511 lines
108 KiB
XML
511 lines
108 KiB
XML
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<mods:title id="17C75B8345C46CDC0AD8EBB2E7D6AAF6">Revision of the freshwater crabs of the Johora tahanensis (Bott, 1966) species group (Crustacea, Brachyura, Potamidae), with a key to the genus</mods:title>
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<mods:namePart id="43E10C293938BE0B5D44B211ED1DEA25">Ng, Peter K. L.</mods:namePart>
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<mods:affiliation id="63444E7130675475A5E13A1A49C72AFA">Lee Kong Chian Natural History Museum, National University of Singapore, 2 Conservatory Drive, Singapore 117377, Singapore</mods:affiliation>
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<mods:date id="E385DE8003790C707F5A5A50FB1C4966">2020</mods:date>
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<treatment id="CEDC1AC617E45B1D9851FF5A4A16DA00" ID-GBIF-Taxon="169094341" LSID="urn:lsid:plazi:treatment:CEDC1AC617E45B1D9851FF5A4A16DA00" httpUri="http://treatment.plazi.org/id/CEDC1AC617E45B1D9851FF5A4A16DA00" lastPageNumber="1" pageId="0" pageNumber="1" scope_family="Potamidae" scope_infraOrder="Brachyura">
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<taxonomicName id="281D5502C8B60EC4850265E7DF4DF8B1" ID-CoL="3QQB7" LSID="CEDC1AC6-17E4-5B1D-9851-FF5A4A16DA00" authority="Bott, 1966" authorityName="Bott" authorityYear="1966" class="Malacostraca" family="Potamidae" genus="Johora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Johora tahanensis" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="tahanensis">Johora tahanensis Bott, 1966</taxonomicName>
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<figureCitation id="91F3515CB7BD6AB9C6351A4AA7148557" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Overall dorsal view A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng & Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure1" httpUri="https://binary.pensoft.net/fig/476312" pageId="0" pageNumber="1">Figures 1B</figureCitation>
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<figureCitation id="60C6B6CD48054B75AD0154ACE5FF9E6C" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal view of carapace A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249) Terengganu D J. hoiseni Ng & Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure2" httpUri="https://binary.pensoft.net/fig/476313" pageId="0" pageNumber="1">, 2B</figureCitation>
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<figureCitation id="BAFB505C952BDCA7C48E0A2E8EDEE600" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Frontal view of cephalothorax A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng & Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure3" httpUri="https://binary.pensoft.net/fig/476314" pageId="0" pageNumber="1">, 3B</figureCitation>
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<figureCitation id="84662A4F5D8B282EBBA90B772EC2B376" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Left third maxilliped A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng & Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure4" httpUri="https://binary.pensoft.net/fig/476315" pageId="0" pageNumber="1">, 4B</figureCitation>
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<figureCitation id="119E19CD38DA904559D4141CCB2F3ECE" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Anterior thoracic sternum and pleon A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng & Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure5" httpUri="https://binary.pensoft.net/fig/476316" pageId="0" pageNumber="1">, 5B</figureCitation>
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<figureCitation id="ADB3ECB0CA9450D8C7312D3B2B10ED53" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. A-D Johora tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang E-H J. hoiseni Ng & Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan A, E left G 1 (ventral view) B, F distal part of left G 1 (ventral view) C, G distal part of left G 1 (dorsal view) D, H left G 2 (ventral view) E-H after Ng and Takeda (1992: fig. 3 B, D-F). Scale bars: 1.0 mm (A, D, E, H); 0.5 mm (B, C, F, G)." figureDoi="10.3897/zookeys.994.56810.figure8" httpUri="https://binary.pensoft.net/fig/476319" pageId="0" pageNumber="1">, 8A-D</figureCitation>
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<figureCitation id="A14E2AE4837B2BE298BC745969FA1E05" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Overall dorsal view A Johora intermedia Ng, 1986, female (17.9 x 14.7 mm) (ZRC 2001.1002), Selangor B J. tahanensis (Bott, 1966), female (24.4 x 21.0 mm) (ZRC 1984.6765), Pahang C J. thoi Ng, 1990, female (31.7 x 25.5 mm) (ZRC 2001.1167), Terengganu D J. hoiseni Ng & Takeda, 1992, paratype female (22.1 x 19.0 mm) (ZRC 1984.6675), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype female (21.0 x 17.4 mm) (ZRC 2006.0053), Thailand F J. booliati sp. nov., paratype female (40.4 x 34.4 mm) (ZRC 1995.270), Pahang G J. erici sp. nov., paratype female (32.8 x 26.9 mm) (ZRC 1995.268), Perak H J. michaeli sp. nov., paratype female (29.9 x 25.0 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure12" httpUri="https://binary.pensoft.net/fig/476323" pageId="0" pageNumber="1">, 12B</figureCitation>
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<figureCitation id="7945E8C2A19EFAAB04610B0AB406C97F" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Female pleon A Johora intermedia Ng, 1986, female (17.9 x 14.7 mm) (ZRC 2001.1002), Selangor B J. tahanensis (Bott, 1966), female (24.4 x 21.0 mm) (ZRC 1984.6765), Pahang C J. thoi Ng, 1990, female (31.7 x 25.5 mm) (ZRC 2001.1167), Terengganu D J. hoiseni Ng & Takeda, 1992, paratype female (22.1 x 19.0 mm) (ZRC 1984.6675), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype female (21.0 x 17.4 mm) (ZRC 2006.0053), Thailand F J. booliati sp. nov., paratype female (40.4 x 34.4 mm) (ZRC 1995.270), Pahang G J. erici sp. nov., paratype female (32.8 x 26.9 mm) (ZRC 1995.268), Perak H J. michaeli sp. nov., paratype female (29.9 x 25.0 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure13" httpUri="https://binary.pensoft.net/fig/476324" pageId="0" pageNumber="1">, 13B</figureCitation>
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<figureCitation id="4FCCE61CC5B293D0B9D1343CE396239E" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Sternopleonal cavity and vulvae A Johora intermedia Ng, 1986, female (17.9 x 14.7 mm) (ZRC 2001.1002), Selangor B J. tahanensis (Bott, 1966), female (24.4 x 21.0 mm) (ZRC 1984.6765), Pahang C J. thoi Ng, 1990, female (31.7 x 25.5 mm) (ZRC 2001.1167), Terengganu D J. hoiseni Ng & Takeda, 1992, paratype female (22.1 x 19.0 mm) (ZRC 1984.6675), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype female (21.0 x 17.4 mm) (ZRC 2006.0053), Thailand F J. booliati sp. nov., paratype female (40.4 x 34.4 mm) (ZRC 1995.270), Pahang G J. erici sp. nov., paratype female (32.8 x 26.9 mm) (ZRC 1995.268), Perak H J. michaeli sp. nov., paratype female (29.9 x 25.0 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure14" httpUri="https://binary.pensoft.net/fig/476325" pageId="0" pageNumber="1">, 14B</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="48CF39ECC3D6B46FDE28AC39BDBE4F2A" pageId="0" pageNumber="1" type="reference_group">
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<paragraph id="472216D74CF660E691C618A3B5321E40" pageId="0" pageNumber="1">
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<taxonomicName id="1D87C8F2AD6C7FF538F106156EE2CC45" authorityName="Bott" authorityYear="1966" class="Malacostraca" family="Potamidae" genus="Potamiscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Potamiscus (Johora) johorensis subsp. tahanensis" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="subSpecies" species="johorensis" subGenus="Johora" subSpecies="tahanensis">Potamiscus (Johora) johorensis tahanensis</taxonomicName>
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Bott, 1966: 495 (part), pl. 21 fig. 15.
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</paragraph>
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<paragraph id="5386949383DBB0BC19C61AD4DAA68E7D" pageId="0" pageNumber="1">
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<taxonomicName id="3AC58A126C7C05D9E35EFAFA04EDC3C5" class="Malacostraca" family="Potamidae" genus="Stoliczia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stoliczia (Johora) johorensis subsp. tahanensis" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="subSpecies" species="johorensis" subGenus="Johora" subSpecies="tahanensis">Stoliczia (Johora) johorensis tahanensis</taxonomicName>
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-
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<bibRefCitation id="553CF61276471B92610FADF9921DE9E3" author="Bott, R" journalOrPublisher="Abhandlungen der senckenbergischen naturforschenden Gesellschaft" pageId="0" pageNumber="1" pagination="1 - 338" refId="B2" refString="Bott, R, 1970. Die Suesswasserkrabben von Europa, Asien, Australien und ihre Stammesgeschichte. Eine Revision der Potamoidea und der Parathelphusoidea (Crustacea, Decapoda). Abhandlungen der senckenbergischen naturforschenden Gesellschaft 526: 1 - 338" title="Die Suesswasserkrabben von Europa, Asien, Australien und ihre Stammesgeschichte. Eine Revision der Potamoidea und der Parathelphusoidea (Crustacea, Decapoda)." volume="526" year="1970">Bott 1970</bibRefCitation>
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: 181 (part), pl. 50 fig. 50.
