year="2019 a">Sands et al. (2019a)</bibRefCitation>
found substantial differentiation on the molecular level, indicating a sister relation between
<taxonomicName id="3589D5D01D27572F14CC9C8399FB8504" lsidName="T. fluviatilis" pageId="0" pageNumber="25" rank="species" species="fluviatilis">
<emphasis id="551A4D2E8A013CD4D8F966C7576E3BA2" italics="true" pageId="0" pageNumber="25">T. fluviatilis</emphasis>
</taxonomicName>
and
<taxonomicName id="4BF3BD34227322482CC37747891C3B52" lsidName="T. velox" pageId="0" pageNumber="25" rank="species" species="velox">
<emphasis id="7A5888519780F44E7EBA10D864A8BDDF" italics="true" pageId="0" pageNumber="25">T. velox</emphasis>
</taxonomicName>
(also see Fig.
<figureCitation id="2709103C8EB901577350A0FC06745C37" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dated phylogeny of the genus Theodoxus constructed with BEAST, based on existing GenBank and newly incorporated data for COI, 16 S and ATPα (Suppl. material 1: Table S 1). Morphospecies where associations are queried are highlighted by a question mark after the species name (see the &quot; Systematic checklist &quot; for more information). Small green (PP ≥ 0.95) and red (PP &lt;0.95) dots on nodes indicate the relative posterior probability support of divergence events. Node labels (1 - 32) correspond to divergence dates and HPD intervals in millions of years ago (Mya) for selected nodes (see Suppl. material 1: Table S 3). Dashed dark blue and light blue lines represent the minimum interspecific and maximum intraspecific variation determined by Sands et al. (2019 a) to aid Theodoxus spp. delimitation (see see Methods, Phylogenetics and species delimitation and that study for more information on the delimitation methods)." figureDoi="10.3897/zse.96.48312.figure2" httpUri="https://binary.pensoft.net/fig/376451" pageId="0" pageNumber="25">2</figureCitation>
). Although it may be challenging to differentiate these species morphologically in some instances, there appears to be a stronger callus on the operculum in
<taxonomicName id="273AF680995ACD9A29BF6AA5BE67847D" lsidName="T. fluviatilis" pageId="0" pageNumber="25" rank="species" species="fluviatilis">
<emphasis id="A0DD680AC75707350CCA933A8A48F5FD" italics="true" pageId="0" pageNumber="25">T. fluviatilis</emphasis>
</taxonomicName>
and greater phenotypic plasticity in shell shape and periostracum colouration as compared to
<taxonomicName id="82966086B68CB0F07ED6ABC262D7B4D1" lsidName="T. velox" pageId="0" pageNumber="25" rank="species" species="velox">
<emphasis id="F8ECE32B544FEF2083E704A5C0DCE137" italics="true" pageId="0" pageNumber="25">T. velox</emphasis>
</taxonomicName>
(Figs
<figureCitation id="8DE9877A2CD0B2649A83414718C06522" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Theodoxus fluviatilis (Linnaeus, 1758). A - D. Specimen collected close to Oued Laabid, Morocco (UGSB 18106) (Fig. 2); E - H. Specimen from the Danube River, Woerth an der Donau, Germany (UGSB 24173); I - L. Specimen (T. euxinus - morphotype) collected in the Papuc Creek, Kiyikoey, Turkey (UGSB 24172) (Fig. 2); M - P. Specimen (T. euxinus - morphotype) from Ovidiopol, Ukraine (UGSB 18124); Q - S. Specimen (T. heldreichi - morphotype) collected in Lake Egirdir, Turkey (UGSB 20361); T, U. Specimen (T. danasteri - morphotype) collected in Krasna Kosa, Ukraine (UGSB 18417) (Fig. 2); V. Opercula of a T. danasteri - morphotype from Lake Skadar, Montenegro (UGSB 24171). Specimens A-D, I-L and T-U were used in the phylogeny (Fig. 2). Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure9" httpUri="https://binary.pensoft.net/fig/376458" pageId="0" pageNumber="25">9</figureCitation>
,
<figureCitation id="EDA9EC62C740F76D692AF140C019EB30" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Theodoxus fluviatilis (Linnaeus, 1758). A - D. Specimen (conforming to T. sarmaticus) from the Dnieper-Bug estuary, Ukraine (UGSB 18328); E - H. Specimen (T. subthermalis - morphotype) from Novy Afon, Abkhazia, Georgia (UGSB 18402); I - K. Syntype of N. saulcyi from near the Penteli Monastery, Greece (MHNG-MOLL- 111736); L. Specimen (T. saulcyi - morphotype) from Stilos, Crete, Greece (UGSB 20519) incorporated into the phylogeny (Fig. 2). Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure10" httpUri="https://binary.pensoft.net/fig/376459" pageId="0" pageNumber="25">10</figureCitation>
,
<figureCitation id="822F3880995E7BF0AD381048A92096A7" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Theodoxus velox V. Anistratenko in O. Anistratenko et al., 1999. A - D. Specimen from the Dnieper River, Kiev, Ukraine (UGSB 24186); E - H. Specimen collected in the Danube delta, Vylkove, Ukraine (UGSB 24184) used in the phylogeny (Fig. 2); I, J. Holotype from Dnieper Delta, Zbur'ivka Liman, Ukraine (IZAN); K - M. Specimen from Lake Sapanca, Yeni Esme, Turkey (UGSB 20691). Theodoxus sarmaticus (Lindholm, 1901). N - P; Q - S; T - U. Three syntypes from the Oskol River, near Golubino, Russia stored in ZIN (coll. no. 6099 / 24). Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure22" httpUri="https://binary.pensoft.net/fig/376471" pageId="0" pageNumber="25">22</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="8E7E2CB4C53CAC83F675037D25401AA2" pageId="0" pageNumber="25" type="distribution">
<paragraph id="31FCE22632C842E376B7034720035BCC" pageId="0" pageNumber="25">Distribution.</paragraph>
<paragraph id="C1FDCB59800F98986C9EA1E604281399" pageId="0" pageNumber="25">
<taxonomicName id="5A066CD8FD5EF53FBBCD1D992C02A53D" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus fluviatilis" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="fluviatilis">
<emphasis id="D2F5715D793CEE3A984748935DB1476F" italics="true" pageId="0" pageNumber="25">Theodoxus fluviatilis</emphasis>
</taxonomicName>
is widely distributed across Europe, northern Africa, and western Asia. Records of this species and its synonyms are noted as far as Ireland to the West (
<bibRefCitation id="C3A45158DB2C2D02218D7AAB7627B68B" author="Anderson, R" journalOrPublisher="Journal of Conchology" pageId="0" pageNumber="25" pagination="607 - 638" refId="B12" refString="Anderson, R, 2005. An annotated list of the non-marine molluscs of Britain and Ireland. Journal of Conchology 38 (6): 607 - 638" title="An annotated list of the non-marine molluscs of Britain and Ireland." volume="38" year="2005">Anderson 2005</bibRefCitation>
), Morocco in the South (
<bibRefCitation id="2DAF928605A50662A57BD2FB12BA82FE" author="Taybi, AF" journalOrPublisher="Journal of Materials and Enviromental Sciences" pageId="0" pageNumber="25" pagination="1401 - 1416" publicationUrl="https://www.jmaterenvironsci.com/Document/vol8/vol8_N4/149-JMES-2926-Taybi.pdf" refId="B153" refString="Taybi, AF, Mabrouki, Y, Ghamizi, M, Berrahou, A, 2017. The freshwater malacological composition of Moulouya's watershed and Oriental Morocco. Journal of Materials and Enviromental Sciences 8 (4): 1401 - 1416, https://www.jmaterenvironsci.com/Document/vol8/vol8_N4/149-JMES-2926-Taybi.pdf" title="The freshwater malacological composition of Moulouya's watershed and Oriental Morocco." url="https://www.jmaterenvironsci.com/Document/vol8/vol8_N4/149-JMES-2926-Taybi.pdf" volume="8" year="2017">Taybi et al. 2017</bibRefCitation>
), Sweden and Finland in the North (
<bibRefCitation id="C3E361A8FB3D03D67DB652608897AA1C" DOI="https://doi.org/10.1016/0302-3524(78)90131-7" author="Kangas, P" journalOrPublisher="Estuarine and Coastal Marine Science" pageId="0" pageNumber="25" pagination="409 - 416" refId="B77" refString="Kangas, P, Skoog, G, 1978. Salinity tolerance of Theodoxus fluviatilis (Mollusca, Gastropoda) from freshwater and from different salinity regimes in the Baltic Sea. Estuarine and Coastal Marine Science 6 (4): 409 - 416, DOI: https://doi.org/10.1016/0302-3524(78)90131-7" title="Salinity tolerance of Theodoxus fluviatilis (Mollusca, Gastropoda) from freshwater and from different salinity regimes in the Baltic Sea." url="https://doi.org/10.1016/0302-3524(78)90131-7" volume="6" year="1978">Kangas and Skoog 1978</bibRefCitation>
), and European Russia in the East (
<bibRefCitation id="19BF7409A7A6B2B02AFA38073F59E8E4" author="Vinarski, MV" journalOrPublisher="Severtsov Institute of Ecology and Evolution of RAS, Moscow" pageId="0" pageNumber="25" refId="B157" refString="Vinarski, MV, Kantor, YI, 2016. Analytical catalogue of fresh and brackish water molluscs of Russia and adjacent countries. A.N. Severtsov Institute of Ecology and Evolution of RAS, Moscow" title="Analytical catalogue of fresh and brackish water molluscs of Russia and adjacent countries. A. N." year="2016">Vinarski and Kantor 2016</bibRefCitation>
). In addition, there are numerous records from across Anatolia (
<bibRefCitation id="AECB7AFF84B34E147AD61BBE13368ED1" author="Schuett, H" journalOrPublisher="Basteria" pageId="0" pageNumber="25" pagination="39 - 46" refId="B145" refString="Schuett, H, Sesen, R, 1989b. Theodoxus in South-eastern Anatolia, Turkey (Gastropoda Prosobranchia, Neritidae). Basteria 53 (1-3): 39 - 46" title="Theodoxus in South-eastern Anatolia, Turkey (Gastropoda Prosobranchia, Neritidae)." volume="53" year="1989 b">
<normalizedToken id="7A7E8E4CED18EF1A058FBACA5F58EAA9" originalValue="Schütt">Schuett</normalizedToken>
and
<normalizedToken id="3D94FC3FD2EE5195B8D40D1F8A332AD2" originalValue="Şeşen">Sesen</normalizedToken>
1989b
</bibRefCitation>
,
<bibRefCitation id="99E9EA5CD74C3585960288D3F6A5C0D4" DOI="https://doi.org/10.1080/09397140.1992.10637614" author="Schuett, H" journalOrPublisher="Zoology in the Middle East" pageId="0" pageNumber="25" pagination="63 - 68" refId="B146" refString="Schuett, H, Sesen, R, 1992. The genus Theodoxus in South-western Anatolia, Turkey (Gastropoda, Prosobranchia, Neritidae). Zoology in the Middle East 6 (1): 63 - 68, DOI: https://doi.org/10.1080/09397140.1992.10637614" title="The genus Theodoxus in South-western Anatolia, Turkey (Gastropoda, Prosobranchia, Neritidae)." url="https://doi.org/10.1080/09397140.1992.10637614" volume="6" year="1992">1992</bibRefCitation>
;
<bibRefCitation id="9A85C70D3786FADD7E11F9F075981B4A" author="Yildirim, MZ" journalOrPublisher="Turkish Journal of Zoology" pageId="0" pageNumber="25" pagination="197 - 204" refId="B164" refString="Yildirim, MZ, Koca, SB, Kebapci, Ue, 2006. Supplement to the Prosobranchia (Mollusca: Gastropoda) Fauna of Fresh and Brackish Waters of Turkey. Turkish Journal of Zoology 30: 197 - 204" title="Supplement to the Prosobranchia (Mollusca: Gastropoda) Fauna of Fresh and Brackish Waters of Turkey." volume="30" year="2006">
<normalizedToken id="E2B5E7CDA72DE57129BC6C81D1DB366A" originalValue="Yıldırım">Yildirim</normalizedToken>
et al. 2006
</bibRefCitation>
;
<bibRefCitation id="0ACBDE3FDF386A14B6F010CD8BD2A0D1" author="Guerlek, ME" journalOrPublisher="Fresenius Enviromental Bulletin" pageId="0" pageNumber="25" pagination="2992 - 3013" refId="B69" refString="Guerlek, ME, Sahin, SK, Doekuemcue, N, Yildirim, MZ, 2019. Checklist of the freshwater mollusca of Turkey (Mollusca: Gastropoda, Bivalvia). Fresenius Enviromental Bulletin 28 (4): 2992 - 3013" title="Checklist of the freshwater mollusca of Turkey (Mollusca: Gastropoda, Bivalvia)." volume="28" year="2019">
<normalizedToken id="560F057F5DB1165B155B5EEF66A97D9D" originalValue="Gürlek">Guerlek</normalizedToken>
et al. 2019
</bibRefCitation>
) and Georgia (
<bibRefCitation id="A266C49368F61E6350A397F11253E94D" author="Vinarski, MV" journalOrPublisher="Severtsov Institute of Ecology and Evolution of RAS, Moscow" pageId="0" pageNumber="25" refId="B157" refString="Vinarski, MV, Kantor, YI, 2016. Analytical catalogue of fresh and brackish water molluscs of Russia and adjacent countries. A.N. Severtsov Institute of Ecology and Evolution of RAS, Moscow" title="Analytical catalogue of fresh and brackish water molluscs of Russia and adjacent countries. A. N." year="2016">Vinarski and Kantor 2016</bibRefCitation>
) within the study region. The wide distribution range has been substantiated through molecular studies (
<bibRefCitation id="B562A4275F5951A0C634B7EBE1F7F18B" DOI="https://doi.org/10.1111/j.1365-294X.2005.02703.x" author="Bunje, PME" journalOrPublisher="Molecular Ecology" pageId="0" pageNumber="25" pagination="4323 - 4340" refId="B34" refString="Bunje, PME, 2005. Pan-European phylogeography of the aquatic snail Theodoxus fluviatilis (Gastropoda: Neritidae). Molecular Ecology 14 (14): 4323 - 4340, DOI: https://doi.org/10.1111/j.1365-294X.2005.02703.x" title="Pan-European phylogeography of the aquatic snail Theodoxus fluviatilis (Gastropoda: Neritidae)." url="https://doi.org/10.1111/j.1365-294X.2005.02703.x" volume="14" year="2005">Bunje 2005</bibRefCitation>
;
<bibRefCitation id="9F3A2F803AE293F72EC8CC9FAB9DC13E" DOI="https://doi.org/10.1016/j.ympev.2006.06.026" author="Bunje, PME" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="0" pageNumber="25" pagination="373 - 387" refId="B35" refString="Bunje, PME, Lindberg, DR, 2007. Lineage divergence of a freshwater snail clade associated with post-Tethys marine basin development. Molecular Phylogenetics and Evolution 42 (2): 373 - 387, DOI: https://doi.org/10.1016/j.ympev.2006.06.026" title="Lineage divergence of a freshwater snail clade associated with post-Tethys marine basin development." url="https://doi.org/10.1016/j.ympev.2006.06.026" volume="42" year="2007">Bunje and Lindberg 2007</bibRefCitation>
;
<bibRefCitation id="563406A232F703480E4272A370895A4F" DOI="https://doi.org/10.1111/jbi.13590" author="Sands, AF" journalOrPublisher="Journal of Biogeography" pageId="0" pageNumber="25" pagination="1274 - 1286" refId="B138" refString="Sands, AF, Sereda, SV, Stelbrink, B, Neubauer, TA, Lazarev, S, Wilke, T, Albrecht, C, 2019a. Contributions of biogeographical functions to species accumulation may change over time in refugial regions. Journal of Biogeography 46 (6): 1274 - 1286, DOI: https://doi.org/10.1111/jbi.13590" title="Contributions of biogeographical functions to species accumulation may change over time in refugial regions." url="https://doi.org/10.1111/jbi.13590" volume="46" year="2019 a">Sands et al. 2019a</bibRefCitation>
; Fig.
<figureCitation id="B7877AD294C06ADBAAF4B7CA77DA09BE" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Occurrence map for Theodoxus spp. in Asia. All points conform to specimens used in the phylogenetic analyses (Suppl. material 1: Table S 1), except for T. gloeri for which no molecular data are available and only the type and paratype localities are indicated. Species are partitioned across four identical maps (A - D) to limit overlap of points as far as possible. Barring T. gloeri, closer related species are grouped together (Fig. 2)." figureDoi="10.3897/zse.96.48312.figure3" httpUri="https://binary.pensoft.net/fig/376452" pageId="0" pageNumber="25">3C</figureCitation>
). Some authors suggested that this species also occurs in Iran (
<bibRefCitation id="F8F6E4E828DB6D2A2E42D54E98363F86" author="Roth, G" editor="Krupp, F" journalOrPublisher="Dr. ludwig Reichert Verlag, Wiesbaden, Germany" pageId="0" pageNumber="25" pagination="72 - 79" refId="B136" refString="Roth, G, 1987. Data on the distribution and faunal history of the genus Theodoxus in the Middle East (Gastropoda: Neritidae). In: Krupp, F, Schneider, W, Kinzelbach, R, Eds., Proceedings of the Symposium on the Fauna and Zoogeography of the Middle East. Dr. ludwig Reichert Verlag, Wiesbaden, Germany: 72 - 79" title="Data on the distribution and faunal history of the genus Theodoxus in the Middle East (Gastropoda: Neritidae)." volumeTitle="Proceedings of the Symposium on the Fauna and Zoogeography of the Middle East." year="1987">Roth 1987</bibRefCitation>
;
<bibRefCitation id="D0D8B11B8F3C7AF2FE3D7D39AF0F884E" DOI="https://doi.org/10.3897/zookeys.219.3406" author="Gloeer, P" journalOrPublisher="ZooKeys" pageId="0" pageNumber="25" pagination="11 - 61" refId="B63" refString="Gloeer, P, Pesic, V, 2012. The freshwater snails (Gastropoda) of Iran, with descriptions of two new genera and eight new species. ZooKeys 219: 11 - 61, DOI: https://doi.org/10.3897/zookeys.219.3406" title="The freshwater snails (Gastropoda) of Iran, with descriptions of two new genera and eight new species." url="https://doi.org/10.3897/zookeys.219.3406" volume="219" year="2012">
<normalizedToken id="D9EB8A693677AF71FD855C87D62D544A" originalValue="Glöer">Gloeer</normalizedToken>
and
<normalizedToken id="4A0484599640C867B90F03EF48ABACA7" originalValue="Pešić">Pesic</normalizedToken>
2012
</bibRefCitation>
), but molecular analyses of material from these parts do not support this hypothesis (
<bibRefCitation id="51BADAA45EA2E3D9AE889006292620C7" DOI="https://doi.org/10.1111/jbi.13590" author="Sands, AF" journalOrPublisher="Journal of Biogeography" pageId="0" pageNumber="25" pagination="1274 - 1286" refId="B138" refString="Sands, AF, Sereda, SV, Stelbrink, B, Neubauer, TA, Lazarev, S, Wilke, T, Albrecht, C, 2019a. Contributions of biogeographical functions to species accumulation may change over time in refugial regions. Journal of Biogeography 46 (6): 1274 - 1286, DOI: https://doi.org/10.1111/jbi.13590" title="Contributions of biogeographical functions to species accumulation may change over time in refugial regions." url="https://doi.org/10.1111/jbi.13590" volume="46" year="2019 a">Sands et al. 2019a</bibRefCitation>
,
<bibRefCitation id="2B0FD0C4437CBE409F1F0299922DE00F" DOI="https://doi.org/10.1098/rsos.190965" author="Sands, AF" journalOrPublisher="Royal Society Open Science" pageId="0" pageNumber="25" pagination=": 190965" refId="B139" refString="Sands, AF, Neubauer, TA, Nasibi, S, Fasihi Harandi, M, Anistratenko, VV, Wilke, T, Albrecht, C, 2019b. Old lake vs. young taxa: a comparative phylogeographic perspective on the evolution of Caspian Sea gastropods (Neritidae: Theodoxus). Royal Society Open Science 6 (10): 190965, DOI: https://doi.org/10.1098/rsos.190965" title="Old lake vs. young taxa: a comparative phylogeographic perspective on the evolution of Caspian Sea gastropods (Neritidae: Theodoxus)." url="https://doi.org/10.1098/rsos.190965" volume="6" year="2019 b">2019b</bibRefCitation>
; personal observation A.F.S.).
</paragraph>
</subSubSection>
</treatment>
</document>