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<document id="7CB8E134DE1545C78761154172BD9C05" ID-DOI="10.11646/zootaxa.5230.3.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="7561385" ID-ZooBank="993813D9-1D74-4B6F-ACB4-EBE0183D5617" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1674479375851" checkinUser="plazi" docAuthor="Martín, Guillermo San, Lucas, Yolanda &amp; Hutchings, Pat" docDate="2023" docId="6C2887987E57B0444CBB2BC9B15CA5E1" docLanguage="en" docName="zootaxa.5230.3.1.pdf" docOrigin="Zootaxa 5230 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5230.3.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Syllis imajimai Martín &amp; Lucas &amp; Hutchings 2023, n. sp." docType="treatment" docVersion="4" lastPageNumber="266" masterDocId="9011FFE07E5AB04B4C2C2947B326A414" masterDocTitle="The genus Syllis Savigny in Lamarck, 1881 (Annelida: Syllidae: Syllinae) from Australia (Part 3): new species and redescription of previously described species" masterLastPageNumber="295" masterPageNumber="251" pageNumber="264" updateTime="1699447600003" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="79C0867C23FA8F49F5DDB44E4E0B9846" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="062EBB05946E6910B011687C61F06E47">The genus Syllis Savigny in Lamarck, 1881 (Annelida: Syllidae: Syllinae) from Australia (Part 3): new species and redescription of previously described species</mods:title>
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<mods:namePart id="5F25BBBC423EFFD27050F95C66528175">Martín, Guillermo San</mods:namePart>
<mods:affiliation id="C99EC0F43B5F6A1DE4E7FFEEA901C28C">Departamento de Biología (Zoología), Centro de Investigación en Biodiversidad y Cambio global (CIBC-UAM), Facultad de Ciencias, Universidad Autónoma de Madrid, calle Darwin, 2, Canto Blanco, 28049 Madrid, Spain.</mods:affiliation>
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<mods:namePart id="E8EA35B730FE92F5279FBC887CEEDA31">Lucas, Yolanda</mods:namePart>
<mods:affiliation id="2E2BAC814F5E9C8DE2F46FDA27239C8A">Departamento de Biología (Zoología), Centro de Investigación en Biodiversidad y Cambio global (CIBC-UAM), Facultad de Ciencias, Universidad Autónoma de Madrid, calle Darwin, 2, Canto Blanco, 28049 Madrid, Spain.</mods:affiliation>
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<mods:namePart id="3346C5D905A22DB2AA85003593B1676E">Hutchings, Pat</mods:namePart>
<mods:nameIdentifier id="CF71F8F67372BD7BE1708CE53F2B0814" type="ORCID">0000-0001-7521-3930</mods:nameIdentifier>
<mods:affiliation id="5E6611F79B296F2EE4C0B223584B19E3">Australian Museum Research Institute, Australian Museum, 1, William Street, Sydney, NSW, 2010, Australia. pat. hutchings @ austmus. gov. au; https: // orcid. org / 0000 - 0001 - 7521 - 3930 &amp; Biological Sciences, Macquarie University, North Ryde, 2109, Australia.</mods:affiliation>
<mods:nameIdentifier id="871027D54FC2868A07655FD159882F96" type="email">pat.hutchings@austmus.gov.au</mods:nameIdentifier>
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<mods:typeOfResource id="CE0CF1E118D211A9C19687BCA7547A30">text</mods:typeOfResource>
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<mods:title id="554C4638BF758E4B769716A6477C1C66">Zootaxa</mods:title>
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<mods:date id="A94C207F5293FB308E9B47AEA35315D9">2023</mods:date>
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<mods:number id="F052A3454139592B3C3746D7746E4C3A">2023-01-23</mods:number>
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<mods:number id="4AEC4A934A4BAF61C1149EA1D444B452">5230</mods:number>
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<mods:number id="673A06292C891F3DFD7EC19E313D08BC">3</mods:number>
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<treatment id="6C2887987E57B0444CBB2BC9B15CA5E1" ID-DOI="http://doi.org/10.5281/zenodo.7561343" ID-Zenodo-Dep="7561343" LSID="urn:lsid:plazi:treatment:6C2887987E57B0444CBB2BC9B15CA5E1" httpUri="http://treatment.plazi.org/id/6C2887987E57B0444CBB2BC9B15CA5E1" lastPageId="15" lastPageNumber="266" pageId="13" pageNumber="264">
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<emphasis id="D6F5EA9C7E57B0464CBB2BC9B2B3A6BD" bold="true" box="[151,405,654,681]" pageId="13" pageNumber="264">
<taxonomicName id="23814D0D7E57B0464CBB2BC9B26AA6BD" ID-CoL="178acf7a-fe0f-483f-b161-429785788613" authority="Martín &amp; Lucas &amp; Hutchings, 2023" authorityName="Martín &amp; Lucas &amp; Hutchings" authorityYear="2023" box="[151,332,654,681]" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="13" pageNumber="264" phylum="Annelida" rank="species" species="imajimai" status="sp. nov.">
<emphasis id="D6F5EA9C7E57B0464CBB2BC9B26AA6BD" bold="true" box="[151,332,654,681]" italics="true" pageId="13" pageNumber="264">Syllis imajimai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CDC657E77E57B0464D7F2BC8B2B3A6BD" box="[339,405,655,681]" pageId="13" pageNumber="264" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<paragraph id="E43E368E7E57B0464CBB2BF4B3D3A6D9" blockId="13.[151,405,654,717]" box="[151,245,691,717]" pageId="13" pageNumber="264">
<figureCitation id="7CBA2A0B7E57B0464CBB2BF4B3D3A6D9" box="[151,245,691,717]" captionStart="FIGURE 6" captionStartId="14.[152,255,1887,1911]" captionTargetBox="[172,1417,394,1842]" captionTargetId="figure-116@14.[151,1436,381,1864]" captionTargetPageId="14" captionText="FIGURE 6. Syllis imajimai n. sp. Holotype.AM W.196660.A, anterior end, dorsal view. B, midbody parapodium. C, falcigers, anterior parapodium. D, falcigers, midbody parapodium. E, falcigers, posterior parapodium. F, dorsal simple chaeta, midbody parapodium. G, dorsal simple chaeta, posterior parapodium. H, ventral simple chaeta. I, aciculae, anterior parapodium. J, aciculae, midbody parapodium. K, acicula, posterior parapodium. Scale. A: 0.2 mm. B: 50 µm. C–K: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.7561401" httpUri="https://zenodo.org/record/7561401/files/figure.png" pageId="13" pageNumber="264">Figure 6</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="AC9B65057E57B0464CBB2BB0B7F5A71B" box="[151,1235,759,784]" pageId="13" pageNumber="264" type="reference_group">
<paragraph id="E43E368E7E57B0464CBB2BB0B7F5A71B" blockId="13.[151,1235,759,784]" box="[151,1235,759,784]" pageId="13" pageNumber="264">
<treatmentCitationGroup id="C49111A07E57B0464CBB2BB0B7F5A71B" box="[151,1235,759,784]" pageId="13" pageNumber="264">
<taxonomicName id="23814D0D7E57B0464CBB2BB0B27DA71B" ID-CoL="59YLD" authorityName="Krohn" authorityYear="1852" box="[151,347,759,783]" class="Polychaeta" family="Syllidae" genus="Typosyllis" kingdom="Animalia" order="Phyllodocida" pageId="13" pageNumber="264" phylum="Annelida" rank="species" species="prolifera">
<emphasis id="D6F5EA9C7E57B0464CBB2BB0B27DA71B" box="[151,347,759,783]" italics="true" pageId="13" pageNumber="264">Typosyllis prolifera</emphasis>
</taxonomicName>
.—
<treatmentCitation id="6520109F7E57B0464D542BB0B118A71B" author="Imajima, M." box="[376,574,759,783]" page="292" pageId="13" pageNumber="264" year="1966">
<bibRefCitation id="80104B7F7E57B0464D542BB0B118A71B" author="Imajima, M." box="[376,574,759,783]" pageId="13" pageNumber="264" pagination="253 - 294" refId="ref27277" refString="Imajima, M. (1966) The Syllidae (Polychaetous Annelids) from Japan (V). Syllinae (2). Publications of the Seto Marine Biological Laboratory, 14 (4), 253 - 294. https: // doi. org / 10.5134 / 175446" type="journal article" year="1966">Imajima 1966: 292</bibRefCitation>
</treatmentCitation>
, text-fig. 65.—
<treatmentCitation id="6520109F7E57B0464EFA2BB0B736A71B" author="Hutchings, P. &amp; Murray, A." box="[726,1040,759,784]" page="34" pageId="13" pageNumber="264" year="1984">
<bibRefCitation id="80104B7F7E57B0464EFA2BB0B736A71B" author="Hutchings, P. &amp; Murray, A." box="[726,1040,759,784]" pageId="13" pageNumber="264" pagination="1 - 118" refId="ref27131" refString="Hutchings, P. &amp; Murray, A. (1984) Taxonomy of polychaetes from the Hawkesbury River and the southern estuaries of New South Wales, Australia. Records of the Australian Museum, Supplement 3, 1 - 118. https: // doi. org / 10.3853 / j. 0812 - 7387.3.1984.101" type="journal article" year="1984">Hutchings &amp; Murray 1984: 34</bibRefCitation>
</treatmentCitation>
. Non
<bibRefCitation id="80104B7F7E57B04648602BB0B7E9A71B" author="Krohn, A." box="[1100,1231,759,783]" pageId="13" pageNumber="264" pagination="66 - 76" refId="ref27367" refString="Krohn, A. (1852) Ueber die Erscheinungen bei der Fortpflanzung von Syllis prolifera und Autolytus prolifer. Archiv fur Naturgeschichte, 18, 66 - 76." type="journal article" year="1852">Krohn, 1852</bibRefCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="AC9B65057E57B0464CBB2A7AB7F1A78B" pageId="13" pageNumber="264" type="materials_examined">
<paragraph id="E43E368E7E57B0464CBB2A7AB7F1A78B" blockId="13.[151,1437,829,2008]" pageId="13" pageNumber="264">
<emphasis id="D6F5EA9C7E57B0464CBB2A7AB25AA743" bold="true" box="[151,380,829,855]" pageId="13" pageNumber="264">Material examined</emphasis>
.
<materialsCitation id="54E93CD37E57B0464DA72A7AB7F1A78B" collectingDate="1979-05-11" collectorName="Hutchings &amp; Murray" country="Australia" elevation="40" latitude="-33.44167" location="Colo River" longLatPrecision="772" longitude="150.88333" municipality="Hawkesbury River" pageId="13" pageNumber="264" specimenCount="1" stateProvince="New South Wales" typeStatus="holotype">
<collectingCountry id="9C96761E7E57B0464DA72A7AB10BA743" box="[395,557,829,855]" name="Australia" pageId="13" pageNumber="264">AUSTRALIA</collectingCountry>
.
<collectingRegion id="2645F86C7E57B0464E162A79B06CA743" box="[570,842,829,856]" country="Australia" name="New South Wales" pageId="13" pageNumber="264">NEW SOUTH WALES</collectingRegion>
.
