142 lines
17 KiB
XML
142 lines
17 KiB
XML
<document id="9B88C04AE760CEB6A8720CA125C1E765" ID-DOI="10.11646/zootaxa.2462.1.1" ID-ISSN="11755334" ID-Zenodo-Dep="5312710" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629676798216" checkinUser="felipe" docAuthor="Hyman, Isabel T. & Ponder, Winston F." docDate="2010" docId="6413F378FF856A19F28B7336FB0DFE29" docLanguage="en" docName="zt02462p148.pdf" docOrigin="Zootaxa 2462 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2462.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Helicarionoidea Bourguignat 1877" docType="treatment" docVersion="4" lastPageNumber="20" masterDocId="982A8B00FF976A0AF21C766BFFEBFF8B" masterDocTitle="A morphological phylogenetic analysis and generic revision of Australian Helicarionidae (Gastropoda: Pulmonata: Stylommatophora), and an assessment of the relationships of the family 2462" masterLastPageNumber="148" masterPageNumber="1" pageNumber="19" updateTime="1699094437759" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="4C71F8C49A69C5949395AA217AFB37BB">A morphological phylogenetic analysis and generic revision of Australian Helicarionidae (Gastropoda: Pulmonata: Stylommatophora), and an assessment of the relationships of the family 2462</mods:title>
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<mods:namePart id="26B5A8DC8138101A09C5A45987902324">Hyman, Isabel T.</mods:namePart>
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<mods:namePart id="151FE0D6DA7065B4FF30252BFD2CB9AB">Ponder, Winston F.</mods:namePart>
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<mods:title id="0DF4E1E1E78060DDC363267F86EFE19E">Zootaxa</mods:title>
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<mods:part id="8EC73BF5E92F51974C7A0E24E9259651">
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<mods:date id="8893DFA7BE94660E8A8EFA6FFDF24C31">2010</mods:date>
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<mods:number id="145AA75066D4F0832D493E5F5A054BE3">2010-05-14</mods:number>
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<mods:number id="471315B28172959B97FF02F7F74B5333">2462</mods:number>
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<mods:url id="2B14634FB980BD17F6AC8B3E2B66E16F">https://biotaxa.org/Zootaxa/article/view/zootaxa.2462.1.1</mods:url>
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<mods:classification id="DC4722D004CE119A21F11462AB710A67">journal article</mods:classification>
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<mods:identifier id="EE2A8296D22B7AFBB4F0ED120E5C81A3" type="DOI">10.11646/zootaxa.2462.1.1</mods:identifier>
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<mods:identifier id="D5C7E65E86D0106050042F1E1EF73412" type="ISSN">11755334</mods:identifier>
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<treatment id="6413F378FF856A19F28B7336FB0DFE29" ID-DOI="http://doi.org/10.5281/zenodo.5320974" ID-Zenodo-Dep="5320974" LSID="urn:lsid:plazi:treatment:6413F378FF856A19F28B7336FB0DFE29" httpUri="http://treatment.plazi.org/id/6413F378FF856A19F28B7336FB0DFE29" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
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<subSubSection id="A4A011E5FF856A18F28B7336FD06FAFC" box="[151,749,1373,1399]" pageId="18" pageNumber="19" type="nomenclature">
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<paragraph id="EC05426EFF856A18F28B7336FD06FAFC" blockId="18.[151,749,1373,1399]" box="[151,749,1373,1399]" pageId="18" pageNumber="19">
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<heading id="B74DF502FF856A18F28B7336FD06FAFC" bold="true" box="[151,749,1373,1399]" fontSize="11" level="1" pageId="18" pageNumber="19" reason="1">
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<emphasis id="DECE9E7CFF856A18F28B7336FD06FAFC" bold="true" box="[151,749,1373,1399]" pageId="18" pageNumber="19">
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Superfamily
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<taxonomicName id="2BBA39EDFF856A18F3257336FD06FAFC" ID-CoL="7NGNR" authority="Bourguignat, 1877" authorityName="Bourguignat" authorityYear="1877" box="[313,749,1373,1399]" class="Gastropoda" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="19" phylum="Mollusca" rank="superFamily" superFamily="Helicarionoidea">
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Helicarionoidea
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<bibRefCitation id="882B3F9FFF856A18F01B7336FD06FAFC" author="Bourguignat, J. - R." box="[519,749,1373,1399]" pageId="18" pageNumber="19" pagination="49 - 101" refId="ref73235" refString="Bourguignat, J. - R. (1877) Description de deux nouveaux genres algeriens, suivi d'une classification des familles et des genres de Mollusques terrestres et fluviatiles du systeme europeen. Bulletin de la Societe des Sciences Physiques et Naturelles de Toulouse, 3, 49 - 101." type="journal article" year="1877">Bourguignat, 1877</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="A4A011E5FF856A18F28B73C9FDA8F879" pageId="18" pageNumber="19" type="discussion">
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<paragraph id="EC05426EFF856A18F28B73C9FE3CF95F" blockId="18.[151,1436,1442,2034]" pageId="18" pageNumber="19">
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<emphasis id="DECE9E7CFF856A18F28B73C9FEFBFA37" bold="true" box="[151,272,1442,1468]" pageId="18" pageNumber="19">Remarks.</emphasis>
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<taxonomicName id="2BBA39EDFF856A18F30573C9FE3FFA37" authorityName="Bourguignat" authorityYear="1877" box="[281,468,1442,1468]" class="Gastropoda" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="19" phylum="Mollusca" rank="superFamily" superFamily="Helicarionoidea">Helicarionoidea</taxonomicName>
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was defined by
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<bibRefCitation id="882B3F9FFF856A18F08773C9FCB4FA37" author="Hausdorf, B." box="[667,863,1442,1468]" pageId="18" pageNumber="19" pagination="35 - 66" refId="ref74906" refString="Hausdorf, B. (1998) Phylogeny of the Limacoidea sensu lato (Gastropoda: Stylommatophora). Journal of Molluscan Studies, 64, 35 - 66." type="journal article" year="1998">Hausdorf (1998)</bibRefCitation>
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as containing taxa with an epiphallic caecum, a flagellum with an axial filament and mantle lobes (although mantle lobes were also present in several other groups, including
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<taxonomicName id="2BBA39EDFF856A18F3737399FDECF987" box="[367,519,1522,1548]" class="Gastropoda" family="Euconulidae" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="19" phylum="Mollusca" rank="family">Euconulidae</taxonomicName>
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). The superfamily was made up of three groups,
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<taxonomicName id="2BBA39EDFF856A18F6487399FB14F987" authorityName="Godwin-Austen" authorityYear="1883" box="[1108,1279,1522,1548]" class="Gastropoda" family="Ariophantidae" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="19" phylum="Mollusca" rank="family">Ariophantidae</taxonomicName>
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,
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<taxonomicName id="2BBA39EDFF856A18F7117399FA77F987" box="[1293,1436,1522,1548]" class="Gastropoda" family="Urocyclidae" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="19" phylum="Mollusca" rank="family">Urocyclidae</taxonomicName>
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and
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<taxonomicName id="2BBA39EDFF856A18F2D07071FE97F9BF" authorityName="Bourguignat" authorityYear="1877" box="[204,380,1562,1588]" class="Gastropoda" family="Helicarionidae" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="19" phylum="Mollusca" rank="family">Helicarionidae</taxonomicName>
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.
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<bibRefCitation id="882B3F9FFF856A18F3977071FDBAF9BF" author="Hausdorf, B." box="[395,593,1562,1588]" pageId="18" pageNumber="19" pagination="35 - 66" refId="ref74906" refString="Hausdorf, B. (1998) Phylogeny of the Limacoidea sensu lato (Gastropoda: Stylommatophora). Journal of Molluscan Studies, 64, 35 - 66." type="journal article" year="1998">Hausdorf (1998)</bibRefCitation>
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showed that the three family-group taxa can be divided by only one character: in
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<taxonomicName id="2BBA39EDFF856A18F32E7029FE35F9D7" authorityName="Bourguignat" authorityYear="1877" box="[306,478,1602,1628]" class="Gastropoda" family="Helicarionidae" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="19" phylum="Mollusca" rank="family">Helicarionidae</taxonomicName>
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the penial sheath is open at the distal end (as in other
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<taxonomicName id="2BBA39EDFF856A18F6797029FB04F9D7" authorityName="Lamarck" authorityYear="1801" box="[1125,1263,1602,1628]" pageId="18" pageNumber="8" rank="superFamily" superFamily="Limacoidea">Limacoidea</taxonomicName>
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<emphasis id="DECE9E7CFF856A18F6E47028FAC9F9D7" box="[1272,1314,1603,1628]" italics="true" pageId="18" pageNumber="19">s. l.</emphasis>
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), while in
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<taxonomicName id="2BBA39EDFF856A18F28B7001FED4F90F" authorityName="Godwin-Austen" authorityYear="1883" box="[151,319,1642,1668]" class="Gastropoda" family="Ariophantidae" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="19" phylum="Mollusca" rank="family">Ariophantidae</taxonomicName>
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and
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<taxonomicName id="2BBA39EDFF856A18F3607001FDE0F90F" box="[380,523,1642,1668]" class="Gastropoda" family="Urocyclidae" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="19" phylum="Mollusca" rank="family">Urocyclidae</taxonomicName>
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the distal end of the penial sheath is fused to the epiphallus. Based on this,
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<bibRefCitation id="882B3F9FFF856A18F28B70F9FEB0F927" author="Hausdorf, B." box="[151,347,1682,1708]" pageId="18" pageNumber="19" pagination="35 - 66" refId="ref74906" refString="Hausdorf, B. (1998) Phylogeny of the Limacoidea sensu lato (Gastropoda: Stylommatophora). Journal of Molluscan Studies, 64, 35 - 66." type="journal article" year="1998">Hausdorf (1998)</bibRefCitation>
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moved several taxa from
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<taxonomicName id="2BBA39EDFF856A18F08770F9FCA8F927" authorityName="Godwin-Austen" authorityYear="1883" box="[667,835,1682,1708]" class="Gastropoda" family="Ariophantidae" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="19" phylum="Mollusca" rank="family">Ariophantidae</taxonomicName>
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to
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<taxonomicName id="2BBA39EDFF856A18F17270F9FBF1F927" authorityName="Bourguignat" authorityYear="1877" box="[878,1050,1682,1708]" class="Gastropoda" family="Helicarionidae" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="19" phylum="Mollusca" rank="family">Helicarionidae</taxonomicName>
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(Durgellinae, Girasiinae,
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<emphasis id="DECE9E7CFF856A18F74870F8FA7EF927" box="[1364,1429,1683,1708]" italics="true" pageId="18" pageNumber="19">Xesta</emphasis>
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,
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<taxonomicName id="2BBA39EDFF856A18F28B70D0FF0CF95F" authorityName="Albers" authorityYear="1860" box="[151,231,1723,1748]" class="Gastropoda" family="Ariophantidae" genus="Sesara" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="19" phylum="Mollusca" rank="genus">
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<emphasis id="DECE9E7CFF856A18F28B70D0FF0CF95F" box="[151,231,1723,1748]" italics="true" pageId="18" pageNumber="19">Sesara</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="2BBA39EDFF856A18F2E970D0FEAAF95F" authorityName="Godwin-Austen" authorityYear="1907" box="[245,321,1723,1748]" class="Gastropoda" family="Ariophantidae" genus="Sarika" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="19" phylum="Mollusca" rank="genus">
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<emphasis id="DECE9E7CFF856A18F2E970D0FEAAF95F" box="[245,321,1723,1748]" italics="true" pageId="18" pageNumber="19">Sarika</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="2BBA39EDFF856A18F36770D0FE2DF95F" authorityName="Solem" authorityYear="1966" box="[379,454,1723,1748]" class="Gastropoda" family="Helicarionidae" genus="Teraia" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="19" phylum="Mollusca" rank="genus">
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<emphasis id="DECE9E7CFF856A18F36770D0FE2DF95F" box="[379,454,1723,1748]" italics="true" pageId="18" pageNumber="19">Teraia</emphasis>
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</taxonomicName>
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).
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</paragraph>
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<paragraph id="EC05426EFF856A18F2DA7089FDA8F879" blockId="18.[151,1436,1442,2034]" pageId="18" pageNumber="19">
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Based on the results of our analyses, we define
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<taxonomicName id="2BBA39EDFF856A18F1157089FC2FF977" authorityName="Bourguignat" authorityYear="1877" box="[777,964,1762,1788]" class="Gastropoda" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="19" phylum="Mollusca" rank="superFamily" superFamily="Helicarionoidea">Helicarionoidea</taxonomicName>
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as follows: Shell complete or reduced. Mantle usually with visible minor blood vessels. Stimulator absent or present; if present, stimulator section A
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<subScript id="703E402BFF856A18F7887170FA77F8A2" attach="left" box="[1428,1436,1819,1833]" fontSize="6" pageId="18" pageNumber="19">5</subScript>
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absent or present; sections A
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<subScript id="703E402BFF856A18F3F6712EFE19F8D8" attach="left" box="[490,498,1861,1875]" fontSize="6" pageId="18" pageNumber="19">3</subScript>
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and A
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<subScript id="703E402BFF856A18F05E712EFDA1F8D8" attach="left" box="[578,586,1861,1875]" fontSize="6" pageId="18" pageNumber="19">4</subScript>
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absent; section A
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<subScript id="703E402BFF856A18F101712EFCCEF8D8" attach="left" box="[797,805,1861,1875]" fontSize="6" pageId="18" pageNumber="19">2</subScript>
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usually with papilla (see appendix for explanation of stimulator section abbreviations), dart absent or present. Penial sheath open (
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<taxonomicName id="2BBA39EDFF856A18F63C710BFB3BF8F1" authorityName="Bourguignat" authorityYear="1877" box="[1056,1232,1888,1914]" class="Gastropoda" family="Helicarionidae" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="19" phylum="Mollusca" rank="family">Helicarionidae</taxonomicName>
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) or closed (
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<taxonomicName id="2BBA39EDFF856A18F746710BFEE7F829" authorityName="Godwin-Austen" authorityYear="1883" class="Gastropoda" family="Ariophantidae" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="19" phylum="Mollusca" rank="family">Ariophantidae</taxonomicName>
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,
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<taxonomicName id="2BBA39EDFF856A18F30471E3FE42F829" box="[280,425,1928,1954]" class="Gastropoda" family="Urocyclidae" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="19" phylum="Mollusca" rank="family">Urocyclidae</taxonomicName>
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) at distal end. Epiphallic retractor caecum and epiphallic flagellum absent or present. Bursa copulatrix inserted on vagina or if vagina absent, at junction of penis and free oviduct. Spermatophore soft capsule with long firm tail-pipe.
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</paragraph>
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</subSubSection>
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<subSubSection id="A4A011E5FF846A19F2DA76FCFB0DFE29" pageId="19" pageNumber="20" type="description">
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<paragraph id="EC05426EFF846A19F2DA76FCFB0DFE29" blockId="19.[151,1436,151,418]" pageId="19" pageNumber="20">
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<emphasis id="DECE9E7CFF846A19F2DA76FCFEB5FF3A" bold="true" box="[198,350,151,177]" pageId="19" pageNumber="20">Description.</emphasis>
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Shell present, complete or reduced. Shell shape variable, sculpture, if present, typically weak; mantle laps and lobes variable. Sole of foot tripartite; caudal apparatus present, formed from curled up sole. Kidney unilobed; mantle gland absent. Genital system oviparous; oviduct glandular. Bursa copulatrix inserted on vagina or if vagina absent, at junction of free oviduct and penis. Stimulator absent or present. Penial sheath present, sometimes not covering entire penis; open (
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<taxonomicName id="2BBA39EDFF846A19F1D77753FB90FED9" authorityName="Bourguignat" authorityYear="1877" box="[971,1147,312,338]" class="Gastropoda" family="Helicarionidae" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="20" phylum="Mollusca" rank="family">Helicarionidae</taxonomicName>
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) or fused to epiphallus (
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<taxonomicName id="2BBA39EDFF846A19F283770BFEA7FEF1" authorityName="Godwin-Austen" authorityYear="1883" box="[159,332,352,378]" class="Gastropoda" family="Ariophantidae" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="20" phylum="Mollusca" rank="family">Ariophantidae</taxonomicName>
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,
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<taxonomicName id="2BBA39EDFF846A19F34B770BFE03FEF1" box="[343,488,352,378]" class="Gastropoda" family="Urocyclidae" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="20" phylum="Mollusca" rank="family">Urocyclidae</taxonomicName>
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) at distal end. Epiphallic retractor caecum absent or present; epiphallic flagellum absent or present. Spermatophore soft capsule with long, firm tail-pipe with various sculpture.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |