treatments-xml/data/03/F4/87/03F487D4FF8C1A72FF04F93CBC26F9B7.xml
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<document id="702152DED5097110E0E00189CD5395A8" ID-DOI="10.11646/zootaxa.4032.4.1" ID-GBIF-Dataset="1459e41f-b82d-4c88-be4d-ee86e329f9c7" ID-ISSN="1175-5326" ID-Zenodo-Dep="243485" ID-ZooBank="AFEF8567-B7C6-4F54-AC51-4C990389BF1D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461294283733" checkinUser="plazi" docAuthor="Kustov, Semen &amp; Zherebilo, Danil" docDate="2015" docId="03F487D4FF8C1A72FF04F93CBC26F9B7" docLanguage="en" docName="zt04032p369.pdf" docOrigin="Zootaxa 4032 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Wiedemannia pseguashae Kustov &amp; Zherebilo, 2015, sp. nov." docType="treatment" docVersion="9" lastPageNumber="364" masterDocId="FFCDFFACFF871A7FFF93FFA0BD01FFCD" masterDocTitle="New data on the genus Wiedemannia Zetterstedt (Diptera: Empididae) from the Caucasus with description of four new species" masterLastPageNumber="369" masterPageNumber="351" pageNumber="362" updateTime="1698616197258" updateUser="plazi">
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<mods:title id="1D48158E2D090232D7A2B151B895FF37">New data on the genus Wiedemannia Zetterstedt (Diptera: Empididae) from the Caucasus with description of four new species</mods:title>
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<mods:namePart id="EFD5102C1F292BB90D11496D03BAAD0F">Kustov, Semen</mods:namePart>
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<mods:namePart id="E1C3D18B18322F7FFB179A789A7E26EA">Zherebilo, Danil</mods:namePart>
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<mods:typeOfResource id="36C8BE03A91E863E463532213A42C14E">text</mods:typeOfResource>
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<mods:title id="A6298C14615691DAA3EE3D7398D41CB3">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="C705E5A9DC4228D8885EE768667FD28B">
<mods:date id="9DFB82C0BE7101DDC4220470D5D7F481">2015</mods:date>
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<treatment id="03F487D4FF8C1A72FF04F93CBC26F9B7" ID-DOI="http://doi.org/10.5281/zenodo.5691037" ID-GBIF-Taxon="119642272" ID-Zenodo-Dep="5691037" LSID="urn:lsid:plazi:treatment:03F487D4FF8C1A72FF04F93CBC26F9B7" httpUri="http://treatment.plazi.org/id/03F487D4FF8C1A72FF04F93CBC26F9B7" lastPageId="13" lastPageNumber="364" pageId="11" pageNumber="362">
<subSubSection id="C3476549FF8C1A74FF04F93CBDF9F91A" pageId="11" pageNumber="362" type="nomenclature">
<paragraph id="8BE236C2FF8C1A74FF04F93CBF37F97B" blockId="11.[151,566,1692,1751]" box="[151,566,1692,1718]" pageId="11" pageNumber="362">
<heading id="D0AA81AEFF8C1A74FF04F93CBF37F97B" bold="true" box="[151,566,1692,1718]" fontSize="11" level="1" pageId="11" pageNumber="362" reason="1">
<emphasis id="B929EAD0FF8C1A74FF04F93CBF37F97B" bold="true" box="[151,566,1692,1718]" pageId="11" pageNumber="362">
<taxonomicName id="4C5D4D41FF8C1A74FF04F93CBCD0F97B" ID-CoL="9HT8J" box="[151,465,1692,1718]" class="Insecta" family="Empididae" genus="Wiedemannia" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="362" phylum="Arthropoda" rank="species" species="pseguashae" status="sp. nov.">
<emphasis id="B929EAD0FF8C1A74FF04F93CBCD0F97B" bold="true" box="[151,465,1692,1718]" italics="true" pageId="11" pageNumber="362">Wiedemannia pseguashae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21A57ABFF8C1A74FE4BF93CBF37F97B" box="[472,566,1692,1718]" pageId="11" pageNumber="362" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BE236C2FF8C1A74FF04F91EBDF9F91A" blockId="11.[151,566,1692,1751]" box="[151,248,1726,1751]" pageId="11" pageNumber="362">
(
<figureCitation id="13662A47FF8C1A74FF0CF91EBDF0F91A" box="[159,241,1726,1751]" captionStart="FIGURES 12 17" captionStartId="12.[151,263,1937,1959]" captionTargetBox="[247,1322,214,1871]" captionTargetId="figure@12.[229,1337,193,1891]" captionTargetPageId="12" captionText="FIGURES 12 17. Male terminalia of Wiedemannia, lateral view. 12, W. pseguashae sp. nov.; 13, W. pseudovaillanti Joost; 14, W. shamshevi Kustov &amp; Zherebilo; 15, W. sinclairi Kustov &amp; Zherebilo; 16, W. vaillanti Joost; 17, W. zetterstedti (Fallén). Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/243488/files/figure.png" pageId="11" pageNumber="362">Fig. 12</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3476549FF8C1A72FF04F8A1B857FF19" lastPageId="13" lastPageNumber="364" pageId="11" pageNumber="362" type="materials_examined">
<paragraph id="8BE236C2FF8C1A72FF04F8A1B857FF19" blockId="11.[151,1437,1793,1998]" lastBlockId="13.[151,1437,151,1658]" lastPageId="13" lastPageNumber="364" pageId="11" pageNumber="362">
<emphasis id="B929EAD0FF8C1A74FF04F8A1BC3FF8D7" bold="true" box="[151,318,1793,1818]" pageId="11" pageNumber="362">
<typeStatus id="54E68860FF8C1A74FF04F8A1BDD3F8D7" box="[151,210,1793,1818]" pageId="11" pageNumber="362">Type</typeStatus>
material
</emphasis>
.
<typeStatus id="54E68860FF8C1A74FEDFF8A1BCEBF8D7" box="[332,490,1793,1818]" pageId="11" pageNumber="362" type="holotype">
<emphasis id="B929EAD0FF8C1A74FEDFF8A1BCEBF8D7" bold="true" box="[332,490,1793,1818]" pageId="11" pageNumber="362">HOLOTYPE</emphasis>
</typeStatus>
, ♂ labelled [printed in Cyrillic], [
<collectingCountry id="F34A7652FF8C1A74FCF9F8A1B94CF8D7" box="[874,1101,1793,1818]" name="South Ossetia" pageId="11" pageNumber="362">
<emphasis id="B929EAD0FF8C1A74FCF9F8A1B94CF8D7" bold="true" box="[874,1101,1793,1818]" pageId="11" pageNumber="362">SOUTH OSSETIA</emphasis>
</collectingCountry>
]: “C. Kavkaz, Ju. Ossetia, hr. [ridge] Britauli / C-B skloni g. [north-east slopes of Mountain Britavtsak / lev.bereg r. [left bank of River] Britadon / Vis. [high]
<quantity id="4CA59B27FF8C1A74FEBEF8E9BC84F8AC" box="[301,389,1865,1890]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="11" pageNumber="362" unit="m" value="2500.0">2500 m</quantity>
[a.s.l.] /
<geoCoordinate id="EE695005FF8C1A74FE61F8EABF8CF8AF" box="[498,653,1866,1890]" direction="north" orientation="latitude" pageId="11" pageNumber="362" precision="1" value="42.487057">42°29'13.4&quot;N</geoCoordinate>
,
<geoCoordinate id="EE695005FF8C1A74FD0FF8E9BE48F8AC" box="[668,841,1865,1890]" direction="east" orientation="longitude" pageId="11" pageNumber="362" precision="1" value="44.176277">044°10'34.6&quot; E</geoCoordinate>
/
<date id="FFE31002FF8C1A74FCF6F8E9BEE6F8AF" box="[869,999,1865,1890]" pageId="11" pageNumber="362" value="2013-07-21">21.vii.2013</date>
, D. Zherebilo” (
<collectionCode id="ED4CAE07FF8C1A74FB35F8EAB9DAF8AF" box="[1190,1243,1866,1890]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="11" pageNumber="362">ZIN</collectionCode>
).
