treatments-xml/data/B2/6F/42/B26F4236273B5D53BF06D29EBBBFCB92.xml

<document ID-CLB-Dataset="124456" ID-DOI="http://dx.doi.org/10.3897/asp.80.e81736" ID-GBIF-Dataset="a3d72dfa-8129-4755-a46f-27e2eaa0bca7" ID-Pensoft-Pub="1864-8312-80-411" ID-Pensoft-UUID="775020E4F468563291A208DB113C16F3" ID-ZooBank="48AB3E3B392849CAB26D4B22839FA7D5" ModsDocID="1864-8312-80-411" checkinTime="1660884374599" checkinUser="pensoft" docAuthor="Li, Yan-Da, Zhang, Yu-Bo, Szawaryn, Karol, Huang, Di-Ying &amp; Cai, Chen-Yang" docDate="2022" docId="B26F4236273B5D53BF06D29EBBBFCB92" docLanguage="en" docName="ArthropodSystPhyl 80: 411-422" docOrigin="Arthropod Systematics &amp; amp; Phylogeny 80" docPubDate="2022-08-18" docSource="http://dx.doi.org/10.3897/asp.80.e81736" docTitle="Xenostanus jiangkuni Li, Szawaryn &amp; Cai 2022, sp. nov." docType="treatment" docUuid="DF781A4B-9A46-460E-AD27-BDC09BCB1958" docUuidSource="ZooBank" docVersion="5" id="775020E4F468563291A208DB113C16F3" lastPageNumber="411" masterDocId="775020E4F468563291A208DB113C16F3" masterDocTitle="Earliest fossil record of Corylophidae from Burmese amber and phylogeny of Corylophidae (Coleoptera: Coccinelloidea)" masterLastPageNumber="422" masterPageNumber="411" pageNumber="411" updateTime="1732850350211" updateUser="ExternalLinkService">
<mods:mods id="67FE985AA9640D6162047F05D60BC2EA" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D028EB53DA7FC34CD08A045ED7FB9260">
<mods:title id="BFB827C0F7C3EB79030734B318FC2183">Earliest fossil record of Corylophidae from Burmese amber and phylogeny of Corylophidae (Coleoptera: Coccinelloidea)</mods:title>
</mods:titleInfo>
<mods:name id="2BC1A4BC2F877F4F451BDCA9A7055D1E" type="personal">
<mods:role id="40A76AF05A2B39B5BAE156F20DBB9FB2">
<mods:roleTerm id="0062C6735E013CEC9C10572CCFEC66A7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="53BBD0FD41EB7B30634CB4DC76EAA5EF">Li, Yan-Da</mods:namePart>
<mods:nameIdentifier id="D2EF84E475818EC8CB2905CB288F5653" type="ORCID">https://orcid.org/0000-0002-9439-202X</mods:nameIdentifier>
<mods:affiliation id="E664B0C52BAAE75752B3D2132F2367EE">State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, and Center for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, Nanjing 210008, China; Yan-Da Li [ydli @ pku. edu. cn]; Di-Ying Huang [dyhuang @ nigpas. ac. cn] &amp; School of Earth Sciences, University of Bristol, Life Sciences Building, Tyndall Avenue, Bristol, BS 8 1 TQ, United Kingdom</mods:affiliation>
</mods:name>
<mods:name id="847492C582FA538BAB9A4F9660773788" type="personal">
<mods:role id="BD64F9D79D58F783E445DDC5DBD902CB">
<mods:roleTerm id="7B188A70B69CB1845863EDDF28CD9C96">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="561D2B8CC043C7E072D39F165A9DCFD7">Zhang, Yu-Bo</mods:namePart>
<mods:affiliation id="260CAFFE065C7C1B13EA19427F3174B2">State Key Laboratory of Protein and Plant Gene Research, and Peking-Tsinghua Center for Life Sciences, Academy for Advanced Interdisciplinary Studies, Peking University, Beijing 100871, China; Yu-Bo Zhang [yubozhang @ pku. edu. cn]</mods:affiliation>
</mods:name>
<mods:name id="FF6A8759E84EC8B2FF6E9C0849C838F1" type="personal">
<mods:role id="A585A10ABE1F201D0DDD4BA69C7381EF">
<mods:roleTerm id="12F4B34D994F7CE2FCC8FFCC4D849B78">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9FB7C731AE8673D5A258BC2848349C62">Szawaryn, Karol</mods:namePart>
<mods:nameIdentifier id="781741CADB8D42E5F85A0593F24506CD" type="ORCID">https://orcid.org/0000-0002-9329-4268</mods:nameIdentifier>
<mods:affiliation id="8BD5A240EB620801EB4BFA0892682F4C">Museum and Institute of Zoology, Polish Academy of Sciences, Wilcza 64, Warsaw, Poland; Karol Szawaryn [k. szawaryn @ gmail. com]</mods:affiliation>
</mods:name>
<mods:name id="D93AD36DF06264FD1A53ECCC4D96E36C" type="personal">
<mods:role id="FC143F1B4AC9D691BD284B7B33BC9D5A">
<mods:roleTerm id="34BDC830B80BEDC0B540BBD4BE56CF96">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AFB3705015D06C4867D3B57E15E578FE">Huang, Di-Ying</mods:namePart>
<mods:nameIdentifier id="B25EB4B95989FDFF5E8656226D563E95" type="ORCID">https://orcid.org/0000-0002-5637-4867</mods:nameIdentifier>
<mods:affiliation id="76079D6A9BE9EA20D4D7D194C4CABDD9">State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, and Center for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, Nanjing 210008, China; Yan-Da Li [ydli @ pku. edu. cn]; Di-Ying Huang [dyhuang @ nigpas. ac. cn]</mods:affiliation>
</mods:name>
<mods:name id="E62710E1FDBCA0FD4892AADF6F56A78A" type="personal">
<mods:role id="2B883DEC99E77B93DBE0D67BC99C7834">
