treatments-xml/data/4C/33/87/4C338790FF8DFFA261FDFC726EC24AEF.xml

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<mods:title id="08646E9746142F0016D67B1584D5E568">On the diagnostic characters of the genus Stygonitocrella (Copepoda, Harpacticoida), with descriptions of seven new species from Australian subterranean waters 2324</mods:title>
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<subSubSection id="8C80650DFF8DFFDE61FDFC726CA54D7B" box="[151,658,1002,1028]" pageId="61" pageNumber="62" type="nomenclature">
<paragraph id="C4253686FF8DFFDE61FDFC726CA54D7B" blockId="61.[151,658,1002,1063]" box="[151,658,1002,1028]" pageId="61" pageNumber="62">
<heading id="9F6D81EAFF8DFFDE61FDFC726CA54D7B" bold="true" box="[151,658,1002,1028]" fontSize="11" level="1" pageId="61" pageNumber="62" reason="1">
<emphasis id="F6EEEA94FF8DFFDE61FDFC726CA54D7B" bold="true" box="[151,658,1002,1028]" pageId="61" pageNumber="62">
<taxonomicName id="039A4D05FF8DFFDE61FDFC726C1A4D7B" authority="Karanovic &amp; Hancock, 2009" authorityName="Karanovic &amp; Hancock" authorityYear="2009" box="[151,557,1002,1028]" class="Hexanauplia" family="Ameiridae" genus="Megastygonitocrella" kingdom="Animalia" order="Harpacticoida" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="pagusregalis" status="sp. nov.">
<emphasis id="F6EEEA94FF8DFFDE61FDFC726C1A4D7B" bold="true" box="[151,557,1002,1028]" italics="true" pageId="61" pageNumber="62">Megastygonitocrella pagusregalis</emphasis>
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<subSubSection id="8C80650DFF8DFFDE61FDFB956F4E4D58" box="[151,377,1037,1063]" pageId="61" pageNumber="62" type="description">
<paragraph id="C4253686FF8DFFDE61FDFB956F4E4D58" blockId="61.[151,658,1002,1063]" box="[151,377,1037,1063]" pageId="61" pageNumber="62">
(
<figureCitation id="5CA12A03FF8DFFDE61F5FB956EC04D58" box="[159,247,1037,1063]" captionStart="FIGURE 15" captionStartId="64.[151,255,1922,1946]" captionTargetBox="[202,1389,198,1860]" captionTargetId="figure-60@64.[151,1436,168,1942]" captionTargetPageId="64" captionText="FIGURE 15. Megastygonitocrella pagusregalis gen. et sp. nov., holotype female: A—habitus, lateral view; B—habitus, dorsal view; C—urosome, ventral view; D—anal somite and left caudal ramus, lateral view; E—antennula; F—antenna; G—labrum; H—paragnaths; I—mandibula; J—maxillula; K—maxilla; L—maxilliped. Scales = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5351289" httpUri="https://zenodo.org/record/5351289/files/figure.png" pageId="61" pageNumber="62">Figs 15</figureCitation>
&amp; 
<figureCitation id="5CA12A03FF8DFFDE6077FB956F454D58" box="[285,370,1037,1063]" captionStart="FIGURE 16" captionStartId="66.[151,256,1834,1858]" captionTargetBox="[238,1380,198,1799]" captionTargetId="figure-119@66.[182,1416,166,1869]" captionTargetPageId="66" captionText="FIGURE 16. Megastygonitocrella pagusregalis gen. et sp. nov., A–E, holotype female; Megastygonitocrella kryptos gen. et sp. nov., F–M, holotype male; N &amp; O, topotype male (WAM C37347; 0.427 mm): A—first swimming leg; B— second swimming leg without second and third exopodal segments; C—third swimming leg; D—fourth swimming leg; E—left fifth leg; F—anal somite and caudal rami, dorsal view; G—basis of first swimming leg; H—endopod of second swimming leg; I—endopod of fourth swimming leg; J—left fifth leg; K—left sixth leg; L—spermatophore; M— antennula; N—urosome, ventral view; O—third exopodal segment of fourth swimming leg. Scale = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5351291" httpUri="https://zenodo.org/record/5351291/files/figure.png" pageId="61" pageNumber="62">16A–E</figureCitation>
)
</paragraph>
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<emphasis id="F6EEEA94FF8DFFDE61FDFBC26F614D0B" bold="true" box="[151,342,1114,1140]" pageId="61" pageNumber="62">Type material.</emphasis>
<materialsCitation id="74F23CDBFF8DFFDE600BFBC26B4D4DE4" accessionNumber="C37345" collectingDate="2003-08-07" collectionCode="WAM" collectorName="P. Hancock" country="Australia" elevation="10" latitude="-21.326946" location="Pioneer Valley" longLatPrecision="21" longitude="149.00166" pageId="61" pageNumber="62" specimenCount="1" stateProvince="Queensland" typeStatus="holotype">
<typeStatus id="1B218824FF8DFFDE600BFBC26FE04D0B" box="[353,471,1114,1140]" pageId="61" pageNumber="62" type="holotype">Holotype</typeStatus>
, adult female dissected on one slide (
<collectionCode id="A28BAE43FF8DFFDE62C6FBC26DCF4D0B" box="[940,1016,1114,1140]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="61" pageNumber="62" type="Museum">WAM</collectionCode>
C37345); 
<collectingCountry id="BC8D7616FF8DFFDE65EAFBC26AC24D0B" box="[1152,1269,1114,1140]" name="Australia" pageId="61" pageNumber="62">Australia</collectingCountry>
, 
<collectingRegion id="065EF864FF8DFFDE6468FBC26BA04D0B" box="[1282,1431,1114,1140]" country="Australia" name="Queensland" pageId="61" pageNumber="62">Queensland</collectingRegion>
, 
<location id="C145605DFF8DFFDE61FDFB196F734DE4" LSID="urn:lsid:plazi:treatment:4C338790FF8DFFA261FDFC726EC24AEF:C145605DFF8DFFDE61FDFB196F734DE4" box="[151,324,1153,1179]" country="Australia" latitude="-21.326946" longLatPrecision="21" longitude="149.00166" name="Pioneer Valley" pageId="61" pageNumber="62" stateProvince="Queensland">Pioneer Valley</location>
, bore 12600065, depth 
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<elevation id="4FB7D1B5FF8DFFDE633CFB196C9D4DE4" box="[598,682,1153,1179]" metricMagnitude="0" metricUnit="m" metricValue="9.57" pageId="61" pageNumber="62" unit="m" value="9.57">9.57 m</elevation>
</quantity>
, 
<date id="B0241046FF8DFFDE63DDFB196D434DE4" box="[695,884,1153,1179]" pageId="61" pageNumber="62" value="2003-08-07">
<collectingDate id="A060E9AEFF8DFFDE63DDFB196D434DE4" box="[695,884,1153,1179]" pageId="61" pageNumber="62" value="2003-08-07">07 August 2003</collectingDate>
</date>
, leg. 
<collectorName id="696F5350FF8DFFDE62D9FB196A0D4DE4" box="[947,1082,1153,1179]" pageId="61" pageNumber="62">P. Hancock</collectorName>
, 
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<geoCoordinate id="A1AE5041FF8DFFDE65B2FB196B414DE4" box="[1240,1398,1152,1179]" degrees="149" direction="east" minutes="00" orientation="longitude" pageId="61" pageNumber="62" precision="15" seconds="06" value="149.00166">149°00’06”E</geoCoordinate>
.
</materialsCitation>
</paragraph>
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<subSubSection id="8C80650DFF8DFFDC61AFFB3F6B6E4E04" lastPageId="63" lastPageNumber="64" pageId="61" pageNumber="62" type="description">
<paragraph id="C4253686FF8DFFDE61AFFB3F6C114CAF" blockId="61.[151,1436,1114,2030]" pageId="61" pageNumber="62">
<emphasis id="F6EEEA94FF8DFFDE61AFFB3F6F524DBE" bold="true" box="[197,357,1191,1217]" pageId="61" pageNumber="62">Description.</emphasis>
FEMALE (
<typeStatus id="1B218824FF8DFFDE6091FB306C924DBD" box="[507,677,1192,1218]" pageId="61" pageNumber="62" type="holotype">HOLOTYPE</typeStatus>
). Body length, excluding caudal setae, 
<quantity id="03629B63FF8DFFDE65F8FB306B224DBD" box="[1170,1301,1192,1218]" metricMagnitude="-4" metricUnit="m" metricValue="4.06" pageId="61" pageNumber="62" unit="mm" value="0.406">0.406 mm</quantity>
. Preserved specimen colourless. Nauplius eye absent. Habitus (
<figureCitation id="5CA12A03FF8DFFDE624DFB566DF54D97" box="[807,962,1230,1256]" captionStart="FIGURE 15" captionStartId="64.[151,255,1922,1946]" captionTargetBox="[202,1389,198,1860]" captionTargetId="figure-60@64.[151,1436,168,1942]" captionTargetPageId="64" captionText="FIGURE 15. Megastygonitocrella pagusregalis gen. et sp. nov., holotype female: A—habitus, lateral view; B—habitus, dorsal view; C—urosome, ventral view; D—anal somite and left caudal ramus, lateral view; E—antennula; F—antenna; G—labrum; H—paragnaths; I—mandibula; J—maxillula; K—maxilla; L—maxilliped. Scales = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5351289" httpUri="https://zenodo.org/record/5351289/files/figure.png" pageId="61" pageNumber="62">Figs 15A–B</figureCitation>
) cylindrical, slender, without distinct demarcation between prosome and urosome; prosome/urosome ratio 0.9; greatest width at posterior part of cephalothorax, which is only slightly wider than first and second free prosomites. Body length/width ratio about 5.8; cephalothorax only slightly wider than genital double-somite (1.06 times). Free pedigerous somites without pronounced lateral or dorsal expansions. Integument weakly chitinized and without any cuticular windows. Rostrum (
<figureCitation id="5CA12A03FF8DFFDE60E7FA086FCC4CD5" box="[397,507,1424,1450]" captionStart="FIGURE 15" captionStartId="64.[151,255,1922,1946]" captionTargetBox="[202,1389,198,1860]" captionTargetId="figure-60@64.[151,1436,168,1942]" captionTargetPageId="64" captionText="FIGURE 15. Megastygonitocrella pagusregalis gen. et sp. nov., holotype female: A—habitus, lateral view; B—habitus, dorsal view; C—urosome, ventral view; D—anal somite and left caudal ramus, lateral view; E—antennula; F—antenna; G—labrum; H—paragnaths; I—mandibula; J—maxillula; K—maxilla; L—maxilliped. Scales = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5351289" httpUri="https://zenodo.org/record/5351289/files/figure.png" pageId="61" pageNumber="62">Fig. 15A</figureCitation>
) very small and membranous, not demarcated at base; ornamented with two dorsal sensilla on anterior margin.
</paragraph>
<paragraph id="C4253686FF8DFFDE61AFFA456C1A4FA0" blockId="61.[151,1436,1114,2030]" pageId="61" pageNumber="62">
Cephalothorax (
<figureCitation id="5CA12A03FF8DFFDE60E9FA456C224C88" box="[387,533,1501,1527]" captionStart="FIGURE 15" captionStartId="64.[151,255,1922,1946]" captionTargetBox="[202,1389,198,1860]" captionTargetId="figure-60@64.[151,1436,168,1942]" captionTargetPageId="64" captionText="FIGURE 15. Megastygonitocrella pagusregalis gen. et sp. nov., holotype female: A—habitus, lateral view; B—habitus, dorsal view; C—urosome, ventral view; D—anal somite and left caudal ramus, lateral view; E—antennula; F—antenna; G—labrum; H—paragnaths; I—mandibula; J—maxillula; K—maxilla; L—maxilliped. Scales = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5351289" httpUri="https://zenodo.org/record/5351289/files/figure.png" pageId="61" pageNumber="62">Figs 15A–B</figureCitation>
) with completely incorporated first pedigerous somite, almost cylindrical in dorsal view, 1.2 times as long as wide; represents 21% of total body length. Surface of cephalic shield and tergites of three free pedigerous somites ornamented only with several large sensilla. Hyaline fringe of all prosomites narrow and smooth. Fifth pedigerous (first urosomal) somite ornamented only with four dorsal sensilla (two posteriorly and two at midlength); hyaline fringe smooth both dorsally and laterally (
<figureCitation id="5CA12A03FF8DFFDE6475F9E06E994FC7" captionStart="FIGURE 15" captionStartId="64.[151,255,1922,1946]" captionTargetBox="[202,1389,198,1860]" captionTargetId="figure-60@64.[151,1436,168,1942]" captionTargetPageId="64" captionText="FIGURE 15. Megastygonitocrella pagusregalis gen. et sp. nov., holotype female: A—habitus, lateral view; B—habitus, dorsal view; C—urosome, ventral view; D—anal somite and left caudal ramus, lateral view; E—antennula; F—antenna; G—labrum; H—paragnaths; I—mandibula; J—maxillula; K—maxilla; L—maxilliped. Scales = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5351289" httpUri="https://zenodo.org/record/5351289/files/figure.png" pageId="61" pageNumber="62">Figs 15A– B</figureCitation>
). Large sclerotized joint (
<figureCitation id="5CA12A03FF8DFFDE608DF9066C4B4FC7" box="[487,636,1694,1720]" captionStart="FIGURE 15" captionStartId="64.[151,255,1922,1946]" captionTargetBox="[202,1389,198,1860]" captionTargetId="figure-60@64.[151,1436,168,1942]" captionTargetPageId="64" captionText="FIGURE 15. Megastygonitocrella pagusregalis gen. et sp. nov., holotype female: A—habitus, lateral view; B—habitus, dorsal view; C—urosome, ventral view; D—anal somite and left caudal ramus, lateral view; E—antennula; F—antenna; G—labrum; H—paragnaths; I—mandibula; J—maxillula; K—maxilla; L—maxilliped. Scales = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5351289" httpUri="https://zenodo.org/record/5351289/files/figure.png" pageId="61" pageNumber="62">Figs 15A–C</figureCitation>
) present between fifth pedigerous and genital double somites and visible both ventrally and dorsally.
</paragraph>
<paragraph id="C4253686FF8DFFDD61AFF9746F7D4898" blockId="61.[151,1436,1114,2030]" lastBlockId="62.[151,1437,152,2034]" lastPageId="62" lastPageNumber="63" pageId="61" pageNumber="62">
Genital double-somite (
<figureCitation id="5CA12A03FF8DFFDE608BF9746C7B4E79" box="[481,588,1772,1798]" captionStart="FIGURE 15" captionStartId="64.[151,255,1922,1946]" captionTargetBox="[202,1389,198,1860]" captionTargetId="figure-60@64.[151,1436,168,1942]" captionTargetPageId="64" captionText="FIGURE 15. Megastygonitocrella pagusregalis gen. et sp. nov., holotype female: A—habitus, lateral view; B—habitus, dorsal view; C—urosome, ventral view; D—anal somite and left caudal ramus, lateral view; E—antennula; F—antenna; G—labrum; H—paragnaths; I—mandibula; J—maxillula; K—maxilla; L—maxilliped. Scales = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5351289" httpUri="https://zenodo.org/record/5351289/files/figure.png" pageId="61" pageNumber="62">Fig. 15C</figureCitation>
) approximately 0.8 times as long as wide (ventral view), only slightly wider at posterior part, with suture marking original segmentation clearly visible dorsally and laterally; ornamented with six large sensilla along suture, two posterior dorsal sensilla, four posterior ventral sensilla and one irregular row of relatively large posterior ventral spinules. Hyaline fringe smooth and very narrow. Genital field with single small copulatory pore, sclerotized narrow copulatory duct and two very small, ellipsoid seminal receptacles, posterior part of which not reaching to last third of copulatory duct. Copulatory pore situated at proximal third of double-somite length. Very small genital aperture covered by fused reduced sixth legs, represents 26% of somite width. Third urosomite (
<figureCitation id="5CA12A03FF8EFFDD6235FF006DD949CD" box="[863,1006,152,178]" captionStart="FIGURE 15" captionStartId="64.[151,255,1922,1946]" captionTargetBox="[202,1389,198,1860]" captionTargetId="figure-60@64.[151,1436,168,1942]" captionTargetPageId="64" captionText="FIGURE 15. Megastygonitocrella pagusregalis gen. et sp. nov., holotype female: A—habitus, lateral view; B—habitus, dorsal view; C—urosome, ventral view; D—anal somite and left caudal ramus, lateral view; E—antennula; F—antenna; G—labrum; H—paragnaths; I—mandibula; J—maxillula; K—maxilla; L—maxilliped. Scales = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5351289" httpUri="https://zenodo.org/record/5351289/files/figure.png" pageId="62" pageNumber="63">Figs 15B–C</figureCitation>
) ornamented with six large posterior sensilla (two dorsal and four ventral) and uninterrupted but irregular posterior ventral row of spinules; hyaline fringe smooth. Preanal somite significantly shorter than previous one, more than twice as wide as long and without surface ornamentation. Anal somite (
<figureCitation id="5CA12A03FF8EFFDD63C6FE946D094859" box="[684,830,268,294]" captionStart="FIGURE 15" captionStartId="64.[151,255,1922,1946]" captionTargetBox="[202,1389,198,1860]" captionTargetId="figure-60@64.[151,1436,168,1942]" captionTargetPageId="64" captionText="FIGURE 15. Megastygonitocrella pagusregalis gen. et sp. nov., holotype female: A—habitus, lateral view; B—habitus, dorsal view; C—urosome, ventral view; D—anal somite and left caudal ramus, lateral view; E—antennula; F—antenna; G—labrum; H—paragnaths; I—mandibula; J—maxillula; K—maxilla; L—maxilliped. Scales = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5351289" httpUri="https://zenodo.org/record/5351289/files/figure.png" pageId="62" pageNumber="63">Figs 15C–D</figureCitation>
) ornamented with pair of large dorsal sensilla, four ventral cuticular pores, interrupted and arched row of spinules ventrally at 1/3 and posterior row of somewhat larger spinules interrupted dorsolaterally and ventrally between caudal rami. Anal operculum convex, not reaching to posterior end of anal somite, represents 49% of somite's width; ornamented near posterior margin with six spinules of about same size as spinules on posterior lateral margin of somite. Anal sinus smooth and widely opened.
</paragraph>
<paragraph id="C4253686FF8EFFDD61AFFE6C6B094A3C" blockId="62.[151,1437,152,2034]" pageId="62" pageNumber="63">
Caudal rami (
<figureCitation id="5CA12A03FF8EFFDD6001FE6C6C354B71" box="[363,514,500,526]" captionStart="FIGURE 15" captionStartId="64.[151,255,1922,1946]" captionTargetBox="[202,1389,198,1860]" captionTargetId="figure-60@64.[151,1436,168,1942]" captionTargetPageId="64" captionText="FIGURE 15. Megastygonitocrella pagusregalis gen. et sp. nov., holotype female: A—habitus, lateral view; B—habitus, dorsal view; C—urosome, ventral view; D—anal somite and left caudal ramus, lateral view; E—antennula; F—antenna; G—labrum; H—paragnaths; I—mandibula; J—maxillula; K—maxilla; L—maxilliped. Scales = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5351289" httpUri="https://zenodo.org/record/5351289/files/figure.png" pageId="62" pageNumber="63">Figs 15B–D</figureCitation>
) short and conical, shorter than their greatest width (ventral view), slightly divergent, with space between them more than one ramus width; with small diagonal chitinous ridge dorsally and armed with seven setae (three lateral, three apical and one dorsal). Ornamentation consists of two spinules each at base of dorsal and distal lateral setae, one dorsal and one lateral pore and posterior ventral row of five large spinules. Dorsal seta inserted nearly at posterior end and close to inner margin, 2.3 times as long as caudal ramus, triarticulate at its base and smooth. Proximal lateral seta arising somewhat dorsolaterally, 0.7 times as long as dorsal one and 1.7 times as long as distal lateral seta, which arises at 3/4 of ramus length. Inner apical seta very slender and small, smooth, about 0.8 times as long as ramus. Both principal setae inserted slightly more dorsally than terminally, with breaking plane and sparsely pinnate at distal end.
</paragraph>
<paragraph id="C4253686FF8EFFDD61AFFCC86EF44D07" blockId="62.[151,1437,152,2034]" pageId="62" pageNumber="63">
Antennula (
<figureCitation id="5CA12A03FF8EFFDD6024FCC86F824A15" box="[334,437,848,874]" captionStart="FIGURE 15" captionStartId="64.[151,255,1922,1946]" captionTargetBox="[202,1389,198,1860]" captionTargetId="figure-60@64.[151,1436,168,1942]" captionTargetPageId="64" captionText="FIGURE 15. Megastygonitocrella pagusregalis gen. et sp. nov., holotype female: A—habitus, lateral view; B—habitus, dorsal view; C—urosome, ventral view; D—anal somite and left caudal ramus, lateral view; E—antennula; F—antenna; G—labrum; H—paragnaths; I—mandibula; J—maxillula; K—maxilla; L—maxilliped. Scales = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5351289" httpUri="https://zenodo.org/record/5351289/files/figure.png" pageId="62" pageNumber="63">Fig. 15E</figureCitation>
) eight-segmented, but ancestral seventh and eighth segments mostly fused, unusually short and stout, only about as long as cephalothorax, unornamented. Relatively slender aesthetasc on fourth segment reaches beyond tip of appendage for length of more than last four segments combined; much smaller and even more slender apical aesthetasc on eighth segment fused basally to two apical setae. Setal formula: 1.8.6.3.2.2.4.7. All segments, except second and eighth, wider than long. All setae smooth, slender and without breaking plane. Only four setae on eighth segment articulating on basal part. Length ratio of antennular segments, from proximal to distal end and along caudal margin, 1: 1.6: 0.7: 0.8: 0.6: 0.6: 0.3: 0.8.
</paragraph>
<paragraph id="C4253686FF8EFFDD61AFFB1D6DD44C84" blockId="62.[151,1437,152,2034]" pageId="62" pageNumber="63">
Antenna (
<figureCitation id="5CA12A03FF8EFFDD6053FB1D6FA84DE0" box="[313,415,1157,1183]" captionStart="FIGURE 15" captionStartId="64.[151,255,1922,1946]" captionTargetBox="[202,1389,198,1860]" captionTargetId="figure-60@64.[151,1436,168,1942]" captionTargetPageId="64" captionText="FIGURE 15. Megastygonitocrella pagusregalis gen. et sp. nov., holotype female: A—habitus, lateral view; B—habitus, dorsal view; C—urosome, ventral view; D—anal somite and left caudal ramus, lateral view; E—antennula; F—antenna; G—labrum; H—paragnaths; I—mandibula; J—maxillula; K—maxilla; L—maxilliped. Scales = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5351289" httpUri="https://zenodo.org/record/5351289/files/figure.png" pageId="62" pageNumber="63">Fig. 15F</figureCitation>
) composed of coxa, basis, two-segmented endopod and one-segmented exopod. Coxa very short, unornamented. Basis about 1.3 times as long as wide, unarmed but ornamented with short row of spinules on anterior distal margin. First endopodal segment 1.5 times as long as basis, nearly twice as long as wide, unornamented and unarmed. Second endopodal segment longest, 1.4 times as long as first and 3.1 times as long as wide, armed laterally with two smooth spines flanking thin seta; apical armature consisting of five geniculate setae, longest one fused basally to additional smaller seta bearing proximal tuft of fine setules; ornamentation consists of few spinules along anterior surface and two fringes on posterior surface. Exopod one-segmented, 0.7 times as long as basis and 2.3 times as long as wide, unornamented but armed with three setae; inner (anterior) seta bipinnate, slender, arising apically and 1.5 times as long as other two setae, which are both slightly curved, spiniform and unipinnate at anterior distal part.
</paragraph>
<paragraph id="C4253686FF8EFFDD61AFF9906EC44FE9" blockId="62.[151,1437,152,2034]" pageId="62" pageNumber="63">
Labrum (
<figureCitation id="5CA12A03FF8EFFDD6058F9906FAA4F5D" box="[306,413,1544,1570]" captionStart="FIGURE 15" captionStartId="64.[151,255,1922,1946]" captionTargetBox="[202,1389,198,1860]" captionTargetId="figure-60@64.[151,1436,168,1942]" captionTargetPageId="64" captionText="FIGURE 15. Megastygonitocrella pagusregalis gen. et sp. nov., holotype female: A—habitus, lateral view; B—habitus, dorsal view; C—urosome, ventral view; D—anal somite and left caudal ramus, lateral view; E—antennula; F—antenna; G—labrum; H—paragnaths; I—mandibula; J—maxillula; K—maxilla; L—maxilliped. Scales = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5351289" httpUri="https://zenodo.org/record/5351289/files/figure.png" pageId="62" pageNumber="63">Fig. 15G</figureCitation>
) not very large when compared to cephalothorax, trapezoidal, rigidly sclerotized, with relatively broad and convex cutting edge, ornamented with one apical row of smaller spinules in between two subapical rows of strong spinules. Two ellipsoid fields of gustatory papillae visible on dorsal (posterior) surface.
</paragraph>
<paragraph id="C4253686FF8EFFDD61AFF93B6BBE4FC2" blockId="62.[151,1437,152,2034]" box="[197,1417,1699,1725]" pageId="62" pageNumber="63">Paragnaths very similar to previous species, but not mounted satisfactorily enough to allow for drawing.</paragraph>
<paragraph id="C4253686FF8EFFDD61AFF9516FD74E01" blockId="62.[151,1437,152,2034]" pageId="62" pageNumber="63">
Mandibula (
<figureCitation id="5CA12A03FF8EFFDD603CF9516F804F9C" box="[342,439,1737,1763]" captionStart="FIGURE 15" captionStartId="64.[151,255,1922,1946]" captionTargetBox="[202,1389,198,1860]" captionTargetId="figure-60@64.[151,1436,168,1942]" captionTargetPageId="64" captionText="FIGURE 15. Megastygonitocrella pagusregalis gen. et sp. nov., holotype female: A—habitus, lateral view; B—habitus, dorsal view; C—urosome, ventral view; D—anal somite and left caudal ramus, lateral view; E—antennula; F—antenna; G—labrum; H—paragnaths; I—mandibula; J—maxillula; K—maxilla; L—maxilliped. Scales = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5351289" httpUri="https://zenodo.org/record/5351289/files/figure.png" pageId="62" pageNumber="63">Fig. 15I</figureCitation>
) very small, with wide cutting edge on relatively short coxa, armed with numerous small teeth in between three coarse ventral teeth and one dorsal unipinnate seta. Palp uniramous, comprising basis and one-segmented endopod. Basis short, unarmed and unornamented, 1.4 times as long as wide and 1.2 times as long as endopod. Endopod small and unornamented, about 1.5 times as long as wide; armed with five slender smooth apical setae.
</paragraph>
<paragraph id="C4253686FF8EFFDC61AFF8136DEB4980" blockId="62.[151,1437,152,2034]" lastBlockId="63.[151,1436,152,846]" lastPageId="63" lastPageNumber="64" pageId="62" pageNumber="63">
Maxillula (
<figureCitation id="5CA12A03FF8EFFDD6024F8136F844EDA" box="[334,435,1931,1957]" captionStart="FIGURE 15" captionStartId="64.[151,255,1922,1946]" captionTargetBox="[202,1389,198,1860]" captionTargetId="figure-60@64.[151,1436,168,1942]" captionTargetPageId="64" captionText="FIGURE 15. Megastygonitocrella pagusregalis gen. et sp. nov., holotype female: A—habitus, lateral view; B—habitus, dorsal view; C—urosome, ventral view; D—anal somite and left caudal ramus, lateral view; E—antennula; F—antenna; G—labrum; H—paragnaths; I—mandibula; J—maxillula; K—maxilla; L—maxilliped. Scales = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5351289" httpUri="https://zenodo.org/record/5351289/files/figure.png" pageId="62" pageNumber="63">Fig. 15J</figureCitation>
) very small compared to body size, with large praecoxa; arthrite rectangular, not movable, unornamented, without smooth setae on anterior surface, armed with three minute setae on dorsal margin and four apical elements (probably three spines and one seta). Coxal endite slightly shorter than praecoxal arthrite, armed apically with one pinnate and curved and two slender smooth setae, all of about same length. Basis about as long as coxal endite, armed with four smooth setae apically and one subapically. Endopod a minute but distinct segment, armed with smooth apical seta.
</paragraph>
<paragraph id="C4253686FF8FFFDC61AFFE946D3948E5" blockId="63.[151,1436,152,846]" pageId="63" pageNumber="64">
Maxilla (
<figureCitation id="5CA12A03FF8FFFDC605CFE946F914859" box="[310,422,268,294]" captionStart="FIGURE 15" captionStartId="64.[151,255,1922,1946]" captionTargetBox="[202,1389,198,1860]" captionTargetId="figure-60@64.[151,1436,168,1942]" captionTargetPageId="64" captionText="FIGURE 15. Megastygonitocrella pagusregalis gen. et sp. nov., holotype female: A—habitus, lateral view; B—habitus, dorsal view; C—urosome, ventral view; D—anal somite and left caudal ramus, lateral view; E—antennula; F—antenna; G—labrum; H—paragnaths; I—mandibula; J—maxillula; K—maxilla; L—maxilliped. Scales = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5351289" httpUri="https://zenodo.org/record/5351289/files/figure.png" pageId="63" pageNumber="64">Fig. 15K</figureCitation>
) small, unornamented, with proximal endite on syncoxa absent; distal endite well developed, highly mobile, armed with one pinnate spine and two smooth setae of about same length. Basis drawn out into long claw, with shorter spiniform seta at its base. Endopod represented by minute segment, armed with two smooth and equally long apical setae.
</paragraph>
<paragraph id="C4253686FF8FFFDC61AFFE3E6B084B71" blockId="63.[151,1436,152,846]" pageId="63" pageNumber="64">
Maxilliped (
<figureCitation id="5CA12A03FF8FFFDC603FFE3E6F8B48BF" box="[341,444,422,448]" captionStart="FIGURE 15" captionStartId="64.[151,255,1922,1946]" captionTargetBox="[202,1389,198,1860]" captionTargetId="figure-60@64.[151,1436,168,1942]" captionTargetPageId="64" captionText="FIGURE 15. Megastygonitocrella pagusregalis gen. et sp. nov., holotype female: A—habitus, lateral view; B—habitus, dorsal view; C—urosome, ventral view; D—anal somite and left caudal ramus, lateral view; E—antennula; F—antenna; G—labrum; H—paragnaths; I—mandibula; J—maxillula; K—maxilla; L—maxilliped. Scales = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5351289" httpUri="https://zenodo.org/record/5351289/files/figure.png" pageId="63" pageNumber="64">Fig. 15L</figureCitation>
) unornamented, with short syncoxa armed with single smooth seta, 1.4 times as long as wide. Basis 2.4 times as long as wide and 1.8 times as long as syncoxa, unarmed. Endopod represented by long curved claw, ornamented distally with row of spinules along concave side; with thin seta at base.
</paragraph>
<paragraph id="C4253686FF8FFFDC61AFFD826D624BFD" blockId="63.[151,1436,152,846]" pageId="63" pageNumber="64">
All swimming legs with three-segmented exopod; endopod of first leg three-segmented (
<figureCitation id="5CA12A03FF8FFFDC6470FD826BBA4B4B" box="[1306,1421,538,564]" captionStart="FIGURE 16" captionStartId="66.[151,256,1834,1858]" captionTargetBox="[238,1380,198,1799]" captionTargetId="figure-119@66.[182,1416,166,1869]" captionTargetPageId="66" captionText="FIGURE 16. Megastygonitocrella pagusregalis gen. et sp. nov., A–E, holotype female; Megastygonitocrella kryptos gen. et sp. nov., F–M, holotype male; N &amp; O, topotype male (WAM C37347; 0.427 mm): A—first swimming leg; B— second swimming leg without second and third exopodal segments; C—third swimming leg; D—fourth swimming leg; E—left fifth leg; F—anal somite and caudal rami, dorsal view; G—basis of first swimming leg; H—endopod of second swimming leg; I—endopod of fourth swimming leg; J—left fifth leg; K—left sixth leg; L—spermatophore; M— antennula; N—urosome, ventral view; O—third exopodal segment of fourth swimming leg. Scale = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5351291" httpUri="https://zenodo.org/record/5351291/files/figure.png" pageId="63" pageNumber="64">Fig. 16A</figureCitation>
), endopod of other swimming legs one-segmented (
<figureCitation id="5CA12A03FF8FFFDC6268FDD96DA94B24" box="[770,926,577,603]" captionStart="FIGURE 16" captionStartId="66.[151,256,1834,1858]" captionTargetBox="[238,1380,198,1799]" captionTargetId="figure-119@66.[182,1416,166,1869]" captionTargetPageId="66" captionText="FIGURE 16. Megastygonitocrella pagusregalis gen. et sp. nov., A–E, holotype female; Megastygonitocrella kryptos gen. et sp. nov., F–M, holotype male; N &amp; O, topotype male (WAM C37347; 0.427 mm): A—first swimming leg; B— second swimming leg without second and third exopodal segments; C—third swimming leg; D—fourth swimming leg; E—left fifth leg; F—anal somite and caudal rami, dorsal view; G—basis of first swimming leg; H—endopod of second swimming leg; I—endopod of fourth swimming leg; J—left fifth leg; K—left sixth leg; L—spermatophore; M— antennula; N—urosome, ventral view; O—third exopodal segment of fourth swimming leg. Scale = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5351291" httpUri="https://zenodo.org/record/5351291/files/figure.png" pageId="63" pageNumber="64">Figs 16B–D</figureCitation>
). Armature formula of swimming legs as follows (inner/outer element; inner/terminal/outer element):
</paragraph>
<paragraph id="C4253686FF8FFFDC6006FD166DD34BDA" blockId="63.[151,1436,152,846]" box="[364,996,654,677]" pageId="63" pageNumber="64">Exopod Endopod</paragraph>
<paragraph id="C4253686FF8FFFDC61FDFD366A684BBA" blockId="63.[151,1436,152,846]" box="[151,1119,686,709]" pageId="63" pageNumber="64">Segments 1 2 3 1 2 3</paragraph>
<paragraph id="C4253686FF8FFFDC61FDFD566ABE4B9A" blockId="63.[151,1436,152,846]" box="[151,1161,718,741]" pageId="63" pageNumber="64">First leg 0/1 0/1 0/2/2 1/0 0/0 1/1/1</paragraph>
<paragraph id="C4253686FF8FFFDC61FDFD686A6C4A78" blockId="63.[151,1436,152,846]" box="[151,1115,750,776]" pageId="63" pageNumber="64">Second leg 0/1 1/1 0/2/2 0/1/0 - -</paragraph>
<paragraph id="C4253686FF8FFFDC61FDFC8B6A6C4A55" blockId="63.[151,1436,152,846]" box="[151,1115,785,811]" pageId="63" pageNumber="64">Third leg 0/1 1/1 0/2/2 1/1/0 - -</paragraph>
<paragraph id="C4253686FF8FFFDC61FDFCAD6A6C4A33" blockId="63.[151,1436,152,846]" box="[151,1115,820,846]" pageId="63" pageNumber="64">Fourth leg 0/1 1/1 0/2/2 0/1/0 - -</paragraph>
<paragraph id="C4253686FF8FFFDC61AFFCEC6F3B4F60" blockId="63.[151,1436,884,2031]" pageId="63" pageNumber="64">
Intercoxal sclerite of all swimming legs small, with concave distal margin and without surface ornamentation. Praecoxae short and smooth. Coxa of third leg with two diagonal rows of spinules near outer margin, that of other legs smooth; all coxae unarmed. Basis of each leg ornamented with spinules near outer margin at base of exopod and with smaller ones at base of endopod; first leg additionally with several minute spinules at base of inner spine; armed with small outer spine on first and second swimming legs and smooth outer seta on third and fourth legs; first leg with stout spine on inner margin at about 3/5 of segment length. All exopodal and endopodal segments ornamented with strong spinules along outer margin and on outer distal corner; endopod and some exopodal segments with spinules along inner margin as well; inner distal corner of first and second exopodal segments with frilled membrane. All exopodal segments of about same length. First endopodal segment of first swimming leg (
<figureCitation id="5CA12A03FF8FFFDC63F2FB486D334D95" box="[664,772,1232,1258]" captionStart="FIGURE 16" captionStartId="66.[151,256,1834,1858]" captionTargetBox="[238,1380,198,1799]" captionTargetId="figure-119@66.[182,1416,166,1869]" captionTargetPageId="66" captionText="FIGURE 16. Megastygonitocrella pagusregalis gen. et sp. nov., A–E, holotype female; Megastygonitocrella kryptos gen. et sp. nov., F–M, holotype male; N &amp; O, topotype male (WAM C37347; 0.427 mm): A—first swimming leg; B— second swimming leg without second and third exopodal segments; C—third swimming leg; D—fourth swimming leg; E—left fifth leg; F—anal somite and caudal rami, dorsal view; G—basis of first swimming leg; H—endopod of second swimming leg; I—endopod of fourth swimming leg; J—left fifth leg; K—left sixth leg; L—spermatophore; M— antennula; N—urosome, ventral view; O—third exopodal segment of fourth swimming leg. Scale = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5351291" httpUri="https://zenodo.org/record/5351291/files/figure.png" pageId="63" pageNumber="64">Fig. 16A</figureCitation>
) about 2.5 times as long as wide and reaching to 2/3 of second exopodal segment; endopod significantly longer than exopod. Endopod of second and third swimming legs about as long as first exopodal segment; that of fourth leg somewhat shorter. Apical armature element(s) on first leg geniculate, pinnate on outer (concave) side and plumose on inner side. Inner apical seta on third exopodal segment of other legs plumose; outer apical seta plumose on inner margin, pinnate on outer (transitional stage between seta and spine). Apical endopodal element of second, third and fourth legs short and spiniform; inner seta on endopod of third leg slender, smooth and about as long as apical element. Inner element on second exopodal segment short and spiniform, while all outer exopodal spines strong and bipinnate.
