467 lines
52 KiB
XML
467 lines
52 KiB
XML
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<mods:title id="78354D70392B03ECFCC37CBE629CF053">A new leaf-tailed gecko of the Uroplatus ebenaui group (Squamata: Gekkonidae) from Madagascar's central eastern rainforests</mods:title>
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<mods:namePart id="1B27505D5B14D9543D1CE700AE387335">Ratsoavina, Fanomezana Mihaja</mods:namePart>
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<mods:namePart id="15B78B2B38070DFA4C43E00065331864">Ranjanaharisoa, Fiadanantsoa Andrianja</mods:namePart>
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<mods:namePart id="5CEEE54954FF6DC84AABE6CCFFB44A28">Glaw, Frank</mods:namePart>
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<mods:namePart id="661566A2DDBF6692FF6984FE1239A30B">Raselimanana, Achille P.</mods:namePart>
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<mods:namePart id="836267FE4A87B446157DE33A54F1EB2B">Miralles, Aurélien</mods:namePart>
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<mods:namePart id="DD9E9C646AD50E23C619C6B16018268B">Vences, Miguel</mods:namePart>
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<mods:title id="8AD3737C551754E46FF077A49D4E3469">Zootaxa</mods:title>
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<mods:date id="13FCC89E22D66BF6C46B5CB0879D6AF6">2015</mods:date>
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<mods:number id="41F354F2BB6DC1F89B7C10E6EDF47F8A">4006</mods:number>
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<treatment id="C6553303FF80FFB6FF414240FD5066A1" ID-DOI="http://doi.org/10.5281/zenodo.6097408" ID-GBIF-Taxon="127711705" ID-Zenodo-Dep="6097408" LSID="urn:lsid:plazi:treatment:C6553303FF80FFB6FF414240FD5066A1" httpUri="http://treatment.plazi.org/id/C6553303FF80FFB6FF414240FD5066A1" lastPageId="15" lastPageNumber="158" pageId="11" pageNumber="154">
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<subSubSection id="06E6D19EFF80FFB2FF414240FEFE6553" pageId="11" pageNumber="154" type="nomenclature">
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<paragraph id="4E438215FF80FFB2FF414240FE656532" blockId="11.[151,437,547,607]" box="[151,437,547,574]" pageId="11" pageNumber="154">
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<heading id="150B3579FF80FFB2FF414240FE656532" bold="true" box="[151,437,547,574]" fontSize="11" level="1" pageId="11" pageNumber="154" reason="1">
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<taxonomicName id="89FCF996FF80FFB2FF414240FE806531" box="[151,336,547,573]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="154" phylum="Chordata" rank="species" species="fiera" status="sp. nov.">
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<emphasis id="7C885E07FF80FFB2FF414240FE806531" bold="true" box="[151,336,547,573]" italics="true" pageId="11" pageNumber="154">Uroplatus fiera</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="67BBE37CFF80FFB2FE814247FE656532" box="[343,437,548,574]" pageId="11" pageNumber="154" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="4E438215FF80FFB2FF414225FEFE6553" blockId="11.[151,437,547,607]" box="[151,302,582,607]" pageId="11" pageNumber="154">(Figures 7–8)</paragraph>
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</subSubSection>
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<subSubSection id="06E6D19EFF80FFB2FF4142EDFC1E641F" pageId="11" pageNumber="154" type="discussion">
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<paragraph id="4E438215FF80FFB2FF4142EDFC1E641F" blockId="11.[151,1437,654,2011]" pageId="11" pageNumber="154">
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<emphasis id="7C885E07FF80FFB2FF4142EDFED065AB" bold="true" box="[151,256,654,679]" pageId="11" pageNumber="154">Remark.</emphasis>
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This new species was previously referred to as
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<taxonomicName id="89FCF996FF80FFB2FCE142EDFBDB65AA" box="[823,1035,654,678]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="154" phylum="Chordata" rank="species" species="ebenaui">
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<emphasis id="7C885E07FF80FFB2FCE142EDFBDB65AA" box="[823,1035,654,678]" italics="true" pageId="11" pageNumber="154">Uroplatus ebenaui</emphasis>
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</taxonomicName>
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[Ca7] by
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<bibRefCitation id="2A6DFFE4FF80FFB2FB5042EDFA4765AB" author="Ratsoavina" box="[1158,1431,654,679]" pageId="11" pageNumber="154" refString="Ratsoavina, F. M., Vences, M. & Louis, E. E. Jr. (2012) Phylogeny and phylogeography of the Malagasy leaf-tailed geckos in the Uroplatus ebenaui group. African Journal of Herpetology, 61, 143 - 158. http: // dx. doi. org / 10.1080 / 21564574.2012.729761" type="journal article" year="2012">
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Ratsoavina
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<emphasis id="7C885E07FF80FFB2FADB42ECFA9965AA" box="[1293,1353,654,678]" italics="true" pageId="11" pageNumber="154">et al.</emphasis>
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(2012
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</bibRefCitation>
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,
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<bibRefCitation id="2A6DFFE4FF80FFB2FF4142D0FF0365C7" author="Ratsoavina" box="[151,211,691,715]" pageId="11" pageNumber="154" refString="Ratsoavina, F. M., Raminosoa, N. R., Louis, E. E. Jr., Raselimanana, A. P., Glaw, F. & Vences, M. (2013) An overview of Madagascar's leaf tailed geckos (genus Uroplatus): species boundaries, candidate species and review of geographical distribution based on molecular data. Salamandra, 49, 115 - 148." type="journal article" year="2013">2013</bibRefCitation>
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), and samples from the locality Fierenana preliminarily as
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<emphasis id="7C885E07FF80FFB2FC7F42D0FBCA65C7" box="[937,1050,691,715]" italics="true" pageId="11" pageNumber="154">Uroplatus</emphasis>
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</taxonomicName>
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||
sp. 1 (
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<bibRefCitation id="2A6DFFE4FF80FFB2FBAC42D1FA4065C7" author="Ratsoavina" box="[1146,1424,690,715]" pageId="11" pageNumber="154" refString="Ratsoavina, F. M., Louis, E. E. Jr., Crottini, A., Randrianiaina, R. D., Glaw, F. & Vences, M. (2011) A new leaf tailed gecko species from northern Madagascar with a preliminary assessment of molecular and morphological variability in the Uroplatus ebenaui group. Zootaxa, 3022, 39 - 57." type="journal article" year="2011">
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||
Ratsoavina
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||
<emphasis id="7C885E07FF80FFB2FAD042D0FA9665C7" box="[1286,1350,691,715]" italics="true" pageId="11" pageNumber="154">et al.</emphasis>
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2011
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||
</bibRefCitation>
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||
).
|
||
<bibRefCitation id="2A6DFFE4FF80FFB2FF4142B5FE7E65E3" box="[151,430,726,751]" pageId="11" pageNumber="154" refString="Ratsoavina, F. M., Raminosoa, N. R., Louis, E. E. Jr., Raselimanana, A. P., Glaw, F. & Vences, M. (2013) An overview of Madagascar's leaf tailed geckos (genus Uroplatus): species boundaries, candidate species and review of geographical distribution based on molecular data. Salamandra, 49, 115 - 148." type="journal article">
|
||
Ratsoavina
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||
<emphasis id="7C885E07FF80FFB2FEC842B4FE8965E2" box="[286,345,726,750]" italics="true" pageId="11" pageNumber="154">et al.</emphasis>
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(2013)
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||
</bibRefCitation>
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||
assigned one additional specimen that lacks precise locality to their
|
||
<taxonomicName id="89FCF996FF80FFB2FB0542B4FA8265E2" box="[1235,1362,726,750]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="154" phylum="Chordata" rank="species" species="ebenaui">
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||
<emphasis id="7C885E07FF80FFB2FB0542B4FA8265E2" box="[1235,1362,726,750]" italics="true" pageId="11" pageNumber="154">U. ebenaui</emphasis>
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</taxonomicName>
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||
[Ca7] (specimen RAX 4012, named
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||
<taxonomicName id="89FCF996FF80FFB2FE3D4298FD8C641F" box="[491,604,763,787]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="154" phylum="Chordata" rank="genus">
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||
<emphasis id="7C885E07FF80FFB2FE3D4298FD8C641F" box="[491,604,763,787]" italics="true" pageId="11" pageNumber="154">Uroplatus</emphasis>
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||
</taxonomicName>
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sp. F by
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<bibRefCitation id="2A6DFFE4FF80FFB2FD144299FC11641F" author="Raxworthy" box="[706,961,762,787]" pageId="11" pageNumber="154" refString="Raxworthy, C. J., Pearson, R. G., Zimkus, B. M., Reddy, S., Deo, A. J., Nussbaum, R. A. & Ingram, C. M. (2008) Continental speciation in the tropics: contrasting biogeographic patterns of divergence in the Uroplatus leaf-tailed gecko radiation of Madagascar. Journal of Zoology, 275, 423 - 440. http: // dx. doi. org / 10.1111 / j. 1469 - 7998.2008.00460. x" type="journal article" year="2008">
|
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Raxworthy
|
||
<emphasis id="7C885E07FF80FFB2FC904298FCAE641F" box="[838,894,763,787]" italics="true" pageId="11" pageNumber="154">et al.</emphasis>
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2008
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</bibRefCitation>
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).
|
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</paragraph>
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</subSubSection>
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<emphasis id="7C885E07FF80FFB2FF11437DFEEB643B" bold="true" box="[199,315,798,823]" pageId="11" pageNumber="154">
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<typeStatus id="91473CB7FF80FFB2FF11437DFEE6643B" box="[199,310,798,823]" pageId="11" pageNumber="154" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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<collectionCode id="28ED1AD0FF80FFB2FE91437CFE51643B" box="[327,385,799,823]" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="11" pageNumber="154">ZSM</collectionCode>
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213/2014 (field number
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<collectionCode id="28ED1AD0FF80FFB2FD78437CFD33643B" box="[686,739,799,823]" httpUri="http://grbio.org/cool/z9c1-g2b0" name="Fusarium Research Center" pageId="11" pageNumber="154">FRC</collectionCode>
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619), adult male with everted hemipenes, collected in the Marohonkona forest of the Fierenana region,
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,
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,
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above sea level, on the night of
|
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<date id="3A42A4D5FF80FFB2FF634304FEEF6473" box="[181,319,871,895]" pageId="11" pageNumber="154" value="2013-05-02">2 May 2013</date>
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, by F. M. Ratsoavina, F. A. Ranjanaharisoa, and A. P. Raselimanana.
|
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</paragraph>
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<paragraph id="4E438215FF80FFB2FF1143E9FC57635B" blockId="11.[151,1437,654,2011]" pageId="11" pageNumber="154">
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<emphasis id="7C885E07FF80FFB2FF1143E9FE9664AF" bold="true" box="[199,326,906,931]" pageId="11" pageNumber="154">
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<typeStatus id="91473CB7FF80FFB2FF1143E9FE9164AF" box="[199,321,906,931]" pageId="11" pageNumber="154" type="paratype">Paratypes</typeStatus>
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.
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</emphasis>
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<collectionCode id="28ED1AD0FF80FFB2FE9A43E8FE5664AF" box="[332,390,907,931]" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="11" pageNumber="154">ZSM</collectionCode>
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212/2014 (
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<collectionCode id="28ED1AD0FF80FFB2FDD343E8FDEC64AF" box="[517,572,907,931]" httpUri="http://grbio.org/cool/z9c1-g2b0" name="Fusarium Research Center" pageId="11" pageNumber="154">FRC</collectionCode>
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618), adult male, with same collecting data as the
|
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<typeStatus id="91473CB7FF80FFB2FBA243E9FB0964AF" box="[1140,1241,906,931]" pageId="11" pageNumber="154" type="holotype">holotype</typeStatus>
|
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;
|
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<collectionCode id="28ED1AD0FF80FFB2FB3343E8FACF64AF" box="[1253,1311,907,931]" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="11" pageNumber="154">ZSM</collectionCode>
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||
1126/2003 (
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||
<collectionCode id="28ED1AD0FF80FFB2FF4843CCFF2164CB" box="[158,241,943,967]" pageId="11" pageNumber="154">FGMV</collectionCode>
|
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2002.3097), adult male, collected by local collectors in the Fierenana region (no precise locality) in
|
||
<date id="3A42A4D5FF80FFB2FF4143B1FE9064E7" box="[151,320,978,1003]" pageId="11" pageNumber="154" value="2003-02">February 2003</date>
|
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;
|
||
<collectionCode id="28ED1AD0FF80FFB2FE9843B0FE5864E7" box="[334,392,979,1003]" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="11" pageNumber="154">ZSM</collectionCode>
|
||
211/2002 (MV 2001-1403), male, collected by local collectors in the Fierenana region (no precise locality) in
|
||
<date id="3A42A4D5FF80FFB2FEA74395FDF86303" box="[369,552,1014,1039]" pageId="11" pageNumber="154" value="2001-12">December 2001</date>
|
||
; UADBA-R 70849 (
|
||
<collectionCode id="28ED1AD0FF80FFB2FCC64394FC976303" box="[784,839,1015,1039]" httpUri="http://grbio.org/cool/z9c1-g2b0" name="Fusarium Research Center" pageId="11" pageNumber="154">FRC</collectionCode>
|
||
650), adult male, and UADBA-R 70850 (
|
||
<collectionCode id="28ED1AD0FF80FFB2FAF54394FA8A6303" box="[1315,1370,1015,1039]" httpUri="http://grbio.org/cool/z9c1-g2b0" name="Fusarium Research Center" pageId="11" pageNumber="154">FRC</collectionCode>
|
||
651), adult female, collected by F. A. Ranjanaharisoa in 2012 at a site in the forest corridor between Fierenana and Ambatovy,
|
||
<geoCoordinate id="2BC8E4D2FF80FFB2FECF445CFE13635B" box="[281,451,1087,1111]" direction="south" orientation="latitude" pageId="11" pageNumber="154" precision="1" value="-18.79686">18° 47' 48.7'' S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="2BC8E4D2FF80FFB2FE19445CFDAA635A" box="[463,634,1086,1111]" direction="east" orientation="longitude" pageId="11" pageNumber="154" precision="1" value="48.371166">48° 22' 16.2'' E</geoCoordinate>
|
||
,
|
||
<quantity id="89042FF0FF80FFB2FD50445DFD1D635A" box="[646,717,1086,1111]" metricMagnitude="2" metricUnit="m" metricValue="9.62" pageId="11" pageNumber="154" unit="m" value="962.0">962 m</quantity>
|
||
above sea level.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="06E6D19EFF80FFB2FF114401FC1A617F" pageId="11" pageNumber="154" type="diagnosis">
|
||
<paragraph id="4E438215FF80FFB2FF114401FB5F625F" blockId="11.[151,1437,654,2011]" pageId="11" pageNumber="154">
|
||
<emphasis id="7C885E07FF80FFB2FF114401FE916377" bold="true" box="[199,321,1122,1147]" pageId="11" pageNumber="154">Diagnosis.</emphasis>
|
||
<taxonomicName id="89FCF996FF80FFB2FE9B4400FE2E6377" box="[333,510,1123,1147]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="154" phylum="Chordata" rank="species" species="fiera" status="sp. nov.">
|
||
<emphasis id="7C885E07FF80FFB2FE9B4400FE2E6377" box="[333,510,1123,1147]" italics="true" pageId="11" pageNumber="154">Uroplatus fiera</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="67BBE37CFF80FFB2FDDC4401FDB86377" box="[522,616,1122,1147]" pageId="11" pageNumber="154" rank="species">
|
||
<emphasis id="7C885E07FF80FFB2FDDC4401FDB86377" bold="true" box="[522,616,1122,1147]" pageId="11" pageNumber="154">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is included in the
|
||
<taxonomicName id="89FCF996FF80FFB2FC984400FBF36377" box="[846,1059,1123,1147]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="154" phylum="Chordata" rank="species" species="ebenaui">
|
||
<emphasis id="7C885E07FF80FFB2FC984400FBF36377" box="[846,1059,1123,1147]" italics="true" pageId="11" pageNumber="154">Uroplatus ebenaui</emphasis>
|
||
</taxonomicName>
|
||
group of small-sized leaf-tailed geckos due to its triangular head with supraocular spines, laterally compressed body, and short tail. It differs from all members of the
|
||
<taxonomicName id="89FCF996FF80FFB2FEA644C8FDD563CF" box="[368,517,1195,1219]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="154" phylum="Chordata" rank="species" species="fimbriatus">
|
||
<emphasis id="7C885E07FF80FFB2FEA644C8FDD563CF" box="[368,517,1195,1219]" italics="true" pageId="11" pageNumber="154">U. fimbriatus</emphasis>
|
||
</taxonomicName>
|
||
group (
|
||
<taxonomicName id="89FCF996FF80FFB2FD8844C8FD2463CF" box="[606,756,1195,1219]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="154" phylum="Chordata" rank="species" species="fimbriatus">
|
||
<emphasis id="7C885E07FF80FFB2FD8844C8FD2463CF" box="[606,756,1195,1219]" italics="true" pageId="11" pageNumber="154">U. fimbriatus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="89FCF996FF80FFB2FCD444C8FC4263CE" box="[770,914,1195,1218]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="154" phylum="Chordata" rank="species" species="giganteus">
|
||
<emphasis id="7C885E07FF80FFB2FCD444C8FC4263CE" box="[770,914,1195,1218]" italics="true" pageId="11" pageNumber="154">U. giganteus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="89FCF996FF80FFB2FC7644C8FBC363CF" box="[928,1043,1195,1219]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="154" phylum="Chordata" rank="species" species="henkeli">
|
||
<emphasis id="7C885E07FF80FFB2FC7644C8FBC363CF" box="[928,1043,1195,1219]" italics="true" pageId="11" pageNumber="154">U. henkeli</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="89FCF996FF80FFB2FBF744C8FB4763CF" box="[1057,1175,1195,1219]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="154" phylum="Chordata" rank="species" species="sikorae">
|
||
<emphasis id="7C885E07FF80FFB2FBF744C8FB4763CF" box="[1057,1175,1195,1219]" italics="true" pageId="11" pageNumber="154">U. sikorae</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="89FCF996FF80FFB2FB1B44C8FA9163CE" box="[1229,1345,1195,1218]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="154" phylum="Chordata" rank="species" species="sameiti">
|
||
<emphasis id="7C885E07FF80FFB2FB1B44C8FA9163CE" box="[1229,1345,1195,1218]" italics="true" pageId="11" pageNumber="154">U. sameiti</emphasis>
|
||
</taxonomicName>
|
||
) and
|
||
<taxonomicName id="89FCF996FF80FFB2FA5744C8FF2063EA" class="Reptilia" family="Gekkonidae" genus="Uroplatus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="154" phylum="Chordata" rank="species" species="lineatus">
|
||
<emphasis id="7C885E07FF80FFB2FA5744C8FF2063EA" italics="true" pageId="11" pageNumber="154">U. lineatus</emphasis>
|
||
</taxonomicName>
|
||
by its much smaller size (adult SVL
|
||
<quantity id="89042FF0FF80FFB2FD4C44ADFCEE63EA" box="[666,830,1230,1255]" metricMagnitude="-2" metricUnit="m" metricValue="6.154999999999999" metricValueMax="6.72" metricValueMin="5.59" pageId="11" pageNumber="154" unit="mm" value="61.55" valueMax="67.2" valueMin="55.9">55.9–67.2 mm</quantity>
|
||
versus at least
|
||
<quantity id="89042FF0FF80FFB2FC3844ADFB9263EB" box="[1006,1090,1230,1255]" metricMagnitude="-2" metricUnit="m" metricValue="8.5" pageId="11" pageNumber="154" unit="mm" value="85.0">85 mm</quantity>
|
||
), lack of lateral integumentary fringing on any part of the body, and lateral compression of the body (versus depressed body shape with lateral integumentary fringes), and a relatively much shorter tail. The evident triangular head, rather smooth skin and short tail distinguish
|
||
<taxonomicName id="89FCF996FF80FFB2FE924558FE4B625F" box="[324,411,1339,1363]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="154" phylum="Chordata" rank="species" species="fiera">
|
||
<emphasis id="7C885E07FF80FFB2FE924558FE4B625F" box="[324,411,1339,1363]" italics="true" pageId="11" pageNumber="154">U. fiera</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<emphasis id="7C885E07FF80FFB2FE094558FC17625F" box="[479,967,1339,1363]" italics="true" pageId="11" pageNumber="154">
|
||
<taxonomicName id="89FCF996FF80FFB2FE094558FDB1625F" box="[479,609,1339,1363]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="154" phylum="Chordata" rank="species" species="alluaudi">U. alluaudi</taxonomicName>
|
||
,
|
||
<taxonomicName id="89FCF996FF80FFB2FDBB4558FD2D625F" box="[621,765,1339,1363]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="154" phylum="Chordata" rank="species" species="guentheri">U. guentheri</taxonomicName>
|
||
,
|
||
<taxonomicName id="89FCF996FF80FFB2FCDE4558FC12625F" box="[776,962,1339,1363]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="154" phylum="Chordata" rank="species" species="pietschmanni">U. pietschmanni</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and
|
||
<emphasis id="7C885E07FF80FFB2FC2B4558FB5F625F" box="[1021,1167,1339,1363]" italics="true" pageId="11" pageNumber="154">
|
||
<taxonomicName id="89FCF996FF80FFB2FC2B4558FB5B625F" box="[1021,1163,1339,1363]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="154" phylum="Chordata" rank="species" species="malahelo">U. malahelo</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="4E438215FF80FFB2FF41453DFC1A617F" blockId="11.[151,1437,654,2011]" pageId="11" pageNumber="154">
|
||
Within the
|
||
<taxonomicName id="89FCF996FF80FFB2FECD453CFE4A627A" box="[283,410,1374,1398]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="154" phylum="Chordata" rank="species" species="ebenaui">
|
||
<emphasis id="7C885E07FF80FFB2FECD453CFE4A627A" box="[283,410,1374,1398]" italics="true" pageId="11" pageNumber="154">U. ebenaui</emphasis>
|
||
</taxonomicName>
|
||
group, the new species differs from
|
||
<taxonomicName id="89FCF996FF80FFB2FC9A453CFC1C627A" box="[844,972,1374,1398]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="154" phylum="Chordata" rank="species" species="malama">
|
||
<emphasis id="7C885E07FF80FFB2FC9A453CFC1C627A" box="[844,972,1374,1398]" italics="true" pageId="11" pageNumber="154">U. malama</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="89FCF996FF80FFB2FC08453CFB42627A" box="[990,1170,1374,1398]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="154" phylum="Chordata" rank="species" species="phantasticus">
|
||
<emphasis id="7C885E07FF80FFB2FC08453CFB42627A" box="[990,1170,1374,1398]" italics="true" pageId="11" pageNumber="154">U. phantasticus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="89FCF996FF80FFB2FB19453CFA8E627B" box="[1231,1374,1375,1399]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="154" phylum="Chordata" rank="species" species="finiavana">
|
||
<emphasis id="7C885E07FF80FFB2FB19453CFA8E627B" box="[1231,1374,1375,1399]" italics="true" pageId="11" pageNumber="154">U. finiavana</emphasis>
|
||
</taxonomicName>
|
||
by a shorter (adult TAL/SVL 0.28–0.34 versus
|
||
<quantity id="89042FF0FF80FFB2FDA045E0FD156297" box="[630,709,1410,1435]" metricMagnitude="-2" metricUnit="m" metricValue="1.8288" pageId="11" pageNumber="154" unit="in" value="0.72">0.72 in</quantity>
|
||
<taxonomicName id="89FCF996FF80FFB2FD1B45E0FC9A6297" box="[717,842,1411,1435]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="154" phylum="Chordata" rank="species" species="malama">
|
||
<emphasis id="7C885E07FF80FFB2FD1B45E0FC9A6297" box="[717,842,1411,1435]" italics="true" pageId="11" pageNumber="154">U. malama</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<quantity id="89042FF0FF80FFB2FC8F45E1FC376297" box="[857,999,1410,1435]" metricMagnitude="-2" metricUnit="m" metricValue="1.7526" metricValueMax="1.9304" metricValueMin="1.5748" pageId="11" pageNumber="154" unit="in" value="0.69" valueMax="0.76" valueMin="0.62">0.62–0.76 in</quantity>
|
||
<emphasis id="7C885E07FF80FFB2FC3945E0FB776297" box="[1007,1191,1411,1435]" italics="true" pageId="11" pageNumber="154">
|
||
<taxonomicName id="89FCF996FF80FFB2FC3945E0FB726297" box="[1007,1186,1411,1435]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="154" phylum="Chordata" rank="species" species="phantasticus">U. phantasticus</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
<quantity id="89042FF0FF80FFB2FB7945E0FB2E6297" box="[1199,1278,1410,1435]" metricMagnitude="-2" metricUnit="m" metricValue="1.0668" pageId="11" pageNumber="154" unit="in" value="0.42">0.42 in</quantity>
|
||
<taxonomicName id="89FCF996FF80FFB2FAD345E0FA436297" box="[1285,1427,1411,1435]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="154" phylum="Chordata" rank="species" species="finiavana">
|
||
<emphasis id="7C885E07FF80FFB2FAD345E0FA436297" box="[1285,1427,1411,1435]" italics="true" pageId="11" pageNumber="154">U. finiavana</emphasis>
|
||
</taxonomicName>
|
||
) and narrower tail (TAW/SVL 0.06–0.07 versus
|
||
<quantity id="89042FF0FF80FFB2FD7B45C4FD2B62B3" box="[685,763,1446,1471]" metricMagnitude="-3" metricUnit="m" metricValue="6.096" pageId="11" pageNumber="154" unit="in" value="0.24">0.24 in</quantity>
|
||
<taxonomicName id="89FCF996FF80FFB2FCD445C4FCAE62B2" box="[770,894,1446,1470]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="154" phylum="Chordata" rank="species" species="malama">
|
||
<emphasis id="7C885E07FF80FFB2FCD445C4FCAE62B2" box="[770,894,1446,1470]" italics="true" pageId="11" pageNumber="154">U. malama</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<quantity id="89042FF0FF80FFB2FC5A45C5FBCA62B3" box="[908,1050,1446,1471]" metricMagnitude="-3" metricUnit="m" metricValue="4.572" metricValueMax="5.08" metricValueMin="4.064" pageId="11" pageNumber="154" unit="in" value="0.18" valueMax="0.2" valueMin="0.16">0.16–0.20 in</quantity>
|
||
<emphasis id="7C885E07FF80FFB2FBF745C4FB0862B2" box="[1057,1240,1446,1470]" italics="true" pageId="11" pageNumber="154">
|
||
<taxonomicName id="89FCF996FF80FFB2FBF745C4FB0462B2" box="[1057,1236,1446,1470]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="154" phylum="Chordata" rank="species" species="phantasticus">U. phantasticus</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and
|
||
<quantity id="89042FF0FF80FFB2FAD945C5FA4C62B3" box="[1295,1436,1446,1471]" metricMagnitude="-3" metricUnit="m" metricValue="3.81" metricValueMax="4.064" metricValueMin="3.556" pageId="11" pageNumber="154" unit="in" value="0.15" valueMax="0.16" valueMin="0.14">0.14–0.16 in</quantity>
|
||
<taxonomicName id="89FCF996FF80FFB2FF4145A8FEF462EF" box="[151,292,1483,1507]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="154" phylum="Chordata" rank="species" species="finiavana">
|
||
<emphasis id="7C885E07FF80FFB2FF4145A8FEF462EF" box="[151,292,1483,1507]" italics="true" pageId="11" pageNumber="154">U. finiavana</emphasis>
|
||
</taxonomicName>
|
||
). It is further distinguished from
|
||
<emphasis id="7C885E07FF80FFB2FD4945A8FC0462EF" box="[671,980,1483,1507]" italics="true" pageId="11" pageNumber="154">
|
||
<taxonomicName id="89FCF996FF80FFB2FD4945A8FCF362EF" box="[671,803,1483,1507]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="154" phylum="Chordata" rank="species" species="malama">U. malama</taxonomicName>
|
||
,
|
||
<taxonomicName id="89FCF996FF80FFB2FCF145A8FC0462EF" box="[807,980,1483,1507]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="154" phylum="Chordata" rank="species" species="phantasticus">U. phantasticus</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="89FCF996FF80FFB2FBDA45A8FB5862EF" box="[1036,1160,1483,1507]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="154" phylum="Chordata" rank="species" species="ebenaui">
|
||
<emphasis id="7C885E07FF80FFB2FBDA45A8FB5862EF" box="[1036,1160,1483,1507]" italics="true" pageId="11" pageNumber="154">U. ebenaui</emphasis>
|
||
</taxonomicName>
|
||
by its unpigmented oral mucosa (versus blackish pigmented oral mucosa). Furthermore,
|
||
<taxonomicName id="89FCF996FF80FFB2FCB5458CFC69610B" box="[867,953,1519,1543]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="154" phylum="Chordata" rank="species" species="fiera" status="sp. nov.">
|
||
<emphasis id="7C885E07FF80FFB2FCB5458CFC69610B" box="[867,953,1519,1543]" italics="true" pageId="11" pageNumber="154">U. fiera</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="67BBE37CFF80FFB2FC16458DFBC9610B" box="[960,1049,1518,1543]" pageId="11" pageNumber="154" rank="species">
|
||
<emphasis id="7C885E07FF80FFB2FC16458DFBC9610B" bold="true" box="[960,1049,1518,1543]" pageId="11" pageNumber="154">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="89FCF996FF80FFB2FB7A458CFAF7610A" box="[1196,1319,1518,1542]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="154" phylum="Chordata" rank="species" species="ebenaui">
|
||
<emphasis id="7C885E07FF80FFB2FB7A458CFAF7610A" box="[1196,1319,1518,1542]" italics="true" pageId="11" pageNumber="154">U. ebenaui</emphasis>
|
||
</taxonomicName>
|
||
by having a greater number of lamellae under the third toe (7–8 versus 5–6), and apparently by larger body size (SVL
|
||
<quantity id="89042FF0FF80FFB2FA804671FF136142" metricMagnitude="-2" metricUnit="m" metricValue="6.7" metricValueMax="7.8" metricValueMin="5.6" pageId="11" pageNumber="154" unit="mm" value="67.0" valueMax="78.0" valueMin="56.0">56–78 mm</quantity>
|
||
versus
|
||
<quantity id="89042FF0FF80FFB2FECF4655FE476143" box="[281,407,1590,1615]" metricMagnitude="-2" metricUnit="m" metricValue="5.65" metricValueMax="6.3" metricValueMin="5.0" pageId="11" pageNumber="154" unit="mm" value="56.5" valueMax="63.0" valueMin="50.0">50–63 mm</quantity>
|
||
). Additionally,
|
||
<taxonomicName id="89FCF996FF80FFB2FD954654FD4B6143" box="[579,667,1591,1615]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="154" phylum="Chordata" rank="species" species="fiera" status="sp. nov.">
|
||
<emphasis id="7C885E07FF80FFB2FD954654FD4B6143" box="[579,667,1591,1615]" italics="true" pageId="11" pageNumber="154">U. fiera</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="67BBE37CFF80FFB2FD744655FD2C6143" box="[674,764,1590,1615]" pageId="11" pageNumber="154" rank="species">
|
||
<emphasis id="7C885E07FF80FFB2FD744655FD2C6143" bold="true" box="[674,764,1590,1615]" pageId="11" pageNumber="154">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
differs from all the other nominal species in the
|
||
<taxonomicName id="89FCF996FF80FFB2FAF74654FA4C6142" box="[1313,1436,1590,1614]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="154" phylum="Chordata" rank="species" species="ebenaui">
|
||
<emphasis id="7C885E07FF80FFB2FAF74654FA4C6142" box="[1313,1436,1590,1614]" italics="true" pageId="11" pageNumber="154">U. ebenaui</emphasis>
|
||
</taxonomicName>
|
||
group by its strong genetic divergence (
|
||
<emphasis id="7C885E07FF80FFB2FD854638FDB1617E" box="[595,609,1627,1650]" italics="true" pageId="11" pageNumber="154">p</emphasis>
|
||
-distances of 16S rDNA>8.5%).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="06E6D19EFF80FFB7FF11461DFDAE60CE" lastPageId="14" lastPageNumber="157" pageId="11" pageNumber="154" type="description">
|
||
<paragraph id="4E438215FF80FFB5FF11461DFB4965C1" blockId="11.[151,1437,654,2011]" lastBlockId="12.[151,1436,151,717]" lastPageId="12" lastPageNumber="155" pageId="11" pageNumber="154">
|
||
<emphasis id="7C885E07FF80FFB2FF11461DFDC4619B" bold="true" box="[199,532,1662,1687]" pageId="11" pageNumber="154">
|
||
Description of the
|
||
<typeStatus id="91473CB7FF80FFB2FE70461DFDDF619B" box="[422,527,1662,1687]" pageId="11" pageNumber="154" type="holotype">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Adult male in good condition with intact tail and everted hemipenes. SVL
|
||
<quantity id="89042FF0FF80FFB2FABD461DFF1561B6" metricMagnitude="-2" metricUnit="m" metricValue="6.62" pageId="11" pageNumber="154" unit="mm" value="66.2">66.2 mm</quantity>
|
||
, tail length
|
||
<quantity id="89042FF0FF80FFB2FE9B46C0FE6461B6" box="[333,436,1699,1723]" metricMagnitude="-2" metricUnit="m" metricValue="1.9899999999999998" pageId="11" pageNumber="154" unit="mm" value="19.9">19.9 mm</quantity>
|
||
, maximum tail width
|
||
<quantity id="89042FF0FF80FFB2FD6646C0FCDA61B6" box="[688,778,1699,1723]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="11" pageNumber="154" unit="mm" value="3.8">3.8 mm</quantity>
|
||
, for further measurements see Table 1. Head triangular in dorsal view, postorbital region
|
||
<quantity id="89042FF0FF80FFB2FE2A46A5FD8761D2" box="[508,599,1734,1759]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="11" pageNumber="154" unit="mm" value="6.8">6.8 mm</quantity>
|
||
; snout length
|
||
<quantity id="89042FF0FF80FFB2FD2C46A4FC8561D2" box="[762,853,1735,1759]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" pageId="11" pageNumber="154" unit="mm" value="7.3">7.3 mm</quantity>
|
||
; canthus rostralis indistinct; snout sloping strongly and continuously downward anteriorly; snout depressed, short (1.2 times longer than eye diameter); eyes large (eye diameter
|
||
<quantity id="89042FF0FF80FFB2FED7476DFE8D602B" box="[257,349,1806,1831]" metricMagnitude="-3" metricUnit="m" metricValue="6.1" pageId="11" pageNumber="154" unit="mm" value="6.1">6.1 mm</quantity>
|
||
), bulging slightly above dorsal surface of cranium, directed laterally, pupil vertical with crenate borders; ear opening very small (horizontal diameter 1.0–
|
||
<quantity id="89042FF0FF80FFB2FCE14750FC426047" box="[823,914,1843,1867]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="11" pageNumber="154" unit="mm" value="1.1">1.1 mm</quantity>
|
||
), its opening facing posterolaterally, but also posteroventrally (ear opening clearly visible in ventral view but not in dorsal view); nostrils laterally oriented; body somewhat laterally compressed, without lateral fringes; limbs well developed, without fringes, forelimb reaches beyond tip of snout when adpressed forward and almost to the groin when adpressed backwards along body (forelimb length/axilla-groin distance 29.1/
|
||
<quantity id="89042FF0FF80FFB2FD5047A1FD3F60D6" box="[646,751,1986,2011]" metricMagnitude="-2" metricUnit="m" metricValue="2.57" pageId="11" pageNumber="154" unit="mm" value="25.7">25.7 mm</quantity>
|
||
=113%), hind limb reaches beyond axilla when adpressed forward along body (hind limb length/axilla-groin distance 36.3/
|
||
<quantity id="89042FF0FF87FFB5FCA040F4FC0D67A3" box="[886,989,151,176]" metricMagnitude="-2" metricUnit="m" metricValue="3.4899999999999998" pageId="12" pageNumber="155" unit="mm" value="34.9">34.9 mm</quantity>
|
||
= 104%); tail length 30% of snout-vent length, membranous borders of the tail narrow and completely absent from the distal tip of the tail, tail with 6 lateral spine-like indentations. Nares separated from each other by at least six small granular scales, from the first supralabial by one scale, and from the rostral scale by one scale; first supralabial taller than the others; rostral entire, much wider than tall; mental scale very small, not differentiated from infralabial scales (total series of infralabials on one side of the head, without mental scale and counting tiny scales at the jaw commissure, equals 28; supralabials yield the same count of 28 on one side); no enlarged postmental scales or chin shields; dorsal and ventral scales of head, neck, body, limbs, and tail small, granular, juxtaposed and largely of uniform size, except for the irregular lines on the head and body which consist of series of slightly enlarged scales. Two curved lines (rows of slightly enlarged scales) extending from the posterolateral parts of the head (nuchal region) converge on the neck forming a V-shaped pattern (neck triangular line). A similar, curved, moderately distinct and posteriorly directed line (also formed by a row of slightly enlarged scales) is present between the eyes and connects the supraciliary spines. Three additional transverse lines, consisting of slightly enlarged scales, are recognizable on the frontal region of the head. Several spines on the posterior part of the head (ca. 16), on hind limbs (ca. 11 per limb), two spines on knee, spines absent from elbow and forelimb; a prominent pointed flap on the posterior portion of each upper eyelid; upper eyelid becomes broader as it approaches the parietal region of the head.
