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<document ID-DOI="10.11646/zootaxa.5277.2.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="7889695" ID-ZooBank="73ED69F3-3887-460D-86DE-C9F1302C8EC3" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1683099613832" checkinUser="plazi" docAuthor="Kluge, Nikita, Srinivasan, Pandiarajan, Sivaruban, T., Barathy, S. &amp; Isack, Rajasekaran" docDate="2023" docId="03F487D0A3524D63A2DBFEBAFD7CE925" docLanguage="en" docName="zootaxa.5277.2.1.pdf" docOrigin="Zootaxa 5277 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5277.2.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Baetis (Tenuibaetis) bialatus Kluge &amp; Srinivasan &amp; Sivaruban &amp; Barathy &amp; Isack 2023, sp. n." docType="treatment" docVersion="2" lastPageNumber="246" masterDocId="FFCDFFA8A3764D4EA24CFFD4FFABE864" masterDocTitle="Contribution to the knowledge of the subgenus Tenuibaetis Kang &amp; Yang 1994 (Ephemeroptera, Baetidae, Baetis s. l.)" masterLastPageNumber="258" masterPageNumber="201" pageNumber="237" updateTime="1683100342607" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Contribution to the knowledge of the subgenus Tenuibaetis Kang &amp; Yang 1994 (Ephemeroptera, Baetidae, Baetis s. l.)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kluge, Nikita</mods:namePart>
<mods:affiliation>Department of Entomology, Saint Petersburg State University, Universitetskaya nab., 7 / 9, Saint Petersburg, 199034, Russia. Website: http: // insecta. bio. spbu. ru</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Srinivasan, Pandiarajan</mods:namePart>
<mods:affiliation>PG and Research department of Zoology, the American College, Madurai- 625002, India</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sivaruban, T.</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0001-8997-9355</mods:nameIdentifier>
<mods:affiliation>PG and Research department of Zoology, the American College, Madurai- 625002, India &amp; sivaruban 270 @ gmail. com; https: // orcid. org / 0000 - 0001 - 8997 - 9355</mods:affiliation>
<mods:nameIdentifier type="email">sivaruban270@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Barathy, S.</mods:namePart>
<mods:affiliation>Department of Zoology, Fatima College, Madurai- 625018, India</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Isack, Rajasekaran</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-9952-4335</mods:nameIdentifier>
<mods:affiliation>PG and Research department of Zoology, the American College, Madurai- 625002, India &amp; iceisack 143 @ gmail. com; https: // orcid. org / 0000 - 0002 - 9952 - 4335</mods:affiliation>
<mods:nameIdentifier type="email">iceisack143@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zootaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-05-02</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>5277</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>2</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>201</mods:start>
<mods:end>258</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.11646/zootaxa.5277.2.1</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/zootaxa.5277.2.1</mods:identifier>
<mods:identifier type="ISSN">1175-5326</mods:identifier>
<mods:identifier type="Zenodo-Dep">7889695</mods:identifier>
<mods:identifier type="ZooBank">73ED69F3-3887-460D-86DE-C9F1302C8EC3</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7889717" ID-Zenodo-Dep="7889717" LSID="urn:lsid:plazi:treatment:03F487D0A3524D63A2DBFEBAFD7CE925" httpUri="http://treatment.plazi.org/id/03F487D0A3524D63A2DBFEBAFD7CE925" lastPageId="45" lastPageNumber="246" pageId="36" pageNumber="237">
<subSubSection box="[151,598,366,393]" pageId="36" pageNumber="237" type="nomenclature">
<paragraph blockId="36.[151,598,366,429]" box="[151,598,366,393]" pageId="36" pageNumber="237">
<heading bold="true" box="[151,598,366,393]" fontSize="11" level="1" pageId="36" pageNumber="237" reason="1">
<emphasis bold="true" box="[151,598,366,393]" pageId="36" pageNumber="237">
6.
<taxonomicName authority="Kluge &amp; Srinivasan &amp; Sivaruban &amp; Barathy &amp; Isack, 2023" authorityName="Kluge &amp; Srinivasan &amp; Sivaruban &amp; Barathy &amp; Isack" authorityYear="2023" box="[181,525,366,393]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="36" pageNumber="237" phylum="Arthropoda" rank="species" species="bialatus" status="sp. n." subGenus="Tenuibaetis">
<emphasis bold="true" box="[181,256,367,393]" italics="true" pageId="36" pageNumber="237">Baetis</emphasis>
(
<emphasis bold="true" box="[273,412,366,393]" italics="true" pageId="36" pageNumber="237">Tenuibaetis</emphasis>
)
<emphasis bold="true" box="[429,525,366,393]" italics="true" pageId="36" pageNumber="237">bialatus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[532,598,367,393]" pageId="36" pageNumber="237" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[151,463,403,429]" pageId="36" pageNumber="237" type="description">
<paragraph blockId="36.[151,598,366,429]" box="[151,463,403,429]" pageId="36" pageNumber="237">
(
<figureCitation box="[159,259,403,429]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1916,1940]" captionTargetBox="[151,1436,885,1892]" captionTargetId="figure-386@2.[151,1436,885,1892]" captionTargetPageId="2" captionText="FIGURES 1–4. Setae on larval legs of Baetis (Tenuibaetis) bialatus sp. n.. 1, two-channel and hirudiform setae on outer side of middle tibia under scanning electron microscope; 2–3, hirudiform seta; 4, two-channel seta on anterior side of femur in transmitted light. Abbreviations: 2-can, two-channel seta; hirud, hirudiform seta." figureDoi="http://doi.org/10.5281/zenodo.7889697" httpUri="https://zenodo.org/record/7889697/files/figure.png" pageId="36" pageNumber="237">Figs 1–4</figureCitation>
,
<figureCitation box="[272,344,403,429]" captionStart="FIGURES 13–21" captionStartId="5.[151,264,1840,1864]" captionTargetBox="[151,1436,181,1816]" captionTargetId="figure-19@5.[151,1436,181,1816]" captionTargetPageId="5" captionText="FIGURES 13–21. Imaginal hind wings and larval protoptera (under the same magnification). 13–14, female Baetis (Tenuibaetis) frequentus from India; 15–17, male B. (T.) octomaculatus sp. n.; 18–19, female of same species; 20–21, male B. (T.) bialatus sp. n. (13, 15, 20, metathorax of imago with hind wing; 14, 21, larval exuviae of metanotum; 16, 18, hind wing of imago; 17, 19, cuticle from metathorax of larva ready to molt to subimago, with subimaginal wing inside protopteron: crumpled on Fig. 17, non-crumpled on Fig. 19)." figureDoi="http://doi.org/10.5281/zenodo.7889705" httpUri="https://zenodo.org/record/7889705/files/figure.png" pageId="36" pageNumber="237">20–21</figureCitation>
,
<figureCitation box="[356,455,403,429]" captionStart-0="FIGURES 184–190" captionStart-1="FIGURES 191–194" captionStart-2="FIGURES 195–201" captionStart-3="FIGURES 202–207" captionStart-4="FIGURES 208–210" captionStart-5="FIGURES211–214" captionStart-6="FIGURES 215–224" captionStart-7="FIGURES 225–229" captionStartId-0="37.[151,264,1745,1769]" captionStartId-1="38.[151,264,1802,1826]" captionStartId-2="39.[151,264,1840,1864]" captionStartId-3="40.[151,264,1840,1864]" captionStartId-4="41.[151,264,1840,1864]" captionStartId-5="42.[151,264,1498,1522]" captionStartId-6="43.[151,264,1770,1794]" captionStartId-7="44.[151,264,1840,1864]" captionTargetBox-0="[151,1436,181,1721]" captionTargetBox-1="[151,1430,181,1778]" captionTargetBox-2="[151,1436,190,1817]" captionTargetBox-3="[151,1396,181,1811]" captionTargetBox-4="[151,1435,181,1817]" captionTargetBox-5="[151,1436,181,1469]" captionTargetBox-6="[151,1435,181,1746]" captionTargetBox-7="[151,1436,185,1817]" captionTargetId-0="figure-19@37.[151,1436,181,1721]" captionTargetId-1="figure-36@38.[151,1436,181,1778]" captionTargetId-2="figure-19@39.[151,1436,181,1817]" captionTargetId-3="figure-36@40.[151,1436,181,1816]" captionTargetId-4="figure-19@41.[151,1435,181,1817]" captionTargetId-5="figure-36@42.[151,1436,181,1474]" captionTargetId-6="figure-19@43.[151,1435,181,1746]" captionTargetId-7="figure-36@44.[151,1436,181,1817]" captionTargetPageId-0="37" captionTargetPageId-1="38" captionTargetPageId-2="39" captionTargetPageId-3="40" captionTargetPageId-4="41" captionTargetPageId-5="42" captionTargetPageId-6="43" captionTargetPageId-7="44" captionText-0="FIGURES 184–190. Baetis (Tenuibaetis) bialatus sp. n., larval exuviae (one and the same magnification). 184–185, abdomen; 186, head with labrum; 187, half of pronotum and mesonotum; 188–190, fore, middle and hind legs." captionText-1="FIGURES 191–194. Baetis (Tenuibaetis) bialatus sp. n., larval mouth parts. 191, labrum; 192, hypopharynx with superlinguae; 193, maxilla; 194, labium." captionText-2="FIGURES 195–201. Baetis (Tenuibaetis) bialatus sp. n., larvae. 195–196, left and right mandibles with developing mandibles of next instar inside; 197–198, the same of another individual; 199, exuviae of abdominal tergum IX; 200, abdominal tergum II; 201, paraprocts. Abbreviations: hirud, hirudiform seta; sq, scale." captionText-3="FIGURES 202–207. Baetis (Tenuibaetis) bialatus sp. n., larval legs. 202, middle femur (anterior side); 203, fore tibia (anterior side); 204, middle tibia (anterior side); 205, middle tibia (outer side); 206–207, tibia and tarsus of fore and middle legs (anterior side) (202–204, 206–207, holotype). Abbreviations: 2-can, two-channel seta." captionText-4="FIGURES 208–210. Baetis (Tenuibaetis) bialatus sp. n., proximal, middle and distal portions of larval femur (view from outer side).Abbreviations: 2-can, two-channel seta; ant-ap, anterior-apical margin of femur; hirud, hirudiform seta; post-ap, posteriorapical margin of femur." captionText-5="FIGURES211–214. Baetis (Tenuibaetis) bialatus sp.n., larvaltibiae(view fromouter side).211–213,fore tibia;214,middle tibia." captionText-6="FIGURES 215–224. Baetis (Tenuibaetis) bialatus sp. n. 215, subimaginal exuviae of mesopleuron; 216, subimaginal exuviae of half of mesonotum; 217–218, exuviae of fore and hind legs of male subimago; 219–220, fore and middle legs of male imago; 221, male imago; 222, female imago; 223, subimaginal exuviae of abdominal segments VIII–IX; 224, egg (215–220, holotype)." captionText-7="FIGURES 225–229. Baetis (Tenuibaetis) bialatus sp. n., genitalia of male imago. 225, 227, ventral view; 226, 229, ventral view with focus on sterno-styligeral muscle; 228, enlarged fragment of sterno-styligeral muscle to show cross-striation (225–226, holotype). Abbreviations: gv, gonovectis; s.s, sterno-styligeral muscle." figureDoi-0="http://doi.org/10.5281/zenodo.7889781" figureDoi-1="http://doi.org/10.5281/zenodo.7889783" figureDoi-2="http://doi.org/10.5281/zenodo.7889785" figureDoi-3="http://doi.org/10.5281/zenodo.7889787" figureDoi-4="http://doi.org/10.5281/zenodo.7889789" figureDoi-5="http://doi.org/10.5281/zenodo.7889791" figureDoi-6="http://doi.org/10.5281/zenodo.7889793" figureDoi-7="http://doi.org/10.5281/zenodo.7889797" httpUri-0="https://zenodo.org/record/7889781/files/figure.png" httpUri-1="https://zenodo.org/record/7889783/files/figure.png" httpUri-2="https://zenodo.org/record/7889785/files/figure.png" httpUri-3="https://zenodo.org/record/7889787/files/figure.png" httpUri-4="https://zenodo.org/record/7889789/files/figure.png" httpUri-5="https://zenodo.org/record/7889791/files/figure.png" httpUri-6="https://zenodo.org/record/7889793/files/figure.png" httpUri-7="https://zenodo.org/record/7889797/files/figure.png" pageId="36" pageNumber="237">184–229</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection box="[151,1241,475,501]" pageId="36" pageNumber="237" type="etymology">
<paragraph blockId="36.[151,1437,475,2013]" box="[151,1241,475,501]" pageId="36" pageNumber="237">
<emphasis bold="true" box="[151,286,475,501]" pageId="36" pageNumber="237">Etymology.</emphasis>
Allusion of complete absence of hind wings, so that only two fore wings are present.