<typeStatus id="54E68860FF8C1A74FB63F8E9B893F8AF" box="[1264,1426,1865,1890]" pageId="11" pageNumber="362" type="paratype">
<emphasis id="B929EAD0FF8C1A74FB63F8E9B893F8AF" bold="true" box="[1264,1426,1865,1890]" pageId="11" pageNumber="362">PARATYPES</emphasis>
</typeStatus>
:
<collectingCountry id="F34A7652FF8C1A74FF04F8CDBC7AF84B" box="[151,379,1901,1926]" name="South Ossetia" pageId="11" pageNumber="362">
<emphasis id="B929EAD0FF8C1A74FF04F8CDBC7AF84B" bold="true" box="[151,379,1901,1926]" pageId="11" pageNumber="362">SOUTH OSSETIA</emphasis>
</collectingCountry>
: same data as
<typeStatus id="54E68860FF8C1A74FDBBF8CDBF8BF84B" box="[552,650,1901,1926]" pageId="11" pageNumber="362" type="holotype">holotype</typeStatus>
(
<specimenCount id="9D5BFD4BFF8C1A74FD0FF8CEBFC8F848" box="[668,713,1902,1926]" pageId="11" pageNumber="362" type="generic">7 ♂</specimenCount>
, 5 ♀,
<collectionCode id="ED4CAE07FF8C1A74FC85F8CEBE49F848" box="[790,840,1902,1925]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="11" pageNumber="362">ZIN</collectionCode>
;
<specimenCount id="9D5BFD4BFF8C1A74FCC5F8CEBE85F848" box="[854,900,1902,1925]" pageId="11" pageNumber="362" type="generic">7 ♂</specimenCount>
, 5 ♀,
<collectionCode id="ED4CAE07FF8C1A74FC5CF8CEB90DF84B" box="[975,1036,1902,1926]" pageId="11" pageNumber="362">KSU</collectionCode>
); Central Caucasus,
<collectingCountry id="F34A7652FF8C1A74FB6BF8CDB899F84B" box="[1272,1432,1901,1926]" name="South Ossetia" pageId="11" pageNumber="362">South Ossetia</collectingCountry>
, Gudiysskiy ridge, source of River Britadon, northern slope of the Mountain Mangavtsek,
<quantity id="4CA59B27FF8C1A74FB61F832B850F864" box="[1266,1361,1938,1962]" metricMagnitude="3" metricUnit="m" metricValue="2.7" pageId="11" pageNumber="362" unit="m" value="2700.0">2700 m</quantity>
a.s.l.,
<date id="FFE31002FF8C1A74FF04F815BC18F803" box="[151,281,1973,1998]" pageId="11" pageNumber="362" value="2013-07-21">21.vii.2013</date>
, D. Zherebilo (
<specimenCount id="9D5BFD4BFF8C1A74FE50F816BCEEF800" box="[451,495,1974,1998]" pageId="11" pageNumber="362" type="generic">2 ♂</specimenCount>
, 5 ♀,
<collectionCode id="ED4CAE07FF8C1A74FDA0F816BF71F803" box="[563,624,1974,1998]" pageId="11" pageNumber="362">KSU</collectionCode>
).
<collectingCountry id="F34A7652FF8C1A74FD11F815BE08F803" box="[642,777,1973,1998]" name="Georgia" pageId="11" pageNumber="362">
<emphasis id="B929EAD0FF8C1A74FD11F815BE08F803" bold="true" box="[642,777,1973,1998]" pageId="11" pageNumber="362">GEORGIA</emphasis>
</collectingCountry>
: Khevsureti, south-western slope of Mountain Tanie,
<quantity id="4CA59B27FF8C1A72FAF7F815BDACFF62" lastPageId="13" lastPageNumber="364" metricMagnitude="3" metricUnit="m" metricValue="2.85" pageId="11" pageNumber="362" unit="m" value="2850.0">2850 m</quantity>
a.s.l.,
<date id="FFE31002FF8A1A72FF6BFF37BC7BFF7D" box="[248,378,151,176]" pageId="13" pageNumber="364" value="2014-07-19">19.vii.2014</date>
, D. Zherebilo (
<specimenCount id="9D5BFD4BFF8A1A72FDBAFF38BF55FF62" box="[553,596,152,176]" pageId="13" pageNumber="364" type="generic">8 ♂</specimenCount>
, 3 ♀,
<collectionCode id="ED4CAE07FF8A1A72FD0EFF38BFDBFF7D" box="[669,730,152,176]" pageId="13" pageNumber="364">KSU</collectionCode>
); Khevsureti, the northern slope of Kistanistavi,
<quantity id="4CA59B27FF8A1A72FA91FF38B858FF62" box="[1282,1369,152,176]" metricMagnitude="3" metricUnit="m" metricValue="2.83" pageId="13" pageNumber="364" unit="m" value="2830.0">2830 m</quantity>
a.s.l.,
<date id="FFE31002FF8A1A72FF04FF1CBC18FF18" box="[151,281,188,213]" pageId="13" pageNumber="364" value="2014-07-21">21.vii.2014</date>
, D. Zherebilo (7 ♀,
<collectionCode id="ED4CAE07FF8A1A72FE6FFF1CBF38FF19" box="[508,569,188,212]" pageId="13" pageNumber="364">KSU</collectionCode>
); same lokality,
<quantity id="4CA59B27FF8A1A72FD7DFF1CBE42FF19" box="[750,835,188,212]" metricMagnitude="3" metricUnit="m" metricValue="2.9" pageId="13" pageNumber="364" unit="m" value="2900.0">2900 m</quantity>
a.s.l.,
<date id="FFE31002FF8A1A72FC1FFF1CB90FFF18" box="[908,1038,188,213]" pageId="13" pageNumber="364" value="2013-07-21">21.vii.2013</date>
, D. Zherebilo (104 ♀,
<collectionCode id="ED4CAE07FF8A1A72FA9EFF1CB84BFF19" box="[1293,1354,188,212]" pageId="13" pageNumber="364">KSU</collectionCode>
).
</paragraph>
</subSubSection>
<caption id="DF22664AFF8B1A73FF04F831BC65F829" httpUri="https://zenodo.org/record/243488/files/figure.png" pageId="12" pageNumber="363" targetBox="[247,1322,214,1871]" targetPageId="12">
<paragraph id="8BE236C2FF8B1A73FF04F831BC65F829" blockId="12.[151,1437,1937,2020]" pageId="12" pageNumber="363">
<emphasis id="B929EAD0FF8B1A73FF04F831BC4DF86A" bold="true" box="[151,332,1937,1959]" pageId="12" pageNumber="363">FIGURES 1217</emphasis>
. Male terminalia of
<taxonomicName id="4C5D4D41FF8B1A73FD8FF831BFA7F86A" box="[540,678,1937,1959]" class="Insecta" family="Empididae" genus="Wiedemannia" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="363" phylum="Arthropoda" rank="genus">
<emphasis id="B929EAD0FF8B1A73FD8FF831BFA7F86A" box="[540,678,1937,1959]" italics="true" pageId="12" pageNumber="363">Wiedemannia</emphasis>
</taxonomicName>
, lateral view.
<emphasis id="B929EAD0FF8B1A73FCA7F831BE4CF86A" bold="true" box="[820,845,1937,1959]" pageId="12" pageNumber="363">12</emphasis>
,
<taxonomicName id="4C5D4D41FF8B1A73FCCAF832BEEFF86A" box="[857,1006,1937,1959]" class="Insecta" family="Empididae" genus="Wiedemannia" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="363" phylum="Arthropoda" rank="species" species="pseguashae" status="sp. nov.">
<emphasis id="B929EAD0FF8B1A73FCCAF832BEEFF86A" box="[857,1006,1937,1959]" italics="true" pageId="12" pageNumber="363">W. pseguashae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21A57ABFF8B1A73FC66F831B947F86A" box="[1013,1094,1937,1959]" pageId="12" pageNumber="363" rank="species">
<emphasis id="B929EAD0FF8B1A73FC66F831B947F86A" bold="true" box="[1013,1094,1937,1959]" pageId="12" pageNumber="363">sp. nov.</emphasis>
</taxonomicNameLabel>
;
<emphasis id="B929EAD0FF8B1A73FBC0F831B96DF86A" bold="true" box="[1107,1132,1937,1959]" pageId="12" pageNumber="363">13</emphasis>
,
<taxonomicName id="4C5D4D41FF8B1A73FBEAF832B873F86A" authority="Joost" authorityName="Joost" box="[1145,1394,1937,1959]" class="Insecta" family="Empididae" genus="Wiedemannia" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="363" phylum="Arthropoda" rank="species" species="pseudovaillanti">
<emphasis id="B929EAD0FF8B1A73FBEAF832B834F86A" box="[1145,1333,1937,1959]" italics="true" pageId="12" pageNumber="363">W. pseudovaillanti</emphasis>
Joost
</taxonomicName>
;
