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<document id="26F66CB55FE2AADA5F87A8F05D94A287" ID-CLB-Dataset="29071" ID-DOI="10.11646/zootaxa.4457.2.10" ID-GBIF-Dataset="014008ab-6070-47e8-b051-002d4c24b07b" ID-ISSN="1175-5326" ID-Zenodo-Dep="1457686" ID-ZooBank="07406594-120E-41D4-B1CE-CA13B274DA02" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1539265245780" checkinUser="plazi" docAuthor="Heiss, Ernst &amp; Guilbert, Eric" docDate="2018" docId="800887D5FFB79664ABFAFDEB10CCFBBB" docLanguage="en" docName="zootaxa.4457.2.10.pdf" docOrigin="Zootaxa 4457 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Burmacader lativentris Heiss &amp; Guilbert 2018, sp.nov." docType="treatment" docVersion="7" lastPageNumber="344" masterDocId="7C31FFADFFB59661AB6DFF951469FFE3" masterDocTitle="A new species of Tingidae from Cretaceous amber from Myanmar (Hemiptera, Heteroptera)" masterLastPageNumber="345" masterPageNumber="339" pageNumber="341" updateTime="1698576900079" updateUser="ExternalLinkService">
<mods:mods id="FD965CBF6CE4B208EE494934BA14CC1B" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E6078E18C61829AD42F6F207579C7219">
<mods:title id="E876DFFDECEE61561E4A127CDE9EF76C">A new species of Tingidae from Cretaceous amber from Myanmar (Hemiptera, Heteroptera)</mods:title>
</mods:titleInfo>
<mods:name id="31AD3FE3C4A7E18E11CD9B5868084758" type="personal">
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<mods:roleTerm id="BFCA2C763E6E2E11018D11A931759EBE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B0B56B58E31A78CB9E36C6E056C42A4A">Heiss, Ernst</mods:namePart>
</mods:name>
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<mods:role id="C8EED4C5C5F49609116B85AB50F14DEA">
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<mods:namePart id="1A5AB00870AE35DE6FE7991EC847871B">Guilbert, Eric</mods:namePart>
</mods:name>
<mods:typeOfResource id="67A3FC6A1E57B74F2A5A9B982DD36F3F">text</mods:typeOfResource>
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<mods:title id="89B0F34668B64CC2700BDA12423E1306">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="50E724BE1BF08229068D91E045258A00">
<mods:date id="5E2C42FBEEE286EB3B4A84465266D0B5">2018</mods:date>
<mods:detail id="A523BBCF3B386F132FB847A03D70C837" type="pubDate">
<mods:number id="3B79D43FB3967A3E54418AD1C284FD12">2018-08-08</mods:number>
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<mods:number id="1EBB61F33F5570E3D0D12B12433FF32C">4457</mods:number>
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<mods:classification id="5D5CDFBBCA229B93415CB3CF54DCFC89">journal article</mods:classification>
<mods:identifier id="C6DF68954A8FB78F3DDDE997031262A6" type="CLB-Dataset">29071</mods:identifier>
<mods:identifier id="F3E68CAE84E03BBD7CFD13A34A184871" type="DOI">10.11646/zootaxa.4457.2.10</mods:identifier>
<mods:identifier id="A8EB096AFE6E257D2041338A823F71B8" type="GBIF-Dataset">014008ab-6070-47e8-b051-002d4c24b07b</mods:identifier>
<mods:identifier id="251EDC9D3CAE5E51D79422BEA268520D" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="5FA76F4F1DE61C53AE5D69EF199856BD" type="Zenodo-Dep">1457686</mods:identifier>
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<treatment id="800887D5FFB79664ABFAFDEB10CCFBBB" ID-DOI="http://doi.org/10.5281/zenodo.5957734" ID-GBIF-Taxon="148692042" ID-Zenodo-Dep="5957734" LSID="urn:lsid:plazi:treatment:800887D5FFB79664ABFAFDEB10CCFBBB" httpUri="http://treatment.plazi.org/id/800887D5FFB79664ABFAFDEB10CCFBBB" lastPageId="5" lastPageNumber="344" pageId="2" pageNumber="341">
<subSubSection id="40BB6548FFB79663ABFAFDEB157BFD59" pageId="2" pageNumber="341" type="nomenclature">
<paragraph id="081E36C3FFB79663ABFAFDEB1662FD7B" blockId="2.[151,523,638,698]" box="[151,523,638,664]" pageId="2" pageNumber="341">
<heading id="535681AFFFB79663ABFAFDEB1662FD7B" bold="true" box="[151,523,638,664]" fontSize="11" level="1" pageId="2" pageNumber="341" reason="1">
<emphasis id="3AD5EAD1FFB79663ABFAFDEB1662FD7B" bold="true" box="[151,523,638,664]" pageId="2" pageNumber="341">
<taxonomicName id="CFA14D40FFB79663ABFAFDEB15C7FD7B" authority="Heiss &amp; Guilbert, 2018" authorityName="Heiss &amp; Guilbert" authorityYear="2018" box="[151,430,638,664]" class="Insecta" family="Tingidae" genus="Burmacader" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="341" phylum="Arthropoda" rank="species" species="lativentris" status="sp. nov.">
<emphasis id="3AD5EAD1FFB79663ABFAFDEB15C7FD7B" bold="true" box="[151,430,638,664]" italics="true" pageId="2" pageNumber="341">Burmacader lativentris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="21E657AAFFB79663AAD8FDEB1662FD7B" box="[437,523,638,664]" pageId="2" pageNumber="341" rank="species">sp.nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="081E36C3FFB79663ABFAFD34157BFD59" blockId="2.[151,523,638,698]" box="[151,274,673,698]" pageId="2" pageNumber="341">
(
<figureCitation id="909A2A46FFB79663ABF2FD341560FD59" box="[159,265,673,698]" captionStart-0="FIGURES 1–2" captionStart-1="FIGURES 3–5" captionStart-2="FIGURES 6–9" captionStartId-0="1.[151,264,1896,1919]" captionStartId-1="3.[151,264,1754,1777]" captionStartId-2="4.[151,264,1955,1978]" captionTargetBox-1="[170,1426,216,1701]" captionTargetBox-2="[202,1397,209,1938]" captionTargetId-1="figure@3.[152,1435,193,1733]" captionTargetId-2="figure@4.[178,1415,193,1942]" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURES 1–2. Burmacader reconstruction of habitus. 1—Burmacader multivenosus; 2—Burmacader lativentris sp.nov. Abbreviations: ca—costal area, da—discoidal area; mb—membrane; pa—paranotum; sa—sutural area; sca—subcostal area. Scale 0.5mm." captionText-1="FIGURES 3–5. Burmacader lativentris sp.nov., amber inclusion. 3—holotype dorsal view; 4—dito ventral view. 5—head and pronotum." captionText-2="FIGURES 6–9. Burmacader lativentris sp.nov. 5—head, pronotum, scutellum dorsal view; 6—head, thorax ventral view; 7—hemelytra dorsal; 8—abdomen, hemelytra ventral." httpUri-1="https://zenodo.org/record/1457688/files/figure.png" httpUri-2="https://zenodo.org/record/1457690/files/figure.png" pageId="2" pageNumber="341">Figs. 2–9</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="40BB6548FFB79663ABFAFD7C17BCFCA9" pageId="2" pageNumber="341" type="materials_examined">
<paragraph id="081E36C3FFB79663ABFAFD7C17BCFCA9" blockId="2.[151,1437,745,2030]" pageId="2" pageNumber="341">
<emphasis id="3AD5EAD1FFB79663ABFAFD7C152EFCE1" bold="true" box="[151,327,745,770]" pageId="2" pageNumber="341">
<typeStatus id="D71A8861FFB79663ABFAFD7C14BBFCE1" box="[151,210,745,770]" pageId="2" pageNumber="341">Type</typeStatus>
material:
</emphasis>
<typeStatus id="D71A8861FFB79663AA22FD7C15DEFCE1" box="[335,439,745,770]" pageId="2" pageNumber="341" type="holotype">Holotype</typeStatus>
female in a
<date id="7C1F1003FFB79663A928FD7F16E0FCE1" box="[581,649,746,770]" pageId="2" pageNumber="341" value="2012-10-18">18x12</date>
x
<quantity id="CF599B26FFB79663A9F5FD7F16B3FCE2" box="[664,730,746,770]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="2" pageNumber="341" unit="mm" value="4.0">4 mm</quantity>
cabochon shaped piece of Burmese Amber, dorsal and ventral surface structures partly obscured by impurities and small air bubbles. This specimen is designated as
<typeStatus id="D71A8861FFB79663AE66FC981104FCC5" box="[1291,1389,781,806]" pageId="2" pageNumber="341" type="holotype">holotype</typeStatus>
and is deposited in the collection of the first author (
<collectionCode id="6EB0AE06FFB79663A9DDFCA7169DFCA9" box="[688,756,818,842]" pageId="2" pageNumber="341">CEHI</collectionCode>
) as BUB-TING-06.
</paragraph>
</subSubSection>
<subSubSection id="40BB6548FFB79663ABAAFCC016AAFBA5" pageId="2" pageNumber="341" type="diagnosis">
<paragraph id="081E36C3FFB79663ABAAFCC0162FFC71" blockId="2.[151,1437,745,2030]" pageId="2" pageNumber="341">
<emphasis id="3AD5EAD1FFB79663ABAAFCC01553FC8D" bold="true" box="[199,314,853,878]" pageId="2" pageNumber="341">Diagnosis</emphasis>
. This taxon shares the following diagnostic characters of
<taxonomicName id="CFA14D40FFB79663A8A3FCC01033FC8D" authorityName="Heiss &amp; Guilbert" authorityYear="2018" box="[974,1114,853,878]" class="Insecta" family="Tingidae" genus="Burmacader" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="341" phylum="Arthropoda" rank="genus">
<emphasis id="3AD5EAD1FFB79663A8A3FCC01033FC8D" box="[974,1114,853,878]" italics="true" pageId="2" pageNumber="341">Burmacader</emphasis>
</taxonomicName>
and is therefore assigned as the second species to this fossil genus.
</paragraph>
<paragraph id="081E36C3FFB79663ABAAFC0B16AAFBA5" blockId="2.[151,1437,745,2030]" pageId="2" pageNumber="341">Macropterous. Body elongate, surface punctate and areolate, lateral margins setigerous; head short without spines; antennae short about 2.2x as long as width of head, segments III and IV longest; pronotum with anterolaterally expanded paranota, disk without longitudinal carinae; scutellum distinct; hemelytra divided into costal, subcostal and discoidal areas, subcostal one with about ten, discoidal one with three to four transverse veins, stenocostal area absent; membrane without veins.</paragraph>
</subSubSection>
<subSubSection id="40BB6548FFB79664ABAAFBC4114BFF37" lastPageId="5" lastPageNumber="344" pageId="2" pageNumber="341" type="description">
<paragraph id="081E36C3FFB79663ABAAFBC415DDFB51" blockId="2.[151,1437,745,2030]" pageId="2" pageNumber="341">
<emphasis id="3AD5EAD1FFB79663ABAAFBC41538FB89" bold="true" box="[199,337,1105,1130]" pageId="2" pageNumber="341">Description</emphasis>
. Body length
<quantity id="CF599B26FFB79663AA94FBC41609FB8A" box="[505,608,1105,1130]" metricMagnitude="-3" metricUnit="m" metricValue="3.15" pageId="2" pageNumber="341" unit="mm" value="3.15">3.15 mm</quantity>
of oval outline, lateral margins of pronotum and hemelytra with sparse small setigerous tubercles, few erect setae also present on main hemelytral veins; head, antennae and pronotum with sparse erect pilosity.
</paragraph>
<paragraph id="081E36C3FFB79663ABAAFB28156DFA4D" blockId="2.[151,1437,745,2030]" pageId="2" pageNumber="341">
<emphasis id="3AD5EAD1FFB79663ABAAFB28156DFB35" box="[199,260,1213,1238]" italics="true" pageId="2" pageNumber="341">Head</emphasis>
. Short and truncate, 3.67 times as wide as long, apices of genae slightly exceeding apically rounded clypeus; surface punctured without distinct spines or larger tubercles, eyes large and globose; antennae slender about 2.23x as long as width of head and longer than pronotum width; antennal segment I shortest and thickest (visible from below), II+III thinner, II slightly tapering toward base, III cylindrical, IV elongate spindle-shaped; length of antennal segment I/II/III/IV = 0.15?/0.225/0.425/
<quantity id="CF599B26FFB79663A859FAD817C8FA85" box="[820,929,1357,1382]" metricMagnitude="-4" metricUnit="m" metricValue="4.25" pageId="2" pageNumber="341" unit="mm" value="0.425">0.425mm</quantity>
. Labial groove delimited by lateral bucculae on anterior half of head, there semiroundly expanded ventrally with two to three rows of cells; length of rostrum obscured.
</paragraph>
<paragraph id="081E36C3FFB79663ABAAFA2F1579F965" blockId="2.[151,1437,745,2030]" pageId="2" pageNumber="341">
<emphasis id="3AD5EAD1FFB79663ABAAFA2F155FFA32" box="[199,310,1466,1489]" italics="true" pageId="2" pageNumber="341">Pronotum</emphasis>
. Surface with deep cell-like punctures, without longitudinal carinae, lateral margins rounded at humeri then converging anteriorly, anterolaterally with large rounded expanded paranota with four rows of cells on widest part, these somewhat larger than those of pronotum but of the same size as areoles of hemelytra; anterior lobe of disk constricted, produced and overlapping the head between paranota; posterior lobe strongly elevated, separated from anterior lobe by a transverse depression, posterior margin slightly sinuate without posterior projection.
</paragraph>
<paragraph id="081E36C3FFB79663ABAAF904173EF949" blockId="2.[151,1437,745,2030]" box="[199,855,1681,1706]" pageId="2" pageNumber="341">
<emphasis id="3AD5EAD1FFB79663ABAAF904155CF949" box="[199,309,1681,1706]" italics="true" pageId="2" pageNumber="341">Scutellum</emphasis>
. Triangular with knob-like apex, surface rugose.
</paragraph>
<paragraph id="081E36C3FFB79663ABAAF92015DCF80E" blockId="2.[151,1437,745,2030]" pageId="2" pageNumber="341">
<emphasis id="3AD5EAD1FFB79663ABAAF9201554F92D" box="[199,317,1717,1742]" italics="true" pageId="2" pageNumber="341">Hemelytra</emphasis>
. Macropterous, distinctly surpassing apex of abdomen by one quarter of their length; clavus triangular with a row of cells along outer margin, irregularly punctured elsewhere, apex reaching about half of length of hemelytra; costal area with three rows of cells at middle, one row posteriorly and four to five cells at sinuate expansion anteriorly; subcostal area divided by about ten transverse veins with six to seven transverse rows of cells at its widest part; discoidal area flat with three to four partly indistinct transverse veins and seven to eight cells at the median widest area; sutural area basally bordering clavus with one row of cells, following inner margin of R+M veins with one to two cells along membrane to its posterior end; membrane without veins; a short hypocostal lamina with one row of cells on anterior half is ventrally developed along insinuation of costal area reaching to anterior apex.
</paragraph>
<caption id="5CDE664BFFB69662ABFAF94F1568F8EC" httpUri="https://zenodo.org/record/1457688/files/figure.png" pageId="3" pageNumber="342" startId="3.[151,264,1754,1777]" targetBox="[170,1426,216,1701]" targetPageId="3">
<paragraph id="081E36C3FFB69662ABFAF94F1568F8EC" blockId="3.[151,1436,1754,1807]" pageId="3" pageNumber="342">
<emphasis id="3AD5EAD1FFB69662ABFAF94F1553F913" bold="true" box="[151,314,1754,1777]" pageId="3" pageNumber="342">FIGURES 3–5.</emphasis>
<taxonomicName id="CFA14D40FFB69662AA2DF94E1645F913" authority="Heiss &amp; Guilbert, 2018" authorityName="Heiss &amp; Guilbert" authorityYear="2018" box="[320,556,1755,1776]" class="Insecta" family="Tingidae" genus="Burmacader" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="342" phylum="Arthropoda" rank="species" species="lativentris" status="sp. nov.">
<emphasis id="3AD5EAD1FFB69662AA2DF94E1645F913" box="[320,556,1755,1776]" italics="true" pageId="3" pageNumber="342">Burmacader lativentris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="21E657AAFFB69662A95FF94E1614F913" box="[562,637,1755,1776]" pageId="3" pageNumber="342" rank="species">
<emphasis id="3AD5EAD1FFB69662A95FF94E1614F913" bold="true" box="[562,637,1755,1776]" pageId="3" pageNumber="342">sp.nov.</emphasis>
</taxonomicNameLabel>
, amber inclusion.
<emphasis id="3AD5EAD1FFB69662A855F94E172CF913" bold="true" box="[824,837,1755,1776]" pageId="3" pageNumber="342">3</emphasis>
—holotype dorsal view;
<emphasis id="3AD5EAD1FFB69662AF52F94E1025F913" bold="true" box="[1087,1100,1755,1776]" pageId="3" pageNumber="342">4</emphasis>
—dito ventral view.
<emphasis id="3AD5EAD1FFB69662AE76F94E1128F913" bold="true" box="[1307,1345,1755,1776]" pageId="3" pageNumber="342">5—</emphasis>
head and pronotum.
</paragraph>
</caption>
<paragraph id="081E36C3FFB69662ABAAF8A81062F87E" blockId="3.[151,1437,1852,2021]" pageId="3" pageNumber="342">
<emphasis id="3AD5EAD1FFB69662ABAAF8A81565F8B7" box="[199,268,1853,1876]" italics="true" pageId="3" pageNumber="342">Venter</emphasis>
. Pro-, meso and metasternum with a wide median labial groove delimited by carinate lateral margins which are subparallel on meso- and metasternum and conically converging on prosternum; sternite VII with a median posteriorly directed projection (subgenital plate,
<bibRefCitation id="6C304B32FFB69662A87FF8111796F87E" box="[786,1023,1924,1949]" pageId="3" pageNumber="342" refId="ref2018" refString="Drake, C. J. &amp; Davis, N. T. (1960) The morphology, phylogeny, and higher classification of the family Tingidae, including description of a new genus and species of the subfamily Vianaidinae (Hemiptera: Heteroptera). Entomologica Americana, 39, 1 - 100." type="journal article">Drake &amp; Davis 1960</bibRefCitation>
).
</paragraph>
<paragraph id="081E36C3FFB69662ABAAF83C1570F806" blockId="3.[151,1437,1852,2021]" pageId="3" pageNumber="342">
<emphasis id="3AD5EAD1FFB69662ABAAF83C1495F823" box="[199,252,1961,1984]" italics="true" pageId="3" pageNumber="342">Legs</emphasis>
. Long and slender, femora not incrassate, tibiae thin and cylindrical, tarsi two-segmented, claws with a basal tooth.
</paragraph>
<caption id="5CDE664BFFB19665ABFAF83616D5F83E" httpUri="https://zenodo.org/record/1457690/files/figure.png" pageId="4" pageNumber="343" startId="4.[151,264,1955,1978]" targetBox="[202,1397,209,1938]" targetPageId="4">
<paragraph id="081E36C3FFB19665ABFAF83616D5F83E" blockId="4.[151,1436,1955,2013]" pageId="4" pageNumber="343">
<emphasis id="3AD5EAD1FFB19665ABFAF8361554F85A" bold="true" box="[151,317,1955,1978]" pageId="4" pageNumber="343">FIGURES 6–9.</emphasis>
<taxonomicName id="CFA14D40FFB19665AA2AF831165EF85A" authority="Heiss &amp; Guilbert, 2018" authorityName="Heiss &amp; Guilbert" authorityYear="2018" box="[327,567,1956,1977]" class="Insecta" family="Tingidae" genus="Burmacader" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="343" phylum="Arthropoda" rank="species" species="lativentris" status="sp. nov.">
<emphasis id="3AD5EAD1FFB19665AA2AF831165EF85A" box="[327,567,1956,1977]" italics="true" pageId="4" pageNumber="343">Burmacader lativentris</emphasis>
</taxonomicName>
<emphasis id="3AD5EAD1FFB19665A92DF83116D3F85A" bold="true" box="[576,698,1956,1977]" pageId="4" pageNumber="343">
<taxonomicNameLabel id="21E657AAFFB19665A92DF83116E2F85A" box="[576,651,1956,1977]" pageId="4" pageNumber="343" rank="species">sp.nov.</taxonomicNameLabel>
5—
</emphasis>
head, pronotum, scutellum dorsal view;
<emphasis id="3AD5EAD1FFB19665AF0EF83110E1F85A" bold="true" box="[1123,1160,1956,1977]" pageId="4" pageNumber="343">6—</emphasis>
head, thorax ventral view;
<emphasis id="3AD5EAD1FFB19665ABFAF85214CDF83F" bold="true" box="[151,164,1991,2012]" pageId="4" pageNumber="343">7</emphasis>
—hemelytra dorsal;
<emphasis id="3AD5EAD1FFB19665AA19F85215F0F83F" bold="true" box="[372,409,1991,2012]" pageId="4" pageNumber="343">8—</emphasis>
abdomen, hemelytra ventral.
</paragraph>
</caption>
<paragraph id="081E36C3FFB09664ABAAFF0C114BFF37" blockId="5.[151,1437,151,1112]" pageId="5" pageNumber="344">
<emphasis id="3AD5EAD1FFB09664ABAAFF0C1500FF53" box="[199,361,153,176]" italics="true" pageId="5" pageNumber="344">Measurements</emphasis>
. Length
<quantity id="CF599B26FFB09664AAB6FF021655FF53" box="[475,572,151,176]" metricMagnitude="-3" metricUnit="m" metricValue="3.15" pageId="5" pageNumber="344" unit="mm" value="3.15">3.15mm</quantity>
; length / width of head 0.15/
<quantity id="CF599B26FFB09664A8DFFF02107BFF53" box="[946,1042,151,176]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" pageId="5" pageNumber="344" unit="mm" value="0.55">0.55mm</quantity>
; pronotum length / width across anterolateral expansions 0.65/
<quantity id="CF599B26FFB09664AA85FF29162FFF36" box="[488,582,188,213]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="5" pageNumber="344" unit="mm" value="1.2">1.20mm</quantity>
; scutellum length / width 0.3/
<quantity id="CF599B26FFB09664A8FEFF29178CFF37" box="[915,997,188,212]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="5" pageNumber="344" unit="mm" value="0.4">0.4mm</quantity>
; width of hemelytra
<quantity id="CF599B26FFB09664AFA1FF291177FF37" box="[1228,1310,188,212]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="5" pageNumber="344" unit="mm" value="1.8">1.8mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="40BB6548FFB09664ABAAFF4A1042FF14" box="[199,1067,223,248]" pageId="5" pageNumber="344" type="etymology">
<paragraph id="081E36C3FFB09664ABAAFF4A1042FF14" blockId="5.[151,1437,151,1112]" box="[199,1067,223,248]" pageId="5" pageNumber="344">
<emphasis id="3AD5EAD1FFB09664ABAAFF4A1527FF1B" bold="true" box="[199,334,223,248]" pageId="5" pageNumber="344">Etymology.</emphasis>
Named after its wide body from&gt;latus&lt;(Latin) wide and venter.
</paragraph>
</subSubSection>
<subSubSection id="40BB6548FFB09664ABAAFE9110CCFBBB" pageId="5" pageNumber="344" type="discussion">
<paragraph id="081E36C3FFB09664ABAAFE911752FE6B" blockId="5.[151,1437,151,1112]" pageId="5" pageNumber="344">
<emphasis id="3AD5EAD1FFB09664ABAAFE91152CFEFE" bold="true" box="[199,325,260,285]" pageId="5" pageNumber="344">Discussion</emphasis>
.
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<emphasis id="3AD5EAD1FFB09664AA38FE911634FEFE" box="[341,605,260,285]" italics="true" pageId="5" pageNumber="344">Burmacader lativentris</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="CFA14D40FFB09664A80FFE91179DFEFE" baseAuthorityName="Heiss &amp; Guilbert" baseAuthorityYear="2013" box="[866,1012,260,285]" class="Insecta" family="Tingidae" genus="Burmacader" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="344" phylum="Arthropoda" rank="species" species="multivenosus">
<emphasis id="3AD5EAD1FFB09664A80FFE91179DFEFE" box="[866,1012,260,285]" italics="true" pageId="5" pageNumber="344">multivenosus</emphasis>
</taxonomicName>
by its larger size (3.15/
<quantity id="CF599B26FFB09664AE64FE911101FEFE" box="[1289,1384,260,285]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="5" pageNumber="344" unit="mm" value="2.8">2.80mm</quantity>
) the much wider habitus (width of abdomen 1.80/
<quantity id="CF599B26FFB09664A9CDFEBD1769FEA3" box="[672,768,296,320]" metricMagnitude="-3" metricUnit="m" metricValue="1.32" pageId="5" pageNumber="344" unit="mm" value="1.32">1.32mm</quantity>
), the shorter antennae (ratio length of antennae / width of head 2.23/2.50), the more rounded shape of anterolateral paranotal expansions and the anterior part of costal area of hemelytra showing four to five cells (three in
<taxonomicName id="CFA14D40FFB09664A9F5FEFA1743FE6B" baseAuthorityName="Heiss &amp; Guilbert" baseAuthorityYear="2013" box="[664,810,367,392]" class="Insecta" family="Tingidae" genus="Burmacader" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="344" phylum="Arthropoda" rank="species" species="multivenosus">
<emphasis id="3AD5EAD1FFB09664A9F5FEFA1743FE6B" box="[664,810,367,392]" italics="true" pageId="5" pageNumber="344">multivenosus</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="081E36C3FFB09664ABAAFE0110ABFDFB" blockId="5.[151,1437,151,1112]" pageId="5" pageNumber="344">
The greatest difference seems to concern the size and shape of eyes, which are large and globular in
<taxonomicName id="CFA14D40FFB09664AE47FE0111F5FE4E" authorityName="Heiss &amp; Guilbert" authorityYear="2018" box="[1322,1436,404,429]" class="Insecta" family="Tingidae" genus="Burmacader" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="344" phylum="Arthropoda" rank="species" species="lativentris">
<emphasis id="3AD5EAD1FFB09664AE47FE0111F5FE4E" box="[1322,1436,404,429]" italics="true" pageId="5" pageNumber="344">lativentris</emphasis>
</taxonomicName>
and versus the much smaller size documented in the reconstruction of
<taxonomicName id="CFA14D40FFB09664A8AEFE22103CFE33" baseAuthorityName="Heiss &amp; Guilbert" baseAuthorityYear="2013" box="[963,1109,439,464]" class="Insecta" family="Tingidae" genus="Burmacader" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="344" phylum="Arthropoda" rank="species" species="multivenosus">
<emphasis id="3AD5EAD1FFB09664A8AEFE22103CFE33" box="[963,1109,439,464]" italics="true" pageId="5" pageNumber="344">multivenosus</emphasis>
</taxonomicName>
(see
<figureCitation id="909A2A46FFB09664AFF9FE2210B9FE33" box="[1172,1232,439,464]" captionStart="FIGURES 3–5" captionStartId="3.[151,264,1754,1777]" captionTargetBox="[170,1426,216,1701]" captionTargetId="figure@3.[152,1435,193,1733]" captionTargetPageId="3" captionText="FIGURES 3–5. Burmacader lativentris sp.nov., amber inclusion. 3—holotype dorsal view; 4—dito ventral view. 5—head and pronotum." httpUri="https://zenodo.org/record/1457688/files/figure.png" pageId="5" pageNumber="344">Fig.4</figureCitation>
<bibRefCitation id="6C304B32FFB09664AFB5FE2214BAFE17" pageId="5" pageNumber="344" refId="ref2189" refString="Heiss, E. &amp; Guilbert, E. (2013) Two new genera and species of Tingidae from Cretaceous amber from Myanmar (Burma) (Hemiptera: Heteroptera). Zootaxa, 3736 (4), 379 - 386. https: // doi. org / 10.11646 / zootaxa. 3736.4.5" type="journal article">Heiss &amp; Guilbert 2013</bibRefCitation>
), where its eyes were damaged and displaced. Re-examination of the
<typeStatus id="D71A8861FFB09664AF43FE481037FE16" box="[1070,1118,477,501]" pageId="5" pageNumber="344">type</typeStatus>
specimen of
<taxonomicName id="CFA14D40FFB09664AE67FE4911F5FE16" baseAuthorityName="Heiss &amp; Guilbert" baseAuthorityYear="2013" box="[1290,1436,476,501]" class="Insecta" family="Tingidae" genus="Burmacader" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="344" phylum="Arthropoda" rank="species" species="multivenosus">
<emphasis id="3AD5EAD1FFB09664AE67FE4911F5FE16" box="[1290,1436,476,501]" italics="true" pageId="5" pageNumber="344">multivenosus</emphasis>
</taxonomicName>
confirmed the wrong assumption of the eyes size and position, which are rediagnosed in
<figureCitation id="909A2A46FFB09664AF14FE6A10D5FDFB" box="[1145,1212,511,536]" captionStart="FIGURES 1–2" captionStartId="1.[151,264,1896,1919]" captionText="FIGURES 1–2. Burmacader reconstruction of habitus. 1—Burmacader multivenosus; 2—Burmacader lativentris sp.nov. Abbreviations: ca—costal area, da—discoidal area; mb—membrane; pa—paranotum; sa—sutural area; sca—subcostal area. Scale 0.5mm." pageId="5" pageNumber="344">Fig. 1</figureCitation>
.
</paragraph>
<paragraph id="081E36C3FFB09664ABAAFDB11082FC2B" blockId="5.[151,1437,151,1112]" pageId="5" pageNumber="344">
As stated in
<bibRefCitation id="6C304B32FFB09664AA3AFDB11619FDDF" box="[343,624,548,573]" pageId="5" pageNumber="344" refId="ref2189" refString="Heiss, E. &amp; Guilbert, E. (2013) Two new genera and species of Tingidae from Cretaceous amber from Myanmar (Burma) (Hemiptera: Heteroptera). Zootaxa, 3736 (4), 379 - 386. https: // doi. org / 10.11646 / zootaxa. 3736.4.5" type="journal article">Heiss &amp; Guilbert (2013)</bibRefCitation>
,
<taxonomicName id="CFA14D40FFB09664A910FDB11760FDDE" authorityName="Heiss &amp; Guilbert" authorityYear="2018" box="[637,777,548,573]" class="Insecta" family="Tingidae" genus="Burmacader" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="344" phylum="Arthropoda" rank="genus">
<emphasis id="3AD5EAD1FFB09664A910FDB11760FDDE" box="[637,777,548,573]" italics="true" pageId="5" pageNumber="344">Burmacader</emphasis>
</taxonomicName>
has a labial groove on the thoracic sternum and bucculae extending from the clypeus to the posterior margin of the head, two characters apomorphic to the
<taxonomicName id="CFA14D40FFB09664AE7CFDD2111EFD83" authorityName="Laporte" authorityYear="1832" box="[1297,1399,583,608]" class="Insecta" family="Tingidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="344" phylum="Arthropoda" rank="family">Tingidae</taxonomicName>
. In addition, the habitus of
<taxonomicName id="CFA14D40FFB09664AAF2FDF91642FD66" authorityName="Heiss &amp; Guilbert" authorityYear="2018" box="[415,555,620,645]" class="Insecta" family="Tingidae" genus="Burmacader" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="344" phylum="Arthropoda" rank="genus">
<emphasis id="3AD5EAD1FFB09664AAF2FDF91642FD66" box="[415,555,620,645]" italics="true" pageId="5" pageNumber="344">Burmacader</emphasis>
</taxonomicName>
is clearly the one of tingid body form with the relatively flat shape with dorsal inflations on the head, pronotum and wings that correspond with the areolae.
<taxonomicName id="CFA14D40FFB09664AF31FD1A1081FD4B" authorityName="Heiss &amp; Guilbert" authorityYear="2018" box="[1116,1256,655,680]" class="Insecta" family="Tingidae" genus="Burmacader" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="344" phylum="Arthropoda" rank="genus">
<emphasis id="3AD5EAD1FFB09664AF31FD1A1081FD4B" box="[1116,1256,655,680]" italics="true" pageId="5" pageNumber="344">Burmacader</emphasis>
</taxonomicName>
shares several characteristics with the
<taxonomicName id="CFA14D40FFB09664AAC0FD21165EFD2E" box="[429,567,692,717]" class="Insecta" family="Tingidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="344" phylum="Arthropoda" rank="subFamily" subFamily="Vianaidinae">Vianaidinae</taxonomicName>
: the length of antennal segment II subequal to segment III, and the welldeveloped scutellum. It also shares characters with
<emphasis id="3AD5EAD1FFB09664A992FD4D1045FD13" box="[767,1068,727,752]" italics="true" pageId="5" pageNumber="344">Annomatocoris bolivianus</emphasis>
Schuh, Cassis &amp; Guilbert and
<taxonomicName id="CFA14D40FFB09664ABFAFD681603FCF7" authority="Golub &amp; Popov" authorityName="Golub &amp; Popov" authorityYear="2000" box="[151,618,764,789]" class="Insecta" family="Tingidae" genus="Vianagramma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="344" phylum="Arthropoda" rank="species" species="goldmani">
<emphasis id="3AD5EAD1FFB09664ABFAFD6815C0FCF6" box="[151,425,764,789]" italics="true" pageId="5" pageNumber="344">Vianagramma goldmani</emphasis>
Golub &amp; Popov
</taxonomicName>
, such as the membranous surface on the sutural area of the hemelytra. However,
<taxonomicName id="CFA14D40FFB09664AA7DFC8A15F5FCDB" authorityName="Heiss &amp; Guilbert" authorityYear="2018" box="[272,412,799,824]" class="Insecta" family="Tingidae" genus="Burmacader" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="344" phylum="Arthropoda" rank="genus">
<emphasis id="3AD5EAD1FFB09664AA7DFC8A15F5FCDB" box="[272,412,799,824]" italics="true" pageId="5" pageNumber="344">Burmacader</emphasis>
</taxonomicName>
and the
<taxonomicName id="CFA14D40FFB09664A961FC8A16FBFCDB" box="[524,658,799,824]" class="Insecta" family="Tingidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="344" phylum="Arthropoda" rank="subFamily" subFamily="Vianaidinae">Vianaidinae</taxonomicName>
differ by the short head not surpassing first antennal segment in
<taxonomicName id="CFA14D40FFB09664ABFAFCD1154BFCBE" authorityName="Heiss &amp; Guilbert" authorityYear="2018" box="[151,290,836,861]" class="Insecta" family="Tingidae" genus="Burmacader" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="344" phylum="Arthropoda" rank="genus">
<emphasis id="3AD5EAD1FFB09664ABFAFCD1154BFCBE" box="[151,290,836,861]" italics="true" pageId="5" pageNumber="344">Burmacader</emphasis>
</taxonomicName>
, the R+M and Cu (cubital) veins are separated at the base of hemelytra (a diagnostic feature of
<taxonomicName id="CFA14D40FFB09664ABFAFCF21493FC63" box="[151,250,871,896]" class="Insecta" family="Tingidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="344" phylum="Arthropoda" rank="subFamily" subFamily="Tinginae">Tinginae</taxonomicName>
+
<taxonomicName id="CFA14D40FFB09664AA7AFCF215A1FC63" baseAuthorityName="Stal" baseAuthorityYear="1873" box="[279,456,871,896]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="344" phylum="Arthropoda" rank="subFamily" subFamily="Cantacaderinae">Cantacaderinae</taxonomicName>
), the discoidal area with transversal veins, the broad costal area, and the small irregular areolae on hemelytra in
<emphasis id="3AD5EAD1FFB09664AAD4FC191623FC46" box="[441,586,908,933]" italics="true" pageId="5" pageNumber="344">
<taxonomicName id="CFA14D40FFB09664AAD4FC19162FFC46" authorityName="Heiss &amp; Guilbert" authorityYear="2018" box="[441,582,908,933]" class="Insecta" family="Tingidae" genus="Burmacader" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="344" phylum="Arthropoda" rank="genus">Burmacader</taxonomicName>
.