</paragraph>
</subSubSection>
<subSubSection lastPageId="37" lastPageNumber="238" pageId="36" pageNumber="237" type="materials_examined">
<paragraph blockId="36.[151,1437,475,2013]" pageId="36" pageNumber="237">
<emphasis bold="true" box="[199,425,510,536]" pageId="36" pageNumber="237">Material examined</emphasis>
(
<collectionCode box="[439,491,511,537]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="36" pageNumber="237" type="Museum">ZIN</collectionCode>
).
<materialsCitation collectingDate="2016-02-06" collectorName="N. Kluge &amp; L. Sheyko." country="India" county="Palni Hills" location="Levinge" municipality="Kodaikanal" pageId="36" pageNumber="237" specimenCount="1" stateProvince="Tamil Nadu" typeStatus="holotype">
<emphasis bold="true" box="[510,628,511,537]" pageId="36" pageNumber="237">
<typeStatus box="[510,623,511,537]" pageId="36" pageNumber="237">Holotype</typeStatus>
:
</emphasis>
L-S-I♁ {specimen [XV](2)2016},
<collectingCountry box="[1023,1105,511,537]" name="India" pageId="36" pageNumber="237">INDIA</collectingCountry>
,
<collectingRegion box="[1115,1307,511,537]" country="India" name="Tamil Nadu" pageId="36" pageNumber="237">Tamil Nadu state</collectingRegion>
,
<collectingCounty box="[1318,1436,511,537]" pageId="36" pageNumber="237">Palni Hills</collectingCounty>
near
<collectingMunicipality box="[206,338,547,573]" pageId="36" pageNumber="237">Kodaikanal</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:03F487D0A3524D63A2DBFEBAFD7CE925:8E82601DA3524D6AA310FDF7FE13EA59" box="[348,440,547,573]" country="India" county="Palni Hills" municipality="Kodaikanal" name="Levinge" pageId="36" pageNumber="237" stateProvince="Tamil Nadu">Levinge</location>
stream downstream of
<location LSID="urn:lsid:plazi:treatment:03F487D0A3524D63A2DBFEBAFD7CE925:8E82601DA3524D6AA0F1FDF7FCF6EA59" box="[701,861,547,573]" country="India" county="Palni Hills" municipality="Kodaikanal" name="Pamparpuram" pageId="36" pageNumber="237" stateProvince="Tamil Nadu">Pamparpuram</location>
,
<date box="[871,976,547,573]" pageId="36" pageNumber="237" value="2016-02-06">
<collectingDate box="[871,976,547,573]" pageId="36" pageNumber="237" value="2016-02-06">6.II.2016</collectingDate>
</date>
, coll.
<collectorName box="[1042,1144,547,573]" pageId="36" pageNumber="237">N. Kluge</collectorName>
&amp;
<collectorName box="[1179,1301,547,573]" pageId="36" pageNumber="237">L. Sheyko.</collectorName>
</materialsCitation>
<materialsCitation collectingDate="2016-01-31" collectingDateMax="2016-02-06" collectingDateMin="2016-01-31" collectorName="N. Kluge &amp; L. Sheyko." country="India" county="Palni Hills" location="Levinge" municipality="Kodaikanal" pageId="36" pageNumber="237" specimenCount="52" specimenCount-female="1" specimenCount-larva="51" stateProvince="Tamil Nadu" typeStatus="paratype">
<emphasis bold="true" box="[1307,1436,547,573]" pageId="36" pageNumber="237">
<typeStatus box="[1307,1431,547,573]" pageId="36" pageNumber="237">Paratypes</typeStatus>
:
</emphasis>
the same locality and collectors,
<date box="[525,690,583,609]" pageId="36" pageNumber="237" value="2016-01-31" valueMax="2016-02-06" valueMin="2016-01-31">
<collectingDate box="[525,690,583,609]" pageId="36" pageNumber="237" value="2016-01-31" valueMax="2016-02-06" valueMin="2016-01-31">31.I–6.II.2016</collectingDate>
</date>
: 4 L-S-I♁, 2 L-S♁, 5 L-S-I
<specimenCount box="[1015,1029,583,609]" pageId="36" pageNumber="237" type="female">♀</specimenCount>
,
<specimenCount box="[1042,1151,582,609]" pageId="36" pageNumber="237" type="larva">51 larvae</specimenCount>
</materialsCitation>
);
<materialsCitation collectingDate="2016-02-05" collectorName="N. Kluge &amp; L. Sheyko" country="India" location="Perumal Malai" municipality="river Palar" pageId="36" pageNumber="237" specimenCount="4" specimenCount-larva="4" stateProvince="Tamil Nadu" typeStatus="paratype">
<collectingMunicipality box="[1172,1291,583,609]" pageId="36" pageNumber="237">river Palar</collectingMunicipality>
downstream
<location LSID="urn:lsid:plazi:treatment:03F487D0A3524D63A2DBFEBAFD7CE925:8E82601DA3524D6AA2DBFDBFFEEBEAE1" box="[151,320,619,645]" country="India" municipality="river Palar" name="Perumal Malai" pageId="36" pageNumber="237" stateProvince="Tamil Nadu">Perumal Malai</location>
,
<date box="[331,436,618,645]" pageId="36" pageNumber="237" value="2016-02-05">
<collectingDate box="[331,436,618,645]" pageId="36" pageNumber="237" value="2016-02-05">5.II.2016</collectingDate>
</date>
, coll.
<collectorName box="[503,606,619,645]" pageId="36" pageNumber="237">N. Kluge</collectorName>
&amp;
<collectorName box="[642,761,619,645]" pageId="36" pageNumber="237">L. Sheyko</collectorName>
:
<specimenCount box="[772,865,619,645]" pageId="36" pageNumber="237" type="larva">4 larvae</specimenCount>
</materialsCitation>
.
</paragraph>
<paragraph blockId="36.[151,1437,475,2013]" box="[199,348,654,680]" pageId="36" pageNumber="237">
<emphasis bold="true" box="[199,348,654,680]" pageId="36" pageNumber="237">Descriptions</emphasis>
</paragraph>
<paragraph blockId="36.[151,1437,475,2013]" pageId="36" pageNumber="237">
<emphasis bold="true" box="[199,275,691,717]" pageId="36" pageNumber="237">
<emphasis bold="true" box="[199,267,691,717]" italics="true" pageId="36" pageNumber="237">Larva</emphasis>
.
</emphasis>
CUTICULAR COLORATION: Head brown (
<figureCitation box="[802,898,691,717]" captionStart="FIGURES 184–190" captionStartId="37.[151,264,1745,1769]" captionTargetBox="[151,1436,181,1721]" captionTargetId="figure-19@37.[151,1436,181,1721]" captionTargetPageId="37" captionText="FIGURES 184–190. Baetis (Tenuibaetis) bialatus sp. n., larval exuviae (one and the same magnification). 184–185, abdomen; 186, head with labrum; 187, half of pronotum and mesonotum; 188–190, fore, middle and hind legs." figureDoi="http://doi.org/10.5281/zenodo.7889781" httpUri="https://zenodo.org/record/7889781/files/figure.png" pageId="36" pageNumber="237">Fig. 186</figureCitation>
). Pronotum nearly uniformly brown; mesonotum brown, darker medially and medio-anteriorly (
<figureCitation box="[678,778,727,753]" captionStart="FIGURES 184–190" captionStartId="37.[151,264,1745,1769]" captionTargetBox="[151,1436,181,1721]" captionTargetId="figure-19@37.[151,1436,181,1721]" captionTargetPageId="37" captionText="FIGURES 184–190. Baetis (Tenuibaetis) bialatus sp. n., larval exuviae (one and the same magnification). 184–185, abdomen; 186, head with labrum; 187, half of pronotum and mesonotum; 188–190, fore, middle and hind legs." figureDoi="http://doi.org/10.5281/zenodo.7889781" httpUri="https://zenodo.org/record/7889781/files/figure.png" pageId="36" pageNumber="237">Fig. 187</figureCitation>
). Thoracic pleura brownish, sterna colorless. Legs brown; fore femur with proximal blank oval (
<figureCitation box="[571,730,763,789]" captionStart="FIGURES 184–190" captionStartId="37.[151,264,1745,1769]" captionTargetBox="[151,1436,181,1721]" captionTargetId="figure-19@37.[151,1436,181,1721]" captionTargetPageId="37" captionText="FIGURES 184–190. Baetis (Tenuibaetis) bialatus sp. n., larval exuviae (one and the same magnification). 184–185, abdomen; 186, head with labrum; 187, half of pronotum and mesonotum; 188–190, fore, middle and hind legs." figureDoi="http://doi.org/10.5281/zenodo.7889781" httpUri="https://zenodo.org/record/7889781/files/figure.png" pageId="36" pageNumber="237">Figs 188–190</figureCitation>
). Abdominal terga nearly uniformly brown; tergum
<collectionCode box="[1307,1337,763,789]" pageId="36" pageNumber="237">IV</collectionCode>
either as dark as others (
<figureCitation box="[322,419,799,825]" captionStart="FIGURES 184–190" captionStartId="37.[151,264,1745,1769]" captionTargetBox="[151,1436,181,1721]" captionTargetId="figure-19@37.[151,1436,181,1721]" captionTargetPageId="37" captionText="FIGURES 184–190. Baetis (Tenuibaetis) bialatus sp. n., larval exuviae (one and the same magnification). 184–185, abdomen; 186, head with labrum; 187, half of pronotum and mesonotum; 188–190, fore, middle and hind legs." figureDoi="http://doi.org/10.5281/zenodo.7889781" httpUri="https://zenodo.org/record/7889781/files/figure.png" pageId="36" pageNumber="237">Fig. 184</figureCitation>
), or laterally lighter than neighboring ones (
<figureCitation box="[913,1011,798,825]" captionStart="FIGURES 184–190" captionStartId="37.[151,264,1745,1769]" captionTargetBox="[151,1436,181,1721]" captionTargetId="figure-19@37.[151,1436,181,1721]" captionTargetPageId="37" captionText="FIGURES 184–190. Baetis (Tenuibaetis) bialatus sp. n., larval exuviae (one and the same magnification). 184–185, abdomen; 186, head with labrum; 187, half of pronotum and mesonotum; 188–190, fore, middle and hind legs." figureDoi="http://doi.org/10.5281/zenodo.7889781" httpUri="https://zenodo.org/record/7889781/files/figure.png" pageId="36" pageNumber="237">Fig. 185</figureCitation>
). Caudalii uniformly brown.
</paragraph>
<paragraph blockId="36.[151,1437,475,2013]" pageId="36" pageNumber="237">
HYPODERMAL COLORATION: Either not expressed, or abdominal terga with dark brown line on posterior margin as in imago (as in
<figureCitation box="[442,600,871,897]" captionStart="FIGURES 215–224" captionStartId="43.[151,264,1770,1794]" captionTargetBox="[151,1435,181,1746]" captionTargetId="figure-19@43.[151,1435,181,1746]" captionTargetPageId="43" captionText="FIGURES 215–224. Baetis (Tenuibaetis) bialatus sp. n. 215, subimaginal exuviae of mesopleuron; 216, subimaginal exuviae of half of mesonotum; 217–218, exuviae of fore and hind legs of male subimago; 219–220, fore and middle legs of male imago; 221, male imago; 222, female imago; 223, subimaginal exuviae of abdominal segments VIII–IX; 224, egg (215–220, holotype)." figureDoi="http://doi.org/10.5281/zenodo.7889793" httpUri="https://zenodo.org/record/7889793/files/figure.png" pageId="36" pageNumber="237">Figs 221–222</figureCitation>
).
</paragraph>
<paragraph blockId="36.[151,1437,475,2013]" pageId="36" pageNumber="237">
<collectionCode box="[200,511,907,933]" pageId="36" pageNumber="237">SHAPE AND SETATION</collectionCode>
: Labrum elongate, narrowing anteriorly (
<figureCitation box="[988,1087,907,933]" captionStart="FIGURES 191–194" captionStartId="38.[151,264,1802,1826]" captionTargetBox="[151,1430,181,1778]" captionTargetId="figure-36@38.[151,1436,181,1778]" captionTargetPageId="38" captionText="FIGURES 191–194. Baetis (Tenuibaetis) bialatus sp. n., larval mouth parts. 191, labrum; 192, hypopharynx with superlinguae; 193, maxilla; 194, labium." figureDoi="http://doi.org/10.5281/zenodo.7889783" httpUri="https://zenodo.org/record/7889783/files/figure.png" pageId="36" pageNumber="237">Fig. 191</figureCitation>
). Mandibles —
<figureCitation box="[1274,1433,906,933]" captionStart="FIGURES 195–201" captionStartId="39.[151,264,1840,1864]" captionTargetBox="[151,1436,190,1817]" captionTargetId="figure-19@39.[151,1436,181,1817]" captionTargetPageId="39" captionText="FIGURES 195–201. Baetis (Tenuibaetis) bialatus sp. n., larvae. 195–196, left and right mandibles with developing mandibles of next instar inside; 197–198, the same of another individual; 199, exuviae of abdominal tergum IX; 200, abdominal tergum II; 201, paraprocts. Abbreviations: hirud, hirudiform seta; sq, scale." figureDoi="http://doi.org/10.5281/zenodo.7889785" httpUri="https://zenodo.org/record/7889785/files/figure.png" pageId="36" pageNumber="237">Figs 195–198</figureCitation>
. Labial glossae, paraglossae and palps relatively long and narrow (
<figureCitation box="[889,987,943,969]" captionStart="FIGURES 191–194" captionStartId="38.[151,264,1802,1826]" captionTargetBox="[151,1430,181,1778]" captionTargetId="figure-36@38.[151,1436,181,1778]" captionTargetPageId="38" captionText="FIGURES 191–194. Baetis (Tenuibaetis) bialatus sp. n., larval mouth parts. 191, labrum; 192, hypopharynx with superlinguae; 193, maxilla; 194, labium." figureDoi="http://doi.org/10.5281/zenodo.7889783" httpUri="https://zenodo.org/record/7889783/files/figure.png" pageId="36" pageNumber="237">Fig. 194</figureCitation>
).
</paragraph>
<paragraph blockId="36.[151,1437,475,2013]" box="[199,699,979,1005]" pageId="36" pageNumber="237">
Hind protoptera completely absent (
<figureCitation box="[603,686,979,1005]" captionStart="FIGURES 13–21" captionStartId="5.[151,264,1840,1864]" captionTargetBox="[151,1436,181,1816]" captionTargetId="figure-19@5.[151,1436,181,1816]" captionTargetPageId="5" captionText="FIGURES 13–21. Imaginal hind wings and larval protoptera (under the same magnification). 13–14, female Baetis (Tenuibaetis) frequentus from India; 15–17, male B. (T.) octomaculatus sp. n.; 18–19, female of same species; 20–21, male B. (T.) bialatus sp. n. (13, 15, 20, metathorax of imago with hind wing; 14, 21, larval exuviae of metanotum; 16, 18, hind wing of imago; 17, 19, cuticle from metathorax of larva ready to molt to subimago, with subimaginal wing inside protopteron: crumpled on Fig. 17, non-crumpled on Fig. 19)." figureDoi="http://doi.org/10.5281/zenodo.7889705" httpUri="https://zenodo.org/record/7889705/files/figure.png" pageId="36" pageNumber="237">Fig. 21</figureCitation>
).
</paragraph>
<paragraph blockId="36.[151,1437,475,2013]" pageId="36" pageNumber="237">
Stout elongate setae forming row along outer side of femur (including two subapical setae), round in cross section and narrowed toward apex (
<figureCitation box="[567,729,1051,1077]" captionStart="FIGURES 208–210" captionStartId="41.[151,264,1840,1864]" captionTargetBox="[151,1435,181,1817]" captionTargetId="figure-19@41.[151,1435,181,1817]" captionTargetPageId="41" captionText="FIGURES 208–210. Baetis (Tenuibaetis) bialatus sp. n., proximal, middle and distal portions of larval femur (view from outer side).Abbreviations: 2-can, two-channel seta; ant-ap, anterior-apical margin of femur; hirud, hirudiform seta; post-ap, posteriorapical margin of femur." figureDoi="http://doi.org/10.5281/zenodo.7889789" httpUri="https://zenodo.org/record/7889789/files/figure.png" pageId="36" pageNumber="237">Figs 208–210</figureCitation>
), distally with single channel, proximally with two channels. Anterior-apical margin of femur with short, blunt two-channel setae (
<figureCitation box="[915,1013,1087,1113]" captionStart="FIGURES 208–210" captionStartId="41.[151,264,1840,1864]" captionTargetBox="[151,1435,181,1817]" captionTargetId="figure-19@41.[151,1435,181,1817]" captionTargetPageId="41" captionText="FIGURES 208–210. Baetis (Tenuibaetis) bialatus sp. n., proximal, middle and distal portions of larval femur (view from outer side).Abbreviations: 2-can, two-channel seta; ant-ap, anterior-apical margin of femur; hirud, hirudiform seta; post-ap, posteriorapical margin of femur." figureDoi="http://doi.org/10.5281/zenodo.7889789" httpUri="https://zenodo.org/record/7889789/files/figure.png" pageId="36" pageNumber="237">Fig. 210</figureCitation>
).Anterior side of femur with colorless oval scales in semicircular sockets (as on abdomen). Stout two-channel setae on anterior side of femur short (
<figureCitation captionStart="FIGURES 202–207" captionStartId="40.[151,264,1840,1864]" captionTargetBox="[151,1396,181,1811]" captionTargetId="figure-36@40.[151,1436,181,1816]" captionTargetPageId="40" captionText="FIGURES 202–207. Baetis (Tenuibaetis) bialatus sp. n., larval legs. 202, middle femur (anterior side); 203, fore tibia (anterior side); 204, middle tibia (anterior side); 205, middle tibia (outer side); 206–207, tibia and tarsus of fore and middle legs (anterior side) (202–204, 206–207, holotype). Abbreviations: 2-can, two-channel seta." figureDoi="http://doi.org/10.5281/zenodo.7889787" httpUri="https://zenodo.org/record/7889787/files/figure.png" pageId="36" pageNumber="237">Fig. 202</figureCitation>
).
</paragraph>
<paragraph blockId="36.[151,1437,475,2013]" pageId="36" pageNumber="237">
Stout, two-channel setae on outer side of fore tibia small (
<figureCitation box="[866,1027,1195,1221]" captionStart="FIGURES 202–207" captionStartId="40.[151,264,1840,1864]" captionTargetBox="[151,1396,181,1811]" captionTargetId="figure-36@40.[151,1436,181,1816]" captionTargetPageId="40" captionText="FIGURES 202–207. Baetis (Tenuibaetis) bialatus sp. n., larval legs. 202, middle femur (anterior side); 203, fore tibia (anterior side); 204, middle tibia (anterior side); 205, middle tibia (outer side); 206–207, tibia and tarsus of fore and middle legs (anterior side) (202–204, 206–207, holotype). Abbreviations: 2-can, two-channel seta." figureDoi="http://doi.org/10.5281/zenodo.7889787" httpUri="https://zenodo.org/record/7889787/files/figure.png" pageId="36" pageNumber="237">Figs 203, 206</figureCitation>
,
<figureCitation box="[1040,1142,1195,1221]" captionStart="FIGURES211–214" captionStartId="42.[151,264,1498,1522]" captionTargetBox="[151,1436,181,1469]" captionTargetId="figure-36@42.[151,1436,181,1474]" captionTargetPageId="42" captionText="FIGURES211–214. Baetis (Tenuibaetis) bialatus sp.n., larvaltibiae(view fromouter side).211–213,fore tibia;214,middle tibia." figureDoi="http://doi.org/10.5281/zenodo.7889791" httpUri="https://zenodo.org/record/7889791/files/figure.png" pageId="36" pageNumber="237">211–213</figureCitation>
); stout, two-channel setae on outer sides of middle and hind tibiae significantly larger (
<figureCitation box="[830,1044,1230,1257]" captionStart="FIGURES 202–207" captionStartId="40.[151,264,1840,1864]" captionTargetBox="[151,1396,181,1811]" captionTargetId="figure-36@40.[151,1436,181,1816]" captionTargetPageId="40" captionText="FIGURES 202–207. Baetis (Tenuibaetis) bialatus sp. n., larval legs. 202, middle femur (anterior side); 203, fore tibia (anterior side); 204, middle tibia (anterior side); 205, middle tibia (outer side); 206–207, tibia and tarsus of fore and middle legs (anterior side) (202–204, 206–207, holotype). Abbreviations: 2-can, two-channel seta." figureDoi="http://doi.org/10.5281/zenodo.7889787" httpUri="https://zenodo.org/record/7889787/files/figure.png" pageId="36" pageNumber="237">Figs 204–205, 207</figureCitation>
,
<figureCitation box="[1055,1102,1231,1257]" captionStart="FIGURES211–214" captionStartId="42.[151,264,1498,1522]" captionTargetBox="[151,1436,181,1469]" captionTargetId="figure-36@42.[151,1436,181,1474]" captionTargetPageId="42" captionText="FIGURES211–214. Baetis (Tenuibaetis) bialatus sp.n., larvaltibiae(view fromouter side).211–213,fore tibia;214,middle tibia." figureDoi="http://doi.org/10.5281/zenodo.7889791" httpUri="https://zenodo.org/record/7889791/files/figure.png" pageId="36" pageNumber="237">214</figureCitation>
). Each tibia with similar stout two-channel setae on inner side and along patella-tibial suture.
</paragraph>
<paragraph blockId="36.[151,1437,475,2013]" pageId="36" pageNumber="237">
Denticles on posterior margins of abdominal terga bluntly triangular (
<figureCitation box="[988,1087,1303,1329]" captionStart="FIGURES 195–201" captionStartId="39.[151,264,1840,1864]" captionTargetBox="[151,1436,190,1817]" captionTargetId="figure-19@39.[151,1436,181,1817]" captionTargetPageId="39" captionText="FIGURES 195–201. Baetis (Tenuibaetis) bialatus sp. n., larvae. 195–196, left and right mandibles with developing mandibles of next instar inside; 197–198, the same of another individual; 199, exuviae of abdominal tergum IX; 200, abdominal tergum II; 201, paraprocts. Abbreviations: hirud, hirudiform seta; sq, scale." figureDoi="http://doi.org/10.5281/zenodo.7889785" httpUri="https://zenodo.org/record/7889785/files/figure.png" pageId="36" pageNumber="237">Fig. 199</figureCitation>
); absent or sparse on tergum I. Denticles on posterior margins of abdominal sterna absent. Paraproct with small denticles (
<figureCitation box="[1175,1274,1339,1365]" captionStart="FIGURES 195–201" captionStartId="39.[151,264,1840,1864]" captionTargetBox="[151,1436,190,1817]" captionTargetId="figure-19@39.[151,1436,181,1817]" captionTargetPageId="39" captionText="FIGURES 195–201. Baetis (Tenuibaetis) bialatus sp. n., larvae. 195–196, left and right mandibles with developing mandibles of next instar inside; 197–198, the same of another individual; 199, exuviae of abdominal tergum IX; 200, abdominal tergum II; 201, paraprocts. Abbreviations: hirud, hirudiform seta; sq, scale." figureDoi="http://doi.org/10.5281/zenodo.7889785" httpUri="https://zenodo.org/record/7889785/files/figure.png" pageId="36" pageNumber="237">Fig. 201</figureCitation>
). Tergalii with short scales in semicircular sockets. Tergalius I much smaller than others (
<figureCitation box="[988,1148,1374,1401]" captionStart="FIGURES 184–190" captionStartId="37.[151,264,1745,1769]" captionTargetBox="[151,1436,181,1721]" captionTargetId="figure-19@37.[151,1436,181,1721]" captionTargetPageId="37" captionText="FIGURES 184–190. Baetis (Tenuibaetis) bialatus sp. n., larval exuviae (one and the same magnification). 184–185, abdomen; 186, head with labrum; 187, half of pronotum and mesonotum; 188–190, fore, middle and hind legs." figureDoi="http://doi.org/10.5281/zenodo.7889781" httpUri="https://zenodo.org/record/7889781/files/figure.png" pageId="36" pageNumber="237">Figs 184–185</figureCitation>
). Paracercus significantly smaller than cerci.
</paragraph>
<paragraph blockId="36.[151,1437,475,2013]" pageId="36" pageNumber="237">
<emphasis bold="true" box="[199,323,1446,1473]" pageId="36" pageNumber="237">
<emphasis bold="true" box="[199,316,1446,1473]" italics="true" pageId="36" pageNumber="237">Subimago</emphasis>
.
</emphasis>
CUTICULAR COLORATION: Head colorless, antennae brown. Pronotum with light brown and colorless areas. Mesonotum with ochre, light brownish and dark brown areas (
<figureCitation box="[1023,1121,1483,1509]" captionStart="FIGURES 215–224" captionStartId="43.[151,264,1770,1794]" captionTargetBox="[151,1435,181,1746]" captionTargetId="figure-19@43.[151,1435,181,1746]" captionTargetPageId="43" captionText="FIGURES 215–224. Baetis (Tenuibaetis) bialatus sp. n. 215, subimaginal exuviae of mesopleuron; 216, subimaginal exuviae of half of mesonotum; 217–218, exuviae of fore and hind legs of male subimago; 219–220, fore and middle legs of male imago; 221, male imago; 222, female imago; 223, subimaginal exuviae of abdominal segments VIII–IX; 224, egg (215–220, holotype)." figureDoi="http://doi.org/10.5281/zenodo.7889793" httpUri="https://zenodo.org/record/7889793/files/figure.png" pageId="36" pageNumber="237">Fig. 216</figureCitation>
). Thoracic pleura and sterna with light brownish and dark brown areas (
<figureCitation box="[626,724,1518,1545]" captionStart="FIGURES 215–224" captionStartId="43.[151,264,1770,1794]" captionTargetBox="[151,1435,181,1746]" captionTargetId="figure-19@43.[151,1435,181,1746]" captionTargetPageId="43" captionText="FIGURES 215–224. Baetis (Tenuibaetis) bialatus sp. n. 215, subimaginal exuviae of mesopleuron; 216, subimaginal exuviae of half of mesonotum; 217–218, exuviae of fore and hind legs of male subimago; 219–220, fore and middle legs of male imago; 221, male imago; 222, female imago; 223, subimaginal exuviae of abdominal segments VIII–IX; 224, egg (215–220, holotype)." figureDoi="http://doi.org/10.5281/zenodo.7889793" httpUri="https://zenodo.org/record/7889793/files/figure.png" pageId="36" pageNumber="237">Fig. 215</figureCitation>
). Legs brownish (
<figureCitation box="[921,1080,1519,1545]" captionStart="FIGURES 215–224" captionStartId="43.[151,264,1770,1794]" captionTargetBox="[151,1435,181,1746]" captionTargetId="figure-19@43.[151,1435,181,1746]" captionTargetPageId="43" captionText="FIGURES 215–224. Baetis (Tenuibaetis) bialatus sp. n. 215, subimaginal exuviae of mesopleuron; 216, subimaginal exuviae of half of mesonotum; 217–218, exuviae of fore and hind legs of male subimago; 219–220, fore and middle legs of male imago; 221, male imago; 222, female imago; 223, subimaginal exuviae of abdominal segments VIII–IX; 224, egg (215–220, holotype)." figureDoi="http://doi.org/10.5281/zenodo.7889793" httpUri="https://zenodo.org/record/7889793/files/figure.png" pageId="36" pageNumber="237">Figs 217–218</figureCitation>
). All abdominal terga and sterna I–X light brownish with lighter sigilla (
<figureCitation box="[592,690,1555,1581]" captionStart="FIGURES 215–224" captionStartId="43.[151,264,1770,1794]" captionTargetBox="[151,1435,181,1746]" captionTargetId="figure-19@43.[151,1435,181,1746]" captionTargetPageId="43" captionText="FIGURES 215–224. Baetis (Tenuibaetis) bialatus sp. n. 215, subimaginal exuviae of mesopleuron; 216, subimaginal exuviae of half of mesonotum; 217–218, exuviae of fore and hind legs of male subimago; 219–220, fore and middle legs of male imago; 221, male imago; 222, female imago; 223, subimaginal exuviae of abdominal segments VIII–IX; 224, egg (215–220, holotype)." figureDoi="http://doi.org/10.5281/zenodo.7889793" httpUri="https://zenodo.org/record/7889793/files/figure.png" pageId="36" pageNumber="237">Fig. 223</figureCitation>
).
</paragraph>
<paragraph blockId="36.[151,1437,475,2013]" pageId="36" pageNumber="237">
HYPODERMAL COLORATION: Both in male and female, head, thorax, legs and abdomen ochre; abdominal terga I–
<collectionCode box="[237,267,1627,1653]" pageId="36" pageNumber="237">IX</collectionCode>
with contrasting reddish-brown band bordering posterior margin and not reaching lateral margins (as in female imago—
<figureCitation box="[335,431,1663,1689]" captionStart="FIGURES 215–224" captionStartId="43.[151,264,1770,1794]" captionTargetBox="[151,1435,181,1746]" captionTargetId="figure-19@43.[151,1435,181,1746]" captionTargetPageId="43" captionText="FIGURES 215–224. Baetis (Tenuibaetis) bialatus sp. n. 215, subimaginal exuviae of mesopleuron; 216, subimaginal exuviae of half of mesonotum; 217–218, exuviae of fore and hind legs of male subimago; 219–220, fore and middle legs of male imago; 221, male imago; 222, female imago; 223, subimaginal exuviae of abdominal segments VIII–IX; 224, egg (215–220, holotype)." figureDoi="http://doi.org/10.5281/zenodo.7889793" httpUri="https://zenodo.org/record/7889793/files/figure.png" pageId="36" pageNumber="237">Fig. 222</figureCitation>
).
</paragraph>
<paragraph blockId="36.[151,1437,475,2013]" pageId="36" pageNumber="237">
<emphasis bold="true" box="[199,345,1698,1725]" italics="true" pageId="36" pageNumber="237">Imago, male</emphasis>
(
<figureCitation box="[362,461,1699,1725]" captionStart="FIGURES 215–224" captionStartId="43.[151,264,1770,1794]" captionTargetBox="[151,1435,181,1746]" captionTargetId="figure-19@43.[151,1435,181,1746]" captionTargetPageId="43" captionText="FIGURES 215–224. Baetis (Tenuibaetis) bialatus sp. n. 215, subimaginal exuviae of mesopleuron; 216, subimaginal exuviae of half of mesonotum; 217–218, exuviae of fore and hind legs of male subimago; 219–220, fore and middle legs of male imago; 221, male imago; 222, female imago; 223, subimaginal exuviae of abdominal segments VIII–IX; 224, egg (215–220, holotype)." figureDoi="http://doi.org/10.5281/zenodo.7889793" httpUri="https://zenodo.org/record/7889793/files/figure.png" pageId="36" pageNumber="237">Fig. 221</figureCitation>
). Head brown. Antennae brown. Turbinate eyes ochre. Thorax brown. Fore wing with membrane colorless or slightly tinged with brownish; veins from ochre (nearer to wing base) to brown; pterostigma with oblique cross veins and disconnected vein fragments. Hind wings entirely absent (
<figureCitation box="[1153,1239,1771,1797]" captionStart="FIGURES 13–21" captionStartId="5.[151,264,1840,1864]" captionTargetBox="[151,1436,181,1816]" captionTargetId="figure-19@5.[151,1436,181,1816]" captionTargetPageId="5" captionText="FIGURES 13–21. Imaginal hind wings and larval protoptera (under the same magnification). 13–14, female Baetis (Tenuibaetis) frequentus from India; 15–17, male B. (T.) octomaculatus sp. n.; 18–19, female of same species; 20–21, male B. (T.) bialatus sp. n. (13, 15, 20, metathorax of imago with hind wing; 14, 21, larval exuviae of metanotum; 16, 18, hind wing of imago; 17, 19, cuticle from metathorax of larva ready to molt to subimago, with subimaginal wing inside protopteron: crumpled on Fig. 17, non-crumpled on Fig. 19)." figureDoi="http://doi.org/10.5281/zenodo.7889705" httpUri="https://zenodo.org/record/7889705/files/figure.png" pageId="36" pageNumber="237">Fig. 20</figureCitation>
). Legs uniformly ochre-brownish (
<figureCitation box="[348,513,1807,1833]" captionStart="FIGURES 215–224" captionStartId="43.[151,264,1770,1794]" captionTargetBox="[151,1435,181,1746]" captionTargetId="figure-19@43.[151,1435,181,1746]" captionTargetPageId="43" captionText="FIGURES 215–224. Baetis (Tenuibaetis) bialatus sp. n. 215, subimaginal exuviae of mesopleuron; 216, subimaginal exuviae of half of mesonotum; 217–218, exuviae of fore and hind legs of male subimago; 219–220, fore and middle legs of male imago; 221, male imago; 222, female imago; 223, subimaginal exuviae of abdominal segments VIII–IX; 224, egg (215–220, holotype)." figureDoi="http://doi.org/10.5281/zenodo.7889793" httpUri="https://zenodo.org/record/7889793/files/figure.png" pageId="36" pageNumber="237">Figs 219–220</figureCitation>
). Middle and hind tarsi with 2 apical spines (on 1st+2nd and 3rd segments). Abdominal terga and sterna either uniformly light brown, or terga diffusively darkened with reddish laterally; terga I–
<collectionCode box="[174,204,1879,1905]" pageId="36" pageNumber="237">IX</collectionCode>
with darker reddish-brown band bordering posterior margin and not reaching lateral margins. Cerci brown. Gonostyli brown; 3rd (terminal) segment short and petiolate (
<figureCitation box="[836,994,1914,1941]" captionStart="FIGURES 225–229" captionStartId="44.[151,264,1840,1864]" captionTargetBox="[151,1436,185,1817]" captionTargetId="figure-36@44.[151,1436,181,1817]" captionTargetPageId="44" captionText="FIGURES 225–229. Baetis (Tenuibaetis) bialatus sp. n., genitalia of male imago. 225, 227, ventral view; 226, 229, ventral view with focus on sterno-styligeral muscle; 228, enlarged fragment of sterno-styligeral muscle to show cross-striation (225–226, holotype). Abbreviations: gv, gonovectis; s.s, sterno-styligeral muscle." figureDoi="http://doi.org/10.5281/zenodo.7889797" httpUri="https://zenodo.org/record/7889797/files/figure.png" pageId="36" pageNumber="237">Figs 225, 227</figureCitation>
). Gonovectes sharply bent, dark brown. Sterno-styligeral muscle vestigial, either without cross-striation (
<figureCitation box="[869,966,1951,1977]" captionStart="FIGURES 225–229" captionStartId="44.[151,264,1840,1864]" captionTargetBox="[151,1436,185,1817]" captionTargetId="figure-36@44.[151,1436,181,1817]" captionTargetPageId="44" captionText="FIGURES 225–229. Baetis (Tenuibaetis) bialatus sp. n., genitalia of male imago. 225, 227, ventral view; 226, 229, ventral view with focus on sterno-styligeral muscle; 228, enlarged fragment of sterno-styligeral muscle to show cross-striation (225–226, holotype). Abbreviations: gv, gonovectis; s.s, sterno-styligeral muscle." figureDoi="http://doi.org/10.5281/zenodo.7889797" httpUri="https://zenodo.org/record/7889797/files/figure.png" pageId="36" pageNumber="237">Fig. 226</figureCitation>
), or with cross-striation of few fibrils (
<figureCitation captionStart="FIGURES 225–229" captionStartId="44.[151,264,1840,1864]" captionTargetBox="[151,1436,185,1817]" captionTargetId="figure-36@44.[151,1436,181,1817]" captionTargetPageId="44" captionText="FIGURES 225–229. Baetis (Tenuibaetis) bialatus sp. n., genitalia of male imago. 225, 227, ventral view; 226, 229, ventral view with focus on sterno-styligeral muscle; 228, enlarged fragment of sterno-styligeral muscle to show cross-striation (225–226, holotype). Abbreviations: gv, gonovectis; s.s, sterno-styligeral muscle." figureDoi="http://doi.org/10.5281/zenodo.7889797" httpUri="https://zenodo.org/record/7889797/files/figure.png" pageId="36" pageNumber="237">Fig. 228</figureCitation>
).
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7889781" ID-Zenodo-Dep="7889781" httpUri="https://zenodo.org/record/7889781/files/figure.png" pageId="37" pageNumber="238" startId="37.[151,264,1745,1769]" targetBox="[151,1436,181,1721]" targetPageId="37" targetType="figure">
<paragraph blockId="37.[151,1437,1745,1805]" pageId="37" pageNumber="238">
<emphasis bold="true" box="[151,365,1745,1769]" pageId="37" pageNumber="238">FIGURES 184–190.</emphasis>
<taxonomicName authority="Kluge &amp; Srinivasan &amp; Sivaruban &amp; Barathy &amp; Isack, 2023" authorityName="Kluge &amp; Srinivasan &amp; Sivaruban &amp; Barathy &amp; Isack" authorityYear="2023" box="[371,661,1745,1769]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="37" pageNumber="238" phylum="Arthropoda" rank="species" species="bialatus" status="sp. n." subGenus="Tenuibaetis">
<emphasis box="[371,434,1746,1769]" italics="true" pageId="37" pageNumber="238">Baetis</emphasis>
(
<emphasis box="[449,565,1745,1769]" italics="true" pageId="37" pageNumber="238">Tenuibaetis</emphasis>
)
<emphasis box="[579,661,1745,1769]" italics="true" pageId="37" pageNumber="238">bialatus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[667,723,1745,1769]" pageId="37" pageNumber="238" rank="species">
<emphasis bold="true" box="[667,717,1745,1769]" pageId="37" pageNumber="238">sp. n</emphasis>
.
</taxonomicNameLabel>
, larval exuviae (one and the same magnification). 184–185, abdomen; 186, head with labrum; 187, half of pronotum and mesonotum; 188–190, fore, middle and hind legs.
</paragraph>
</caption>
<paragraph blockId="37.[151,1437,1851,2022]" pageId="37" pageNumber="238">
<emphasis bold="true" box="[199,369,1851,1878]" italics="true" pageId="37" pageNumber="238">Imago, female</emphasis>
(
<figureCitation box="[387,490,1851,1877]" captionStart="FIGURES 215–224" captionStartId="43.[151,264,1770,1794]" captionTargetBox="[151,1435,181,1746]" captionTargetId="figure-19@43.[151,1435,181,1746]" captionTargetPageId="43" captionText="FIGURES 215–224. Baetis (Tenuibaetis) bialatus sp. n. 215, subimaginal exuviae of mesopleuron; 216, subimaginal exuviae of half of mesonotum; 217–218, exuviae of fore and hind legs of male subimago; 219–220, fore and middle legs of male imago; 221, male imago; 222, female imago; 223, subimaginal exuviae of abdominal segments VIII–IX; 224, egg (215–220, holotype)." figureDoi="http://doi.org/10.5281/zenodo.7889793" httpUri="https://zenodo.org/record/7889793/files/figure.png" pageId="37" pageNumber="238">Fig. 222</figureCitation>
). Head, thorax, legs, abdomen and cerci ochre; abdominal terga I–
<collectionCode box="[1265,1295,1852,1878]" pageId="37" pageNumber="238">IX</collectionCode>
with darker reddish-brown band bordering posterior margin and not reaching lateral margins. All tarsi with 2 apical spines (fore tarsus on 2nd and 3rd segments, middle and hind tarsi on 1st+2nd and 3rd segments).
</paragraph>
<paragraph blockId="37.[151,1437,1851,2022]" box="[199,604,1959,1986]" pageId="37" pageNumber="238">
<emphasis bold="true" box="[199,334,1959,1985]" pageId="37" pageNumber="238">Dimension.</emphasis>
Forewing length
<quantity box="[532,600,1959,1986]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="37" pageNumber="238" unit="mm" value="6.0">6 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="42" lastPageNumber="243" pageId="37" pageNumber="238" type="distribution">
<paragraph blockId="37.[151,1437,1851,2022]" box="[199,1083,1995,2022]" pageId="37" pageNumber="238">
<emphasis bold="true" box="[199,353,1995,2021]" pageId="37" pageNumber="238">Distribution.</emphasis>
Mountains of the Western Ghats in southern
<collectingCountry box="[860,918,1995,2021]" name="India" pageId="37" pageNumber="238">India</collectingCountry>
(
<collectingRegion box="[934,1072,1995,2021]" country="India" name="Tamil Nadu" pageId="37" pageNumber="238">Tamil Nadu</collectingRegion>
).
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7889783" ID-Zenodo-Dep="7889783" httpUri="https://zenodo.org/record/7889783/files/figure.png" pageId="38" pageNumber="239" startId="38.[151,264,1802,1826]" targetBox="[151,1430,181,1778]" targetPageId="38" targetType="figure">
<paragraph blockId="38.[151,1437,1802,1863]" pageId="38" pageNumber="239">
<emphasis bold="true" box="[151,365,1802,1826]" pageId="38" pageNumber="239">FIGURES 191–194.</emphasis>
<taxonomicName authority="Kluge &amp; Srinivasan &amp; Sivaruban &amp; Barathy &amp; Isack, 2023" authorityName="Kluge &amp; Srinivasan &amp; Sivaruban &amp; Barathy &amp; Isack" authorityYear="2023" box="[370,659,1802,1827]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="38" pageNumber="239" phylum="Arthropoda" rank="species" species="bialatus" status="sp. n." subGenus="Tenuibaetis">
<emphasis box="[370,433,1803,1826]" italics="true" pageId="38" pageNumber="239">Baetis</emphasis>
(
<emphasis box="[447,563,1802,1826]" italics="true" pageId="38" pageNumber="239">Tenuibaetis</emphasis>
)
<emphasis box="[577,659,1802,1826]" italics="true" pageId="38" pageNumber="239">bialatus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[664,720,1803,1827]" pageId="38" pageNumber="239" rank="species">
<emphasis bold="true" box="[664,714,1803,1827]" pageId="38" pageNumber="239">sp. n</emphasis>
.
</taxonomicNameLabel>
, larval mouth parts. 191, labrum; 192, hypopharynx with superlinguae; 193, maxilla; 194, labium.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7889785" ID-Zenodo-Dep="7889785" httpUri="https://zenodo.org/record/7889785/files/figure.png" pageId="39" pageNumber="240" startId="39.[151,264,1840,1864]" targetBox="[151,1436,190,1817]" targetPageId="39" targetType="figure">
<paragraph blockId="39.[151,1436,1840,1937]" pageId="39" pageNumber="240">
<emphasis bold="true" box="[151,366,1840,1865]" pageId="39" pageNumber="240">FIGURES 195–201.</emphasis>
<taxonomicName authority="Kluge &amp; Srinivasan &amp; Sivaruban &amp; Barathy &amp; Isack, 2023" authorityName="Kluge &amp; Srinivasan &amp; Sivaruban &amp; Barathy &amp; Isack" authorityYear="2023" box="[372,662,1841,1865]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="39" pageNumber="240" phylum="Arthropoda" rank="species" species="bialatus" status="sp. n." subGenus="Tenuibaetis">
<emphasis box="[372,435,1841,1864]" italics="true" pageId="39" pageNumber="240">Baetis</emphasis>
(
<emphasis box="[450,566,1841,1865]" italics="true" pageId="39" pageNumber="240">Tenuibaetis</emphasis>
)
<emphasis box="[580,662,1841,1865]" italics="true" pageId="39" pageNumber="240">bialatus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[668,725,1841,1865]" pageId="39" pageNumber="240" rank="species">
<emphasis bold="true" box="[668,719,1841,1865]" pageId="39" pageNumber="240">sp. n</emphasis>
.
</taxonomicNameLabel>
, larvae. 195–196, left and right mandibles with developing mandibles of next instar inside; 197–198, the same of another individual; 199, exuviae of abdominal tergum IX; 200, abdominal tergum II; 201, paraprocts. Abbreviations: hirud, hirudiform seta; sq, scale.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7889787" ID-Zenodo-Dep="7889787" httpUri="https://zenodo.org/record/7889787/files/figure.png" pageId="40" pageNumber="241" startId="40.[151,264,1840,1864]" targetBox="[151,1396,181,1811]" targetPageId="40" targetType="figure">
<paragraph blockId="40.[151,1437,1840,1937]" pageId="40" pageNumber="241">
<emphasis bold="true" box="[151,364,1840,1864]" pageId="40" pageNumber="241">FIGURES 202–207.</emphasis>
<taxonomicName authority="Kluge &amp; Srinivasan &amp; Sivaruban &amp; Barathy &amp; Isack, 2023" authorityName="Kluge &amp; Srinivasan &amp; Sivaruban &amp; Barathy &amp; Isack" authorityYear="2023" box="[371,661,1840,1865]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="40" pageNumber="241" phylum="Arthropoda" rank="species" species="bialatus" status="sp. n." subGenus="Tenuibaetis">
<emphasis box="[371,434,1841,1864]" italics="true" pageId="40" pageNumber="241">Baetis</emphasis>
(
<emphasis box="[449,565,1840,1864]" italics="true" pageId="40" pageNumber="241">Tenuibaetis</emphasis>
)
<emphasis box="[579,661,1840,1864]" italics="true" pageId="40" pageNumber="241">bialatus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[666,723,1841,1865]" pageId="40" pageNumber="241" rank="species">
<emphasis bold="true" box="[666,717,1841,1865]" pageId="40" pageNumber="241">sp. n</emphasis>
.
</taxonomicNameLabel>
, larval legs. 202, middle femur (anterior side); 203, fore tibia (anterior side); 204, middle tibia (anterior side); 205, middle tibia (outer side); 206–207, tibia and tarsus of fore and middle legs (anterior side) (202–204, 206–207, holotype). Abbreviations: 2-can, two-channel seta.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7889789" ID-Zenodo-Dep="7889789" httpUri="https://zenodo.org/record/7889789/files/figure.png" pageId="41" pageNumber="242" startId="41.[151,264,1840,1864]" targetBox="[151,1435,181,1817]" targetPageId="41" targetType="figure">
<paragraph blockId="41.[151,1436,1840,1936]" pageId="41" pageNumber="242">
<emphasis bold="true" box="[151,364,1840,1864]" pageId="41" pageNumber="242">FIGURES 208–210.</emphasis>
<taxonomicName authority="Kluge &amp; Srinivasan &amp; Sivaruban &amp; Barathy &amp; Isack, 2023" authorityName="Kluge &amp; Srinivasan &amp; Sivaruban &amp; Barathy &amp; Isack" authorityYear="2023" box="[371,661,1840,1865]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="41" pageNumber="242" phylum="Arthropoda" rank="species" species="bialatus" status="sp. n." subGenus="Tenuibaetis">
<emphasis box="[371,434,1841,1864]" italics="true" pageId="41" pageNumber="242">Baetis</emphasis>
(
<emphasis box="[449,565,1840,1864]" italics="true" pageId="41" pageNumber="242">Tenuibaetis</emphasis>
)
<emphasis box="[579,661,1840,1864]" italics="true" pageId="41" pageNumber="242">bialatus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[666,723,1841,1865]" pageId="41" pageNumber="242" rank="species">
<emphasis bold="true" box="[666,717,1841,1865]" pageId="41" pageNumber="242">sp. n</emphasis>
.
</taxonomicNameLabel>
, proximal, middle and distal portions of larval femur (view from outer side). Abbreviations: 2-can, two-channel seta; ant-ap, anterior-apical margin of femur; hirud, hirudiform seta; post-ap, posteriorapical margin of femur.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7889791" ID-Zenodo-Dep="7889791" box="[151,1437,1498,1522]" httpUri="https://zenodo.org/record/7889791/files/figure.png" pageId="42" pageNumber="243" startId="42.[151,264,1498,1522]" targetBox="[151,1436,181,1469]" targetPageId="42" targetType="figure">
<paragraph blockId="42.[151,1437,1498,1522]" box="[151,1437,1498,1522]" pageId="42" pageNumber="243">
<emphasis bold="true" box="[151,360,1498,1522]" pageId="42" pageNumber="243">FIGURES 211–214.</emphasis>
<taxonomicName authority="Kluge &amp; Srinivasan &amp; Sivaruban &amp; Barathy &amp; Isack, 2023" authorityName="Kluge &amp; Srinivasan &amp; Sivaruban &amp; Barathy &amp; Isack" authorityYear="2023" box="[364,649,1498,1522]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="42" pageNumber="243" phylum="Arthropoda" rank="species" species="bialatus" status="sp. n." subGenus="Tenuibaetis">
<emphasis box="[364,427,1499,1522]" italics="true" pageId="42" pageNumber="243">Baetis</emphasis>
(
<emphasis box="[440,556,1498,1522]" italics="true" pageId="42" pageNumber="243">Tenuibaetis</emphasis>
)
<emphasis box="[567,649,1498,1522]" italics="true" pageId="42" pageNumber="243">bialatus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[653,706,1498,1522]" pageId="42" pageNumber="243" rank="species">
<emphasis bold="true" box="[653,700,1498,1522]" pageId="42" pageNumber="243">sp. n</emphasis>
.
</taxonomicNameLabel>
, larval tibiae (view from outer side). 211–213, fore tibia; 214, middle tibia.
</paragraph>
</caption>
<paragraph blockId="42.[151,1437,1568,1990]" pageId="42" pageNumber="243">
<emphasis bold="true" box="[199,354,1568,1594]" pageId="42" pageNumber="243">Comparison.</emphasis>
<taxonomicName authority="Kluge &amp; Srinivasan &amp; Sivaruban &amp; Barathy &amp; Isack, 2023" authorityName="Kluge &amp; Srinivasan &amp; Sivaruban &amp; Barathy &amp; Isack" authorityYear="2023" box="[362,686,1568,1594]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="42" pageNumber="243" phylum="Arthropoda" rank="species" species="bialatus" status="sp. n." subGenus="Tenuibaetis">
<emphasis box="[362,432,1569,1594]" italics="true" pageId="42" pageNumber="243">Baetis</emphasis>
(
<emphasis box="[450,578,1568,1594]" italics="true" pageId="42" pageNumber="243">Tenuibaetis</emphasis>
)
<emphasis box="[596,686,1568,1594]" italics="true" pageId="42" pageNumber="243">bialatus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[694,759,1568,1595]" pageId="42" pageNumber="243" rank="species">
<emphasis bold="true" box="[694,752,1568,1594]" pageId="42" pageNumber="243">sp. n</emphasis>
.
</taxonomicNameLabel>
differs from most other species of
<taxonomicName authorityName=": Fujitani, Hirowatari &amp; Tanida" authorityYear="2003" box="[1161,1289,1568,1594]" class="Insecta" family="Baetidae" genus="Tenuibaetis" kingdom="Animalia" order="Ephemeroptera" pageId="42" pageNumber="243" phylum="Arthropoda" rank="genus">
<emphasis box="[1161,1289,1568,1594]" italics="true" pageId="42" pageNumber="243">Tenuibaetis</emphasis>
</taxonomicName>
by complete absence of hind wings in winged stages and by complete absence of hind protoptera in larvae of all instars (
<figureCitation captionStart="FIGURES 13–21" captionStartId="5.[151,264,1840,1864]" captionTargetBox="[151,1436,181,1816]" captionTargetId="figure-19@5.[151,1436,181,1816]" captionTargetPageId="5" captionText="FIGURES 13–21. Imaginal hind wings and larval protoptera (under the same magnification). 13–14, female Baetis (Tenuibaetis) frequentus from India; 15–17, male B. (T.) octomaculatus sp. n.; 18–19, female of same species; 20–21, male B. (T.) bialatus sp. n. (13, 15, 20, metathorax of imago with hind wing; 14, 21, larval exuviae of metanotum; 16, 18, hind wing of imago; 17, 19, cuticle from metathorax of larva ready to molt to subimago, with subimaginal wing inside protopteron: crumpled on Fig. 17, non-crumpled on Fig. 19)." figureDoi="http://doi.org/10.5281/zenodo.7889705" httpUri="https://zenodo.org/record/7889705/files/figure.png" pageId="42" pageNumber="243">Figs 20–21</figureCitation>
). Larva of
<taxonomicName authority="Kluge &amp; Srinivasan &amp; Sivaruban &amp; Barathy &amp; Isack, 2023" authorityName="Kluge &amp; Srinivasan &amp; Sivaruban &amp; Barathy &amp; Isack" authorityYear="2023" box="[355,530,1640,1667]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="42" pageNumber="246" phylum="Arthropoda" rank="species" species="bialatus" status="sp. n." subGenus="Tenuibaetis">
<emphasis box="[355,372,1641,1666]" italics="true" pageId="42" pageNumber="243">B</emphasis>
. (
<emphasis box="[399,415,1641,1666]" italics="true" pageId="42" pageNumber="243">T</emphasis>
.)
<emphasis box="[440,530,1640,1666]" italics="true" pageId="42" pageNumber="243">bialatus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[540,606,1640,1667]" pageId="42" pageNumber="243" rank="species">
<emphasis bold="true" box="[540,600,1640,1666]" pageId="42" pageNumber="243">sp. n</emphasis>
.
</taxonomicNameLabel>
differs from another two-winged species,
<taxonomicName authorityName="Kluge &amp; Srinivasan &amp; Sivaruban &amp; Barathy &amp; Isack" authorityYear="2023" baseAuthorityName="Kluge &amp; Srinivasan &amp; Sivaruban &amp; Barathy &amp; Isack" baseAuthorityYear="2022" box="[1097,1260,1640,1667]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="42" pageNumber="208" phylum="Arthropoda" rank="species" species="panhai" subGenus="Tenuibaetis">
<emphasis box="[1097,1114,1641,1666]" italics="true" pageId="42" pageNumber="243">B</emphasis>
. (
<emphasis box="[1141,1157,1641,1666]" italics="true" pageId="42" pageNumber="243">T</emphasis>
.)
<emphasis box="[1182,1260,1640,1666]" italics="true" pageId="42" pageNumber="243">panhai</emphasis>
</taxonomicName>
, by longer and narrower labrum, labial glossae, paraglossae and palps (
<figureCitation box="[769,924,1676,1702]" captionStart="FIGURES 191–194" captionStartId="38.[151,264,1802,1826]" captionTargetBox="[151,1430,181,1778]" captionTargetId="figure-36@38.[151,1436,181,1778]" captionTargetPageId="38" captionText="FIGURES 191–194. Baetis (Tenuibaetis) bialatus sp. n., larval mouth parts. 191, labrum; 192, hypopharynx with superlinguae; 193, maxilla; 194, labium." figureDoi="http://doi.org/10.5281/zenodo.7889783" httpUri="https://zenodo.org/record/7889783/files/figure.png" pageId="42" pageNumber="243">Figs 191, 194</figureCitation>
), by more uniform brown coloration of cuticle on mesonotum and abdominal terga, by pointed and non-flattened setae of outer row on each femur (
<figureCitation box="[1273,1429,1712,1738]" captionStart="FIGURES 208–210" captionStartId="41.[151,264,1840,1864]" captionTargetBox="[151,1435,181,1817]" captionTargetId="figure-19@41.[151,1435,181,1817]" captionTargetPageId="41" captionText="FIGURES 208–210. Baetis (Tenuibaetis) bialatus sp. n., proximal, middle and distal portions of larval femur (view from outer side).Abbreviations: 2-can, two-channel seta; ant-ap, anterior-apical margin of femur; hirud, hirudiform seta; post-ap, posteriorapical margin of femur." figureDoi="http://doi.org/10.5281/zenodo.7889789" httpUri="https://zenodo.org/record/7889789/files/figure.png" pageId="42" pageNumber="243">Figs 208–210</figureCitation>
) and by smaller stout setae on anterior surface of each femur.
</paragraph>
<paragraph blockId="42.[151,1437,1568,1990]" pageId="42" pageNumber="243">
From the sympatric species
<taxonomicName authorityName="Muller-Liebenau &amp; Hubbard" authorityYear="1985" box="[522,720,1784,1810]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="42" pageNumber="243" phylum="Arthropoda" rank="species" species="frequentus" subGenus="Tenuibaetis">
<emphasis box="[522,546,1785,1810]" italics="true" pageId="42" pageNumber="243">B.</emphasis>
(
<emphasis box="[564,586,1785,1810]" italics="true" pageId="42" pageNumber="243">T.</emphasis>
)
<emphasis box="[603,720,1784,1810]" italics="true" pageId="42" pageNumber="243">frequentus</emphasis>
</taxonomicName>
, larva of
<taxonomicName authority="Kluge &amp; Srinivasan &amp; Sivaruban &amp; Barathy &amp; Isack, 2023" authorityName="Kluge &amp; Srinivasan &amp; Sivaruban &amp; Barathy &amp; Isack" authorityYear="2023" box="[834,1008,1784,1811]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="42" pageNumber="246" phylum="Arthropoda" rank="species" species="bialatus" status="sp. n." subGenus="Tenuibaetis">
<emphasis box="[834,851,1785,1810]" italics="true" pageId="42" pageNumber="243">B</emphasis>
. (
<emphasis box="[877,893,1785,1810]" italics="true" pageId="42" pageNumber="243">T</emphasis>
.)
<emphasis box="[918,1008,1784,1810]" italics="true" pageId="42" pageNumber="243">bialatus</emphasis>
</taxonomicName>
<emphasis bold="true" box="[1017,1083,1784,1810]" pageId="42" pageNumber="243">
<taxonomicNameLabel box="[1017,1083,1784,1810]" pageId="42" pageNumber="243" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
differs, besides the absence of hind wings, by two-channel setae on tibiae (rather than pseudo-bifurcate ones) and by pointed and non-flattened stout setae on outers sides of femora; its proximal blank on fore femur is oval (
<figureCitation box="[1049,1147,1856,1882]" captionStart="FIGURES 184–190" captionStartId="37.[151,264,1745,1769]" captionTargetBox="[151,1436,181,1721]" captionTargetId="figure-19@37.[151,1436,181,1721]" captionTargetPageId="37" captionText="FIGURES 184–190. Baetis (Tenuibaetis) bialatus sp. n., larval exuviae (one and the same magnification). 184–185, abdomen; 186, head with labrum; 187, half of pronotum and mesonotum; 188–190, fore, middle and hind legs." figureDoi="http://doi.org/10.5281/zenodo.7889781" httpUri="https://zenodo.org/record/7889781/files/figure.png" pageId="42" pageNumber="243">Fig. 188</figureCitation>
), rather than triangular in
<taxonomicName authorityName="Muller-Liebenau &amp; Hubbard" authorityYear="1985" box="[151,347,1892,1919]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="42" pageNumber="246" phylum="Arthropoda" rank="species" species="frequentus" subGenus="Tenuibaetis">
<emphasis box="[151,168,1893,1918]" italics="true" pageId="42" pageNumber="243">B</emphasis>
. (
<emphasis box="[191,207,1893,1918]" italics="true" pageId="42" pageNumber="243">T</emphasis>
.)
<emphasis box="[230,347,1892,1918]" italics="true" pageId="42" pageNumber="243">frequentus</emphasis>
</taxonomicName>
(
<figureCitation box="[363,447,1892,1919]" captionStart="FIGURES 69–77" captionStartId="18.[151,264,1844,1868]" captionTargetBox="[151,1425,181,1820]" captionTargetId="figure-36@18.[151,1436,181,1820]" captionTargetPageId="18" captionText="FIGURES 69–77. Baetis (Tenuibaetis) frequentus, larval exuviae. 69–75, from India (69–74, at one magnification): 69, head; 70–72, fore, middle and hind legs; 73, abdomen; 74, half of pronotum and mesonotum, 75, femur of dark individual. 76–77, posterior margin of abdominal tergum VI of two individuals from Haputale (Sri Lanka)." figureDoi="http://doi.org/10.5281/zenodo.7889739" httpUri="https://zenodo.org/record/7889739/files/figure.png" pageId="42" pageNumber="243">Fig. 75</figureCitation>
). In contrast to
<taxonomicName authorityName="Muller-Liebenau &amp; Hubbard" authorityYear="1985" box="[621,817,1892,1919]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="42" pageNumber="246" phylum="Arthropoda" rank="species" species="frequentus" subGenus="Tenuibaetis">
<emphasis box="[621,638,1893,1918]" italics="true" pageId="42" pageNumber="243">B</emphasis>
. (
<emphasis box="[661,677,1893,1918]" italics="true" pageId="42" pageNumber="243">T</emphasis>
.)
