131 lines
16 KiB
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131 lines
16 KiB
XML
<document id="6AA667DFD8EBF366A433B3F9D534D41D" ID-CLB-Dataset="33935" ID-DOI="http://dx.doi.org/10.3897/zookeys.131.1921" ID-GBIF-Dataset="72a81528-63ef-4531-9f9c-ade551a651d8" ID-PMC="PMC3208436" ID-Pensoft-Pub="1313-2970-131-1" ID-PubMed="22140329" ModsDocAuthor="" ModsDocDate="2011" ModsDocID="1313-2970-131-1" ModsDocOrigin="ZooKeys 131" ModsDocTitle="Revision of the Southeast Asian millipede genus Orthomorpha Bollman, 1893, with the proposal of a new genus (Diplopoda, Polydesmida, Paradoxosomatidae)" checkinTime="1451249839150" checkinUser="pensoft" docAuthor="Likhitrakarn, Natdanai, Golovatch, Sergei I. & Panha, Somsak" docDate="2011" docId="3E1E41D8DEC4B45A86478202FA560A06" docLanguage="en" docName="ZooKeys 131: 1-161" docOrigin="ZooKeys 131" docSource="http://dx.doi.org/10.3897/zookeys.131.1921" docTitle="Orthomorpha cambodjana Attems 1953" docType="treatment" docVersion="8" lastPageNumber="57" masterDocId="FFFF0F54FF88BD1C371C965D22453928" masterDocTitle="Revision of the Southeast Asian millipede genus Orthomorpha Bollman, 1893, with the proposal of a new genus (Diplopoda, Polydesmida, Paradoxosomatidae)" masterLastPageNumber="161" masterPageNumber="1" pageNumber="54" updateTime="1732923565725" updateUser="ExternalLinkService">
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<mods:title id="71973CF7C8FFAA68E2C870A46D75D054">Revision of the Southeast Asian millipede genus Orthomorpha Bollman, 1893, with the proposal of a new genus (Diplopoda, Polydesmida, Paradoxosomatidae)</mods:title>
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<mods:namePart id="2A9EC72EE93B4402645FADF4A705037D">Likhitrakarn, Natdanai</mods:namePart>
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<mods:namePart id="9A7BC6C1534C5E2998CCFF44C1AC885C">Golovatch, Sergei I.</mods:namePart>
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<mods:namePart id="43C7652D440116FDDDE0106113437156">Panha, Somsak</mods:namePart>
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<mods:title id="A8A37AE0AB465FF9AB223BFC10FCC1C4">ZooKeys</mods:title>
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<mods:date id="B320571F7D62D2CBAF51946314C3180B">2011</mods:date>
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<mods:detail id="15B508F6CC4F1698DF6C4AA7DD0ABE3D" type="volume">
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<mods:number id="BAC63F81843EDD02B876BE03CF1CEC1B">131</mods:number>
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<mods:url id="582743A99FDD217C5030D4659B0CD528">http://dx.doi.org/10.3897/zookeys.131.1921</mods:url>
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<mods:classification id="03BCB142AFFD29B23570C81F79EC8829">journal article</mods:classification>
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<mods:identifier id="7DC078483D5F71251E64746D0E4766D5" type="DOI">http://dx.doi.org/10.3897/zookeys.131.1921</mods:identifier>
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<treatment id="3E1E41D8DEC4B45A86478202FA560A06" ID-DOI="http://doi.org/10.5281/zenodo.3500878" ID-GBIF-Taxon="152031141" ID-Zenodo-Dep="3500878" LSID="urn:lsid:plazi:treatment:3E1E41D8DEC4B45A86478202FA560A06" httpUri="http://treatment.plazi.org/id/3E1E41D8DEC4B45A86478202FA560A06" lastPageId="56" lastPageNumber="57" pageId="53" pageNumber="54" scope_class="Diplopoda" scope_family="Paradoxosomatidae" scope_order="Polydesmida">
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<subSubSection id="3BDAAE7141A19F82EC85E679AAF9AF69" pageId="53" pageNumber="54" type="nomenclature">
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<paragraph id="7B53481ECD089BB235BF3C7A85940151" pageId="53" pageNumber="54">
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<taxonomicName id="5BFBFDFEAFAB79E7449E1392CB18BAE1" ID-CoL="6T8P5" LSID="http://species-id.net/wiki/Orthomorpha_cambodjana" authority="Attems, 1953" authorityName="Attems" authorityYear="1953" class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha cambodjana" order="Polydesmida" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="cambodjana">Orthomorpha cambodjana (Attems, 1953)</taxonomicName>
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Figs 4748
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</paragraph>
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</subSubSection>
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<subSubSection id="76EADF0B700983AAF39976CD7DF5E9B1" pageId="53" pageNumber="54" type="reference_group">
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<paragraph id="281BC04270985A8909F4A30477A73E96" pageId="53" pageNumber="54">
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<taxonomicName id="E6C907F66F23E3A2FECF403C4D8EA21D" class="Diplopoda" family="Paradoxosomatidae" genus="Pratinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pratinus cambodjanus" order="Polydesmida" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="cambodjanus">Pratinus cambodjanus</taxonomicName>
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<bibRefCitation id="B8B79E362C35F034CC0F8079EB0B685B" author="Attems, C" journalOrPublisher="Memoires du Museum national d'Histoire naturelle, Serie A, Zoologie" pageId="114" pageNumber="115" pagination="133 - 230" title="Myriopoden von Indochina. Expedition von Dr. C. Dawydoff (1938 - 1939)." volume="5" year="1953">Attems 1953</bibRefCitation>
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: 168 (D).