<mods:roleTerm id="E9E1D46FAE8F5BF9D7D600F7A96E50D9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="59E8550AA5A11EA8E67478DA30C98797">Cai, Chen-Yang</mods:namePart>
<mods:nameIdentifier id="F608F707AB53F60CAB89A59CD8D60524" type="ORCID">https://orcid.org/0000-0002-9283-8323</mods:nameIdentifier>
<mods:affiliation id="4429830B0F3EDAE33BF588D9BBD8696F">State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, and Center for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, Nanjing 210008, China; Yan-Da Li [ydli @ pku. edu. cn]; Di-Ying Huang [dyhuang @ nigpas. ac. cn] &amp; School of Earth Sciences, University of Bristol, Life Sciences Building, Tyndall Avenue, Bristol, BS 8 1 TQ, United Kingdom</mods:affiliation>
<mods:nameIdentifier id="6346084BC33C355C9329823B5F123ED9" type="email">cycai@nigpas.ac.cn</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="67B8816BE922C3FC8EEA71B2802A38C2">text</mods:typeOfResource>
<mods:relatedItem id="3D70553186FF8EA629BE5EECB3849759" type="host">
<mods:titleInfo id="BB8E1EAB27826573A47E003CF39C5003">
<mods:title id="D1074F8E629A04130FEBD64F9B6186AD">Arthropod Systematics &amp; amp; Phylogeny</mods:title>
</mods:titleInfo>
<mods:part id="36510ED1089673F52F581560A80D39F0">
<mods:date id="DC3E2CD5F036B06DFDB73FDB8909C0D3">2022</mods:date>
<mods:detail id="648025A3367AE19336A2298636CADE8B" type="pubDate">
<mods:number id="4C85F1081537BB9B252215AEC3C6BD9D">2022-08-18</mods:number>
</mods:detail>
<mods:detail id="179ACF9653A4B1D0A262C5F91CDE1921" type="volume">
<mods:number id="B1925AE0BA6DB5809E7B9AC238EC3827">80</mods:number>
</mods:detail>
<mods:extent id="774FDE6AE30C2A81C61359388943246A" unit="page">
<mods:start id="CEF866253065941DD77BC69C175CB2DB">411</mods:start>
<mods:end id="FDB6FE7DBCEF3E8402800B475DFA89B6">422</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="F53B593E90E2D2193E23BB7496734D12">
<mods:url id="278069DC813A4F45D7E476E2E506803B">http://dx.doi.org/10.3897/asp.80.e81736</mods:url>
</mods:location>
<mods:classification id="0B6792F3AE7C6A8CA05F4BC88950773F">journal article</mods:classification>
<mods:identifier id="E139ADF5B899A21E01536CB8FA4DE969" type="DOI">http://dx.doi.org/10.3897/asp.80.e81736</mods:identifier>
<mods:identifier id="72308D63635BAF29F96C54699F8952B4" type="Pensoft-Pub">1864-8312-80-411</mods:identifier>
<mods:identifier id="FA82990D95282688725A2589AB575757" type="ZooBank">48AB3E3B392849CAB26D4B22839FA7D5</mods:identifier>
<mods:identifier id="2994CB2F501E35B9A605EFFC7440C9FC" type="Pensoft-UUID">775020E4F468563291A208DB113C16F3</mods:identifier>
</mods:mods>
<subSection id="913E8BC67F27C70E0F79A50B5464001E" pageId="0" pageNumber="411" type="3. systematic paleontology">
<treatment id="B26F4236273B5D53BF06D29EBBBFCB92" ID-GBIF-Taxon="199504930" LSID="urn:lsid:zoobank.org:act:DF781A4B-9A46-460E-AD27-BDC09BCB1958" httpUri="http://treatment.plazi.org/id/B26F4236273B5D53BF06D29EBBBFCB92" lastPageNumber="411" pageId="0" pageNumber="411" scope_family="Corylophidae" scope_order="Coleoptera" scope_subOrder="Polyphaga" scope_superFamily="Coccinelloidea" scope_tribe="Xenostanini">
<subSubSection id="63014A531F20D0B7BC0078D48DBD2676" pageId="0" pageNumber="411" type="nomenclature">
<paragraph id="39E4106BF9856B05EE0B48B0E66AB846" pageId="0" pageNumber="411">
<taxonomicName id="CAEF5B69D180228323463171C389713E" LSID="http://zoobank.org/DF781A4B-9A46-460E-AD27-BDC09BCB1958" authority="Li, Szawaryn &amp; Cai" authorityName="Li, Szawaryn &amp; Cai" authorityYear="2022" class="Insecta" family="Corylophidae" genus="Xenostanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xenostanus jiangkuni" order="Coleoptera" pageId="0" pageNumber="411" phylum="Arthropoda" rank="species" species="jiangkuni" status="sp. nov.">Xenostanus jiangkuni Li, Szawaryn &amp; Cai</taxonomicName>
<taxonomicNameLabel id="783F4D946F8BB3A128A8A056F89B111C" pageId="0" pageNumber="411">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="6E4B2E44331AEDDE68CAB906FEE6F2B3" pageId="0" pageNumber="411" type="description">
<paragraph id="9EDD8CDFEA1CF3D3E4F0130C5A39DBB7" pageId="0" pageNumber="411">
<figureCitation id="147E5D47A021AB0C01E91DEEB54E29A8" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. General habitus of Xenostanus jiangkuni Li, Szawaryn &amp; Cai gen. et sp. nov., holotype, NIGP 177782, under incident light. A Dorsal view. B Ventral view. Scale bars: 400 μm." figureDoi="10.3897/asp.80.e81736.figure1" httpUri="https://binary.pensoft.net/fig/730768" pageId="0" pageNumber="411">Figs 1</figureCitation>
