treatments-xml/data/B2/76/DC/B276DC1EFFA5FFEEFF0DF8D5FBC7003D.xml
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<document id="B8C88E3A89B9EAF89775E0AB2A5BFEBA" ID-CLB-Dataset="28091" ID-DOI="10.11646/zootaxa.4504.1.3" ID-GBIF-Dataset="0b803003-244e-4511-aaf4-5e9c3f863861" ID-ISSN="1175-5326" ID-Zenodo-Dep="2606164" ID-ZooBank="52BA28B6-4280-4BD0-8B38-C794F04E4A66" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553577945224" checkinUser="plazi" docAuthor="Abdalla, Ishtiag H., Deschodt, Christian M., Scholtz, Clarke H. &amp; Sole, Catherine L." docDate="2018" docId="B276DC1EFFA5FFEEFF0DF8D5FBC7003D" docLanguage="en" docName="zootaxa.4504.1.3.pdf" docOrigin="Zootaxa 4504 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Macroderes cederbergensis Abdalla &amp; Deschodt &amp; Scholtz &amp; Sole 2018, new species" docType="treatment" docUuid="9FED0DC0-CFF6-424C-A3AE-092656B24972" docUuidSource="ZooBank" docVersion="9" lastPageNumber="49" masterDocId="4E4FA466FFA2FFE6FF9AFFF0FFD20545" masterDocTitle="An update to the taxonomy of the genus Macroderes Westwood 1842 (Coleoptera: Scarabaeidae: Scarabaeinae) with descriptions of new species from South Africa" masterLastPageNumber="75" masterPageNumber="41" pageNumber="48" updateTime="1698712718888" updateUser="ExternalLinkService">
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<mods:title id="4DCC4F5494854159BB5E13743987E446">An update to the taxonomy of the genus Macroderes Westwood 1842 (Coleoptera: Scarabaeidae: Scarabaeinae) with descriptions of new species from South Africa</mods:title>
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<mods:namePart id="5784723EA3C0B99DBD68FAF1F2605C9C">Abdalla, Ishtiag H.</mods:namePart>
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<mods:namePart id="8A2AE01BA84758543718977531517D28">Deschodt, Christian M.</mods:namePart>
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<mods:namePart id="7645411D3BC06125DC72C2E0F73600F5">Scholtz, Clarke H.</mods:namePart>
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<mods:namePart id="248A68A08B9915D6424F54F84DC92C6C">Sole, Catherine L.</mods:namePart>
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<treatment id="B276DC1EFFA5FFEEFF0DF8D5FBC7003D" ID-DOI="http://doi.org/10.5281/zenodo.5996726" ID-GBIF-Taxon="154494804" ID-Zenodo-Dep="5996726" LSID="urn:lsid:zoobank.org:act:9FED0DC0-CFF6-424C-A3AE-092656B24972" httpUri="http://treatment.plazi.org/id/B276DC1EFFA5FFEEFF0DF8D5FBC7003D" lastPageId="8" lastPageNumber="49" pageId="7" pageNumber="48">
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<paragraph id="3A606D08FFA5FFE1FF0DF8D5FC560205" blockId="7.[151,900,1829,1889]" box="[151,900,1829,1856]" pageId="7" pageNumber="48">
<heading id="6128DA64FFA5FFE1FF0DF8D5FC560205" bold="true" box="[151,900,1829,1856]" fontSize="11" level="1" pageId="7" pageNumber="48" reason="1">
<emphasis id="08ABB11AFFA5FFE1FF0DF8D5FC560205" bold="true" box="[151,900,1829,1856]" pageId="7" pageNumber="48">
<taxonomicName id="FDDF168BFFA5FFE1FF0DF8D5FD350205" ID-CoL="3X4TF" authority="Abdalla &amp; Deschodt" authorityName="Abdalla &amp; Deschodt &amp; Scholtz &amp; Sole" authorityYear="2018" box="[151,743,1829,1856]" class="Insecta" family="Scarabaeidae" genus="Macroderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="48" phylum="Arthropoda" rank="species" species="cederbergensis" status="sp. nov.">
<emphasis id="08ABB11AFFA5FFE1FF0DF8D5FE32027A" bold="true" box="[151,480,1829,1855]" italics="true" pageId="7" pageNumber="48">Macroderes cederbergensis</emphasis>
Abdalla &amp; Deschodt
</taxonomicName>
,
<taxonomicNameLabel id="13980C61FFA5FFE1FD6EF8D6FC560205" box="[756,900,1830,1856]" pageId="7" pageNumber="48" rank="species">new species</taxonomicNameLabel>
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<subSubSection id="72C53E83FFA5FFE1FF0DF8B8FCAF02E4" pageId="7" pageNumber="48" type="description">
<paragraph id="3A606D08FFA5FFE1FF0DF8B8FE7E0224" blockId="7.[151,900,1829,1889]" box="[151,428,1864,1889]" pageId="7" pageNumber="48">
(
<figureCitation id="A2E4718DFFA5FFE1FF05F8B8FF2E0224" box="[159,252,1864,1889]" captionStart="FIGURES 1923" captionStartId="9.[151,264,1928,1951]" captionTargetBox="[411,1172,706,1874]" captionTargetId="figure@9.[337,1297,671,1926]" captionTargetPageId="9" captionText="FIGURES 1923. Habitus of Macroderes species. 19, M. oreatus Abdalla &amp; Deschodt, new species; 20, M. cederbergensis Abdalla &amp; Deschodt, new species; 21, M. soleiana Abdalla &amp; Deschodt, new species; 22, M. tourtosus Abdalla &amp; Scholtz, new species; 23, M. leipoldti Abdalla &amp; Deschodt, new species." httpUri="https://zenodo.org/record/2606176/files/figure.png" pageId="7" pageNumber="48">Figs. 20</figureCitation>
,
<figureCitation id="A2E4718DFFA5FFE1FE92F8B9FEF40224" box="[264,294,1865,1889]" captionStart="FIGURES 2439" captionStartId="10.[151,264,1343,1366]" captionTargetBox="[323,1250,205,1286]" captionTargetId="figure@10.[323,1283,195,1303]" captionTargetPageId="10" captionText="FIGURES 2439. Internal sac of aedeagus (24), aedeagus (25) and internal sac sclerite (26-39) of Macroderes species. 26, M. endroedyi Frolov &amp; Scholtz, 2005; 27, M. amplior Frolov &amp; Scholtz, 2005; 28, M. bias (Olivier, 1789); 29, M. cederbergensis Abdalla &amp; Deschodt, new species; 30, M. cornutus Frolov &amp; Scholtz, 2005; 31, M. foveatus Frolov &amp; Scholtz, 2005; 32, M. gifboomi Abdalla &amp; Scholtz, new species; 33, M. leipoldti Abdalla &amp; Deschodt, new species; 34, M. mutilans Kolbe, 1908; 35, M. namakwanus Frolov &amp; Scholtz, 2005; 36, M. oreatus Abdalla &amp; Deschodt, new species; 37, M. porselinus Abdalla, new species; 38, M. soleiana Abdalla &amp; Deschodt, new species; 39, M. tourtosus Abdalla &amp; Scholtz, new species." httpUri="https://zenodo.org/record/2606178/files/figure.png" pageId="7" pageNumber="48">29</figureCitation>
,
<figureCitation id="A2E4718DFFA5FFE1FEA8F8B8FE830224" box="[306,337,1864,1889]" captionStart="FIGURES 7681" captionStartId="20.[151,264,1637,1660]" captionTargetBox="[171,1423,205,1613]" captionTargetId="figure@20.[171,1423,205,1613]" captionTargetPageId="20" captionText="FIGURES 7681. Punctuation of the base of pronotum of Macroderes species. 76, M. cederbergensis Abdalla &amp; Deschodt, new species; 77, M. gifboomi Abdalla &amp; Scholtz, new species; 78, M. oreatus Abdalla &amp; Deschodt, new species; 79, M. porselinus Abdalla, new species; 80, M. soleiana Abdalla &amp; Deschodt, new species; 81, M. leipoldti Abdalla &amp; Deschodt, new species." httpUri="https://zenodo.org/record/2606198/files/figure.png" pageId="7" pageNumber="48">76</figureCitation>
,
<figureCitation id="A2E4718DFFA5FFE1FEC7F8B9FEA90224" box="[349,379,1865,1889]" captionStart="FIGURES 8287" captionStartId="21.[151,264,1505,1528]" captionTargetBox="[316,1276,406,1475]" captionTargetId="figure@21.[316,1276,406,1475]" captionTargetPageId="21" captionText="FIGURES 8287. Pronotum punctures shape of Macroderes species. 82, M. tourtosus Abdalla &amp; Scholtz, new species; 83, M. cederbergensis Abdalla &amp; Deschodt, new species; 84, M. gifboomi Abdalla &amp; Scholtz, new species; 85, M. leipoldti Abdalla &amp; Deschodt, new species; 86, M. oreatus Abdalla &amp; Deschodt, new species; 87, M. porselinus Abdalla, new species." httpUri="https://zenodo.org/record/2606200/files/figure.png" pageId="7" pageNumber="48">83</figureCitation>
,
<figureCitation id="A2E4718DFFA5FFE1FE1CF8B9FE710224" box="[390,419,1865,1889]" captionStart="FIGURES 9499" captionStartId="24.[151,264,1299,1322]" captionTargetBox="[316,1273,193,1274]" captionTargetId="figure@24.[316,1273,193,1274]" captionTargetPageId="24" captionText="FIGURES 9499. Sculpture of second elytra intervals of Macroderes species. 94, M. cederbergensis Abdalla &amp; Deschodt, new species; 95, M. leipoldti Abdalla &amp; Deschodt, new species; 96, M. oreatus Abdalla &amp; Deschodt, new species; 97, M. porselinus Abdalla, new species; 98, M. soleiana Abdalla &amp; Deschodt, new species; 99, M. tourtosus Abdalla &amp; Scholtz, new species." httpUri="https://zenodo.org/record/2606204/files/figure.png" pageId="7" pageNumber="48">94</figureCitation>
)
</paragraph>
<paragraph id="3A606D08FFA5FFE1FF0DF87BFCAF02E4" blockId="7.[151,893,1931,1984]" box="[151,893,1931,1953]" pageId="7" pageNumber="48">
<uri id="4E4E610AFFA5FFE1FF0DF87BFCAF02E4" box="[151,893,1931,1953]" pageId="7" pageNumber="48">
urn:lsid:zoobank.org:act:
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9FED0DC0-CFF6-424C-A3AE-
<date id="4E614BC8FFA5FFE1FD40F87BFCF802E4" box="[730,810,1931,1953]" pageId="7" pageNumber="48" value="56-09-26">092656</date>
B24972
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<treatmentCitationGroup id="1ACF4A26FFA5FFE1FF0DF85AFD3D0285" box="[151,751,1962,1984]" pageId="7" pageNumber="48">
<taxonomicName id="FDDF168BFFA5FFE1FF0DF85AFD5C0285" ID-CoL="3X4TQ" authority="Frolov &amp; Scholtz, 2005: 385" authorityName="in Frolov &amp; Scholtz" authorityPageNumber="385" authorityYear="2005" box="[151,654,1962,1984]" class="Insecta" family="Scarabaeidae" genus="Macroderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="48" phylum="Arthropoda" rank="species" species="minutus">
<emphasis id="08ABB11AFFA5FFE1FF0DF85AFEB70285" box="[151,357,1962,1984]" italics="true" pageId="7" pageNumber="48">Macroderes minutus</emphasis>
<treatmentCitation id="BB7E4B19FFA5FFE1FEF6F85AFD5C0285" author="Frolov, A. V. &amp; Scholtz, C. H." box="[364,654,1962,1984]" page="385" pageId="7" pageNumber="48" year="2005">
<bibRefCitation id="5E4E10F9FFA5FFE1FEF6F85AFD8D0285" author="Frolov, A. V. &amp; Scholtz, C. H." box="[364,607,1962,1984]" pageId="7" pageNumber="48" pagination="373 - 393" refId="ref21694" refString="Frolov, A. V. &amp; Scholtz, C. H. (2005) Revision of the southern African genus Macroderes Westwood (Coleoptera: Scarabaeidae: Scarabaeinae). Annales de la Societe Entomologique de France, 40 (3 - 4), 373 - 393. https: // doi. org / 10.1080 / 00379271.2004.10697429" type="journal article" year="2005">Frolov &amp; Scholtz, 2005</bibRefCitation>
: 385
</treatmentCitation>
</taxonomicName>
(partim).
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</paragraph>
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<subSubSection id="72C53E83FFAAFFEEFF0DFF67FEEE07B5" pageId="8" pageNumber="49" type="materials_examined">
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<emphasis id="08ABB11AFFAAFFEEFF0DFF67FEE705F5" bold="true" box="[151,309,151,176]" pageId="8" pageNumber="49">Type locality.</emphasis>
Cederberg Mts, Jeep Track,
<geoCoordinate id="5FEB0BCFFFAAFFEEFDE2FF67FD0205F5" box="[632,720,151,176]" direction="south" orientation="latitude" pageId="8" pageNumber="49" precision="925" value="-32.433334">32°26'S</geoCoordinate>
<geoCoordinate id="5FEB0BCFFFAAFFEEFD4CFF68FCE105F5" box="[726,819,152,176]" direction="east" orientation="longitude" pageId="8" pageNumber="49" precision="925" value="19.216667">19°13'E</geoCoordinate>
.
</paragraph>
<paragraph id="3A606D08FFAAFFEEFF5DFF4CFAF80778" blockId="8.[151,1436,151,1401]" pageId="8" pageNumber="49">
<emphasis id="08ABB11AFFAAFFEEFF5DFF4CFEAB0590" bold="true" box="[199,377,188,213]" pageId="8" pageNumber="49">Type material.</emphasis>
<materialsCitation id="8AB76755FFAAFFEEFE1FFF4CFCBE0458" ID-GBIF-Occurrence="2028137725" collectingDate="1981-09-01" collectionCode="TMSA" collectorName="Younga" latitude="-32.433334" location="Cederberg Mts" longLatPrecision="1211" longitude="19.216667" municipality="Jeep Track" pageId="8" pageNumber="49" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="E564D3AAFFAAFFEEFE1FFF4CFE220590" box="[389,496,188,213]" pageId="8" pageNumber="49" type="holotype">Holotype</typeStatus>
,
<specimenCount id="2CD9A681FFAAFFEEFD9AFF4DFDC70591" box="[512,533,189,212]" pageId="8" pageNumber="49" type="male"></specimenCount>
(
<collectionCode id="5CCEF5CDFFAAFFEEFDB3FF4CFDAC0591" box="[553,638,188,212]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="8" pageNumber="49">TMSA</collectionCode>
): “
<location id="3F003BD3FFAAFFEEFD05FF4CFC9C0591" LSID="urn:lsid:plazi:treatment:B276DC1EFFA5FFEEFF0DF8D5FBC7003D:3F003BD3FFAAFFEEFD05FF4CFC9C0591" box="[671,846,188,213]" latitude="-32.433334" longLatPrecision="1211" longitude="19.216667" municipality="Jeep Track" name="Cederberg Mts" pageId="8" pageNumber="49">Cederberg Mts</location>
/
<collectingMunicipality id="DA04F772FFAAFFEEFCF7FF4DFC390590" box="[877,1003,188,213]" pageId="8" pageNumber="49">Jeep Track</collectingMunicipality>
/
<geoCoordinate id="5FEB0BCFFFAAFFEEFB91FF4CFBA80591" box="[1035,1146,188,213]" direction="south" orientation="latitude" pageId="8" pageNumber="49" precision="925" value="-32.433334">32° 26' S</geoCoordinate>
<geoCoordinate id="5FEB0BCFFFAAFFEEFB1CFF4CFB250591" box="[1158,1271,188,212]" direction="east" orientation="longitude" pageId="8" pageNumber="49" precision="925" value="19.216667">19° 13' E</geoCoordinate>
/
<date id="4E614BC8FFAAFFEEFA8DFF4CFA5A0591" box="[1303,1416,188,212]" pageId="8" pageNumber="49" value="1981-09-01">
<collectingDate id="5E25B220FFAAFFEEFA8DFF4CFA5A0591" box="[1303,1416,188,212]" pageId="8" pageNumber="49" value="1981-09-01">1.IX.1981</collectingDate>
</date>
/ groundtraps with faeces and meat bait / 63 days / Endrödy
<collectorName id="972A08DEFFAAFFEEFCCDFF10FC7F05BD" box="[855,941,224,248]" pageId="8" pageNumber="49">Younga</collectorName>
leg [printed] //
<typeStatus id="E564D3AAFFAAFFEEFBF2FF10FB2E05BD" box="[1128,1276,224,248]" pageId="8" pageNumber="49" type="holotype">HOLOTYPE</typeStatus>
/
<emphasis id="08ABB11AFFAAFFEEFA8DFF10FE9E045E" italics="true" pageId="8" pageNumber="49">
<taxonomicName id="FDDF168BFFAAFFEEFA8DFF10FE920459" authorityName="Abdalla &amp; Deschodt &amp; Scholtz &amp; Sole" authorityYear="2018" class="Insecta" family="Scarabaeidae" genus="Macroderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="49" phylum="Arthropoda" rank="species" species="cederbergensis">Macroderes cederbergensis</taxonomicName>
/
</emphasis>
Abdalla &amp; Deschodt / 2018 [red label, printed]”
</materialsCitation>
.
<materialsCitation id="8AB76755FFAAFFEEFCEFFEF5FA810420" ID-GBIF-Occurrence="2028137711" collectingDate="1981-09-01" collectionCode="TMSA" collectorName="Younga" latitude="-32.433334" location="Cederberg Mts" longLatPrecision="1211" longitude="19.216667" municipality="Jeep Track" pageId="8" pageNumber="49" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<typeStatus id="E564D3AAFFAAFFEEFCEFFEF5FC340458" box="[885,998,261,285]" pageId="8" pageNumber="49" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="2CD9A681FFAAFFEEFC6AFEF4FBFC0459" box="[1008,1070,260,284]" pageId="8" pageNumber="49" type="female">2 ♀♀</specimenCount>
(
<collectionCode id="5CCEF5CDFFAAFFEEFBA7FEF4FB430459" box="[1085,1169,260,284]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="8" pageNumber="49">TMSA</collectionCode>
): “
<location id="3F003BD3FFAAFFEEFB37FEF4FA840459" LSID="urn:lsid:plazi:treatment:B276DC1EFFA5FFEEFF0DF8D5FBC7003D:3F003BD3FFAAFFEEFB37FEF4FA840459" box="[1197,1366,260,285]" latitude="-32.433334" longLatPrecision="1211" longitude="19.216667" municipality="Jeep Track" name="Cederberg Mts" pageId="8" pageNumber="49">Cederberg Mts</location>
/
<collectingMunicipality id="DA04F772FFAAFFEEFAF0FEF5FF050405" pageId="8" pageNumber="49">Jeep Track</collectingMunicipality>
/
<geoCoordinate id="5FEB0BCFFFAAFFEEFF75FED8FE8A0405" box="[239,344,295,320]" direction="south" orientation="latitude" pageId="8" pageNumber="49" precision="925" value="-32.433334">32° 26' S</geoCoordinate>
<geoCoordinate id="5FEB0BCFFFAAFFEEFEFAFED8FE18047A" box="[352,458,296,320]" direction="east" orientation="longitude" pageId="8" pageNumber="49" precision="925" value="19.216667">19° 13' E</geoCoordinate>
/
<date id="4E614BC8FFAAFFEEFE79FED8FD860405" box="[483,596,296,320]" pageId="8" pageNumber="49" value="1981-09-01">
<collectingDate id="5E25B220FFAAFFEEFE79FED8FD860405" box="[483,596,296,320]" pageId="8" pageNumber="49" value="1981-09-01">1.IX.1981</collectingDate>
</date>
/ groundtraps with faeces and meat bait / 63 days / Endrody
<collectorName id="972A08DEFFAAFFEEFA86FED8FAA00405" box="[1308,1394,296,320]" pageId="8" pageNumber="49">Younga</collectorName>
leg [printed] //
<typeStatus id="E564D3AAFFAAFFEEFE84FEBCFE7B0421" box="[286,425,332,356]" pageId="8" pageNumber="49" type="paratype">PARATYPE</typeStatus>
/
<emphasis id="08ABB11AFFAAFFEEFE5DFEBCFCC10426" box="[455,787,332,356]" italics="true" pageId="8" pageNumber="49">
<taxonomicName id="FDDF168BFFAAFFEEFE5DFEBCFCD20421" authorityName="Abdalla &amp; Deschodt &amp; Scholtz &amp; Sole" authorityYear="2018" box="[455,768,332,356]" class="Insecta" family="Scarabaeidae" genus="Macroderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="49" phylum="Arthropoda" rank="species" species="cederbergensis">Macroderes cederbergensis</taxonomicName>
/
</emphasis>
Abdalla &amp; Deschodt / 2018 [red label, printed]”
</materialsCitation>
;
<materialsCitation id="8AB76755FFAAFFEEFAFCFEBCFF2F0495" ID-GBIF-Occurrence="2028137719" collectingDate="1981-09-01" collectorName="Younga" latitude="-32.383335" location="Cederberg Mts." longLatPrecision="1211" longitude="19.4" municipality="Jeep Track" pageId="8" pageNumber="49" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="2CD9A681FFAAFFEEFAFCFEBCFA450421" box="[1382,1431,332,356]" pageId="8" pageNumber="49" type="male">1 ♂</specimenCount>
: “
<location id="3F003BD3FFAAFFEEFF3BFE9FFE8004C2" LSID="urn:lsid:plazi:treatment:B276DC1EFFA5FFEEFF0DF8D5FBC7003D:3F003BD3FFAAFFEEFF3BFE9FFE8004C2" box="[161,338,367,392]" latitude="-32.383335" longLatPrecision="1211" longitude="19.4" municipality="Jeep Track" name="Cederberg Mts." pageId="8" pageNumber="49">Cederberg Mts.</location>
/
<collectingMunicipality id="DA04F772FFAAFFEEFEF3FE80FE3104CD" box="[361,483,367,392]" pageId="8" pageNumber="49">Jeep Track</collectingMunicipality>
/
<geoCoordinate id="5FEB0BCFFFAAFFEEFE61FE80FDB104CD" box="[507,611,368,392]" direction="south" orientation="latitude" pageId="8" pageNumber="49" precision="925" value="-32.383335">32° 23' S</geoCoordinate>
<geoCoordinate id="5FEB0BCFFFAAFFEEFDF1FE80FD0604C2" box="[619,724,368,392]" direction="east" orientation="longitude" pageId="8" pageNumber="49" precision="925" value="19.4">19° 24' E</geoCoordinate>
/
<date id="4E614BC8FFAAFFEEFD71FE80FC8F04CD" box="[747,861,368,392]" pageId="8" pageNumber="49" value="1981-09-01">
<collectingDate id="5E25B220FFAAFFEEFD71FE80FC8F04CD" box="[747,861,368,392]" pageId="8" pageNumber="49" value="1981-09-01">1.IX.1981</collectingDate>
</date>
/ groundtrap with meat bait / 63 days / Endrödy—
<collectorName id="972A08DEFFAAFFEEFF0DFE65FF3F04E8" box="[151,237,405,429]" pageId="8" pageNumber="49">Younga</collectorName>
leg [printed] //
<typeStatus id="E564D3AAFFAAFFEEFE22FE64FD9104E9" box="[440,579,404,428]" pageId="8" pageNumber="49" type="paratype">PARATYPE</typeStatus>
/
<emphasis id="08ABB11AFFAAFFEEFDFDFE64FC6A04EE" box="[615,952,404,428]" italics="true" pageId="8" pageNumber="49">
<taxonomicName id="FDDF168BFFAAFFEEFDFDFE64FC7004E9" authorityName="Abdalla &amp; Deschodt &amp; Scholtz &amp; Sole" authorityYear="2018" box="[615,930,404,428]" class="Insecta" family="Scarabaeidae" genus="Macroderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="49" phylum="Arthropoda" rank="species" species="cederbergensis">Macroderes cederbergensis</taxonomicName>
/
</emphasis>
Abdalla &amp; Deschodt / 2018 [red label, printed]”
</materialsCitation>
;
<materialsCitation id="8AB76755FFAAFFEEFE97FE48FA9C075D" ID-GBIF-Occurrence="2028137718" collectingDate="1981-09-01" collectionCode="TMSA" collectorName="Endrody-Younga" latitude="-32.4" location="Cederberg Mts" longLatPrecision="1211" longitude="19.166666" municipality="Jeep Track" pageId="8" pageNumber="49" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="2CD9A681FFAAFFEEFE97FE48FEEB048A" box="[269,313,440,464]" pageId="8" pageNumber="49" type="male">1 ♂</specimenCount>
and
<specimenCount id="2CD9A681FFAAFFEEFEEEFE48FE72048A" box="[372,416,440,464]" pageId="8" pageNumber="49" type="female">1 ♀</specimenCount>
(
<collectionCode id="5CCEF5CDFFAAFFEEFE2BFE48FDD40495" box="[433,518,440,464]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="8" pageNumber="49">TMSA</collectionCode>
): “
<location id="3F003BD3FFAAFFEEFDBEFE47FD02048A" LSID="urn:lsid:plazi:treatment:B276DC1EFFA5FFEEFF0DF8D5FBC7003D:3F003BD3FFAAFFEEFDBEFE47FD02048A" box="[548,720,439,464]" latitude="-32.4" longLatPrecision="1211" longitude="19.166666" municipality="Jeep Track" name="Cederberg Mts" pageId="8" pageNumber="49">Cederberg Mts</location>
/
<collectingMunicipality id="DA04F772FFAAFFEEFD73FE48FCB60495" box="[745,868,439,464]" pageId="8" pageNumber="49">Jeep Track</collectingMunicipality>
/
<geoCoordinate id="5FEB0BCFFFAAFFEEFCE4FE48FC3A0495" box="[894,1000,440,464]" direction="south" orientation="latitude" pageId="8" pageNumber="49" precision="925" value="-32.4">32° 24' S</geoCoordinate>
<geoCoordinate id="5FEB0BCFFFAAFFEEFC6BFE48FB8F048A" box="[1009,1117,440,464]" direction="east" orientation="longitude" pageId="8" pageNumber="49" precision="925" value="19.166666">19° 10' E</geoCoordinate>
/
<date id="4E614BC8FFAAFFEEFBEDFE48FB3A0495" box="[1143,1256,440,464]" pageId="8" pageNumber="49" value="1981-09-01">
<collectingDate id="5E25B220FFAAFFEEFBEDFE48FB3A0495" box="[1143,1256,440,464]" pageId="8" pageNumber="49" value="1981-09-01">1.IX.1981</collectingDate>
</date>
/
<taxonomicName id="FDDF168BFFAAFFEEFA98FE48FA4E0495" box="[1282,1436,440,464]" class="Mammalia" family="Bovidae" genus="Redunca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="8" pageNumber="49" phylum="Chordata" rank="species" species="rufo">
<emphasis id="08ABB11AFFAAFFEEFA98FE48FA4E0495" box="[1282,1436,440,464]" italics="true" pageId="8" pageNumber="49">Redunca rufo</emphasis>
</taxonomicName>
[name could not be traced, probably
<taxonomicName id="FDDF168BFFAAFFEEFDA7FE2CFC7704B1" authority="(Forster, 1790)" authorityName="Forster" authorityYear="1790" baseAuthorityName="Forster" baseAuthorityYear="1790" box="[573,933,476,500]" class="Mammalia" family="Bovidae" genus="Pelea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="8" pageNumber="49" phylum="Chordata" rank="species" species="capreolus">
<emphasis id="08ABB11AFFAAFFEEFDA7FE2CFD2104B1" box="[573,755,476,500]" italics="true" pageId="8" pageNumber="49">Pelea capreolus</emphasis>
(Forster, 1790)
</taxonomicName>
or Grey Rhebok] dung /
<collectorName id="972A08DEFFAAFFEEFB50FE2CFA4E04B0" box="[1226,1436,476,501]" pageId="8" pageNumber="49">Endrödy—Younga</collectorName>
leg [printed] //
<typeStatus id="E564D3AAFFAAFFEEFED8FDF0FE1F075D" box="[322,461,512,536]" pageId="8" pageNumber="49" type="paratype">PARATYPE</typeStatus>
/
<emphasis id="08ABB11AFFAAFFEEFE7EFDF0FCFA0752" box="[484,808,512,536]" italics="true" pageId="8" pageNumber="49">
<taxonomicName id="FDDF168BFFAAFFEEFE7EFDF0FCCA075D" authorityName="Abdalla &amp; Deschodt &amp; Scholtz &amp; Sole" authorityYear="2018" box="[484,792,512,536]" class="Insecta" family="Scarabaeidae" genus="Macroderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="49" phylum="Arthropoda" rank="species" species="cederbergensis">Macroderes cederbergensis</taxonomicName>
/
</emphasis>
Abdalla &amp; Deschodt / 2018 [red label, printed]”
</materialsCitation>
;
<materialsCitation id="8AB76755FFAAFFEEFAC6FDF0FAF40778" ID-GBIF-Occurrence="2028137724" collectingDate="1981-09-01" collectionCode="TMSA" collectorName="Endrody-Younga" latitude="-32.4" location="Cederberg Mts" longLatPrecision="1211" longitude="19.166666" municipality="Jeep Track" pageId="8" pageNumber="49" specimenCount="2" specimenCount-male="2" typeStatus="paratype">
<specimenCount id="2CD9A681FFAAFFEEFAC6FDF0FA4E0752" box="[1372,1436,512,536]" pageId="8" pageNumber="49" type="male">2 ♂♂</specimenCount>
(
<collectionCode id="5CCEF5CDFFAAFFEEFF04FDD4FF3D0779" box="[158,239,548,572]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="8" pageNumber="49">TMSA</collectionCode>
) with the same data but collected from groundtraps with meat and feaces bait set up for 63 days
</materialsCitation>
.
</paragraph>
<paragraph id="3A606D08FFAAFFEEFF5DFDB7FBBB071A" blockId="8.[151,1436,151,1401]" box="[199,1129,583,608]" pageId="8" pageNumber="49">
<emphasis id="08ABB11AFFAAFFEEFF5DFDB7FE1A0725" bold="true" box="[199,456,583,608]" pageId="8" pageNumber="49">
<typeStatus id="E564D3AAFFAAFFEEFF5DFDB7FEE60725" box="[199,308,583,608]" pageId="8" pageNumber="49" type="holotype">Holotype</typeStatus>
description.
</emphasis>
<typeStatus id="E564D3AAFFAAFFEEFE54FDB7FDEB0725" box="[462,569,583,608]" pageId="8" pageNumber="49" type="holotype">Holotype</typeStatus>
,
<specimenCount id="2CD9A681FFAAFFEEFDDFFDB8FD88071A" box="[581,602,584,607]" pageId="8" pageNumber="49" type="male"></specimenCount>
(
<figureCitation id="A2E4718DFFAAFFEEFDF3FDB7FD6C0725" box="[617,702,583,608]" captionStart="FIGURES 1923" captionStartId="9.[151,264,1928,1951]" captionTargetBox="[411,1172,706,1874]" captionTargetId="figure@9.[337,1297,671,1926]" captionTargetPageId="9" captionText="FIGURES 1923. Habitus of Macroderes species. 19, M. oreatus Abdalla &amp; Deschodt, new species; 20, M. cederbergensis Abdalla &amp; Deschodt, new species; 21, M. soleiana Abdalla &amp; Deschodt, new species; 22, M. tourtosus Abdalla &amp; Scholtz, new species; 23, M. leipoldti Abdalla &amp; Deschodt, new species." httpUri="https://zenodo.org/record/2606176/files/figure.png" pageId="8" pageNumber="49">Fig. 20</figureCitation>
). Body length
<quantity id="FD27C0EDFFAAFFEEFCFBFDB8FC6B071A" box="[865,953,584,608]" metricMagnitude="-3" metricUnit="m" metricValue="9.4" pageId="8" pageNumber="49" unit="mm" value="9.4">9.4 mm</quantity>
, width
<quantity id="FD27C0EDFFAAFFEEFB96FDB7FBB7071A" box="[1036,1125,583,608]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="8" pageNumber="49" unit="mm" value="6.8">6.8 mm</quantity>
.
</paragraph>
<paragraph id="3A606D08FFAAFFEEFF5DFD9CFEEE07B5" blockId="8.[151,1436,151,1401]" pageId="8" pageNumber="49">
<emphasis id="08ABB11AFFAAFFEEFF5DFD9CFEDF07C0" bold="true" box="[199,269,620,645]" pageId="8" pageNumber="49">Head.</emphasis>
Genae rightangled. Frontal suture visible, not raised, strongly curved, no tubercle. Genal sutures barely discernible. Dorsal surface of clypeus behind frontal suture punctate punctures small, irregular and separated by 1 3 puncture diameters, area between punctures slightly rugose proximally of frontal suture. Dorsal part of eyes clearly visible.
</paragraph>
</subSubSection>
<subSubSection id="72C53E83FFAAFFEEFF5DFD0CFD860138" pageId="8" pageNumber="49" type="description">
<paragraph id="3A606D08FFAAFFEEFF5DFD0CFCA60619" blockId="8.[151,1436,151,1401]" pageId="8" pageNumber="49">
<emphasis id="08ABB11AFFAAFFEEFF5DFD0CFE950650" bold="true" box="[199,327,764,789]" pageId="8" pageNumber="49">Pronotum.</emphasis>
Slightly convex. Anterior lateral angles almost rightangled, posterior lateral angles fairly rounded. No excavation antero-laterally. Lateral border not punctate. Dorsal surface smooth, with regular punctures separated by 0.51.5 puncture diameters (
<figureCitation id="A2E4718DFFAAFFEEFD45FCB4FCEE0618" box="[735,828,836,861]" captionStart="FIGURES 7681" captionStartId="20.[151,264,1637,1660]" captionTargetBox="[171,1423,205,1613]" captionTargetId="figure@20.[171,1423,205,1613]" captionTargetPageId="20" captionText="FIGURES 7681. Punctuation of the base of pronotum of Macroderes species. 76, M. cederbergensis Abdalla &amp; Deschodt, new species; 77, M. gifboomi Abdalla &amp; Scholtz, new species; 78, M. oreatus Abdalla &amp; Deschodt, new species; 79, M. porselinus Abdalla, new species; 80, M. soleiana Abdalla &amp; Deschodt, new species; 81, M. leipoldti Abdalla &amp; Deschodt, new species." httpUri="https://zenodo.org/record/2606198/files/figure.png" pageId="8" pageNumber="49">Figs. 76</figureCitation>
,
<figureCitation id="A2E4718DFFAAFFEEFCD2FCB4FCB50619" box="[840,871,836,860]" captionStart="FIGURES 8287" captionStartId="21.[151,264,1505,1528]" captionTargetBox="[316,1276,406,1475]" captionTargetId="figure@21.[316,1276,406,1475]" captionTargetPageId="21" captionText="FIGURES 8287. Pronotum punctures shape of Macroderes species. 82, M. tourtosus Abdalla &amp; Scholtz, new species; 83, M. cederbergensis Abdalla &amp; Deschodt, new species; 84, M. gifboomi Abdalla &amp; Scholtz, new species; 85, M. leipoldti Abdalla &amp; Deschodt, new species; 86, M. oreatus Abdalla &amp; Deschodt, new species; 87, M. porselinus Abdalla, new species." httpUri="https://zenodo.org/record/2606200/files/figure.png" pageId="8" pageNumber="49">83</figureCitation>
).
</paragraph>
<paragraph id="3A606D08FFAAFFEEFF5DFC97FB2F06A8" blockId="8.[151,1436,151,1401]" pageId="8" pageNumber="49">
<emphasis id="08ABB11AFFAAFFEEFF5DFC97FEC806C5" bold="true" box="[199,282,871,896]" pageId="8" pageNumber="49">Elytra.</emphasis>
Elytral interstriae 18 flat, shagreened, irregularly punctate, punctures irregularly sized, separated by 0.53 puncture diameters, margins of punctures not carinate (
<figureCitation id="A2E4718DFFAAFFEEFCD0FC7CFC4D06E1" box="[842,927,908,933]" captionStart="FIGURES 9499" captionStartId="24.[151,264,1299,1322]" captionTargetBox="[316,1273,193,1274]" captionTargetId="figure@24.[316,1273,193,1274]" captionTargetPageId="24" captionText="FIGURES 9499. Sculpture of second elytra intervals of Macroderes species. 94, M. cederbergensis Abdalla &amp; Deschodt, new species; 95, M. leipoldti Abdalla &amp; Deschodt, new species; 96, M. oreatus Abdalla &amp; Deschodt, new species; 97, M. porselinus Abdalla, new species; 98, M. soleiana Abdalla &amp; Deschodt, new species; 99, M. tourtosus Abdalla &amp; Scholtz, new species." httpUri="https://zenodo.org/record/2606204/files/figure.png" pageId="8" pageNumber="49">Fig. 94</figureCitation>
). Striae punctate with punctures separated by 24 puncture diameters. Sutural interstria narrow with single row of irregularly spaced punctures. Elytral stria 9 2/ 3 the length of elytron; interstria 10 about 5 times narrower than interstria
<quantity id="FD27C0EDFFAAFFEEFC42FC24FBD106A8" box="[984,1027,980,1005]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" pageId="8" pageNumber="49" unit="in" value="9.0">9 in</quantity>
the middle of elytron.
</paragraph>
<paragraph id="3A606D08FFAAFFEEFF5DFC07FB0B0155" blockId="8.[151,1436,151,1401]" box="[199,1241,1015,1040]" pageId="8" pageNumber="49">
<emphasis id="08ABB11AFFAAFFEEFF5DFC07FE930155" bold="true" box="[199,321,1015,1040]" pageId="8" pageNumber="49">Aedeagus.</emphasis>
The sclerite of internal sac is curved, with small lateral process apically (
<figureCitation id="A2E4718DFFAAFFEEFBE2FC07FB1F0155" box="[1144,1229,1015,1040]" captionStart="FIGURES 2439" captionStartId="10.[151,264,1343,1366]" captionTargetBox="[323,1250,205,1286]" captionTargetId="figure@10.[323,1283,195,1303]" captionTargetPageId="10" captionText="FIGURES 2439. Internal sac of aedeagus (24), aedeagus (25) and internal sac sclerite (26-39) of Macroderes species. 26, M. endroedyi Frolov &amp; Scholtz, 2005; 27, M. amplior Frolov &amp; Scholtz, 2005; 28, M. bias (Olivier, 1789); 29, M. cederbergensis Abdalla &amp; Deschodt, new species; 30, M. cornutus Frolov &amp; Scholtz, 2005; 31, M. foveatus Frolov &amp; Scholtz, 2005; 32, M. gifboomi Abdalla &amp; Scholtz, new species; 33, M. leipoldti Abdalla &amp; Deschodt, new species; 34, M. mutilans Kolbe, 1908; 35, M. namakwanus Frolov &amp; Scholtz, 2005; 36, M. oreatus Abdalla &amp; Deschodt, new species; 37, M. porselinus Abdalla, new species; 38, M. soleiana Abdalla &amp; Deschodt, new species; 39, M. tourtosus Abdalla &amp; Scholtz, new species." httpUri="https://zenodo.org/record/2606178/files/figure.png" pageId="8" pageNumber="49">Fig. 29</figureCitation>
).
</paragraph>
<paragraph id="3A606D08FFAAFFEEFF5DFBECFA5F0171" blockId="8.[151,1436,151,1401]" box="[199,1421,1052,1077]" pageId="8" pageNumber="49">
<emphasis id="08ABB11AFFAAFFEEFF5DFBECFE9B0170" bold="true" box="[199,329,1052,1077]" pageId="8" pageNumber="49">Size range.</emphasis>
Males length: 7.59.0 mm, width:
<quantity id="FD27C0EDFFAAFFEEFD4EFBECFC8C0171" box="[724,862,1052,1077]" metricMagnitude="-3" metricUnit="m" metricValue="6.15" metricValueMax="6.8" metricValueMin="5.5" pageId="8" pageNumber="49" unit="mm" value="6.15" valueMax="6.8" valueMin="5.5">5.56.8 mm</quantity>
; females length:
<quantity id="FD27C0EDFFAAFFEEFB86FBECFB770171" box="[1052,1189,1052,1077]" metricMagnitude="-3" metricUnit="m" metricValue="8.65" metricValueMax="9.5" metricValueMin="7.8" pageId="8" pageNumber="49" unit="mm" value="8.65" valueMax="9.5" valueMin="7.8">7.89.5 mm</quantity>
, width: 6.0
<quantity id="FD27C0EDFFAAFFEEFAABFBECFA5B0171" box="[1329,1417,1052,1077]" metricMagnitude="-3" metricUnit="m" metricValue="6.3" pageId="8" pageNumber="49" unit="mm" value="6.3">6.3 mm</quantity>
.
</paragraph>
<paragraph id="3A606D08FFAAFFEEFF5DFBCFFD860138" blockId="8.[151,1436,151,1401]" pageId="8" pageNumber="49">
<emphasis id="08ABB11AFFAAFFEEFF5DFBCFFEF7011D" bold="true" box="[199,293,1087,1112]" pageId="8" pageNumber="49">Female.</emphasis>
Female differs from male by having elytral interstriae 18 less shagreened; otherwise variation between male and female is very slight.
</paragraph>
</subSubSection>
<subSubSection id="72C53E83FFAAFFEEFF5DFB77FB2801E5" box="[199,1274,1159,1185]" pageId="8" pageNumber="49" type="etymology">
<paragraph id="3A606D08FFAAFFEEFF5DFB77FB2801E5" blockId="8.[151,1436,151,1401]" box="[199,1274,1159,1185]" pageId="8" pageNumber="49">
<emphasis id="08ABB11AFFAAFFEEFF5DFB77FE9C01E5" bold="true" box="[199,334,1159,1184]" pageId="8" pageNumber="49">Etymology.</emphasis>
The specific epithet is derived from the Cederberg Range where the species occurs.
</paragraph>
</subSubSection>
<subSubSection id="72C53E83FFAAFFEEFF5DFB5CFAE801AD" pageId="8" pageNumber="49" type="diagnosis">
<paragraph id="3A606D08FFAAFFEEFF5DFB5CFAE801AD" blockId="8.[151,1436,151,1401]" pageId="8" pageNumber="49">
<emphasis id="08ABB11AFFAAFFEEFF5DFB5CFE020180" bold="true" box="[199,464,1196,1221]" pageId="8" pageNumber="49">Differential diagnosis.</emphasis>
It is close to
<taxonomicName id="FDDF168BFFAAFFEEFDE9FB5DFD280181" authority="Abdalla &amp; Deschodt &amp; Scholtz &amp; Sole, 2018" authorityName="Abdalla &amp; Deschodt &amp; Scholtz &amp; Sole" authorityYear="2018" box="[627,762,1196,1220]" class="Insecta" family="Scarabaeidae" genus="Macroderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="49" phylum="Arthropoda" rank="species" species="soleiana" status="sp. nov.">
<emphasis id="08ABB11AFFAAFFEEFDE9FB5DFD280181" box="[627,762,1196,1220]" italics="true" pageId="8" pageNumber="49">M. soleiana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="13980C61FFAAFFEEFC9EFB5DFC5F0180" box="[772,909,1196,1221]" pageId="8" pageNumber="49" rank="species">new species</taxonomicNameLabel>
but can be separated by having flatter elytral intervals (
<figureCitation id="A2E4718DFFAAFFEEFE9CFB3FFEB601AD" box="[262,356,1231,1256]" captionStart="FIGURES 1923" captionStartId="9.[151,264,1928,1951]" captionTargetBox="[411,1172,706,1874]" captionTargetId="figure@9.[337,1297,671,1926]" captionTargetPageId="9" captionText="FIGURES 1923. Habitus of Macroderes species. 19, M. oreatus Abdalla &amp; Deschodt, new species; 20, M. cederbergensis Abdalla &amp; Deschodt, new species; 21, M. soleiana Abdalla &amp; Deschodt, new species; 22, M. tourtosus Abdalla &amp; Scholtz, new species; 23, M. leipoldti Abdalla &amp; Deschodt, new species." httpUri="https://zenodo.org/record/2606176/files/figure.png" pageId="8" pageNumber="49">Figs. 20</figureCitation>
,
<figureCitation id="A2E4718DFFAAFFEEFEEAFB20FE5F01AD" box="[368,397,1232,1256]" captionStart="FIGURES 9499" captionStartId="24.[151,264,1299,1322]" captionTargetBox="[316,1273,193,1274]" captionTargetId="figure@24.[316,1273,193,1274]" captionTargetPageId="24" captionText="FIGURES 9499. Sculpture of second elytra intervals of Macroderes species. 94, M. cederbergensis Abdalla &amp; Deschodt, new species; 95, M. leipoldti Abdalla &amp; Deschodt, new species; 96, M. oreatus Abdalla &amp; Deschodt, new species; 97, M. porselinus Abdalla, new species; 98, M. soleiana Abdalla &amp; Deschodt, new species; 99, M. tourtosus Abdalla &amp; Scholtz, new species." httpUri="https://zenodo.org/record/2606204/files/figure.png" pageId="8" pageNumber="49">94</figureCitation>
) and the striae being slightly arcuate and not straight as in
<taxonomicName id="FDDF168BFFAAFFEEFBBBFB21FB7701AD" authority="Abdalla &amp; Deschodt &amp; Scholtz &amp; Sole, 2018" authorityName="Abdalla &amp; Deschodt &amp; Scholtz &amp; Sole" authorityYear="2018" box="[1057,1189,1232,1256]" class="Insecta" family="Scarabaeidae" genus="Macroderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="49" phylum="Arthropoda" rank="species" species="soleiana" status="sp. nov.">
<emphasis id="08ABB11AFFAAFFEEFBBBFB21FB7701AD" box="[1057,1189,1232,1256]" italics="true" pageId="8" pageNumber="49">M. soleiana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="13980C61FFAAFFEEFB36FB21FAE401AD" box="[1196,1334,1231,1256]" pageId="8" pageNumber="49" rank="species">new species</taxonomicNameLabel>
.
</paragraph>
</subSubSection>
<subSubSection id="72C53E83FFAAFFEEFF5DFB04FA840010" pageId="8" pageNumber="49" type="description">
<paragraph id="3A606D08FFAAFFEEFF5DFB04FA840010" blockId="8.[151,1436,151,1401]" pageId="8" pageNumber="49">
<emphasis id="08ABB11AFFAAFFEEFF5DFB04FE270048" bold="true" box="[199,501,1268,1293]" pageId="8" pageNumber="49">Habitat and distribution.</emphasis>
This species was collected on the highest plateau in the Cederberg range (
<figureCitation id="A2E4718DFFAAFFEEFAD2FB04FA5C0049" box="[1352,1422,1268,1293]" captionStart="FIGURE 2" captionStartId="2.[151,250,1574,1596]" captionText="FIGURE 2. Distribution map of Macroderes species." pageId="8" pageNumber="49">Fig. 2</figureCitation>
), above
<quantity id="FD27C0EDFFAAFFEEFF79FAE8FEEB0075" box="[227,313,1304,1328]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="8" pageNumber="49" unit="m" value="1100.0">1100 m</quantity>
a.s.l. (metres above sea level), characterised by clay soils and classified as Northern Inland Shale Band Vegetation (
<bibRefCitation id="5E4E10F9FFAAFFEEFEC5FACCFD4B0010" author="Mucina, L. &amp; Rutherford, M. C." box="[351,665,1340,1365]" pageId="8" pageNumber="49" pagination="1 - 807" refId="ref21930" refString="Mucina, L. &amp; Rutherford, M. C. (Eds.) (2006) The Vegetation of South Africa, Lesotho and Swaziland. Strelitzia, 19, 1 - 807." type="journal article" year="2006">Mucina &amp; Rutherford 2006</bibRefCitation>
).
<taxonomicName id="FDDF168BFFAAFFEEFD31FACCFC440011" authority="Abdalla &amp; Deschodt &amp; Scholtz &amp; Sole, 2018" authorityName="Abdalla &amp; Deschodt &amp; Scholtz &amp; Sole" authorityYear="2018" box="[683,918,1340,1364]" class="Insecta" family="Scarabaeidae" genus="Macroderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="49" phylum="Arthropoda" rank="species" species="soleiana" status="sp. nov.">
<emphasis id="08ABB11AFFAAFFEEFD31FACCFC440011" box="[683,918,1340,1364]" italics="true" pageId="8" pageNumber="49">Macroderes soleiana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="13980C61FFAAFFEEFC07FACDFBF60010" box="[925,1060,1340,1365]" pageId="8" pageNumber="49" rank="species">new species</taxonomicNameLabel>
occurs below
<quantity id="FD27C0EDFFAAFFEEFB5CFACCFAC90011" box="[1222,1307,1340,1364]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="8" pageNumber="49" unit="m" value="1100.0">1100 m</quantity>
a.s.l.
</paragraph>
</subSubSection>
<subSubSection id="72C53E83FFAAFFEEFF5DFAAFFBC7003D" box="[199,1045,1375,1401]" pageId="8" pageNumber="49" type="discussion">
<paragraph id="3A606D08FFAAFFEEFF5DFAAFFBC7003D" blockId="8.[151,1436,151,1401]" box="[199,1045,1375,1401]" pageId="8" pageNumber="49">
<emphasis id="08ABB11AFFAAFFEEFF5DFAAFFEE2003D" bold="true" box="[199,304,1375,1400]" pageId="8" pageNumber="49">Remark.</emphasis>
See
<taxonomicName id="FDDF168BFFAAFFEEFEFCFA91FE370032" authorityName="in Frolov &amp; Scholtz" authorityYear="2005" box="[358,485,1376,1400]" class="Insecta" family="Scarabaeidae" genus="Macroderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="49" phylum="Arthropoda" rank="species" species="minutus">
<emphasis id="08ABB11AFFAAFFEEFEFCFA91FE370032" box="[358,485,1376,1400]" italics="true" pageId="8" pageNumber="49">M. minutus</emphasis>
</taxonomicName>
for an explanation of the origin of the
<typeStatus id="E564D3AAFFAAFFEEFC0DFA91FC15003C" box="[919,967,1377,1401]" pageId="8" pageNumber="49">type</typeStatus>
series.
</paragraph>
</subSubSection>
</treatment>
</document>