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</paragraph>
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<paragraph id="B23A540158FD221FC971C00858A4FD5E" pageId="53" pageNumber="54">
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<taxonomicName id="D8664359B0BEA9430C8E07F616E94194" class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha cambodjana" order="Polydesmida" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="cambodjana">Orthomorpha cambodjana</taxonomicName>
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-
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<bibRefCitation id="202497698A6C56371054672644171206" author="Jeekel, CAW" journalOrPublisher="Tijdschrift voor Entomologie" pageId="114" pageNumber="115" pagination="205 - 283" title="Paradoxosomatidae from Borneo (Diplopoda, Polydesmida)." volume="106" year="1963">Jeekel 1963</bibRefCitation>
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: 265 (M);
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<bibRefCitation id="AF48E9C15788FF4868ECEEF1A161996B" author="Jeekel, CAW" journalOrPublisher="Tijdschrift voor Entomologie" pageId="114" pageNumber="115" pagination="355 - 364" title="A new species of Orthomorpha Bollman from Thailand observed in migration, with taxonomic notes on the genus (Diplopoda)." volume="107" year="1964">1964</bibRefCitation>
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: 361 (M, D);
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<bibRefCitation id="C498A19DC52974DB891E6061CCFA470D" author="Jeekel, CAW" journalOrPublisher="Academisch Proefschrift, Rotterdam" pageId="114" pageNumber="115" title="On the classification and geographical distribution of the family Paradoxosomatidae (Diplopoda, Polydesmida)." year="1968">1968</bibRefCitation>
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: 56 (M);
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<bibRefCitation id="EBC1025174358219DE24D6F745CA8880" author="Hoffman, RL" journalOrPublisher="Revue suisse de Zoologie" pageId="114" pageNumber="115" pagination="699 - 719" title="Diplopoda from Malayan caves, collected by M. Pierre Strinati." volume="84" year="1977">Hoffman 1977</bibRefCitation>
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: 700 (M);
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<bibRefCitation id="6245D5B47DDDB6F1C6F0FE120DDBA32F" pageId="53" pageNumber="54">Golovatch 1998</bibRefCitation>
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: 42 (M, D).
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</paragraph>
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</subSubSection>
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<subSubSection id="D260DFB3393AF86E2BE79DE0007A4ABF" pageId="53" pageNumber="54" type="lectotype">
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<paragraph id="C8D987E9F72D1467BC118CE1310BD141" pageId="53" pageNumber="54">Lectotype.</paragraph>
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<paragraph id="8ACD6653A96AD57814BC530BE7910E3E" pageId="53" pageNumber="54">♂ (NHMW-3501), Cambodia, Kampot Prov., Ream, Sre-Umbell, 1938-39, leg. C. Dawydoff.</paragraph>
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</subSubSection>
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<subSubSection id="306FA015A9BF545758950B750FEB598F" pageId="53" pageNumber="54" type="paralectotype">
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<paragraph id="29B341551684837BFB31519B30C23715" pageId="53" pageNumber="54">Paralectotype.</paragraph>
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<paragraph id="91FA0128794A3219C60C47373F329036" pageId="53" pageNumber="54">1 ♀ (NHMW-3501), same locality, together with lectotype.</paragraph>
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<paragraph id="F7A71C95ACB7036D5C00FFADE41E983A" pageId="53" pageNumber="54">Lectotype designation proposed herewith is necessary to ensure the species is based on a complete male.</paragraph>
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</subSubSection>
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<subSubSection id="7CAE35A6369A1762C028DE07083DB879" lastPageId="56" lastPageNumber="57" pageId="53" pageNumber="54" type="redescription">
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<paragraph id="7514639A8B316187A53DB291BBDC2158" pageId="53" pageNumber="54">Redescription.</paragraph>
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<paragraph id="E867E11DEEA0EB5B8A67A4F8CF51DD22" pageId="53" pageNumber="54">
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Length ca 31 mm (♂), 32 mm (♀), width of midbody pro- and metazona 2.2 and 3.4 mm (♂), 3.1 and 4.3 mm (♀), respectively (vs 3.2-4.0 mm in width, as given in the original description (
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<bibRefCitation id="9B250599E9A57EE8EA7D380AB12581AB" author="Attems, C" journalOrPublisher="Memoires du Museum national d'Histoire naturelle, Serie A, Zoologie" pageId="114" pageNumber="115" pagination="133 - 230" title="Myriopoden von Indochina. Expedition von Dr. C. Dawydoff (1938 - 1939)." volume="5" year="1953">Attems 1953</bibRefCitation>
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)). Coloration of alcohol material upon long-term preservation rather uniformly brown (Fig. 47) (vs dark castaneous with venter, legs and paraterga yellow-brown, and antennae light brown, as given in the original description (
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<bibRefCitation id="851A76A0867676BC02C0F40766EEDDD6" author="Attems, C" journalOrPublisher="Memoires du Museum national d'Histoire naturelle, Serie A, Zoologie" pageId="114" pageNumber="115" pagination="133 - 230" title="Myriopoden von Indochina. Expedition von Dr. C. Dawydoff (1938 - 1939)." volume="5" year="1953">Attems 1953</bibRefCitation>
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)).
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</paragraph>
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<paragraph id="D9AFBE65CAD9B029F6028483B5E03FDB" lastPageId="56" lastPageNumber="57" pageId="54" pageNumber="55">
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<pageBreakToken id="0736B445246EDCE6514A56DBD0113173" pageId="54" pageNumber="55" start="start">Head</pageBreakToken>
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usual, clypeolabral region densely setose, surface of vertex smooth, with 1+1 setae flanking a distinct epicranial suture. Antennae rather long and slender (Fig. 47B & J), extending behind segment 3 (♂, ♀) dorsally. Head in width <collum <segments 3 and 4 <segment 2 <segments 5-16 (♂, ♀), gently and gradually tapering thereafter. Collum smooth, with three transverse rows of setae, 4+4 anterior, 2+2 intermediate, and 3+3 posterior setae; caudal corner of paraterga acutangular (ca 70°), nearly pointed (Fig. 47A & J). Tegument smooth and shining; metaterga very faintly rugulose, each postcollum one with two rows of mostly abraded, medium-sized setae: 3+3 in front row and 3+3 or, on segments 12-19, 4+4 in caudal row; prozona very finely shagreened,
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<pageBreakToken id="521A6470BC2CB0FA36E406481B75C6D1" pageId="55" pageNumber="56" start="start">surface</pageBreakToken>
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below paraterga finely microgranulate. Axial line faint, but visible, starting from collum. Paraterga very strongly developed (Fig. 47A-G & J-L), set high (at ca 1/4 metazonital height), mostly slightly upturned, all lying below dorsum, thin in lateral view, like blunt blades, a little thicker only on pore-bearing segments, on postcollum segments extending increasingly beyond rear tergal margin, nearly pointed to pointed, caudal tip on paraterga 16-19 evidently curved mesad. Calluses on paraterga 2 delimited by a sulcus only dorsally, following paraterga by a sulcus both dorsally and ventrally, rather broad. Paraterga 2 broad, anterior edge angulate, lateral edge with three small incisions in anterior half; posterior edge evidently concave (Fig. 47A, B & J). Paraterga 3 and 4 subequal, like subsequent paraterga, anterior edge broadly rounded, bordered and fused to callus, lateral edge with two small incisions. Ozopores evident, lateral, lying inside an ovoid groove at about 1/4 metazonital length. Transverse sulcus complete on metaterga 5-18, incomplete on metatergum 4, shallow, not reaching bases of paraterga, beaded at bottom, slightly sinuate anteromedially (Fig. 47A, C, F
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<pageBreakToken id="F1166399E618A9FFB4E82279E784DD69" pageId="56" pageNumber="57" start="start">&</pageBreakToken>
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amp; J-L). Stricture between pro- and metazona narrow, deep, ribbed at bottom down to base of paraterga. Pleurosternal carinae complete crests only on segments 2-4 (Fig. 47B, D & E), each with an evident sharp denticle caudally, thereafter increasingly strongly reduced until segment 14 (♂) or 11 (♀). Epiproct (Fig. 47E-G & L) conical, flattened dorsoventrally, apical papillae small, dentiform, directed caudoventrally; tip subtruncate; pre-apical papillae small, lying close to tip. Hypoproct (Fig. 47G) roundly subtrapeziform, setiferous knobs at caudal margin large and well-separated.
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</paragraph>
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<paragraph id="CF4DBBC322256CBB0C3A1DC7269FE798" pageId="56" pageNumber="57">Sterna sparsely setose, without modifications, but with two large, rounded, fully separated, setose cones between ♂ coxae 4 (Fig. 47H & I). A paramedian pair of small tubercles in front of gonopod aperture. Legs long and slender, midbody ones ca 1.2-1.3 (♂) or 0.8-0.9 (♀) as long as body height, prefemora without modifications, tarsal brushes present until ♂ legs 5.</paragraph>
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<paragraph id="5967DF1D6351F607ECF2700080242213" pageId="56" pageNumber="57">
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Gonopods (Fig. 48) simple. Coxa long and slender, with several setae distodorsally. Prefemur densely setose, nearly 3 times shorter than femorite +
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<normalizedToken id="33D7C2C37A7D9292E65548C83AF7BD8E" originalValue="“postfemoral”">"postfemoral"</normalizedToken>
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part. Femorite very slender, slightly curved, very slightly enlarged distad,
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<normalizedToken id="F3AE7CCE7D3FC3398EB249A2D53C6C80" originalValue="“postfemoral”">"postfemoral"</normalizedToken>
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part demarcated by an oblique lateral sulcus; tip of solenophore small, trifid, with two denticles (terminal tooth larger, middle one smaller) and a small subterminal lobule.
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</paragraph>
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<caption id="FA954DF791F61C15ED473AC150325033" pageId="56" pageNumber="57">
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<paragraph id="2E91DCC27E481CF8BFE0FC656A2B5C11" pageId="56" pageNumber="57">
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Figure 47.
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<taxonomicName id="4D91C731B80D3796EEA21E8BB2E37777" class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha cambodijana" order="Polydesmida" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="cambodijana">Orthomorpha cambodijana</taxonomicName>
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(Attems, 1953), ♂ lectotype (
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<normalizedToken id="1B3632ACC594EE5A82514BC94B6CCB36" originalValue="A–I">A-I</normalizedToken>
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), ♀ paralectotype (
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<normalizedToken id="2938E4570A9A82A9761B97FA12352F8C" originalValue="J–L">J-L</normalizedToken>
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). A, B, J anterior part of body, dorsal, lateral and dorsal views, respectively C, K segments 10 and 11, dorsal view D segments 11 and 12 lateral view
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<normalizedToken id="11AB8CBC68F35DCF7F4523A56FC072AD" originalValue="E–F">E-F</normalizedToken>
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, L posterior part of body, lateral, dorsal, ventral and dorsal views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively.
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</paragraph>
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</caption>
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<caption id="7726FD5F00898BDB1EF06D2AE7809AA3" pageId="56" pageNumber="57">
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<paragraph id="411C58194B99BF4B245EC34326177DE4" pageId="56" pageNumber="57">
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Figure 48.
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<taxonomicName id="0F45A12D850922E566D87A1A0CF76B03" class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha cambodijana" order="Polydesmida" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="cambodijana">Orthomorpha cambodijana</taxonomicName>
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(Attems, 1953), ♂ lectotype. A, B right gonopod, lateral and mesal views, respectively.
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</paragraph>
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</caption>
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</subSubSection>
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</treatment>
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</document> |