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</paragraph>
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<paragraph id="198FB84E8A466130401BCB50170E72F8" pageId="0" pageNumber="1">
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<taxonomicName id="D49BCA72B8D38803618B080D8789AA14" authorityName="Bott" authorityYear="1966" class="Malacostraca" family="Potamidae" genus="Johora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Johora tahanensis" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="tahanensis">Johora tahanensis</taxonomicName>
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-
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<bibRefCitation id="5B24B5B1B1A97A24D9701130B41E6497" author="Ng, PKL" journalOrPublisher="Malayan Nature Journal" pageId="0" pageNumber="1" pagination="13 - 44" refId="B11" refString="Ng, PKL, 1987. A revision of the Malayan freshwater crabs of the genus Johora Bott, 1966 stat. nov. (Decapoda: Brachyura: Potamidae). Malayan Nature Journal 41: 13 - 44" title="A revision of the Malayan freshwater crabs of the genus Johora Bott, 1966 stat. nov. (Decapoda: Brachyura: Potamidae)." volume="41" year="1987">Ng 1987</bibRefCitation>
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: 33 (part), fig. 9A;
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<bibRefCitation id="FF4B6B10CFC8F1C606D5FC1E9B44AACE" author="Ng, PKL" journalOrPublisher="Department of Zoology, National University of Singapore, Shinglee Press" pageId="0" pageNumber="1" refId="B12" refString="Ng, PKL, 1988. The Freshwater Crabs of Peninsular Malaysia and Singapore. Department of Zoology, National University of Singapore, Shinglee Press" title="The Freshwater Crabs of Peninsular Malaysia and Singapore." year="1988">Ng 1988</bibRefCitation>
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: 42 (part), fig. C, F;
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<bibRefCitation id="A6A44DA9F1A64A8C6991D9DDAC35C0B2" author="Ng, PKL" journalOrPublisher="Bulletin of the National Science Museum, Tokyo, Series A" pageId="0" pageNumber="1" pagination="103 - 116" refId="B19" refString="Ng, PKL, Takeda, M, 1992. On some freshwater crabs (Crustacea, Brachyura, Potamidae, Parathelphusidae and Grapsidae) from Peninsular Malaysia. Bulletin of the National Science Museum, Tokyo, Series A 18 (3): 103 - 116" title="On some freshwater crabs (Crustacea, Brachyura, Potamidae, Parathelphusidae and Grapsidae) from Peninsular Malaysia." volume="18" year="1992">Ng and Takeda 1992</bibRefCitation>
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: 107 (part);
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<bibRefCitation id="E032DF8418BB053C6701BD9FB36FF046" author="Ng, PKL" editor="Yule, C" journalOrPublisher="Malaysian Academy of Sciences, city" pageId="0" pageNumber="1" pagination="311 - 336" refId="B14" refString="Ng, PKL, 2004. Crustacea: Decapoda, Brachyura. In: Yule, C, Yong, HS, Eds., Freshwater Invertebrates of the Malaysian Region. Malaysian Academy of Sciences, city: 311 - 336" title="Crustacea: Decapoda, Brachyura." volumeTitle="Freshwater Invertebrates of the Malaysian Region." year="2004">Ng 2004</bibRefCitation>
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||
: 321;
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<bibRefCitation id="5C21EAC0087C0210C947678BB371D44E" author="Ng, PKL" editor="Chua, L" journalOrPublisher="28 - 30 June 2005, Forest Research Institute Malaysia, Kepong" pageId="0" pageNumber="1" pagination="95 - 120" refId="B21" refString="Ng, PKL, Yeo, DCJ, 2007. Malaysian freshwater crabs: conservation prospects and challenges. In: Chua, L, Ed., Proceedings of the Seminar on the Status of Biological Diversity in Malaysia & Threat Assessment of Plant Species in Malaysia. 28-30 June 2005, Forest Research Institute Malaysia, Kepong: 95 - 120" title="Malaysian freshwater crabs: conservation prospects and challenges." volumeTitle="Proceedings of the Seminar on the Status of Biological Diversity in Malaysia & Threat Assessment of Plant Species in Malaysia." year="2007">Ng and Yeo 2007</bibRefCitation>
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: 102;
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<bibRefCitation id="3E849FC0629307387B26C938EB983C32" author="Ng, PKL" journalOrPublisher="Raffles Bulletin of Zoology, Supplement" pageId="0" pageNumber="1" pagination="1 - 286" refId="B18" refString="Ng, PKL, Guinot, D, Davie, PJF, 2008. Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, Supplement 17: 1 - 286" title="Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world." volume="17" year="2008">Ng et al. 2008</bibRefCitation>
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: 163;
|
||
<bibRefCitation id="77B2799FAE6AD3A5F021E98E3C8B0E33" DOI="https://doi.org/10.1016/j.biocon.2009.02.038" author="Cumberlidge, N" journalOrPublisher="Biological Conservation" pageId="0" pageNumber="1" pagination="1665 - 1673" refId="B3" refString="Cumberlidge, N, Ng, PKL, Yeo, DCJ, Magalhaes, C, Campos, MR, Alvarez, F, Naruse, T, Daniels, SR, Esser, LJ, Attipoe, FYK, Clotilde-Ba, F-L, Darwall, W, Mcivor, A, Collen, B, Ram, M, 2009. Freshwater crabs and the biodiversity crisis: importance, threats, status, and conservation challenges. Biological Conservation 142: 1665 - 1673, DOI: https://doi.org/10.1016/j.biocon.2009.02.038" title="Freshwater crabs and the biodiversity crisis: importance, threats, status, and conservation challenges." url="https://doi.org/10.1016/j.biocon.2009.02.038" volume="142" year="2009">Cumberlidge et al. 2009</bibRefCitation>
|
||
: table.
|
||
</paragraph>
|
||
<paragraph id="05E3FB2EF74B28A35E39EB5BC01A7878" pageId="0" pageNumber="1">
|
||
<taxonomicName id="0117AAE40BAE42F97FE9522E7DC373E7" class="Malacostraca" family="Potamidae" genus="Stoliczia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stoliczia johorensis subsp. tahanensis" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="subSpecies" species="johorensis" subSpecies="tahanensis">Stoliczia johorensis tahanensis</taxonomicName>
|
||
-
|
||
<bibRefCitation id="9DEA309F8DD1C7B470328E45C3B9C81D" author="Takeda, M" editor="Kawashima, K" journalOrPublisher="Paragonimus Research Report, Kyushu University" pageId="0" pageNumber="1" pagination="89 - 106" refId="B25" refString="Takeda, M, 1987. Freshwater crabs from Peninsular Malaysia, with notes on some species from the Philippines. In: Kawashima, K, Ed., Paragonimus in Asia: Biology, Genetic Variation and Speciation. Paragonimus Research Report, Kyushu University 1: 89 - 106" title="Freshwater crabs from Peninsular Malaysia, with notes on some species from the Philippines." volume="1" volumeTitle="Paragonimus in Asia: Biology, Genetic Variation and Speciation." year="1987">Takeda 1987</bibRefCitation>
|
||
: 92, pl. X (center).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="8FF6E51647B79B3ED2727141737F889B" pageId="0" pageNumber="1" type="materials_examined">
|
||
<paragraph id="0383F8E6D4117F9140AE9CAEA0CA7567" pageId="0" pageNumber="1">Material examined.</paragraph>
|
||
<paragraph id="66FD384A0E5C92904A7CF1FF584023CB" pageId="0" pageNumber="1">
|
||
13 males (largest 27.2
|
||
<normalizedToken id="87AAD4C6792A748782FDA4422F91880A" originalValue="×">x</normalizedToken>
|
||
22.9 mm), 13 females (largest 24.4
|
||
<normalizedToken id="264C97F3DF19DA14A1C6655A789FB261" originalValue="×">x</normalizedToken>
|
||
21.0 mm), 4 juveniles (ZRC 1984.6764-6793), Kuala Tahan, Taman Negara National Park, Pahang, coll. Tweedie MWF, April 1940; 1 male (30.7
|
||
<normalizedToken id="9D7EB5C626083EC0D38C66EF471E615D" originalValue="×">x</normalizedToken>
|
||
25.5 mm), 2 females (ZRC 1984.6795-6797), rivulet of Sungei Tahan, Kuala Tahan, Taman Negara National Park, Pahang, coll. Alfred ER, 23 March 1956; 1 male (ZRC 1989.2086), Changah Siveh, Sungei Tahan, Taman Negara National Park, Pahang, coll. Alfred ER, 3 March 1948; 1 female (ZRC 1989.3749), Jenka, Sungei Tekam, Pahang,
|
||
<geoCoordinate id="1DCE328719686A28C889809E49318947" degrees="4" direction="north" minutes="13" orientation="latitude" precision="925" value="4.2166667">4°13'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="77F61168D324C37A181E0810127D1127" degrees="102" direction="east" minutes="39" orientation="longitude" precision="925" value="102.65">102°39'E</geoCoordinate>
|
||
, coll. Lim RP, 19 March 1981; 1 young male, 1 female (ZRC 1989.2144-2145), from University of Malaya collections, 1960s; 2 juveniles (ZRC 1989.3358-3359), station F63/20, tributary of Sungei Telom (= Sungei Telum), Pahang, coll. University of Malaya, 9 March 1963; 1 female (ZRC 1989.3686), coll. University of Malaya, no date. All locations in Peninsular Malaysia.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A2FFB95F59E8E78CDF59221752CDA2AC" pageId="0" pageNumber="1" type="diagnosis">
|
||
<paragraph id="B43148FC62E6FAF4CAB41F517D357F82" pageId="0" pageNumber="1">Diagnosis.</paragraph>
|
||
<paragraph id="DCB654791FA0D6DC4A264762241440D6" pageId="0" pageNumber="1">
|
||
Adult carapace width to length ratio 1.16-1.19 (Figs
|
||
<figureCitation id="0BA859891321CD15D026B4579A1A05D8" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Overall dorsal view A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng & Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure1" httpUri="https://binary.pensoft.net/fig/476312" pageId="0" pageNumber="1">1B</figureCitation>
|
||
,
|
||
<figureCitation id="FA2471E4C8B5C883A8574AE99C7E2D44" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal view of carapace A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249) Terengganu D J. hoiseni Ng & Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure2" httpUri="https://binary.pensoft.net/fig/476313" pageId="0" pageNumber="1">2B</figureCitation>
|
||
,
|
||
<figureCitation id="6974716F71D0F382ADCF22193694CBBA" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Overall dorsal view A Johora intermedia Ng, 1986, female (17.9 x 14.7 mm) (ZRC 2001.1002), Selangor B J. tahanensis (Bott, 1966), female (24.4 x 21.0 mm) (ZRC 1984.6765), Pahang C J. thoi Ng, 1990, female (31.7 x 25.5 mm) (ZRC 2001.1167), Terengganu D J. hoiseni Ng & Takeda, 1992, paratype female (22.1 x 19.0 mm) (ZRC 1984.6675), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype female (21.0 x 17.4 mm) (ZRC 2006.0053), Thailand F J. booliati sp. nov., paratype female (40.4 x 34.4 mm) (ZRC 1995.270), Pahang G J. erici sp. nov., paratype female (32.8 x 26.9 mm) (ZRC 1995.268), Perak H J. michaeli sp. nov., paratype female (29.9 x 25.0 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure12" httpUri="https://binary.pensoft.net/fig/476323" pageId="0" pageNumber="1">12B</figureCitation>
|
||
); dorsal surface gently convex in frontal view, not inflated (Fig.
|
||
<figureCitation id="5024020CA92ECF36929864A8B17A6D1B" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Frontal view of cephalothorax A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng & Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure3" httpUri="https://binary.pensoft.net/fig/476314" pageId="0" pageNumber="1">3B</figureCitation>
|
||
); frontal margin almost straight (Fig.
|
||
<figureCitation id="EAFAA77899BD588BC3118BF3ED0887C4" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal view of carapace A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249) Terengganu D J. hoiseni Ng & Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure2" httpUri="https://binary.pensoft.net/fig/476313" pageId="0" pageNumber="1">2B</figureCitation>
|
||
); suborbital, pterygostomial and sub-branchial regions rugose, pterygostomial region covered with dense setae (Fig.
|
||
<figureCitation id="8C10B1F94F0EEDD9E8FD740A3A6D42DA" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Frontal view of cephalothorax A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng & Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure3" httpUri="https://binary.pensoft.net/fig/476314" pageId="0" pageNumber="1">3B</figureCitation>
|
||
); epigastric cristae distinct, distinctly anterior to sharp postorbital cristae, postorbital cristae with lateral edges low, joining lateral margin through oblique striae (Fig.
|
||
<figureCitation id="4F858AC32464468AF361310EB42472F7" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal view of carapace A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249) Terengganu D J. hoiseni Ng & Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure2" httpUri="https://binary.pensoft.net/fig/476313" pageId="0" pageNumber="1">2B</figureCitation>
|
||
); external orbital tooth separated from epibranchial tooth by distinct cleft, epibranchial tooth sharp, distinct (Fig.
|
||
<figureCitation id="927B71471EA7F41E518BBE331A22FAEA" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal view of carapace A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249) Terengganu D J. hoiseni Ng & Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure2" httpUri="https://binary.pensoft.net/fig/476313" pageId="0" pageNumber="1">2B</figureCitation>
|
||
); anterolateral margin distinctly convex (Fig.
|
||
<figureCitation id="266DB4E57CBE9275C0D796EA6C72105E" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal view of carapace A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249) Terengganu D J. hoiseni Ng & Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure2" httpUri="https://binary.pensoft.net/fig/476313" pageId="0" pageNumber="1">2B</figureCitation>
|
||
); posterolateral margin gently sinuous to almost straight, distinctly converging towards gently convex, entire posterior carapace margin (Fig.
|
||
<figureCitation id="BFFEB2ABF1270347A6120B0DD9A73E93" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal view of carapace A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249) Terengganu D J. hoiseni Ng & Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure2" httpUri="https://binary.pensoft.net/fig/476313" pageId="0" pageNumber="1">2B</figureCitation>
|
||
); posterior margin of epistome with triangular median triangle, lateral margin obliquely sloping (Fig.
|
||
<figureCitation id="0DD16312C6C9CF146C93A72719C7638F" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Frontal view of cephalothorax A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng & Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure3" httpUri="https://binary.pensoft.net/fig/476314" pageId="0" pageNumber="1">3B</figureCitation>
|
||
); outer surfaces of third maxillipeds with dense, long stiff setae; ischium subrectangular, with shallow median oblique groove (Figs
|
||
<figureCitation id="391A0FF8AA550C13CE5E5A13F9A14E54" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Frontal view of cephalothorax A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng & Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure3" httpUri="https://binary.pensoft.net/fig/476314" pageId="0" pageNumber="1">3B</figureCitation>
|
||
,
|
||
<figureCitation id="4EE0DF3BF0BDF9DA39F7199BADB5171C" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Left third maxilliped A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng & Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure4" httpUri="https://binary.pensoft.net/fig/476315" pageId="0" pageNumber="1">4B</figureCitation>
|
||
); ambulatory legs not elongate, length to width ratio of merus of fourth ambulatory leg 2.7-2.8 (Figs
|
||
<figureCitation id="36313AAAFA84D6C6C3B5A7E27F14C4A1" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Overall dorsal view A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng & Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure1" httpUri="https://binary.pensoft.net/fig/476312" pageId="0" pageNumber="1">1B</figureCitation>
|
||
,
|
||
<figureCitation id="64BAC244C2FC526431B2C3A187EB53E2" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Overall dorsal view A Johora intermedia Ng, 1986, female (17.9 x 14.7 mm) (ZRC 2001.1002), Selangor B J. tahanensis (Bott, 1966), female (24.4 x 21.0 mm) (ZRC 1984.6765), Pahang C J. thoi Ng, 1990, female (31.7 x 25.5 mm) (ZRC 2001.1167), Terengganu D J. hoiseni Ng & Takeda, 1992, paratype female (22.1 x 19.0 mm) (ZRC 1984.6675), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype female (21.0 x 17.4 mm) (ZRC 2006.0053), Thailand F J. booliati sp. nov., paratype female (40.4 x 34.4 mm) (ZRC 1995.270), Pahang G J. erici sp. nov., paratype female (32.8 x 26.9 mm) (ZRC 1995.268), Perak H J. michaeli sp. nov., paratype female (29.9 x 25.0 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure12" httpUri="https://binary.pensoft.net/fig/476323" pageId="0" pageNumber="1">12B</figureCitation>
|
||
); G1 subterminal segment gradually tapering from broad proximal part to slender distal part, without distinct shelf-like structure along gently concave outer margin; terminal segment gently curved outwards (from median part of sternum), ca. half length of subterminal segment, surface with numerous short setae (Fig.
|
||
<figureCitation id="788961F0C1B8665BBAB37D97809CC204" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. A-D Johora tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang E-H J. hoiseni Ng & Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan A, E left G 1 (ventral view) B, F distal part of left G 1 (ventral view) C, G distal part of left G 1 (dorsal view) D, H left G 2 (ventral view) E-H after Ng and Takeda (1992: fig. 3 B, D-F). Scale bars: 1.0 mm (A, D, E, H); 0.5 mm (B, C, F, G)." figureDoi="10.3897/zookeys.994.56810.figure8" httpUri="https://binary.pensoft.net/fig/476319" pageId="0" pageNumber="1">8A-C</figureCitation>
|
||
); G2 slightly longer than G1, distal segment long, about half length of basal segment (Fig.
|
||
<figureCitation id="3AE2582A03B24D9B01A1A5B5118D4A35" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. A-D Johora tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang E-H J. hoiseni Ng & Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan A, E left G 1 (ventral view) B, F distal part of left G 1 (ventral view) C, G distal part of left G 1 (dorsal view) D, H left G 2 (ventral view) E-H after Ng and Takeda (1992: fig. 3 B, D-F). Scale bars: 1.0 mm (A, D, E, H); 0.5 mm (B, C, F, G)." figureDoi="10.3897/zookeys.994.56810.figure8" httpUri="https://binary.pensoft.net/fig/476319" pageId="0" pageNumber="1">8D</figureCitation>
|
||
). Female pleon longitudinally ovate; somites 3-6 progressively narrower; telson subtriangular (Fig.
|
||
<figureCitation id="55717742C903C7D6B06F42E0B920A3A4" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Female pleon A Johora intermedia Ng, 1986, female (17.9 x 14.7 mm) (ZRC 2001.1002), Selangor B J. tahanensis (Bott, 1966), female (24.4 x 21.0 mm) (ZRC 1984.6765), Pahang C J. thoi Ng, 1990, female (31.7 x 25.5 mm) (ZRC 2001.1167), Terengganu D J. hoiseni Ng & Takeda, 1992, paratype female (22.1 x 19.0 mm) (ZRC 1984.6675), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype female (21.0 x 17.4 mm) (ZRC 2006.0053), Thailand F J. booliati sp. nov., paratype female (40.4 x 34.4 mm) (ZRC 1995.270), Pahang G J. erici sp. nov., paratype female (32.8 x 26.9 mm) (ZRC 1995.268), Perak H J. michaeli sp. nov., paratype female (29.9 x 25.0 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure13" httpUri="https://binary.pensoft.net/fig/476324" pageId="0" pageNumber="1">13B</figureCitation>
|
||
). Vulvae large, on anterior half of sternite 6, slightly pushing into suture with sternite 5, lateral sternal vulvar cover semicircular (Fig.
|
||
<figureCitation id="3048AF3E0325D57AD684D04E1878AE02" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Sternopleonal cavity and vulvae A Johora intermedia Ng, 1986, female (17.9 x 14.7 mm) (ZRC 2001.1002), Selangor B J. tahanensis (Bott, 1966), female (24.4 x 21.0 mm) (ZRC 1984.6765), Pahang C J. thoi Ng, 1990, female (31.7 x 25.5 mm) (ZRC 2001.1167), Terengganu D J. hoiseni Ng & Takeda, 1992, paratype female (22.1 x 19.0 mm) (ZRC 1984.6675), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype female (21.0 x 17.4 mm) (ZRC 2006.0053), Thailand F J. booliati sp. nov., paratype female (40.4 x 34.4 mm) (ZRC 1995.270), Pahang G J. erici sp. nov., paratype female (32.8 x 26.9 mm) (ZRC 1995.268), Perak H J. michaeli sp. nov., paratype female (29.9 x 25.0 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure14" httpUri="https://binary.pensoft.net/fig/476325" pageId="0" pageNumber="1">14B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="2BE61B5925C146794C897903CF85A6BF" doi="10.3897/zookeys.994.56810.figure3" httpUri="https://binary.pensoft.net/fig/476314" pageId="0" pageNumber="1" start="Figure 3" startId="F3">
|
||
<paragraph id="046EFA73C7109EBFD8028BE264F2A528" pageId="0" pageNumber="1">
|
||
<emphasis id="1D83364AE4E7FFBDA551DBD8EE680F0C" bold="true" pageId="0" pageNumber="1">Figure 3.</emphasis>
|
||
Frontal view of cephalothorax
|
||
<emphasis id="4FEB361740D4DB98D3A6D8E998B6F452" bold="true" pageId="0" pageNumber="1">A</emphasis>
|
||
<taxonomicName id="1883A518DEFE7A1524CAAD42FE5E49D1" baseAuthorityName="Ng" baseAuthorityYear="1986" class="Malacostraca" family="Potamidae" genus="Johora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Johora intermedia" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="intermedia">
|
||
<emphasis id="748C16A637E46A54723A957B2EE7D24E" italics="true" pageId="0" pageNumber="1">Johora intermedia</emphasis>
|
||
</taxonomicName>
|
||
Ng, 1986, holotype male (19.8
|
||
<normalizedToken id="0934CDEE1903F75EAF4340BAD6D7F082" originalValue="×">x</normalizedToken>
|
||
15.1 mm) (ZRC 1984.6529), Pahang
|
||
<emphasis id="AEBA5907E890ED342F5718E1634C5F49" bold="true" pageId="0" pageNumber="1">B</emphasis>
|
||
<taxonomicName id="760A9868A516AA99140E2DDBC1051E39" lsidName="J. tahanensis" pageId="0" pageNumber="1" rank="species" species="tahanensis">
|
||
<emphasis id="9BABEA11323EE6F1D2BAB8F3197A7AAA" italics="true" pageId="0" pageNumber="1">J. tahanensis</emphasis>
|
||
</taxonomicName>
|
||
(Bott, 1966), male (30.7
|
||
<normalizedToken id="7730D5A6942F6DD6E503CEEB5CABF738" originalValue="×">x</normalizedToken>
|
||
25.5 mm) (ZRC 1984.6795), Pahang
|
||
<emphasis id="08E40FFC2CDAA976BAC3C12B83466DCD" bold="true" pageId="0" pageNumber="1">C</emphasis>
|
||
<taxonomicName id="A2BD8A8DBF5222FF6BF091E52B585256" lsidName="J. thoi" pageId="0" pageNumber="1" rank="species" species="thoi">
|
||
<emphasis id="2F4851653C34594F5B981996BE780C68" italics="true" pageId="0" pageNumber="1">J. thoi</emphasis>
|
||
</taxonomicName>
|
||
Ng, 1990, holotype male (41.2
|
||
<normalizedToken id="89CEB5FE9E90544369CC6FB21C567EEA" originalValue="×">x</normalizedToken>
|
||
33.3 mm) (ZRC 1989.2249), Terengganu
|
||
<emphasis id="6AE2D384516FB5E2D9DBCDD2DFB9FF96" bold="true" pageId="0" pageNumber="1">D</emphasis>
|
||
<taxonomicName id="E4C94A8929DCB1384E5B08301D1AC1FF" lsidName="J. hoiseni" pageId="0" pageNumber="1" rank="species" species="hoiseni">
|
||
<emphasis id="D743AA411699B676606DEDD435D081CE" italics="true" pageId="0" pageNumber="1">J. hoiseni</emphasis>
|
||
</taxonomicName>
|
||
Ng & Takeda, 1992, holotype male (25.1
|
||
<normalizedToken id="B888032A9127AB75BE7DE2A892865449" originalValue="×">x</normalizedToken>
|
||
21.1 mm) (ZRC 1984.6673), Kelantan
|
||
<emphasis id="3C534CDFD7DB04B2561B9323CE1C6E54" bold="true" pageId="0" pageNumber="1">E</emphasis>
|
||
<taxonomicName id="95BD62EAE3EB248D5FEF04C35959070B" lsidName="J. thaiana" pageId="0" pageNumber="1" rank="species" species="thaiana">
|
||
<emphasis id="F06F41D8E9CCD8B0160D06E8024ECDC5" italics="true" pageId="0" pageNumber="1">J. thaiana</emphasis>
|
||
</taxonomicName>
|
||
Leelawathanagoon, Lheknim & Ng, 2005, paratype male (22.2
|
||
<normalizedToken id="6B76FA92AC3A4ED5E8D5E0DF16B003C5" originalValue="×">x</normalizedToken>
|
||
18.6 mm) (ZRC 2006.0052), Thailand
|
||
<emphasis id="B1C0226CD7951D1C25722348E81F0164" bold="true" pageId="0" pageNumber="1">F</emphasis>
|
||
<taxonomicName id="5E5C9F16E8F359EF930F450C82B887C9" lsidName="J. booliati" pageId="0" pageNumber="1" rank="species" species="booliati">
|
||
<emphasis id="3618D31B54B966DC3B2FFD440F76BFAA" italics="true" pageId="0" pageNumber="1">J. booliati</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype male (34.6
|
||
<normalizedToken id="DC3639950ACF35F8F69ACCC69B6133B5" originalValue="×">x</normalizedToken>
|
||
30.0 mm) (ZRC 2020.0072), Pahang
|
||
<emphasis id="14D4C6E7443842813EC83C278C816BF2" bold="true" pageId="0" pageNumber="1">G</emphasis>
|
||
<taxonomicName id="26464837276EF390DF1D194CD5783E84" lsidName="J. erici" pageId="0" pageNumber="1" rank="species" species="erici">
|
||
<emphasis id="E5136F02AD0E88C2585F3E99751C6EBA" italics="true" pageId="0" pageNumber="1">J. erici</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype male (38.3
|
||
<normalizedToken id="BC0A2B2F1B9FA2570C41E9119ED838B6" originalValue="×">x</normalizedToken>
|
||
32.0 mm) (ZRC 2020.0360), Perak
|
||
<emphasis id="8AD09E5EDDE65364ACEF32D94519A1FB" bold="true" pageId="0" pageNumber="1">H</emphasis>
|
||
<taxonomicName id="AFA0B31435BD7F25ED3A9729A5E7990A" lsidName="J. michaeli" pageId="0" pageNumber="1" rank="species" species="michaeli">
|
||
<emphasis id="844EF10AEAF3C2755044F61D9D22CED1" italics="true" pageId="0" pageNumber="1">J. michaeli</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype male (22.7
|
||
<normalizedToken id="1C0DFEF8A60EFDF411A95E53F12DF337" originalValue="×">x</normalizedToken>
|
||
19.2 mm) (ZRC 2010.0047), Terengganu.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="661F67CA8F0ECF3B8A14727587690AF0" pageId="0" pageNumber="1" type="description">
|
||
<paragraph id="4C08061B69C76F2D5A8FB007CB01D284" pageId="0" pageNumber="1">Description of male.</paragraph>
|
||
<paragraph id="0C3AE466DCE374690D1809913502997E" pageId="0" pageNumber="1">
|
||
Carapace subrectangular broader than long, width to length ratio 1.16-1.19; dorsal surface gently convex in frontal view (Figs
|
||
<figureCitation id="9933C468ABDF379CC92790C6DD417127" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Overall dorsal view A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng & Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure1" httpUri="https://binary.pensoft.net/fig/476312" pageId="0" pageNumber="1">1B</figureCitation>
|
||
,
|
||
<figureCitation id="E46BA87E6E25F55FB98AB02AAB1D208A" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal view of carapace A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249) Terengganu D J. hoiseni Ng & Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure2" httpUri="https://binary.pensoft.net/fig/476313" pageId="0" pageNumber="1">2B</figureCitation>
|
||
,
|
||
<figureCitation id="7D39DEB44A21B0748F012C094862FAED" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Frontal view of cephalothorax A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng & Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure3" httpUri="https://binary.pensoft.net/fig/476314" pageId="0" pageNumber="1">3B</figureCitation>
|
||
). Frontal margin almost straight; frontal region, dorsal surface, lateral parts of anterolateral and branchial regions rugose, covered with small granules and striae; regions clearly indicated, median H-shaped gastro-cardiac groove deep; cervical grooves broad, shallow; suborbital, pterygostomial and sub-branchial regions rugose, covered with dense setae (Figs
|
||
<figureCitation id="FA2618739E9A1A1F6D2D69582FEF68B8" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal view of carapace A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249) Terengganu D J. hoiseni Ng & Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure2" httpUri="https://binary.pensoft.net/fig/476313" pageId="0" pageNumber="1">2B</figureCitation>
|
||
,
|
||
<figureCitation id="56D2A568301FC3BC95B38A50C08253DE" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Frontal view of cephalothorax A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng & Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure3" httpUri="https://binary.pensoft.net/fig/476314" pageId="0" pageNumber="1">3B</figureCitation>
|
||
). Epigastric cristae distinct, marked by transverse striae, not cristate, separated by median groove; postorbital cristae sharp, prominent, positioned distinctly posterior to and separated from epigastric cristae, lateral edges reaching beginning of cervical groove, reaching lateral margin through series of short, oblique striae (Figs
|
||
<figureCitation id="BDFF4CB29F847AB706B358D21F65EB32" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Overall dorsal view A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng & Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure1" httpUri="https://binary.pensoft.net/fig/476312" pageId="0" pageNumber="1">1B</figureCitation>
|
||
,
|
||
<figureCitation id="EC65EB76C5C9D6890ADB26E7C1CD7A4A" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal view of carapace A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249) Terengganu D J. hoiseni Ng & Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure2" httpUri="https://binary.pensoft.net/fig/476313" pageId="0" pageNumber="1">2B</figureCitation>
|
||
). Frontal margin entire; separated from supraorbital margin by rounded angle (Fig.
|
||
<figureCitation id="7654F45010ECAE536AC9C5F6A72E65CB" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal view of carapace A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249) Terengganu D J. hoiseni Ng & Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure2" httpUri="https://binary.pensoft.net/fig/476313" pageId="0" pageNumber="1">2B</figureCitation>
|
||
). External orbital tooth triangular, outer margin twice length of inner margin; epibranchial tooth sharp, distinct, separated from anterolateral margin by distinct cleft (Figs
|
||
<figureCitation id="BF5E275B24706246887C756FA9DCB645" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Overall dorsal view A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng & Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure1" httpUri="https://binary.pensoft.net/fig/476312" pageId="0" pageNumber="1">1B</figureCitation>
|
||
,
|
||
<figureCitation id="60805EB0CFA9A2F12F013216CEAFFE07" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal view of carapace A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249) Terengganu D J. hoiseni Ng & Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure2" httpUri="https://binary.pensoft.net/fig/476313" pageId="0" pageNumber="1">2B</figureCitation>
|
||
). Anterolateral margins convex, cristate, granulated (Fig.
|
||
<figureCitation id="0E8817A4E30BA71949AD1F2B278AD07B" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal view of carapace A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249) Terengganu D J. hoiseni Ng & Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure2" httpUri="https://binary.pensoft.net/fig/476313" pageId="0" pageNumber="1">2B</figureCitation>
|
||
). Posterolateral margin gently sinuous to almost straight, distinctly converging towards gently convex, entire posterior carapace margin (Figs
|
||
<figureCitation id="CFAA75278D188EC00785DB0AAFF67406" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Overall dorsal view A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng & Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure1" httpUri="https://binary.pensoft.net/fig/476312" pageId="0" pageNumber="1">1B</figureCitation>
|
||
,
|
||
<figureCitation id="7489D3F8AEA50872289E7118863AF170" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal view of carapace A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249) Terengganu D J. hoiseni Ng & Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure2" httpUri="https://binary.pensoft.net/fig/476313" pageId="0" pageNumber="1">2B</figureCitation>
|
||
). Orbits subovate; eye filling up most of orbital space; ocular peduncle relatively short, stout; cornea large, pigmented (Fig.
|
||
<figureCitation id="D623ADAB05892C1754A9430306962654" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Frontal view of cephalothorax A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng & Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure3" httpUri="https://binary.pensoft.net/fig/476314" pageId="0" pageNumber="1">3B</figureCitation>
|
||
). Supraorbital margin concave, entire (Fig.
|
||
<figureCitation id="4F6773A42405E4B962EAB3BC99920A20" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal view of carapace A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249) Terengganu D J. hoiseni Ng & Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure2" httpUri="https://binary.pensoft.net/fig/476313" pageId="0" pageNumber="1">2B</figureCitation>
|
||
). Suborbital margin concave, complete, cristate (Fig.
|
||
<figureCitation id="465B3F83EA5F311E6D4896A37980E79A" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Frontal view of cephalothorax A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng & Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure3" httpUri="https://binary.pensoft.net/fig/476314" pageId="0" pageNumber="1">3B</figureCitation>
|
||
). Antennules short, folding transversely in narrow fossa; antennae very short, not reaching cornea of eyes (Fig.
|
||
<figureCitation id="5E28B2AF9F8BF920614565CA4EABE702" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Frontal view of cephalothorax A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng & Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure3" httpUri="https://binary.pensoft.net/fig/476314" pageId="0" pageNumber="1">3B</figureCitation>
|
||
). Posterior margin of epistome with triangular median lobe; lateral part obliquely sloping, with 2 distinct concave margins (Fig.
|
||
<figureCitation id="186822F5D4E2340F82FB0A5B44CBE735" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Frontal view of cephalothorax A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng & Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure3" httpUri="https://binary.pensoft.net/fig/476314" pageId="0" pageNumber="1">3B</figureCitation>
|
||
). Mandibular palp with 3 distinct articles, terminal article single lobed.
|
||
</paragraph>
|
||
<paragraph id="41853E6CF837A2A0FF7B927627B64E76" pageId="0" pageNumber="1">
|
||
Third maxillipeds covering most of buccal cavity when closed; surfaces of merus, ischium and exopod with numerous long stiff setae; ischium subrectangular, with shallow median oblique groove; merus subquadrate, slightly wider than long, anteroexternal angle not expanded; exopod slender, reaching half length of merus, with long flagellum (Figs
|
||
<figureCitation id="8121A507D6CB6D243D22880C28722DB6" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Frontal view of cephalothorax A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng & Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure3" httpUri="https://binary.pensoft.net/fig/476314" pageId="0" pageNumber="1">3B</figureCitation>
|
||
,
|
||
<figureCitation id="CCAE92995AF48D1211C3BCA12710CB8E" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Left third maxilliped A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng & Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure4" httpUri="https://binary.pensoft.net/fig/476315" pageId="0" pageNumber="1">4B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="53248FF8BB07A170246117B8EECF063F" doi="10.3897/zookeys.994.56810.figure4" httpUri="https://binary.pensoft.net/fig/476315" pageId="0" pageNumber="1" start="Figure 4" startId="F4">
|
||
<paragraph id="47D098307D7D072167BE45EAC1036E39" pageId="0" pageNumber="1">
|
||
<emphasis id="27766B205CF8EF8A53251FA4AF482DAE" bold="true" pageId="0" pageNumber="1">Figure 4.</emphasis>
|
||
Left third maxilliped
|
||
<emphasis id="60A635708CBF4438B2AA076419C1C9FB" bold="true" pageId="0" pageNumber="1">A</emphasis>
|
||
<taxonomicName id="883624B8BF5034F452491EF712E01BA1" baseAuthorityName="Ng" baseAuthorityYear="1986" class="Malacostraca" family="Potamidae" genus="Johora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Johora intermedia" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="intermedia">
|
||
<emphasis id="489FAE7E777C5775C187CAD333DC80FF" italics="true" pageId="0" pageNumber="1">Johora intermedia</emphasis>
|
||
</taxonomicName>
|
||
Ng, 1986, holotype male (19.8
|
||
<normalizedToken id="F9D11826E8EE45F408D5F70FB05C3F47" originalValue="×">x</normalizedToken>
|
||
15.1 mm) (ZRC 1984.6529), Pahang
|
||
<emphasis id="448F26B7ECA55D950314EEE75ACDE3DE" bold="true" pageId="0" pageNumber="1">B</emphasis>
|
||
<taxonomicName id="F8E160D88DA9745CB31B9E13113FB0F6" lsidName="J. tahanensis" pageId="0" pageNumber="1" rank="species" species="tahanensis">
|
||
<emphasis id="B960CA07E7E832E17A2984531648D307" italics="true" pageId="0" pageNumber="1">J. tahanensis</emphasis>
|
||
</taxonomicName>
|
||
(Bott, 1966), male (30.7
|
||
<normalizedToken id="1FD1479090277A61FBA4E24EBDEC8217" originalValue="×">x</normalizedToken>
|
||
25.5 mm) (ZRC 1984.6795), Pahang
|
||
<emphasis id="D7EEEB994D2E6A9C46D7D7240DE5BD05" bold="true" pageId="0" pageNumber="1">C</emphasis>
|
||
<taxonomicName id="7D0B99C01DCEEDEDBF8CD8B6091E9F78" lsidName="J. thoi" pageId="0" pageNumber="1" rank="species" species="thoi">
|
||
<emphasis id="E8BFE87D4662424B24E9775D4844EDCB" italics="true" pageId="0" pageNumber="1">J. thoi</emphasis>
|
||
</taxonomicName>
|
||
Ng, 1990, holotype male (41.2
|
||
<normalizedToken id="BE74CFD61574B2A53700FD1EFD112AB8" originalValue="×">x</normalizedToken>
|
||
33.3 mm) (ZRC 1989.2249), Terengganu
|
||
<emphasis id="052F5526FBDBF2A1E74B125CAACB28CB" bold="true" pageId="0" pageNumber="1">D</emphasis>
|
||
<taxonomicName id="7EB596E81862EEB1729F7A1D35762C67" lsidName="J. hoiseni" pageId="0" pageNumber="1" rank="species" species="hoiseni">
|
||
<emphasis id="EBC1E4D81954945AFB7D8507CE2CBF54" italics="true" pageId="0" pageNumber="1">J. hoiseni</emphasis>
|
||
</taxonomicName>
|
||
Ng & Takeda, 1992, holotype male (25.1
|
||
<normalizedToken id="3F9E27299FD224733B011B72DB794E16" originalValue="×">x</normalizedToken>
|
||
21.1 mm) (ZRC 1984.6673), Kelantan
|
||
<emphasis id="D7B157C1EDC87B09CE1640D0F899EAD9" bold="true" pageId="0" pageNumber="1">E</emphasis>
|
||
<taxonomicName id="C24CEC88FFFCF6AA70077C3A87CC5A31" lsidName="J. thaiana" pageId="0" pageNumber="1" rank="species" species="thaiana">
|
||
<emphasis id="4A40E01269B2EB00CF42A0AC5508E166" italics="true" pageId="0" pageNumber="1">J. thaiana</emphasis>
|
||
</taxonomicName>
|
||
Leelawathanagoon, Lheknim & Ng, 2005, paratype male (22.2
|
||
<normalizedToken id="5C09A6699C6AA13060F5894E022FE035" originalValue="×">x</normalizedToken>
|
||
18.6 mm) (ZRC 2006.0052), Thailand
|
||
<emphasis id="4E0A7FB7D2A7BF6DB323A9C4DB7FC6FB" bold="true" pageId="0" pageNumber="1">F</emphasis>
|
||
<taxonomicName id="EB4E5971246D22B5F75679652253A07D" lsidName="J. booliati" pageId="0" pageNumber="1" rank="species" species="booliati">
|
||
<emphasis id="53C021CEF41008A6AA1440E30CE43158" italics="true" pageId="0" pageNumber="1">J. booliati</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype male (34.6
|
||
<normalizedToken id="66DCD97D1AF7CC0AB958853F8F3A6F2E" originalValue="×">x</normalizedToken>
|
||
30.0 mm) (ZRC 2020.0072), Pahang
|
||
<emphasis id="A42691BBF16ED81D1172102313EE6C39" bold="true" pageId="0" pageNumber="1">G</emphasis>
|
||
<taxonomicName id="699A53106EA31AB1280EDC294DB70FAC" lsidName="J. erici" pageId="0" pageNumber="1" rank="species" species="erici">
|
||
<emphasis id="1CE166F00A40129449746C3CAF11C50F" italics="true" pageId="0" pageNumber="1">J. erici</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype male (38.3
|
||
<normalizedToken id="193BF1206EF691540D1BC582E3E6AE6F" originalValue="×">x</normalizedToken>
|
||
32.0 mm) (ZRC 2020.0360), Perak
|
||
<emphasis id="31B9DFD5ED77B39DF45749E1F8240F67" bold="true" pageId="0" pageNumber="1">H</emphasis>
|
||
<taxonomicName id="EC6B7EA25875430650B752E1EFF53667" lsidName="J. michaeli" pageId="0" pageNumber="1" rank="species" species="michaeli">
|
||
<emphasis id="9E39EDDA9A82A65485C1A422337F9EB0" italics="true" pageId="0" pageNumber="1">J. michaeli</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype male (22.7
|
||
<normalizedToken id="E4CFEA9CB62E35D23EDA3150E6D89342" originalValue="×">x</normalizedToken>
|
||
19.2 mm) (ZRC 2010.0047), Terengganu.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="96AD4F82921675F0FFF0599CE42D5BB0" pageId="0" pageNumber="1">
|
||
Chelipeds asymmetrical (Fig.
|
||
<figureCitation id="726B4C6D7880F647D094309693865AB9" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Overall dorsal view A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng & Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure1" httpUri="https://binary.pensoft.net/fig/476312" pageId="0" pageNumber="1">1B</figureCitation>
|
||
). Anterior margin of basis-ischium smooth; margins of merus rugose, uneven; inner margin lined with dense setae. Outer surface of carpus rugose, inner distal angle with sharp tooth (Fig.
|
||
<figureCitation id="E51DDAE6B2D92EE10447F5078CDD0252" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Overall dorsal view A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng & Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure1" httpUri="https://binary.pensoft.net/fig/476312" pageId="0" pageNumber="1">1B</figureCitation>
|
||
). Outer surfaces of chelae rugose; chela stout (Fig.
|
||
<figureCitation id="A6703145AD6C21E64BD57B45B23E89F4" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Overall dorsal view A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng & Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure1" httpUri="https://binary.pensoft.net/fig/476312" pageId="0" pageNumber="1">1B</figureCitation>
|
||
). Fingers of chela almost straight, longer than palm; cutting edges of both fingers with variously sized teeth and denticles; fingers of slightly smaller chela similar (Fig.
|
||
<figureCitation id="A7F996E74C6C39392BAE607999211469" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Overall dorsal view A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng & Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure1" httpUri="https://binary.pensoft.net/fig/476312" pageId="0" pageNumber="1">1B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="9484EA1F09E526D8D0F22A0A97C459C6" pageId="0" pageNumber="1">
|
||
Ambulatory legs not elongate, length to width ratio of merus of fourth ambulatory leg 2.7-2.8; second pair longest, fourth pair shortest (Fig.
|
||
<figureCitation id="BEC3400CF3989FB5969C77A39BC5166F" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Overall dorsal view A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng & Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure1" httpUri="https://binary.pensoft.net/fig/476312" pageId="0" pageNumber="1">1B</figureCitation>
|
||
). Surface of merus gently rugose, dorsal margin weakly cristate, slightly uneven, appears serrated, without subdistal spine or tooth; carpus rugose with short setae; propodus laterally flattened, margins with short setae; dactylus gently curved, setose, margins with short spines (Fig.
|
||
<figureCitation id="1945C9D1744336BE3A707865A42ADF0B" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Overall dorsal view A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng & Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure1" httpUri="https://binary.pensoft.net/fig/476312" pageId="0" pageNumber="1">1B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="56F366DBFB16D4245A87941E142F5383" pageId="0" pageNumber="1">
|
||
Thoracic sternum (notably sternites 3, 4) with shallow pits to smooth (Fig.
|
||
<figureCitation id="408ED02D6378030F558FC19797A70D76" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Anterior thoracic sternum and pleon A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng & Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure5" httpUri="https://binary.pensoft.net/fig/476316" pageId="0" pageNumber="1">5B</figureCitation>
|
||
). Sternites 1, 2 completely fused to form triangular plate with convex margins; separated from sternite 3 by distinct, gently concave suture lined with short setae; sternites 3, 4 completely fused except for shallow oblique depression between sternites, lined with short stiff setae (Fig.
|
||
<figureCitation id="A0E314D9764A965711155339FCAF10BF" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Anterior thoracic sternum and pleon A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng & Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure5" httpUri="https://binary.pensoft.net/fig/476316" pageId="0" pageNumber="1">5B</figureCitation>
|
||
); sutures between sternites 4/5, 5/6, 6/7 medially interrupted; suture between sternites 7, 8 complete; deep longitudinal groove on sternite 8 extending to most of sternite 7. Penis on condyle of coxa of fourth ambulatory leg. Sternopleonal cavity deep, reaching to imaginary line connecting submedian parts of cheliped coxae (Fig.
|
||
<figureCitation id="E6BD342AFD45F2F27A841BBB3A03F6F3" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Anterior thoracic sternum and pleon A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng & Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure5" httpUri="https://binary.pensoft.net/fig/476316" pageId="0" pageNumber="1">5B</figureCitation>
|
||
). Male pleonal locking tubercle relatively large, low, round, positioned on posterior edge of sternite 5, just adjacent to sternite 6.
|
||
</paragraph>
|
||
<caption id="849407157328C3BCE4871817F7F5FBFF" doi="10.3897/zookeys.994.56810.figure5" httpUri="https://binary.pensoft.net/fig/476316" pageId="0" pageNumber="1" start="Figure 5" startId="F5">
|
||
<paragraph id="C0BAFE19A838A527D09BAC8BD44688F5" pageId="0" pageNumber="1">
|
||
<emphasis id="287F684653AB0AA98AF51249E8E591D2" bold="true" pageId="0" pageNumber="1">Figure 5.</emphasis>
|
||
Anterior thoracic sternum and pleon
|
||
<emphasis id="F2AF39B9B2FB851166B281B6DB7EEDD5" bold="true" pageId="0" pageNumber="1">A</emphasis>
|
||
<taxonomicName id="318A8C97C04821AC5AB53340842C6388" baseAuthorityName="Ng" baseAuthorityYear="1986" class="Malacostraca" family="Potamidae" genus="Johora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Johora intermedia" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="intermedia">
|
||
<emphasis id="C8303035A18E147F0FB59465E7190B62" italics="true" pageId="0" pageNumber="1">Johora intermedia</emphasis>
|
||
</taxonomicName>
|
||
Ng, 1986, holotype male (19.8
|
||
<normalizedToken id="F97A5E14AC58AB1281CED84994673D99" originalValue="×">x</normalizedToken>
|
||
15.1 mm) (ZRC 1984.6529), Pahang
|
||
<emphasis id="8875BF8E307AEC6DB7683012D7CB7282" bold="true" pageId="0" pageNumber="1">B</emphasis>
|
||
<taxonomicName id="A90C05CEF5FEE080541114CBF9F5A52A" lsidName="J. tahanensis" pageId="0" pageNumber="1" rank="species" species="tahanensis">
|
||
<emphasis id="630939E15AD9B42CF507D4EBC86BBEC9" italics="true" pageId="0" pageNumber="1">J. tahanensis</emphasis>
|
||
</taxonomicName>
|
||
(Bott, 1966), male (30.7
|
||
<normalizedToken id="B7B3885BA2E7B3FBFDA0D9E9DAD56AA9" originalValue="×">x</normalizedToken>
|
||
25.5 mm) (ZRC 1984.6795), Pahang
|
||
<emphasis id="6C09AAC037D5B88A0EDD636DA85C42AC" bold="true" pageId="0" pageNumber="1">C</emphasis>
|
||
<taxonomicName id="568488202482A35169F65F44AF0354B4" lsidName="J. thoi" pageId="0" pageNumber="1" rank="species" species="thoi">
|
||
<emphasis id="FCCDFD20D98F15ADF0BC5CE2E770B0DA" italics="true" pageId="0" pageNumber="1">J. thoi</emphasis>
|
||
</taxonomicName>
|
||
Ng, 1990, holotype male (41.2
|
||
<normalizedToken id="3E47118B8B53CB207F755F730B5BB8C9" originalValue="×">x</normalizedToken>
|
||
33.3 mm) (ZRC 1989.2249), Terengganu
|
||
<emphasis id="4499255F9924645DDBE003E3E2D16EE8" bold="true" pageId="0" pageNumber="1">D</emphasis>
|
||
<taxonomicName id="69012F7E7FF1C631CAAA785B65E9E736" lsidName="J. hoiseni" pageId="0" pageNumber="1" rank="species" species="hoiseni">
|
||
<emphasis id="054BB7532F61CC29F840BED2694696AF" italics="true" pageId="0" pageNumber="1">J. hoiseni</emphasis>
|
||
</taxonomicName>
|
||
Ng & Takeda, 1992, holotype male (25.1
|
||
<normalizedToken id="1F7245C1C14DAA9EC9C902D69C2B96A4" originalValue="×">x</normalizedToken>
|
||
21.1 mm) (ZRC 1984.6673), Kelantan
|
||
<emphasis id="1D9C78E4BD8EAEA41F903A22DDB7C107" bold="true" pageId="0" pageNumber="1">E</emphasis>
|
||
<taxonomicName id="87762395D7AC69A61D102930016A46A5" lsidName="J. thaiana" pageId="0" pageNumber="1" rank="species" species="thaiana">
|
||
<emphasis id="3F46F777ED26D8A7DFA0414B035FB6F8" italics="true" pageId="0" pageNumber="1">J. thaiana</emphasis>
|
||
</taxonomicName>
|
||
Leelawathanagoon, Lheknim & Ng, 2005, paratype male (22.2
|
||
<normalizedToken id="0E93F1C6A9C9CB79EDF2FD6892D851BC" originalValue="×">x</normalizedToken>
|
||
18.6 mm) (ZRC 2006.0052), Thailand
|
||
<emphasis id="2649B88754CE0C709B49F3701F23E362" bold="true" pageId="0" pageNumber="1">F</emphasis>
|
||
<taxonomicName id="C732C9D0A021494E0624B4435F382C70" lsidName="J. booliati" pageId="0" pageNumber="1" rank="species" species="booliati">
|
||
<emphasis id="F4E8C162A99DD16CC5AABF1F05B8AF72" italics="true" pageId="0" pageNumber="1">J. booliati</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype male (34.6
|
||
<normalizedToken id="84ECD97B4AD630C5DB704182B1CE5AB5" originalValue="×">x</normalizedToken>
|
||
30.0 mm) (ZRC 2020.0072), Pahang
|
||
<emphasis id="12A377235D869473037A4DEA893C75A6" bold="true" pageId="0" pageNumber="1">G</emphasis>
|
||
<taxonomicName id="EA314CA9D7CCF499EAEE757C8457053A" lsidName="J. erici" pageId="0" pageNumber="1" rank="species" species="erici">
|
||
<emphasis id="4D4E064C968856AF332053B13637737D" italics="true" pageId="0" pageNumber="1">J. erici</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype male (38.3
|
||
<normalizedToken id="F12FCA57B787591AFA59324C6EBF8002" originalValue="×">x</normalizedToken>
|
||
32.0 mm) (ZRC 2020.0360), Perak
|
||
<emphasis id="B42567EF16D3DC63BE9F36D2F957841D" bold="true" pageId="0" pageNumber="1">H</emphasis>
|
||
<taxonomicName id="6BF5BE20FA79ED82EEBAA4E43C39052F" lsidName="J. michaeli" pageId="0" pageNumber="1" rank="species" species="michaeli">
|
||
<emphasis id="2D8D314540F65F2D4C83B074640BDE28" italics="true" pageId="0" pageNumber="1">J. michaeli</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype male (22.7
|
||
<normalizedToken id="02D0A0538A903F123779055CCB2F91D8" originalValue="×">x</normalizedToken>
|
||
19.2 mm) (ZRC 2010.0047), Terengganu.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="55A8B5AC80F8A80FC111F51B597960A1" pageId="0" pageNumber="1">
|
||
Pleon triangular, all somites, telson free; telson triangular, lateral margins almost straight to gently sinuous; somite 6 subtrapezoidal, distinctly wider than long, lateral margins gently sinuous; somites 3-5 trapezoidal, gradually decreasing in width; somites 1, 2 subrectangular, narrow, very wide, reaching to bases of coxae of fourth ambulatory legs, thoracic sternite 8 not visible when pleon closed (Fig.
|
||
<figureCitation id="6CEBBA4158A9D58D674E310B0D3DC41D" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Anterior thoracic sternum and pleon A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng & Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure5" httpUri="https://binary.pensoft.net/fig/476316" pageId="0" pageNumber="1">5B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="A8DA97579D07A75D06A4D48CE70DAEAB" pageId="0" pageNumber="1">
|
||
G1 subterminal segment gradually tapering from broad proximal part to slender distal part, without distinct shelf-like structure along gently concave outer margin; terminal segment gently curved outwards (from median part of sternum), ca. half length of subterminal segment, surfaces with numerous short setae (Fig.
|
||
<figureCitation id="981B55B0295370FA6CA7C07EE4E7E3E0" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. A-D Johora tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang E-H J. hoiseni Ng & Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan A, E left G 1 (ventral view) B, F distal part of left G 1 (ventral view) C, G distal part of left G 1 (dorsal view) D, H left G 2 (ventral view) E-H after Ng and Takeda (1992: fig. 3 B, D-F). Scale bars: 1.0 mm (A, D, E, H); 0.5 mm (B, C, F, G)." figureDoi="10.3897/zookeys.994.56810.figure8" httpUri="https://binary.pensoft.net/fig/476319" pageId="0" pageNumber="1">8A-C</figureCitation>
|
||
); G2 slightly longer than G1, distal segment long, about half length of basal segment (Fig.
|
||
<figureCitation id="2982DADF2F463F89DCAC311368A6F684" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. A-D Johora tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang E-H J. hoiseni Ng & Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan A, E left G 1 (ventral view) B, F distal part of left G 1 (ventral view) C, G distal part of left G 1 (dorsal view) D, H left G 2 (ventral view) E-H after Ng and Takeda (1992: fig. 3 B, D-F). Scale bars: 1.0 mm (A, D, E, H); 0.5 mm (B, C, F, G)." figureDoi="10.3897/zookeys.994.56810.figure8" httpUri="https://binary.pensoft.net/fig/476319" pageId="0" pageNumber="1">8D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="63122FF9720857FEA759B2A637778EC0" pageId="0" pageNumber="1">
|
||
<emphasis id="893A4EE861E6958FCCACE3C0ABCA2C27" bold="true" pageId="0" pageNumber="1">Female</emphasis>
|
||
. Similar to male in most non-sexual features; chelipeds symmetrical or only slightly asymmetrical (Fig.
|
||
<figureCitation id="39183BA7FB277E175CD6AAB796C4CF3A" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Overall dorsal view A Johora intermedia Ng, 1986, female (17.9 x 14.7 mm) (ZRC 2001.1002), Selangor B J. tahanensis (Bott, 1966), female (24.4 x 21.0 mm) (ZRC 1984.6765), Pahang C J. thoi Ng, 1990, female (31.7 x 25.5 mm) (ZRC 2001.1167), Terengganu D J. hoiseni Ng & Takeda, 1992, paratype female (22.1 x 19.0 mm) (ZRC 1984.6675), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype female (21.0 x 17.4 mm) (ZRC 2006.0053), Thailand F J. booliati sp. nov., paratype female (40.4 x 34.4 mm) (ZRC 1995.270), Pahang G J. erici sp. nov., paratype female (32.8 x 26.9 mm) (ZRC 1995.268), Perak H J. michaeli sp. nov., paratype female (29.9 x 25.0 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure12" httpUri="https://binary.pensoft.net/fig/476323" pageId="0" pageNumber="1">12B</figureCitation>
|
||
). Pleon longitudinally ovate, covering most of thoracic sternal surface, all somites and telson free; somites 3-6 progressively narrower; telson subtriangular (Fig.
|
||
<figureCitation id="D5BFE2BD896E501ABD8F3D6341350847" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Female pleon A Johora intermedia Ng, 1986, female (17.9 x 14.7 mm) (ZRC 2001.1002), Selangor B J. tahanensis (Bott, 1966), female (24.4 x 21.0 mm) (ZRC 1984.6765), Pahang C J. thoi Ng, 1990, female (31.7 x 25.5 mm) (ZRC 2001.1167), Terengganu D J. hoiseni Ng & Takeda, 1992, paratype female (22.1 x 19.0 mm) (ZRC 1984.6675), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype female (21.0 x 17.4 mm) (ZRC 2006.0053), Thailand F J. booliati sp. nov., paratype female (40.4 x 34.4 mm) (ZRC 1995.270), Pahang G J. erici sp. nov., paratype female (32.8 x 26.9 mm) (ZRC 1995.268), Perak H J. michaeli sp. nov., paratype female (29.9 x 25.0 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure13" httpUri="https://binary.pensoft.net/fig/476324" pageId="0" pageNumber="1">13B</figureCitation>
|
||
). Vulvae large, covering anterior half of sternite 6, slightly pushing into suture with sternite 5, lateral sternal vulvar cover semicircular (Fig.
|
||
<figureCitation id="EDA832C65895446752B2D9CA17A403D0" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Sternopleonal cavity and vulvae A Johora intermedia Ng, 1986, female (17.9 x 14.7 mm) (ZRC 2001.1002), Selangor B J. tahanensis (Bott, 1966), female (24.4 x 21.0 mm) (ZRC 1984.6765), Pahang C J. thoi Ng, 1990, female (31.7 x 25.5 mm) (ZRC 2001.1167), Terengganu D J. hoiseni Ng & Takeda, 1992, paratype female (22.1 x 19.0 mm) (ZRC 1984.6675), Kelantan E J. thaiana Leelawathanagoon, Lheknim & Ng, 2005, paratype female (21.0 x 17.4 mm) (ZRC 2006.0053), Thailand F J. booliati sp. nov., paratype female (40.4 x 34.4 mm) (ZRC 1995.270), Pahang G J. erici sp. nov., paratype female (32.8 x 26.9 mm) (ZRC 1995.268), Perak H J. michaeli sp. nov., paratype female (29.9 x 25.0 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure14" httpUri="https://binary.pensoft.net/fig/476325" pageId="0" pageNumber="1">14B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="EADB2AE3EAEBBBB14A3A56CBD0024EC8" pageId="0" pageNumber="1" type="remarks">
|
||
<paragraph id="75842457843116857508C42042EE375A" pageId="0" pageNumber="1">Remarks.</paragraph>
|
||
<paragraph id="6DE171B2B7DD847826F72946C5C9E118" pageId="0" pageNumber="1">
|
||
<bibRefCitation id="47DD8A4B306E2D7B122B13208DEC5DAC" author="Ng, PKL" journalOrPublisher="Bulletin of the National Science Museum, Tokyo, Series A" pageId="0" pageNumber="1" pagination="103 - 116" refId="B19" refString="Ng, PKL, Takeda, M, 1992. On some freshwater crabs (Crustacea, Brachyura, Potamidae, Parathelphusidae and Grapsidae) from Peninsular Malaysia. Bulletin of the National Science Museum, Tokyo, Series A 18 (3): 103 - 116" title="On some freshwater crabs (Crustacea, Brachyura, Potamidae, Parathelphusidae and Grapsidae) from Peninsular Malaysia." volume="18" year="1992">Ng and Takeda (1992</bibRefCitation>
|
||
: 108) discussed the taxonomy of
|
||
<taxonomicName id="948A10FF483049EBA8AC2D231F4D55AA" lsidName="J. tahanensis" pageId="0" pageNumber="1" rank="species" species="tahanensis">
|
||
<emphasis id="DFB95CEF14D0B6A5F2AE086488B25D73" italics="true" pageId="0" pageNumber="1">J. tahanensis</emphasis>
|
||
</taxonomicName>
|
||
at length, showing that the material from two parts of Taman Negara National Park in Malaysia belonged to two species and that the type series is mixed.
|
||
<bibRefCitation id="8FE1FDAC2AC121676A0C89950B601ADC" author="Bott, R" journalOrPublisher="Senckenbergiana biologica" pageId="0" pageNumber="1" pagination="469 - 509" refId="B1" refString="Bott, R, 1966. Potamiden aus Asien (Potamon Savigny and Potamiscus Alcock) (Crustacea, Decapoda). Senckenbergiana biologica 47: 469 - 509" title="Potamiden aus Asien (Potamon Savigny and Potamiscus Alcock) (Crustacea, Decapoda)." volume="47" year="1966">Bott (1966</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="B8613DCCA36D2255A9B40B39439187AF" author="Bott, R" journalOrPublisher="Abhandlungen der senckenbergischen naturforschenden Gesellschaft" pageId="0" pageNumber="1" pagination="1 - 338" refId="B2" refString="Bott, R, 1970. Die Suesswasserkrabben von Europa, Asien, Australien und ihre Stammesgeschichte. Eine Revision der Potamoidea und der Parathelphusoidea (Crustacea, Decapoda). Abhandlungen der senckenbergischen naturforschenden Gesellschaft 526: 1 - 338" title="Die Suesswasserkrabben von Europa, Asien, Australien und ihre Stammesgeschichte. Eine Revision der Potamoidea und der Parathelphusoidea (Crustacea, Decapoda)." volume="526" year="1970">1970</bibRefCitation>
|
||
) and
|
||
<bibRefCitation id="BC9E348A216FC78B2D63E3BAF42D6A04" author="Ng, PKL" journalOrPublisher="Malayan Nature Journal" pageId="0" pageNumber="1" pagination="13 - 44" refId="B11" refString="Ng, PKL, 1987. A revision of the Malayan freshwater crabs of the genus Johora Bott, 1966 stat. nov. (Decapoda: Brachyura: Potamidae). Malayan Nature Journal 41: 13 - 44" title="A revision of the Malayan freshwater crabs of the genus Johora Bott, 1966 stat. nov. (Decapoda: Brachyura: Potamidae)." volume="41" year="1987">Ng (1987</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="5D9BE562B238288DC56AA758D95B48DF" author="Ng, PKL" journalOrPublisher="Department of Zoology, National University of Singapore, Shinglee Press" pageId="0" pageNumber="1" refId="B12" refString="Ng, PKL, 1988. The Freshwater Crabs of Peninsular Malaysia and Singapore. Department of Zoology, National University of Singapore, Shinglee Press" title="The Freshwater Crabs of Peninsular Malaysia and Singapore." year="1988">1988</bibRefCitation>
|
||
) had regarded them as one taxon, incorrectly noting that the G1 structure was variable. The holotype of
|
||
<taxonomicName id="A9482A4ACF458EF42934667A2C59E3EC" lsidName="J. tahanensis" pageId="0" pageNumber="1" rank="species" species="tahanensis">
|
||
<emphasis id="C73280E64BA4318CF45B670BBDFB6824" italics="true" pageId="0" pageNumber="1">J. tahanensis</emphasis>
|
||
</taxonomicName>
|
||
was from Kuala Tahan in the state of Pahang, in the southern part of the national park, and the G1 of this specimen (in the Senckenberg Museum, Frankfurt) as well as the topotypic material we have from that location has a terminal segment which is gently curved (Fig.
|
||
<figureCitation id="E8172AC93C61BCC999FF7F68A3CDE07B" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. A-D Johora tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang E-H J. hoiseni Ng & Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan A, E left G 1 (ventral view) B, F distal part of left G 1 (ventral view) C, G distal part of left G 1 (dorsal view) D, H left G 2 (ventral view) E-H after Ng and Takeda (1992: fig. 3 B, D-F). Scale bars: 1.0 mm (A, D, E, H); 0.5 mm (B, C, F, G)." figureDoi="10.3897/zookeys.994.56810.figure8" httpUri="https://binary.pensoft.net/fig/476319" pageId="0" pageNumber="1">8A</figureCitation>
|
||
). Material from the northern part of the park from the adjacent state of Kelantan had a G1 terminal segment that is straight (Fig.
|
||
<figureCitation id="A57EC2DAF3B5D7DD6CFA38321B5DE7C5" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. A-D Johora tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang E-H J. hoiseni Ng & Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan A, E left G 1 (ventral view) B, F distal part of left G 1 (ventral view) C, G distal part of left G 1 (dorsal view) D, H left G 2 (ventral view) E-H after Ng and Takeda (1992: fig. 3 B, D-F). Scale bars: 1.0 mm (A, D, E, H); 0.5 mm (B, C, F, G)." figureDoi="10.3897/zookeys.994.56810.figure8" httpUri="https://binary.pensoft.net/fig/476319" pageId="0" pageNumber="1">8E</figureCitation>
|
||
).
|
||
<bibRefCitation id="84E0F960D2085DFB353820CDE2178534" author="Ng, PKL" journalOrPublisher="Bulletin of the National Science Museum, Tokyo, Series A" pageId="0" pageNumber="1" pagination="103 - 116" refId="B19" refString="Ng, PKL, Takeda, M, 1992. On some freshwater crabs (Crustacea, Brachyura, Potamidae, Parathelphusidae and Grapsidae) from Peninsular Malaysia. Bulletin of the National Science Museum, Tokyo, Series A 18 (3): 103 - 116" title="On some freshwater crabs (Crustacea, Brachyura, Potamidae, Parathelphusidae and Grapsidae) from Peninsular Malaysia." volume="18" year="1992">Ng and Takeda (1992)</bibRefCitation>
|
||
showed that the G1 differences of the Pahang and Kelantan specimens are consistent and the Kelantan material was referred to a separate species,
|
||
<taxonomicName id="C33D060D273327708110EA98875FEDC2" lsidName="J. hoiseni" pageId="0" pageNumber="1" rank="species" species="hoiseni">
|
||
<emphasis id="8E2D91FCEEF5B5F6C28033CA973A261A" italics="true" pageId="0" pageNumber="1">J. hoiseni</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="6C6D8298F57A6C08FF09C321C5C53879" pageId="0" pageNumber="1">
|
||
The material reported as "
|
||
<taxonomicName id="09B4A97233A950E914CD889E387EA084" lsidName="J. tahanensis" pageId="0" pageNumber="1" rank="species" species="tahanensis">
|
||
<emphasis id="D450DB2EC7EABBF6FBB6F2206169D630" italics="true" pageId="0" pageNumber="1">J. tahanensis</emphasis>
|
||
</taxonomicName>
|
||
" by
|
||
<bibRefCitation id="63B6A89C853AEB73CB829A717CB542EF" author="Ng, PKL" journalOrPublisher="Bulletin of the National Science Museum, Tokyo, Series A" pageId="0" pageNumber="1" pagination="103 - 116" refId="B19" refString="Ng, PKL, Takeda, M, 1992. On some freshwater crabs (Crustacea, Brachyura, Potamidae, Parathelphusidae and Grapsidae) from Peninsular Malaysia. Bulletin of the National Science Museum, Tokyo, Series A 18 (3): 103 - 116" title="On some freshwater crabs (Crustacea, Brachyura, Potamidae, Parathelphusidae and Grapsidae) from Peninsular Malaysia." volume="18" year="1992">Ng and Takeda (1992</bibRefCitation>
|
||
: 108) from and around
|
||
<normalizedToken id="34364DFA20E7956BE818E71DE1D4BA8A" originalValue="Fraser’s">Fraser's</normalizedToken>
|
||
Hill in Selangor and Pahang should be now referred to
|
||
<taxonomicName id="F0E858F1B66D613622E2ECB6FBEEAC58" lsidName="J. booliati" pageId="0" pageNumber="1" rank="species" species="booliati">
|
||
<emphasis id="D20133569C1B5D86D482E1063B0BCD84" italics="true" pageId="0" pageNumber="1">J. booliati</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. (see remarks for this species).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="292B1AE81515A5E68E00D775B8079D18" pageId="0" pageNumber="1" type="distribution">
|
||
<paragraph id="EC22FA4934AFE59432286E42F0CE027B" pageId="0" pageNumber="1">Distribution.</paragraph>
|
||
<paragraph id="63E9927D45F1C73A2B56E62C321CB149" pageId="0" pageNumber="1">
|
||
<taxonomicName id="7A1A5D4F3DC7B1F8D1BFA8BFA4242A40" authorityName="Bott" authorityYear="1966" class="Malacostraca" family="Potamidae" genus="Johora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Johora tahanensis" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="tahanensis">
|
||
<emphasis id="2C028C0B599A82EDA4FFB4BC3F2BB3A7" italics="true" pageId="0" pageNumber="1">Johora tahanensis</emphasis>
|
||
</taxonomicName>
|
||
is known from the tributaries around Sungei Tahan in southern Pahang, at the southern part of Taman Negara National Park (Fig.
|
||
<figureCitation id="AAD4F4706E1790DEA30C9816AF955274" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Distribution of species of the Johora tahanensis species group 1 J. tahanensis (Bott, 1966) 2 J. thoi Ng, 1990 3 J. hoiseni Ng & Takeda, 1992 4 J. thaiana Leelawathanagoon, Lheknim & Ng, 2005 5 J. booliati sp. nov. 6 J. erici sp. nov. 7 J. michaeli sp. nov. Map data 2020 Google." figureDoi="10.3897/zookeys.994.56810.figure15" httpUri="https://binary.pensoft.net/fig/476326" pageId="0" pageNumber="1">15</figureCitation>
|
||
). It lives among rocks and submerged vegetation in the relatively fast flowing streams in the forest.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="EE9612341E38561F3F89DF5A1B2A172D" pageId="0" pageNumber="1" type="conservation">
|
||
<paragraph id="1CBEEE6A22C89DAD2781A812011F699D" pageId="0" pageNumber="1">Conservation.</paragraph>
|
||
<paragraph id="57288267B18819C03F23AAB1EBCBE319" pageId="0" pageNumber="1">
|
||
The species is not under any immediate threat as it is found in
|
||
<normalizedToken id="73E46C014A61FE190E8A587ADAEA9D4A" originalValue="Malaysia’s">Malaysia's</normalizedToken>
|
||
oldest and largest national park.
|
||
<bibRefCitation id="BA46714D479B65AED8AE089E66900780" author="Ng, PKL" editor="Chua, L" journalOrPublisher="28 - 30 June 2005, Forest Research Institute Malaysia, Kepong" pageId="0" pageNumber="1" pagination="95 - 120" refId="B21" refString="Ng, PKL, Yeo, DCJ, 2007. Malaysian freshwater crabs: conservation prospects and challenges. In: Chua, L, Ed., Proceedings of the Seminar on the Status of Biological Diversity in Malaysia & Threat Assessment of Plant Species in Malaysia. 28-30 June 2005, Forest Research Institute Malaysia, Kepong: 95 - 120" title="Malaysian freshwater crabs: conservation prospects and challenges." volumeTitle="Proceedings of the Seminar on the Status of Biological Diversity in Malaysia & Threat Assessment of Plant Species in Malaysia." year="2007">Ng and Yeo (2007)</bibRefCitation>
|
||
treated
|
||
<taxonomicName id="9FE206E69D754C3B102268952456282E" lsidName="J. tahanensis" pageId="0" pageNumber="1" rank="species" species="tahanensis">
|
||
<emphasis id="623B7C2E93D5E5F36F1BB8EBCB23EA19" italics="true" pageId="0" pageNumber="1">J. tahanensis</emphasis>
|
||
</taxonomicName>
|
||
s. str. as vulnerable as it is not known from a relatively wide geographic area but it was reappraised and regarded as of least concern in
|
||
<bibRefCitation id="5A783A35CAFBEFFE2DBCBCD83C818332" DOI="https://doi.org/10.1016/j.biocon.2009.02.038" author="Cumberlidge, N" journalOrPublisher="Biological Conservation" pageId="0" pageNumber="1" pagination="1665 - 1673" refId="B3" refString="Cumberlidge, N, Ng, PKL, Yeo, DCJ, Magalhaes, C, Campos, MR, Alvarez, F, Naruse, T, Daniels, SR, Esser, LJ, Attipoe, FYK, Clotilde-Ba, F-L, Darwall, W, Mcivor, A, Collen, B, Ram, M, 2009. Freshwater crabs and the biodiversity crisis: importance, threats, status, and conservation challenges. Biological Conservation 142: 1665 - 1673, DOI: https://doi.org/10.1016/j.biocon.2009.02.038" title="Freshwater crabs and the biodiversity crisis: importance, threats, status, and conservation challenges." url="https://doi.org/10.1016/j.biocon.2009.02.038" volume="142" year="2009">Cumberlidge et al. (2009)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |