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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1027.54888" ID-GBIF-Dataset="a108311a-65d1-431d-b108-40e585c2646b" ID-PMC="PMC8035127" ID-Pensoft-Pub="1313-2970-1027-1" ID-Pensoft-UUID="C5103AABD06058C595280B912CBE9B8B" ID-PubMed="33867800" ID-ZooBank="7D179ED7D7A540A2A972E07BC648F3B9" ModsDocID="1313-2970-1027-1" checkinTime="1617435236653" checkinUser="pensoft" docAuthor="Godwin, Rebecca L. &amp; Bond, Jason E." docDate="2021" docId="4C77DF7122B356E6BC1EF403ACDA341F" docLanguage="en" docName="ZooKeys 1027: 1-165" docOrigin="ZooKeys 1027" docPubDate="2021-04-02" docSource="http://dx.doi.org/10.3897/zookeys.1027.54888" docTitle="Ummidia quijichacaca Godwin &amp; Bond 2021, sp. nov." docType="treatment" docUuid="36DF98D3-1C6D-43A0-85D2-61FFF3E03A8A" docUuidSource="ZooBank" docVersion="5" id="C5103AABD06058C595280B912CBE9B8B" lastPageNumber="1" masterDocId="C5103AABD06058C595280B912CBE9B8B" masterDocTitle="Taxonomic revision of the New World members of the trapdoor spider genus Ummidia Thorell (Araneae, Mygalomorphae, Halonoproctidae)" masterLastPageNumber="165" masterPageNumber="1" pageNumber="1" updateTime="1730571528781" updateUser="felipe">
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<mods:title id="7AAC9BA9ED015B83EE61CBE031804DB7">Taxonomic revision of the New World members of the trapdoor spider genus Ummidia Thorell (Araneae, Mygalomorphae, Halonoproctidae)</mods:title>
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<mods:namePart id="676878A6F44F700EFA1F3C43115D6872">Godwin, Rebecca L.</mods:namePart>
<mods:nameIdentifier id="4DD0DA2E520C3186068BB371700F9CD0" type="ORCID">https://orcid.org/0000-0002-2439-2868</mods:nameIdentifier>
<mods:affiliation id="3C389FEF4FE0BB73C2C294A5FA712B20">Department of Entomology and Nematology, University of California, Davis 1 Shields Ave, Davis, CA, 95616</mods:affiliation>
<mods:nameIdentifier id="C3A6F180A87ADDB90879191538C0BEC9" type="email">rlc0004@tigermail.auburn.edu</mods:nameIdentifier>
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<mods:namePart id="53D728881B8EF5A39AC93E54BE089479">Bond, Jason E.</mods:namePart>
<mods:affiliation id="2EF36FBFE58D70F30D6ACF00C78C59A7">Department of Entomology and Nematology, University of California, Davis 1 Shields Ave, Davis, CA, 95616</mods:affiliation>
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<mods:date id="F0AF9631B5A8FFC47370EC78B5F92FBE">2021</mods:date>
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<mods:number id="2487AA3BFE329F1A128170CBD8A20D96">1027</mods:number>
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<mods:classification id="0046E5BA7D89F8136E952CA7BFF9535B">journal article</mods:classification>
<mods:identifier id="9EFFBCC26E211DCBCE63CFED513DBE65" type="DOI">http://dx.doi.org/10.3897/zookeys.1027.54888</mods:identifier>
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<treatment id="4C77DF7122B356E6BC1EF403ACDA341F" ID-GBIF-Taxon="180850642" LSID="urn:lsid:zoobank.org:act:36DF98D3-1C6D-43A0-85D2-61FFF3E03A8A" httpUri="http://treatment.plazi.org/id/4C77DF7122B356E6BC1EF403ACDA341F" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection id="3140E917F7F7023EDF77B09D3E146D13" pageId="0" pageNumber="1" type="nomenclature">
<paragraph id="BD79FB621C0764487055CBC117D3027A" pageId="0" pageNumber="1">
<taxonomicName id="BDF3D5705AF0F22DD00896F9EF974180" ID-CoL="98Z3J" LSID="http://zoobank.org/36DF98D3-1C6D-43A0-85D2-61FFF3E03A8A" authority="Godwin &amp; Bond, 2021" authorityName="Godwin &amp; Bond" authorityYear="2021" class="Arachnida" family="Halonoproctidae" genus="Ummidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ummidia quijichacaca" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="quijichacaca" status="sp. nov.">Ummidia quijichacaca</taxonomicName>
<taxonomicNameLabel id="AD25C639FC7044FEB562022235ACA12A" pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
<figureCitation id="E61967592E9D6D24CB56D0F6A4BCF2A7" captionStart="Figure 66" captionStartId="F71" captionText="Figure 66. Ummidia quijichacaca sp. nov. female holotype (UMM 230) from Monserrate, Colombia A female habitus illustration B cleared spermathecae. Scale bars: 4.0 mm (A), 1.0 mm (B)." figureDoi="10.3897/zookeys.1027.54888.figure66" httpUri="https://binary.pensoft.net/fig/527531" pageId="0" pageNumber="1">Fig. 66</figureCitation>
<figureCitation id="C9253563647498F1BF076BDFB1906E4F" captionStart="Map 6" captionStartId="F72" captionText="Map 6. Locality records for South American and Caribbean species." figureDoi="10.3897/zookeys.1027.54888.map6" httpUri="https://binary.pensoft.net/fig/527530" pageId="0" pageNumber="1">, Map 6</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="63B994B83D33DC7152691782ED6B2D22" pageId="0" pageNumber="1" type="type material">
<paragraph id="F644ECD176F6CCC248CDADF4C97A0983" pageId="0" pageNumber="1">Type material.</paragraph>
<paragraph id="AAF0DF86E3D2EEB53C944346076363DC" pageId="0" pageNumber="1">
HOLOTYPE: 1 ♀ (UMM230) from Monserrate above
<normalizedToken id="EBFDF5414F61BEEDD0B9B3F7C9B1A99A" originalValue="Bogotá">Bogota</normalizedToken>
, Cundinamarca, Colombia,
<geoCoordinate id="AE3D6FA4204B698B0DF7AA3FC4B019C0" degrees="4.6049" direction="north" orientation="latitude" precision="5" value="4.6049">4.6049</geoCoordinate>
<geoCoordinate id="C0B2C051D35258C0FF9B7832ADBE3318" degrees="74.0565" direction="west" orientation="longitude" precision="5" value="-74.0565">-74.0565</geoCoordinate>
5, 3152 m a.s.l., 27.iv.1965, coll. PR Craig, DL Craig, AMNH.
</paragraph>
</subSubSection>
<subSubSection id="73C290A58D28115227E5286FC3B46F9E" pageId="0" pageNumber="1" type="etymology">
<paragraph id="24A347E34A6E1AC380D87889C244D33A" pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph id="47AF14698069AD4E30AF602A05F3ACDC" pageId="0" pageNumber="1">
The specific epithet is a noun taken in apposition. It is the Muisca name for Monserrate, Quijichacaca, which translates to
<normalizedToken id="19877B5B45C195771F9598F4025376BB" originalValue="“grandmothers">&quot;grandmother's</normalizedToken>
foot&quot;.
</paragraph>
<caption id="35DA477F31EB23763965B158E5BA0D9E" doi="10.3897/zookeys.1027.54888.map6" httpUri="https://binary.pensoft.net/fig/527530" pageId="0" pageNumber="1" start="Map 6" startId="F72">
<paragraph id="6A8B75D3E9ECF8572C17E6BCB2694A2D" pageId="0" pageNumber="1">
<emphasis id="A968F3C0127D5002FB5F7758E461D31B" bold="true" pageId="0" pageNumber="1">Map 6.</emphasis>
Locality records for South American and Caribbean species.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="196B22DB39AE42822500537E704EB2B6" pageId="0" pageNumber="1" type="diagnosis">
<paragraph id="B99CC73D2A414C7D5E113FB98152876C" pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph id="8B456B8A30BFE462ACE300DD5B638953" pageId="0" pageNumber="1">
<taxonomicName id="D7EE4B84F6849F87BB430CC83A0EA9E3" authorityName="Godwin &amp; Bond" authorityYear="2021" class="Arachnida" family="Halonoproctidae" genus="Ummidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ummidia quijichacaca" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="quijichacaca">
<emphasis id="539DD2736F80968ACD8766C84D791BDA" italics="true" pageId="0" pageNumber="1">Ummidia quijichacaca</emphasis>
</taxonomicName>
can be differentiated from all geographically proximate species by having the carapace angular and longer than wide and legs which are darker distally. Females can be differentiated from
<taxonomicName id="9F84CD4A8A81C6AF4E81164105CEAD60" class="Arachnida" family="Halonoproctidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="U. tibacuy" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="tibacuy">
<emphasis id="79C56AF3C7674DAD747211F3218973F5" italics="true" pageId="0" pageNumber="1">U. tibacuy</emphasis>
</taxonomicName>
,
<taxonomicName id="4BD43CB42A16C9E0E90BB8DA7964EACD" class="Arachnida" family="Halonoproctidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="U. asperula" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="asperula">
<emphasis id="546EA5471F9BDB3B06F16777E0E7C089" italics="true" pageId="0" pageNumber="1">U. asperula</emphasis>
</taxonomicName>
, and
<taxonomicName id="A3ABD85B97AF8E131B63ED4FDB2AE37E" class="Arachnida" family="Halonoproctidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="U. neblina" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="neblina">
<emphasis id="09C2E3B38FDFA1BE2836BB3DE73EE52D" italics="true" pageId="0" pageNumber="1">U. neblina</emphasis>
</taxonomicName>
by the presence of a defined comb of long spinules on the retrolateral face of tarsus IV. Females can be further distinguished from all geographically proximate species by having spermathecae which are straight with wide, flat, anteriorly facing bulbs.
</paragraph>
<caption id="D5D35A4D89D2D3C834349A27783B429E" doi="10.3897/zookeys.1027.54888.figure66" httpUri="https://binary.pensoft.net/fig/527531" pageId="0" pageNumber="1" start="Figure 66" startId="F71">
<paragraph id="EA1AF8E1CE349BB43B85B543939498CE" pageId="0" pageNumber="1">
<emphasis id="8A11AE51DCE968096A6D5EF70FA41367" bold="true" pageId="0" pageNumber="1">Figure 66.</emphasis>
<taxonomicName id="5D8E7B0B53B61BC658E76D69994DAF87" authorityName="Godwin &amp; Bond" authorityYear="2021" class="Arachnida" family="Halonoproctidae" genus="Ummidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ummidia quijichacaca" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="quijichacaca">
<emphasis id="D2A095D772F11CDA9915083EB1845BE2" italics="true" pageId="0" pageNumber="1">Ummidia quijichacaca</emphasis>
</taxonomicName>
sp. nov. female holotype (UMM230) from Monserrate, Colombia
<emphasis id="A4413ECD8C27D686A3B6B5233706B331" bold="true" pageId="0" pageNumber="1">A</emphasis>
female habitus illustration
<emphasis id="028BF4259A84320682DC3434FA2A597C" bold="true" pageId="0" pageNumber="1">B</emphasis>
cleared spermathecae. Scale bars: 4.0 mm (
<emphasis id="B00CB62E8E36454901602B769F07C98D" bold="true" pageId="0" pageNumber="1">A</emphasis>
), 1.0 mm (
<emphasis id="B75CC2D30B85C8FA1CA4AF010F36C3EF" bold="true" pageId="0" pageNumber="1">B</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="139BDD1EA64F1F32FD524EA0567558DB" pageId="0" pageNumber="1" type="description">
<paragraph id="7D606F08F49D36299D0E88A0294DD6B3" pageId="0" pageNumber="1">Description of female holotype.</paragraph>
<paragraph id="8EF8A05287464E134F57E6687B19C5B8" pageId="0" pageNumber="1">
<emphasis id="94C42BCFC6C3629ABD837907A1A0FF18" italics="true" pageId="0" pageNumber="1">Specimen preparation and condition</emphasis>
. Specimen preserved in 80% EtOH. Spermathecae removed, cleared, in vial with specimen.
<emphasis id="66CCBFD2948FCDAF321E62D74ACCD6A9" italics="true" pageId="0" pageNumber="1">General coloration</emphasis>
. Carapace, chelicerae, and legs dark reddish brown 5YR 3/4. Abdomen brown 7.5YR 4/2, spinnerets brownish yellow 10YR 6/6.
<emphasis id="78D5E19B7BC5F6BBA400BAE4D7391FEE" italics="true" pageId="0" pageNumber="1">Cephalothorax</emphasis>
. Carapace 6.83 long, 6.24 wide. Pars cephalica 4.58 long. Foveal groove procurved, 0.46 long, 1.46 wide. All eyes on low tubercle. AER procurved. PER straight. Eye group 1.06 long, 1.54 wide, AME 0.35, PME 0.4, ALE 0.56, PLE 0.41. Sternum with posterior fringe, sparsely setose anteriorly, STRl 4.05, STRw 4.32. Chelicerae with anterior row comprising five teeth, posterior margin with six teeth. Palpal endites with 27 cuspules spread across proximal half of endite and 24 cuspules distally; equal in size to proximal cuspules anteriorly, smaller posteriorly, ENDw 1.57, ENDl 2.7. Labium with seven cuspules, LBw 1.7, LBl 1.24. Rastellum with many strong spines on process and up cheliceral face for ~ 2
<normalizedToken id="385EB894F1A6DE9F84F37B17895BEFD6" originalValue="×">x</normalizedToken>
length of process.
<emphasis id="E68F9F022DC7864ECDE35DCC82A8903F" italics="true" pageId="0" pageNumber="1">Abdomen</emphasis>
. Evenly setose.
<emphasis id="A1AFD4B00228A3E3C3642F272192A130" italics="true" pageId="0" pageNumber="1">Legs</emphasis>
. F1 4.67; F1w 1.63; P1 2.98; Ti1 2.87; Mt1 2.02; Tr1 1.06; F3 3.75; F3w 2.08; P3 2.38; Ti3 2.24, Sd3 1.55; Mt3 1.93; Tr3 1.44; F4 5.05; F4w 2.05; P4 2.69; Ti4 2.93; Mt4 3.1; Tr4 1.43. Retrolateral face tarsus IV with defined comb of long spinules over length of tarsus.
<emphasis id="0FFD72A833263863E09BE5C421569F8C" italics="true" pageId="0" pageNumber="1">Pedipalps</emphasis>
. PF 4.22, PP 2.45, PTi 2.49, PTr 2.3. Spermathecae simple, straight, bulbs facing anteriorly.
</paragraph>
</subSubSection>
<subSubSection id="57B8F3FD9A660D61A5C2BC42E6D80487" pageId="0" pageNumber="1" type="variation">
<paragraph id="50D031DFC3CEE6510CA446D09D7E8FF1" pageId="0" pageNumber="1">Variation, females.</paragraph>
<paragraph id="91D7D56F46975E2A29B03E1F06D6DB9D" pageId="0" pageNumber="1">Known only from female holotype.</paragraph>
</subSubSection>
<subSubSection id="57B7EEFF7708693876FF46CAE1684938" pageId="0" pageNumber="1" type="males">
<paragraph id="1774A4AFAD74D80FFBA39BC90B793D47" pageId="0" pageNumber="1">Males.</paragraph>
<paragraph id="619E7D7F91581DDC9E0809B09CB1930C" pageId="0" pageNumber="1">Unknown.</paragraph>
</subSubSection>
</treatment>
</document>