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<document id="3E1C2845BFFCDFB17670BADE01A9AF70" ID-DOI="10.11646/zootaxa.4755.2.1" ID-GBIF-Dataset="dc3a4f00-2ae5-4844-a803-7d8dabd65435" ID-ISSN="1175-5334" ID-Zenodo-Dep="3731414" ID-ZooBank="urn:lsid:zoobank.org:pub:CAD4295B-2456-48EE-98F6-723FDEF5C0EB" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1585327475241" checkinUser="carolina" docAuthor="Santos, Allan Paulo Moreira" docDate="2020" docId="D40B8780CA5DFFFBD7F1FE3AFD1BF965" docLanguage="en" docName="zootaxa.4755.2.1.pdf.imf" docOrigin="Zootaxa 4755 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Acostatrichia ujasa Olah &amp; Flint 2012" docType="treatment" docVersion="5" lastPageNumber="225" masterDocId="2832FFF8CA45FFE3D766FFCFFF85FF88" masterDocTitle="A review of the Neotropical microcaddisfly genus Acostatrichia Mosely, 1939 with description of a new species from Brazil (Trichoptera: Hydroptilidae: Leucotrichiinae)" masterLastPageNumber="230" masterPageNumber="201" pageNumber="225" updateTime="1698732921180" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="774FF785EAA46CD13F141D086BA7BED9">A review of the Neotropical microcaddisfly genus Acostatrichia Mosely, 1939 with description of a new species from Brazil (Trichoptera: Hydroptilidae: Leucotrichiinae)</mods:title>
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<mods:namePart id="FFABEEAB05A4FDDDD8E97E1F1912ABB4">Santos, Allan Paulo Moreira</mods:namePart>
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<treatment id="D40B8780CA5DFFFBD7F1FE3AFD1BF965" ID-DOI="http://doi.org/10.5281/zenodo.3812833" ID-GBIF-Taxon="163300399" ID-Zenodo-Dep="3812833" LSID="urn:lsid:plazi:treatment:D40B8780CA5DFFFBD7F1FE3AFD1BF965" httpUri="http://treatment.plazi.org/id/D40B8780CA5DFFFBD7F1FE3AFD1BF965" lastPageNumber="225" pageId="24" pageNumber="225">
<subSubSection id="14B8651DCA5DFFFBD7F1FE3AFDE9FD98" box="[151,620,501,528]" pageId="24" pageNumber="225" type="nomenclature">
<paragraph id="5C1D3696CA5DFFFBD7F1FE3AFDE9FD98" blockId="24.[151,620,501,564]" box="[151,620,501,528]" pageId="24" pageNumber="225">
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<taxonomicName id="9BA24D15CA5DFFFBD7F1FE3AFDE9FD98" authority="Olah &amp; Flint 2012" authorityName="Olah &amp; Flint" authorityYear="2012" box="[151,620,501,528]" class="Insecta" family="Hydroptilidae" genus="Acostatrichia" kingdom="Animalia" order="Trichoptera" pageId="24" pageNumber="225" phylum="Arthropoda" rank="species" species="ujasa">
<emphasis id="6ED6EA84CA5DFFFBD7F1FE3AFDE9FD98" bold="true" box="[151,620,501,528]" pageId="24" pageNumber="225">
<emphasis id="6ED6EA84CA5DFFFBD7F1FE3AFE04FD98" bold="true" box="[151,385,501,528]" italics="true" pageId="24" pageNumber="225">Acostatrichia ujasa</emphasis>
<bibRefCitation id="38334B67CA5DFFFBD6EEFE3AFDE9FD98" author="Olah, J. &amp; Flint, O. S. Jr." box="[392,620,501,528]" firstAuthor="Olah" journalOrPublisher="Annales Historico-Naturales Musei Nationalis Hungarici" pageId="24" pageNumber="225" pagination="131 - 213" part="104" refId="ref15616" refString="Olah, J. &amp; Flint, O. S. Jr. (2012) Description of new species in the Leucotrichiini tribe (Trichoptera: Hydroptilidae). Annales Historico-Naturales Musei Nationalis Hungarici, 104, 131 - 213." title="Description of new species in the Leucotrichiini tribe (Trichoptera: Hydroptilidae)" type="journal article" year="2012">Oláh &amp; Flint 2012</bibRefCitation>
</emphasis>
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</heading>
</paragraph>
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<subSubSection id="14B8651DCA5DFFFBD7F1FDD5FEDDFDBC" box="[151,344,537,564]" pageId="24" pageNumber="225" type="description">
<paragraph id="5C1D3696CA5DFFFBD7F1FDD5FEDDFDBC" blockId="24.[151,620,501,564]" box="[151,344,537,564]" pageId="24" pageNumber="225">
<figureCitation id="C4992A13CA5DFFFBD7F1FDD5FF76FDBC" box="[151,243,538,564]" captionStart="FIGURE 14" captionStartId="25.[151,250,1952,1977]" captionTargetBox="[163,1424,182,1927]" captionTargetId="figure@25.[162,1425,181,1927]" captionTargetPageId="25" captionText="FIGURE 14. Acostatrichia ujasa Oláh &amp; Flint 2012, paratype male, genitalia. 14A, ventral; 14B, dorsal; 14C, left lateral; 14D, phallus, dorsal; 14E, phallus, left lateral. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3731446" httpUri="https://zenodo.org/record/3731446/files/figure.png" pageId="24" pageNumber="225">Figs. 14</figureCitation>