<collectingMunicipality id="045AACF47E57B0464F7B2A7AB708A743" box="[855,1070,829,855]" pageId="13" pageNumber="264">Hawkesbury River</collectingMunicipality>
, just downstream from junction with
<location id="E15E60557E57B0464CFD2A26B269A76F" LSID="urn:lsid:plazi:treatment:6C2887987E57B0444CBB2BC9B15CA5E1:E15E60557E57B0464CFD2A26B269A76F" box="[209,335,865,891]" country="Australia" latitude="-33.44167" longLatPrecision="772" longitude="150.88333" municipality="Hawkesbury River" name="Colo River" pageId="13" pageNumber="264" stateProvince="New South Wales">Colo River</location>
,
<geoCoordinate id="81B550497E57B0464D762A26B2CDA76F" box="[346,491,865,891]" degrees="33" direction="south" minutes="26" orientation="latitude" pageId="13" pageNumber="264" precision="15" seconds="30" value="-33.44167">33º 26’ 30”S</geoCoordinate>
,
<geoCoordinate id="81B550497E57B0464DD42A26B141A768" box="[504,615,865,892]" degrees="150" direction="east" minutes="53" orientation="longitude" pageId="13" pageNumber="264" precision="925" value="150.88333">
150
<superScript id="13F49BC67E57B0464E0E2A26B101A77B" attach="left" box="[546,551,865,879]" fontSize="6" pageId="13" pageNumber="264">º</superScript>
53’E
</geoCoordinate>
, small beach, 40 m off beach, 4 m, fine clean sand sediment,
<collectorName id="497453587E57B04649322A25B67AA768" box="[1310,1372,866,892]" pageId="13" pageNumber="264">Jones</collectorName>
<emphasis id="D6F5EA9C7E57B04649482A25B6BBA76F" box="[1380,1437,865,891]" italics="true" pageId="13" pageNumber="264">et al.</emphasis>
coll.,
<date id="903F104E7E57B0464CFA2AC2B24BA78B" box="[214,365,901,928]" pageId="13" pageNumber="264" value="1979-05-11">
<collectingDate id="807BE9A67E57B0464CFA2AC2B24BA78B" box="[214,365,901,928]" pageId="13" pageNumber="264" value="1979-05-11">11 May 1979</collectingDate>
</date>
, id. as
<taxonomicName id="23814D0D7E57B0464D972AC2B1B2A78B" authorityName="Krohn" authorityYear="1852" box="[443,660,901,927]" class="Polychaeta" family="Syllidae" genus="Typosyllis" kingdom="Animalia" order="Phyllodocida" pageId="13" pageNumber="264" phylum="Annelida" rank="species" species="prolifera">
<emphasis id="D6F5EA9C7E57B0464D972AC2B1B2A78B" box="[443,660,901,927]" italics="true" pageId="13" pageNumber="264">Typosyllis prolifera</emphasis>
</taxonomicName>
by
<collectorName id="497453587E57B0464E922AC2B017A78B" box="[702,817,901,927]" pageId="13" pageNumber="264">Hutchings</collectorName>
&amp;
<collectorName id="497453587E57B0464F782AC1B08FA7B4" box="[852,937,902,928]" pageId="13" pageNumber="264">Murray</collectorName>
, Holotype, AM W.196660.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="AC9B65057E57B0464CEB2AEEB60CA7F3" pageId="13" pageNumber="264" type="diagnosis">
<paragraph id="E43E368E7E57B0464CEB2AEEB60CA7F3" blockId="13.[151,1437,829,2008]" pageId="13" pageNumber="264">
<emphasis id="D6F5EA9C7E57B0464CEB2AEEB264A7D7" bold="true" box="[199,322,937,963]" pageId="13" pageNumber="264">Diagnosis.</emphasis>
Body robust, dark, reddish-brown, with numerous rounded hyaline dermal inclusions. Dorsal cirri slender and short. Compound chaetae with short, bidentate blades. Posterior aciculae slightly acuminate.
</paragraph>
</subSubSection>
<subSubSection id="AC9B65057E57B0464CEB2AB6B032A353" pageId="13" pageNumber="264" type="description">
<paragraph id="E43E368E7E57B0464CEB2AB6B02DA337" blockId="13.[151,1437,829,2008]" pageId="13" pageNumber="264">
<emphasis id="D6F5EA9C7E57B0464CEB2AB6B277A01F" bold="true" box="[199,337,1009,1035]" pageId="13" pageNumber="264">Description</emphasis>
.
<typeStatus id="3B3A882C7E57B0464D4F2AB6B2EDA01F" box="[355,459,1009,1035]" pageId="13" pageNumber="264" type="holotype">Holotype</typeStatus>
<quantity id="23799B6B7E57B0464DFA2AB6B13CA018" box="[470,538,1009,1036]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="13" pageNumber="264" unit="mm" value="8.0">8 mm</quantity>
long, 0.61 mm wide, for 54 chaetigers. Body broad, robust, contracted, very dark and opaque, reddish-brown, especially on anterior segments, except peristomium, with numerous minute rounded hyaline inclusions on dorsum (
<figureCitation id="7CBA2A0B7E57B0464E602D7EB183A047" box="[588,677,1081,1107]" captionStart="FIGURE 6" captionStartId="14.[152,255,1887,1911]" captionTargetBox="[172,1417,394,1842]" captionTargetId="figure-116@14.[151,1436,381,1864]" captionTargetPageId="14" captionText="FIGURE 6. Syllis imajimai n. sp. Holotype.AM W.196660.A, anterior end, dorsal view. B, midbody parapodium. C, falcigers, anterior parapodium. D, falcigers, midbody parapodium. E, falcigers, posterior parapodium. F, dorsal simple chaeta, midbody parapodium. G, dorsal simple chaeta, posterior parapodium. H, ventral simple chaeta. I, aciculae, anterior parapodium. J, aciculae, midbody parapodium. K, acicula, posterior parapodium. Scale. A: 0.2 mm. B: 50 µm. C–K: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.7561401" httpUri="https://zenodo.org/record/7561401/files/figure.png" pageId="13" pageNumber="264">Fig. 6A</figureCitation>
), extending also to ventral side although less numerous. Prostomium pentagonal to oval, with some pigment spots; four eyes in trapezoidal arrangement and two anterior eyespots. Palps robust, similar in length to prostomium. Median antenna inserted on posterior part of prostomium, between posterior eyes, with 20 articles, longer than combined length of prostomium and palps; lateral antennae shorter than median one, with 14–15 articles each, shorter than median antenna. Peristomium shorter than subsequent segments, covering posterior part of prostomium, less pigmented than subsequent segments (
<figureCitation id="7CBA2A0B7E57B0464FEC2DAAB73FA113" box="[960,1049,1261,1287]" captionStart="FIGURE 6" captionStartId="14.[152,255,1887,1911]" captionTargetBox="[172,1417,394,1842]" captionTargetId="figure-116@14.[151,1436,381,1864]" captionTargetPageId="14" captionText="FIGURE 6. Syllis imajimai n. sp. Holotype.AM W.196660.A, anterior end, dorsal view. B, midbody parapodium. C, falcigers, anterior parapodium. D, falcigers, midbody parapodium. E, falcigers, posterior parapodium. F, dorsal simple chaeta, midbody parapodium. G, dorsal simple chaeta, posterior parapodium. H, ventral simple chaeta. I, aciculae, anterior parapodium. J, aciculae, midbody parapodium. K, acicula, posterior parapodium. Scale. A: 0.2 mm. B: 50 µm. C–K: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.7561401" httpUri="https://zenodo.org/record/7561401/files/figure.png" pageId="13" pageNumber="264">Fig. 6A</figureCitation>
). Dorsal tentacular cirri with about 15 articles each; ventral tentacular cirri with about 10 articles. Dorsal cirri shorter than body width, somewhat longer on anterior segments; dorsal cirri of chaetiger 1, distinctly longer, with 19 articles, with well defined cirrophores and articles; in midbody, with about 20–23 articles, usually coiled (
<figureCitation id="7CBA2A0B7E57B0464F4A2C1EB0E7A167" box="[870,961,1369,1395]" captionStart="FIGURE 6" captionStartId="14.[152,255,1887,1911]" captionTargetBox="[172,1417,394,1842]" captionTargetId="figure-116@14.[151,1436,381,1864]" captionTargetPageId="14" captionText="FIGURE 6. Syllis imajimai n. sp. Holotype.AM W.196660.A, anterior end, dorsal view. B, midbody parapodium. C, falcigers, anterior parapodium. D, falcigers, midbody parapodium. E, falcigers, posterior parapodium. F, dorsal simple chaeta, midbody parapodium. G, dorsal simple chaeta, posterior parapodium. H, ventral simple chaeta. I, aciculae, anterior parapodium. J, aciculae, midbody parapodium. K, acicula, posterior parapodium. Scale. A: 0.2 mm. B: 50 µm. C–K: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.7561401" httpUri="https://zenodo.org/record/7561401/files/figure.png" pageId="13" pageNumber="264">Fig. 6A</figureCitation>
). Parapodia conical, distally bilobed (
<figureCitation id="7CBA2A0B7E57B04649432C1EB39AA183" captionStart="FIGURE 6" captionStartId="14.[152,255,1887,1911]" captionTargetBox="[172,1417,394,1842]" captionTargetId="figure-116@14.[151,1436,381,1864]" captionTargetPageId="14" captionText="FIGURE 6. Syllis imajimai n. sp. Holotype.AM W.196660.A, anterior end, dorsal view. B, midbody parapodium. C, falcigers, anterior parapodium. D, falcigers, midbody parapodium. E, falcigers, posterior parapodium. F, dorsal simple chaeta, midbody parapodium. G, dorsal simple chaeta, posterior parapodium. H, ventral simple chaeta. I, aciculae, anterior parapodium. J, aciculae, midbody parapodium. K, acicula, posterior parapodium. Scale. A: 0.2 mm. B: 50 µm. C–K: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.7561401" httpUri="https://zenodo.org/record/7561401/files/figure.png" pageId="13" pageNumber="264">Fig. 6B</figureCitation>
). Ventral cirri digitiform. All compound chaetae heterogomph falcigers with relatively short, markedly bidentate blades and small dorsoventral gradation in length and shape (
<figureCitation id="7CBA2A0B7E57B0464F102CE6B0F5A1AF" box="[828,979,1441,1468]" captionStart="FIGURE 6" captionStartId="14.[152,255,1887,1911]" captionTargetBox="[172,1417,394,1842]" captionTargetId="figure-116@14.[151,1436,381,1864]" captionTargetPageId="14" captionText="FIGURE 6. Syllis imajimai n. sp. Holotype.AM W.196660.A, anterior end, dorsal view. B, midbody parapodium. C, falcigers, anterior parapodium. D, falcigers, midbody parapodium. E, falcigers, posterior parapodium. F, dorsal simple chaeta, midbody parapodium. G, dorsal simple chaeta, posterior parapodium. H, ventral simple chaeta. I, aciculae, anterior parapodium. J, aciculae, midbody parapodium. K, acicula, posterior parapodium. Scale. A: 0.2 mm. B: 50 µm. C–K: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.7561401" httpUri="https://zenodo.org/record/7561401/files/figure.png" pageId="13" pageNumber="264">Fig. 6C, D, E</figureCitation>
). Anterior parapodia each with about 16– 18 chaetae, blades elongated, proximal tooth similar to distal one, and short spines on margin (
<figureCitation id="7CBA2A0B7E57B046488B2C82B7D8A1CB" box="[1191,1278,1477,1503]" captionStart="FIGURE 6" captionStartId="14.[152,255,1887,1911]" captionTargetBox="[172,1417,394,1842]" captionTargetId="figure-116@14.[151,1436,381,1864]" captionTargetPageId="14" captionText="FIGURE 6. Syllis imajimai n. sp. Holotype.AM W.196660.A, anterior end, dorsal view. B, midbody parapodium. C, falcigers, anterior parapodium. D, falcigers, midbody parapodium. E, falcigers, posterior parapodium. F, dorsal simple chaeta, midbody parapodium. G, dorsal simple chaeta, posterior parapodium. H, ventral simple chaeta. I, aciculae, anterior parapodium. J, aciculae, midbody parapodium. K, acicula, posterior parapodium. Scale. A: 0.2 mm. B: 50 µm. C–K: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.7561401" httpUri="https://zenodo.org/record/7561401/files/figure.png" pageId="13" pageNumber="264">Fig. 6C</figureCitation>
), blades about 24–21 μm long; midbody parapodia with 13–14 compound chaetae (
<figureCitation id="7CBA2A0B7E57B0464FB42CAEB0D4A217" box="[920,1010,1513,1539]" captionStart="FIGURE 6" captionStartId="14.[152,255,1887,1911]" captionTargetBox="[172,1417,394,1842]" captionTargetId="figure-116@14.[151,1436,381,1864]" captionTargetPageId="14" captionText="FIGURE 6. Syllis imajimai n. sp. Holotype.AM W.196660.A, anterior end, dorsal view. B, midbody parapodium. C, falcigers, anterior parapodium. D, falcigers, midbody parapodium. E, falcigers, posterior parapodium. F, dorsal simple chaeta, midbody parapodium. G, dorsal simple chaeta, posterior parapodium. H, ventral simple chaeta. I, aciculae, anterior parapodium. J, aciculae, midbody parapodium. K, acicula, posterior parapodium. Scale. A: 0.2 mm. B: 50 µm. C–K: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.7561401" httpUri="https://zenodo.org/record/7561401/files/figure.png" pageId="13" pageNumber="264">Fig. 6D</figureCitation>
), with thick shafts and short bidentate blades, all similar, with short spines on margin, 20–17 μm long; posterior parapodia with eight compound chaetae (
<figureCitation id="7CBA2A0B7E57B0464CB32F76B3DFA25F" box="[159,249,1585,1611]" captionStart="FIGURE 6" captionStartId="14.[152,255,1887,1911]" captionTargetBox="[172,1417,394,1842]" captionTargetId="figure-116@14.[151,1436,381,1864]" captionTargetPageId="14" captionText="FIGURE 6. Syllis imajimai n. sp. Holotype.AM W.196660.A, anterior end, dorsal view. B, midbody parapodium. C, falcigers, anterior parapodium. D, falcigers, midbody parapodium. E, falcigers, posterior parapodium. F, dorsal simple chaeta, midbody parapodium. G, dorsal simple chaeta, posterior parapodium. H, ventral simple chaeta. I, aciculae, anterior parapodium. J, aciculae, midbody parapodium. K, acicula, posterior parapodium. Scale. A: 0.2 mm. B: 50 µm. C–K: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.7561401" httpUri="https://zenodo.org/record/7561401/files/figure.png" pageId="13" pageNumber="264">Fig. 6E</figureCitation>
), similar to those of anterior parapodia but with thicker shafts and more marked proximal teeth, about 17 μm long. Dorsal simple chaeta from midbody parapodia, markedly bidentate, apparently smooth (
<figureCitation id="7CBA2A0B7E57B046493A2F12B6A9A27B" box="[1302,1423,1621,1647]" captionStart="FIGURE 6" captionStartId="14.[152,255,1887,1911]" captionTargetBox="[172,1417,394,1842]" captionTargetId="figure-116@14.[151,1436,381,1864]" captionTargetPageId="14" captionText="FIGURE 6. Syllis imajimai n. sp. Holotype.AM W.196660.A, anterior end, dorsal view. B, midbody parapodium. C, falcigers, anterior parapodium. D, falcigers, midbody parapodium. E, falcigers, posterior parapodium. F, dorsal simple chaeta, midbody parapodium. G, dorsal simple chaeta, posterior parapodium. H, ventral simple chaeta. I, aciculae, anterior parapodium. J, aciculae, midbody parapodium. K, acicula, posterior parapodium. Scale. A: 0.2 mm. B: 50 µm. C–K: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.7561401" httpUri="https://zenodo.org/record/7561401/files/figure.png" pageId="13" pageNumber="264">Fig. 6F, G</figureCitation>
); ventral simple chaetae smooth, bidentate (
<figureCitation id="7CBA2A0B7E57B0464E472F3EB1E2A287" box="[619,708,1657,1683]" captionStart="FIGURE 6" captionStartId="14.[152,255,1887,1911]" captionTargetBox="[172,1417,394,1842]" captionTargetId="figure-116@14.[151,1436,381,1864]" captionTargetPageId="14" captionText="FIGURE 6. Syllis imajimai n. sp. Holotype.AM W.196660.A, anterior end, dorsal view. B, midbody parapodium. C, falcigers, anterior parapodium. D, falcigers, midbody parapodium. E, falcigers, posterior parapodium. F, dorsal simple chaeta, midbody parapodium. G, dorsal simple chaeta, posterior parapodium. H, ventral simple chaeta. I, aciculae, anterior parapodium. J, aciculae, midbody parapodium. K, acicula, posterior parapodium. Scale. A: 0.2 mm. B: 50 µm. C–K: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.7561401" httpUri="https://zenodo.org/record/7561401/files/figure.png" pageId="13" pageNumber="264">Fig. 6H</figureCitation>
), on far posterior segments. Anterior parapodia with three slender aciculae aciculae (
<figureCitation id="7CBA2A0B7E57B0464D4B2FDAB293A2A3" box="[359,437,1693,1719]" captionStart="FIGURE 6" captionStartId="14.[152,255,1887,1911]" captionTargetBox="[172,1417,394,1842]" captionTargetId="figure-116@14.[151,1436,381,1864]" captionTargetPageId="14" captionText="FIGURE 6. Syllis imajimai n. sp. Holotype.AM W.196660.A, anterior end, dorsal view. B, midbody parapodium. C, falcigers, anterior parapodium. D, falcigers, midbody parapodium. E, falcigers, posterior parapodium. F, dorsal simple chaeta, midbody parapodium. G, dorsal simple chaeta, posterior parapodium. H, ventral simple chaeta. I, aciculae, anterior parapodium. J, aciculae, midbody parapodium. K, acicula, posterior parapodium. Scale. A: 0.2 mm. B: 50 µm. C–K: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.7561401" httpUri="https://zenodo.org/record/7561401/files/figure.png" pageId="13" pageNumber="264">Fig. 6I</figureCitation>
), two at midbody (
<figureCitation id="7CBA2A0B7E57B0464EA22FDAB1F8A2A3" box="[654,734,1693,1719]" captionStart="FIGURE 6" captionStartId="14.[152,255,1887,1911]" captionTargetBox="[172,1417,394,1842]" captionTargetId="figure-116@14.[151,1436,381,1864]" captionTargetPageId="14" captionText="FIGURE 6. Syllis imajimai n. sp. Holotype.AM W.196660.A, anterior end, dorsal view. B, midbody parapodium. C, falcigers, anterior parapodium. D, falcigers, midbody parapodium. E, falcigers, posterior parapodium. F, dorsal simple chaeta, midbody parapodium. G, dorsal simple chaeta, posterior parapodium. H, ventral simple chaeta. I, aciculae, anterior parapodium. J, aciculae, midbody parapodium. K, acicula, posterior parapodium. Scale. A: 0.2 mm. B: 50 µm. C–K: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.7561401" httpUri="https://zenodo.org/record/7561401/files/figure.png" pageId="13" pageNumber="264">Fig. 6J</figureCitation>
), and single, slightly acuminate acicula in posterior parapodia (
<figureCitation id="7CBA2A0B7E57B0464CB32F86B3D8A2CF" box="[159,254,1729,1755]" captionStart="FIGURE 6" captionStartId="14.[152,255,1887,1911]" captionTargetBox="[172,1417,394,1842]" captionTargetId="figure-116@14.[151,1436,381,1864]" captionTargetPageId="14" captionText="FIGURE 6. Syllis imajimai n. sp. Holotype.AM W.196660.A, anterior end, dorsal view. B, midbody parapodium. C, falcigers, anterior parapodium. D, falcigers, midbody parapodium. E, falcigers, posterior parapodium. F, dorsal simple chaeta, midbody parapodium. G, dorsal simple chaeta, posterior parapodium. H, ventral simple chaeta. I, aciculae, anterior parapodium. J, aciculae, midbody parapodium. K, acicula, posterior parapodium. Scale. A: 0.2 mm. B: 50 µm. C–K: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.7561401" httpUri="https://zenodo.org/record/7561401/files/figure.png" pageId="13" pageNumber="264">Fig. 6K</figureCitation>
). Pharynx long, extending through about six segments; pharyngeal tooth not seen because of opacity of anterior segments (
<figureCitation id="7CBA2A0B7E57B0464DBA2FA2B2D2A2EB" box="[406,500,1765,1791]" captionStart="FIGURE 6" captionStartId="14.[152,255,1887,1911]" captionTargetBox="[172,1417,394,1842]" captionTargetId="figure-116@14.[151,1436,381,1864]" captionTargetPageId="14" captionText="FIGURE 6. Syllis imajimai n. sp. Holotype.AM W.196660.A, anterior end, dorsal view. B, midbody parapodium. C, falcigers, anterior parapodium. D, falcigers, midbody parapodium. E, falcigers, posterior parapodium. F, dorsal simple chaeta, midbody parapodium. G, dorsal simple chaeta, posterior parapodium. H, ventral simple chaeta. I, aciculae, anterior parapodium. J, aciculae, midbody parapodium. K, acicula, posterior parapodium. Scale. A: 0.2 mm. B: 50 µm. C–K: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.7561401" httpUri="https://zenodo.org/record/7561401/files/figure.png" pageId="13" pageNumber="264">Fig. 6A</figureCitation>
). Proventricle, through about six segments, with 28 muscle cell rows. Pygidium triangular, with two articulated anal cirri and one stylus.
</paragraph>
<paragraph id="E43E368E7E57B0464CEB2E6AB032A353" blockId="13.[151,1437,829,2008]" box="[199,788,1837,1864]" pageId="13" pageNumber="264">
The
<typeStatus id="3B3A882C7E57B0464CD62E6AB27AA353" box="[250,348,1837,1863]" pageId="13" pageNumber="264" type="holotype">holotype</typeStatus>
is developing a sexual dicerous stolon.
</paragraph>
</subSubSection>
<subSubSection id="AC9B65057E57B0444CEB2E16B71AA571" lastPageId="15" lastPageNumber="266" pageId="13" pageNumber="264" type="discussion">
<paragraph id="E43E368E7E57B0454CEB2E16B218A571" blockId="13.[151,1437,829,2008]" lastBlockId="14.[151,1437,151,357]" lastPageId="14" lastPageNumber="265" pageId="13" pageNumber="264">
<emphasis id="D6F5EA9C7E57B0464CEB2E16B212A37F" bold="true" box="[199,308,1873,1899]" pageId="13" pageNumber="264">Remarks</emphasis>
. This species is characterised by its colour pattern, reddish-brown with small hyaline, refringent dermal inclusions, especially marked on anterior segments, dorsal cirri similar in length or shorter than body width, and compound chaetae with relatively short falcigers, with strong, acute proximal and distal teeth, especially from midbody onwards. The Australian examined specimen agrees very well with the Imajima’ description of
<taxonomicName id="23814D0D7E57B04549012EFAB3DCA4A5" authorityName="Krohn" authorityYear="1852" class="Polychaeta" family="Syllidae" genus="Typosyllis" kingdom="Animalia" lastPageId="14" lastPageNumber="265" order="Phyllodocida" pageId="13" pageNumber="264" phylum="Annelida" rank="species" species="prolifera">
<emphasis id="D6F5EA9C7E57B04549012EFAB3DCA4A5" italics="true" lastPageId="14" lastPageNumber="265" pageId="13" pageNumber="264">Typosyllis prolifera</emphasis>
</taxonomicName>
, but differs in many aspects with the descriptions of this species from European coasts, a non pigmented species, with aciculae ending in rounded, hollow tips, and pharyngeal tooth located just behind the anterior margin of pharynx (see
<bibRefCitation id="80104B7F7E54B0454D622998B2FBA4ED" author="Fauvel, P." box="[334,477,223,249]" pageId="14" pageNumber="265" refId="ref25833" refString="Fauvel, P. (1923) Polychetes Errantes. En: Faune de France. Vol. 5. Le Chevalier, Paris, 486 pp." type="book" year="1923">Fauvel 1923</bibRefCitation>
; Campoy 1982, San Martín 2003).
<bibRefCitation id="80104B7F7E54B0454F402998B73DA4ED" author="Imajima, M." box="[876,1051,223,249]" pageId="14" pageNumber="265" pagination="253 - 294" refId="ref27277" refString="Imajima, M. (1966) The Syllidae (Polychaetous Annelids) from Japan (V). Syllinae (2). Publications of the Seto Marine Biological Laboratory, 14 (4), 253 - 294. https: // doi. org / 10.5134 / 175446" type="journal article" year="1966">Imajima (1966)</bibRefCitation>
states that the pharyngeal tooth is located on the anterior margin. Also, the dorsal simple chaetae are very characteristic, markedly bidentate, with the proximal teeth prominent on his material which is why we have recognized his material as our new species
<taxonomicName id="23814D0D7E54B045494C2860B3DDA570" authority="Martín &amp; Lucas &amp; Hutchings, 2023" authorityName="Martín &amp; Lucas &amp; Hutchings" authorityYear="2023" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="14" pageNumber="265" phylum="Annelida" rank="species" species="imajimai" status="sp. nov.">
<emphasis id="D6F5EA9C7E54B045494C2860B3DDA570" italics="true" pageId="14" pageNumber="265">Syllis imajimai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CDC657E77E54B0454D2E280CB218A571" box="[258,318,331,357]" pageId="14" pageNumber="265" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
<caption id="B0FE66067E54B0454CB42E18B7C3A3F0" ID-DOI="http://doi.org/10.5281/zenodo.7561401" ID-Zenodo-Dep="7561401" httpUri="https://zenodo.org/record/7561401/files/figure.png" pageId="14" pageNumber="265" startId="14.[152,255,1887,1911]" targetBox="[172,1417,394,1842]" targetPageId="14">
<paragraph id="E43E368E7E54B0454CB42E18B7C3A3F0" blockId="14.[151,1437,1887,2020]" pageId="14" pageNumber="265">
<emphasis id="D6F5EA9C7E54B0454CB42E18B231A363" bold="true" box="[152,279,1887,1911]" pageId="14" pageNumber="265">FIGURE 6.</emphasis>
<taxonomicName id="23814D0D7E54B0454D312E18B292A363" authority="Martín &amp; Lucas &amp; Hutchings, 2023" authorityName="Martín &amp; Lucas &amp; Hutchings" authorityYear="2023" box="[285,436,1887,1911]" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="14" pageNumber="265" phylum="Annelida" rank="species" species="imajimai" status="sp. nov.">
<emphasis id="D6F5EA9C7E54B0454D312E18B292A363" box="[285,436,1887,1911]" italics="true" pageId="14" pageNumber="265">Syllis imajimai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CDC657E77E54B0454D962E27B2D6A363" box="[442,496,1888,1911]" pageId="14" pageNumber="265" rank="species">n. sp.</taxonomicNameLabel>
Holotype. AM W.196660. A, anterior end, dorsal view. B, midbody parapodium. C, falcigers, anterior parapodium. D, falcigers, midbody parapodium. E, falcigers, posterior parapodium. F, dorsal simple chaeta, midbody parapodium. G, dorsal simple chaeta, posterior parapodium. H, ventral simple chaeta. I, aciculae, anterior parapodium. J, aciculae, midbody parapodium. K, acicula, posterior parapodium. Scale. A: 0.2 mm. B: 50 µm. C–K: 20 µm.
</paragraph>
</caption>
<paragraph id="E43E368E7E55B0444CEB29D0B71AA571" blockId="15.[151,1437,151,501]" pageId="15" pageNumber="266">
Other species with hyaline, refringent dermal inclusion are
<taxonomicName id="23814D0D7E55B0444F7A29D0B6B1A4A5" authority="(Hartmann-Schroder, 1979)" authorityName="Martín &amp; Lucas &amp; Hutchings" authorityYear="2023" baseAuthorityName="Hartmann-Schroder" baseAuthorityYear="1979" box="[854,1431,151,177]" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="15" pageNumber="266" phylum="Annelida" rank="species" species="pharobroomensis" status="comb. nov.">
<emphasis id="D6F5EA9C7E55B0444F7A29D0B77DA4A5" box="[854,1115,151,177]" italics="true" pageId="15" pageNumber="266">Syllis pharobroomensis</emphasis>
(
<bibRefCitation id="80104B7F7E55B044484429D0B6B6A4A5" author="Hartmann-Schroder, G." box="[1128,1424,151,177]" pageId="15" pageNumber="266" pagination="75 - 218" refId="ref26423" refString="Hartmann-Schroder, G. (1979) Teil 2. Die Polychaeten der tropischen Nordwestk ¸ ste Australiens (zwischen Port Samson in Norden und Port Hedland in S ¸ den). Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut, 76, 75 - 218." type="journal article" year="1979">Hartmann-Schröder, 1979</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="CDC657E77E55B0444CBB29FCB3DEA4C1" box="[151,248,187,213]" pageId="15" pageNumber="266" rank="species">n. comb.</taxonomicNameLabel>
(see this paper),
<taxonomicName id="23814D0D7E55B0444D9F29FCB7ABA4C1" authority="Salcedo-Oropeza, San Martin &amp; Solis-Weiss, 2012" authorityName="Salcedo-Oropeza, San Martin &amp; Solis-Weiss" authorityYear="2012" box="[435,1165,187,213]" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="15" pageNumber="266" phylum="Annelida" rank="species" species="deleoini">
<emphasis id="D6F5EA9C7E55B0444D9F29FCB176A4C1" box="[435,592,187,213]" italics="true" pageId="15" pageNumber="266">Syllis deleoini</emphasis>
<bibRefCitation id="80104B7F7E55B0444E7A29FCB7ABA4C1" author="Salcedo-Oropeza, D. L. &amp; San Martin, G. &amp; Solis-Weiss, V." box="[598,1165,187,213]" pageId="15" pageNumber="266" pagination="47 - 62" refId="ref28111" refString="Salcedo-Oropeza, D. L., San Martin, G. &amp; Solis-Weiss, V. (2012) The genus Syllis (Polychaeta: Syllidae: Syllinae) in the southern Mexican Pacific, with the description of two new species and three new records. Zootaxa, 3263 (1), 47 - 62. https: // doi. org / 10.11646 / zootaxa. 3263.1.2" type="journal article" year="2012">Salcedo-Oropeza, San Martín &amp; Solís-Weiss, 2012</bibRefCitation>
</taxonomicName>
, from southern Mexican Pacific, and
<taxonomicName id="23814D0D7E55B0444D052998B16CA4ED" authority="Cinar, 2005" authorityName="Cinar" authorityYear="2005" box="[297,586,222,249]" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="15" pageNumber="266" phylum="Annelida" rank="species" species="ergeni">
<emphasis id="D6F5EA9C7E55B0444D052998B291A4EC" box="[297,439,223,249]" italics="true" pageId="15" pageNumber="266">Syllis ergeni</emphasis>
Çinar, 2005
</taxonomicName>
, from the Aegean Sea,
<collectingCountry id="9C96761E7E55B0444F712998B089A4ED" box="[861,943,223,249]" name="Turkey" pageId="15" pageNumber="266">Turkey</collectingCountry>
.
<taxonomicName id="23814D0D7E55B0444F912998B76DA4EC" authorityName="Cinar" authorityYear="2005" box="[957,1099,223,249]" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="15" pageNumber="266" phylum="Annelida" rank="species" species="ergeni">
<emphasis id="D6F5EA9C7E55B0444F912998B76DA4EC" box="[957,1099,223,249]" italics="true" pageId="15" pageNumber="266">Syllis ergeni</emphasis>
</taxonomicName>
is also a strongly pigmented species, the compound chaetae and dorsal simple chaetae are not as markedly bidentate as in
<taxonomicName id="23814D0D7E55B04448B62844B63FA508" authority="Martín &amp; Lucas &amp; Hutchings, 2023" authorityName="Martín &amp; Lucas &amp; Hutchings" authorityYear="2023" box="[1178,1305,259,285]" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="15" pageNumber="266" phylum="Annelida" rank="species" species="imajimai" status="sp. nov.">
<emphasis id="D6F5EA9C7E55B04448B62844B63FA508" box="[1178,1305,259,285]" italics="true" pageId="15" pageNumber="266">S. imajimai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CDC657E77E55B04449332844B67CA509" box="[1311,1370,259,285]" pageId="15" pageNumber="266" rank="species">n. sp.</taxonomicNameLabel>
<taxonomicName id="23814D0D7E55B044494C2844B3CFA555" authorityName="Salcedo Oropeza, San Martin &amp; Solis-Weiss" authorityYear="2012" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="15" pageNumber="266" phylum="Annelida" rank="species" species="deleoni">
<emphasis id="D6F5EA9C7E55B044494C2844B3CFA555" italics="true" pageId="15" pageNumber="266">Syllis deleoni</emphasis>
</taxonomicName>
lacks colour pattern, the compound chaetae are similar, with relatively short bidentate blades, but both teeth are short, somewhat broader and close to each other (
<bibRefCitation id="80104B7F7E55B0444EC6280CB716A571" author="Salcedo-Oropeza, D. L. &amp; San Martin, G. &amp; Solis-Weiss, V." box="[746,1072,331,357]" pageId="15" pageNumber="266" pagination="47 - 62" refId="ref28111" refString="Salcedo-Oropeza, D. L., San Martin, G. &amp; Solis-Weiss, V. (2012) The genus Syllis (Polychaeta: Syllidae: Syllinae) in the southern Mexican Pacific, with the description of two new species and three new records. Zootaxa, 3263 (1), 47 - 62. https: // doi. org / 10.11646 / zootaxa. 3263.1.2" type="journal article" year="2012">
Salcedo-Oropeza
<emphasis id="D6F5EA9C7E55B0444F98280CB0C3A571" box="[948,997,331,357]" italics="true" pageId="15" pageNumber="266">et al</emphasis>
. 2012
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="AC9B65057E55B0444CEB2828B083A5C5" pageId="15" pageNumber="266" type="etymology">
<paragraph id="E43E368E7E55B0444CEB2828B1DEA5B9" blockId="15.[151,1437,151,501]" pageId="15" pageNumber="266">
<emphasis id="D6F5EA9C7E55B0444CEB2828B26EA59D" bold="true" box="[199,328,367,393]" pageId="15" pageNumber="266">Etymology</emphasis>
. The species is named after Dr Minoru Imajima, a Japanese polychaetologist, who made important contributions to our knowledge of the family
<taxonomicName id="23814D0D7E55B0444EB928D4B1D2A5B9" authorityName="Grube" authorityYear="1850" box="[661,756,403,429]" class="Polychaeta" family="Syllidae" kingdom="Animalia" order="Phyllodocida" pageId="15" pageNumber="266" phylum="Annelida" rank="family">Syllidae</taxonomicName>
.
</paragraph>
<paragraph id="E43E368E7E55B0444CEB28F1B083A5C5" blockId="15.[151,1437,151,501]" box="[199,933,438,465]" pageId="15" pageNumber="266">
<emphasis id="D6F5EA9C7E55B0444CEB28F1B205A5C4" bold="true" box="[199,291,438,464]" pageId="15" pageNumber="266">Habitat</emphasis>
. Fine clean sand sediment. Intertidal and shallow depths.
</paragraph>
</subSubSection>
<subSubSection id="AC9B65057E55B0444CEB289DB15CA5E1" box="[199,634,474,501]" pageId="15" pageNumber="266" type="distribution">
<paragraph id="E43E368E7E55B0444CEB289DB15CA5E1" blockId="15.[151,1437,151,501]" box="[199,634,474,501]" pageId="15" pageNumber="266">
<emphasis id="D6F5EA9C7E55B0444CEB289DB27FA5E0" bold="true" box="[199,345,474,500]" pageId="15" pageNumber="266">Distribution</emphasis>
.
<collectingCountry id="9C96761E7E55B0444D4A289CB2EBA5E1" box="[358,461,475,501]" name="Australia" pageId="15" pageNumber="266">Australia</collectingCountry>
(NSW).
<collectingCountry id="9C96761E7E55B0444E1E289CB153A5E1" box="[562,629,475,501]" name="Japan" pageId="15" pageNumber="266">Japan</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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