|
||
</paragraph>
|
||
<paragraph id="4E438215FF87FFB5FF414600FD316196" blockId="12.[151,1435,1635,1690]" pageId="12" pageNumber="155">
|
||
<emphasis id="7C885E07FF87FFB5FF414600FEC56174" bold="true" box="[151,277,1635,1657]" pageId="12" pageNumber="155">FIGURE 7.</emphasis>
|
||
Photos of the male
|
||
<typeStatus id="91473CB7FF87FFB5FE3D4600FD946175" box="[491,580,1635,1657]" pageId="12" pageNumber="155" type="holotype">holotype</typeStatus>
|
||
ZSM 213/2014 (FRC 619) of
|
||
<taxonomicName id="89FCF996FF87FFB5FC5F4600FBF86174" box="[905,1064,1635,1656]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="155" phylum="Chordata" rank="species" species="fiera" status="sp. nov.">
|
||
<emphasis id="7C885E07FF87FFB5FC5F4600FBF86174" box="[905,1064,1635,1656]" italics="true" pageId="12" pageNumber="155">Uroplatus fiera</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="67BBE37CFF87FFB5FBE64600FB536174" box="[1072,1155,1635,1656]" pageId="12" pageNumber="155" rank="species">
|
||
<emphasis id="7C885E07FF87FFB5FBE64600FB536174" bold="true" box="[1072,1155,1635,1656]" pageId="12" pageNumber="155">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
in life from Marohonkona forest near Fierenana, taken in the night of
|
||
<date id="3A42A4D5FF87FFB5FD9D46E6FD0D6196" box="[587,733,1665,1690]" pageId="12" pageNumber="155" value="2013-05-02">May 2nd, 2013</date>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="4E438215FF87FFB7FF1146A5FD8A62E3" blockId="12.[151,1437,1734,2011]" lastBlockId="14.[151,1437,1350,2023]" lastPageId="14" lastPageNumber="157" pageId="12" pageNumber="155">
|
||
<emphasis id="7C885E07FF87FFB5FF1146A5FE9861D3" bold="true" box="[199,328,1734,1759]" pageId="12" pageNumber="155">Coloration</emphasis>
|
||
(
|
||
<figureCitation id="D6C79E90FF87FFB5FE8E46A5FE4E61D3" box="[344,414,1734,1759]" captionStart="FIGURE 8" captionStartId="13.[151,250,1877,1899]" captionTargetBox="[196,1381,196,1837]" captionTargetId="figure@13.[196,1383,196,1838]" captionTargetPageId="13" captionText="FIGURE 8. Preserved holotype (ZSM 213 / 2014 - FRC 619) of Uroplatus fiera sp. nov." httpUri="https://zenodo.org/record/232801/files/figure.png" pageId="12" pageNumber="155">Fig. 8</figureCitation>
|
||
). After one year of preservation in 70% ethanol the colour pattern remains the same as in the living animal (Fig. 7) but its vivacity and intensity have faded. All dorsal surfaces are beige to light brown except on the dorsal sides of the head, neck, forelimbs, in the cloacal region, and on the hind limbs, where the colour is darker, showing an irregular light pattern. The tail is uniformly light brown mottled with few dark spots. The mottling of the body is configured as a series of fine posteriorly directed markings along the dorsum. Two whitish spots are present below each eye. The chin is darker than the throat, which is beige with a distinct blackish wide V-marking, the arms of which merge to form an indistinct dark longitudinal line that fades on the throat. The venter is light brown with irregular dots, which become more numerous towards the posterior half of the body, where a distinct dark line delimits the ventral and the dorsal side of the body. The lower hind limbs and feet are slightly darker than other ventral surfaces. The postpygal portion of the tail is marked with a whitish spot. A dorsolateral dark line runs from each forelimb insertion backwards, both lines joining at the tail insertion; just above this dark line four obvious spines are visible on each side of the body. The oral mucosa is unpigmented (whitish in preservative, pinkish in life).
|
||
</paragraph>
|
||
<caption id="1A83D29DFF86FFB4FF414736FBC26066" box="[151,1042,1877,1899]" httpUri="https://zenodo.org/record/232801/files/figure.png" pageId="13" pageNumber="156" targetBox="[196,1381,196,1837]" targetPageId="13">
|
||
<paragraph id="4E438215FF86FFB4FF414736FBC26066" blockId="13.[151,1042,1877,1899]" box="[151,1042,1877,1899]" pageId="13" pageNumber="156">
|
||
<emphasis id="7C885E07FF86FFB4FF414736FEC26066" bold="true" box="[151,274,1877,1899]" pageId="13" pageNumber="156">FIGURE 8.</emphasis>
|
||
Preserved holotype (ZSM 213/2014 - FRC 619) of
|
||
<taxonomicName id="89FCF996FF86FFB4FCF64736FC6C6066" box="[800,956,1877,1898]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="156" phylum="Chordata" rank="species" species="fiera" status="sp. nov.">
|
||
<emphasis id="7C885E07FF86FFB4FCF64736FC6C6066" box="[800,956,1877,1898]" italics="true" pageId="13" pageNumber="156">Uroplatus fiera</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="67BBE37CFF86FFB4FC144736FBC26066" box="[962,1042,1877,1898]" pageId="13" pageNumber="156" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="1A83D29DFF85FFB7FF414486FDE16216" httpUri="https://zenodo.org/record/232802/files/figure.png" pageId="14" pageNumber="157" targetBox="[269,1333,198,1235]" targetPageId="14">
|
||
<paragraph id="4E438215FF85FFB7FF414486FDE16216" blockId="14.[151,1435,1253,1306]" pageId="14" pageNumber="157">
|
||
<emphasis id="7C885E07FF85FFB7FF414486FEC463F6" bold="true" box="[151,276,1253,1275]" pageId="14" pageNumber="157">FIGURE 9.</emphasis>
|
||
Photos of everted hemipenes of two males of
|
||
<taxonomicName id="89FCF996FF85FFB7FD224486FC4163F6" box="[756,913,1253,1274]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="157" phylum="Chordata" rank="species" species="fiera">
|
||
<emphasis id="7C885E07FF85FFB7FD224486FC4163F6" box="[756,913,1253,1274]" italics="true" pageId="14" pageNumber="157">Uroplatus fiera</emphasis>
|
||
</taxonomicName>
|
||
: holotype ZSM 213/2014 (FRC 619) and paratype ZSM 212/2014 (FRC 618). Not to scale.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="4E438215FF85FFB7FF114599FDE06003" blockId="14.[151,1437,1350,2023]" pageId="14" pageNumber="157">
|
||
<emphasis id="7C885E07FF85FFB7FF114599FEEB611F" bold="true" box="[199,315,1530,1555]" pageId="14" pageNumber="157">Variation.</emphasis>
|
||
In general, the
|
||
<typeStatus id="91473CB7FF85FFB7FE3B4598FD89611F" box="[493,601,1531,1555]" pageId="14" pageNumber="157" type="paratype">paratypes</typeStatus>
|
||
agree well with the
|
||
<typeStatus id="91473CB7FF85FFB7FCEA4599FC4E611F" box="[828,926,1530,1555]" pageId="14" pageNumber="157" type="holotype">holotype</typeStatus>
|
||
in morphology. For measurements, see Table 1. The number of head spines varies from 4 (FRC 651) to 25 (ZSM 211/2002). FRC 650 also has spines on elbow (n=1) and forelimb (n=4). The oral mucosa is unpigmented in all specimens. The number of recognizable transverse lines across the head (in addition to the one between supraciliaries) varies from zero (ZSM 211/2002) to three (in various specimens). FRC 650 is the largest specimens (SVL 78.0 mm; Table 1). The colour pattern is highly variable. FRC 650 is rather strongly patterned including an isolated small dark brown patch on the neck and a second one on the dorsum. Other specimens such as ZSM 212/2014 are dorsally almost uniform pale brown and ventrally with very limited mottling.
|
||
</paragraph>
|
||
<paragraph id="4E438215FF85FFB7FF114779FDAE60CE" blockId="14.[151,1437,1350,2023]" pageId="14" pageNumber="157">
|
||
<emphasis id="7C885E07FF85FFB7FF114779FE14603F" bold="true" box="[199,452,1818,1843]" pageId="14" pageNumber="157">Hemipenis structure.</emphasis>
|
||
The everted hemipenes of the
|
||
<typeStatus id="91473CB7FF85FFB7FCF24779FC56603F" box="[804,902,1818,1843]" pageId="14" pageNumber="157" type="holotype">holotype</typeStatus>
|
||
(ZSM 213/2014) and one
|
||
<typeStatus id="91473CB7FF85FFB7FB614778FAC7603F" box="[1207,1303,1819,1843]" pageId="14" pageNumber="157" type="paratype">paratype</typeStatus>
|
||
(ZSM 212/ 2014) were examined (
|
||
<figureCitation id="D6C79E90FF85FFB7FE4C475DFE0E605B" box="[410,478,1854,1879]" captionStart="FIGURE 9" captionStartId="14.[151,250,1253,1275]" captionTargetBox="[269,1333,198,1235]" captionTargetId="figure@14.[269,1334,198,1235]" captionTargetPageId="14" captionText="FIGURE 9. Photos of everted hemipenes of two males of Uroplatus fiera: holotype ZSM 213 / 2014 (FRC 619) and paratype ZSM 212 / 2014 (FRC 618). Not to scale." httpUri="https://zenodo.org/record/232802/files/figure.png" pageId="14" pageNumber="157">Fig. 9</figureCitation>
|
||
). In both specimens, and as in other species of the
|
||
<taxonomicName id="89FCF996FF85FFB7FBCA4723FB47605B" box="[1052,1175,1855,1879]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="157" phylum="Chordata" rank="species" species="ebenaui">
|
||
<emphasis id="7C885E07FF85FFB7FBCA4723FB47605B" box="[1052,1175,1855,1879]" italics="true" pageId="14" pageNumber="157">U. ebenaui</emphasis>
|
||
</taxonomicName>
|
||
group, each hemipenis has two lobes that become more obvious towards the apical region. The calyx displays a protuberance with a honeycomb appearance, especially on the asulcal side. The area of the sulcus spermaticus is smooth. Each lobe has a dense field of pointed papillae at its apex.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="06E6D19EFF85FFB6FF1147ADFDED67F4" lastPageId="15" lastPageNumber="158" pageId="14" pageNumber="157" type="etymology">
|
||
<paragraph id="4E438215FF85FFB6FF1147ADFDED67F4" blockId="14.[151,1437,1350,2023]" lastBlockId="15.[151,1437,151,429]" lastPageId="15" lastPageNumber="158" pageId="14" pageNumber="157">
|
||
<emphasis id="7C885E07FF85FFB7FF1147ADFE9E60EB" bold="true" box="[199,334,1998,2023]" pageId="14" pageNumber="157">Etymology.</emphasis>
|
||
The species name
|
||
<taxonomicName id="89FCF996FF85FFB7FDF747ACFD8660EB" box="[545,598,1999,2023]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="157" phylum="Chordata" rank="species" species="fiera">
|
||
<emphasis id="7C885E07FF85FFB7FDF747ACFD8660EB" box="[545,598,1999,2023]" italics="true" pageId="14" pageNumber="157">fiera</emphasis>
|
||
</taxonomicName>
|
||
is used as noun in apposition to the generic name. It was chosen because it alludes, on one hand, to an euphonious abbreviation of the species'
|
||
<typeStatus id="91473CB7FF84FFB6FC5040FBFC6667BC" box="[902,950,152,176]" pageId="15" pageNumber="158">type</typeStatus>
|
||
locality Fierenana, and on the other hand it corresponds to the Spanish noun
|
||
<taxonomicName id="89FCF996FF84FFB6FDDF40DFFDEC67D9" box="[521,572,188,213]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="158" phylum="Chordata" rank="species" species="fiera">fiera</taxonomicName>
|
||
= beast, thus making allusion to the devil-like appearance of this species and other small
|
||
<taxonomicName id="89FCF996FF84FFB6FECD4083FE5C67F4" box="[283,396,224,248]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="158" phylum="Chordata" rank="genus">
|
||
<emphasis id="7C885E07FF84FFB6FECD4083FE5C67F4" box="[283,396,224,248]" italics="true" pageId="15" pageNumber="158">Uroplatus</emphasis>
|
||
</taxonomicName>
|
||
in frontal view.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="06E6D19EFF84FFB6FF114167FD5066A1" pageId="15" pageNumber="158" type="distribution">
|
||
<paragraph id="4E438215FF84FFB6FF114167FAA16611" blockId="15.[151,1437,151,429]" box="[199,1393,260,285]" pageId="15" pageNumber="158">
|
||
<emphasis id="7C885E07FF84FFB6FF114167FEB06611" bold="true" box="[199,352,260,285]" pageId="15" pageNumber="158">Distribution.</emphasis>
|
||
Known from the Fierenana and Ambatovy regions, of Northern Central East of
|
||
<collectingCountry id="36EBC285FF84FFB6FB354167FABD6611" box="[1251,1389,260,285]" name="Madagascar" pageId="15" pageNumber="158">Madagascar</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="4E438215FF84FFB6FF114144FD5066A1" blockId="15.[151,1437,151,429]" pageId="15" pageNumber="158">
|
||
<emphasis id="7C885E07FF84FFB6FF114144FE11664C" bold="true" box="[199,449,295,320]" pageId="15" pageNumber="158">Habitat and habits.</emphasis>
|
||
The
|
||
<typeStatus id="91473CB7FF84FFB6FDDA4144FDBE664C" box="[524,622,295,320]" pageId="15" pageNumber="158" type="holotype">holotype</typeStatus>
|
||
was collected in the Fierenana region in a primary forest called Marohonkona. Part of the Northern Central East of
|
||
<collectingCountry id="36EBC285FF84FFB6FD0F412FFCB26669" box="[729,866,332,357]" name="Madagascar" pageId="15" pageNumber="158">Madagascar</collectingCountry>
|
||
, the
|
||
<typeStatus id="91473CB7FF84FFB6FC40412EFC166669" box="[918,966,333,357]" pageId="15" pageNumber="158">type</typeStatus>
|
||
locality has a humid tropical climate, and its vegetation at is dominated by moist evergreen forest. The
|
||
<typeStatus id="91473CB7FF84FFB6FCB3410CFC176684" box="[869,967,367,392]" pageId="15" pageNumber="158" type="holotype">holotype</typeStatus>
|
||
was found active during a night search, perched on a fern tree at
|
||
<quantity id="89042FF0FF84FFB6FE7B41F7FE0866A0" box="[429,472,404,428]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="15" pageNumber="158" unit="m" value="2.0">2 m</quantity>
|
||
above ground.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |