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<document id="81FD8E2536540A11FEB294F9C57C7430" ID-CLB-Dataset="28091" ID-DOI="10.11646/zootaxa.4504.1.3" ID-GBIF-Dataset="0b803003-244e-4511-aaf4-5e9c3f863861" ID-ISSN="1175-5326" ID-Zenodo-Dep="2606164" ID-ZooBank="52BA28B6-4280-4BD0-8B38-C794F04E4A66" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553577945224" checkinUser="plazi" docAuthor="Abdalla, Ishtiag H., Deschodt, Christian M., Scholtz, Clarke H. &amp; Sole, Catherine L." docDate="2018" docId="B276DC1EFFBDFFF9FF0DFE52FEE506CD" docLanguage="en" docName="zootaxa.4504.1.3.pdf" docOrigin="Zootaxa 4504 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Macroderes dubius Peringuey 1901" docType="treatment" docVersion="8" lastPageNumber="72" masterDocId="4E4FA466FFA2FFE6FF9AFFF0FFD20545" masterDocTitle="An update to the taxonomy of the genus Macroderes Westwood 1842 (Coleoptera: Scarabaeidae: Scarabaeinae) with descriptions of new species from South Africa" masterLastPageNumber="75" masterPageNumber="41" pageNumber="72" updateTime="1698712718888" updateUser="ExternalLinkService">
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<mods:title id="12286E9AA10500988ED4B02B910DF93D">An update to the taxonomy of the genus Macroderes Westwood 1842 (Coleoptera: Scarabaeidae: Scarabaeinae) with descriptions of new species from South Africa</mods:title>
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<mods:namePart id="D2C2F845B8A174C9FF061FCE9F8CEA32">Abdalla, Ishtiag H.</mods:namePart>
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<mods:namePart id="4C9A881744A6858547C9F87B64F05BF9">Deschodt, Christian M.</mods:namePart>
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<mods:namePart id="A6A458D98DAD4C68F0A1236A71EE7C02">Scholtz, Clarke H.</mods:namePart>
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<mods:namePart id="B922C98E289989806C1DE4947832B9BE">Sole, Catherine L.</mods:namePart>
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<paragraph id="3A606D08FFBDFFF9FF0DFE52FD9F04F9" blockId="31.[151,589,418,444]" box="[151,589,418,444]" pageId="31" pageNumber="72">
<heading id="6128DA64FFBDFFF9FF0DFE52FD9F04F9" bold="true" box="[151,589,418,444]" fontSize="11" level="1" pageId="31" pageNumber="72" reason="1">
<taxonomicName id="FDDF168BFFBDFFF9FF0DFE52FD9F04F9" ID-CoL="3X4TH" authority="Peringuey, 1901" authorityName="Peringuey" authorityYear="1901" box="[151,589,418,444]" class="Insecta" family="Scarabaeidae" genus="Macroderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="72" phylum="Arthropoda" rank="species" species="dubius">
<emphasis id="08ABB11AFFBDFFF9FF0DFE52FD9F04F9" bold="true" box="[151,589,418,444]" pageId="31" pageNumber="72">
<emphasis id="08ABB11AFFBDFFF9FF0DFE52FEAC04F9" bold="true" box="[151,382,418,444]" italics="true" pageId="31" pageNumber="72">Macroderes dubius</emphasis>
<bibRefCitation id="5E4E10F9FFBDFFF9FE1CFE52FD9F04F9" author="Peringuey, L." box="[390,589,418,444]" pageId="31" pageNumber="72" pagination="1 - 563" refId="ref22154" refString="Peringuey, L. (1901) Descriptive catalogue of the Coleoptera of South Africa (Lucanidae and Scarabaeidae). Transactions of the South African Philosophical Society, 12 (1), 1 - 563. https: // doi. org / 10.5962 / bhl. title. 8963" type="journal article" year="1901">Péringuey, 1901</bibRefCitation>
</emphasis>
</taxonomicName>
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</paragraph>
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<subSubSection id="72C53E83FFBDFFF9FF0DFE19FB2404BA" box="[151,1270,489,511]" pageId="31" pageNumber="72" type="reference_group">
<paragraph id="3A606D08FFBDFFF9FF0DFE19FB2404BA" blockId="31.[151,1270,489,511]" box="[151,1270,489,511]" pageId="31" pageNumber="72">
<treatmentCitationGroup id="1ACF4A26FFBDFFF9FF0DFE19FB2404BA" box="[151,1270,489,511]" pageId="31" pageNumber="72">
<taxonomicName id="FDDF168BFFBDFFF9FF0DFE19FDE804BA" ID-CoL="3X4TH" authority="Peringuey, 1901: 301" authorityName="Peringuey" authorityPageNumber="301" authorityYear="1901" box="[151,570,489,511]" class="Insecta" family="Scarabaeidae" genus="Macroderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="72" phylum="Arthropoda" rank="species" species="dubius">
<emphasis id="08ABB11AFFBDFFF9FF0DFE19FE8B04BA" box="[151,345,489,511]" italics="true" pageId="31" pageNumber="72">Macroderes dubius</emphasis>
<treatmentCitation id="BB7E4B19FFBDFFF9FEC5FE19FDE804BA" author="Peringuey, L." box="[351,570,489,511]" page="301" pageId="31" pageNumber="72" year="1901">
<bibRefCitation id="5E4E10F9FFBDFFF9FEC5FE19FDD504BA" author="Peringuey, L." box="[351,519,489,511]" pageId="31" pageNumber="72" pagination="1 - 563" refId="ref22154" refString="Peringuey, L. (1901) Descriptive catalogue of the Coleoptera of South Africa (Lucanidae and Scarabaeidae). Transactions of the South African Philosophical Society, 12 (1), 1 - 563. https: // doi. org / 10.5962 / bhl. title. 8963" type="journal article" year="1901">Péringuey, 1901</bibRefCitation>
: 301
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</taxonomicName>
;
<treatmentCitation id="BB7E4B19FFBDFFF9FDDEFE1AFD2C04BA" author="Janssens, A." box="[580,766,489,511]" page="29" pageId="31" pageNumber="72" year="1939">
<bibRefCitation id="5E4E10F9FFBDFFF9FDDEFE1AFD0A04BA" author="Janssens, A." box="[580,728,489,511]" pageId="31" pageNumber="72" pagination="1 - 104" refId="ref21780" refString="Janssens, A. (1939) Coprini. Exploration du Parc National Albert. Mission G. F. de Witte, 29, 1 - 104." type="journal article" year="1939">Janssens 1939</bibRefCitation>
: 29
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;
<treatmentCitation id="BB7E4B19FFBDFFF9FC93FE19FC1A04BA" author="Ferreira, M." box="[777,968,489,511]" page="320" pageId="31" pageNumber="72" year="1969">
<bibRefCitation id="5E4E10F9FFBDFFF9FC93FE19FC4704BA" author="Ferreira, M." box="[777,917,489,511]" pageId="31" pageNumber="72" pagination="5 - 1088" refId="ref21605" refString="Ferreira, M. (1969) Os escarabideos de Africa (Sul do Saara), I. Revista de Entomologia de Mocambique, 11, 5 - 1088." type="journal article" year="1969">Ferreira 1969</bibRefCitation>
: 320
</treatmentCitation>
;
<treatmentCitation id="BB7E4B19FFBDFFF9FC48FE19FB2304BA" author="Frolov, A. V. &amp; Scholtz, C. H." box="[978,1265,489,511]" page="392" pageId="31" pageNumber="72" year="2005">
<bibRefCitation id="5E4E10F9FFBDFFF9FC48FE19FB6C04BA" author="Frolov, A. V. &amp; Scholtz, C. H." box="[978,1214,489,511]" pageId="31" pageNumber="72" pagination="373 - 393" refId="ref21694" refString="Frolov, A. V. &amp; Scholtz, C. H. (2005) Revision of the southern African genus Macroderes Westwood (Coleoptera: Scarabaeidae: Scarabaeinae). Annales de la Societe Entomologique de France, 40 (3 - 4), 373 - 393. https: // doi. org / 10.1080 / 00379271.2004.10697429" type="journal article" year="2005">Frolov &amp; Scholtz 2005</bibRefCitation>
: 392
</treatmentCitation>
.
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</paragraph>
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<paragraph id="3A606D08FFBDFFF9FF0DFDDCFD7B0700" blockId="31.[151,1436,556,904]" box="[151,681,556,581]" pageId="31" pageNumber="72">
<materialsCitation id="8AB76755FFBDFFF9FF0DFDDCFD770700" ID-GBIF-Occurrence="2028137835" box="[151,677,556,581]" location="Colony" pageId="31" pageNumber="72" specimenCount="1" stateProvince="Western Cape" typeStatus="holotype">
<emphasis id="08ABB11AFFBDFFF9FF0DFDDCFEE70700" bold="true" box="[151,309,556,581]" pageId="31" pageNumber="72">
<typeStatus id="E564D3AAFFBDFFF9FF0DFDDCFF000700" box="[151,210,556,581]" pageId="31" pageNumber="72">Type</typeStatus>
locality.
</emphasis>
<collectingRegion id="F81BA3EAFFBDFFF9FEA6FDDCFEA40701" box="[316,374,556,580]" country="South Africa" name="Western Cape" pageId="31" pageNumber="72">Cape</collectingRegion>
<location id="3F003BD3FFBDFFF9FEE7FDDCFE020700" LSID="urn:lsid:plazi:treatment:B276DC1EFFBDFFF9FF0DFE52FEE506CD:3F003BD3FFBDFFF9FEE7FDDCFE020700" box="[381,464,556,581]" name="Colony" pageId="31" pageNumber="72" stateProvince="Western Cape">Colony</location>
(no exact locality)
</materialsCitation>
.
</paragraph>
<paragraph id="3A606D08FFBDFFF9FF5DFDBFFEC907C8" blockId="31.[151,1436,556,904]" pageId="31" pageNumber="72">
<emphasis id="08ABB11AFFBDFFF9FF5DFDBFFE28072D" bold="true" box="[199,506,591,616]" pageId="31" pageNumber="72">
<typeStatus id="E564D3AAFFBDFFF9FF5DFDBFFED0072D" box="[199,258,591,616]" pageId="31" pageNumber="72">Type</typeStatus>
material examined.
</emphasis>
None. No indication of where the
<typeStatus id="E564D3AAFFBDFFF9FC3FFDA0FC07072D" box="[933,981,592,616]" pageId="31" pageNumber="72">type</typeStatus>
was deposited in Péringueys original description.
</paragraph>
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<subSubSection id="72C53E83FFBDFFF9FF5DFD67FEE506CD" pageId="31" pageNumber="72" type="discussion">
<paragraph id="3A606D08FFBDFFF9FF5DFD67FEE506CD" blockId="31.[151,1436,556,904]" pageId="31" pageNumber="72">
<emphasis id="08ABB11AFFBDFFF9FF5DFD67FEE907F5" bold="true" box="[199,315,663,688]" pageId="31" pageNumber="72">Remarks.</emphasis>
Unfortunately we could not locate the
<typeStatus id="E564D3AAFFBDFFF9FD69FD67FC8707F5" box="[755,853,663,688]" pageId="31" pageNumber="72" type="holotype">holotype</typeStatus>
of this species, but according to
<bibRefCitation id="5E4E10F9FFBDFFF9FB52FD67FF180791" author="Peringuey, L." pageId="31" pageNumber="72" pagination="1 - 563" refId="ref22154" refString="Peringuey, L. (1901) Descriptive catalogue of the Coleoptera of South Africa (Lucanidae and Scarabaeidae). Transactions of the South African Philosophical Society, 12 (1), 1 - 563. https: // doi. org / 10.5962 / bhl. title. 8963" type="journal article" year="1901">Péringueys (1901: 301)</bibRefCitation>
original description, body size is
<quantity id="FD27C0EDFFBDFFF9FDD3FD4CFD4A0791" box="[585,664,700,724]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="31" pageNumber="72" unit="mm" value="11.0">11 mm</quantity>
in length and
<quantity id="FD27C0EDFFBDFFF9FCA0FD4CFC430791" box="[826,913,700,725]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="31" pageNumber="72" unit="mm" value="7.5">7.5 mm</quantity>
in width and it resembles
<taxonomicName id="FDDF168BFFBDFFF9FB26FD4DFA960791" authorityName="Preudhomme de Borre" authorityYear="1880" box="[1212,1348,700,724]" class="Insecta" family="Scarabaeidae" genus="Macroderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="72" phylum="Arthropoda" rank="species" species="politulus">
<emphasis id="08ABB11AFFBDFFF9FB26FD4DFA960791" box="[1212,1348,700,724]" italics="true" pageId="31" pageNumber="72">M. politulus</emphasis>
</taxonomicName>
in size, shape, head, and punctuation of the pronotum.
<bibRefCitation id="5E4E10F9FFBDFFF9FD52FD2FFC4207BD" author="Peringuey, L." box="[712,912,735,760]" pageId="31" pageNumber="72" pagination="1 - 563" refId="ref22154" refString="Peringuey, L. (1901) Descriptive catalogue of the Coleoptera of South Africa (Lucanidae and Scarabaeidae). Transactions of the South African Philosophical Society, 12 (1), 1 - 563. https: // doi. org / 10.5962 / bhl. title. 8963" type="journal article" year="1901">Péringuey (1901)</bibRefCitation>
mentioned that this species differs from
<taxonomicName id="FDDF168BFFBDFFF9FAE7FD11FF2B0659" authorityName="Preudhomme de Borre" authorityYear="1880" class="Insecta" family="Scarabaeidae" genus="Macroderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="72" phylum="Arthropoda" rank="species" species="politulus">
<emphasis id="08ABB11AFFBDFFF9FAE7FD11FF2B0659" italics="true" pageId="31" pageNumber="72">M. politulus</emphasis>
</taxonomicName>
by bearing a more convex and rounded pronotum without any observed excavation antero-laterally. In addition, the species has even elytral intervals covered by finer and shallower punctures than
<taxonomicName id="FDDF168BFFBDFFF9FB2DFCD9FAED0605" authorityName="Preudhomme de Borre" authorityYear="1880" box="[1207,1343,808,832]" class="Insecta" family="Scarabaeidae" genus="Macroderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="72" phylum="Arthropoda" rank="species" species="politulus">
<emphasis id="08ABB11AFFBDFFF9FB2DFCD9FAED0605" box="[1207,1343,808,832]" italics="true" pageId="31" pageNumber="72">M. politulus</emphasis>
</taxonomicName>
, costate and punctate striae. He also observed an unusual elytral shape in the specimen he studied and attributed this to a malformation.
</paragraph>
</subSubSection>
</treatment>
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