,
<figureCitation id="C4992A13CA5DFFFBD799FDD6FEB4FDBC" box="[255,305,537,564]" captionStart="FIGURE 15" captionStartId="26.[151,250,1003,1028]" captionTargetBox="[155,1431,185,974]" captionTargetId="figure@26.[151,1435,181,977]" captionTargetPageId="26" captionText="FIGURE 15. Acostatrichia spp., adult male habitus. 15A, Acostatrichia darda Oláh &amp; Flint 2012 (paratype male of synonym A. pika Oláh &amp; Flint 2012); 15B, Acostatrichia kihara Oláh &amp; Flint 2012, paratype male; 15C, Acostatrichia ujasa Oláh &amp; Flint 2012, paratype male. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3731448" httpUri="https://zenodo.org/record/3731448/files/figure.png" pageId="24" pageNumber="225">15C</figureCitation>
,
<figureCitation id="C4992A13CA5DFFFBD65AFDD5FEDDFDBC" box="[316,344,538,564]" captionStart="FIGURE 16" captionStartId="27.[151,250,1798,1823]" captionTargetBox="[191,1387,622,1773]" captionTargetId="figure@27.[191,1397,622,1773]" captionTargetPageId="27" captionText="FIGURE 16. Distribution map of the Acostatrichia species in South America, based on material examined here and previously published records. When latitude and longitude were not available, an approximate location was used. Circle, species in the Acostatrichia plaumanni Group; square, species in the Acostatrichia brevipenis Group; triangle, species in the Acostatrichia cerna Group." figureDoi="http://doi.org/10.5281/zenodo.3731450" httpUri="https://zenodo.org/record/3731450/files/figure.png" pageId="24" pageNumber="225">16</figureCitation>
</paragraph>
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<subSubSection id="14B8651DCA5DFFFBD7F1FD91FC6CFD3C" pageId="24" pageNumber="225" type="reference_group">
<paragraph id="5C1D3696CA5DFFFBD7F1FD91FD3AFD1C" blockId="24.[151,1436,605,692]" pageId="24" pageNumber="225">
<treatmentCitationGroup id="7CB211B8CA5DFFFBD7F1FD91FD3AFD1C" pageId="24" pageNumber="225">
<treatmentCitation id="DD031087CA5DFFFBD7F1FD91FDD3FDFE" author="Olah, J. &amp; Flint, O. S. Jr." box="[151,598,605,630]" page="158" pageId="24" pageNumber="225" year="2012">
<taxonomicName id="9BA24D15CA5DFFFBD7F1FD91FDD3FDFE" authority="Olah &amp; Flint 2012: 158" authorityName="Olah &amp; Flint" authorityPageNumber="158" authorityYear="2012" box="[151,598,605,630]" class="Insecta" family="Hydroptilidae" genus="Acostatrichia" kingdom="Animalia" order="Trichoptera" pageId="24" pageNumber="225" phylum="Arthropoda" rank="species" species="ujasa">
<emphasis id="6ED6EA84CA5DFFFBD7F1FD91FED8FDFE" box="[151,349,606,630]" italics="true" pageId="24" pageNumber="225">Acostatrichia ujasa</emphasis>
<bibRefCitation id="38334B67CA5DFFFBD602FD92FDD3FDFE" author="Olah, J. &amp; Flint, O. S. Jr." box="[356,598,605,630]" firstAuthor="Olah" journalOrPublisher="Annales Historico-Naturales Musei Nationalis Hungarici" pageId="24" pageNumber="225" pagination="131 - 213" part="104" refId="ref15616" refString="Olah, J. &amp; Flint, O. S. Jr. (2012) Description of new species in the Leucotrichiini tribe (Trichoptera: Hydroptilidae). Annales Historico-Naturales Musei Nationalis Hungarici, 104, 131 - 213." title="Description of new species in the Leucotrichiini tribe (Trichoptera: Hydroptilidae)" type="journal article" year="2012">Oláh &amp; Flint 2012: 158</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, figs. 41–43, male;
<typeStatus id="83198834CA5DFFFBD479FD90FCCEFDFF" box="[799,843,607,631]" pageId="24" pageNumber="225">type</typeStatus>
locality:
<collectingCountry id="24B57606CA5DFFFBD4CAFD92FB81FDFE" box="[940,1028,605,630]" name="Ecuador" pageId="24" pageNumber="225">Ecuador</collectingCountry>
:
<collectingRegion id="9E66F874CA5DFFFBD368FD90FB3BFDFE" box="[1038,1214,605,630]" country="Ecuador" name="Pastaza" pageId="24" pageNumber="225">Pastaza Province</collectingRegion>
, Puyo (
<quantity id="9B5A9B73CA5DFFFBD26CFD91FACEFDFE" box="[1290,1355,605,630]" metricMagnitude="4" metricUnit="m" metricValue="2.7" pageId="24" pageNumber="225" unit="km" value="27.0">27 km</quantity>
North), Estacion Fluviometrica;
<typeStatus id="83198834CA5DFFFBD6A6FDB2FE69FD1D" box="[448,492,637,661]" pageId="24" pageNumber="225">type</typeStatus>
depository: NMNH.