</emphasis>
For these reasons
<taxonomicName id="CFA14D40FFB09664A840FC1917D0FC46" authorityName="Heiss &amp; Guilbert" authorityYear="2018" box="[813,953,908,933]" class="Insecta" family="Tingidae" genus="Burmacader" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="344" phylum="Arthropoda" rank="genus">
<emphasis id="3AD5EAD1FFB09664A840FC1917D0FC46" box="[813,953,908,933]" italics="true" pageId="5" pageNumber="344">Burmacader</emphasis>
</taxonomicName>
is definitely neither a
<taxonomicName id="CFA14D40FFB09664AFA0FC19113AFC46" box="[1229,1363,908,933]" class="Insecta" family="Tingidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="344" phylum="Arthropoda" rank="subFamily" subFamily="Vianaidinae">Vianaidinae</taxonomicName>
nor a
<taxonomicName id="CFA14D40FFB09664ABFAFC3A1493FC2B" box="[151,250,943,968]" class="Insecta" family="Tingidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="344" phylum="Arthropoda" rank="subFamily" subFamily="Tinginae">Tinginae</taxonomicName>
or
<taxonomicName id="CFA14D40FFB09664AA72FC3A15A5FC2B" baseAuthorityName="Stal" baseAuthorityYear="1873" box="[287,460,943,968]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="344" phylum="Arthropoda" rank="subFamily" subFamily="Cantacaderinae">Cantacaderinae</taxonomicName>
but may represent an extinct clade basal to
<taxonomicName id="CFA14D40FFB09664A8DBFC3A1070FC2B" box="[950,1049,943,968]" class="Insecta" family="Tingidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="344" phylum="Arthropoda" rank="subFamily" subFamily="Tinginae">Tinginae</taxonomicName>
+
<taxonomicName id="CFA14D40FFB09664AF5BFC3A108FFC2B" baseAuthorityName="Stal" baseAuthorityYear="1873" box="[1078,1254,943,968]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="344" phylum="Arthropoda" rank="subFamily" subFamily="Cantacaderinae">Cantacaderinae</taxonomicName>
.
</paragraph>
<paragraph id="081E36C3FFB09664ABAAFC4110CCFBBB" blockId="5.[151,1437,151,1112]" pageId="5" pageNumber="344">
The placement of these cretaceous amber species, which are by far older than other known
<taxonomicName id="CFA14D40FFB09664AF9DFC4111F4FC0E" baseAuthorityName="Stal" baseAuthorityYear="1873" box="[1264,1437,980,1005]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="344" phylum="Arthropoda" rank="subFamily" subFamily="Cantacaderinae">Cantacaderinae</taxonomicName>
fossils
<bibRefCitation id="6C304B32FFB09664AB8BFC621587FBF3" box="[230,494,1015,1040]" pageId="5" pageNumber="344" refId="ref1909" refString="Cassis, G. &amp; Schuh, R. T. (2010) Systematic methods, fossils, and relationships within Heteroptera (Insecta). Cladistics, 26, 262 - 280. https: // doi. org / 10.1111 / j. 1096 - 0031.2009.00283. x" type="journal article">Cassis &amp; Schuh (2010)</bibRefCitation>
,
<bibRefCitation id="6C304B32FFB09664AA97FC6216C5FBF3" box="[506,684,1015,1040]" pageId="5" pageNumber="344" refId="ref2149" refString="Guilbert, E. (2012) Biogeography of Cantacaderinae Stal (Insecta: Heteroptera: Tingidae) revisited. Invertebrate Systematics, 26, 316 - 322. https: // doi. org / 10.1071 / IS 12010" type="journal article">Guilbert (2012)</bibRefCitation>
,
<bibRefCitation id="6C304B32FFB09664A9D4FC6217DCFBF3" box="[697,949,1015,1040]" pageId="5" pageNumber="344" refId="ref2296" refString="Perrichot, V., Nel, A., Guilbert, E. &amp; Neraudeau, D. (2006) Fossil Tingoidea (Heteroptera: Cimicomorpha) from French Cretaceous amber, including Tingidae and a new family, Ebboidae. Zootaxa, 1203, 57 - 68." type="journal article">
Perrichot
<emphasis id="3AD5EAD1FFB09664A84AFC6C1709FBF3" box="[807,864,1015,1040]" italics="true" pageId="5" pageNumber="344">et al.</emphasis>
(2006)
</bibRefCitation>
,
<bibRefCitation id="6C304B32FFB09664A8ACFC6210F3FBF3" box="[961,1178,1015,1040]" pageId="5" pageNumber="344" refId="ref2347" refString="Schuh, R. T., Cassis, G. &amp; Guilbert, E. (2006) Description of the first recent macropterous species of Vianaidinae (Heteroptera: Tingidae) with comments on the phylogenetic relationships of the family within the Cimicomorpha. Journal of the New York Entomological Society, 114, 38 - 53. https: // doi. org / 10.1664 / 0028 - 7199 (2006) 114 [38: DOTFRM] 2.0. CO; 2" type="journal article">
Schuh
<emphasis id="3AD5EAD1FFB09664AF62FC6C1028FBF3" box="[1039,1089,1015,1040]" italics="true" pageId="5" pageNumber="344">et al</emphasis>
. (2006)
</bibRefCitation>
still needs to be tested in the frame of phylogenetic analyses including biogeographic data. However, still few species are known from Cretaceous amber that this is a significant contribution to our knowledge of tingid evolution.
</paragraph>
</subSubSection>
</treatment>
</document>
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