<emphasis id="B929EAD0FF8B1A73FAEFF831B894F86A" bold="true" box="[1404,1429,1937,1959]" pageId="12" pageNumber="363">14</emphasis>
,
<taxonomicName id="4C5D4D41FF8B1A73FF04F811BF01F80B" authority="Kustov &amp; Zherebilo" authorityName="Kustov &amp; Zherebilo" box="[151,512,1968,1990]" class="Insecta" family="Empididae" genus="Wiedemannia" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="363" phylum="Arthropoda" rank="species" species="shamshevi">
<emphasis id="B929EAD0FF8B1A73FF04F811BC23F80B" box="[151,290,1968,1990]" italics="true" pageId="12" pageNumber="363">W. shamshevi</emphasis>
Kustov &amp; Zherebilo
</taxonomicName>
;
<emphasis id="B929EAD0FF8B1A73FD9EF810BF27F80B" bold="true" box="[525,550,1968,1990]" pageId="12" pageNumber="363">15</emphasis>
,
<taxonomicName id="4C5D4D41FF8B1A73FDA6F811BE8BF80B" authority="Kustov &amp; Zherebilo" authorityName="Kustov &amp; Zherebilo" box="[565,906,1968,1990]" class="Insecta" family="Empididae" genus="Wiedemannia" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="363" phylum="Arthropoda" rank="species" species="sinclairi">
<emphasis id="B929EAD0FF8B1A73FDA6F811BFADF80B" box="[565,684,1968,1990]" italics="true" pageId="12" pageNumber="363">W. sinclairi</emphasis>
Kustov &amp; Zherebilo
</taxonomicName>
;
<emphasis id="B929EAD0FF8B1A73FC05F810BEAEF80B" bold="true" box="[918,943,1968,1990]" pageId="12" pageNumber="363">16</emphasis>
,
<taxonomicName id="4C5D4D41FF8B1A73FC2CF811B974F80B" authority="Joost" authorityName="Joost" box="[959,1141,1968,1990]" class="Insecta" family="Empididae" genus="Wiedemannia" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="363" phylum="Arthropoda" rank="species" species="vaillanti">
<emphasis id="B929EAD0FF8B1A73FC2CF811B934F80B" box="[959,1077,1968,1990]" italics="true" pageId="12" pageNumber="363">W. vaillanti</emphasis>
Joost
</taxonomicName>
;
<emphasis id="B929EAD0FF8B1A73FB10F810B99DF80B" bold="true" box="[1155,1180,1968,1990]" pageId="12" pageNumber="363">17</emphasis>
,
<taxonomicName id="4C5D4D41FF8B1A73FB38F811B899F80B" authority="Fallen" authorityName="Fallen" box="[1195,1432,1968,1990]" class="Insecta" family="Empididae" genus="Wiedemannia" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="363" phylum="Arthropoda" rank="species" species="zetterstedti">
<emphasis id="B929EAD0FF8B1A73FB38F811B9C4F80B" box="[1195,1221,1969,1990]" italics="true" pageId="12" pageNumber="363">W.</emphasis>
<emphasis id="B929EAD0FF8B1A73FB5EF810B83DF80B" box="[1229,1340,1968,1990]" italics="true" pageId="12" pageNumber="363">zetterstedti</emphasis>
(Fallén)
</taxonomicName>
. Scale bar = 0.1 mm.
</paragraph>
</caption>
<subSubSection id="C3476549FF8A1A72FF54FF7FB918FEA8" pageId="13" pageNumber="364" type="diagnosis">
<paragraph id="8BE236C2FF8A1A72FF54FF7FB918FEA8" blockId="13.[151,1437,151,1658]" pageId="13" pageNumber="364">
<emphasis id="B929EAD0FF8A1A72FF54FF7FBC3BFF35" bold="true" box="[199,314,223,248]" pageId="13" pageNumber="364">Diagnosis</emphasis>
. This species is distinguished by acrostichals divergent around the prescutellar depression, 67 pairs of dorsocentrals with interspersed numerous shorter setae, dark brown scutum with light brown pruinescence on prescutellar depression; 2 anterior setae on fore femur and unilobed, elongate, narrowed apically clasping cercus with inner numerous setae on posterior margin, becoming longer on lower third.
</paragraph>
</subSubSection>
<subSubSection id="C3476549FF8A1A72FF54FECFBC7AFAFB" pageId="13" pageNumber="364" type="description">
<paragraph id="8BE236C2FF8A1A72FF54FECFBC7AFAFB" blockId="13.[151,1437,151,1658]" pageId="13" pageNumber="364">
<emphasis id="B929EAD0FF8A1A72FF54FECFBC50FE45" bold="true" box="[199,337,367,392]" pageId="13" pageNumber="364">Description</emphasis>
.
<emphasis id="B929EAD0FF8A1A72FEF2FECFBC9CFE45" bold="true" box="[353,413,367,392]" pageId="13" pageNumber="364">Male</emphasis>
. Body length
<quantity id="4CA59B27FF8A1A72FDD3FECFBFCDFE4A" box="[576,716,367,392]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" metricValueMax="5.1" metricValueMin="4.5" pageId="13" pageNumber="364" unit="mm" value="4.8" valueMax="5.1" valueMin="4.5">4.55.1 mm</quantity>
; wing length 5.06.0 mm.
<emphasis id="B929EAD0FF8A1A72FC6DFECFB93CFE45" bold="true" box="[1022,1085,367,392]" pageId="13" pageNumber="364">Head</emphasis>
brown, face, occiput and gena with blue pruinescence, frons and vertex with olive-brown pruinescence. Gena broad, about 1/4 height of eye. Face wide, distinct carina on lower margin. Pair of strong ocellar setae (equal postpronotal setae) and pair of vertical setae, several strong occipitals similarly sized and black, with interspersed shorter postoculars. Lower postocular setae much finer and paler merging with pale, longer setae on middle and lower occiput. Few dark setulae present on vertex and between ocellar area and eye margin. Antenna dark brown to black, postpedicel and stylus minutely pubescent; scape with setulae dorsally, pedicel almost equal in size to scape, with complete circlet of apical setae; postpedicel longer, apically pointed, stylus twice as long as postpedicel, tip blunt. Palpus with numerous dark dorsal setae.
<emphasis id="B929EAD0FF8A1A72FEBDFD2FBC98FD65" bold="true" box="[302,409,655,680]" pageId="13" pageNumber="364">Thoracic</emphasis>
pleura with blue pruinescence; scutum dark brown with indistinct darker strips between rows of acrostichals and dorsocentrals; prescutellar depression with light brown pruinescence, extending onto concolourous scutellum; postpronotal lobe and lower margin of notopleuron with lighter pruinescence. Antepronotum with pair of dark setae and some fine setulae. Proepisternum with whitish fine setulae. Acrostichals small, irregularly biserial, divergent around prescutellar depression; 67 pairs of dorsocentrals with numerous interspersed shorter setae; 1 postpronotal seta (with several short setae), 2 notopleural setae (upper stronger), 1 presutural supra-alar seta and with numerous scattered setulae; 1 postalar seta; 1 strong scutellar setae and 2 weak marginal setae. Laterotergite with patch of fine, dark setulae.
<emphasis id="B929EAD0FF8A1A72FCC7FC2CBE8DFC68" bold="true" box="[852,908,908,933]" pageId="13" pageNumber="364">Legs</emphasis>
entirely dark brown to black, coxae with blue pruinescence. All legs with evenly distributed setae; hind tibia with longer dorsal and ventral setae on apical third. All coxae with longer setae anteriorly. Fore femur narrowed apically, with 2 stout, closely approximated anterior setae on apical third and 1 same seta near tip.
<emphasis id="B929EAD0FF8A1A72FD5AFC57BE0BFBDD" bold="true" box="[713,778,1015,1040]" pageId="13" pageNumber="364">Wing</emphasis>
membrane infuscate, veins brown; pterostigma distinct, elongate, brown; single basal costal seta. Origin of veins M1 and M2 widely separated at end of cell dm (some specimens with veins M1 and M2 form additional cell after dm). R4 and R5 straight and divergent. Halter dark brown with blue pruinescence.
<emphasis id="B929EAD0FF8A1A72FE39FBC9BF31FB4F" bold="true" box="[426,560,1129,1154]" pageId="13" pageNumber="364">Abdominal</emphasis>
sclerites with blue pruinescence. Terminalia (
<figureCitation id="13662A47FF8A1A72FBABFBC9B98FFB4F" box="[1080,1166,1129,1154]" captionStart="FIGURES 12 17" captionStartId="12.[151,263,1937,1959]" captionTargetBox="[247,1322,214,1871]" captionTargetId="figure@12.[229,1337,193,1891]" captionTargetPageId="12" captionText="FIGURES 12 17. Male terminalia of Wiedemannia, lateral view. 12, W. pseguashae sp. nov.; 13, W. pseudovaillanti Joost; 14, W. shamshevi Kustov &amp; Zherebilo; 15, W. sinclairi Kustov &amp; Zherebilo; 16, W. vaillanti Joost; 17, W. zetterstedti (Fallén). Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/243488/files/figure.png" pageId="13" pageNumber="364">Fig. 12</figureCitation>
): Hypandrium subequal in length to epandrium, bearing several setae. Epandrium rhomboid, covered with numerous dark long setae especially ventrally and laterally. Clasping cercus unilobed, elongated, narrowed apically clothed with fine setulae; inner surface of posterior margin with numerous setae, becoming longer on lower third. Phallus more or less linear, brownish; distiphallus without swelling in middle.
<emphasis id="B929EAD0FF8A1A72FD4BFB59BE2EFADF" bold="true" box="[728,815,1273,1298]" pageId="13" pageNumber="364">Female</emphasis>
. Similar to male, except terminalia; cercus short ovate and minutely pilose.
</paragraph>
</subSubSection>
<subSubSection id="C3476549FF8A1A72FF54FAE1BE3FFA97" box="[199,830,1345,1370]" pageId="13" pageNumber="364" type="distribution">
<paragraph id="8BE236C2FF8A1A72FF54FAE1BE3FFA97" blockId="13.[151,1437,151,1658]" box="[199,830,1345,1370]" pageId="13" pageNumber="364">
<emphasis id="B929EAD0FF8A1A72FF54FAE1BC58FA97" bold="true" box="[199,345,1345,1370]" pageId="13" pageNumber="364">Distribution</emphasis>
. PALAEARCTIC:
<collectingCountry id="F34A7652FF8A1A72FDA7FAE1BFD0FA97" box="[564,721,1345,1370]" name="South Ossetia" pageId="13" pageNumber="364">South Ossetia</collectingCountry>
,
<collectingCountry id="F34A7652FF8A1A72FD4EFAE1BE3BFA97" box="[733,826,1345,1370]" name="Georgia" pageId="13" pageNumber="364">Georgia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3476549FF8A1A72FF54FAC5BC7DF9FF" pageId="13" pageNumber="364" type="discussion">
<paragraph id="8BE236C2FF8A1A72FF54FAC5BE24FA27" blockId="13.[151,1437,151,1658]" pageId="13" pageNumber="364">
<emphasis id="B929EAD0FF8A1A72FF54FAC5BC35FAB3" bold="true" box="[199,308,1381,1406]" pageId="13" pageNumber="364">Remarks</emphasis>
.
<taxonomicName id="4C5D4D41FF8A1A72FED6FAC5BF6BFAB0" box="[325,618,1381,1405]" class="Insecta" family="Empididae" genus="Wiedemannia" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="364" phylum="Arthropoda" rank="species" species="pseguashae" status="sp. nov.">
<emphasis id="B929EAD0FF8A1A72FED6FAC5BF6BFAB0" box="[325,618,1381,1405]" italics="true" pageId="13" pageNumber="364">Wiedemannia pseguashae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21A57ABFF8A1A72FDE7FAC5BFD0FAB3" box="[628,721,1381,1406]" pageId="13" pageNumber="364" rank="species">
<emphasis id="B929EAD0FF8A1A72FDE7FAC5BFD0FAB3" bold="true" box="[628,721,1381,1406]" pageId="13" pageNumber="364">sp. nov.</emphasis>
</taxonomicNameLabel>
is most similar to
<taxonomicName id="4C5D4D41FF8A1A72FC3CFAC6B89DFAB3" authority="Vaillant 1967" authorityName="Vaillant" authorityYear="1967" box="[943,1436,1381,1406]" class="Insecta" family="Empididae" genus="Wiedemannia" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="364" phylum="Arthropoda" rank="species" species="pyrenaica" subGenus="Chamaedipsia">
<emphasis id="B929EAD0FF8A1A72FC3CFAC6BECBFAB0" box="[943,970,1382,1405]" italics="true" pageId="13" pageNumber="364">W.</emphasis>
(
<emphasis id="B929EAD0FF8A1A72FC47FAC5B977FAB0" box="[980,1142,1381,1405]" italics="true" pageId="13" pageNumber="364">Chamaedipsia</emphasis>
)
<emphasis id="B929EAD0FF8A1A72FB19FAC6B9F8FAB0" box="[1162,1273,1382,1405]" italics="true" pageId="13" pageNumber="364">pyrenaica</emphasis>
<bibRefCitation id="EFCC4B33FF8A1A72FA90FAC5B89DFAB3" author="Vaillant" box="[1283,1436,1381,1406]" pageId="13" pageNumber="364" refString="Vaillant, F. (1967) La repartition des Wiedemannia dans les cours d'eau et leur utilisation comme indicateurs de zones ecologiques (Diptera, Empididae). Annales de Limnologie, 3, 267 - 293. http: // dx. doi. org / 10.1051 / limn / 1967016" type="journal article" year="1967">Vaillant 1967</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C5D4D41FF8A1A72FF59FA2ABF02FA6C" box="[202,515,1417,1441]" class="Insecta" family="Empididae" genus="Wiedemannia" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="364" phylum="Arthropoda" rank="species" species="beckeri" subGenus="Chamaedipsia">
<emphasis id="B929EAD0FF8A1A72FF59FA2ABF02FA6C" box="[202,515,1417,1441]" italics="true" pageId="13" pageNumber="364">W. (Chamaedipsia) beckeri</emphasis>
</taxonomicName>
, but differs by the structure of the male terminalia, especially the form of the clasping cercus and distribution of inner setae.
<taxonomicName id="4C5D4D41FF8A1A72FD32FA0DBEC2FA08" box="[673,963,1453,1477]" class="Insecta" family="Empididae" genus="Wiedemannia" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="364" phylum="Arthropoda" rank="species" species="pseguashae" status="sp. nov.">
<emphasis id="B929EAD0FF8A1A72FD32FA0DBEC2FA08" box="[673,963,1453,1477]" italics="true" pageId="13" pageNumber="364">Wiedemannia pseguashae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21A57ABFF8A1A72FC5AFA0DB923FA0B" box="[969,1058,1453,1478]" pageId="13" pageNumber="364" rank="species">
<emphasis id="B929EAD0FF8A1A72FC5AFA0DB923FA0B" bold="true" box="[969,1058,1453,1478]" pageId="13" pageNumber="364">sp. nov.</emphasis>
</taxonomicNameLabel>
can be differentiated from related new species described in this article as outlined in the key.
</paragraph>
<paragraph id="8BE236C2FF8A1A72FF54FA55BC7DF9FF" blockId="13.[151,1437,151,1658]" pageId="13" pageNumber="364">
The new species is known only from high-altitude streams and rivers of Central Caucasus (
<quantity id="4CA59B27FF8A1A72FB2FFA55B859F9C0" box="[1212,1368,1525,1550]" metricMagnitude="3" metricUnit="m" metricValue="2.75" metricValueMax="3.0" metricValueMin="2.5" pageId="13" pageNumber="364" unit="m" value="2750.0" valueMax="3000.0" valueMin="2500.0">25003000 m</quantity>
a.s.l.) and is active in July.
</paragraph>
</subSubSection>
<subSubSection id="C3476549FF8A1A72FF54F99DBC26F9B7" pageId="13" pageNumber="364" type="etymology">
<paragraph id="8BE236C2FF8A1A72FF54F99DBC26F9B7" blockId="13.[151,1437,151,1658]" pageId="13" pageNumber="364">
<emphasis id="B929EAD0FF8A1A72FF54F99DBC47F99B" bold="true" box="[199,326,1597,1622]" pageId="13" pageNumber="364">Etymology</emphasis>
. The epithet refers to “Pseguash” (maid of rivers), a pagan goddess in the mythology of ancient Cherkessian.
</paragraph>
</subSubSection>
</treatment>
</document>