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<document id="D4AA3B557D2AB111B6237D20B6A5773B" ID-CLB-Dataset="28091" ID-DOI="10.11646/zootaxa.4504.1.3" ID-GBIF-Dataset="0b803003-244e-4511-aaf4-5e9c3f863861" ID-ISSN="1175-5326" ID-Zenodo-Dep="2606164" ID-ZooBank="52BA28B6-4280-4BD0-8B38-C794F04E4A66" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553577945224" checkinUser="plazi" docAuthor="Abdalla, Ishtiag H., Deschodt, Christian M., Scholtz, Clarke H. &amp; Sole, Catherine L." docDate="2018" docId="B276DC1EFFB4FFF0FF0DFBA9FC0B02AA" docLanguage="en" docName="zootaxa.4504.1.3.pdf" docOrigin="Zootaxa 4504 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Macroderes namakwanus Frolov &amp; Scholtz 2005" docType="treatment" docVersion="8" lastPageNumber="63" masterDocId="4E4FA466FFA2FFE6FF9AFFF0FFD20545" masterDocTitle="An update to the taxonomy of the genus Macroderes Westwood 1842 (Coleoptera: Scarabaeidae: Scarabaeinae) with descriptions of new species from South Africa" masterLastPageNumber="75" masterPageNumber="41" pageNumber="63" updateTime="1698712718888" updateUser="ExternalLinkService">
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<mods:title id="4CD90D82B04707AE95C209B21634729B">An update to the taxonomy of the genus Macroderes Westwood 1842 (Coleoptera: Scarabaeidae: Scarabaeinae) with descriptions of new species from South Africa</mods:title>
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<mods:namePart id="6B426E9CD8244719FEED384531F5A387">Abdalla, Ishtiag H.</mods:namePart>
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<mods:namePart id="DC27FC00BA4BF330130CC96AE7CA351A">Deschodt, Christian M.</mods:namePart>
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<mods:namePart id="857E9DB50D2DEF1AD9993B5BB18AE0C4">Scholtz, Clarke H.</mods:namePart>
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<mods:namePart id="CB9682BF2C37B40E4C05A1A052D92299">Sole, Catherine L.</mods:namePart>
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<mods:title id="87BCE384E788DF82CCFCEB8FAA7F38A8">Zootaxa</mods:title>
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<mods:date id="FAC83736C9554FB3B3EBC085D287624E">2018</mods:date>
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<mods:number id="9BE59D2BB8ADC25F0CB493CF38C33348">2018-10-23</mods:number>
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<treatment id="B276DC1EFFB4FFF0FF0DFBA9FC0B02AA" ID-DOI="http://doi.org/10.5281/zenodo.5996748" ID-GBIF-Taxon="154494814" ID-Zenodo-Dep="5996748" LSID="urn:lsid:plazi:treatment:B276DC1EFFB4FFF0FF0DFBA9FC0B02AA" httpUri="http://treatment.plazi.org/id/B276DC1EFFB4FFF0FF0DFBA9FC0B02AA" lastPageNumber="63" pageId="22" pageNumber="63">
<subSubSection id="72C53E83FFB4FFF0FF0DFBA9FD260136" box="[151,756,1113,1139]" pageId="22" pageNumber="63" type="nomenclature">
<paragraph id="3A606D08FFB4FFF0FF0DFBA9FD260136" blockId="22.[151,756,1113,1173]" box="[151,756,1113,1139]" pageId="22" pageNumber="63">
<heading id="6128DA64FFB4FFF0FF0DFBA9FD260136" bold="true" box="[151,756,1113,1139]" fontSize="11" level="1" pageId="22" pageNumber="63" reason="1">
<taxonomicName id="FDDF168BFFB4FFF0FF0DFBA9FD260136" ID-CoL="3X4TS" authority="Frolov &amp; Scholtz, 2005" authorityName="Frolov &amp; Scholtz" authorityYear="2005" box="[151,756,1113,1139]" class="Insecta" family="Scarabaeidae" genus="Macroderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="63" phylum="Arthropoda" rank="species" species="namakwanus">
<emphasis id="08ABB11AFFB4FFF0FF0DFBA9FD260136" bold="true" box="[151,756,1113,1139]" pageId="22" pageNumber="63">
<emphasis id="08ABB11AFFB4FFF0FF0DFBA9FE1C0136" bold="true" box="[151,462,1113,1139]" italics="true" pageId="22" pageNumber="63">Macroderes namakwanus</emphasis>
<bibRefCitation id="5E4E10F9FFB4FFF0FE4CFBA9FD260136" author="Frolov, A. V. &amp; Scholtz, C. H." box="[470,756,1113,1139]" pageId="22" pageNumber="63" pagination="373 - 393" refId="ref21694" refString="Frolov, A. V. &amp; Scholtz, C. H. (2005) Revision of the southern African genus Macroderes Westwood (Coleoptera: Scarabaeidae: Scarabaeinae). Annales de la Societe Entomologique de France, 40 (3 - 4), 373 - 393. https: // doi. org / 10.1080 / 00379271.2004.10697429" type="journal article" year="2005">Frolov &amp; Scholtz, 2005</bibRefCitation>
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</paragraph>
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<subSubSection id="72C53E83FFB4FFF0FF0DFB8CFE7901D0" box="[151,427,1148,1173]" pageId="22" pageNumber="63" type="description">
<paragraph id="3A606D08FFB4FFF0FF0DFB8CFE7901D0" blockId="22.[151,756,1113,1173]" box="[151,427,1148,1173]" pageId="22" pageNumber="63">
(
<figureCitation id="A2E4718DFFB4FFF0FF05FB8CFF2E01D1" box="[159,252,1148,1173]" captionStart="FIGURES 11–14" captionStartId="6.[151,264,1359,1382]" captionTargetBox="[309,1210,215,1332]" captionTargetId="figure@6.[304,1264,195,1332]" captionTargetPageId="6" captionText="FIGURES 11–14. Habitus of Macroderes species. 11, M. minutus Frolov &amp; Scholtz, 2005; 12, M. mutilans Kolbe, 1908; 13, M. namakwanus Frolov &amp; Scholtz, 2005; 14, M. nitidus Harold, 1877." httpUri="https://zenodo.org/record/2606172/files/figure.png" pageId="22" pageNumber="63">Figs. 13</figureCitation>
,
<figureCitation id="A2E4718DFFB4FFF0FE92FB8CFEF401D0" box="[264,294,1148,1173]" captionStart="FIGURES 24–39" captionStartId="10.[151,264,1343,1366]" captionTargetBox="[323,1250,205,1286]" captionTargetId="figure@10.[323,1283,195,1303]" captionTargetPageId="10" captionText="FIGURES 24–39. Internal sac of aedeagus (24), aedeagus (25) and internal sac sclerite (26-39) of Macroderes species. 26, M. endroedyi Frolov &amp; Scholtz, 2005; 27, M. amplior Frolov &amp; Scholtz, 2005; 28, M. bias (Olivier, 1789); 29, M. cederbergensis Abdalla &amp; Deschodt, new species; 30, M. cornutus Frolov &amp; Scholtz, 2005; 31, M. foveatus Frolov &amp; Scholtz, 2005; 32, M. gifboomi Abdalla &amp; Scholtz, new species; 33, M. leipoldti Abdalla &amp; Deschodt, new species; 34, M. mutilans Kolbe, 1908; 35, M. namakwanus Frolov &amp; Scholtz, 2005; 36, M. oreatus Abdalla &amp; Deschodt, new species; 37, M. porselinus Abdalla, new species; 38, M. soleiana Abdalla &amp; Deschodt, new species; 39, M. tourtosus Abdalla &amp; Scholtz, new species." httpUri="https://zenodo.org/record/2606178/files/figure.png" pageId="22" pageNumber="63">35</figureCitation>
,
<figureCitation id="A2E4718DFFB4FFF0FEA8FB8CFE8301D1" box="[306,337,1148,1172]" captionStart="FIGURES 40–45" captionStartId="12.[151,264,1736,1759]" captionTargetBox="[306,1266,609,1690]" captionTargetId="figure@12.[306,1266,609,1690]" captionTargetPageId="12" captionText="FIGURES 40–45. Pronotum of Macroderes species. 40, M. foveatus Frolov &amp; Scholtz, 2005; 41, M. cornutus Frolov &amp; Scholtz, 2005; 42, M. namakwanus Frolov &amp; Scholtz, 2005; 43, M. politulus Preudhomme de Borre, 1880; 44, M. greeni (Kirby, 1818); 45, M. endroedyi Frolov &amp; Scholtz, 2005." httpUri="https://zenodo.org/record/2606180/files/figure.png" pageId="22" pageNumber="63">42</figureCitation>
,
<figureCitation id="A2E4718DFFB4FFF0FEC7FB8CFEA901D0" box="[349,379,1148,1173]" captionStart="FIGURES 58–63" captionStartId="16.[151,264,1848,1871]" captionTargetBox="[165,1430,411,1834]" captionTargetId="figure@16.[165,1430,411,1835]" captionTargetPageId="16" captionText="FIGURES 58–63. Punctuation of the base of pronotum of Macroderes species. 58, M. greeni (Kirby, 1818); 59, M. arrowi Janssens, 1939; 60, M. bias (Olivier, 1789); 61, M. namakwanus Frolov &amp; Scholtz, 2005; 62, M. endroedyi Frolov &amp; Scholtz, 2005; 63, M. politulus Preudhomme de Borre, 1880." httpUri="https://zenodo.org/record/2606188/files/figure.png" pageId="22" pageNumber="63">61</figureCitation>
,
<figureCitation id="A2E4718DFFB4FFF0FE1CFB8CFE7101D0" box="[390,419,1148,1173]" captionStart="FIGURES 70–75" captionStartId="18.[151,264,1624,1647]" captionTargetBox="[159,1435,197,1611]" captionTargetId="figure@18.[159,1435,197,1611]" captionTargetPageId="18" captionText="FIGURES 70–75. Sculpture of second elytra intervals of Macroderes species. 70, M. arrowi Janssens, 1939; 71, M. cornutus Frolov &amp; Scholtz, 2005; 72, M. endroedyi Frolov &amp; Scholtz, 2005;73, M. foveatus Frolov &amp; Scholtz, 2005; 74, M. mutilans Kolbe, 1908; 75, M. namakwanus Frolov &amp; Scholtz, 2005." httpUri="https://zenodo.org/record/2606196/files/figure.png" pageId="22" pageNumber="63">75</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="72C53E83FFB4FFF0FF0DFB34FD11019F" box="[151,707,1220,1242]" pageId="22" pageNumber="63" type="reference_group">
<paragraph id="3A606D08FFB4FFF0FF0DFB34FD11019F" blockId="22.[151,707,1220,1242]" box="[151,707,1220,1242]" pageId="22" pageNumber="63">
<treatmentCitationGroup id="1ACF4A26FFB4FFF0FF0DFB34FD11019F" box="[151,707,1220,1242]" pageId="22" pageNumber="63">
<taxonomicName id="FDDF168BFFB4FFF0FF0DFB34FD6D019F" ID-CoL="3X4TS" authority="Frolov &amp; Scholtz 2005: 388" authorityName="Frolov &amp; Scholtz" authorityPageNumber="388" authorityYear="2005" box="[151,703,1220,1242]" class="Insecta" family="Scarabaeidae" genus="Macroderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="63" phylum="Arthropoda" rank="species" species="namakwanus">
<emphasis id="08ABB11AFFB4FFF0FF0DFB34FE4B019F" box="[151,409,1220,1242]" italics="true" pageId="22" pageNumber="63">Macroderes namakwanus</emphasis>
<treatmentCitation id="BB7E4B19FFB4FFF0FE3AFB34FD6D019F" author="Frolov, A. V. &amp; Scholtz, C. H." box="[416,703,1220,1242]" page="388" pageId="22" pageNumber="63" year="2005">
<bibRefCitation id="5E4E10F9FFB4FFF0FE3AFB34FD5E019F" author="Frolov, A. V. &amp; Scholtz, C. H." box="[416,652,1220,1242]" pageId="22" pageNumber="63" pagination="373 - 393" refId="ref21694" refString="Frolov, A. V. &amp; Scholtz, C. H. (2005) Revision of the southern African genus Macroderes Westwood (Coleoptera: Scarabaeidae: Scarabaeinae). Annales de la Societe Entomologique de France, 40 (3 - 4), 373 - 393. https: // doi. org / 10.1080 / 00379271.2004.10697429" type="journal article" year="2005">Frolov &amp; Scholtz 2005</bibRefCitation>
: 388
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.
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</paragraph>
</subSubSection>
<subSubSection id="72C53E83FFB4FFF0FF0DFAF6FA4C021B" pageId="22" pageNumber="63" type="materials_examined">
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<typeStatus id="E564D3AAFFB4FFF0FF0DFAF6FF00005A" box="[151,210,1286,1311]" pageId="22" pageNumber="63">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="42C82D98FFB4FFF0FEA6FAF7FEA7005A" box="[316,373,1287,1311]" name="South Africa" pageId="22" pageNumber="63">RSA</collectingCountry>
,
<collectingRegion id="F81BA3EAFFB4FFF0FE1AFAF7FDCE005A" box="[384,540,1287,1311]" country="South Africa" name="Western Cape" pageId="22" pageNumber="63">Western Cape</collectingRegion>
<collectingMunicipality id="DA04F772FFB4FFF0FDB9FAF7FD8B005B" box="[547,601,1287,1310]" pageId="22" pageNumber="63">Prov</collectingMunicipality>
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<location id="3F003BD3FFB4FFF0FDFEFAF6FD00005A" LSID="urn:lsid:plazi:treatment:B276DC1EFFB4FFF0FF0DFBA9FC0B02AA:3F003BD3FFB4FFF0FDFEFAF6FD00005A" box="[612,722,1286,1311]" country="South Africa" latitude="-31.157242" longLatPrecision="1" longitude="17.765472" municipality="Prov" name="Hoekbaai" pageId="22" pageNumber="63" stateProvince="Western Cape">Hoekbaai</location>
,
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.
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<paragraph id="3A606D08FFB4FFF0FF5DFADAFD7003B6" blockId="22.[151,1438,1286,2031]" pageId="22" pageNumber="63">
<emphasis id="08ABB11AFFB4FFF0FF5DFADAFE270006" bold="true" box="[199,501,1322,1347]" pageId="22" pageNumber="63">Type material examined.</emphasis>
<materialsCitation id="8AB76755FFB4FFF0FE65FADAFB5100CE" ID-GBIF-Occurrence="2028137793" collectingDate="2003-09-04" collectingDateMax="2003-09-11" collectingDateMin="2003-09-04" collectionCode="TMSA" collectorName="C. Deschodt" country="South Africa" latitude="-31.07391" location="Hoekbaai" longLatPrecision="1" longitude="17.765472" municipality="Prov" pageId="22" pageNumber="63" specimenCount="1" specimenCount-male="1" stateProvince="Western Cape" typeStatus="holotype">
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,
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(
<collectionCode id="5CCEF5CDFFB4FFF0FD05FADBFD210006" box="[671,755,1323,1347]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="22" pageNumber="63">TMSA</collectionCode>
): “
<collectingCountry id="42C82D98FFB4FFF0FC89FADBFC9E0006" box="[787,844,1323,1347]" name="South Africa" pageId="22" pageNumber="63">RSA</collectingCountry>
/
<collectingRegion id="F81BA3EAFFB4FFF0FCF2FADBFBD40006" box="[872,1030,1323,1347]" country="South Africa" name="Western Cape" pageId="22" pageNumber="63">Western Cape</collectingRegion>
<collectingMunicipality id="DA04F772FFB4FFF0FB8BFADBFB940007" box="[1041,1094,1323,1346]" pageId="22" pageNumber="63">Prov</collectingMunicipality>
/
<location id="3F003BD3FFB4FFF0FBFBFADAFB1E0006" LSID="urn:lsid:plazi:treatment:B276DC1EFFB4FFF0FF0DFBA9FC0B02AA:3F003BD3FFB4FFF0FBFBFADAFB1E0006" box="[1121,1228,1322,1347]" country="South Africa" latitude="-31.07391" longLatPrecision="1" longitude="17.765472" municipality="Prov" name="Hoekbaai" pageId="22" pageNumber="63" stateProvince="Western Cape">Hoekbaai</location>
/ A. Frolov &amp;
<collectorName id="972A08DEFFB4FFF0FA18FADBFED30022" pageId="22" pageNumber="63">C. Deschodt</collectorName>
leg [printed] // AF-0039 (19) /
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<collectingDate id="5E25B220FFB4FFF0FD16FABFFC930022" box="[652,833,1359,1383]" pageId="22" pageNumber="63" value="2003-09-04" valueMax="2003-09-11" valueMin="2003-09-04">4–11. IX. 2003</collectingDate>
</date>
/
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<geoCoordinate id="5FEB0BCFFFB4FFF0FBA2FABFFAC20022" box="[1080,1296,1358,1383]" direction="east" orientation="longitude" pageId="22" pageNumber="63" precision="1" value="17.765472">E 017° 45' 55.70''</geoCoordinate>
[printed] //
<typeStatus id="E564D3AAFFB4FFF0FF0DFA83FEEF00CE" box="[151,317,1395,1419]" pageId="22" pageNumber="63" type="holotype">HOLOTYPUS</typeStatus>
/
<taxonomicName id="FDDF168BFFB4FFF0FECEFA83FDA000CE" authorityName="Frolov &amp; Scholtz" authorityYear="2005" box="[340,626,1395,1419]" class="Insecta" family="Scarabaeidae" genus="Macroderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="63" phylum="Arthropoda" rank="species" species="namakwanus">
<emphasis id="08ABB11AFFB4FFF0FECEFA83FDA000CE" box="[340,626,1395,1419]" italics="true" pageId="22" pageNumber="63">Macroderes namakwanus</emphasis>
</taxonomicName>
/ Frolov &amp; Scholtz / 2003 [red label, printed]”
</materialsCitation>
;
<materialsCitation id="8AB76755FFB4FFF0FB14FA83FE9700B2" ID-GBIF-Occurrence="2028137809" collectingDate="1979-08-25" collectionCode="TMSA" collectorName="Endrody-Younga" country="South Africa" county="Afr" location="Brakriver" municipality="Namaquland" pageId="22" pageNumber="63" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<typeStatus id="E564D3AAFFB4FFF0FB14FA83FB2D00CE" box="[1166,1279,1395,1419]" pageId="22" pageNumber="63" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="2CD9A681FFB4FFF0FA90FA83FAFF00CE" box="[1290,1325,1395,1419]" pageId="22" pageNumber="63" type="male">1♂</specimenCount>
(
<collectionCode id="5CCEF5CDFFB4FFF0FAA6FA83FA4300CE" box="[1340,1425,1395,1419]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="22" pageNumber="63">TMSA</collectionCode>
): “S.
<collectingCounty id="D3011584FFB4FFF0FF5BFA66FF3A00EA" box="[193,232,1430,1455]" pageId="22" pageNumber="63">Afr</collectingCounty>
/
<collectingMunicipality id="DA04F772FFB4FFF0FF64FA66FE5F00EA" box="[254,397,1430,1455]" pageId="22" pageNumber="63">Namaquland</collectingMunicipality>
/
<location id="3F003BD3FFB4FFF0FE39FA66FDDC00EA" LSID="urn:lsid:plazi:treatment:B276DC1EFFB4FFF0FF0DFBA9FC0B02AA:3F003BD3FFB4FFF0FE39FA66FDDC00EA" box="[419,526,1430,1455]" country="South Africa" county="Afr" municipality="Namaquland" name="Brakriver" pageId="22" pageNumber="63">Brakriver</location>
mouth / 36°.06' S–17°.44' E [printed] //
<date id="4E614BC8FFB4FFF0FC4CFA66FB8600EA" box="[982,1108,1430,1455]" pageId="22" pageNumber="63" value="1979-08-25">
<collectingDate id="5E25B220FFB4FFF0FC4CFA66FB8600EA" box="[982,1108,1430,1455]" pageId="22" pageNumber="63" value="1979-08-25">25. 8. 1979</collectingDate>
</date>
// E– Y: 1597 / groundtraps / 63 days /
<collectorName id="972A08DEFFB4FFF0FE99FA4AFE070096" box="[259,469,1466,1491]" pageId="22" pageNumber="63">Endrödy—Younga</collectorName>
leg [printed] // PARTYPUS /
<taxonomicName id="FDDF168BFFB4FFF0FCBEFA4BFB910096" authorityName="Frolov &amp; Scholtz" authorityYear="2005" box="[804,1091,1467,1491]" class="Insecta" family="Scarabaeidae" genus="Macroderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="63" phylum="Arthropoda" rank="species" species="namakwanus">
<emphasis id="08ABB11AFFB4FFF0FCBEFA4BFB910096" box="[804,1091,1467,1491]" italics="true" pageId="22" pageNumber="63">Macroderes namakwanus</emphasis>
</taxonomicName>
/ Frolvo &amp; Scholtz / 2003 [red label, printed]”
</materialsCitation>
;
<materialsCitation id="8AB76755FFB4FFF0FECAFA2FFCEF037A" ID-GBIF-Occurrence="2028137797" collectingDate="1979-08-27" collectionCode="TMSA" collectorName="Endrody-Younga" country="South Africa" county="Namaquland" location="Km" municipality="Hoekbaai" pageId="22" pageNumber="63" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="2CD9A681FFB4FFF0FECAFA2FFEA800B3" box="[336,378,1503,1527]" pageId="22" pageNumber="63" type="male">1 ♂</specimenCount>
(
<collectionCode id="5CCEF5CDFFB4FFF0FE13FA2FFE0C00B2" box="[393,478,1503,1527]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="22" pageNumber="63">TMSA</collectionCode>
): “S. AFR /
<collectingCounty id="D3011584FFB4FFF0FDF2FA2EFD2500B2" box="[616,759,1502,1527]" pageId="22" pageNumber="63">Namaquland</collectingCounty>
/
<collectingMunicipality id="DA04F772FFB4FFF0FC94FA2EFCAB00B2" box="[782,889,1502,1527]" pageId="22" pageNumber="63">Hoekbaai</collectingMunicipality>
/ 2
<location id="3F003BD3FFB4FFF0FC3FFA2FFC1D00B3" LSID="urn:lsid:plazi:treatment:B276DC1EFFB4FFF0FF0DFBA9FC0B02AA:3F003BD3FFB4FFF0FC3FFA2FFC1D00B3" box="[933,975,1503,1526]" country="South Africa" county="Namaquland" municipality="Hoekbaai" name="Km" pageId="22" pageNumber="63">Km</location>
/ E NE / 31°. 11' S 17°. 47' E [printed] //
<date id="4E614BC8FFB4FFF0FF0DF9F3FECE035E" box="[151,284,1539,1563]" pageId="22" pageNumber="63" value="1979-08-27">
<collectingDate id="5E25B220FFB4FFF0FF0DF9F3FECE035E" box="[151,284,1539,1563]" pageId="22" pageNumber="63" value="1979-08-27">27. 8. 1979</collectingDate>
</date>
// E–Y: 1612 / white sand / night /
<collectorName id="972A08DEFFB4FFF0FD51F9F2FC4F035E" box="[715,925,1538,1563]" pageId="22" pageNumber="63">Endrödy—Younga</collectorName>
leg [printed] //
<typeStatus id="E564D3AAFFB4FFF0FBC6F9F3FB2B035E" box="[1116,1273,1539,1563]" pageId="22" pageNumber="63" type="paratype">PARATYPUS</typeStatus>
/
<taxonomicName id="FDDF168BFFB4FFF0FA8DF9F3FEFB037B" authorityName="Frolov &amp; Scholtz" authorityYear="2005" class="Insecta" family="Scarabaeidae" genus="Macroderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="63" phylum="Arthropoda" rank="species" species="namakwanus">
<emphasis id="08ABB11AFFB4FFF0FA8DF9F3FEFB037B" italics="true" pageId="22" pageNumber="63">Macroderes namakwanus</emphasis>
</taxonomicName>
/ Frolov &amp; Scholtz / 2003 [red label, printed]”
</materialsCitation>
;
<materialsCitation id="8AB76755FFB4FFF0FCD2F9D7FBA703C2" ID-GBIF-Occurrence="2028137813" collectingDate="1979-08-23" collectionCode="TMSA" collectorName="Endrody-Younga" country="South Africa" county="Afr" latitude="-30.95" location="Kotzerus" municipality="Namaquland" pageId="22" pageNumber="63" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="2CD9A681FFB4FFF0FCD2F9D7FCA1037B" box="[840,883,1575,1599]" pageId="22" pageNumber="63" type="male">1 ♂</specimenCount>
(
<collectionCode id="5CCEF5CDFFB4FFF0FC19F9D7FC05037A" box="[899,983,1575,1599]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="22" pageNumber="63">TMSA</collectionCode>
): “S.
<collectingCounty id="D3011584FFB4FFF0FB8FF9D6FBEE037A" box="[1045,1084,1574,1599]" pageId="22" pageNumber="63">Afr</collectingCounty>
/
<collectingMunicipality id="DA04F772FFB4FFF0FBC9F9D6FB30037A" box="[1107,1250,1574,1599]" pageId="22" pageNumber="63">Namaquland</collectingMunicipality>
/
<location id="3F003BD3FFB4FFF0FB63F9D7FA8C037B" LSID="urn:lsid:plazi:treatment:B276DC1EFFB4FFF0FF0DFBA9FC0B02AA:3F003BD3FFB4FFF0FB63F9D7FA8C037B" box="[1273,1374,1575,1598]" country="South Africa" county="Afr" latitude="-30.95" municipality="Namaquland" name="Kotzerus" pageId="22" pageNumber="63">Kotzerus</location>
/
<geoCoordinate id="5FEB0BCFFFB4FFF0FAEFF9D7FF360326" direction="south" orientation="latitude" pageId="22" pageNumber="63" precision="925" value="-30.95">30° 57' S–</geoCoordinate>
17°.50' E [printed] //
<date id="4E614BC8FFB4FFF0FE7AF9BBFDB50326" box="[480,615,1611,1635]" pageId="22" pageNumber="63" value="1979-08-23">
<collectingDate id="5E25B220FFB4FFF0FE7AF9BBFDB50326" box="[480,615,1611,1635]" pageId="22" pageNumber="63" value="1979-08-23">23. 8. 1979</collectingDate>
</date>
// E–Y: 1581 / white dune / day /.
<collectorName id="972A08DEFFB4FFF0FB8EF9BAFB340326" box="[1044,1254,1610,1635]" pageId="22" pageNumber="63">Endrödy—Younga</collectorName>
leg [printed] //
<typeStatus id="E564D3AAFFB4FFF0FF0DF99FFEE703C2" box="[151,309,1647,1671]" pageId="22" pageNumber="63" type="paratype">PARATYPUS</typeStatus>
/
<taxonomicName id="FDDF168BFFB4FFF0FECEF99EFDA403C3" authorityName="Frolov &amp; Scholtz" authorityYear="2005" box="[340,630,1646,1670]" class="Insecta" family="Scarabaeidae" genus="Macroderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="63" phylum="Arthropoda" rank="species" species="namakwanus">
<emphasis id="08ABB11AFFB4FFF0FECEF99EFDA403C3" box="[340,630,1646,1670]" italics="true" pageId="22" pageNumber="63">Macroderes namakwanus</emphasis>
</taxonomicName>
/ Frolov &amp; Scholtz / 2003 [red– printed]”
</materialsCitation>
;
<materialsCitation id="8AB76755FFB4FFF0FB1FF99FFD4C03B6" ID-GBIF-Occurrence="2028137782" collectingDate="1976-09-08" collectionCode="TMSA" collectorName="Endrody-Younga" country="South Africa" county="Afr" location="Klein Helskloof" municipality="Richtersveld" pageId="22" pageNumber="63" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="2CD9A681FFB4FFF0FB1FF99FFB6103C3" box="[1157,1203,1647,1671]" pageId="22" pageNumber="63" type="female">1 ♀</specimenCount>
(
<collectionCode id="5CCEF5CDFFB4FFF0FB5DF99FFACE03C2" box="[1223,1308,1647,1671]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="22" pageNumber="63">TMSA</collectionCode>
): “S.
<collectingCounty id="D3011584FFB4FFF0FAF8F99EFA5B03C2" box="[1378,1417,1646,1671]" pageId="22" pageNumber="63">Afr</collectingCounty>
/
<collectingMunicipality id="DA04F772FFB4FFF0FF0DF962FEF603EE" box="[151,292,1682,1707]" pageId="22" pageNumber="63">Richtersveld</collectingMunicipality>
/
<location id="3F003BD3FFB4FFF0FEA4F962FE2603EE" LSID="urn:lsid:plazi:treatment:B276DC1EFFB4FFF0FF0DFBA9FC0B02AA:3F003BD3FFB4FFF0FEA4F962FE2603EE" box="[318,500,1682,1707]" country="South Africa" county="Afr" municipality="Richtersveld" name="Klein Helskloof" pageId="22" pageNumber="63">Klein Helskloof</location>
/ 28°.51' S–17°.24' E [printed] //
<date id="4E614BC8FFB4FFF0FCE1F962FC0F03EE" box="[891,989,1682,1707]" pageId="22" pageNumber="63" value="1976-09-08">
<collectingDate id="5E25B220FFB4FFF0FCE1F962FC0F03EE" box="[891,989,1682,1707]" pageId="22" pageNumber="63" value="1976-09-08">8.9.1976</collectingDate>
</date>
// E–Y: 1239 / groundtraps / 32 days /
<collectorName id="972A08DEFFB4FFF0FF0DF946FEBB038A" box="[151,361,1718,1743]" pageId="22" pageNumber="63">Endrödy—Younga</collectorName>
leg [printed] // groundtrap with faeces bait [printed] //
<typeStatus id="E564D3AAFFB4FFF0FC56F947FBB8038A" box="[972,1130,1719,1743]" pageId="22" pageNumber="63" type="paratype">PARATYPUS</typeStatus>
/
<taxonomicName id="FDDF168BFFB4FFF0FBE4F946FA4E038B" authorityName="Frolov &amp; Scholtz" authorityYear="2005" box="[1150,1436,1718,1742]" class="Insecta" family="Scarabaeidae" genus="Macroderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="63" phylum="Arthropoda" rank="species" species="namakwanus">
<emphasis id="08ABB11AFFB4FFF0FBE4F946FA4E038B" box="[1150,1436,1718,1742]" italics="true" pageId="22" pageNumber="63">Macroderes namakwanus</emphasis>
</taxonomicName>
/ Frolov &amp; Scholtz / 2003 [red label, printed]”
</materialsCitation>
.
</paragraph>
<paragraph id="3A606D08FFB4FFF0FF5DF90EFD97027E" blockId="22.[151,1438,1286,2031]" pageId="22" pageNumber="63">
<emphasis id="08ABB11AFFB4FFF0FF5DF90EFDE50252" bold="true" box="[199,567,1790,1815]" pageId="22" pageNumber="63">Additional material examined.</emphasis>
<materialsCitation id="8AB76755FFB4FFF0FDA5F90FFD93027E" ID-GBIF-Occurrence="2028137783" collectingDate="2015-08-12" collectionCode="UPSA" country="South Africa" location="Kotzerus" pageId="22" pageNumber="63" specimenCount="5" specimenCount-female="3" specimenCount-male="2">
<specimenCount id="2CD9A681FFB4FFF0FDA5F90FFD520253" box="[575,640,1791,1815]" pageId="22" pageNumber="63" type="male">2 ♂♂</specimenCount>
and
<specimenCount id="2CD9A681FFB4FFF0FD20F90FFD280253" box="[698,762,1791,1815]" pageId="22" pageNumber="63" type="female">3 ♀♀</specimenCount>
(
<collectionCode id="5CCEF5CDFFB4FFF0FC91F90FFC8B0252" box="[779,857,1791,1815]" country="South Africa" httpUri="http://grbio.org/cool/73qr-ciny" name="University of Pretoria" pageId="22" pageNumber="63">UPSA</collectionCode>
): “
<collectingCountry id="42C82D98FFB4FFF0FCE2F90EFBDE0252" box="[888,1036,1790,1815]" name="South Africa" pageId="22" pageNumber="63">South Africa</collectingCountry>
/
<quantity id="FD27C0EDFFB4FFF0FBBFF90EFB8D0252" box="[1061,1119,1790,1815]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="22" pageNumber="63" unit="km" value="6.0">6 km</quantity>
N of /
<location id="3F003BD3FFB4FFF0FB2FF90FFAC80253" LSID="urn:lsid:plazi:treatment:B276DC1EFFB4FFF0FF0DFBA9FC0B02AA:3F003BD3FFB4FFF0FB2FF90FFAC80253" box="[1205,1306,1791,1814]" country="South Africa" name="Kotzerus" pageId="22" pageNumber="63">Kotzerus</location>
/ S 30° 69' 26.11'' E 017° 86' 9.83'' /
<date id="4E614BC8FFB4FFF0FE29F8D3FDE4027E" box="[435,566,1826,1851]" pageId="22" pageNumber="63" value="2015-08-12">
<collectingDate id="5E25B220FFB4FFF0FE29F8D3FDE4027E" box="[435,566,1826,1851]" pageId="22" pageNumber="63" value="2015-08-12">12. 8. 2015</collectingDate>
</date>
”
</materialsCitation>
.
</paragraph>
<paragraph id="3A606D08FFB4FFF0FF5DF8B6FA4C021B" blockId="22.[151,1438,1286,2031]" box="[199,1438,1862,1887]" pageId="22" pageNumber="63">
<emphasis id="08ABB11AFFB4FFF0FF5DF8B6FE94021A" bold="true" box="[199,326,1862,1887]" pageId="22" pageNumber="63">Size range.</emphasis>
Males length:
<quantity id="FD27C0EDFFB4FFF0FE7DF8B7FD5B021B" box="[487,649,1863,1887]" metricMagnitude="-2" metricUnit="m" metricValue="1.16" metricValueMax="1.28" metricValueMin="1.04" pageId="22" pageNumber="63" unit="mm" value="11.6" valueMax="12.8" valueMin="10.4">10.4–12.8 mm</quantity>
, width: 7.0–
<quantity id="FD27C0EDFFB4FFF0FC8AF8B7FCB7021B" box="[784,869,1863,1887]" metricMagnitude="-3" metricUnit="m" metricValue="8.7" pageId="22" pageNumber="63" unit="mm" value="8.7">8.7 mm</quantity>
; females length:
<quantity id="FD27C0EDFFB4FFF0FB86F8B6FB6F021B" box="[1052,1213,1862,1887]" metricMagnitude="-2" metricUnit="m" metricValue="1.21" metricValueMax="1.25" metricValueMin="1.17" pageId="22" pageNumber="63" unit="mm" value="12.1" valueMax="12.5" valueMin="11.7">11.7–12.5 mm</quantity>
, width: 8.0–9.0 mm.
</paragraph>
</subSubSection>
<subSubSection id="72C53E83FFB4FFF0FF5DF89AFC0B028E" pageId="22" pageNumber="63" type="diagnosis">
<paragraph id="3A606D08FFB4FFF0FF5DF89AFC0B028E" blockId="22.[151,1438,1286,2031]" pageId="22" pageNumber="63">
<emphasis id="08ABB11AFFB4FFF0FF5DF89AFE0702C6" bold="true" box="[199,469,1898,1923]" pageId="22" pageNumber="63">Differential diagnosis.</emphasis>
<taxonomicName id="FDDF168BFFB4FFF0FE79F89BFD7002C6" authorityName="Frolov &amp; Scholtz" authorityYear="2005" box="[483,674,1899,1923]" class="Insecta" family="Scarabaeidae" genus="Macroderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="63" phylum="Arthropoda" rank="species" species="namakwanus">
<emphasis id="08ABB11AFFB4FFF0FE79F89BFD7002C6" box="[483,674,1899,1923]" italics="true" pageId="22" pageNumber="63">M. namakwanus</emphasis>
</taxonomicName>
is in habitus similar to
<taxonomicName id="FDDF168BFFB4FFF0FC4FF89BFBA202C6" authorityName="Frolov &amp; Scholtz" authorityYear="2005" box="[981,1136,1899,1923]" class="Insecta" family="Scarabaeidae" genus="Macroderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="63" phylum="Arthropoda" rank="species" species="endroedyi">
<emphasis id="08ABB11AFFB4FFF0FC4FF89BFBA202C6" box="[981,1136,1899,1923]" italics="true" pageId="22" pageNumber="63">M. endroedyi</emphasis>
</taxonomicName>
(
<figureCitation id="A2E4718DFFB4FFF0FB1DF89AFB3202C6" box="[1159,1248,1898,1923]" captionStart="FIGURES 11–14" captionStartId="6.[151,264,1359,1382]" captionTargetBox="[309,1210,215,1332]" captionTargetId="figure@6.[304,1264,195,1332]" captionTargetPageId="6" captionText="FIGURES 11–14. Habitus of Macroderes species. 11, M. minutus Frolov &amp; Scholtz, 2005; 12, M. mutilans Kolbe, 1908; 13, M. namakwanus Frolov &amp; Scholtz, 2005; 14, M. nitidus Harold, 1877." httpUri="https://zenodo.org/record/2606172/files/figure.png" pageId="22" pageNumber="63">Fig. 13</figureCitation>
) however, it is differentiated by the combination of the following characteristics: Pronotum regularly punctate lacking the distinct tubercle in between (
<figureCitation id="A2E4718DFFB4FFF0FE1BF842FE01028E" box="[385,467,1970,1995]" captionStart="FIGURES 58–63" captionStartId="16.[151,264,1848,1871]" captionTargetBox="[165,1430,411,1834]" captionTargetId="figure@16.[165,1430,411,1835]" captionTargetPageId="16" captionText="FIGURES 58–63. Punctuation of the base of pronotum of Macroderes species. 58, M. greeni (Kirby, 1818); 59, M. arrowi Janssens, 1939; 60, M. bias (Olivier, 1789); 61, M. namakwanus Frolov &amp; Scholtz, 2005; 62, M. endroedyi Frolov &amp; Scholtz, 2005; 63, M. politulus Preudhomme de Borre, 1880." httpUri="https://zenodo.org/record/2606188/files/figure.png" pageId="22" pageNumber="63">Fig. 61</figureCitation>
) and no shiny tubercles on the elytra (
<figureCitation id="A2E4718DFFB4FFF0FC1AF842FC03028E" box="[896,977,1970,1995]" captionStart="FIGURES 70–75" captionStartId="18.[151,264,1624,1647]" captionTargetBox="[159,1435,197,1611]" captionTargetId="figure@18.[159,1435,197,1611]" captionTargetPageId="18" captionText="FIGURES 70–75. Sculpture of second elytra intervals of Macroderes species. 70, M. arrowi Janssens, 1939; 71, M. cornutus Frolov &amp; Scholtz, 2005; 72, M. endroedyi Frolov &amp; Scholtz, 2005;73, M. foveatus Frolov &amp; Scholtz, 2005; 74, M. mutilans Kolbe, 1908; 75, M. namakwanus Frolov &amp; Scholtz, 2005." httpUri="https://zenodo.org/record/2606196/files/figure.png" pageId="22" pageNumber="63">Fig. 75</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="72C53E83FFB4FFF0FF5DF826FC0B02AA" box="[199,985,2006,2031]" pageId="22" pageNumber="63" type="description">
<paragraph id="3A606D08FFB4FFF0FF5DF826FC0B02AA" blockId="22.[151,1438,1286,2031]" box="[199,985,2006,2031]" pageId="22" pageNumber="63">
<emphasis id="08ABB11AFFB4FFF0FF5DF826FE3902AA" bold="true" box="[199,491,2006,2031]" pageId="22" pageNumber="63">Habitat and distribution</emphasis>
. See distribution under
<taxonomicName id="FDDF168BFFB4FFF0FD6DF827FCAF02AB" authorityName="Kolbe" authorityYear="1908" box="[759,893,2006,2030]" class="Insecta" family="Scarabaeidae" genus="Macroderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="63" phylum="Arthropoda" rank="species" species="mutilans">
<emphasis id="08ABB11AFFB4FFF0FD6DF827FCAF02AB" box="[759,893,2006,2030]" italics="true" pageId="22" pageNumber="63">M. mutilans</emphasis>
</taxonomicName>
(
<figureCitation id="A2E4718DFFB4FFF0FC17F826FC0202AA" box="[909,976,2006,2031]" captionStart="FIGURE 2" captionStartId="2.[151,250,1574,1596]" captionText="FIGURE 2. Distribution map of Macroderes species." pageId="22" pageNumber="63">Fig. 2</figureCitation>
)
</paragraph>
</subSubSection>
</treatment>
</document>
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