treatments-xml/data/0C/30/22/0C30227A9311D12CCAB5FDD4FA673F67.xml
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<document id="4894DD0B7F95772121B6134C109222E7" ID-DOI="10.11646/zootaxa.5346.3.5" ID-ISSN="1175-5326" ID-Zenodo-Dep="8390212" ID-ZooBank="BAAC4494-8FD4-4740-AD15-45A56B612421" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1695984437868" checkinUser="plazi" docAuthor="Yin, Zi-Wei &amp; Zeng, Ting-Kai" docDate="2023" docId="0C30227A9311D12CCAB5FDD4FA673F67" docLanguage="en" docName="zootaxa.5346.3.5.pdf" docOrigin="Zootaxa 5346 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5346.3.5" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Batrisiella riparia Yin &amp; Zeng 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="320" masterDocId="F0095A029310D12FCA22FF92FFE7396E" masterDocTitle="Two new riparian Pselaphinae from Hubei, Central China (Coleoptera: taphylinidae)" masterLastPageNumber="324" masterPageNumber="317" pageNumber="318" updateTime="1695984812321" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="55A205FF53F690FD6E96A41B09487414">Two new riparian Pselaphinae from Hubei, Central China (Coleoptera: taphylinidae)</mods:title>
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<mods:namePart id="3C35CD7BAC5CE3FFC61CC1C883BBAC67">Yin, Zi-Wei</mods:namePart>
<mods:affiliation id="F32B8CD0F43BC23FD2BEB94A349994B6">Laboratory of Systematic Entomology, College of Life Sciences, Shanghai Normal University, Shanghai 200234, China.</mods:affiliation>
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<mods:namePart id="89B212476D8DE7F43AEA3E7DEEF508F7">Zeng, Ting-Kai</mods:namePart>
<mods:nameIdentifier id="640B653897BEFDF80E91A6F62A95B4F2" type="ORCID">0009-0009-0790-5017</mods:nameIdentifier>
<mods:affiliation id="CC0A87595119C2A654333DA75F742C4F">Chongyang No. 2 Senior High School, Baini Town, Chongyang County, Xianning 437515, Hubei, China. https: // orcid. org / 0009 - 0009 - 0790 - 5017</mods:affiliation>
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<mods:date id="327424C74DB3687FB653A64645D6DFEE">2023</mods:date>
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<treatment id="0C30227A9311D12CCAB5FDD4FA673F67" LSID="urn:lsid:plazi:treatment:0C30227A9311D12CCAB5FDD4FA673F67" httpUri="http://treatment.plazi.org/id/0C30227A9311D12CCAB5FDD4FA673F67" lastPageId="3" lastPageNumber="320" pageId="1" pageNumber="318">
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<paragraph id="8426936C9311D12ECAB5FDD4FD923B0F" blockId="1.[151,629,582,645]" box="[151,629,582,609]" pageId="1" pageNumber="318">
<heading id="DF6E24009311D12ECAB5FDD4FD923B0F" bold="true" box="[151,629,582,609]" fontSize="11" level="1" pageId="1" pageNumber="318" reason="1">
<emphasis id="B6ED4F7E9311D12ECAB5FDD4FD923B0F" bold="true" box="[151,629,582,609]" pageId="1" pageNumber="318">
<taxonomicName id="4399E8EF9311D12ECAB5FDD4FDED3B0F" authority="Yin &amp; Zeng" authorityName="Yin &amp; Zeng" authorityYear="2023" box="[151,522,582,609]" class="Insecta" family="Staphylinidae" genus="Batrisiella" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="318" phylum="Arthropoda" rank="species" species="riparia" status="sp. nov.">
<emphasis id="B6ED4F7E9311D12ECAB5FDD4FE883B0F" bold="true" box="[151,367,582,609]" italics="true" pageId="1" pageNumber="318">Batrisiella riparia</emphasis>
Yin &amp; Zeng
</taxonomicName>
,
<taxonomicNameLabel id="ADDEF2059311D12EC835FDD5FD923B0F" box="[535,629,583,609]" pageId="1" pageNumber="318" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="CC83C0E79311D12ECAB5FDF9FD963BA5" pageId="1" pageNumber="318" type="description">
<paragraph id="8426936C9311D12ECAB5FDF9FEEC3BEB" blockId="1.[151,629,582,645]" box="[151,267,619,645]" pageId="1" pageNumber="318">
(
<figureCitation id="1CA28FE99311D12ECABDFDF9FF013BEB" box="[159,230,619,645]" captionStart="FIGURE 1" captionStartId="2.[151,248,1734,1758]" captionTargetBox="[167,1413,151,1709]" captionTargetId="figure-150@2.[166,1413,151,1711]" captionTargetPageId="2" captionText="FIGURE 1. Habitus and morphological details of Batrisiella riparia sp. nov., male. A. Dorsal habitus. B. Head and pronotum. C. Scape, arrow indicating lamellar structure at apicolateral corner. D. Metafemur. E. Sternite 7 (IX). FH. Aedeagus, lateral (F), ventral (G), and axial (H). I. Female genital complex. Scale bars: 0.5 mm in A; 0.2 mm in B, D; 0.05 mm in C, E; 0.1 mm in FI." figureDoi="http://doi.org/10.5281/zenodo.8390216" httpUri="https://zenodo.org/record/8390216/files/figure.png" pageId="1" pageNumber="318">Figs 1</figureCitation>
,
<figureCitation id="1CA28FE99311D12ECAD6FDF9FEE53BEB" box="[244,258,619,645]" captionStart="FIGURE 3" captionStartId="6.[151,250,1628,1652]" captionTargetBox="[166,1413,454,1588]" captionTargetId="figure-173@6.[166,1413,454,1589]" captionTargetPageId="6" captionText="FIGURE 3. Collecting environment and habitat of Batrisiella riparia sp. nov. and Nipponobythus varicornis sp. nov." figureDoi="http://doi.org/10.5281/zenodo.8390222" httpUri="https://zenodo.org/record/8390222/files/figure.png" pageId="1" pageNumber="318">3</figureCitation>
)
</paragraph>
<paragraph id="8426936C9311D12ECAB5FD21FD963BA5" blockId="1.[151,625,691,717]" box="[151,625,691,717]" pageId="1" pageNumber="318">
Chinese common name:
<emphasis id="B6ED4F7E9311D12ECB8FFD21FD963BA5" bold="true" box="[429,625,691,715]" pageId="1" pageNumber="318">DZḳṁŧDzfim</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="CC83C0E79311D12ECAB5FD69FA833A33" pageId="1" pageNumber="318" type="materials_examined">
<paragraph id="8426936C9311D12ECAB5FD69FA833A33" blockId="1.[151,1437,762,2013]" pageId="1" pageNumber="318">
<emphasis id="B6ED4F7E9311D12ECAB5FD69FED83A7A" bold="true" box="[151,319,762,789]" pageId="1" pageNumber="318">Type material</emphasis>
(67 exx.).
<materialsCitation id="34F199319311D12ECB9DFD69FE493A33" collectingDate="2023-07-16" collectingDateMax="2023-07-17" collectingDateMin="2023-07-16" collectionCode="SNUC" collectorName="Ting-Kai Zeng, AE" country="China" county="Chongyang Co." elevation="165" latitude="29.30611" location="Xianning City" longLatPrecision="20" longitude="114.17305" pageId="1" pageNumber="318" specimenCount="1" stateProvince="Hubei" typeStatus="holotype">
<emphasis id="B6ED4F7E9311D12ECB9DFD69FD433A7A" bold="true" box="[447,676,762,789]" pageId="1" pageNumber="318">
<typeStatus id="5B222DCE9311D12ECB9DFD69FDD73A7B" box="[447,560,763,789]" pageId="1" pageNumber="318">Holotype</typeStatus>
:
<collectingCountry id="FC8ED3FC9311D12EC81FFD68FD783A7A" box="[573,671,762,788]" name="China" pageId="1" pageNumber="318">CHINA</collectingCountry>
:
</emphasis>
J, China:
<collectingRegion id="465D5D8E9311D12EC90CFD69FC923A7B" box="[814,885,763,789]" country="China" name="Hubei" pageId="1" pageNumber="318">Hubei</collectingRegion>
,
<location id="8146C5B79311D12EC9A0FD69FBC33A7B" LSID="urn:lsid:plazi:treatment:0C30227A9311D12CCAB5FDD4FA673F67:8146C5B79311D12EC9A0FD69FBC33A7B" box="[898,1060,763,789]" country="China" county="Chongyang Co." latitude="29.30611" longLatPrecision="20" longitude="114.17305" name="Xianning City" pageId="1" pageNumber="318" stateProvince="Hubei">Xianning City</location>
,
<collectingCounty id="6D47EBE09311D12ECE13FD69FB023A7B" box="[1073,1253,763,789]" pageId="1" pageNumber="318">Chongyang Co.</collectingCounty>
, Shuikeng Vill.,
<geoCoordinate id="E1ADF5AB9311D12ECAB5FCB2FEF83A55" box="[151,287,800,827]" degrees="29" direction="north" minutes="18" orientation="latitude" pageId="1" pageNumber="318" precision="15" seconds="22" value="29.30611">29°1822″N</geoCoordinate>
,
<geoCoordinate id="E1ADF5AB9311D12ECB08FCB2FE5B3A55" box="[298,444,800,827]" degrees="114" direction="east" minutes="10" orientation="longitude" pageId="1" pageNumber="318" precision="15" seconds="23" value="114.17305">114°1023″E</geoCoordinate>
,
<quantity id="43613E899311D12ECBE4FC8CFDE93A57" box="[454,526,798,825]" metricMagnitude="2" metricUnit="m" metricValue="1.65" pageId="1" pageNumber="318" unit="m" value="165.0">
<elevation id="0FB4745F9311D12ECBE4FC8CFDE93A57" box="[454,526,798,825]" metricMagnitude="2" metricUnit="m" metricValue="1.65" pageId="1" pageNumber="318" unit="m" value="165.0">165 m</elevation>
</quantity>
,
<date id="F027B5AC9311D12EC83BFC8DFD273A57" box="[537,704,799,825]" pageId="1" pageNumber="318" value="2023-07-16" valueMax="2023-07-17" valueMin="2023-07-16">
<collectingDate id="E0634C449311D12EC83BFC8DFD273A57" box="[537,704,799,825]" pageId="1" pageNumber="318" value="2023-07-16" valueMax="2023-07-17" valueMin="2023-07-16">16-17.vii.2023</collectingDate>
</date>
, riverbank, under stone,
<collectorName id="296CF6BA9311D12EC9F0FC8DFB493A58" box="[978,1198,799,825]" pageId="1" pageNumber="318">
Ting-Kai Zeng,
<emphasis id="B6ED4F7E9311D12ECEA6FC8DFB493A58" bold="true" box="[1156,1198,799,822]" pageId="1" pageNumber="318">AE</emphasis>
</collectorName>
ẌṪOiǥŵDzffiṁṻ ḦAEñƃäŀ (
<collectionCode id="E2880BA99311D12ECB77FCD1FE413A33" box="[341,422,835,861]" pageId="1" pageNumber="318">SNUC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="34F199319311D12ECB9BFCD1FA873A33" box="[441,1376,834,861]" collectingDate="2023-07-16" collectingDateMax="2023-07-17" collectingDateMin="2023-07-16" collectionCode="SNUC" collectorName="Ting-Kai Zeng, AE" country="China" county="Chongyang Co." elevation="165" latitude="29.30611" location="Xianning City" longLatPrecision="20" longitude="114.17305" pageId="1" pageNumber="318" specimenCount="29" specimenCount-female="29" stateProvince="Hubei" typeStatus="paratype">
<emphasis id="B6ED4F7E9311D12ECB9BFCD1FD403A32" bold="true" box="[441,679,834,861]" pageId="1" pageNumber="318">
<typeStatus id="5B222DCE9311D12ECB9BFCD1FDD23A33" box="[441,565,835,861]" pageId="1" pageNumber="318">Paratypes</typeStatus>
:
<collectingCountry id="FC8ED3FC9311D12EC863FCD0FD443A32" box="[577,675,834,860]" name="China" pageId="1" pageNumber="318">CHINA</collectingCountry>
:
</emphasis>
37 JJ,
<specimenCount id="929F58E59311D12EC928FCD1FCBF3A33" box="[778,856,835,861]" pageId="1" pageNumber="318" type="female">29 ♀♀</specimenCount>
, same collecting data as for holotype (
<collectionCode id="E2880BA99311D12ECF25FCD1FABF3A33" box="[1287,1368,835,861]" pageId="1" pageNumber="318">SNUC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="CC83C0E79311D12ECAE5FCF4FC953D37" pageId="1" pageNumber="318" type="diagnosis">
<paragraph id="8426936C9311D12ECAE5FCF4FC953D37" blockId="1.[151,1437,762,2013]" pageId="1" pageNumber="318">
<emphasis id="B6ED4F7E9311D12ECAE5FCF4FEA53AEE" bold="true" box="[199,322,870,896]" pageId="1" pageNumber="318">Diagnosis.</emphasis>
<emphasis id="B6ED4F7E9311D12ECB6AFCF5FE653AEF" box="[328,386,871,897]" italics="true" pageId="1" pageNumber="318">Male</emphasis>
. Body length approximately
<quantity id="43613E899311D12EC8EBFCF5FCB43AEF" box="[713,851,871,897]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="1.8" metricValueMin="1.7" pageId="1" pageNumber="318" unit="mm" value="1.75" valueMax="1.8" valueMin="1.7">1.71.8 mm</quantity>
. Head subtruncate at base; vertex with curved transverse sulcus posterior to level of antennal tubercles, with long mediobasal carina, vertexal foveae moderately distinct and asetose; antenna relatively long, antennomeres each elongate, lacking modification. Discal stria of elytron extending posteriorly to approach posterior margin. Metafemur greatly swollen at apical 3/5 of dorsal margin. Abdomen dorsally with tergite 1 (IV) longer than 24 (VVII) combined, simple. Aedeagus strongly asymmetric, ventral stalk of median lobe much narrower than dorsal lobe.
<emphasis id="B6ED4F7E9311D12EC947FB89FC263D5B" box="[869,961,1051,1077]" italics="true" pageId="1" pageNumber="318">Female.</emphasis>
Body length approximately
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, metafemur lacking modification, genital complex as in
<figureCitation id="1CA28FE99311D12EC926FBADFC8A3D37" box="[772,877,1087,1113]" captionStart="FIGURE 1" captionStartId="2.[151,248,1734,1758]" captionTargetBox="[167,1413,151,1709]" captionTargetId="figure-150@2.[166,1413,151,1711]" captionTargetPageId="2" captionText="FIGURE 1. Habitus and morphological details of Batrisiella riparia sp. nov., male. A. Dorsal habitus. B. Head and pronotum. C. Scape, arrow indicating lamellar structure at apicolateral corner. D. Metafemur. E. Sternite 7 (IX). FH. Aedeagus, lateral (F), ventral (G), and axial (H). I. Female genital complex. Scale bars: 0.5 mm in A; 0.2 mm in B, D; 0.05 mm in C, E; 0.1 mm in FI." figureDoi="http://doi.org/10.5281/zenodo.8390216" httpUri="https://zenodo.org/record/8390216/files/figure.png" pageId="1" pageNumber="318">Figure 1I</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="CC83C0E79311D12CCAE5FBF0FB133DCF" lastPageId="3" lastPageNumber="320" pageId="1" pageNumber="318" type="description">
<paragraph id="8426936C9311D12ECAE5FBF0FC523DCF" blockId="1.[151,1437,762,2013]" pageId="1" pageNumber="318">
<emphasis id="B6ED4F7E9311D12ECAE5FBF0FEBE3D12" bold="true" box="[199,345,1122,1148]" pageId="1" pageNumber="318">Description.</emphasis>
<emphasis id="B6ED4F7E9311D12ECB42FBF1FE7D3D13" box="[352,410,1123,1149]" italics="true" pageId="1" pageNumber="318">Male</emphasis>
. Body (
<figureCitation id="1CA28FE99311D12ECBD1FBF1FDAD3D13" box="[499,586,1123,1149]" captionStart="FIGURE 1" captionStartId="2.[151,248,1734,1758]" captionTargetBox="[167,1413,151,1709]" captionTargetId="figure-150@2.[166,1413,151,1711]" captionTargetPageId="2" captionText="FIGURE 1. Habitus and morphological details of Batrisiella riparia sp. nov., male. A. Dorsal habitus. B. Head and pronotum. C. Scape, arrow indicating lamellar structure at apicolateral corner. D. Metafemur. E. Sternite 7 (IX). FH. Aedeagus, lateral (F), ventral (G), and axial (H). I. Female genital complex. Scale bars: 0.5 mm in A; 0.2 mm in B, D; 0.05 mm in C, E; 0.1 mm in FI." figureDoi="http://doi.org/10.5281/zenodo.8390216" httpUri="https://zenodo.org/record/8390216/files/figure.png" pageId="1" pageNumber="318">Fig. 1A</figureCitation>
) length
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; color reddish-brown, tarsi and mouthparts lighter in color. Dorsal surface of body covered with relatively short pubescence.
</paragraph>
<paragraph id="8426936C9311D12ECAE5FB39FC523F1B" blockId="1.[151,1437,762,2013]" pageId="1" pageNumber="318">
Head (
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) subtruncate at base, much wider than long, length
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, width across eyes
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; vertex finely punctate, convex at middle, with moderately distinct, asetose and broadly separated vertexal foveae (dorsal tentorial pits), with short, curved transverse sulcus at level of posterior to antennal tubercles, mediobasal carina extending posteriorly to occipital constriction and anteriorly to sulcus (level of anterior margin of eyes), antennal tubercles moderately raised, surrounding area roughly punctate; frons coarsely punctate broadly and shallowly impressed medially, confluent with clypeus at middle, anterolaterally with thin, oblique carinae; clypeus with rough surface, its anterior margin carinate and moderately raised; ocular-mandibular carinae complete. Venter with tiny gular foveae (posterior tentorial pits) originating from shared small, transverse opening, with distinct median carina extending from opening anteriorly to mouthparts. Compound eyes greatly prominent, each composed of approximately 32 large ommatidia. Antenna 1.0 mm long, lacking modification; antennomere 1 thick and short, anterolateral margin with small hyaline, lamellar structure (
<figureCitation id="1CA28FE99311D12EC8F7F981FCCB3F43" box="[725,812,1555,1581]" captionStart="FIGURE 1" captionStartId="2.[151,248,1734,1758]" captionTargetBox="[167,1413,151,1709]" captionTargetId="figure-150@2.[166,1413,151,1711]" captionTargetPageId="2" captionText="FIGURE 1. Habitus and morphological details of Batrisiella riparia sp. nov., male. A. Dorsal habitus. B. Head and pronotum. C. Scape, arrow indicating lamellar structure at apicolateral corner. D. Metafemur. E. Sternite 7 (IX). FH. Aedeagus, lateral (F), ventral (G), and axial (H). I. Female genital complex. Scale bars: 0.5 mm in A; 0.2 mm in B, D; 0.05 mm in C, E; 0.1 mm in FI." figureDoi="http://doi.org/10.5281/zenodo.8390216" httpUri="https://zenodo.org/record/8390216/files/figure.png" pageId="1" pageNumber="318">Fig. 1C</figureCitation>
), antennomeres 27 each elongate, 7 longer than 6 and 8, 8 shortest, elongate, 911 enlarged, forming indistinct club, 10 as long as but broader than 9, 11 largest, slightly shorter than 9 and 10 combined (18:21), sub-fusiform, truncate at base.
</paragraph>
<paragraph id="8426936C9311D12ECAE5F9EDFB1E3E23" blockId="1.[151,1437,762,2013]" pageId="1" pageNumber="318">
Pronotum (
<figureCitation id="1CA28FE99311D12ECB64F9EDFE7A3FF7" box="[326,413,1663,1689]" captionStart="FIGURE 1" captionStartId="2.[151,248,1734,1758]" captionTargetBox="[167,1413,151,1709]" captionTargetId="figure-150@2.[166,1413,151,1711]" captionTargetPageId="2" captionText="FIGURE 1. Habitus and morphological details of Batrisiella riparia sp. nov., male. A. Dorsal habitus. B. Head and pronotum. C. Scape, arrow indicating lamellar structure at apicolateral corner. D. Metafemur. E. Sternite 7 (IX). FH. Aedeagus, lateral (F), ventral (G), and axial (H). I. Female genital complex. Scale bars: 0.5 mm in A; 0.2 mm in B, D; 0.05 mm in C, E; 0.1 mm in FI." figureDoi="http://doi.org/10.5281/zenodo.8390216" httpUri="https://zenodo.org/record/8390216/files/figure.png" pageId="1" pageNumber="318">Fig. 1B</figureCitation>
) slightly longer than wide, length
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, width
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, widest at middle; lateral margins rounded; disc moderately convex, finely punctate, with median longitudinal sulcus as long as semicircular lateral sulci in dorsal view; lacking median antebasal fovea, with complete, deep transverse antebasal sulcus connecting lateral antebasal foveae, with distinct antebasal spines; outer and inner pair of basolateral foveae small. Prosternum with anterior part slightly longer than coxal part, with small lateral procoxal foveae; hypomeral groove complete, with small lateral antebasal hypomeral impression; margin of coxal cavity thinly carinate.
</paragraph>
<paragraph id="8426936C9311D12ECAE5F8C5FB543EB3" blockId="1.[151,1437,762,2013]" pageId="1" pageNumber="318">
Elytra much wider than long, length
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, width
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; each elytron with two large, widely separated basal foveae, lacking subbasal fovea; humeri roundly protuberant; discal striae long, slightly curved, extending from outer basal foveae to approximately apical 4/5 of elytral length; subhumeral foveae small, carinate marginal striae extending posteriorly from fovea to apex of elytra. Metathoracic wings fully developed.
</paragraph>
<caption id="D0E6C3E49312D12DCAB5F954FA713E48" ID-DOI="http://doi.org/10.5281/zenodo.8390216" ID-Zenodo-Dep="8390216" httpUri="https://zenodo.org/record/8390216/files/figure.png" pageId="2" pageNumber="319" startId="2.[151,248,1734,1758]" targetBox="[167,1413,151,1709]" targetPageId="2" targetType="figure">
<paragraph id="8426936C9312D12DCAB5F954FA713E48" blockId="2.[151,1437,1734,1830]" pageId="2" pageNumber="319">
<emphasis id="B6ED4F7E9312D12DCAB5F954FEF43FB0" bold="true" box="[151,275,1734,1758]" pageId="2" pageNumber="319">FIGURE 1.</emphasis>
Habitus and morphological details of
<taxonomicName id="4399E8EF9312D12DC8B5F954FCAA3FB0" authority="Yin &amp; Zeng, 2023" authorityName="Yin &amp; Zeng" authorityYear="2023" box="[663,845,1734,1758]" class="Insecta" family="Staphylinidae" genus="Batrisiella" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="319" phylum="Arthropoda" rank="species" species="riparia" status="sp. nov.">
<emphasis id="B6ED4F7E9312D12DC8B5F954FCAA3FB0" box="[663,845,1734,1758]" italics="true" pageId="2" pageNumber="319">Batrisiella riparia</emphasis>
</taxonomicName>
<emphasis id="B6ED4F7E9312D12DC976F954FC423FB0" bold="true" box="[852,933,1734,1758]" pageId="2" pageNumber="319">
<taxonomicNameLabel id="ADDEF2059312D12DC976F954FC423FB0" box="[852,933,1734,1758]" pageId="2" pageNumber="319" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male.
<emphasis id="B6ED4F7E9312D12DC9D3F954FBEE3FB0" bold="true" box="[1009,1033,1734,1758]" pageId="2" pageNumber="319">A.</emphasis>
Dorsal habitus.
<emphasis id="B6ED4F7E9312D12DCE8DF954FB213FB0" bold="true" box="[1199,1222,1734,1758]" pageId="2" pageNumber="319">B.</emphasis>
Head and pronotum.
<emphasis id="B6ED4F7E9312D12DCAB5F978FF483E6C" bold="true" box="[151,175,1770,1794]" pageId="2" pageNumber="319">C.</emphasis>
Scape, arrow indicating lamellar structure at apicolateral corner.
<emphasis id="B6ED4F7E9312D12DC91AF978FCB63E6C" bold="true" box="[824,849,1770,1794]" pageId="2" pageNumber="319">D.</emphasis>
Metafemur.
<emphasis id="B6ED4F7E9312D12DC9F2F978FC003E6C" bold="true" box="[976,999,1770,1794]" pageId="2" pageNumber="319">E.</emphasis>
Sternite 7 (IX).
<emphasis id="B6ED4F7E9312D12DCEA9F978FB263E6C" bold="true" box="[1163,1217,1770,1794]" pageId="2" pageNumber="319">FH.</emphasis>
Aedeagus, lateral (F), ventral (G), and axial (H).
<emphasis id="B6ED4F7E9312D12DCBBDF89CFE483E48" bold="true" box="[415,431,1806,1830]" pageId="2" pageNumber="319">I.</emphasis>
Female genital complex. Scale bars: 0.5 mm in A; 0.2 mm in B, D; 0.05 mm in C, E; 0.1 mm in FI.
</paragraph>
</caption>
<paragraph id="8426936C9312D12DCAE5F8C6FA6F3EB4" blockId="2.[151,1437,1876,2011]" pageId="2" pageNumber="319">Mesoventrite short, fully demarcated from metaventrite by oblique lateral carinae, with pair of admesal carinae; setose median mesoventral foveae broadly separated, lateral mesoventral foveae large and setose, forked internally. Metaventrite weakly convex adjacent to middle, with pairs of setose lateral mesocoxal and lateral metaventral foveae located lateral and posterior to coxal cavities, respectively, posterior margin with small and narrow split at middle.</paragraph>
<paragraph id="8426936C9313D12CCAE5FF05FED43997" blockId="3.[151,1437,151,1545]" pageId="3" pageNumber="320">
Legs elongate; mesotibia with small spur at apex; metafemur (
<figureCitation id="1CA28FE99313D12CC9B2FF05FC0F39DF" box="[912,1000,151,177]" captionStart="FIGURE 1" captionStartId="2.[151,248,1734,1758]" captionTargetBox="[167,1413,151,1709]" captionTargetId="figure-150@2.[166,1413,151,1711]" captionTargetPageId="2" captionText="FIGURE 1. Habitus and morphological details of Batrisiella riparia sp. nov., male. A. Dorsal habitus. B. Head and pronotum. C. Scape, arrow indicating lamellar structure at apicolateral corner. D. Metafemur. E. Sternite 7 (IX). FH. Aedeagus, lateral (F), ventral (G), and axial (H). I. Female genital complex. Scale bars: 0.5 mm in A; 0.2 mm in B, D; 0.05 mm in C, E; 0.1 mm in FI." figureDoi="http://doi.org/10.5281/zenodo.8390216" httpUri="https://zenodo.org/record/8390216/files/figure.png" pageId="3" pageNumber="320">Fig. 1D</figureCitation>
) distinctly clavate, greatly swollen on dorsal side, forming two projections and with setose sulcate area between them, proximal area with dense, short sensory setae.
</paragraph>
<paragraph id="8426936C9313D12CCAE5FE91FE9D3B0F" blockId="3.[151,1437,151,1545]" pageId="3" pageNumber="320">
Abdomen slightly narrower than elytra, widest at basolateral margins of tergite 1 (IV), length
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, width
<quantity id="43613E899313D12CCAFFFEB5FEA4382F" box="[221,323,295,321]" metricMagnitude="-4" metricUnit="m" metricValue="6.1" pageId="3" pageNumber="320" unit="mm" value="0.61">0.61 mm</quantity>
; lacking modification. Tergite 1 (IV) in dorsal view longer than 24 (VVII) combined; setose basal sulcus laterally ended at inner pair of basolateral foveae, with outer pair of basolateral foveae lateral to sulcus, discal carinae thin and short, indistinct; tergites 2 and 3 (V and VI) each short, lacking fovea, 4 (VII) as long as 2 and 3 combined along middle, with one pair of small basolateral foveae, 5 (VIII) semicircular, posterior margin broadly and roundly emarginate at middle. Sternite 2 (IV) with one pair of mediobasal foveae and one pair of large basolateral sockets, with one pair of short lateral carinae; midlength of sternite 2 as long as 35 (VVII) combined, 35 each short, successively shorter, lacking fovea, 6 (VIII) greatly transverse, posterior margin evenly roundly curved, sternite 7 (IX) (
<figureCitation id="1CA28FE99313D12CCB60FDB1FE723B53" box="[322,405,547,573]" captionStart="FIGURE 1" captionStartId="2.[151,248,1734,1758]" captionTargetBox="[167,1413,151,1709]" captionTargetId="figure-150@2.[166,1413,151,1711]" captionTargetPageId="2" captionText="FIGURE 1. Habitus and morphological details of Batrisiella riparia sp. nov., male. A. Dorsal habitus. B. Head and pronotum. C. Scape, arrow indicating lamellar structure at apicolateral corner. D. Metafemur. E. Sternite 7 (IX). FH. Aedeagus, lateral (F), ventral (G), and axial (H). I. Female genital complex. Scale bars: 0.5 mm in A; 0.2 mm in B, D; 0.05 mm in C, E; 0.1 mm in FI." figureDoi="http://doi.org/10.5281/zenodo.8390216" httpUri="https://zenodo.org/record/8390216/files/figure.png" pageId="3" pageNumber="320">Fig. 1E</figureCitation>
) suboval, apical half moderately sclerotized, basal half membranous, apical margin with few scattered long setae.
</paragraph>
<paragraph id="8426936C9313D12CCAE5FDF9FE6F3A7B" blockId="3.[151,1437,151,1545]" pageId="3" pageNumber="320">
Aedeagus (
<figureCitation id="1CA28FE99313D12CCB64FDF9FE5B3BEB" box="[326,444,619,645]" captionStart="FIGURE 1" captionStartId="2.[151,248,1734,1758]" captionTargetBox="[167,1413,151,1709]" captionTargetId="figure-150@2.[166,1413,151,1711]" captionTargetPageId="2" captionText="FIGURE 1. Habitus and morphological details of Batrisiella riparia sp. nov., male. A. Dorsal habitus. B. Head and pronotum. C. Scape, arrow indicating lamellar structure at apicolateral corner. D. Metafemur. E. Sternite 7 (IX). FH. Aedeagus, lateral (F), ventral (G), and axial (H). I. Female genital complex. Scale bars: 0.5 mm in A; 0.2 mm in B, D; 0.05 mm in C, E; 0.1 mm in FI." figureDoi="http://doi.org/10.5281/zenodo.8390216" httpUri="https://zenodo.org/record/8390216/files/figure.png" pageId="3" pageNumber="320">Fig. 1FH</figureCitation>
)
<quantity id="43613E899313D12CCBEFFDF9FDD53BEB" box="[461,562,619,645]" metricMagnitude="-4" metricUnit="m" metricValue="2.7" pageId="3" pageNumber="320" unit="mm" value="0.27">0.27 mm</quantity>
long, strongly asymmetric; median lobe with greatly expanded basal capsule and large, suboval foramen, with long, extended basoventral projection, ventral stalk of median lobe shorter and narrower than dorsal lobe, in lateral view sharply narrowing toward apex, in axial view twisted and narrowed at middle; dorsal lobe broadened apically, at apex bifurcated; parameres reduced to small, membranous structure located above foramen.
</paragraph>
<paragraph id="8426936C9313D12CCAE5FC8DFDF73A83" blockId="3.[151,1437,151,1545]" pageId="3" pageNumber="320">
<emphasis id="B6ED4F7E9313D12CCAE5FC8DFEFC3A57" box="[199,283,799,825]" italics="true" pageId="3" pageNumber="320">Female</emphasis>
. Similar to male in external morphology; antenna slightly shorter; metafemur lacking modification; each compound eye composed of approximately 23 ommatidia; humeral prominence small, indistinct; metathoracic wings absent. Measurements (as for male): body length
<quantity id="43613E899313D12CC933FCF5FC503AEF" box="[785,951,871,897]" metricMagnitude="-3" metricUnit="m" metricValue="1.765" metricValueMax="1.81" metricValueMin="1.72" pageId="3" pageNumber="320" unit="mm" value="1.765" valueMax="1.81" valueMin="1.72">1.721.81 mm</quantity>
; length/width of head 0.360.37/
<quantity id="43613E899313D12CCF0EFCF5FF223ACB" metricMagnitude="-4" metricUnit="m" metricValue="3.95" metricValueMax="4.0" metricValueMin="3.9" pageId="3" pageNumber="320" unit="mm" value="0.395" valueMax="0.4" valueMin="0.39">0.390.40 mm</quantity>
, pronotum 0.400.42/
<quantity id="43613E899313D12CCBE1FC19FDCC3ACB" box="[451,555,907,933]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="3" pageNumber="320" unit="mm" value="0.4">0.40 mm</quantity>
, elytra 0.50/
<quantity id="43613E899313D12CC89FFC18FCC13ACB" box="[701,806,906,933]" metricMagnitude="-4" metricUnit="m" metricValue="6.4" pageId="3" pageNumber="320" unit="mm" value="0.64">0.64 mm</quantity>
; abdomen 0.500.52/
<quantity id="43613E899313D12CCE3FFC18FB213ACB" box="[1053,1222,906,933]" metricMagnitude="-4" metricUnit="m" metricValue="5.949999999999999" metricValueMax="6.0" metricValueMin="5.8999999999999995" pageId="3" pageNumber="320" unit="mm" value="0.595" valueMax="0.6" valueMin="0.59">0.590.60 mm</quantity>
; length of antenna
<quantity id="43613E899313D12CCAB5FC3DFEDA3AA7" box="[151,317,943,969]" metricMagnitude="-4" metricUnit="m" metricValue="9.2" metricValueMax="9.3" metricValueMin="9.1" pageId="3" pageNumber="320" unit="mm" value="0.92" valueMax="0.93" valueMin="0.91">0.910.93 mm</quantity>
; maximum width of genital complex (
<figureCitation id="1CA28FE99313D12CC8C9FC3DFCD03AA7" box="[747,823,943,969]" captionStart="FIGURE 1" captionStartId="2.[151,248,1734,1758]" captionTargetBox="[167,1413,151,1709]" captionTargetId="figure-150@2.[166,1413,151,1711]" captionTargetPageId="2" captionText="FIGURE 1. Habitus and morphological details of Batrisiella riparia sp. nov., male. A. Dorsal habitus. B. Head and pronotum. C. Scape, arrow indicating lamellar structure at apicolateral corner. D. Metafemur. E. Sternite 7 (IX). FH. Aedeagus, lateral (F), ventral (G), and axial (H). I. Female genital complex. Scale bars: 0.5 mm in A; 0.2 mm in B, D; 0.05 mm in C, E; 0.1 mm in FI." figureDoi="http://doi.org/10.5281/zenodo.8390216" httpUri="https://zenodo.org/record/8390216/files/figure.png" pageId="3" pageNumber="320">Fig. 1I</figureCitation>
)
<quantity id="43613E899313D12CC965FC3DFC493AA7" box="[839,942,943,969]" metricMagnitude="-4" metricUnit="m" metricValue="2.2" pageId="3" pageNumber="320" unit="mm" value="0.22">0.22 mm</quantity>
, genital plate slightly narrower than sternite 9, lateral arms narrowing distally.
</paragraph>
<paragraph id="8426936C9313D12CCAE5FC64FB133DCF" blockId="3.[151,1437,151,1545]" pageId="3" pageNumber="320">
<emphasis id="B6ED4F7E9313D12CCAE5FC64FE573D7F" bold="true" box="[199,432,1014,1041]" pageId="3" pageNumber="320">Comparative notes.</emphasis>
This species is morphologically most similar to the
<typeStatus id="5B222DCE9313D12CC9D0FC65FBC53D7F" box="[1010,1058,1015,1041]" pageId="3" pageNumber="320">type</typeStatus>
species of
<taxonomicName id="4399E8EF9313D12CCEBEFC65FA7F3D7F" authority="Jeannel" authorityName="Jeannel" authorityYear="1952" box="[1180,1432,1015,1041]" class="Insecta" family="Staphylinidae" genus="Physomerinus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="320" phylum="Arthropoda" rank="genus">
<emphasis id="B6ED4F7E9313D12CCEBEFC65FADC3D7F" box="[1180,1339,1015,1041]" italics="true" pageId="3" pageNumber="320">Physomerinus</emphasis>
Jeannel
</taxonomicName>
,
<taxonomicName id="4399E8EF9313D12CCAB5FB89FDB13D5B" authority="(Schaufuss, L. W.)" baseAuthorityName="Schaufuss, L. W." box="[151,598,1051,1077]" class="Insecta" family="Staphylinidae" genus="Physomerinus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="320" phylum="Arthropoda" rank="species" species="septemfoveolatus">
<emphasis id="B6ED4F7E9313D12CCAB5FB89FE903D5B" box="[151,375,1051,1077]" italics="true" pageId="3" pageNumber="320">P. septemfoveolatus</emphasis>
(Schaufuss, L. W.)
</taxonomicName>
distributed in
<collectingCountry id="FC8ED3FC9313D12CC923FB89FC873D5B" box="[769,864,1051,1077]" name="Vietnam" pageId="3" pageNumber="320">Vietnam</collectingCountry>
and
<collectingCountry id="FC8ED3FC9313D12CC9BEFB89FBE53D5B" box="[924,1026,1051,1077]" name="Thailand" pageId="3" pageNumber="320">Thailand</collectingCountry>
, by the greatly swollen male metafemora. They may be separated by the different shapes of the male femoral modification and the aedeagus. The metafemora of
<taxonomicName id="4399E8EF9313D12CCB62FBF1FDFC3D13" box="[320,539,1123,1149]" class="Insecta" family="Staphylinidae" genus="Physomerinus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="320" phylum="Arthropoda" rank="species" species="septemfoveolatus">
<emphasis id="B6ED4F7E9313D12CCB62FBF1FDFC3D13" box="[320,539,1123,1149]" italics="true" pageId="3" pageNumber="320">P. septemfoveolatus</emphasis>
</taxonomicName>
appear to have only one projection on the dorsal margin, and the surface declines distally following the projection. For the aedeagus of
<taxonomicName id="4399E8EF9313D12CC8D2FB15FC373DCF" box="[752,976,1159,1185]" class="Insecta" family="Staphylinidae" genus="Physomerinus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="320" phylum="Arthropoda" rank="species" species="septemfoveolatus">
<emphasis id="B6ED4F7E9313D12CC8D2FB15FCE63DCE" box="[752,769,1159,1184]" italics="true" pageId="3" pageNumber="320">P</emphasis>
.
<emphasis id="B6ED4F7E9313D12CC92DFB15FC373DCF" box="[783,976,1159,1185]" italics="true" pageId="3" pageNumber="320">septemfoveolatus</emphasis>
</taxonomicName>
see
<bibRefCitation id="E008EE9D9313D12CCE20FB15FB7E3DCF" author="Jeannel, R." box="[1026,1177,1158,1185]" pageId="3" pageNumber="320" pagination="69 - 113" refId="ref4165" refString="Jeannel, R. (1952) Pselaphides de Saigon. Revue Francaise d'Entomologie, 19, 69 - 113." type="journal article" year="1952">Jeannel 1952</bibRefCitation>
: fig. 42.
</paragraph>
</subSubSection>
<subSubSection id="CC83C0E79313D12CCAE5FB38FD7A3D87" pageId="3" pageNumber="320" type="biology_ecology">
<paragraph id="8426936C9313D12CCAE5FB38FD7A3D87" blockId="3.[151,1437,151,1545]" pageId="3" pageNumber="320">
<emphasis id="B6ED4F7E9313D12CCAE5FB38FEC03DAA" bold="true" box="[199,295,1194,1220]" pageId="3" pageNumber="320">Biology.</emphasis>
Most individuals were collected under stones on the river bank, and a few were observed running around on sandy soil between grasses (
<figureCitation id="1CA28FE99313D12CC869FB5DFD773D87" box="[587,656,1231,1257]" captionStart="FIGURE 3" captionStartId="6.[151,250,1628,1652]" captionTargetBox="[166,1413,454,1588]" captionTargetId="figure-173@6.[166,1413,454,1589]" captionTargetPageId="6" captionText="FIGURE 3. Collecting environment and habitat of Batrisiella riparia sp. nov. and Nipponobythus varicornis sp. nov." figureDoi="http://doi.org/10.5281/zenodo.8390222" httpUri="https://zenodo.org/record/8390222/files/figure.png" pageId="3" pageNumber="320">Fig. 3</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="CC83C0E79313D12CCAE5FB60FDB83C63" box="[199,607,1266,1293]" pageId="3" pageNumber="320" type="distribution">
<paragraph id="8426936C9313D12CCAE5FB60FDB83C63" blockId="3.[151,1437,151,1545]" box="[199,607,1266,1293]" pageId="3" pageNumber="320">
<emphasis id="B6ED4F7E9313D12CCAE5FB60FE863C62" bold="true" box="[199,353,1266,1292]" pageId="3" pageNumber="320">Distribution.</emphasis>
Central
<collectingCountry id="FC8ED3FC9313D12CCBE3FB61FDEF3C63" box="[449,520,1267,1293]" name="China" pageId="3" pageNumber="320">China</collectingCountry>
:
<collectingRegion id="465D5D8E9313D12CC831FB61FDBC3C63" box="[531,603,1267,1293]" country="China" name="Hubei" pageId="3" pageNumber="320">Hubei</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="CC83C0E79313D12CCAE5FA85FC813C3B" pageId="3" pageNumber="320" type="etymology">
<paragraph id="8426936C9313D12CCAE5FA85FC813C3B" blockId="3.[151,1437,151,1545]" pageId="3" pageNumber="320">
<emphasis id="B6ED4F7E9313D12CCAE5FA85FEA93C5F" bold="true" box="[199,334,1303,1329]" pageId="3" pageNumber="320">Etymology.</emphasis>
The name
<emphasis id="B6ED4F7E9313D12CCBEEFA85FDC13C5F" box="[460,550,1303,1329]" italics="true" pageId="3" pageNumber="320">rîpârius</emphasis>
(-
<emphasis id="B6ED4F7E9313D12CC862FA85FDA93C5E" box="[576,590,1303,1328]" italics="true" pageId="3" pageNumber="320">a</emphasis>
, -
<emphasis id="B6ED4F7E9313D12CC844FA85FD6F3C5E" box="[614,648,1303,1328]" italics="true" pageId="3" pageNumber="320">um</emphasis>
) is a Latin adjective, meaning, “that inhabits the banks of rivers”, “riparian”, referring to the habitat where the species was collected.
</paragraph>
</subSubSection>
<subSubSection id="CC83C0E79313D12CCAE5FACDFA673F67" pageId="3" pageNumber="320" type="discussion">
<paragraph id="8426936C9313D12CCAE5FACDFA673F67" blockId="3.[151,1437,151,1545]" pageId="3" pageNumber="320">
<emphasis id="B6ED4F7E9313D12CCAE5FACDFEDD3C17" bold="true" box="[199,314,1375,1401]" pageId="3" pageNumber="320">Remarks.</emphasis>
Without examining the aedeagus and lacking a closer look at the scapes, this species would be easily regarded as a member of
<taxonomicName id="4399E8EF9313D12CCB94FA11FDB43CF3" authorityName="Jeannel" authorityYear="1952" box="[438,595,1411,1437]" class="Insecta" family="Staphylinidae" genus="Physomerinus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="320" phylum="Arthropoda" rank="genus">
<emphasis id="B6ED4F7E9313D12CCB94FA11FDB43CF3" box="[438,595,1411,1437]" italics="true" pageId="3" pageNumber="320">Physomerinus</emphasis>
</taxonomicName>
due to the swollen male metafemora. This may be the reason why
<bibRefCitation id="E008EE9D9313D12CCF6BFA11FF3D3CAF" author="Jeannel, R." pageId="3" pageNumber="320" pagination="69 - 113" refId="ref4165" refString="Jeannel, R. (1952) Pselaphides de Saigon. Revue Francaise d'Entomologie, 19, 69 - 113." type="journal article" year="1952">Jeannel (1952</bibRefCitation>
,
<bibRefCitation id="E008EE9D9313D12CCAC7FA34FEF93CAF" author="Jeannel, R." box="[229,286,1446,1473]" pageId="3" pageNumber="320" pagination="1 - 138" refId="ref4186" refString="Jeannel, R. (1958) Revision des Pselaphides du Japon. Memoires du Museum National d'Histoire Naturelle, A: Zoologie, 18, 1 - 138." type="journal article" year="1958">1958</bibRefCitation>
) created
<taxonomicName id="4399E8EF9313D12CCBA3FA35FDF93CAF" authorityName="Jeannel" authorityYear="1952" box="[385,542,1447,1473]" class="Insecta" family="Staphylinidae" genus="Physomerinus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="320" phylum="Arthropoda" rank="genus">
<emphasis id="B6ED4F7E9313D12CCBA3FA35FDF93CAF" box="[385,542,1447,1473]" italics="true" pageId="3" pageNumber="320">Physomerinus</emphasis>
</taxonomicName>
and removed other, unrelated species there when his generic concept of
<taxonomicName id="4399E8EF9313D12CCF17FA35FF5A3C8B" class="Insecta" family="Staphylinidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="320" phylum="Arthropoda" rank="subFamily" subFamily="Pselaphinae">Pselaphinae</taxonomicName>
was largely based on male features. The probable synonymy of
<taxonomicName id="4399E8EF9313D12CC957FA59FBF53C8B" authorityName="Jeannel" authorityYear="1952" box="[885,1042,1483,1509]" class="Insecta" family="Staphylinidae" genus="Physomerinus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="320" phylum="Arthropoda" rank="genus">
<emphasis id="B6ED4F7E9313D12CC957FA59FBF53C8B" box="[885,1042,1483,1509]" italics="true" pageId="3" pageNumber="320">Physomerinus</emphasis>
</taxonomicName>
with
<taxonomicName id="4399E8EF9313D12CCE6CFA59FAFB3C8B" authority="Raffray" authorityName="Raffray" authorityYear="1904" box="[1102,1308,1483,1509]" class="Insecta" family="Staphylinidae" genus="Batrisiella" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="320" phylum="Arthropoda" rank="genus">
<emphasis id="B6ED4F7E9313D12CCE6CFA59FB243C8B" box="[1102,1219,1483,1509]" italics="true" pageId="3" pageNumber="320">Batrisiella</emphasis>
Raffray
</taxonomicName>
(pers. com. with S. Nomura, dated
<date id="F027B5AC9313D12CCBBAFA7DFDFA3F67" box="[408,541,1519,1545]" pageId="3" pageNumber="320" value="2023-07-27">2023-07-27</date>
), and placement of the other species of the genus will be dealt with separately.
</paragraph>
</subSubSection>
</treatment>
</document>