treatments-xml/data/B2/76/DC/B276DC1EFFB9FFFAFF0DFD75FC2304E8.xml
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<mods:title id="64062EFCA18DA3774CF5C203D34FCCA4">An update to the taxonomy of the genus Macroderes Westwood 1842 (Coleoptera: Scarabaeidae: Scarabaeinae) with descriptions of new species from South Africa</mods:title>
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<mods:namePart id="B9C57FE64423916DCCA6817B3E3C33A8">Scholtz, Clarke H.</mods:namePart>
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<emphasis id="08ABB11AFFB9FFFDFF0DFD75FCE407DA" bold="true" box="[151,822,645,671]" pageId="27" pageNumber="68">
<taxonomicName id="FDDF168BFFB9FFFDFF0DFD75FD4B07DA" ID-CoL="3X4TZ" authority="Abdalla &amp; Deschodt" authorityName="Abdalla &amp; Deschodt &amp; Scholtz &amp; Sole" authorityYear="2018" box="[151,665,645,671]" class="Insecta" family="Scarabaeidae" genus="Macroderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="68" phylum="Arthropoda" rank="species" species="soleiana" status="sp. nov.">
<emphasis id="08ABB11AFFB9FFFDFF0DFD75FE4007DA" bold="true" box="[151,402,645,671]" italics="true" pageId="27" pageNumber="68">Macroderes soleiana</emphasis>
Abdalla &amp; Deschodt
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,
<taxonomicNameLabel id="13980C61FFB9FFFDFD3CFD75FCE407DA" box="[678,822,645,671]" pageId="27" pageNumber="68" rank="species">new species</taxonomicNameLabel>
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(
<figureCitation id="A2E4718DFFB9FFFDFF05FD57FF2E0785" box="[159,252,679,704]" captionStart="FIGURES 1923" captionStartId="9.[151,264,1928,1951]" captionTargetBox="[411,1172,706,1874]" captionTargetId="figure@9.[337,1297,671,1926]" captionTargetPageId="9" captionText="FIGURES 1923. Habitus of Macroderes species. 19, M. oreatus Abdalla &amp; Deschodt, new species; 20, M. cederbergensis Abdalla &amp; Deschodt, new species; 21, M. soleiana Abdalla &amp; Deschodt, new species; 22, M. tourtosus Abdalla &amp; Scholtz, new species; 23, M. leipoldti Abdalla &amp; Deschodt, new species." httpUri="https://zenodo.org/record/2606176/files/figure.png" pageId="27" pageNumber="68">Figs. 21</figureCitation>
,
<figureCitation id="A2E4718DFFB9FFFDFE92FD58FEF40785" box="[264,294,680,704]" captionStart="FIGURES 2439" captionStartId="10.[151,264,1343,1366]" captionTargetBox="[323,1250,205,1286]" captionTargetId="figure@10.[323,1283,195,1303]" captionTargetPageId="10" captionText="FIGURES 2439. Internal sac of aedeagus (24), aedeagus (25) and internal sac sclerite (26-39) of Macroderes species. 26, M. endroedyi Frolov &amp; Scholtz, 2005; 27, M. amplior Frolov &amp; Scholtz, 2005; 28, M. bias (Olivier, 1789); 29, M. cederbergensis Abdalla &amp; Deschodt, new species; 30, M. cornutus Frolov &amp; Scholtz, 2005; 31, M. foveatus Frolov &amp; Scholtz, 2005; 32, M. gifboomi Abdalla &amp; Scholtz, new species; 33, M. leipoldti Abdalla &amp; Deschodt, new species; 34, M. mutilans Kolbe, 1908; 35, M. namakwanus Frolov &amp; Scholtz, 2005; 36, M. oreatus Abdalla &amp; Deschodt, new species; 37, M. porselinus Abdalla, new species; 38, M. soleiana Abdalla &amp; Deschodt, new species; 39, M. tourtosus Abdalla &amp; Scholtz, new species." httpUri="https://zenodo.org/record/2606178/files/figure.png" pageId="27" pageNumber="68">38</figureCitation>
,
<figureCitation id="A2E4718DFFB9FFFDFEA8FD58FE830785" box="[306,337,680,704]" captionStart="FIGURES 7681" captionStartId="20.[151,264,1637,1660]" captionTargetBox="[171,1423,205,1613]" captionTargetId="figure@20.[171,1423,205,1613]" captionTargetPageId="20" captionText="FIGURES 7681. Punctuation of the base of pronotum of Macroderes species. 76, M. cederbergensis Abdalla &amp; Deschodt, new species; 77, M. gifboomi Abdalla &amp; Scholtz, new species; 78, M. oreatus Abdalla &amp; Deschodt, new species; 79, M. porselinus Abdalla, new species; 80, M. soleiana Abdalla &amp; Deschodt, new species; 81, M. leipoldti Abdalla &amp; Deschodt, new species." httpUri="https://zenodo.org/record/2606198/files/figure.png" pageId="27" pageNumber="68">80</figureCitation>
,
<figureCitation id="A2E4718DFFB9FFFDFEC7FD58FEA90785" box="[349,379,680,704]" captionStart="FIGURES 8893" captionStartId="23.[151,264,1292,1315]" captionTargetBox="[316,1276,183,1264]" captionTargetId="figure@23.[316,1276,183,1264]" captionTargetPageId="23" captionText="FIGURES 8893. Pronotum punctures shape (88, 89) and sculpture of second elytra intervals (9093) of Macroderes species. 88, M. soleiana Abdalla &amp; Deschodt, new species; 89, M. tourtosus Abdalla and Scholtz, new species; 90, M. minutus Frolov and Scholtz, 2005; 91, M. greeni (Kirby, 1818); 92, M. politulus Preudhomme de Borre, 1880; 93, M. gifboomi Abdalla &amp; Scholtz, new species." httpUri="https://zenodo.org/record/2606202/files/figure.png" pageId="27" pageNumber="68">88</figureCitation>
,
<figureCitation id="A2E4718DFFB9FFFDFE1CFD58FE710785" box="[390,419,680,704]" captionStart="FIGURES 9499" captionStartId="24.[151,264,1299,1322]" captionTargetBox="[316,1273,193,1274]" captionTargetId="figure@24.[316,1273,193,1274]" captionTargetPageId="24" captionText="FIGURES 9499. Sculpture of second elytra intervals of Macroderes species. 94, M. cederbergensis Abdalla &amp; Deschodt, new species; 95, M. leipoldti Abdalla &amp; Deschodt, new species; 96, M. oreatus Abdalla &amp; Deschodt, new species; 97, M. porselinus Abdalla, new species; 98, M. soleiana Abdalla &amp; Deschodt, new species; 99, M. tourtosus Abdalla &amp; Scholtz, new species." httpUri="https://zenodo.org/record/2606204/files/figure.png" pageId="27" pageNumber="68">98</figureCitation>
)
</paragraph>
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urn:lsid:zoobank.org:act:
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<taxonomicName id="FDDF168BFFB9FFFDFF0DFCFEFDC80661" ID-CoL="3X4TT" authority="Harold, 1877: 384" authorityName="Harold /" authorityPageNumber="384" authorityYear="1877" box="[151,538,782,804]" class="Insecta" family="Scarabaeidae" genus="Macroderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="68" phylum="Arthropoda" rank="species" species="nitidus">
<emphasis id="08ABB11AFFB9FFFDFF0DFCFEFE880661" box="[151,346,782,804]" italics="true" pageId="27" pageNumber="68">Macroderes nitidus</emphasis>
<treatmentCitation id="BB7E4B19FFB9FFFDFEFBFCFEFDC80661" author="Harold, E. V." box="[353,538,782,804]" page="384" pageId="27" pageNumber="68" year="1877">
<bibRefCitation id="5E4E10F9FFB9FFFDFEFBFCFEFE380661" author="Harold, E. V." box="[353,490,782,804]" pageId="27" pageNumber="68" pagination="97 - 111" refId="ref21755" refString="Harold, E. V. (1877) Coleopterorum species novae. Mittheilungen des Munchener Entomologischen Uereins, 1, 97 - 111." type="journal article" year="1877">Harold, 1877</bibRefCitation>
: 384
</treatmentCitation>
</taxonomicName>
(partim, sensu
<bibRefCitation id="5E4E10F9FFB9FFFDFD2CFCFEFC720661" author="Frolov, A. V. &amp; Scholtz, C. H." box="[694,928,782,804]" pageId="27" pageNumber="68" pagination="373 - 393" refId="ref21694" refString="Frolov, A. V. &amp; Scholtz, C. H. (2005) Revision of the southern African genus Macroderes Westwood (Coleoptera: Scarabaeidae: Scarabaeinae). Annales de la Societe Entomologique de France, 40 (3 - 4), 373 - 393. https: // doi. org / 10.1080 / 00379271.2004.10697429" type="journal article" year="2005">Frolov &amp; Scholtz 2005</bibRefCitation>
)
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</paragraph>
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<emphasis id="08ABB11AFFB9FFFDFF0DFCA1FEE7062F" bold="true" box="[151,309,849,874]" pageId="27" pageNumber="68">Type locality.</emphasis>
<materialsCitation id="8AB76755FFB9FFFDFEA6FCA1FB21062F" ID-GBIF-Occurrence="2028137854" box="[316,1267,849,874]" country="South Africa" latitude="-32.483334" location="Cederberg Range" longLatPrecision="1210" longitude="19.366667" pageId="27" pageNumber="68" specimenCount="1" stateProvince="Western Cape">
<collectingCountry id="42C82D98FFB9FFFDFEA6FCA1FE1D062F" box="[316,463,849,874]" name="South Africa" pageId="27" pageNumber="68">South Africa</collectingCountry>
,
<collectingRegion id="F81BA3EAFFB9FFFDFE40FCA2FD31062F" box="[474,739,849,874]" country="South Africa" name="Western Cape" pageId="27" pageNumber="68">Western Cape Province</collectingRegion>
,
<location id="3F003BD3FFB9FFFDFD74FCA1FC61062F" LSID="urn:lsid:plazi:treatment:B276DC1EFFB9FFFAFF0DFD75FC2304E8:3F003BD3FFB9FFFDFD74FCA1FC61062F" box="[750,947,849,874]" country="South Africa" latitude="-32.483334" longLatPrecision="1210" longitude="19.366667" name="Cederberg Range" pageId="27" pageNumber="68" stateProvince="Western Cape">Cederberg Range</location>
, east track,
<geoCoordinate id="5FEB0BCFFFB9FFFDFBADFCA2FB5D062F" box="[1079,1167,850,874]" direction="south" orientation="latitude" pageId="27" pageNumber="68" precision="925" value="-32.483334">32°29'S</geoCoordinate>
<geoCoordinate id="5FEB0BCFFFB9FFFDFB0CFCA2FB21062F" box="[1174,1267,850,874]" direction="east" orientation="longitude" pageId="27" pageNumber="68" precision="925" value="19.366667">19°22'E</geoCoordinate>
</materialsCitation>
.
</paragraph>
<paragraph id="3A606D08FFB9FFFDFF5DFC85FED100C3" blockId="27.[151,1436,849,2026]" pageId="27" pageNumber="68">
<emphasis id="08ABB11AFFB9FFFDFF5DFC85FEA706CB" bold="true" box="[199,373,885,910]" pageId="27" pageNumber="68">Type material.</emphasis>
<materialsCitation id="8AB76755FFB9FFFDFEE6FC85FAD40693" ID-GBIF-Occurrence="2028137839" collectingDate="1983-08-21" collectionCode="TMSA" collectorName="Penrith" country="South Africa" elevation="800" latitude="-32.483334" location="Cederberg Range" longLatPrecision="1210" longitude="19.366667" pageId="27" pageNumber="68" specimenCount="1" specimenCount-male="1" stateProvince="Western Cape" typeStatus="holotype">
<typeStatus id="E564D3AAFFB9FFFDFEE6FC85FE3506CB" box="[380,487,885,910]" pageId="27" pageNumber="68" type="holotype">Holotype</typeStatus>
,
<specimenCount id="2CD9A681FFB9FFFDFE68FC86FDD506C8" box="[498,519,886,909]" pageId="27" pageNumber="68" type="male"></specimenCount>
(
<collectionCode id="5CCEF5CDFFB9FFFDFD8CFC86FDB906CB" box="[534,619,886,910]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="27" pageNumber="68">TMSA</collectionCode>
): “
<collectingCountry id="42C82D98FFB9FFFDFD1DFC85FCCB06CB" box="[647,793,885,910]" name="South Africa" pageId="27" pageNumber="68">South Africa</collectingCountry>
/
<collectingRegion id="F81BA3EAFFB9FFFDFCB5FC86FBE706CB" box="[815,1077,885,910]" country="South Africa" name="Western Cape" pageId="27" pageNumber="68">Western Cape Province</collectingRegion>
/
<location id="3F003BD3FFB9FFFDFBD0FC85FADF06CB" LSID="urn:lsid:plazi:treatment:B276DC1EFFB9FFFAFF0DFD75FC2304E8:3F003BD3FFB9FFFDFBD0FC85FADF06CB" box="[1098,1293,885,910]" country="South Africa" latitude="-32.483334" longLatPrecision="1210" longitude="19.366667" name="Cederberg Range" pageId="27" pageNumber="68" stateProvince="Western Cape">Cederberg Range</location>
/ east track /
<quantity id="FD27C0EDFFB9FFFDFF0DFC6AFF0D06F4" box="[151,223,922,946]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="27" pageNumber="68" unit="m" value="800.0">
<elevation id="B1F28A3BFFB9FFFDFF0DFC6AFF0D06F4" box="[151,223,922,946]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="27" pageNumber="68" unit="m" value="800.0">800 m</elevation>
</quantity>
/
<geoCoordinate id="5FEB0BCFFFB9FFFDFF6DFC6AFE8C06F7" box="[247,350,922,946]" direction="south" orientation="latitude" pageId="27" pageNumber="68" precision="925" value="-32.483334">32° 29' S</geoCoordinate>
<geoCoordinate id="5FEB0BCFFFB9FFFDFEFDFC6AFE0206F4" box="[359,464,922,946]" direction="east" orientation="longitude" pageId="27" pageNumber="68" precision="925" value="19.366667">19° 22' E</geoCoordinate>
/
<date id="4E614BC8FFB9FFFDFE72FC6AFDA806F7" box="[488,634,922,946]" pageId="27" pageNumber="68" value="1983-08-21">
<collectingDate id="5E25B220FFB9FFFDFE72FC6AFDA806F7" box="[488,634,922,946]" pageId="27" pageNumber="68" value="1983-08-21">21.VIII.1983</collectingDate>
</date>
/ groundtrap with feaces bait / 66 days / EndrödyYounga and
<collectorName id="972A08DEFFB9FFFDFAD1FC69FA4E06F7" box="[1355,1436,921,946]" pageId="27" pageNumber="68">Penrith</collectorName>
leg [printed] // HOLOTYE /
<emphasis id="08ABB11AFFB9FFFDFE44FC4DFD090690" box="[478,731,957,981]" italics="true" pageId="27" pageNumber="68">
<taxonomicName id="FDDF168BFFB9FFFDFE44FC4DFD190690" authorityName="Abdalla &amp; Deschodt &amp; Scholtz &amp; Sole" authorityYear="2018" box="[478,715,957,981]" class="Insecta" family="Scarabaeidae" genus="Macroderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="68" phylum="Arthropoda" rank="species" species="soleiana">Macroderes soleiana</taxonomicName>
/
</emphasis>
Abdalla &amp; Deschodt / 2018 [red label, printed]”
</materialsCitation>
.
<materialsCitation id="8AB76755FFB9FFFDFA8BFC4EFD7F06BF" ID-GBIF-Occurrence="2028137829" collectingDate="1983-08-21" collectionCode="TMSA" collectorName="Penrith" country="South Africa" elevation="800" latitude="-32.483334" location="Cederberg Range" longLatPrecision="1210" longitude="19.366667" pageId="27" pageNumber="68" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Western Cape" typeStatus="paratype">
<typeStatus id="E564D3AAFFB9FFFDFA8BFC4EFA500693" box="[1297,1410,958,982]" pageId="27" pageNumber="68" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="2CD9A681FFB9FFFDFA14FC4EFF1306BC" pageId="27" pageNumber="68" type="male">3 ♂♂</specimenCount>
and
<specimenCount id="2CD9A681FFB9FFFDFF63FC12FEEB06BC" box="[249,313,994,1018]" pageId="27" pageNumber="68" type="female">2 ♀♀</specimenCount>
(
<collectionCode id="5CCEF5CDFFB9FFFDFED3FC12FE4E06BF" box="[329,412,994,1018]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="27" pageNumber="68">TMSA</collectionCode>
), same data as holotype
</materialsCitation>
;
<materialsCitation id="8AB76755FFB9FFFDFD23FC11FED0015B" ID-GBIF-Occurrence="2028137832" collectingDate="1983-08-21" collectionCode="TMSA" collectorName="Penrith" country="South Africa" elevation="800" latitude="-32.483334" location="Cederberg Range" longLatPrecision="1210" longitude="19.366667" pageId="27" pageNumber="68" specimenCount="26" stateProvince="Western Cape" typeStatus="paratype">
<specimenCount id="2CD9A681FFB9FFFDFD23FC11FC8006BF" box="[697,850,993,1018]" pageId="27" pageNumber="68" type="generic">26 specimens</specimenCount>
with the same data as holotype but coordinates and elevation
</materialsCitation>
:
<materialsCitation id="8AB76755FFB9FFFDFE96FBF6FCB10158" ID-GBIF-Occurrence="2028137840" box="[268,867,1029,1054]" collectionCode="TMSA" country="South Africa" elevation="650" latitude="-32.4" longLatPrecision="1211" longitude="19.416666" pageId="27" pageNumber="68" specimenCount="9" specimenCount-female="8" specimenCount-male="1" stateProvince="Western Cape" typeStatus="paratype">
<specimenCount id="2CD9A681FFB9FFFDFE96FBF6FE980158" box="[268,330,1030,1054]" pageId="27" pageNumber="68" type="male">1 ♂♂</specimenCount>
and
<specimenCount id="2CD9A681FFB9FFFDFE1AFBF6FE6D0158" box="[384,447,1030,1054]" pageId="27" pageNumber="68" type="female">8 ♀♀</specimenCount>
(
<collectionCode id="5CCEF5CDFFB9FFFDFE57FBF6FDF0015B" box="[461,546,1030,1054]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="27" pageNumber="68">TMSA</collectionCode>
):
<geoCoordinate id="5FEB0BCFFFB9FFFDFDAEFBF6FD4B015B" box="[564,665,1030,1054]" direction="south" orientation="latitude" pageId="27" pageNumber="68" precision="925" value="-32.4">32° 24' S</geoCoordinate>
<geoCoordinate id="5FEB0BCFFFB9FFFDFD3AFBF6FCD40158" box="[672,774,1029,1054]" direction="east" orientation="longitude" pageId="27" pageNumber="68" precision="925" value="19.416666">19° 25' E</geoCoordinate>
/
<quantity id="FD27C0EDFFB9FFFDFC81FBF5FCB10158" box="[795,867,1029,1054]" metricMagnitude="2" metricUnit="m" metricValue="6.5" pageId="27" pageNumber="68" unit="m" value="650.0">
<elevation id="B1F28A3BFFB9FFFDFC81FBF5FCB10158" box="[795,867,1029,1054]" metricMagnitude="2" metricUnit="m" metricValue="6.5" pageId="27" pageNumber="68" unit="m" value="650.0">650 m</elevation>
</quantity>
</materialsCitation>
,
<materialsCitation id="8AB76755FFB9FFFDFCF5FBF6FBA2015B" ID-GBIF-Occurrence="2028137798" box="[879,1136,1029,1054]" collectionCode="UPSA" country="South Africa" pageId="27" pageNumber="68" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Western Cape" typeStatus="paratype">
<specimenCount id="2CD9A681FFB9FFFDFCF5FBF6FC4A0158" box="[879,920,1030,1054]" pageId="27" pageNumber="68" type="male">1 ♂</specimenCount>
and
<specimenCount id="2CD9A681FFB9FFFDFC54FBF6FBDE0158" box="[974,1036,1030,1054]" pageId="27" pageNumber="68" type="female">2 ♀♀</specimenCount>
(
<collectionCode id="5CCEF5CDFFB9FFFDFB81FBF6FBBA015B" box="[1051,1128,1030,1054]" country="South Africa" httpUri="http://grbio.org/cool/73qr-ciny" name="University of Pretoria" pageId="27" pageNumber="68">UPSA</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="8AB76755FFB9FFFDFBE1FBF5FAF6015B" ID-GBIF-Occurrence="2028137812" box="[1147,1316,1029,1054]" collectionCode="TMSA" country="South Africa" pageId="27" pageNumber="68" specimenCount="6" specimenCount-female="6" stateProvince="Western Cape" typeStatus="paratype">
<specimenCount id="2CD9A681FFB9FFFDFBE1FBF5FB6B0158" box="[1147,1209,1029,1054]" pageId="27" pageNumber="68" type="female">6 ♀♀</specimenCount>
(
<collectionCode id="5CCEF5CDFFB9FFFDFB52FBF6FACE015B" box="[1224,1308,1030,1054]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="27" pageNumber="68">TMSA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="8AB76755FFB9FFFDFAB5FBF6FD730104" ID-GBIF-Occurrence="2028137820" collectionCode="TMSA" country="South Africa" elevation="100" latitude="-32.45" longLatPrecision="1211" longitude="19.383333" pageId="27" pageNumber="68" specimenCount="5" specimenCount-female="3" specimenCount-male="2" stateProvince="Western Cape" typeStatus="paratype">
<specimenCount id="2CD9A681FFB9FFFDFAB5FBF6FABF0158" box="[1327,1389,1030,1054]" pageId="27" pageNumber="68" type="male">2 ♂♂</specimenCount>
and
<specimenCount id="2CD9A681FFB9FFFDFF0DFBDAFF040104" box="[151,214,1066,1090]" pageId="27" pageNumber="68" type="female">3 ♀♀</specimenCount>
(
<collectionCode id="5CCEF5CDFFB9FFFDFF7EFBDAFEEB0107" box="[228,313,1066,1090]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="27" pageNumber="68">TMSA</collectionCode>
):
<geoCoordinate id="5FEB0BCFFFB9FFFDFED1FBDAFE630107" box="[331,433,1066,1090]" direction="south" orientation="latitude" pageId="27" pageNumber="68" precision="925" value="-32.45">32° 27' S</geoCoordinate>
<geoCoordinate id="5FEB0BCFFFB9FFFDFE2DFBDAFDCC0104" box="[439,542,1066,1090]" direction="east" orientation="longitude" pageId="27" pageNumber="68" precision="925" value="19.383333">19° 23' E</geoCoordinate>
/ 1 /
<quantity id="FD27C0EDFFB9FFFDFDCDFBDAFD730104" box="[599,673,1066,1090]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="27" pageNumber="68" unit="m" value="100.0">
<elevation id="B1F28A3BFFB9FFFDFDCDFBDAFD730104" box="[599,673,1066,1090]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="27" pageNumber="68" unit="m" value="100.0">100 m</elevation>
</quantity>
</materialsCitation>
;
<materialsCitation id="8AB76755FFB9FFFDFD36FBDAFB200104" ID-GBIF-Occurrence="2028137806" box="[684,1266,1065,1090]" collectionCode="TMSA" country="South Africa" elevation="650" latitude="-32.383335" longLatPrecision="1211" longitude="19.4" pageId="27" pageNumber="68" specimenCount="9" specimenCount-female="5" specimenCount-male="4" stateProvince="Western Cape" typeStatus="paratype">
<specimenCount id="2CD9A681FFB9FFFDFD36FBDAFD040104" box="[684,726,1066,1090]" pageId="27" pageNumber="68" type="male">4 ♂</specimenCount>
and
<specimenCount id="2CD9A681FFB9FFFDFC96FBD9FC980104" box="[780,842,1065,1090]" pageId="27" pageNumber="68" type="female">5 ♀♀</specimenCount>
(
<collectionCode id="5CCEF5CDFFB9FFFDFCC3FBDAFC7C0107" box="[857,942,1066,1090]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="27" pageNumber="68">TMSA</collectionCode>
):
<geoCoordinate id="5FEB0BCFFFB9FFFDFC5AFBDAFBF40107" box="[960,1062,1066,1090]" direction="south" orientation="latitude" pageId="27" pageNumber="68" precision="925" value="-32.383335">32° 23' S</geoCoordinate>
<geoCoordinate id="5FEB0BCFFFB9FFFDFBB6FBDAFB410104" box="[1068,1171,1066,1090]" direction="east" orientation="longitude" pageId="27" pageNumber="68" precision="925" value="19.4">19° 24' E</geoCoordinate>
/
<quantity id="FD27C0EDFFB9FFFDFB32FBD9FB200104" box="[1192,1266,1065,1090]" metricMagnitude="2" metricUnit="m" metricValue="6.5" pageId="27" pageNumber="68" unit="m" value="650.0">
<elevation id="B1F28A3BFFB9FFFDFB32FBD9FB200104" box="[1192,1266,1065,1090]" metricMagnitude="2" metricUnit="m" metricValue="6.5" pageId="27" pageNumber="68" unit="m" value="650.0">650 m</elevation>
</quantity>
</materialsCitation>
;
<materialsCitation id="8AB76755FFB9FFFDFB64FBDAFD870120" ID-GBIF-Occurrence="2028137824" collectionCode="TMSA" country="South Africa" elevation="650" latitude="-32.366665" longLatPrecision="1211" longitude="19.4" pageId="27" pageNumber="68" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Western Cape" typeStatus="paratype">
<specimenCount id="2CD9A681FFB9FFFDFB64FBDAFAF50104" box="[1278,1319,1066,1090]" pageId="27" pageNumber="68" type="male">1 ♂</specimenCount>
and
<specimenCount id="2CD9A681FFB9FFFDFAC7FBDAFA4E0104" box="[1373,1436,1066,1090]" pageId="27" pageNumber="68" type="female">2 ♀♀</specimenCount>
(
<collectionCode id="5CCEF5CDFFB9FFFDFF05FBBEFF260123" box="[159,244,1102,1126]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="27" pageNumber="68">TMSA</collectionCode>
):
<geoCoordinate id="5FEB0BCFFFB9FFFDFE93FBBEFEA70123" box="[265,373,1102,1126]" direction="south" orientation="latitude" pageId="27" pageNumber="68" precision="925" value="-32.366665">32° 22' S</geoCoordinate>
<geoCoordinate id="5FEB0BCFFFB9FFFDFEE5FBBEFE3E0120" box="[383,492,1102,1126]" direction="east" orientation="longitude" pageId="27" pageNumber="68" precision="925" value="19.4">19° 24' E</geoCoordinate>
/
<quantity id="FD27C0EDFFB9FFFDFD92FBBDFD870120" box="[520,597,1101,1126]" metricMagnitude="2" metricUnit="m" metricValue="6.5" pageId="27" pageNumber="68" unit="m" value="650.0">
<elevation id="B1F28A3BFFB9FFFDFD92FBBDFD870120" box="[520,597,1101,1126]" metricMagnitude="2" metricUnit="m" metricValue="6.5" pageId="27" pageNumber="68" unit="m" value="650.0">650 m</elevation>
</quantity>
</materialsCitation>
;
<materialsCitation id="8AB76755FFB9FFFDFDF9FBBEFD6D01EB" ID-GBIF-Occurrence="2028137825" collectionCode="UPSA" collectorName="W. P. Strumpher" country="South Africa" location="Ramakraal" longitude="19.43167" pageId="27" pageNumber="68" specimenCount="13" specimenCount-female="6" specimenCount-male="7" typeStatus="paratype">
<specimenCount id="2CD9A681FFB9FFFDFDF9FBBEFD770120" box="[611,677,1102,1125]" pageId="27" pageNumber="68" type="male">7 ♂♂</specimenCount>
and
<specimenCount id="2CD9A681FFB9FFFDFD7BFBBDFCF10120" box="[737,803,1101,1126]" pageId="27" pageNumber="68" type="female">6 ♀♀</specimenCount>
(
<collectionCode id="5CCEF5CDFFB9FFFDFCAFFBBEFC560123" box="[821,900,1102,1126]" country="South Africa" httpUri="http://grbio.org/cool/73qr-ciny" name="University of Pretoria" pageId="27" pageNumber="68">UPSA</collectionCode>
): “
<collectingCountry id="42C82D98FFB9FFFDFC3EFBBDFBEA0123" box="[932,1080,1101,1126]" name="South Africa" pageId="27" pageNumber="68">South Africa</collectingCountry>
/
<location id="3F003BD3FFB9FFFDFBCEFBBDFB1F0123" LSID="urn:lsid:plazi:treatment:B276DC1EFFB9FFFAFF0DFD75FC2304E8:3F003BD3FFB9FFFDFBCEFBBDFB1F0123" box="[1108,1229,1101,1126]" country="South Africa" longitude="19.43167" name="Ramakraal" pageId="27" pageNumber="68">Ramakraal</location>
/ S 32. 41883°
<geoCoordinate id="5FEB0BCFFFB9FFFDFA11FBBEFECB01CF" direction="east" orientation="longitude" pageId="27" pageNumber="68" precision="1" value="19.43167">E 019.43167°</geoCoordinate>
/ 17. 0 8. 2015 / C. Deschodt &amp;
<collectorName id="972A08DEFFB9FFFDFD17FB82FC9401CF" box="[653,838,1137,1162]" pageId="27" pageNumber="68">W. P. Strümpher</collectorName>
leg [printed] //
<typeStatus id="E564D3AAFFB9FFFDFC61FB82FB5401CF" box="[1019,1158,1138,1162]" pageId="27" pageNumber="68" type="paratype">PARATYPE</typeStatus>
/
<taxonomicName id="FDDF168BFFB9FFFDFB05FB81FA5E01CC" authorityName="Abdalla &amp; Deschodt &amp; Scholtz &amp; Sole" authorityYear="2018" box="[1183,1420,1137,1161]" class="Insecta" family="Scarabaeidae" genus="Macroderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="68" phylum="Arthropoda" rank="species" species="soleiana">
<emphasis id="08ABB11AFFB9FFFDFB05FB81FA5E01CC" box="[1183,1420,1137,1161]" italics="true" pageId="27" pageNumber="68">Macroderes soleiana</emphasis>
</taxonomicName>
/ Abdalla &amp; Deschodt / 2018 / [red label, printed]”
</materialsCitation>
;
<materialsCitation id="8AB76755FFB9FFFDFD50FB66FB5001B3" ID-GBIF-Occurrence="2028137815" collectionCode="UPSA" collectorName="W. P. Strumpher" country="South Africa" latitude="-32.45506" location="Leopard Rock" longLatPrecision="1" longitude="19.40608" pageId="27" pageNumber="68" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Western Cape" typeStatus="paratype">
<specimenCount id="2CD9A681FFB9FFFDFD50FB66FD2601E8" box="[714,756,1174,1198]" pageId="27" pageNumber="68" type="male">1 ♂</specimenCount>
and
<specimenCount id="2CD9A681FFB9FFFDFCB3FB66FCBA01E8" box="[809,872,1174,1198]" pageId="27" pageNumber="68" type="female">2 ♀♀</specimenCount>
(
<collectionCode id="5CCEF5CDFFB9FFFDFCEDFB66FC1701EB" box="[887,965,1174,1198]" country="South Africa" httpUri="http://grbio.org/cool/73qr-ciny" name="University of Pretoria" pageId="27" pageNumber="68">UPSA</collectionCode>
): “
<collectingCountry id="42C82D98FFB9FFFDFC7BFB65FBA101EB" box="[993,1139,1173,1198]" name="South Africa" pageId="27" pageNumber="68">South Africa</collectingCountry>
/
<collectingRegion id="F81BA3EAFFB9FFFDFB12FB66FA5C01EB" box="[1160,1422,1173,1198]" country="South Africa" name="Western Cape" pageId="27" pageNumber="68">Western Cape Province</collectingRegion>
/
<location id="3F003BD3FFB9FFFDFF0DFB49FEEE0197" LSID="urn:lsid:plazi:treatment:B276DC1EFFB9FFFAFF0DFD75FC2304E8:3F003BD3FFB9FFFDFF0DFB49FEEE0197" box="[151,316,1209,1234]" country="South Africa" latitude="-32.45506" longLatPrecision="1" longitude="19.40608" name="Leopard Rock" pageId="27" pageNumber="68" stateProvince="Western Cape">Leopard Rock</location>
/
<geoCoordinate id="5FEB0BCFFFB9FFFDFEC4FB49FE300197" box="[350,482,1209,1234]" direction="south" orientation="latitude" pageId="27" pageNumber="68" precision="1" value="-32.45506">S32.45506°</geoCoordinate>
<geoCoordinate id="5FEB0BCFFFB9FFFDFE75FB49FD500197" box="[495,642,1209,1234]" direction="east" orientation="longitude" pageId="27" pageNumber="68" precision="1" value="19.40608">E019.40608°</geoCoordinate>
/ 17. 0 8. 2015 / C. Deschodt &amp;
<collectorName id="972A08DEFFB9FFFDFB85FB4AFB330197" box="[1055,1249,1209,1234]" pageId="27" pageNumber="68">W. P. Strümpher</collectorName>
leg [printed] //
<typeStatus id="E564D3AAFFB9FFFDFF0DFB2EFEF001B3" box="[151,290,1246,1270]" pageId="27" pageNumber="68" type="paratype">PARATYPE</typeStatus>
/
<taxonomicName id="FDDF168BFFB9FFFDFEA6FB2DFDF801B0" authorityName="Abdalla &amp; Deschodt &amp; Scholtz &amp; Sole" authorityYear="2018" box="[316,554,1245,1269]" class="Insecta" family="Scarabaeidae" genus="Macroderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="68" phylum="Arthropoda" rank="species" species="soleiana">
<emphasis id="08ABB11AFFB9FFFDFEA6FB2DFDF801B0" box="[316,554,1245,1269]" italics="true" pageId="27" pageNumber="68">Macroderes soleiana</emphasis>
</taxonomicName>
/ Abdalla &amp; Deschodt / 2018 / [red label, printed]”
</materialsCitation>
;
<materialsCitation id="8AB76755FFB9FFFDFB0AFB2EFCE5007B" ID-GBIF-Occurrence="2028137819" collectingDate="2015-08-21" collectionCode="UPSA" country="South Africa" latitude="-32.483612" location="Matjiesrivier" longLatPrecision="1" longitude="19.36014" municipality="Kleinlee" pageId="27" pageNumber="68" specimenCount="1" specimenCount-male="1" stateProvince="Western Cape" typeStatus="paratype">
<specimenCount id="2CD9A681FFB9FFFDFB0AFB2EFB6E01B0" box="[1168,1212,1246,1270]" pageId="27" pageNumber="68" type="male">1 ♂</specimenCount>
(
<collectionCode id="5CCEF5CDFFB9FFFDFB57FB2EFACE01B3" box="[1229,1308,1246,1270]" country="South Africa" httpUri="http://grbio.org/cool/73qr-ciny" name="University of Pretoria" pageId="27" pageNumber="68">UPSA</collectionCode>
): “
<collectingMunicipality id="DA04F772FFB9FFFDFAA1FB2DFA4E01B3" box="[1339,1436,1245,1270]" pageId="27" pageNumber="68">Kleinlee</collectingMunicipality>
[uu] lak /
<location id="3F003BD3FFB9FFFDFE92FAF1FE4B005F" LSID="urn:lsid:plazi:treatment:B276DC1EFFB9FFFAFF0DFD75FC2304E8:3F003BD3FFB9FFFDFE92FAF1FE4B005F" box="[264,409,1281,1306]" country="South Africa" latitude="-32.483612" longLatPrecision="1" longitude="19.36014" municipality="Kleinlee" name="Matjiesrivier" pageId="27" pageNumber="68" stateProvince="Western Cape">Matjiesrivier</location>
/ NR /
<geoCoordinate id="5FEB0BCFFFB9FFFDFE75FAF2FD49005F" box="[495,667,1282,1306]" direction="south" orientation="latitude" pageId="27" pageNumber="68" precision="1" value="-32.483612">32° 29' 01.0'' S</geoCoordinate>
<geoCoordinate id="5FEB0BCFFFB9FFFDFD39FAF2FC9D005C" box="[675,847,1281,1306]" direction="east" orientation="longitude" pageId="27" pageNumber="68" precision="1" value="19.36014">19° 21' 36.5'' E</geoCoordinate>
/
<date id="4E614BC8FFB9FFFDFCFDFAF1FC39005F" box="[871,1003,1281,1306]" pageId="27" pageNumber="68" value="2015-08-21">
<collectingDate id="5E25B220FFB9FFFDFCFDFAF1FC39005F" box="[871,1003,1281,1306]" pageId="27" pageNumber="68" value="2015-08-21">21-08-2015</collectingDate>
</date>
[printed] //
<typeStatus id="E564D3AAFFB9FFFDFBEEFAF2FB2D005F" box="[1140,1279,1282,1306]" pageId="27" pageNumber="68" type="paratype">PARATYPE</typeStatus>
/
<taxonomicName id="FDDF168BFFB9FFFDFA8DFAF1FF240078" authorityName="Abdalla &amp; Deschodt &amp; Scholtz &amp; Sole" authorityYear="2018" class="Insecta" family="Scarabaeidae" genus="Macroderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="68" phylum="Arthropoda" rank="species" species="soleiana">
<emphasis id="08ABB11AFFB9FFFDFA8DFAF1FF240078" italics="true" pageId="27" pageNumber="68">Macroderes soleiana</emphasis>
</taxonomicName>
/ Abdalla &amp; Deschodt / 2018 / [red label, printed]”
</materialsCitation>
.
<materialsCitation id="8AB76755FFB9FFFDFCDBFAD6FED200C3" ID-GBIF-Occurrence="2028137846" collectionCode="UPSA" country="South Africa" latitude="-32.444973" location="Matjiesrivier" longLatPrecision="1" longitude="19.351473" municipality="Perdewater" pageId="27" pageNumber="68" specimenCount="1" specimenCount-female="1" stateProvince="Western Cape" typeStatus="paratype">
<specimenCount id="2CD9A681FFB9FFFDFCDBFAD6FCB90078" box="[833,875,1318,1342]" pageId="27" pageNumber="68" type="female">1 ♀</specimenCount>
(
<collectionCode id="5CCEF5CDFFB9FFFDFCE0FAD6FC1A007B" box="[890,968,1318,1342]" country="South Africa" httpUri="http://grbio.org/cool/73qr-ciny" name="University of Pretoria" pageId="27" pageNumber="68">UPSA</collectionCode>
): “
<collectingMunicipality id="DA04F772FFB9FFFDFC7EFAD5FBB7007B" box="[996,1125,1317,1342]" pageId="27" pageNumber="68">Perdewater</collectingMunicipality>
/
<location id="3F003BD3FFB9FFFDFBE1FAD5FADE007B" LSID="urn:lsid:plazi:treatment:B276DC1EFFB9FFFAFF0DFD75FC2304E8:3F003BD3FFB9FFFDFBE1FAD5FADE007B" box="[1147,1292,1317,1342]" country="South Africa" latitude="-32.444973" longLatPrecision="1" longitude="19.351473" municipality="Perdewater" name="Matjiesrivier" pageId="27" pageNumber="68" stateProvince="Western Cape">Matjiesrivier</location>
/ NR /
<geoCoordinate id="5FEB0BCFFFB9FFFDFAC4FAD6FF2E0027" direction="south" orientation="latitude" pageId="27" pageNumber="68" precision="1" value="-32.444973">S 32° 26' 41.9''</geoCoordinate>
<geoCoordinate id="5FEB0BCFFFB9FFFDFF61FABAFE700027" box="[251,418,1353,1378]" direction="south" orientation="latitude" pageId="27" pageNumber="68" precision="1" value="-19.351473">
S
<geoCoordinate id="5FEB0BCFFFB9FFFDFE8AFABAFE700027" box="[272,418,1353,1378]" direction="east" orientation="longitude" pageId="27" pageNumber="68" precision="1" value="19.351473">19° 21' 05.3''</geoCoordinate>
</geoCoordinate>
E [printed] //
<typeStatus id="E564D3AAFFB9FFFDFDDAFABAFD190027" box="[576,715,1354,1378]" pageId="27" pageNumber="68" type="paratype">PARATYPE</typeStatus>
/
<taxonomicName id="FDDF168BFFB9FFFDFD7AFAB9FC1E0024" authorityName="Abdalla &amp; Deschodt &amp; Scholtz &amp; Sole" authorityYear="2018" box="[736,972,1353,1377]" class="Insecta" family="Scarabaeidae" genus="Macroderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="68" phylum="Arthropoda" rank="species" species="soleiana">
<emphasis id="08ABB11AFFB9FFFDFD7AFAB9FC1E0024" box="[736,972,1353,1377]" italics="true" pageId="27" pageNumber="68">Macroderes soleiana</emphasis>
</taxonomicName>
/ Abdalla &amp; Deschodt / 2018 / [red label, printed]”
</materialsCitation>
.
</paragraph>
<paragraph id="3A606D08FFB9FFFDFF5DFA61FB7A00EC" blockId="27.[151,1436,849,2026]" box="[199,1192,1425,1450]" pageId="27" pageNumber="68">
<emphasis id="08ABB11AFFB9FFFDFF5DFA61FE1A00EF" bold="true" box="[199,456,1425,1450]" pageId="27" pageNumber="68">
<typeStatus id="E564D3AAFFB9FFFDFF5DFA61FEE600EF" box="[199,308,1425,1450]" pageId="27" pageNumber="68" type="holotype">Holotype</typeStatus>
description.
</emphasis>
<typeStatus id="E564D3AAFFB9FFFDFE54FA61FDEB00EF" box="[462,569,1425,1450]" pageId="27" pageNumber="68" type="holotype">Holotype</typeStatus>
,
<specimenCount id="2CD9A681FFB9FFFDFDDFFA62FD8800EC" box="[581,602,1426,1449]" pageId="27" pageNumber="68" type="male"></specimenCount>
(
<figureCitation id="A2E4718DFFB9FFFDFDF3FA61FD6F00EF" box="[617,701,1425,1450]" captionStart="FIGURES 1923" captionStartId="9.[151,264,1928,1951]" captionTargetBox="[411,1172,706,1874]" captionTargetId="figure@9.[337,1297,671,1926]" captionTargetPageId="9" captionText="FIGURES 1923. Habitus of Macroderes species. 19, M. oreatus Abdalla &amp; Deschodt, new species; 20, M. cederbergensis Abdalla &amp; Deschodt, new species; 21, M. soleiana Abdalla &amp; Deschodt, new species; 22, M. tourtosus Abdalla &amp; Scholtz, new species; 23, M. leipoldti Abdalla &amp; Deschodt, new species." httpUri="https://zenodo.org/record/2606176/files/figure.png" pageId="27" pageNumber="68">Fig. 21</figureCitation>
). Body length
<quantity id="FD27C0EDFFB9FFFDFCFBFA62FC6B00EC" box="[865,953,1426,1450]" metricMagnitude="-3" metricUnit="m" metricValue="8.8" pageId="27" pageNumber="68" unit="mm" value="8.8">8.8 mm</quantity>
, body width
<quantity id="FD27C0EDFFB9FFFDFBD1FA61FB7600EC" box="[1099,1188,1425,1450]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" pageId="27" pageNumber="68" unit="mm" value="6.2">6.2 mm</quantity>
.
</paragraph>
<paragraph id="3A606D08FFB9FFFDFF5DFA45FB3E00B7" blockId="27.[151,1436,849,2026]" pageId="27" pageNumber="68">
<emphasis id="08ABB11AFFB9FFFDFF5DFA45FEDC008B" bold="true" box="[199,270,1461,1486]" pageId="27" pageNumber="68">Head.</emphasis>
Genae sharp angled and sutures indistinct. Frontal suture distinct, angulate medially. Dorsal surface of clypeus rugose anterior of frontal suture, punctate posteriorly. Area between punctures shagreened.
</paragraph>
</subSubSection>
<subSubSection id="72C53E83FFB9FFFDFF5DFA0DFB91023B" pageId="27" pageNumber="68" type="description">
<paragraph id="3A606D08FFB9FFFDFF5DFA0DFAAB037F" blockId="27.[151,1436,849,2026]" pageId="27" pageNumber="68">
<emphasis id="08ABB11AFFB9FFFDFF5DFA0DFE950353" bold="true" box="[199,327,1533,1558]" pageId="27" pageNumber="68">Pronotum.</emphasis>
Convex, anterior angles rounded, posterior angles rounded. Lateral border not punctate. Dorsal surface regularly punctate punctures close together, separated by less than one puncture diameter (
<figureCitation id="A2E4718DFFB9FFFDFB79F9D1FA93037F" box="[1251,1345,1569,1594]" captionStart="FIGURES 7681" captionStartId="20.[151,264,1637,1660]" captionTargetBox="[171,1423,205,1613]" captionTargetId="figure@20.[171,1423,205,1613]" captionTargetPageId="20" captionText="FIGURES 7681. Punctuation of the base of pronotum of Macroderes species. 76, M. cederbergensis Abdalla &amp; Deschodt, new species; 77, M. gifboomi Abdalla &amp; Scholtz, new species; 78, M. oreatus Abdalla &amp; Deschodt, new species; 79, M. porselinus Abdalla, new species; 80, M. soleiana Abdalla &amp; Deschodt, new species; 81, M. leipoldti Abdalla &amp; Deschodt, new species." httpUri="https://zenodo.org/record/2606198/files/figure.png" pageId="27" pageNumber="68">Figs. 80</figureCitation>
,
<figureCitation id="A2E4718DFFB9FFFDFAD6F9D2FAB9037F" box="[1356,1387,1570,1594]" captionStart="FIGURES 8893" captionStartId="23.[151,264,1292,1315]" captionTargetBox="[316,1276,183,1264]" captionTargetId="figure@23.[316,1276,183,1264]" captionTargetPageId="23" captionText="FIGURES 8893. Pronotum punctures shape (88, 89) and sculpture of second elytra intervals (9093) of Macroderes species. 88, M. soleiana Abdalla &amp; Deschodt, new species; 89, M. tourtosus Abdalla and Scholtz, new species; 90, M. minutus Frolov and Scholtz, 2005; 91, M. greeni (Kirby, 1818); 92, M. politulus Preudhomme de Borre, 1880; 93, M. gifboomi Abdalla &amp; Scholtz, new species." httpUri="https://zenodo.org/record/2606202/files/figure.png" pageId="27" pageNumber="68">88</figureCitation>
).
</paragraph>
<paragraph id="3A606D08FFB9FFFDFF5DF9B5FECD03AB" blockId="27.[151,1436,849,2026]" pageId="27" pageNumber="68">
<emphasis id="08ABB11AFFB9FFFDFF5DF9B5FEC8031B" bold="true" box="[199,282,1605,1630]" pageId="27" pageNumber="68">Elytra.</emphasis>
Elytral interstriae 18 slightly raised, punctate, punctures shallow, punctures separated by 12 puncture diameters, each puncture with short associated seta. Elytral margins not carinate. Area between punctures shagreened. Striae clearly bordered, faintly shagreened, punctures very faint (
<figureCitation id="A2E4718DFFB9FFFDFC61F97DFB9D03E3" box="[1019,1103,1677,1702]" captionStart="FIGURES 9499" captionStartId="24.[151,264,1299,1322]" captionTargetBox="[316,1273,193,1274]" captionTargetId="figure@24.[316,1273,193,1274]" captionTargetPageId="24" captionText="FIGURES 9499. Sculpture of second elytra intervals of Macroderes species. 94, M. cederbergensis Abdalla &amp; Deschodt, new species; 95, M. leipoldti Abdalla &amp; Deschodt, new species; 96, M. oreatus Abdalla &amp; Deschodt, new species; 97, M. porselinus Abdalla, new species; 98, M. soleiana Abdalla &amp; Deschodt, new species; 99, M. tourtosus Abdalla &amp; Scholtz, new species." httpUri="https://zenodo.org/record/2606204/files/figure.png" pageId="27" pageNumber="68">Fig. 98</figureCitation>
). Stria 9 disjunct, anterior 1/5 medially between stria 8 and 10, second 2/3 very close to stria 10, interstria 10 almost covered by stria
<quantity id="FD27C0EDFFB9FFFDFADEF942FAA3038F" box="[1348,1393,1713,1738]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" pageId="27" pageNumber="68" unit="in" value="9.0">9 in</quantity>
the anterior 1/3.
</paragraph>
<paragraph id="3A606D08FFB9FFFDFF5DF909FAEA0257" blockId="27.[151,1436,849,2026]" box="[199,1336,1785,1810]" pageId="27" pageNumber="68">
<emphasis id="08ABB11AFFB9FFFDFF5DF909FE930257" bold="true" box="[199,321,1785,1810]" pageId="27" pageNumber="68">Aedeagus.</emphasis>
The sclerite of internal sac elongate and strongly curved, without lateral process (
<figureCitation id="A2E4718DFFB9FFFDFB4DF909FAF90257" box="[1239,1323,1785,1810]" captionStart="FIGURES 2439" captionStartId="10.[151,264,1343,1366]" captionTargetBox="[323,1250,205,1286]" captionTargetId="figure@10.[323,1283,195,1303]" captionTargetPageId="10" captionText="FIGURES 2439. Internal sac of aedeagus (24), aedeagus (25) and internal sac sclerite (26-39) of Macroderes species. 26, M. endroedyi Frolov &amp; Scholtz, 2005; 27, M. amplior Frolov &amp; Scholtz, 2005; 28, M. bias (Olivier, 1789); 29, M. cederbergensis Abdalla &amp; Deschodt, new species; 30, M. cornutus Frolov &amp; Scholtz, 2005; 31, M. foveatus Frolov &amp; Scholtz, 2005; 32, M. gifboomi Abdalla &amp; Scholtz, new species; 33, M. leipoldti Abdalla &amp; Deschodt, new species; 34, M. mutilans Kolbe, 1908; 35, M. namakwanus Frolov &amp; Scholtz, 2005; 36, M. oreatus Abdalla &amp; Deschodt, new species; 37, M. porselinus Abdalla, new species; 38, M. soleiana Abdalla &amp; Deschodt, new species; 39, M. tourtosus Abdalla &amp; Scholtz, new species." httpUri="https://zenodo.org/record/2606178/files/figure.png" pageId="27" pageNumber="68">Fig. 38</figureCitation>
).
</paragraph>
<paragraph id="3A606D08FFB9FFFDFF5DF8EDFA480270" blockId="27.[151,1436,849,2026]" box="[199,1434,1821,1846]" pageId="27" pageNumber="68">
<emphasis id="08ABB11AFFB9FFFDFF5DF8EDFEED0273" bold="true" box="[199,319,1821,1846]" pageId="27" pageNumber="68">Size range</emphasis>
. Males length:
<quantity id="FD27C0EDFFB9FFFDFE73F8EEFDAC0270" box="[489,638,1822,1846]" metricMagnitude="-2" metricUnit="m" metricValue="1.025" metricValueMax="1.23" metricValueMin="0.82" pageId="27" pageNumber="68" unit="mm" value="10.25" valueMax="12.3" valueMin="8.2">8.212.3 mm</quantity>
, width: 5.08.0 mm; females length: 10.0
<quantity id="FD27C0EDFFB9FFFDFBCEF8EDFB6A0270" box="[1108,1208,1821,1846]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" pageId="27" pageNumber="68" unit="mm" value="12.5">12.5 mm</quantity>
, width: 7.0
<quantity id="FD27C0EDFFB9FFFDFADBF8EEFA440270" box="[1345,1430,1822,1846]" metricMagnitude="-3" metricUnit="m" metricValue="8.1" pageId="27" pageNumber="68" unit="mm" value="8.1">8.1 mm</quantity>
.
</paragraph>
<paragraph id="3A606D08FFB9FFFDFF5DF8B1FB91023B" blockId="27.[151,1436,849,2026]" pageId="27" pageNumber="68">Female. Beside the body length variations, female differs from male by having less developed frontoclypeal suture as well as the pronotal laterl marigins more clear in dorsal view than in male.</paragraph>
</subSubSection>
<subSubSection id="72C53E83FFB9FFFDFF5DF879FD9C02AF" pageId="27" pageNumber="68" type="etymology">
<paragraph id="3A606D08FFB9FFFDFF5DF879FD9C02AF" blockId="27.[151,1436,849,2026]" pageId="27" pageNumber="68">
<emphasis id="08ABB11AFFB9FFFDFF5DF879FE9C02E7" bold="true" box="[199,334,1929,1954]" pageId="27" pageNumber="68">Etymology.</emphasis>
This species is named after our colleague Prof. Catherine L. Sole of the Department Zoology and Entomology, University of Pretoria, in recognition of her leadership and her molecular work on the African dung beetles, lacewings and Baboon spiders.
</paragraph>
</subSubSection>
<subSubSection id="72C53E83FFBEFFFAFF5DFF67FB8D05BD" pageId="28" pageNumber="69" type="diagnosis">
<paragraph id="3A606D08FFBEFFFAFF5DFF67FB8D05BD" blockId="28.[151,1436,151,429]" pageId="28" pageNumber="69">
<emphasis id="08ABB11AFFBEFFFAFF5DFF67FE1F05F5" bold="true" box="[199,461,151,176]" pageId="28" pageNumber="69">Differential diagnosis.</emphasis>
This species is most similar to
<taxonomicName id="FDDF168BFFBEFFFAFCB6FF69FC4F05F5" authorityName="Harold /" authorityYear="1877" box="[812,925,152,176]" class="Insecta" family="Scarabaeidae" genus="Macroderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="69" phylum="Arthropoda" rank="species" species="nitidus">
<emphasis id="08ABB11AFFBEFFFAFCB6FF69FC4F05F5" box="[812,925,152,176]" italics="true" pageId="28" pageNumber="69">M. nitidus</emphasis>
</taxonomicName>
and
<taxonomicName id="FDDF168BFFBEFFFAFC49FF69FB7305F5" authority="Abdalla &amp; Deschodt &amp; Scholtz &amp; Sole, 2018" authorityName="Abdalla &amp; Deschodt &amp; Scholtz &amp; Sole" authorityYear="2018" box="[979,1185,152,176]" class="Insecta" family="Scarabaeidae" genus="Macroderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="69" phylum="Arthropoda" rank="species" species="cederbergensis" status="sp. nov.">
<emphasis id="08ABB11AFFBEFFFAFC49FF69FB7305F5" box="[979,1185,152,176]" italics="true" pageId="28" pageNumber="69">M. cederbergensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="13980C61FFBEFFFAFB3DFF68FAE005F5" box="[1191,1330,151,176]" pageId="28" pageNumber="69" rank="species">new species</taxonomicNameLabel>
. It differs from the former in being more shagreened and less shiny on the elytral interstriae (
<figureCitation id="A2E4718DFFBEFFFAFBD9FF4CFB470591" box="[1091,1173,188,213]" captionStart="FIGURES 9499" captionStartId="24.[151,264,1299,1322]" captionTargetBox="[316,1273,193,1274]" captionTargetId="figure@24.[316,1273,193,1274]" captionTargetPageId="24" captionText="FIGURES 9499. Sculpture of second elytra intervals of Macroderes species. 94, M. cederbergensis Abdalla &amp; Deschodt, new species; 95, M. leipoldti Abdalla &amp; Deschodt, new species; 96, M. oreatus Abdalla &amp; Deschodt, new species; 97, M. porselinus Abdalla, new species; 98, M. soleiana Abdalla &amp; Deschodt, new species; 99, M. tourtosus Abdalla &amp; Scholtz, new species." httpUri="https://zenodo.org/record/2606204/files/figure.png" pageId="28" pageNumber="69">Fig. 98</figureCitation>
) and from the latter by being larger and having the pronotal punctures smaller and farther apart (
<figureCitation id="A2E4718DFFBEFFFAFC50FF2FFBFA05BD" box="[970,1064,223,248]" captionStart="FIGURES 7681" captionStartId="20.[151,264,1637,1660]" captionTargetBox="[171,1423,205,1613]" captionTargetId="figure@20.[171,1423,205,1613]" captionTargetPageId="20" captionText="FIGURES 7681. Punctuation of the base of pronotum of Macroderes species. 76, M. cederbergensis Abdalla &amp; Deschodt, new species; 77, M. gifboomi Abdalla &amp; Scholtz, new species; 78, M. oreatus Abdalla &amp; Deschodt, new species; 79, M. porselinus Abdalla, new species; 80, M. soleiana Abdalla &amp; Deschodt, new species; 81, M. leipoldti Abdalla &amp; Deschodt, new species." httpUri="https://zenodo.org/record/2606198/files/figure.png" pageId="28" pageNumber="69">Figs. 80</figureCitation>
,
<figureCitation id="A2E4718DFFBEFFFAFBA9FF10FB8005BD" box="[1075,1106,224,248]" captionStart="FIGURES 8893" captionStartId="23.[151,264,1292,1315]" captionTargetBox="[316,1276,183,1264]" captionTargetId="figure@23.[316,1276,183,1264]" captionTargetPageId="23" captionText="FIGURES 8893. Pronotum punctures shape (88, 89) and sculpture of second elytra intervals (9093) of Macroderes species. 88, M. soleiana Abdalla &amp; Deschodt, new species; 89, M. tourtosus Abdalla and Scholtz, new species; 90, M. minutus Frolov and Scholtz, 2005; 91, M. greeni (Kirby, 1818); 92, M. politulus Preudhomme de Borre, 1880; 93, M. gifboomi Abdalla &amp; Scholtz, new species." httpUri="https://zenodo.org/record/2606202/files/figure.png" pageId="28" pageNumber="69">88</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="72C53E83FFBEFFFAFF5DFEF4FB1A0420" pageId="28" pageNumber="69" type="description">
<paragraph id="3A606D08FFBEFFFAFF5DFEF4FB1A0420" blockId="28.[151,1436,151,429]" pageId="28" pageNumber="69">
<emphasis id="08ABB11AFFBEFFFAFF5DFEF4FE2A0458" bold="true" box="[199,504,260,285]" pageId="28" pageNumber="69">Habitat and distribution.</emphasis>
Cederberg Mountains (
<figureCitation id="A2E4718DFFBEFFFAFC90FEF4FC830459" box="[778,849,260,285]" captionStart="FIGURE 2" captionStartId="2.[151,250,1574,1596]" captionText="FIGURE 2. Distribution map of Macroderes species." pageId="28" pageNumber="69">Fig. 2</figureCitation>
). Specimens have been collected between 600 and
<quantity id="FD27C0EDFFBEFFFAFF0DFED8FF3F047A" box="[151,237,296,320]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="28" pageNumber="69" unit="m" value="1000.0">1000 m</quantity>
a.s.l. and mainly in the Swartruggens Quartzite Fynbos vegetation
<typeStatus id="E564D3AAFFBEFFFAFC70FED8FBCC0405" box="[1002,1054,296,320]" pageId="28" pageNumber="69">type</typeStatus>
. This area receives between
<quantity id="FD27C0EDFFBEFFFAFAFEFED8FF260421" metricMagnitude="-1" metricUnit="m" metricValue="4.1" metricValueMax="6.2" metricValueMin="2.0" pageId="28" pageNumber="69" unit="mm" value="410.0" valueMax="620.0" valueMin="200.0">200 620 mm</quantity>
of precipitation per year. The soils are sandy but skeletal (
<bibRefCitation id="5E4E10F9FFBEFFFAFC18FEBCFB6E0420" author="Mucina, L. &amp; Rutherford, M. C." box="[898,1212,332,357]" pageId="28" pageNumber="69" pagination="1 - 807" refId="ref21930" refString="Mucina, L. &amp; Rutherford, M. C. (Eds.) (2006) The Vegetation of South Africa, Lesotho and Swaziland. Strelitzia, 19, 1 - 807." type="journal article" year="2006">Mucina &amp; Rutherford 2006</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="72C53E83FFBEFFFAFF5DFE9FFC2304E8" pageId="28" pageNumber="69" type="discussion">
<paragraph id="3A606D08FFBEFFFAFF5DFE9FFC2304E8" blockId="28.[151,1436,151,429]" pageId="28" pageNumber="69">
<emphasis id="08ABB11AFFBEFFFAFF5DFE9FFEE204CD" bold="true" box="[199,304,367,392]" pageId="28" pageNumber="69">Remark.</emphasis>
<bibRefCitation id="5E4E10F9FFBEFFFAFEA1FE9FFD8904CD" author="Frolov, A. V. &amp; Scholtz, C. H." box="[315,603,367,392]" pageId="28" pageNumber="69" pagination="373 - 393" refId="ref21694" refString="Frolov, A. V. &amp; Scholtz, C. H. (2005) Revision of the southern African genus Macroderes Westwood (Coleoptera: Scarabaeidae: Scarabaeinae). Annales de la Societe Entomologique de France, 40 (3 - 4), 373 - 393. https: // doi. org / 10.1080 / 00379271.2004.10697429" type="journal article" year="2005">Frolov &amp; Scholtz (2005)</bibRefCitation>
used specimens from this species for their redescription of
<taxonomicName id="FDDF168BFFBEFFFAFA85FE81FA4604CD" authorityName="Harold /" authorityYear="1877" box="[1311,1428,368,392]" class="Insecta" family="Scarabaeidae" genus="Macroderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="69" phylum="Arthropoda" rank="species" species="nitidus">
<emphasis id="08ABB11AFFBEFFFAFA85FE81FA4604CD" box="[1311,1428,368,392]" italics="true" pageId="28" pageNumber="69">M. nitidus</emphasis>
</taxonomicName>
. Please, refer to the redescription of
<taxonomicName id="FDDF168BFFBEFFFAFDBFFE65FD4504E9" authorityName="Harold /" authorityYear="1877" box="[549,663,404,428]" class="Insecta" family="Scarabaeidae" genus="Macroderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="69" phylum="Arthropoda" rank="species" species="nitidus">
<emphasis id="08ABB11AFFBEFFFAFDBFFE65FD4504E9" box="[549,663,404,428]" italics="true" pageId="28" pageNumber="69">M. nitidus</emphasis>
</taxonomicName>
for more detailed explanation.
</paragraph>
</subSubSection>
</treatment>
</document>