</paragraph>
<paragraph id="C4253686FF8FFFDC61AFF9B46DE34F9F" blockId="63.[151,1436,884,2031]" pageId="63" pageNumber="64">
Fifth leg (
<figureCitation id="5CA12A03FF8FFFDC6052F9B46F904F39" box="[312,423,1580,1606]" captionStart="FIGURE 15" captionStartId="64.[151,255,1922,1946]" captionTargetBox="[202,1389,198,1860]" captionTargetId="figure-60@64.[151,1436,168,1942]" captionTargetPageId="64" captionText="FIGURE 15. Megastygonitocrella pagusregalis gen. et sp. nov., holotype female: A—habitus, lateral view; B—habitus, dorsal view; C—urosome, ventral view; D—anal somite and left caudal ramus, lateral view; E—antennula; F—antenna; G—labrum; H—paragnaths; I—mandibula; J—maxillula; K—maxilla; L—maxilliped. Scales = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5351289" httpUri="https://zenodo.org/record/5351289/files/figure.png" pageId="63" pageNumber="64">Figs 15C</figureCitation>
, 
<figureCitation id="5CA12A03FF8FFFDC60D8F9B46FD44F39" box="[434,483,1580,1606]" captionStart="FIGURE 16" captionStartId="66.[151,256,1834,1858]" captionTargetBox="[238,1380,198,1799]" captionTargetId="figure-119@66.[182,1416,166,1869]" captionTargetPageId="66" captionText="FIGURE 16. Megastygonitocrella pagusregalis gen. et sp. nov., A–E, holotype female; Megastygonitocrella kryptos gen. et sp. nov., F–M, holotype male; N &amp; O, topotype male (WAM C37347; 0.427 mm): A—first swimming leg; B— second swimming leg without second and third exopodal segments; C—third swimming leg; D—fourth swimming leg; E—left fifth leg; F—anal somite and caudal rami, dorsal view; G—basis of first swimming leg; H—endopod of second swimming leg; I—endopod of fourth swimming leg; J—left fifth leg; K—left sixth leg; L—spermatophore; M— antennula; N—urosome, ventral view; O—third exopodal segment of fourth swimming leg. Scale = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5351291" httpUri="https://zenodo.org/record/5351291/files/figure.png" pageId="63" pageNumber="64">16E</figureCitation>
) fused almost completely to somite, represented by two unornamented but armed ventrolateral knobs. Inner knob, representing exopod, armed with three subequal, smooth and slender setae. Outer knob represents outer part of basis and armed with single long unipinnate seta; this seta much longer than any seta on inner knob, reaching to 3/4 of genital double-somite length. Inner part of each baseoendopod absent, fused together into smooth and slightly concave hyaline fringe.
</paragraph>
<paragraph id="C4253686FF8FFFDC61AFF9756C824E51" blockId="63.[151,1436,884,2031]" pageId="63" pageNumber="64">
Sixth legs (
<figureCitation id="5CA12A03FF8FFFDC6025F9756F8C4E78" box="[335,443,1773,1799]" captionStart="FIGURE 15" captionStartId="64.[151,255,1922,1946]" captionTargetBox="[202,1389,198,1860]" captionTargetId="figure-60@64.[151,1436,168,1942]" captionTargetPageId="64" captionText="FIGURE 15. Megastygonitocrella pagusregalis gen. et sp. nov., holotype female: A—habitus, lateral view; B—habitus, dorsal view; C—urosome, ventral view; D—anal somite and left caudal ramus, lateral view; E—antennula; F—antenna; G—labrum; H—paragnaths; I—mandibula; J—maxillula; K—maxilla; L—maxilliped. Scales = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5351289" httpUri="https://zenodo.org/record/5351289/files/figure.png" pageId="63" pageNumber="64">Fig. 15C</figureCitation>
) completely fused together, indistinct, forming simple operculum covering single gonopore, without ornamentation or armature.
</paragraph>
<paragraph id="C4253686FF8FFFDC61AFF8A36F954E2A" blockId="63.[151,1436,884,2031]" box="[197,418,1851,1877]" pageId="63" pageNumber="64">MALE. Unknown.</paragraph>
<paragraph id="C4253686FF8FFFDC61AFF8F86B6E4E04" blockId="63.[151,1436,884,2031]" box="[197,1369,1888,1915]" pageId="63" pageNumber="64">
<emphasis id="F6EEEA94FF8FFFDC61AFF8F86F664E05" bold="true" box="[197,337,1888,1914]" pageId="63" pageNumber="64">Variability.</emphasis>
Only 
<specimenCount id="D29CFD0FFF8FFFDC60F0F8F96C2C4E04" box="[410,539,1889,1915]" pageId="63" pageNumber="64" type="female">one female</specimenCount>
was collected and studied, and no asymmetric features were observed.
</paragraph>
</subSubSection>
<subSubSection id="8C80650DFF8FFFDC61AFF81F6CE64E90" pageId="63" pageNumber="64" type="etymology">
<paragraph id="C4253686FF8FFFDC61AFF81F6CE64E90" blockId="63.[151,1436,884,2031]" pageId="63" pageNumber="64">
<emphasis id="F6EEEA94FF8FFFDC61AFF81F6F654EDE" bold="true" box="[197,338,1927,1953]" pageId="63" pageNumber="64">Etymology.</emphasis>
The species name is a combination of a Latin noun “pagus” (meaning “district”, “state”) and a Latin possessive adjective “regalis” (meaning “royal”) and is a rough translation of 
<collectingRegion id="065EF864FF8FFFDC65B8F8366B5F4EB7" box="[1234,1384,1966,1992]" country="Australia" name="Queensland" pageId="63" pageNumber="64">Queensland</collectingRegion>
, the Australian state in which this species was found.
</paragraph>
</subSubSection>
<caption id="90E5660EFFF0FFA361FDF81A6AC04EA0" ID-DOI="http://doi.org/10.5281/zenodo.5351289" ID-Zenodo-Dep="5351289" httpUri="https://zenodo.org/record/5351289/files/figure.png" pageId="64" pageNumber="65" startId="64.[151,255,1922,1946]" targetBox="[202,1389,198,1860]" targetPageId="64">
<paragraph id="C4253686FFF0FFA361FDF81A6AC04EA0" blockId="64.[151,1436,1922,2015]" pageId="64" pageNumber="65">
<emphasis id="F6EEEA94FFF0FFA361FDF81A6F114EE5" bold="true" box="[151,294,1922,1946]" pageId="64" pageNumber="65">FIGURE 15.</emphasis>
<taxonomicName id="039A4D05FFF0FFA36046F81A6CA04EE6" authority="Karanovic &amp; Hancock, 2009" authorityName="Karanovic &amp; Hancock" authorityYear="2009" box="[300,663,1922,1945]" class="Hexanauplia" family="Ameiridae" genus="Megastygonitocrella" kingdom="Animalia" order="Harpacticoida" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="pagusregalis" status="gen. et sp. nov.">
<emphasis id="F6EEEA94FFF0FFA36046F81A6CA04EE6" box="[300,663,1922,1945]" italics="true" pageId="64" pageNumber="65">Megastygonitocrella pagusregalis</emphasis>
</taxonomicName>
<emphasis id="F6EEEA94FFF0FFA363F7F81A6D744EE5" bold="true" box="[669,835,1922,1946]" pageId="64" pageNumber="65">
<taxonomicNameLabel id="EDDD57EFFFF0FFA363F7F81A6D744EE5" box="[669,835,1922,1946]" pageId="64" pageNumber="65" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype female: A—habitus, lateral view; B—habitus, dorsal view; C—urosome, ventral view; D—anal somite and left caudal ramus, lateral view; E—antennula; F—antenna; G—labrum; H—paragnaths; I—mandibula; J—maxillula; K—maxilla; L—maxilliped. Scales = 0.1 mm.
</paragraph>
</caption>
<subSubSection id="8C80650DFFF1FFA261AFFF0F6EC24AEF" pageId="65" pageNumber="66" type="discussion">
<paragraph id="C4253686FFF1FFA261AFFF0F6D8848BF" blockId="65.[151,1436,151,912]" pageId="65" pageNumber="66">
<emphasis id="F6EEEA94FFF1FFA261AFFF0F6F0849CE" bold="true" box="[197,319,151,177]" pageId="65" pageNumber="66">Remarks.</emphasis>
As mentioned in the Remarks section for 
<taxonomicName id="039A4D05FFF1FFA26256FF006A5449CE" authority="Karanovic &amp; Hancock, 2009" authorityName="Karanovic &amp; Hancock" authorityYear="2009" box="[828,1123,152,177]" class="Hexanauplia" family="Ameiridae" genus="Megastygonitocrella" kingdom="Animalia" order="Harpacticoida" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="dec" status="sp. nov.">
<emphasis id="F6EEEA94FFF1FFA26256FF006A5449CE" box="[828,1123,152,177]" italics="true" pageId="65" pageNumber="66">Megastygonitocrella dec</emphasis>
</taxonomicName>
<emphasis id="F6EEEA94FFF1FFA26501FF0F6AFC49CE" bold="true" box="[1131,1227,151,177]" pageId="65" pageNumber="66">
<taxonomicNameLabel id="EDDD57EFFFF1FFA26501FF0F6AFC49CE" box="[1131,1227,151,177]" pageId="65" pageNumber="66" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, 
<taxonomicName id="039A4D05FFF1FFA265B1FF006BAB49CE" authority="Karanovic &amp; Hancock, 2009" authorityName="Karanovic &amp; Hancock" authorityYear="2009" box="[1243,1436,152,177]" class="Hexanauplia" family="Ameiridae" genus="Megastygonitocrella" kingdom="Animalia" order="Harpacticoida" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="pagusregalis" status="sp. nov.">
<emphasis id="F6EEEA94FFF1FFA265B1FF006BAB49CE" box="[1243,1436,152,177]" italics="true" pageId="65" pageNumber="66">M. pagusregalis</emphasis>
</taxonomicName>
<emphasis id="F6EEEA94FFF1FFA261FDFF266EC249A7" bold="true" box="[151,245,190,216]" pageId="65" pageNumber="66">
<taxonomicNameLabel id="EDDD57EFFFF1FFA261FDFF266EC249A7" box="[151,245,190,216]" pageId="65" pageNumber="66" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
belongs to a group of very closely related congeners, all with the same segmentation and almost the same armature formula of the swimming legs: 
<taxonomicName id="039A4D05FFF1FFA263A8FF7D6A7E4980" authority="(Borutzky, 1967)" authorityName="Karanovic &amp; Hancock" authorityYear="2009" baseAuthorityName="Borutzky" baseAuthorityYear="1967" box="[706,1097,229,255]" class="Hexanauplia" family="Ameiridae" genus="Nitocrella" kingdom="Animalia" order="Harpacticoida" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="ljovuschkini" status="comb. nov.">
<emphasis id="F6EEEA94FFF1FFA263A8FF7D6D404981" box="[706,887,229,254]" italics="true" pageId="65" pageNumber="66">M. ljovuschkini</emphasis>
(
<bibRefCitation id="A00B4B77FFF1FFA262E2FF7D6A764980" author="Borutzky, E. V." box="[904,1089,229,255]" pageId="65" pageNumber="66" pagination="32 - 39" refId="ref55876" refString="Borutzky, E. V. (1967) Biospeologica Sovietica 32, On the genus Nitocrella Chappuis (Copepoda, Harpacticoida). Byulleten Moskovskogo Obschestva Ispytatelei Prirody (nov. ser.), 72, 32 - 39. [in Russian]" type="journal article" year="1967">Borutzky, 1967</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="EDDD57EFFFF1FFA2653BFF7D6AE14980" box="[1105,1238,229,255]" pageId="65" pageNumber="66" rank="species">comb. nov.</taxonomicNameLabel>
, 
<taxonomicName id="039A4D05FFF1FFA26589FF7D6B474981" authority="Karanovic &amp; Hancock, 2009" authorityName="Karanovic &amp; Hancock" authorityYear="2009" box="[1251,1392,229,254]" class="Hexanauplia" family="Ameiridae" genus="Megastygonitocrella" kingdom="Animalia" order="Harpacticoida" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="ecowisei" status="sp. nov.">
<emphasis id="F6EEEA94FFF1FFA26589FF7D6B474981" box="[1251,1392,229,254]" italics="true" pageId="65" pageNumber="66">M. ecowisei</emphasis>
</taxonomicName>
<emphasis id="F6EEEA94FFF1FFA26413FF7D6EFB485A" bold="true" pageId="65" pageNumber="66">
<taxonomicNameLabel id="EDDD57EFFFF1FFA26413FF7D6EFB485A" pageId="65" pageNumber="66" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, 
<taxonomicName id="039A4D05FFF1FFA261B5FE946F03485A" authority="Karanovic &amp; Hancock, 2009" authorityName="Karanovic &amp; Hancock" authorityYear="2009" box="[223,308,268,293]" class="Hexanauplia" family="Ameiridae" genus="Megastygonitocrella" kingdom="Animalia" order="Harpacticoida" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="dec" status="sp. nov.">
<emphasis id="F6EEEA94FFF1FFA261B5FE946F03485A" box="[223,308,268,293]" italics="true" pageId="65" pageNumber="66">M. dec</emphasis>
</taxonomicName>
<emphasis id="F6EEEA94FFF1FFA26054FE936F95485A" bold="true" box="[318,418,267,293]" pageId="65" pageNumber="66">
<taxonomicNameLabel id="EDDD57EFFFF1FFA26054FE936F95485A" box="[318,418,267,293]" pageId="65" pageNumber="66" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, 
<taxonomicName id="039A4D05FFF1FFA260DFFE946C0C485A" authority="Karanovic &amp; Hancock, 2009" authorityName="Karanovic &amp; Hancock" authorityYear="2009" box="[437,571,268,293]" class="Hexanauplia" family="Ameiridae" genus="Megastygonitocrella" kingdom="Animalia" order="Harpacticoida" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="kryptos" status="sp. nov.">
<emphasis id="F6EEEA94FFF1FFA260DFFE946C0C485A" box="[437,571,268,293]" italics="true" pageId="65" pageNumber="66">M. kryptos</emphasis>
</taxonomicName>
<emphasis id="F6EEEA94FFF1FFA2632FFE936C9E485A" bold="true" box="[581,681,267,293]" pageId="65" pageNumber="66">
<taxonomicNameLabel id="EDDD57EFFFF1FFA2632FFE936C9E485A" box="[581,681,267,293]" pageId="65" pageNumber="66" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, 
<taxonomicName id="039A4D05FFF1FFA263D6FE946A644859" authority="(Karanovic, 2006)" authorityName="Karanovic &amp; Hancock" authorityYear="2009" baseAuthorityName="Karanovic" baseAuthorityYear="2006" box="[700,1107,268,294]" class="Hexanauplia" family="Ameiridae" genus="Megastygonitocrella" kingdom="Animalia" order="Harpacticoida" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="trispinosa" status="comb. nov.">
<emphasis id="F6EEEA94FFF1FFA263D6FE946D54485A" box="[700,867,268,293]" italics="true" pageId="65" pageNumber="66">M. trispinosa</emphasis>
(
<bibRefCitation id="A00B4B77FFF1FFA2621CFE946A7D4859" author="Karanovic, T." box="[886,1098,268,294]" pageId="65" pageNumber="66" pagination="1 - 239" refId="ref57102" refString="Karanovic, T. (2006) Subterranean copepods (Crustacea, Copepoda) from the Pilbara region in Western Australia. Records of the Western Australian Museum, Supplement 70, 1 - 239." type="journal article" year="2006">Karanovic, 2006</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="EDDD57EFFFF1FFA26537FE946ADC4859" box="[1117,1259,268,294]" pageId="65" pageNumber="66" rank="species">comb. nov.</taxonomicNameLabel>
, 
<taxonomicName id="039A4D05FFF1FFA26590FE946F4A4833" authority="(Karanovic, 2006)" authorityName="Karanovic &amp; Hancock" authorityYear="2009" baseAuthorityName="Karanovic" baseAuthorityYear="2006" class="Hexanauplia" family="Ameiridae" genus="Stygonitocrella" kingdom="Animalia" order="Harpacticoida" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="bispinosa" status="comb. nov.">
<emphasis id="F6EEEA94FFF1FFA26590FE946BAB485A" box="[1274,1436,268,293]" italics="true" pageId="65" pageNumber="66">M. bispinosa</emphasis>
(
<bibRefCitation id="A00B4B77FFF1FFA261F5FEAA6F414833" author="Karanovic, T." box="[159,374,306,332]" pageId="65" pageNumber="66" pagination="1 - 239" refId="ref57102" refString="Karanovic, T. (2006) Subterranean copepods (Crustacea, Copepoda) from the Pilbara region in Western Australia. Records of the Western Australian Museum, Supplement 70, 1 - 239." type="journal article" year="2006">Karanovic, 2006</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="EDDD57EFFFF1FFA260E2FEAA6C244833" box="[392,531,306,332]" pageId="65" pageNumber="66" rank="species">comb. nov.</taxonomicNameLabel>
and 
<taxonomicName id="039A4D05FFF1FFA26339FEAA6DC04833" authority="(Karanovic, 2006)" authorityName="Karanovic &amp; Hancock" authorityYear="2009" baseAuthorityName="Karanovic" baseAuthorityYear="2006" box="[595,1015,306,332]" class="Hexanauplia" family="Ameiridae" genus="Stygonitocrella" kingdom="Animalia" order="Harpacticoida" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="unispinosa" status="comb. nov.">
<emphasis id="F6EEEA94FFF1FFA26339FEAA6D314834" box="[595,774,306,331]" italics="true" pageId="65" pageNumber="66">M. unispinosa</emphasis>
(
<bibRefCitation id="A00B4B77FFF1FFA26273FEAA6DD84833" author="Karanovic, T." box="[793,1007,306,332]" pageId="65" pageNumber="66" pagination="1 - 239" refId="ref57102" refString="Karanovic, T. (2006) Subterranean copepods (Crustacea, Copepoda) from the Pilbara region in Western Australia. Records of the Western Australian Museum, Supplement 70, 1 - 239." type="journal article" year="2006">Karanovic, 2006</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="EDDD57EFFFF1FFA2656BFEAA6ABB4833" box="[1025,1164,306,332]" pageId="65" pageNumber="66" rank="species">comb. nov.</taxonomicNameLabel>
However, it is easily distinguished from all these species by its unusually short antennula and the fifth leg completely fused to the somite. It differs from each species individually by many more characters and we will avoid repeating those differences already stressed in the Remarks sections of other species.
</paragraph>
<paragraph id="C4253686FFF1FFA261AFFE556EC24AEF" blockId="65.[151,1436,151,912]" pageId="65" pageNumber="66">
<taxonomicName id="039A4D05FFF1FFA261AFFE556C444899" authorityName="Karanovic &amp; Hancock" authorityYear="2009" box="[197,627,461,486]" class="Hexanauplia" family="Ameiridae" genus="Megastygonitocrella" kingdom="Animalia" order="Harpacticoida" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="pagusregalis">
<emphasis id="F6EEEA94FFF1FFA261AFFE556C444899" box="[197,627,461,486]" italics="true" pageId="65" pageNumber="66">Megastygonitocrella pagusregalis</emphasis>
</taxonomicName>
is probably most closely related to the only other species from 
<collectingRegion id="065EF864FFF1FFA261FDFE6C6F104B71" box="[151,295,500,526]" country="Australia" name="Queensland" pageId="65" pageNumber="66">Queensland</collectingRegion>
, 
<taxonomicName id="039A4D05FFF1FFA2605EFE6C6F834B72" authority="Karanovic &amp; Hancock, 2009" authorityName="Karanovic &amp; Hancock" authorityYear="2009" box="[308,436,500,525]" class="Hexanauplia" family="Ameiridae" genus="Megastygonitocrella" kingdom="Animalia" order="Harpacticoida" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="kryptos" status="sp. nov.">
<emphasis id="F6EEEA94FFF1FFA2605EFE6C6F834B72" box="[308,436,500,525]" italics="true" pageId="65" pageNumber="66">M. kryptos</emphasis>
</taxonomicName>
<emphasis id="F6EEEA94FFF1FFA260D6FE6B6C2B4B72" bold="true" box="[444,540,499,525]" pageId="65" pageNumber="66">
<taxonomicNameLabel id="EDDD57EFFFF1FFA260D6FE6B6C2B4B72" box="[444,540,499,525]" pageId="65" pageNumber="66" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, and it is unfortunate that the former species is known only from females, while the latter is known only from males, so some characters cannot be compared. Actually, in our preliminary identification of the 
<collectingRegion id="065EF864FFF1FFA2637DFDD96C954B24" box="[535,674,577,603]" country="Australia" name="Queensland" pageId="65" pageNumber="66">Queensland</collectingRegion>
material, we entertained for a short time the possibility that they belonged to the same species because of their very similar habitus and swimming legs. However, differences in the ornamentation of the urosomites and anal operculum as well as the shape of the anal operculum and caudal rami all point towards their separate specific status. Also, the fifth leg is not fused to the somite in 
<emphasis id="F6EEEA94FFF1FFA26416FD2D6EC04B8A" italics="true" pageId="65" pageNumber="66">
<taxonomicName id="039A4D05FFF1FFA26416FD2D6EC54B8A" authorityName="Karanovic &amp; Hancock" authorityYear="2009" class="Hexanauplia" family="Ameiridae" genus="Megastygonitocrella" kingdom="Animalia" order="Harpacticoida" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="kryptos">M. kryptos</taxonomicName>
,
</emphasis>
although the two species have homologous armature elements on this appendage (that of male 
<taxonomicName id="039A4D05FFF1FFA26416FD446ED94A64" authorityName="Karanovic &amp; Hancock" authorityYear="2009" class="Hexanauplia" family="Ameiridae" genus="Megastygonitocrella" kingdom="Animalia" order="Harpacticoida" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="kryptos">
<emphasis id="F6EEEA94FFF1FFA26416FD446ED94A64" italics="true" pageId="65" pageNumber="66">M. kryptos</emphasis>
</taxonomicName>
with an additional innermost seta as a normal form of sexual dimorphism in this group of freshwater ameirids). The third exopodal segment of the fouth swimming leg carries one inner seta in 
<taxonomicName id="039A4D05FFF1FFA2658DFCB16B5F4A3D" authorityName="Karanovic &amp; Hancock" authorityYear="2009" box="[1255,1384,809,834]" class="Hexanauplia" family="Ameiridae" genus="Megastygonitocrella" kingdom="Animalia" order="Harpacticoida" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="kryptos">
<emphasis id="F6EEEA94FFF1FFA2658DFCB16B5F4A3D" box="[1255,1384,809,834]" italics="true" pageId="65" pageNumber="66">M. kryptos</emphasis>
</taxonomicName>
and none in 
<taxonomicName id="039A4D05FFF1FFA26192FCC86F8D4A16" authorityName="Karanovic &amp; Hancock" authorityYear="2009" box="[248,442,848,873]" class="Hexanauplia" family="Ameiridae" genus="Megastygonitocrella" kingdom="Animalia" order="Harpacticoida" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="pagusregalis">
<emphasis id="F6EEEA94FFF1FFA26192FCC86F8D4A16" box="[248,442,848,873]" italics="true" pageId="65" pageNumber="66">M. pagusregalis</emphasis>
</taxonomicName>
, but this character is known to be variable in some species (see, for example, 
<figureCitation id="5CA12A03FFF1FFA26402FCC86EDF4AEF" captionStart="FIGURE 11" captionStartId="51.[151,257,1836,1860]" captionTargetBox="[234,1376,203,1798]" captionTargetId="figure-103@51.[180,1414,170,1873]" captionTargetPageId="51" captionText="FIGURE 11. Megastygonitocrella dec gen. et sp. nov., A–G, holotype female; H–Q, allotype male: A—left caudal ramus, lateral view; B—first swimming leg; C—endopod of second swimming leg; D—third swimming leg; E—third exopodal segment of right fourth leg; F—left fourth swimming leg; G—right fifth leg; H—habitus, dorsal view; I— antennula; J—exopod of antenna; K—basis of first swimming leg; L—endopod of second swimming leg; M—endopod of third swimming leg; N—endopod of fourth swimming leg; O—right fifth leg; P—left fifth leg; Q—left sixth leg. Scales = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5313068" httpUri="https://zenodo.org/record/5313068/files/figure.png" pageId="65" pageNumber="66">Figs 11E–F</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>