<figureCitation id="8B777B6E78B0EEC73988F5C18351CFF4" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. General habitus of Xenostanus jiangkuni Li, Szawaryn &amp; Cai gen. et sp. nov., holotype, NIGP 177782, under confocal microscopy. A Dorsal view. B Ventral view, with arrowheads indicating the metaventral and abdominal postcoxal lines. Scale bars: 400 μm." figureDoi="10.3897/asp.80.e81736.figure2" httpUri="https://binary.pensoft.net/fig/730769" pageId="0" pageNumber="411">, 2</figureCitation>
<figureCitation id="1F15592B8D37887CC1B533107F18B7C2" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Details of Xenostanus jiangkuni Li, Szawaryn &amp; Cai gen. et sp. nov., holotype, NIGP 177782, under confocal microscopy. A Head and prothorax, ventral view. B Head, dorsal view. C Head and prothorax, lateral view. D Abdominal base, ventral view. Abbreviations: an, antenna; ey, compound eye; lb, labrum; lbp, labial palp; md, mandible; msf, mesofemur; msts, mesotarsus; mtf, metafemur; mttb, metatibia; mtts, metatarsus; mtv, metaventrite; mxp, maxillary palp; pc, procoxa; pf, profemur; pn, pronotum; ps, prosternum; pts, protarsus; v 1 - 3, ventrites 1 - 3. Scale bars: 200 μm." figureDoi="10.3897/asp.80.e81736.figure3" httpUri="https://binary.pensoft.net/fig/730770" pageId="0" pageNumber="411">, 3</figureCitation>
<figureCitation id="1EEAB48B61FF7181748250B18FE892D0" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. X-ray microtomographic reconstruction of Xenostanus jiangkuni Li, Szawaryn &amp; Cai gen. et sp. nov., holotype, NIGP 177782. A Dorsal view. B Ventral view. C Lateral view. Scale bar: 400 μm." figureDoi="10.3897/asp.80.e81736.figure4" httpUri="https://binary.pensoft.net/fig/730771" pageId="0" pageNumber="411">, 4</figureCitation>
<figureCitation id="0AEB44DFF09ADE9AA489EDBCEF3D6C81" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. X-ray microtomographic reconstruction of Xenostanus jiangkuni Li, Szawaryn &amp; Cai gen. et sp. nov., holotype, NIGP 177782. A Ventral view, with legs removed. B Ventral view, rendered under &quot; Sum along Ray &quot; mode. C, D Antenna. E Head and prothorax, ventral view. F Head and prothorax, dorsal view. Scale bar: 400 μm." figureDoi="10.3897/asp.80.e81736.figure5" httpUri="https://binary.pensoft.net/fig/730772" pageId="0" pageNumber="411">, 5, S1</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="BD44D4EA4A974F101EED05087AAD00EF" pageId="0" pageNumber="411" type="etymology">
<paragraph id="BB54B496663E45B95093485CBFD4C172" pageId="0" pageNumber="411">Etymology.</paragraph>
<paragraph id="41471B9E0933C290BD963BC8CAD32565" pageId="0" pageNumber="411">The species is named after Mr. Kun Jiang, who kindly donated many fossils for our research.</paragraph>
</subSubSection>
<subSubSection id="ECCD2EA2982605C71F2209D187323C29" pageId="0" pageNumber="411" type="type material">
<paragraph id="24DA388C8E059F6F093FAD6FF22953AB" pageId="0" pageNumber="411">Type materials.</paragraph>
<paragraph id="7D85524A804C5117EF36FAFE7497ECEB" pageId="0" pageNumber="411">
<emphasis id="605B233FE16ED10DBA9645A67A3F2646" bold="true" italics="true" pageId="0" pageNumber="411">Holotype</emphasis>
: NIGP177782. Two 
<emphasis id="0D4C948A882D44C87ABB51F6FA38AA0B" bold="true" italics="true" pageId="0" pageNumber="411">paratypes</emphasis>
, GPIH no. 5058 (CCGG no. 11948), GPIH no. 5059 (CCGG no. 11105).
</paragraph>
</subSubSection>
<subSubSection id="49545BE092059F5B455B047E6A502706" pageId="0" pageNumber="411" type="type locality">
<paragraph id="D54DECFB334F39CCCB26A6F53E4D8732" pageId="0" pageNumber="411">Type locality and horizon.</paragraph>
<paragraph id="16900978D7D15EB4C96153AFAA1E3810" pageId="0" pageNumber="411">Amber mine located near Noije Bum Village, Tanai Township, Myitkyina District, Kachin State, Myanmar; unnamed horizon, mid-Cretaceous, Upper Albian to Lower Cenomanian.</paragraph>
</subSubSection>
<subSubSection id="988B8CFCEA4F09A35209D360007C5EF2" pageId="0" pageNumber="411" type="diagnosis">
<paragraph id="3639A6372E10491E953C800C4E7E21F7" pageId="0" pageNumber="411">Diagnosis.</paragraph>
<paragraph id="70B87709A130430EF4EA9C96FF77BA9C" pageId="0" pageNumber="411">As for the tribe.</paragraph>
</subSubSection>
<subSubSection id="3A9F725590E94ADC1BB1C8ED0C389B5E" pageId="0" pageNumber="411" type="description">
<paragraph id="B62684799C7A19A61C8A1F0C52F21D02" pageId="0" pageNumber="411">Description.</paragraph>
<paragraph id="4D7BE3D18E96231FDE4E1946545A762E" pageId="0" pageNumber="411">
Body elongate, widest at middle of elytra, very weakly convex. Surface with hair-like setae. - 
<emphasis id="5DE7FED84883F407BB4C3E85D9C3966B" bold="true" italics="true" pageId="0" pageNumber="411">Head</emphasis>
partially exposed and visible from above (Fig. 
<figureCitation id="54F474DE5EED51B9BC604BC8742C8063" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. X-ray microtomographic reconstruction of Xenostanus jiangkuni Li, Szawaryn &amp; Cai gen. et sp. nov., holotype, NIGP 177782. A Ventral view, with legs removed. B Ventral view, rendered under &quot; Sum along Ray &quot; mode. C, D Antenna. E Head and prothorax, ventral view. F Head and prothorax, dorsal view. Scale bar: 400 μm." figureDoi="10.3897/asp.80.e81736.figure5" httpUri="https://binary.pensoft.net/fig/730772" pageId="0" pageNumber="411">5F</figureCitation>
). Eyes coarsely faceted, without interfacetal setae (Fig. 
<figureCitation id="F69758A609B6742C159A330FE3620C85" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Details of Xenostanus jiangkuni Li, Szawaryn &amp; Cai gen. et sp. nov., holotype, NIGP 177782, under confocal microscopy. A Head and prothorax, ventral view. B Head, dorsal view. C Head and prothorax, lateral view. D Abdominal base, ventral view. Abbreviations: an, antenna; ey, compound eye; lb, labrum; lbp, labial palp; md, mandible; msf, mesofemur; msts, mesotarsus; mtf, metafemur; mttb, metatibia; mtts, metatarsus; mtv, metaventrite; mxp, maxillary palp; pc, procoxa; pf, profemur; pn, pronotum; ps, prosternum; pts, protarsus; v 1 - 3, ventrites 1 - 3. Scale bars: 200 μm." figureDoi="10.3897/asp.80.e81736.figure3" httpUri="https://binary.pensoft.net/fig/730770" pageId="0" pageNumber="411">3B,C</figureCitation>
). Frontoclypeal suture absent (Fig. 
<figureCitation id="4034F5A4AF204B0AD101B0CA9C540DB0" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Details of Xenostanus jiangkuni Li, Szawaryn &amp; Cai gen. et sp. nov., holotype, NIGP 177782, under confocal microscopy. A Head and prothorax, ventral view. B Head, dorsal view. C Head and prothorax, lateral view. D Abdominal base, ventral view. Abbreviations: an, antenna; ey, compound eye; lb, labrum; lbp, labial palp; md, mandible; msf, mesofemur; msts, mesotarsus; mtf, metafemur; mttb, metatibia; mtts, metatarsus; mtv, metaventrite; mxp, maxillary palp; pc, procoxa; pf, profemur; pn, pronotum; ps, prosternum; pts, protarsus; v 1 - 3, ventrites 1 - 3. Scale bars: 200 μm." figureDoi="10.3897/asp.80.e81736.figure3" httpUri="https://binary.pensoft.net/fig/730770" pageId="0" pageNumber="411">3B</figureCitation>
). Labrum free, transverse (Fig. 
<figureCitation id="2EA3F76DCA7B4A39272043B454ABD496" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Details of Xenostanus jiangkuni Li, Szawaryn &amp; Cai gen. et sp. nov., holotype, NIGP 177782, under confocal microscopy. A Head and prothorax, ventral view. B Head, dorsal view. C Head and prothorax, lateral view. D Abdominal base, ventral view. Abbreviations: an, antenna; ey, compound eye; lb, labrum; lbp, labial palp; md, mandible; msf, mesofemur; msts, mesotarsus; mtf, metafemur; mttb, metatibia; mtts, metatarsus; mtv, metaventrite; mxp, maxillary palp; pc, procoxa; pf, profemur; pn, pronotum; ps, prosternum; pts, protarsus; v 1 - 3, ventrites 1 - 3. Scale bars: 200 μm." figureDoi="10.3897/asp.80.e81736.figure3" httpUri="https://binary.pensoft.net/fig/730770" pageId="0" pageNumber="411">3B</figureCitation>
). Subantennal grooves absent. Antennae (Fig. 
<figureCitation id="7E4DCB222EAFEBBE48C64C6951F3FF67" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. X-ray microtomographic reconstruction of Xenostanus jiangkuni Li, Szawaryn &amp; Cai gen. et sp. nov., holotype, NIGP 177782. A Dorsal view. B Ventral view. C Lateral view. Scale bar: 400 μm." figureDoi="10.3897/asp.80.e81736.figure4" httpUri="https://binary.pensoft.net/fig/730771" pageId="0" pageNumber="411">4C,D</figureCitation>
) with 10 antennomeres; scape distinctly longer and wider than pedicel; antennomeres 3-7 small, subquadrate to transverse; club (antennomeres 8-10) asymmetrical, as long as antennomeres 3-7 combined. Mandibles short (Fig. 
<figureCitation id="83A83DA606D046FE5A5A1BE809C2259A" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Details of Xenostanus jiangkuni Li, Szawaryn &amp; Cai gen. et sp. nov., holotype, NIGP 177782, under confocal microscopy. A Head and prothorax, ventral view. B Head, dorsal view. C Head and prothorax, lateral view. D Abdominal base, ventral view. Abbreviations: an, antenna; ey, compound eye; lb, labrum; lbp, labial palp; md, mandible; msf, mesofemur; msts, mesotarsus; mtf, metafemur; mttb, metatibia; mtts, metatarsus; mtv, metaventrite; mxp, maxillary palp; pc, procoxa; pf, profemur; pn, pronotum; ps, prosternum; pts, protarsus; v 1 - 3, ventrites 1 - 3. Scale bars: 200 μm." figureDoi="10.3897/asp.80.e81736.figure3" httpUri="https://binary.pensoft.net/fig/730770" pageId="0" pageNumber="411">3A</figureCitation>
). Maxillary palps (Fig. 
<figureCitation id="884C0CF136B6E9B0E20766A08E38102B" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Details of Xenostanus jiangkuni Li, Szawaryn &amp; Cai gen. et sp. nov., holotype, NIGP 177782, under confocal microscopy. A Head and prothorax, ventral view. B Head, dorsal view. C Head and prothorax, lateral view. D Abdominal base, ventral view. Abbreviations: an, antenna; ey, compound eye; lb, labrum; lbp, labial palp; md, mandible; msf, mesofemur; msts, mesotarsus; mtf, metafemur; mttb, metatibia; mtts, metatarsus; mtv, metaventrite; mxp, maxillary palp; pc, procoxa; pf, profemur; pn, pronotum; ps, prosternum; pts, protarsus; v 1 - 3, ventrites 1 - 3. Scale bars: 200 μm." figureDoi="10.3897/asp.80.e81736.figure3" httpUri="https://binary.pensoft.net/fig/730770" pageId="0" pageNumber="411">3A</figureCitation>
) 3-segmented; apical palpomere about twice as long as penultimate one, conical. Labial palps (Fig. 
<figureCitation id="0878646D4C02CB95A83F7EEE53D18781" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Details of Xenostanus jiangkuni Li, Szawaryn &amp; Cai gen. et sp. nov., holotype, NIGP 177782, under confocal microscopy. A Head and prothorax, ventral view. B Head, dorsal view. C Head and prothorax, lateral view. D Abdominal base, ventral view. Abbreviations: an, antenna; ey, compound eye; lb, labrum; lbp, labial palp; md, mandible; msf, mesofemur; msts, mesotarsus; mtf, metafemur; mttb, metatibia; mtts, metatarsus; mtv, metaventrite; mxp, maxillary palp; pc, procoxa; pf, profemur; pn, pronotum; ps, prosternum; pts, protarsus; v 1 - 3, ventrites 1 - 3. Scale bars: 200 μm." figureDoi="10.3897/asp.80.e81736.figure3" httpUri="https://binary.pensoft.net/fig/730770" pageId="0" pageNumber="411">3A</figureCitation>
) 2-segmented; apical palpomere about as long as basal one. Ventral head surface seemingly with a pair of parallel subgenal ridges (Figs 
<figureCitation id="0E26A44C132F784FBF58664DB4DEE68C" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Details of Xenostanus jiangkuni Li, Szawaryn &amp; Cai gen. et sp. nov., holotype, NIGP 177782, under confocal microscopy. A Head and prothorax, ventral view. B Head, dorsal view. C Head and prothorax, lateral view. D Abdominal base, ventral view. Abbreviations: an, antenna; ey, compound eye; lb, labrum; lbp, labial palp; md, mandible; msf, mesofemur; msts, mesotarsus; mtf, metafemur; mttb, metatibia; mtts, metatarsus; mtv, metaventrite; mxp, maxillary palp; pc, procoxa; pf, profemur; pn, pronotum; ps, prosternum; pts, protarsus; v 1 - 3, ventrites 1 - 3. Scale bars: 200 μm." figureDoi="10.3897/asp.80.e81736.figure3" httpUri="https://binary.pensoft.net/fig/730770" pageId="0" pageNumber="411">3A</figureCitation>
, 
<figureCitation id="B009044CEFB1D5148430B8A6162250D6" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. X-ray microtomographic reconstruction of Xenostanus jiangkuni Li, Szawaryn &amp; Cai gen. et sp. nov., holotype, NIGP 177782. A Ventral view, with legs removed. B Ventral view, rendered under &quot; Sum along Ray &quot; mode. C, D Antenna. E Head and prothorax, ventral view. F Head and prothorax, dorsal view. Scale bar: 400 μm." figureDoi="10.3897/asp.80.e81736.figure5" httpUri="https://binary.pensoft.net/fig/730772" pageId="0" pageNumber="411">5E</figureCitation>
). - 
<emphasis id="6C059A5D49323A00EF18B169B7749F8B" bold="true" italics="true" pageId="0" pageNumber="411">Prothorax</emphasis>
: Pronotal disc (Fig. 
<figureCitation id="556EC9C439329065320F60D3DFE14244" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. X-ray microtomographic reconstruction of Xenostanus jiangkuni Li, Szawaryn &amp; Cai gen. et sp. nov., holotype, NIGP 177782. A Ventral view, with legs removed. B Ventral view, rendered under &quot; Sum along Ray &quot; mode. C, D Antenna. E Head and prothorax, ventral view. F Head and prothorax, dorsal view. Scale bar: 400 μm." figureDoi="10.3897/asp.80.e81736.figure5" httpUri="https://binary.pensoft.net/fig/730772" pageId="0" pageNumber="411">5F</figureCitation>
) widest at base; anterior margin straight; lateral margins bordered in posterior part; posterior angles pointed; posterior margin bisinuate. Prosternum (Figs 
<figureCitation id="73E7CB36FEBD0A9D5661701CDE700C6C" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Details of Xenostanus jiangkuni Li, Szawaryn &amp; Cai gen. et sp. nov., holotype, NIGP 177782, under confocal microscopy. A Head and prothorax, ventral view. B Head, dorsal view. C Head and prothorax, lateral view. D Abdominal base, ventral view. Abbreviations: an, antenna; ey, compound eye; lb, labrum; lbp, labial palp; md, mandible; msf, mesofemur; msts, mesotarsus; mtf, metafemur; mttb, metatibia; mtts, metatarsus; mtv, metaventrite; mxp, maxillary palp; pc, procoxa; pf, profemur; pn, pronotum; ps, prosternum; pts, protarsus; v 1 - 3, ventrites 1 - 3. Scale bars: 200 μm." figureDoi="10.3897/asp.80.e81736.figure3" httpUri="https://binary.pensoft.net/fig/730770" pageId="0" pageNumber="411">3A</figureCitation>
, 
<figureCitation id="D3394C6E0229CC8AEE29B09B770EADB5" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. X-ray microtomographic reconstruction of Xenostanus jiangkuni Li, Szawaryn &amp; Cai gen. et sp. nov., holotype, NIGP 177782. A Ventral view, with legs removed. B Ventral view, rendered under &quot; Sum along Ray &quot; mode. C, D Antenna. E Head and prothorax, ventral view. F Head and prothorax, dorsal view. Scale bar: 400 μm." figureDoi="10.3897/asp.80.e81736.figure5" httpUri="https://binary.pensoft.net/fig/730772" pageId="0" pageNumber="411">5E</figureCitation>
) in front of coxae about as long as longitudinal coxal diameter, anteriorly not produced forward; prosternal carinae absent; prosternal process broad, widened beyond front coxae, meeting postcoxal hypomeral projections, truncate at apex. Procoxal cavities externally closed, ovaloid, without lateral slit (Fig. 
<figureCitation id="06E75F798CC3ACFE71B471C6016A2EA4" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. X-ray microtomographic reconstruction of Xenostanus jiangkuni Li, Szawaryn &amp; Cai gen. et sp. nov., holotype, NIGP 177782. A Ventral view, with legs removed. B Ventral view, rendered under &quot; Sum along Ray &quot; mode. C, D Antenna. E Head and prothorax, ventral view. F Head and prothorax, dorsal view. Scale bar: 400 μm." figureDoi="10.3897/asp.80.e81736.figure5" httpUri="https://binary.pensoft.net/fig/730772" pageId="0" pageNumber="411">5E</figureCitation>
). - 
<emphasis id="F1623EA5CAB9FFF27A3B55CABCD4D1B5" bold="true" italics="true" pageId="0" pageNumber="411">Meso- and metathorax</emphasis>
: Scutellar shield strongly transverse. Elytra covering entire abdomen except for part of pygidium (Fig. 
<figureCitation id="B9A2B06BD5436F8E2FE522B8E6134401" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. General habitus of Xenostanus jiangkuni Li, Szawaryn &amp; Cai gen. et sp. nov., holotype, NIGP 177782, under incident light. A Dorsal view. B Ventral view. Scale bars: 400 μm." figureDoi="10.3897/asp.80.e81736.figure1" httpUri="https://binary.pensoft.net/fig/730768" pageId="0" pageNumber="411">1A</figureCitation>
). Mesoventrite flat, without transverse carina (Figs 
<figureCitation id="2C2673C2304F0FFB6DA03DE4300A895B" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. General habitus of Xenostanus jiangkuni Li, Szawaryn &amp; Cai gen. et sp. nov., holotype, NIGP 177782, under incident light. A Dorsal view. B Ventral view. Scale bars: 400 μm." figureDoi="10.3897/asp.80.e81736.figure1" httpUri="https://binary.pensoft.net/fig/730768" pageId="0" pageNumber="411">1B</figureCitation>
, 
<figureCitation id="57EDDE1303EDE98DF07EA2E11600827A" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. X-ray microtomographic reconstruction of Xenostanus jiangkuni Li, Szawaryn &amp; Cai gen. et sp. nov., holotype, NIGP 177782. A Ventral view, with legs removed. B Ventral view, rendered under &quot; Sum along Ray &quot; mode. C, D Antenna. E Head and prothorax, ventral view. F Head and prothorax, dorsal view. Scale bar: 400 μm." figureDoi="10.3897/asp.80.e81736.figure5" httpUri="https://binary.pensoft.net/fig/730772" pageId="0" pageNumber="411">5A</figureCitation>
). Mesocoxal cavities circular, outwardly closed (Figs 
<figureCitation id="1BA748466BCF60645178A1C102A6366E" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. General habitus of Xenostanus jiangkuni Li, Szawaryn &amp; Cai gen. et sp. nov., holotype, NIGP 177782, under incident light. A Dorsal view. B Ventral view. Scale bars: 400 μm." figureDoi="10.3897/asp.80.e81736.figure1" httpUri="https://binary.pensoft.net/fig/730768" pageId="0" pageNumber="411">1B</figureCitation>
, 
<figureCitation id="85B20927B1A57F1E0A21973D87A7737E" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. X-ray microtomographic reconstruction of Xenostanus jiangkuni Li, Szawaryn &amp; Cai gen. et sp. nov., holotype, NIGP 177782. A Ventral view, with legs removed. B Ventral view, rendered under &quot; Sum along Ray &quot; mode. C, D Antenna. E Head and prothorax, ventral view. F Head and prothorax, dorsal view. Scale bar: 400 μm." figureDoi="10.3897/asp.80.e81736.figure5" httpUri="https://binary.pensoft.net/fig/730772" pageId="0" pageNumber="411">5A</figureCitation>
). Meso-metaventral junction nearly straight (though with a small projection medially). Metaventrite with distinct postcoxal lines (Figs 
<figureCitation id="00A334DD7930BD1E0F89DB63E2566F23" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. General habitus of Xenostanus jiangkuni Li, Szawaryn &amp; Cai gen. et sp. nov., holotype, NIGP 177782, under incident light. A Dorsal view. B Ventral view. Scale bars: 400 μm." figureDoi="10.3897/asp.80.e81736.figure1" httpUri="https://binary.pensoft.net/fig/730768" pageId="0" pageNumber="411">1B</figureCitation>
, 
<figureCitation id="98FF8A87050A630BA9C37C1E5CA4F567" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. General habitus of Xenostanus jiangkuni Li, Szawaryn &amp; Cai gen. et sp. nov., holotype, NIGP 177782, under confocal microscopy. A Dorsal view. B Ventral view, with arrowheads indicating the metaventral and abdominal postcoxal lines. Scale bars: 400 μm." figureDoi="10.3897/asp.80.e81736.figure2" httpUri="https://binary.pensoft.net/fig/730769" pageId="0" pageNumber="411">2B</figureCitation>
, 
<figureCitation id="367F645DA704D74B6E234AECA1DF2946" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. X-ray microtomographic reconstruction of Xenostanus jiangkuni Li, Szawaryn &amp; Cai gen. et sp. nov., holotype, NIGP 177782. A Ventral view, with legs removed. B Ventral view, rendered under &quot; Sum along Ray &quot; mode. C, D Antenna. E Head and prothorax, ventral view. F Head and prothorax, dorsal view. Scale bar: 400 μm." figureDoi="10.3897/asp.80.e81736.figure5" httpUri="https://binary.pensoft.net/fig/730772" pageId="0" pageNumber="411">5A</figureCitation>
); discrimen visible in posterior third of metaventrite. Metacoxae transverse, broadly separated. - 
<emphasis id="D46DF6D7225AEE3E475E8193C60A8E33" bold="true" italics="true" pageId="0" pageNumber="411">Legs</emphasis>
: Femora flattened. Tibiae simple, not strongly widened, apically with small denticles; tibial spurs 2-2-2 (Fig. 
<figureCitation id="D4E86E26E1460B26B93A9F4B7C838C49" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Details of Xenostanus jiangkuni Li, Szawaryn &amp; Cai gen. et sp. nov., holotype, NIGP 177782, under confocal microscopy. A Head and prothorax, ventral view. B Head, dorsal view. C Head and prothorax, lateral view. D Abdominal base, ventral view. Abbreviations: an, antenna; ey, compound eye; lb, labrum; lbp, labial palp; md, mandible; msf, mesofemur; msts, mesotarsus; mtf, metafemur; mttb, metatibia; mtts, metatarsus; mtv, metaventrite; mxp, maxillary palp; pc, procoxa; pf, profemur; pn, pronotum; ps, prosternum; pts, protarsus; v 1 - 3, ventrites 1 - 3. Scale bars: 200 μm." figureDoi="10.3897/asp.80.e81736.figure3" httpUri="https://binary.pensoft.net/fig/730770" pageId="0" pageNumber="411">3D</figureCitation>
). Tarsi 4-4-4 (Fig. 
<figureCitation id="50F68CBF6A1859BC04290B28CC3B9450" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Details of Xenostanus jiangkuni Li, Szawaryn &amp; Cai gen. et sp. nov., holotype, NIGP 177782, under confocal microscopy. A Head and prothorax, ventral view. B Head, dorsal view. C Head and prothorax, lateral view. D Abdominal base, ventral view. Abbreviations: an, antenna; ey, compound eye; lb, labrum; lbp, labial palp; md, mandible; msf, mesofemur; msts, mesotarsus; mtf, metafemur; mttb, metatibia; mtts, metatarsus; mtv, metaventrite; mxp, maxillary palp; pc, procoxa; pf, profemur; pn, pronotum; ps, prosternum; pts, protarsus; v 1 - 3, ventrites 1 - 3. Scale bars: 200 μm." figureDoi="10.3897/asp.80.e81736.figure3" httpUri="https://binary.pensoft.net/fig/730770" pageId="0" pageNumber="411">3D</figureCitation>
); tarsomeres 1 and 2 ventrally lobed; tarsomere 3 smaller and simple; tarsomere 4 elongate, as long as 1-3 combined. Pretarsal claws simple, with small basal angulation. - 
<emphasis id="3A456FE3EBC08905DB3CAD02BCA42964" bold="true" italics="true" pageId="0" pageNumber="411">Abdomen</emphasis>
with six freely articulated ventrites. Ventrite 1 longer than 2-4 combined, with strongly arcuate postcoxal lines (Figs 
<figureCitation id="E7B0CC106ABD181DF753A86C3C6F80A9" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. General habitus of Xenostanus jiangkuni Li, Szawaryn &amp; Cai gen. et sp. nov., holotype, NIGP 177782, under incident light. A Dorsal view. B Ventral view. Scale bars: 400 μm." figureDoi="10.3897/asp.80.e81736.figure1" httpUri="https://binary.pensoft.net/fig/730768" pageId="0" pageNumber="411">1B</figureCitation>
, 
<figureCitation id="9871C92D4DD2767D006585C8C8DB1AD3" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. General habitus of Xenostanus jiangkuni Li, Szawaryn &amp; Cai gen. et sp. nov., holotype, NIGP 177782, under confocal microscopy. A Dorsal view. B Ventral view, with arrowheads indicating the metaventral and abdominal postcoxal lines. Scale bars: 400 μm." figureDoi="10.3897/asp.80.e81736.figure2" httpUri="https://binary.pensoft.net/fig/730769" pageId="0" pageNumber="411">2B</figureCitation>
, 
<figureCitation id="78F08AB9A5ECEFC35BF791A6CF1168D2" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. X-ray microtomographic reconstruction of Xenostanus jiangkuni Li, Szawaryn &amp; Cai gen. et sp. nov., holotype, NIGP 177782. A Ventral view, with legs removed. B Ventral view, rendered under &quot; Sum along Ray &quot; mode. C, D Antenna. E Head and prothorax, ventral view. F Head and prothorax, dorsal view. Scale bar: 400 μm." figureDoi="10.3897/asp.80.e81736.figure5" httpUri="https://binary.pensoft.net/fig/730772" pageId="0" pageNumber="411">5A</figureCitation>
), anteriorly complete; intercoxal process very broad and truncate.
</paragraph>
<caption id="C975E2B33B5C5F57CDB7872EABCE838E" doi="10.3897/asp.80.e81736.figure1" httpUri="https://binary.pensoft.net/fig/730768" pageId="0" pageNumber="411" start="Figure 1" startId="F1">
<paragraph id="0DD7B0FED63B86DA8A22CE80EA567902" pageId="0" pageNumber="411">
<emphasis id="65C58EF4B18D9350F7B4BB4B3FDEEBE8" bold="true" pageId="0" pageNumber="411">Figure 1.</emphasis>
General habitus of 
<taxonomicName id="A1966ADD0E7D3826D8FD88B9C20B4B39" authorityName="Li, Szawaryn &amp; Cai" authorityYear="2022" class="Insecta" family="Corylophidae" genus="Xenostanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xenostanus jiangkuni" order="Coleoptera" pageId="0" pageNumber="411" phylum="Arthropoda" rank="species" species="jiangkuni">
<emphasis id="AA48989CD41BCB1EC0E20A5C8D2612F9" italics="true" pageId="0" pageNumber="411">Xenostanus jiangkuni</emphasis>
</taxonomicName>
Li, Szawaryn &amp; Cai 
<emphasis id="E01F117AF917B93AE5DFEBB66400EE5B" bold="true" pageId="0" pageNumber="411">gen. et sp. nov.</emphasis>
, holotype, NIGP177782, under incident light. 
<emphasis id="AE14A8A0CE14E5D96170CC7703EECF67" bold="true" pageId="0" pageNumber="411">A</emphasis>
Dorsal view. 
<emphasis id="9BE6E151E14B0AC1AAF6CE4D2DB24C04" bold="true" pageId="0" pageNumber="411">B</emphasis>
Ventral view. Scale bars: 400 
<normalizedToken id="2459B150F9A4F2FB4ED11E26817ED37A" originalValue="μm">μm</normalizedToken>
.
</paragraph>
</caption>
<caption id="B91CD0BA28A373D553D9FBEE9C0CE7B9" doi="10.3897/asp.80.e81736.figure2" httpUri="https://binary.pensoft.net/fig/730769" pageId="0" pageNumber="411" start="Figure 2" startId="F2">
<paragraph id="0B26A5E14B4AE14F5D509C0FDA560872" pageId="0" pageNumber="411">
<emphasis id="E295723FFD51776C3981CA2CCBE8FA98" bold="true" pageId="0" pageNumber="411">Figure 2.</emphasis>
General habitus of 
<taxonomicName id="F282A2769454D7C65E5B277D242C715E" authorityName="Li, Szawaryn &amp; Cai" authorityYear="2022" class="Insecta" family="Corylophidae" genus="Xenostanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xenostanus jiangkuni" order="Coleoptera" pageId="0" pageNumber="411" phylum="Arthropoda" rank="species" species="jiangkuni">
<emphasis id="5256119077BC1838CD07A75A59D2E4EC" italics="true" pageId="0" pageNumber="411">Xenostanus jiangkuni</emphasis>
</taxonomicName>
Li, Szawaryn &amp; Cai 
<emphasis id="1702C6B6946A90C278C5094D9EAF3188" bold="true" pageId="0" pageNumber="411">gen. et sp. nov.</emphasis>
, holotype, NIGP177782, under confocal microscopy. 
<emphasis id="748B0B6986C93BA82567D88999BF2ACF" bold="true" pageId="0" pageNumber="411">A</emphasis>
Dorsal view. 
<emphasis id="BF6B25559ECF335D2587F0D2A9350B50" bold="true" pageId="0" pageNumber="411">B</emphasis>
Ventral view, with arrowheads indicating the metaventral and abdominal postcoxal lines. Scale bars: 400 
<normalizedToken id="568AFF5DAFB6DDD6B4B346F19A7BA47A" originalValue="μm">μm</normalizedToken>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="F8EF86056E3C46A766AC2A5C91C32B66" pageId="0" pageNumber="411" type="description">
<paragraph id="D2ADBD859505E30A9098EAD5A8BB4FB2" pageId="0" pageNumber="411">Measurements.</paragraph>
<paragraph id="2A57BB47E05214CCA0B28E0CE70D8430" pageId="0" pageNumber="411">
NIGP177782: 
<emphasis id="8BDDF44AD48BAB09CE8D1FB87C1524AE" bold="true" pageId="0" pageNumber="411">BL</emphasis>
1.42 mm, 
<emphasis id="86D9836F2BCC3DF53A10B673565050D5" bold="true" pageId="0" pageNumber="411">BW</emphasis>
0.63 mm, 
<emphasis id="A54D60FAD58AB7B0B9000BF80F37D04F" bold="true" pageId="0" pageNumber="411">HL</emphasis>
0.27 mm, 
<emphasis id="BB0A9E972F2BEA4E56D8833846CFB9CA" bold="true" pageId="0" pageNumber="411">HW</emphasis>
0.31 mm, 
<emphasis id="16682D7780E2AFF6BF2BA8D6C2C005D8" bold="true" pageId="0" pageNumber="411">PL</emphasis>
0.34 mm, 
<emphasis id="D420145E45190C34D81D5F1C93E694EF" bold="true" pageId="0" pageNumber="411">PW</emphasis>
0.51 mm, 
<emphasis id="395E239D642AC7E7478C52831CC41610" bold="true" pageId="0" pageNumber="411">EL</emphasis>
0.96 mm. GPIH no. 5058: 
<emphasis id="806F256659450B6E7B05084C124AA9CE" bold="true" pageId="0" pageNumber="411">BL</emphasis>
1.88 mm, 
<emphasis id="F5A240FB28A98CF00FA5D4D677C713BC" bold="true" pageId="0" pageNumber="411">BW</emphasis>
0.74 mm. GPIH no. 5059: 
<emphasis id="9B2DC78B2AB002DC24714BAF2604CA86" bold="true" pageId="0" pageNumber="411">BL</emphasis>
1.30 mm, 
<emphasis id="CC42E9D4F239AD1B36D1477EA4E83D32" bold="true" pageId="0" pageNumber="411">BW</emphasis>
0.62 mm
</paragraph>
</subSubSection>
</treatment>
</subSection>
</document>