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<document ID-CLB-Dataset="106374" ID-DOI="http://dx.doi.org/10.3897/dez.69.84080" ID-GBIF-Dataset="7d18e551-4fca-4cf5-9857-11b5e2dd4a5c" ID-Pensoft-Pub="1860-1324-2-151" ID-Pensoft-UUID="B7D8CC48BD32502C819369D75ADA50C8" ID-ZooBank="3B245B5371564A3F96672F2CD756779A" ModsDocID="1860-1324-69-2-151" checkinTime="1658871268840" checkinUser="pensoft" docAuthor="Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia &amp; Prous, Marko" docDate="2022" docId="A169625F1E8A5801822B7CD36645130C" docLanguage="en" docName="DeutEntomolZeit 69(2): 151-218" docOrigin="Deutsche Entomologische Zeitschrift 69 (2)" docPubDate="2022-07-26" docSource="http://dx.doi.org/10.3897/dez.69.84080" docTitle="Dolerus timidus, sp. rev." docType="treatment" docVersion="8" id="B7D8CC48BD32502C819369D75ADA50C8" lastPageNumber="151" masterDocId="B7D8CC48BD32502C819369D75ADA50C8" masterDocTitle="Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta)" masterLastPageNumber="218" masterPageNumber="151" pageNumber="151" updateTime="1737031476642" updateUser="carolina">
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<mods:title id="6A839A45DAF220A0D12F93FCB37365AB">Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta)</mods:title>
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<mods:namePart id="DBDABE4D095C5DFC4FA4669951F88BC3">Liston, Andrew</mods:namePart>
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<mods:affiliation id="516F9DC0EC30AC4EC795BFD1EC0248FD">Senckenberg Deutsches Entomologisches Institut, Eberswalder Str. 90, 15374 Muencheberg, Germany</mods:affiliation>
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<mods:namePart id="D6080E4493B465F5022C64266E51FEF9">Mutanen, Marko</mods:namePart>
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<mods:affiliation id="3C1FA09E9D849981B39A466C0ECFAAC6">Ecology and Genetics Research Unit, PO Box 3000, 90014 University of Oulu, Oulu, Finland</mods:affiliation>
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<mods:namePart id="7D5FE71FBA002B08630F8C3A854C7DFB">Heidemaa, Mikk</mods:namePart>
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<mods:namePart id="77237FAAE0ECBD44A1044864CD3166B3">Blank, Stephan M.</mods:namePart>
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<mods:affiliation id="53C812FA30536AF09263D3EF345E6CE9">Senckenberg Deutsches Entomologisches Institut, Eberswalder Str. 90, 15374 Muencheberg, Germany</mods:affiliation>
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<mods:affiliation id="62ABF91D34E0B4D988FC06EC5E4BE16F">Department of Environmental and Biological Sciences, University of Eastern Finland, 80101 Joensuu, Finland</mods:affiliation>
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<mods:namePart id="3936714FF6569E777101B9A73D2E2311">Prous, Marko</mods:namePart>
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<mods:affiliation id="A3C1ACFB46DB78ECB6BF63CAEE531E65">Senckenberg Deutsches Entomologisches Institut, Eberswalder Str. 90, 15374 Muencheberg, Germany &amp; Ecology and Genetics Research Unit, PO Box 3000, 90014 University of Oulu, Oulu, Finland &amp; Department of Zoology, Institute of Ecology and Earth Sciences, University of Tartu, Vanemuise 46, 51014 Tartu, Estonia</mods:affiliation>
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<treatment id="A169625F1E8A5801822B7CD36645130C" ID-DOI="http://doi.org/10.5281/zenodo.7019641" ID-GBIF-Taxon="197857907" ID-Zenodo-Dep="7019641" LSID="urn:lsid:plazi:treatment:A169625F1E8A5801822B7CD36645130C" httpUri="http://treatment.plazi.org/id/A169625F1E8A5801822B7CD36645130C" lastPageNumber="151" pageId="0" pageNumber="151" scope_order="Hymenoptera">
<subSubSection id="C0E2377DDD6276C7C33D8D8A92C3CD0E" pageId="0" pageNumber="151" type="nomenclature">
<paragraph id="B4CBBEC090375A169971D758A6A0E5DE" pageId="0" pageNumber="151">
<taxonomicName id="F8BB42403BDA9901B9133E23E459D58D" LSID="A169625F-1E8A-5801-822B-7CD36645130C" authority="(Klug, 1818)" baseAuthorityName="Klug" baseAuthorityYear="1818" class="Insecta" family="Tenthredinidae" genus="Dolerus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dolerus timidus" order="Hymenoptera" pageId="0" pageNumber="151" phylum="Arthropoda" rank="species" species="timidus" status="sp. rev.">Dolerus timidus (Klug, 1818)</taxonomicName>
<taxonomicNameLabel id="E8F68882E147ABC1DD2469972BC29187" pageId="0" pageNumber="151">sp. rev.</taxonomicNameLabel>
</paragraph>
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<subSubSection id="7A91E5D860D4BDD1C032085C2BB121C9" pageId="0" pageNumber="151" type="description">
<paragraph id="B9E4BCC8FFDD31C553F74307AE82821D" pageId="0" pageNumber="151">
<figureCitation id="6DF4A34F59605CD80DB8371027AF716E" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Dolerus pratensis (Linnaeus, 1758) ♀ (ZMUO. 045279). A. Dorsal habitus; B. Lateral habitus. Dolerus timidus (Klug, 1818) ♀ (ZMUO. 032881); C. Dorsal habitus; D. Lateral habitus." figureDoi="10.3897/dez.69.84080.figure13" httpUri="https://binary.pensoft.net/fig/719524" pageId="0" pageNumber="151">Figs 13C, D</figureCitation>
<figureCitation id="2A0747374EFFE300BD05FA1A45558C23" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Dolerus timidus (Klug, 1818), D. variator Enslin, 1927 and D. pratensis (Linnaeus, 1758). A. Penis valve, lectotype Tenthredo timida; B. Penis valve, D. variator; C. Metepimeron (arrow), D. pratensis (ZMUO. 032881); D. Metepimeron (arrow), D. timidus (ZMUO. 029409), mirrored; E. Lancet, D. timidus; F. Lancet, D. pratensis." figureDoi="10.3897/dez.69.84080.figure14" httpUri="https://binary.pensoft.net/fig/719525" pageId="0" pageNumber="151">, 14A</figureCitation>
<figureCitation id="7FF5F64B8A4FCE336977F32F75F98CEC" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Dolerus timidus (Klug, 1818), D. variator Enslin, 1927 and D. pratensis (Linnaeus, 1758). A. Penis valve, lectotype Tenthredo timida; B. Penis valve, D. variator; C. Metepimeron (arrow), D. pratensis (ZMUO. 032881); D. Metepimeron (arrow), D. timidus (ZMUO. 029409), mirrored; E. Lancet, D. timidus; F. Lancet, D. pratensis." figureDoi="10.3897/dez.69.84080.figure14" httpUri="https://binary.pensoft.net/fig/719525" pageId="0" pageNumber="151">, 14D</figureCitation>
<figureCitation id="F9A1F32A14326F85746148549B4B08EB" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Dolerus timidus (Klug, 1818), D. variator Enslin, 1927 and D. pratensis (Linnaeus, 1758). A. Penis valve, lectotype Tenthredo timida; B. Penis valve, D. variator; C. Metepimeron (arrow), D. pratensis (ZMUO. 032881); D. Metepimeron (arrow), D. timidus (ZMUO. 029409), mirrored; E. Lancet, D. timidus; F. Lancet, D. pratensis." figureDoi="10.3897/dez.69.84080.figure14" httpUri="https://binary.pensoft.net/fig/719525" pageId="0" pageNumber="151">, 14E</figureCitation>
</paragraph>
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<subSubSection id="878F55735B8E3191200F0573F640199D" pageId="0" pageNumber="151" type="reference_group">
<paragraph id="1326989ED9BE718D54EC22498E33B4FB" pageId="0" pageNumber="151">
<taxonomicName id="7461B02B2EB7D6382001F7D76630A510" authority="Klug, 1818" class="Insecta" family="Tenthredinidae" genus="Tenthredo" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tenthredo (Dolerus) timida" order="Hymenoptera" pageId="0" pageNumber="151" phylum="Arthropoda" rank="species" species="timida" status="sp. rev." subGenus="Dolerus">Tenthredo (Dolerus) timida</taxonomicName>
Klug, 1818: 300. ♀, ♂. Syntypes. Published type locality:
<normalizedToken id="2C137B3D6BF9C8A5530C85020B19F6C6" originalValue="“Deutschland”">&quot;Deutschland&quot;</normalizedToken>
[Germany]. Lectotype designated below.
</paragraph>
<paragraph id="F6AA294E060BF3D777CC8D976D13DF74" pageId="0" pageNumber="151">
<taxonomicName id="47E9ABB1F2C39B8FC7868E20685E4C72" authority="Klug, 1818" class="Insecta" family="Tenthredinidae" genus="Tenthredo" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tenthredo (Dolerus) dubia" order="Hymenoptera" pageId="0" pageNumber="151" phylum="Arthropoda" rank="species" species="dubia" status="sp. rev." subGenus="Dolerus">Tenthredo (Dolerus) dubia</taxonomicName>
Klug, 1818: 299-300. ♀, ♂. Syntypes. Type locality: Germany. Lectotype designated below. Primary homonym of
<taxonomicName id="619765248AAE3A23F8116FB0EE77CE22" authority="Ström, 1768" class="Insecta" family="Tenthredinidae" genus="Tenthredo" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tenthredo dubia" order="Hymenoptera" pageId="0" pageNumber="151" phylum="Arthropoda" rank="species" species="dubia" status="sp. rev.">Tenthredo dubia</taxonomicName>
<normalizedToken id="112DD5BCE092CF31ACA34C2C97D9A4F4" originalValue="Ström">Stroem</normalizedToken>
, 1768 [=
<taxonomicName id="CF7E1FD112FEEF4CC306A096451AC427" authority="Linnaeus, 1758" class="Insecta" family="Tenthredinidae" genus="Tenthredo" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tenthredo (Tenthredella) livida" order="Hymenoptera" pageId="0" pageNumber="151" phylum="Arthropoda" rank="species" species="livida" status="sp. rev." subGenus="Tenthredella">Tenthredo (Tenthredella) livida</taxonomicName>
Linnaeus, 1758].
</paragraph>
<paragraph id="C6FD3C7C59A32257E3C8EA10C91F625F" pageId="0" pageNumber="151">
<taxonomicName id="1A141E55114FC52FB828355EE74C8604" authority="Enslin, 1913" class="Insecta" family="Tenthredinidae" genus="Dolerus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dolerus dubius subsp. var. var. atratus" order="Hymenoptera" pageId="0" pageNumber="151" phylum="Arthropoda" rank="variety" species="dubius" status="sp. rev." variety="atratus">Dolerus dubius var. atratus</taxonomicName>
Enslin, 1913: 159. ♂. Syntypes (assumed). Type locality: Switzerland. Lectotype designated below.
</paragraph>
</subSubSection>
<subSubSection id="0EAAA892CE17C3F538E51000ABD69174" pageId="0" pageNumber="151" type="type material">
<paragraph id="A46C87E9A0FCD9644E781701D8ADC30A" pageId="0" pageNumber="151">Type material examined and taxonomic notes.</paragraph>
<paragraph id="D0651E37B7A8BB261B5BA610AE72D866" pageId="0" pageNumber="151">
<taxonomicName id="68A8E03DCE5E027B8FBD03A26C36BCBD" authorityName="Klug, 1818" class="Insecta" family="Tenthredinidae" genus="Tenthredo" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tenthredo (Dolerus) timida" order="Hymenoptera" pageId="0" pageNumber="151" phylum="Arthropoda" rank="species" species="timida" subGenus="Dolerus">Tenthredo (Dolerus) timida</taxonomicName>
Klug, 1818. Lectotype ♂ hereby designated, labelled:
<normalizedToken id="D29EF8F953A26C2DED9F64E4F9D3C7D0" originalValue="“14199”">&quot;14199&quot;</normalizedToken>
[printed catalog no.], &quot;Var.
<taxonomicName id="D64681B37DC5C93CBD19EF9714865DA3" lsidName="T. timida" pageId="0" pageNumber="151" rank="species" species="timida">
<emphasis id="837E9371F583B669344C4E17433C0DAA" italics="true" pageId="0" pageNumber="151">T. timida</emphasis>
</taxonomicName>
Kl. [
<normalizedToken id="DCCC1DA40765A06CD0D9C1F53F3B1AA9" originalValue="Klug’s">Klug's</normalizedToken>
handwriting]&quot;, &quot;GBIF-GISHym 2317&quot;, &quot;Zool. Mus. Berlin&quot; [printed], &quot;
<taxonomicName id="EAC112435C9C895539BD67FF4BC8D988" genus="Lectotypus" lsidName="Lectotypus" pageId="0" pageNumber="151" rank="genus">
<emphasis id="D73A876762991B29B46A04A910584B99" italics="true" pageId="0" pageNumber="151">Lectotypus</emphasis>
</taxonomicName>
[printed] ♂
<taxonomicName id="6771FF1C1787076100861F122D33209F" authority="Klug, 1818" class="Insecta" family="Tenthredinidae" genus="Tenthredo" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tenthredo (Dolerus) timida" order="Hymenoptera" pageId="0" pageNumber="151" phylum="Arthropoda" rank="species" species="timida" subGenus="Dolerus">Tenthredo (Dolerus) timida</taxonomicName>
Klug, 1818 [handwritten in block letters] M.Heidemaa design. [printed]&quot; [red], &quot;
<taxonomicName id="CB096B4FED44E82A6A869724E15CA167" authority="(Klug, 1818)" baseAuthorityName="Klug" baseAuthorityYear="1818" class="Insecta" family="Tenthredinidae" genus="Dolerus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dolerus timidus" order="Hymenoptera" pageId="0" pageNumber="151" phylum="Arthropoda" rank="species" species="timidus">
<emphasis id="579A48A96226E50F22E00623053CE6C5" italics="true" pageId="0" pageNumber="151">Dolerus timidus</emphasis>
</taxonomicName>
(Klug, 1818) M.Heidemaa det.&quot; [printed]. Good condition, penis valve dissected, pinned with the specimen (Fig.
<figureCitation id="97397E9ED2090F5193F57BFE65407BC5" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Dolerus timidus (Klug, 1818), D. variator Enslin, 1927 and D. pratensis (Linnaeus, 1758). A. Penis valve, lectotype Tenthredo timida; B. Penis valve, D. variator; C. Metepimeron (arrow), D. pratensis (ZMUO. 032881); D. Metepimeron (arrow), D. timidus (ZMUO. 029409), mirrored; E. Lancet, D. timidus; F. Lancet, D. pratensis." figureDoi="10.3897/dez.69.84080.figure14" httpUri="https://binary.pensoft.net/fig/719525" pageId="0" pageNumber="151">14A</figureCitation>
). Paralectotypes: 4♂ 5♀ (GBIF-GISHym: 2312, 2318, 2320-2326). All in ZMHB.
</paragraph>
<paragraph id="B5435B6D31E2EFE6B03A299C0956E156" pageId="0" pageNumber="151">
<taxonomicName id="EFEF26133802F3F6EF41A8E9330BB1CC" authority="Klug, 1818" class="Insecta" family="Tenthredinidae" genus="Tenthredo" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tenthredo (Dolerus) dubia" order="Hymenoptera" pageId="0" pageNumber="151" phylum="Arthropoda" rank="species" species="dubia" subGenus="Dolerus">Tenthredo (Dolerus) dubia</taxonomicName>
Klug, 1818. Lectotype ♂ hereby designated, labelled:
<normalizedToken id="5C2782781B3E0A487B917CF68771375E" originalValue="“14198”">&quot;14198&quot;</normalizedToken>
[catalog no.], &quot;
<taxonomicName id="A5FCE78E4F8F21D2763F40D49A9D0B11" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Insecta" family="Tenthredinidae" genus="Dolerus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dolerus pratensis" order="Hymenoptera" pageId="0" pageNumber="151" phylum="Arthropoda" rank="species" species="pratensis">
<emphasis id="EB8B1B8B069A076F5E19623C22102DEE" italics="true" pageId="0" pageNumber="151">Dolerus pratensis</emphasis>
</taxonomicName>
(L.) ♂ [handwritten] det. F. Koch [printed]&quot;, &quot;GBIF-GISHym 2315&quot; [printed], &quot;Zool. Mus. Berlin&quot; [printed], &quot;
<taxonomicName id="A330003FA676639BC474778E189391D4" genus="Lectotypus" lsidName="Lectotypus" pageId="0" pageNumber="151" rank="genus">
<emphasis id="B3A0F100778034096139AF9BACBACAC8" italics="true" pageId="0" pageNumber="151">Lectotypus</emphasis>
</taxonomicName>
[printed] ♂
<taxonomicName id="79A4C95E9107B97657A6453B32D88824" authority="Klug, 1818" class="Insecta" family="Tenthredinidae" genus="Tenthredo" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tenthredo (Dolerus) dubia" order="Hymenoptera" pageId="0" pageNumber="151" phylum="Arthropoda" rank="species" species="dubia" subGenus="Dolerus">Tenthredo (Dolerus) dubia</taxonomicName>
Klug, 1818 [handwritten in block letters] M.Heidemaa design. [printed]&quot; [red], &quot;
<taxonomicName id="1C7F95CD43BD5ED44D221807B3FA73DC" authority="(Klug, 1818)" baseAuthorityName="Klug" baseAuthorityYear="1818" class="Insecta" family="Tenthredinidae" genus="Dolerus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dolerus timidus" order="Hymenoptera" pageId="0" pageNumber="151" phylum="Arthropoda" rank="species" species="timidus">
<emphasis id="01CC89A751407AC9F4189BEA55C9258D" italics="true" pageId="0" pageNumber="151">Dolerus timidus</emphasis>
</taxonomicName>
(Klug, 1818) M.Heidemaa det.&quot; [printed], Condition satisfactory, genitalia dissected, penis valves on a slide pinned with the specimen (they fit
<taxonomicName id="69381A0F6A64435A364243A5254156CA" class="Insecta" family="Tenthredinidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="D. timidus" order="Hymenoptera" pageId="0" pageNumber="151" phylum="Arthropoda" rank="species" species="timidus">
<emphasis id="2DF2E25A5179A9F16B5001FDE78F2181" italics="true" pageId="0" pageNumber="151">D. timidus</emphasis>
</taxonomicName>
). Paralectotypes: 1♂, 1♀ (&quot;GBIF-GISHym 2313&quot;,
<normalizedToken id="DFCFBEA4389102D22C5CD9E8E00CE2DB" originalValue="“…">&quot;...</normalizedToken>
2314&quot;). All in ZMHB.
</paragraph>
<paragraph id="0DA749CAFB643BACB1984C2E82558EA8" pageId="0" pageNumber="151">
<taxonomicName id="B50525972C5FCA41F75D4522C03279A7" authority="Enslin, 1913" class="Insecta" family="Tenthredinidae" genus="Dolerus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dolerus dubius subsp. var. var. atratus" order="Hymenoptera" pageId="0" pageNumber="151" phylum="Arthropoda" rank="variety" species="dubius" subSpecies="var." variety="atratus">Dolerus dubius var. atratus</taxonomicName>
Enslin, 1913. Lectotype ♂ hereby designated, labelled:
<normalizedToken id="E439FF494A4B0009ECC8452E012C758E" originalValue="“Helvet.[ia]”">&quot;Helvet.[ia]&quot;</normalizedToken>
,
<normalizedToken id="FB3D887AEA3866D974BB22A9DAC7DFBB" originalValue="“Type”">&quot;Type&quot;</normalizedToken>
[pink, printed], &quot;
<taxonomicName id="454FE7F7461F7C11F1E613A8EA3E7271" class="Insecta" family="Tenthredinidae" genus="Dolerus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dolerus" order="Hymenoptera" pageId="0" pageNumber="151" phylum="Arthropoda" rank="genus">
<emphasis id="C330617E164BC36411A8C3DB919DA4A6" italics="true" pageId="0" pageNumber="151">Dolerus</emphasis>
</taxonomicName>
♂
<taxonomicName id="77445F70FBDADDB9120DF80129906D46" authorityName="Klug" authorityYear="1818" class="Insecta" family="Tenthredinidae" genus="Dolerus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dolerus dubius" order="Hymenoptera" pageId="0" pageNumber="151" phylum="Arthropoda" rank="species" species="dubius">
<emphasis id="EB2C956D27F9E05115B2BFE8CA359F0F" italics="true" pageId="0" pageNumber="151">Dolerus dubius</emphasis>
</taxonomicName>
<taxonomicName id="BC431C7177F0482D76074594EF3F10C3" authority=" Enslin, 1913" class="Insecta" family="Tenthredinidae" genus="Dolerus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dolerus dubius subsp. var var. atratus" order="Hymenoptera" pageId="0" pageNumber="151" phylum="Arthropoda" rank="variety" species="dubius" variety="atratus">Dolerus dubius var atratus</taxonomicName>
[handwritten] Dr. Enslin det. [printed]&quot;, &quot;Sammlung Dr. Enslin&quot; [printed], &quot;GBIF-GISHym 3102&quot;, &quot;LECTOTYPE ♂
<taxonomicName id="EFD8F4902BF3D57C459AD721137E6D5C" authority="Enslin, 1913" class="Insecta" family="Tenthredinidae" genus="Dolerus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dolerus dubius subsp. var. var. atratus" order="Hymenoptera" pageId="0" pageNumber="151" phylum="Arthropoda" rank="variety" species="dubius" subSpecies="var." variety="atratus">Dolerus dubius var. atratus</taxonomicName>
Enslin, 1913 designated M. Heidemaa 2022&quot; [red, printed]. Some damage: left flagellum missing, abdomen glued to a card together with the dissected penis valves and pinned with the specimen. ZSM.
</paragraph>
<paragraph id="499201693CBBC9C575EE1FCEFDFD7D61" pageId="0" pageNumber="151">
This melanic color form was identified as
<taxonomicName id="234D28D4DA7E905D9E4D8411BFA19354" class="Insecta" family="Tenthredinidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="D. timidus" order="Hymenoptera" pageId="0" pageNumber="151" phylum="Arthropoda" rank="species" species="timidus">
<emphasis id="9FAAC0D31830669136D614723F91ABE4" italics="true" pageId="0" pageNumber="151">D. timidus</emphasis>
</taxonomicName>
based on its penis valve structure and the sculpture of the metepimeron.
</paragraph>
</subSubSection>
</treatment>
</document>
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