</treatmentCitationGroup>
</paragraph>
<paragraph id="5C1D3696CA5DFFFBD7F1FD54FC6CFD3C" blockId="24.[151,1436,605,692]" box="[151,1001,667,692]" pageId="24" pageNumber="225">
<treatmentCitationGroup id="7CB211B8CA5DFFFBD7F1FD54FC6CFD3C" box="[151,1001,667,692]" pageId="24" pageNumber="225">
<taxonomicName id="9BA24D15CA5DFFFBD7F1FD54FEB3FD3C" box="[151,310,667,692]" class="Insecta" family="Hydroptilidae" kingdom="Animalia" order="Trichoptera" pageId="24" pageNumber="225" phylum="Arthropoda" rank="subFamily" subFamily="Leucotrichiinae">Leucotrichiinae</taxonomicName>
genus B
<treatmentCitation id="DD031087CA5DFFFBD6F3FD54FD05FD3C" author="Santos, A. P. M. &amp; Nessimian, J. L. &amp; Takiya, D. M." box="[405,640,667,692]" page="462" pageId="24" pageNumber="225" year="2016">
<bibRefCitation id="38334B67CA5DFFFBD6F3FD54FD05FD3C" author="Santos, A. P. M. &amp; Nessimian, J. L. &amp; Takiya, D. M." box="[405,640,667,692]" etAl="et al." firstAuthor="Santos" journalOrPublisher="Systematic Entomology" pageId="24" pageNumber="225" pagination="458 - 480" part="41" refId="ref15814" refString="Santos, A. P. M., Nessimian, J. L. &amp; Takiya, D. M. (2016) Revised classification and evolution of leucotrichiine microcaddisflies (Trichoptera: Hydroptilidae) based on morphological and molecular data. Systematic Entomology, 41, 458 - 480. https: // doi. org / 10.1111 / syen. 12168" title="Revised classification and evolution of leucotrichiine microcaddisflies (Trichoptera: Hydroptilidae) based on morphological and molecular data" type="journal article" year="2016">
Santos
<emphasis id="6ED6EA84CA5DFFFBD6B9FD53FD89FD3B" box="[479,524,667,691]" italics="true" pageId="24" pageNumber="225">et al</emphasis>
. 2016: 462
</bibRefCitation>
</treatmentCitation>
, figs. 4, 5; phylogenetic placement.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="14B8651DCA5DFFFBD7F1FD2EFC88FA0C" pageId="24" pageNumber="225" type="description">
<paragraph id="5C1D3696CA5DFFFBD7F1FD2EFC88FA0C" blockId="24.[151,1437,737,1773]" pageId="24" pageNumber="225">
<emphasis id="6ED6EA84CA5DFFFBD7F1FD2EFEC0FD74" bold="true" box="[151,325,737,764]" pageId="24" pageNumber="225">Redescription.</emphasis>
Length from front of head to tips of folded forewings 4.0–
<quantity id="9B5A9B73CA5DFFFBD4B6FD2EFBA0FD74" box="[976,1061,737,764]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="24" pageNumber="225" unit="mm" value="4.5">4.5 mm</quantity>
(n = 6). General color, in alcohol, brown. Pinned specimen brown, with spots of green setae on head, thorax, and wings (
<figureCitation id="C4992A13CA5DFFFBD31DFCC9FB66FCA8" box="[1147,1251,773,800]" captionStart="FIGURE 15" captionStartId="26.[151,250,1003,1028]" captionTargetBox="[155,1431,185,974]" captionTargetId="figure@26.[151,1435,181,977]" captionTargetPageId="26" captionText="FIGURE 15. Acostatrichia spp., adult male habitus. 15A, Acostatrichia darda Oláh &amp; Flint 2012 (paratype male of synonym A. pika Oláh &amp; Flint 2012); 15B, Acostatrichia kihara Oláh &amp; Flint 2012, paratype male; 15C, Acostatrichia ujasa Oláh &amp; Flint 2012, paratype male. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3731448" httpUri="https://zenodo.org/record/3731448/files/figure.png" pageId="24" pageNumber="225">Fig. 15C</figureCitation>
). Head unmodified. Ocelli 3. Antennae each 21-articulated; scape cylindrical, twice as long as wide, inner margin not produced; pedicel cylindrical; flagellomeres cylindrical, unmodified. Forewings each with costal vein simple, unmodified. Abdominal segment VII bearing long ventromesal process, with acute apex (
<figureCitation id="C4992A13CA5DFFFBD488FCBDFB18FC04" box="[1006,1181,882,908]" captionStart="FIGURE 14" captionStartId="25.[151,250,1952,1977]" captionTargetBox="[163,1424,182,1927]" captionTargetId="figure@25.[162,1425,181,1927]" captionTargetPageId="25" captionText="FIGURE 14. Acostatrichia ujasa Oláh &amp; Flint 2012, paratype male, genitalia. 14A, ventral; 14B, dorsal; 14C, left lateral; 14D, phallus, dorsal; 14E, phallus, left lateral. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3731446" httpUri="https://zenodo.org/record/3731446/files/figure.png" pageId="24" pageNumber="225">Figs. 14A, 14C</figureCitation>
). Segment VIII shorter dorsally than ventrally (
<figureCitation id="C4992A13CA5DFFFBD6C6FC59FD80FC38" box="[416,517,918,944]" captionStart="FIGURE 14" captionStartId="25.[151,250,1952,1977]" captionTargetBox="[163,1424,182,1927]" captionTargetId="figure@25.[162,1425,181,1927]" captionTargetPageId="25" captionText="FIGURE 14. Acostatrichia ujasa Oláh &amp; Flint 2012, paratype male, genitalia. 14A, ventral; 14B, dorsal; 14C, left lateral; 14D, phallus, dorsal; 14E, phallus, left lateral. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3731446" httpUri="https://zenodo.org/record/3731446/files/figure.png" pageId="24" pageNumber="225">Fig. 14C</figureCitation>
); in ventral view, posterior margin of sternum with two lobes forming deep median U-shaped incision surrounded by short setae (
<figureCitation id="C4992A13CA5DFFFBD5C0FC75FC95FC5C" box="[678,784,954,980]" captionStart="FIGURE 14" captionStartId="25.[151,250,1952,1977]" captionTargetBox="[163,1424,182,1927]" captionTargetId="figure@25.[162,1425,181,1927]" captionTargetPageId="25" captionText="FIGURE 14. Acostatrichia ujasa Oláh &amp; Flint 2012, paratype male, genitalia. 14A, ventral; 14B, dorsal; 14C, left lateral; 14D, phallus, dorsal; 14E, phallus, left lateral. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3731446" httpUri="https://zenodo.org/record/3731446/files/figure.png" pageId="24" pageNumber="225">Fig. 14A</figureCitation>
); without lateral processes, but with apicoventral margin projecting in lateral view (
<figureCitation id="C4992A13CA5DFFFBD6DCFC11FD9AFC70" box="[442,543,990,1016]" captionStart="FIGURE 14" captionStartId="25.[151,250,1952,1977]" captionTargetBox="[163,1424,182,1927]" captionTargetId="figure@25.[162,1425,181,1927]" captionTargetPageId="25" captionText="FIGURE 14. Acostatrichia ujasa Oláh &amp; Flint 2012, paratype male, genitalia. 14A, ventral; 14B, dorsal; 14C, left lateral; 14D, phallus, dorsal; 14E, phallus, left lateral. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3731446" httpUri="https://zenodo.org/record/3731446/files/figure.png" pageId="24" pageNumber="225">Fig. 14C</figureCitation>
); tergum with transverse row of long setae (
<figureCitation id="C4992A13CA5DFFFBD366FC11FBE3FC70" box="[1024,1126,990,1016]" captionStart="FIGURE 14" captionStartId="25.[151,250,1952,1977]" captionTargetBox="[163,1424,182,1927]" captionTargetId="figure@25.[162,1425,181,1927]" captionTargetPageId="25" captionText="FIGURE 14. Acostatrichia ujasa Oláh &amp; Flint 2012, paratype male, genitalia. 14A, ventral; 14B, dorsal; 14C, left lateral; 14D, phallus, dorsal; 14E, phallus, left lateral. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3731446" httpUri="https://zenodo.org/record/3731446/files/figure.png" pageId="24" pageNumber="225">Fig. 14B</figureCitation>
). Segment IX mostly within segment VIII, ventrally open, with pair of long digitiform dorsolateral processes with slightly dilated apices, slightly curved gradually laterad except apices curved slightly inwards in ventral and dorsal views (
<figureCitation id="C4992A13CA5DFFFBD3EEFBE9FAB0FBC8" box="[1160,1333,1062,1088]" captionStart="FIGURE 14" captionStartId="25.[151,250,1952,1977]" captionTargetBox="[163,1424,182,1927]" captionTargetId="figure@25.[162,1425,181,1927]" captionTargetPageId="25" captionText="FIGURE 14. Acostatrichia ujasa Oláh &amp; Flint 2012, paratype male, genitalia. 14A, ventral; 14B, dorsal; 14C, left lateral; 14D, phallus, dorsal; 14E, phallus, left lateral. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3731446" httpUri="https://zenodo.org/record/3731446/files/figure.png" pageId="24" pageNumber="225">Figs. 14A, 14B</figureCitation>
), slightly upturned in lateral view (
<figureCitation id="C4992A13CA5DFFFBD6DEFB85FDA5FBEC" box="[440,544,1098,1124]" captionStart="FIGURE 14" captionStartId="25.[151,250,1952,1977]" captionTargetBox="[163,1424,182,1927]" captionTargetId="figure@25.[162,1425,181,1927]" captionTargetPageId="25" captionText="FIGURE 14. Acostatrichia ujasa Oláh &amp; Flint 2012, paratype male, genitalia. 14A, ventral; 14B, dorsal; 14C, left lateral; 14D, phallus, dorsal; 14E, phallus, left lateral. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3731446" httpUri="https://zenodo.org/record/3731446/files/figure.png" pageId="24" pageNumber="225">Fig. 14C</figureCitation>
); each with four very long, strong, and curved apical spines (
<figureCitation id="C4992A13CA5DFFFBD3B8FB85FABEFBEC" box="[1246,1339,1098,1124]" captionStart="FIGURE 14" captionStartId="25.[151,250,1952,1977]" captionTargetBox="[163,1424,182,1927]" captionTargetId="figure@25.[162,1425,181,1927]" captionTargetPageId="25" captionText="FIGURE 14. Acostatrichia ujasa Oláh &amp; Flint 2012, paratype male, genitalia. 14A, ventral; 14B, dorsal; 14C, left lateral; 14D, phallus, dorsal; 14E, phallus, left lateral. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3731446" httpUri="https://zenodo.org/record/3731446/files/figure.png" pageId="24" pageNumber="225">Figs. 14</figureCitation>
A–14C). Preanal process absent (
<figureCitation id="C4992A13CA5DFFFBD6C5FBA1FD8CFB00" box="[419,521,1134,1160]" captionStart="FIGURE 14" captionStartId="25.[151,250,1952,1977]" captionTargetBox="[163,1424,182,1927]" captionTargetId="figure@25.[162,1425,181,1927]" captionTargetPageId="25" captionText="FIGURE 14. Acostatrichia ujasa Oláh &amp; Flint 2012, paratype male, genitalia. 14A, ventral; 14B, dorsal; 14C, left lateral; 14D, phallus, dorsal; 14E, phallus, left lateral. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3731446" httpUri="https://zenodo.org/record/3731446/files/figure.png" pageId="24" pageNumber="225">Fig. 14B</figureCitation>
). Inferior appendages fused with each other only basally, distal portions free, each short, apically rounded and with subapical incision on inner margin (
<figureCitation id="C4992A13CA5DFFFBD4FBFB5DFB80FB24" box="[925,1029,1170,1196]" captionStart="FIGURE 14" captionStartId="25.[151,250,1952,1977]" captionTargetBox="[163,1424,182,1927]" captionTargetId="figure@25.[162,1425,181,1927]" captionTargetPageId="25" captionText="FIGURE 14. Acostatrichia ujasa Oláh &amp; Flint 2012, paratype male, genitalia. 14A, ventral; 14B, dorsal; 14C, left lateral; 14D, phallus, dorsal; 14E, phallus, left lateral. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3731446" httpUri="https://zenodo.org/record/3731446/files/figure.png" pageId="24" pageNumber="225">Fig. 14A</figureCitation>
); with pair of very long rod-like lateral processes rising from basal area, dilatated at apices (
<figureCitation id="C4992A13CA5DFFFBD477FB79FCFCFB58" box="[785,889,1206,1232]" captionStart="FIGURE 14" captionStartId="25.[151,250,1952,1977]" captionTargetBox="[163,1424,182,1927]" captionTargetId="figure@25.[162,1425,181,1927]" captionTargetPageId="25" captionText="FIGURE 14. Acostatrichia ujasa Oláh &amp; Flint 2012, paratype male, genitalia. 14A, ventral; 14B, dorsal; 14C, left lateral; 14D, phallus, dorsal; 14E, phallus, left lateral. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3731446" httpUri="https://zenodo.org/record/3731446/files/figure.png" pageId="24" pageNumber="225">Fig. 14A</figureCitation>
); in lateral view, upturned (
<figureCitation id="C4992A13CA5DFFFBD3CDFB79FA94FB58" box="[1195,1297,1206,1232]" captionStart="FIGURE 14" captionStartId="25.[151,250,1952,1977]" captionTargetBox="[163,1424,182,1927]" captionTargetId="figure@25.[162,1425,181,1927]" captionTargetPageId="25" captionText="FIGURE 14. Acostatrichia ujasa Oláh &amp; Flint 2012, paratype male, genitalia. 14A, ventral; 14B, dorsal; 14C, left lateral; 14D, phallus, dorsal; 14E, phallus, left lateral. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3731446" httpUri="https://zenodo.org/record/3731446/files/figure.png" pageId="24" pageNumber="225">Fig. 14C</figureCitation>
). Subgenital plate, in ventral view, broad, with sclerotized belt basally and with V-shaped incision on posterior margin (
<figureCitation id="C4992A13CA5DFFFBD209FB15FF4EFA90" captionStart="FIGURE 14" captionStartId="25.[151,250,1952,1977]" captionTargetBox="[163,1424,182,1927]" captionTargetId="figure@25.[162,1425,181,1927]" captionTargetPageId="25" captionText="FIGURE 14. Acostatrichia ujasa Oláh &amp; Flint 2012, paratype male, genitalia. 14A, ventral; 14B, dorsal; 14C, left lateral; 14D, phallus, dorsal; 14E, phallus, left lateral. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3731446" httpUri="https://zenodo.org/record/3731446/files/figure.png" pageId="24" pageNumber="225">Fig. 14A</figureCitation>
); in lateral view, directed posterad and rounded apically (
<figureCitation id="C4992A13CA5DFFFBD43DFB31FC41FA90" box="[859,964,1278,1304]" captionStart="FIGURE 14" captionStartId="25.[151,250,1952,1977]" captionTargetBox="[163,1424,182,1927]" captionTargetId="figure@25.[162,1425,181,1927]" captionTargetPageId="25" captionText="FIGURE 14. Acostatrichia ujasa Oláh &amp; Flint 2012, paratype male, genitalia. 14A, ventral; 14B, dorsal; 14C, left lateral; 14D, phallus, dorsal; 14E, phallus, left lateral. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3731446" httpUri="https://zenodo.org/record/3731446/files/figure.png" pageId="24" pageNumber="225">Fig. 14C</figureCitation>
). Tergum X membranous, bilobed, dorsal lobe quadrangular, ventral lobe pentagonal in dorsal view (
<figureCitation id="C4992A13CA5DFFFBD45BFAEDFC20FAB4" box="[829,933,1314,1340]" captionStart="FIGURE 14" captionStartId="25.[151,250,1952,1977]" captionTargetBox="[163,1424,182,1927]" captionTargetId="figure@25.[162,1425,181,1927]" captionTargetPageId="25" captionText="FIGURE 14. Acostatrichia ujasa Oláh &amp; Flint 2012, paratype male, genitalia. 14A, ventral; 14B, dorsal; 14C, left lateral; 14D, phallus, dorsal; 14E, phallus, left lateral. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3731446" httpUri="https://zenodo.org/record/3731446/files/figure.png" pageId="24" pageNumber="225">Fig. 14B</figureCitation>
). Phallus tubular basally, bearing midlength complex, with dorsal window and basal loop as long as basal portion (
<figureCitation id="C4992A13CA5DFFFBD4FBFA89FB80FAE8" box="[925,1029,1350,1376]" captionStart="FIGURE 14" captionStartId="25.[151,250,1952,1977]" captionTargetBox="[163,1424,182,1927]" captionTargetId="figure@25.[162,1425,181,1927]" captionTargetPageId="25" captionText="FIGURE 14. Acostatrichia ujasa Oláh &amp; Flint 2012, paratype male, genitalia. 14A, ventral; 14B, dorsal; 14C, left lateral; 14D, phallus, dorsal; 14E, phallus, left lateral. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3731446" httpUri="https://zenodo.org/record/3731446/files/figure.png" pageId="24" pageNumber="225">Fig. 14D</figureCitation>
); apical portion with plate-like sclerite and more than eight internal spines (
<figureCitation id="C4992A13CA5DFFFBD534FAA5FC84FA0C" box="[594,769,1386,1412]" captionStart="FIGURE 14" captionStartId="25.[151,250,1952,1977]" captionTargetBox="[163,1424,182,1927]" captionTargetId="figure@25.[162,1425,181,1927]" captionTargetPageId="25" captionText="FIGURE 14. Acostatrichia ujasa Oláh &amp; Flint 2012, paratype male, genitalia. 14A, ventral; 14B, dorsal; 14C, left lateral; 14D, phallus, dorsal; 14E, phallus, left lateral. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3731446" httpUri="https://zenodo.org/record/3731446/files/figure.png" pageId="24" pageNumber="225">Figs. 14D, 14E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="14B8651DCA5DFFFBD7A1FA42FD7CF99C" pageId="24" pageNumber="225" type="materials_examined">
<paragraph id="5C1D3696CA5DFFFBD7A1FA42FBE0FA44" blockId="24.[151,1437,737,1773]" pageId="24" pageNumber="225">
<emphasis id="6ED6EA84CA5DFFFBD7A1FA42FE36FA20" bold="true" box="[199,435,1421,1448]" pageId="24" pageNumber="225">Material examined.</emphasis>
<materialsCitation id="ECCA3CCBCA5DFFFBD6DDFA42FDA2FA44" ID-GBIF-Occurrence="2592188793" collectingDate="1976-02-04" collectionCode="NMNH" collectorName="Kms. N." country="Ecuador" location="Ecuador" pageId="24" pageNumber="225" specimenCount="1" typeStatus="holotype">
<emphasis id="6ED6EA84CA5DFFFBD6DDFA42FD23FA20" bold="true" box="[443,678,1421,1448]" pageId="24" pageNumber="225">
<typeStatus id="83198834CA5DFFFBD6DDFA42FDDFFA20" box="[443,602,1421,1448]" pageId="24" pageNumber="225" type="holotype">HOLOTYPE</typeStatus>
male:
</emphasis>
<collectingCountry id="24B57606CA5DFFFBD5C9FA41FC8BFA20" box="[687,782,1422,1448]" name="Ecuador" pageId="24" pageNumber="225">Ecuador</collectingCountry>
, Past., Puyo (27
<collectorName id="F1575340CA5DFFFBD4B7FA41FBB7FA20" box="[977,1074,1422,1448]" pageId="24" pageNumber="225">Kms. N.</collectorName>
) Est. Fluv. Metrica,
<date id="281C1056CA5DFFFBD246FA41FF57FA44" pageId="24" pageNumber="225" value="1976-02-04">
<collectingDate id="3858E9BECA5DFFFBD246FA41FF57FA44" pageId="24" pageNumber="225" value="1976-02-04">4 February 1976</collectingDate>
</date>
, Spangler
<emphasis id="6ED6EA84CA5DFFFBD62EFA7CFEFCFA44" box="[328,377,1458,1484]" italics="true" pageId="24" pageNumber="225">et al</emphasis>
. leg. (
<collectionCode id="3AB3AE53CA5DFFFBD6D9FA7DFD9EFA44" box="[447,539,1458,1484]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="24" pageNumber="225" type="Museum">NMNH</collectionCode>
);
</materialsCitation>
<materialsCitation id="ECCA3CCBCA5DFFFBD548FA7EFBE0FA44" ID-GBIF-Occurrence="2592188797" box="[558,1125,1457,1484]" collectionCode="NMNH" pageId="24" pageNumber="225" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<emphasis id="6ED6EA84CA5DFFFBD548FA7EFD4EFA44" bold="true" box="[558,715,1457,1484]" pageId="24" pageNumber="225">
<typeStatus id="83198834CA5DFFFBD548FA7EFD47FA44" box="[558,706,1457,1484]" pageId="24" pageNumber="225" type="paratype">PARATYPE</typeStatus>
:
</emphasis>
same data,
<specimenCount id="4AA4FD1FCA5DFFFBD436FA7DFC1EFA44" box="[848,923,1458,1484]" pageId="24" pageNumber="225" type="male">1 male</specimenCount>
pinned (
<collectionCode id="3AB3AE53CA5DFFFBD49BFA7DFBDCFA44" box="[1021,1113,1458,1484]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="24" pageNumber="225" type="Museum">NMNH</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="5C1D3696CA5DFFFBD7A1FA1AFD7CF99C" blockId="24.[151,1437,737,1773]" pageId="24" pageNumber="225">
<materialsCitation id="ECCA3CCBCA5DFFFBD7A1FA1AFD7CF99C" ID-GBIF-Occurrence="2592188798" collectingDate="2012-08-20" collectionCode="DM, DZRJ" country="Peru" elevation="633" latitude="13.2175000°" municipality="Quincemil" pageId="24" pageNumber="225" specimenCount="3" specimenCount-male="3" stateProvince="Cusco">
<emphasis id="6ED6EA84CA5DFFFBD7A1FA1AFE3FFA78" bold="true" box="[199,442,1493,1520]" pageId="24" pageNumber="225">Additional material.</emphasis>
<collectingCountry id="24B57606CA5DFFFBD6A7FA19FE72FA78" box="[449,503,1494,1520]" name="Peru" pageId="24" pageNumber="225">Peru</collectingCountry>
,
<collectingRegion id="9E66F874CA5DFFFBD564FA19FDCEFA78" box="[514,587,1494,1520]" country="Peru" name="Cusco" pageId="24" pageNumber="225">Cusco</collectingRegion>
,
<collectingMunicipality id="BC79ACECCA5DFFFBD531FA19FD55FA78" box="[599,720,1494,1520]" pageId="24" pageNumber="225">Quincemil</collectingMunicipality>
, stream, S
<geoCoordinate id="39965051CA5DFFFBD425FA19FC3DFA78" box="[835,952,1494,1520]" pageId="24" pageNumber="225">13°13’03”</geoCoordinate>
W
<geoCoordinate id="39965051CA5DFFFBD4BBFA19FBD5FA78" box="[989,1104,1494,1520]" pageId="24" pageNumber="225">73°43’40”</geoCoordinate>
, el.
<quantity id="9B5A9B73CA5DFFFBD318FA19FB4DFA78" box="[1150,1224,1494,1520]" metricMagnitude="2" metricUnit="m" metricValue="6.33" pageId="24" pageNumber="225" unit="m" value="633.0">
<elevation id="D78FD1A5CA5DFFFBD318FA19FB4DFA78" box="[1150,1224,1494,1520]" metricMagnitude="2" metricUnit="m" metricValue="6.33" pageId="24" pageNumber="225" unit="m" value="633.0">633 m</elevation>
</quantity>
,
<date id="281C1056CA5DFFFBD3B5FA19FAD8FA78" box="[1235,1373,1494,1520]" pageId="24" pageNumber="225" value="2012-08-20">
<collectingDate id="3858E9BECA5DFFFBD3B5FA19FAD8FA78" box="[1235,1373,1494,1520]" pageId="24" pageNumber="225" value="2012-08-20">20.viii.2012</collectingDate>
</date>
, light trap, APM
<location id="597D604DCA5DFFFBD675FA35FEDBF99C" LSID="urn:lsid:plazi:treatment:D40B8780CA5DFFFBD7F1FE3AFD1BF965:597D604DCA5DFFFBD675FA35FEDBF99C" box="[275,350,1530,1556]" country="Peru" municipality="Quincemil" name="Santos" pageId="24" pageNumber="225" stateProvince="Cusco">Santos</location>
&amp;
<collectionCode id="3AB3AE53CA5DFFFBD6E7FA35FE2BF99C" box="[385,430,1530,1556]" country="New Zealand" name="Dominion Museum" pageId="24" pageNumber="225">DM</collectionCode>
<location id="597D604DCA5DFFFBD6D3FA35FD84F99C" LSID="urn:lsid:plazi:treatment:D40B8780CA5DFFFBD7F1FE3AFD1BF965:597D604DCA5DFFFBD6D3FA35FD84F99C" box="[437,513,1530,1556]" country="Peru" municipality="Quincemil" name="Takiya" pageId="24" pageNumber="225" stateProvince="Cusco">Takiya</location>
leg.,
<specimenCount id="4AA4FD1FCA5DFFFBD559FA35FD10F99C" box="[575,661,1530,1556]" pageId="24" pageNumber="225" type="male">3 males</specimenCount>
(
<collectionCode id="3AB3AE53CA5DFFFBD5C2FA35FD68F99C" box="[676,749,1530,1556]" pageId="24" pageNumber="225">DZRJ</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="14B8651DCA5DFFFBD7A1F9D1FDAEF940" pageId="24" pageNumber="225" type="discussion">
<paragraph id="5C1D3696CA5DFFFBD7A1F9D1FDAEF940" blockId="24.[151,1437,737,1773]" pageId="24" pageNumber="225">
<emphasis id="6ED6EA84CA5DFFFBD7A1F9D1FEBEF9B0" bold="true" box="[199,315,1566,1592]" pageId="24" pageNumber="225">Remarks.</emphasis>
This species shares with
<taxonomicName id="9BA24D15CA5DFFFBD533F9D1FD36F9B0" authorityName="Olah &amp; Flint" authorityYear="2012" box="[597,691,1566,1592]" class="Insecta" family="Hydroptilidae" genus="Acostatrichia" kingdom="Animalia" order="Trichoptera" pageId="24" pageNumber="225" phylum="Arthropoda" rank="species" species="cerna">
<emphasis id="6ED6EA84CA5DFFFBD533F9D1FDE3F9B0" box="[597,614,1566,1592]" italics="true" pageId="24" pageNumber="225">A</emphasis>
.
<emphasis id="6ED6EA84CA5DFFFBD515F9D0FD36F9B0" box="[627,691,1567,1592]" italics="true" pageId="24" pageNumber="225">cerna</emphasis>
</taxonomicName>
similarities in the general aspect of the male genitalia, particularly, the presence of three or four conspicuous apical spines on each dorsolateral process of segment IX (
<figureCitation id="C4992A13CA5DFFFBD384F98DFAD4F9D4" box="[1250,1361,1602,1628]" captionStart="FIGURE 11" captionStartId="19.[151,250,1893,1918]" captionTargetBox="[197,1394,184,1867]" captionTargetId="figure@19.[194,1394,181,1867]" captionTargetPageId="19" captionText="FIGURE 11. Acostatrichia cerna Oláh &amp; Flint 2012, paratype male, genitalia. 11A, ventral; 11B, dorsal; 11C, left lateral; 11D, phallus, dorsal; 11E, phallus, left lateral. Abbreviations: sb. pl. = subgenital plate; inf. app. = inferior appendage (paired); dl. proc. = dorsolateral process of segment IX (paired); lat. proc. = lateral process of inferior appendage (paired). Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3731439" httpUri="https://zenodo.org/record/3731439/files/figure.png" pageId="24" pageNumber="225">Figs. 11A</figureCitation>
,
<figureCitation id="C4992A13CA5DFFFBD23AF98DFA15F9D4" box="[1372,1424,1602,1628]" captionStart="FIGURE 14" captionStartId="25.[151,250,1952,1977]" captionTargetBox="[163,1424,182,1927]" captionTargetId="figure@25.[162,1425,181,1927]" captionTargetPageId="25" captionText="FIGURE 14. Acostatrichia ujasa Oláh &amp; Flint 2012, paratype male, genitalia. 14A, ventral; 14B, dorsal; 14C, left lateral; 14D, phallus, dorsal; 14E, phallus, left lateral. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3731446" httpUri="https://zenodo.org/record/3731446/files/figure.png" pageId="24" pageNumber="225">14A</figureCitation>
). However, these two species can be distinguished from each other by those apical spines, much longer in
<taxonomicName id="9BA24D15CA5DFFFBD25FF9A9FA13F908" authorityName="Olah &amp; Flint" authorityYear="2012" box="[1337,1430,1638,1664]" class="Insecta" family="Hydroptilidae" genus="Acostatrichia" kingdom="Animalia" order="Trichoptera" pageId="24" pageNumber="225" phylum="Arthropoda" rank="species" species="ujasa">
<emphasis id="6ED6EA84CA5DFFFBD25FF9A9FACFF908" box="[1337,1354,1638,1664]" italics="true" pageId="24" pageNumber="225">A</emphasis>
.
<emphasis id="6ED6EA84CA5DFFFBD23FF9A8FA13F908" box="[1369,1430,1639,1664]" italics="true" pageId="24" pageNumber="225">ujasa</emphasis>
</taxonomicName>
, and also by phallus, in
<taxonomicName id="9BA24D15CA5DFFFBD6F1F945FE76F92C" authorityName="Olah &amp; Flint" authorityYear="2012" box="[407,499,1674,1700]" class="Insecta" family="Hydroptilidae" genus="Acostatrichia" kingdom="Animalia" order="Trichoptera" pageId="24" pageNumber="225" phylum="Arthropoda" rank="species" species="ujasa">
<emphasis id="6ED6EA84CA5DFFFBD6F1F945FE2DF92C" box="[407,424,1674,1700]" italics="true" pageId="24" pageNumber="225">A</emphasis>
.
<emphasis id="6ED6EA84CA5DFFFBD6D0F944FE76F92C" box="[438,499,1675,1700]" italics="true" pageId="24" pageNumber="225">ujasa</emphasis>
</taxonomicName>
with more than eight internal spines (
<figureCitation id="C4992A13CA5DFFFBD4FEF945FC7AF92C" box="[920,1023,1674,1700]" captionStart="FIGURE 14" captionStartId="25.[151,250,1952,1977]" captionTargetBox="[163,1424,182,1927]" captionTargetId="figure@25.[162,1425,181,1927]" captionTargetPageId="25" captionText="FIGURE 14. Acostatrichia ujasa Oláh &amp; Flint 2012, paratype male, genitalia. 14A, ventral; 14B, dorsal; 14C, left lateral; 14D, phallus, dorsal; 14E, phallus, left lateral. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3731446" httpUri="https://zenodo.org/record/3731446/files/figure.png" pageId="24" pageNumber="225">Fig. 14D</figureCitation>
), whereas in
<taxonomicName id="9BA24D15CA5DFFFBD3F4F945FB75F92C" authorityName="Olah &amp; Flint" authorityYear="2012" box="[1170,1264,1674,1700]" class="Insecta" family="Hydroptilidae" genus="Acostatrichia" kingdom="Animalia" order="Trichoptera" pageId="24" pageNumber="225" phylum="Arthropoda" rank="species" species="cerna">
<emphasis id="6ED6EA84CA5DFFFBD3F4F945FB26F92C" box="[1170,1187,1674,1700]" italics="true" pageId="24" pageNumber="225">A</emphasis>
.
<emphasis id="6ED6EA84CA5DFFFBD3D6F944FB75F92C" box="[1200,1264,1675,1700]" italics="true" pageId="24" pageNumber="225">cerna</emphasis>
</taxonomicName>
the phallus has four long internal spines (
<figureCitation id="C4992A13CA5DFFFBD6D1F961FD9BF940" box="[439,542,1710,1736]" captionStart="FIGURE 11" captionStartId="19.[151,250,1893,1918]" captionTargetBox="[197,1394,184,1867]" captionTargetId="figure@19.[194,1394,181,1867]" captionTargetPageId="19" captionText="FIGURE 11. Acostatrichia cerna Oláh &amp; Flint 2012, paratype male, genitalia. 11A, ventral; 11B, dorsal; 11C, left lateral; 11D, phallus, dorsal; 11E, phallus, left lateral. Abbreviations: sb. pl. = subgenital plate; inf. app. = inferior appendage (paired); dl. proc. = dorsolateral process of segment IX (paired); lat. proc. = lateral process of inferior appendage (paired). Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3731439" httpUri="https://zenodo.org/record/3731439/files/figure.png" pageId="24" pageNumber="225">Fig. 11D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="14B8651DCA5DFFFBD7A1F91EFD1BF965" box="[199,670,1745,1773]" pageId="24" pageNumber="225" type="distribution">
<paragraph id="5C1D3696CA5DFFFBD7A1F91EFD1BF965" blockId="24.[151,1437,737,1773]" box="[199,670,1745,1773]" pageId="24" pageNumber="225">
<emphasis id="6ED6EA84CA5DFFFBD7A1F91EFEE4F964" bold="true" box="[199,353,1745,1772]" pageId="24" pageNumber="225">Distribution.</emphasis>
<collectingCountry id="24B57606CA5DFFFBD601F91DFE41F964" box="[359,452,1746,1772]" name="Ecuador" pageId="24" pageNumber="225">Ecuador</collectingCountry>
and
<collectingCountry id="24B57606CA5DFFFBD69DF91DFDABF964" box="[507,558,1746,1772]" name="Peru" pageId="24" pageNumber="225">Peru</collectingCountry>
(
<figureCitation id="C4992A13CA5DFFFBD558F91DFD14F965" box="[574,657,1746,1773]" captionStart="FIGURE 16" captionStartId="27.[151,250,1798,1823]" captionTargetBox="[191,1387,622,1773]" captionTargetId="figure@27.[191,1397,622,1773]" captionTargetPageId="27" captionText="FIGURE 16. Distribution map of the Acostatrichia species in South America, based on material examined here and previously published records. When latitude and longitude were not available, an approximate location was used. Circle, species in the Acostatrichia plaumanni Group; square, species in the Acostatrichia brevipenis Group; triangle, species in the Acostatrichia cerna Group." figureDoi="http://doi.org/10.5281/zenodo.3731450" httpUri="https://zenodo.org/record/3731450/files/figure.png" pageId="24" pageNumber="225">Fig. 16</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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