<emphasis box="[700,817,1892,1918]" italics="true" pageId="42" pageNumber="243">frequentus</emphasis>
</taxonomicName>
, stout setae on fore tibiae are significantly smaller than such setae on middle and hind tibiae (
<figureCitation box="[571,731,1928,1954]" captionStart="FIGURES 202–207" captionStartId="40.[151,264,1840,1864]" captionTargetBox="[151,1396,181,1811]" captionTargetId="figure-36@40.[151,1436,181,1816]" captionTargetPageId="40" captionText="FIGURES 202–207. Baetis (Tenuibaetis) bialatus sp. n., larval legs. 202, middle femur (anterior side); 203, fore tibia (anterior side); 204, middle tibia (anterior side); 205, middle tibia (outer side); 206–207, tibia and tarsus of fore and middle legs (anterior side) (202–204, 206–207, holotype). Abbreviations: 2-can, two-channel seta." figureDoi="http://doi.org/10.5281/zenodo.7889787" httpUri="https://zenodo.org/record/7889787/files/figure.png" pageId="42" pageNumber="243">Figs 206–207</figureCitation>
). Male imago of
<taxonomicName authority="Kluge &amp; Srinivasan &amp; Sivaruban &amp; Barathy &amp; Isack, 2023" authorityName="Kluge &amp; Srinivasan &amp; Sivaruban &amp; Barathy &amp; Isack" authorityYear="2023" box="[920,1089,1928,1955]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="42" pageNumber="246" phylum="Arthropoda" rank="species" species="bialatus" status="sp. n." subGenus="Tenuibaetis">
<emphasis box="[920,937,1929,1954]" italics="true" pageId="42" pageNumber="243">B</emphasis>
. (
<emphasis box="[960,976,1929,1954]" italics="true" pageId="42" pageNumber="243">T</emphasis>
.)
<emphasis box="[999,1089,1928,1954]" italics="true" pageId="42" pageNumber="243">bialatus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1096,1157,1928,1955]" pageId="42" pageNumber="243" rank="species">
<emphasis bold="true" box="[1096,1152,1928,1954]" pageId="42" pageNumber="243">sp. n</emphasis>
.
</taxonomicNameLabel>
, besides absence of hind wings, differs from
<taxonomicName authorityName="Muller-Liebenau &amp; Hubbard" authorityYear="1985" box="[373,567,1964,1990]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="42" pageNumber="243" phylum="Arthropoda" rank="species" species="frequentus" subGenus="Tenuibaetis">
<emphasis box="[373,397,1965,1990]" italics="true" pageId="42" pageNumber="243">B.</emphasis>
(
<emphasis box="[414,435,1965,1990]" italics="true" pageId="42" pageNumber="243">T.</emphasis>
)
<emphasis box="[450,567,1964,1990]" italics="true" pageId="42" pageNumber="243">frequentus</emphasis>
</taxonomicName>
by uniformly brown color of abdomen (
<figureCitation box="[1020,1118,1964,1990]" captionStart="FIGURES 215–224" captionStartId="43.[151,264,1770,1794]" captionTargetBox="[151,1435,181,1746]" captionTargetId="figure-19@43.[151,1435,181,1746]" captionTargetPageId="43" captionText="FIGURES 215–224. Baetis (Tenuibaetis) bialatus sp. n. 215, subimaginal exuviae of mesopleuron; 216, subimaginal exuviae of half of mesonotum; 217–218, exuviae of fore and hind legs of male subimago; 219–220, fore and middle legs of male imago; 221, male imago; 222, female imago; 223, subimaginal exuviae of abdominal segments VIII–IX; 224, egg (215–220, holotype)." figureDoi="http://doi.org/10.5281/zenodo.7889793" httpUri="https://zenodo.org/record/7889793/files/figure.png" pageId="42" pageNumber="243">Fig. 221</figureCitation>
).
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7889793" ID-Zenodo-Dep="7889793" httpUri="https://zenodo.org/record/7889793/files/figure.png" pageId="43" pageNumber="244" startId="43.[151,264,1770,1794]" targetBox="[151,1435,181,1746]" targetPageId="43" targetType="figure">
<paragraph blockId="43.[151,1436,1770,1902]" pageId="43" pageNumber="244">
<emphasis bold="true" box="[151,365,1770,1794]" pageId="43" pageNumber="244">FIGURES 215–224.</emphasis>
<taxonomicName authority="Kluge &amp; Srinivasan &amp; Sivaruban &amp; Barathy &amp; Isack, 2023" authorityName="Kluge &amp; Srinivasan &amp; Sivaruban &amp; Barathy &amp; Isack" authorityYear="2023" box="[373,664,1770,1795]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="43" pageNumber="244" phylum="Arthropoda" rank="species" species="bialatus" status="sp. n." subGenus="Tenuibaetis">
<emphasis box="[373,436,1771,1794]" italics="true" pageId="43" pageNumber="244">Baetis</emphasis>
(
<emphasis box="[451,567,1770,1794]" italics="true" pageId="43" pageNumber="244">Tenuibaetis</emphasis>
)
<emphasis box="[582,664,1770,1794]" italics="true" pageId="43" pageNumber="244">bialatus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[671,728,1771,1795]" pageId="43" pageNumber="244" rank="species">
<emphasis bold="true" box="[671,722,1771,1795]" pageId="43" pageNumber="244">sp. n</emphasis>
.
</taxonomicNameLabel>
215, subimaginal exuviae of mesopleuron; 216, subimaginal exuviae of half of mesonotum; 217–218, exuviae of fore and hind legs of male subimago; 219–220, fore and middle legs of male imago; 221, male imago; 222, female imago; 223, subimaginal exuviae of abdominal segments VIII–IX; 224, egg (215–220, holotype).
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7889797" ID-Zenodo-Dep="7889797" httpUri="https://zenodo.org/record/7889797/files/figure.png" pageId="44" pageNumber="245" startId="44.[151,264,1840,1864]" targetBox="[151,1436,185,1817]" targetPageId="44" targetType="figure">
<paragraph blockId="44.[151,1436,1840,1937]" pageId="44" pageNumber="245">
<emphasis bold="true" box="[151,363,1840,1864]" pageId="44" pageNumber="245">FIGURES 225–229.</emphasis>
<taxonomicName authority="Kluge &amp; Srinivasan &amp; Sivaruban &amp; Barathy &amp; Isack, 2023" authorityName="Kluge &amp; Srinivasan &amp; Sivaruban &amp; Barathy &amp; Isack" authorityYear="2023" box="[369,657,1840,1865]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="44" pageNumber="245" phylum="Arthropoda" rank="species" species="bialatus" status="sp. n." subGenus="Tenuibaetis">
<emphasis box="[369,432,1841,1864]" italics="true" pageId="44" pageNumber="245">Baetis</emphasis>
(
<emphasis box="[446,562,1840,1864]" italics="true" pageId="44" pageNumber="245">Tenuibaetis</emphasis>
)
<emphasis box="[575,657,1840,1864]" italics="true" pageId="44" pageNumber="245">bialatus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[662,718,1841,1865]" pageId="44" pageNumber="245" rank="species">
<emphasis bold="true" box="[662,711,1841,1865]" pageId="44" pageNumber="245">sp. n</emphasis>
.
</taxonomicNameLabel>
, genitalia of male imago. 225, 227, ventral view; 226, 229, ventral view with focus on sterno-styligeral muscle; 228, enlarged fragment of sterno-styligeral muscle to show cross-striation (225–226, holotype). Abbreviations: gv, gonovectis; s.s, sterno-styligeral muscle.
</paragraph>
</caption>
<subSubSection pageId="45" pageNumber="246" type="discussion">
<paragraph blockId="45.[151,1436,150,321]" pageId="45" pageNumber="246">
<emphasis bold="true" box="[199,324,150,176]" pageId="45" pageNumber="246">Comment.</emphasis>
Larvae from southern
<collectingCountry box="[600,658,151,177]" name="India" pageId="45" pageNumber="246">India</collectingCountry>
originally described under the name
<taxonomicName authorityName="Kumar, Sundar &amp; Sivaramakrishnan" authorityYear="2012" box="[1102,1436,151,177]" class="Insecta" family="Baetidae" genus="Indobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="45" pageNumber="246" phylum="Arthropoda" rank="species" species="michaelohubbardi">
<emphasis box="[1102,1436,151,177]" italics="true" pageId="45" pageNumber="246">Indobaetis michaelohubbardi</emphasis>
</taxonomicName>
Selva-kumar
<emphasis box="[304,356,187,213]" italics="true" pageId="45" pageNumber="246">et al</emphasis>
., 2012 were said to have ́hindwing pads absent» (Selva-kumar
<emphasis box="[1104,1156,187,213]" italics="true" pageId="45" pageNumber="246">et al</emphasis>
. 2012: 127). Later K.G. Sivaramakrishnan (personal communication) reported that actually the
<typeStatus box="[944,992,223,249]" pageId="45" pageNumber="246">type</typeStatus>
specimens have hind protoptera (
<bibRefCitation author="Kluge, N. J. &amp; Novikova, E. A." pageId="45" pageNumber="246" pagination="209 - 236" refId="ref18518" refString="Kluge, N. J. &amp; Novikova, E. A. (2014) Systematics of Indobaetis Muller-Liebenau &amp; Morihara 1982, and related implications for some other Baetidae genera (Ephemeroptera). Zootaxa, 3835 (2), 209 - 236. https: // doi. org / 10.11646 / zootaxa. 3835.2.3" type="journal article" year="2014">Kluge &amp; Novikova 2014: 215</bibRefCitation>
), so this species is not conspecific with
<taxonomicName authority="Kluge &amp; Srinivasan &amp; Sivaruban &amp; Barathy &amp; Isack, 2023" authorityName="Kluge &amp; Srinivasan &amp; Sivaruban &amp; Barathy &amp; Isack" authorityYear="2023" box="[851,1017,259,285]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="45" pageNumber="246" phylum="Arthropoda" rank="species" species="bialatus" status="sp. n." subGenus="Tenuibaetis">
<emphasis box="[851,876,259,284]" italics="true" pageId="45" pageNumber="246">B.</emphasis>
(
<emphasis box="[891,912,259,284]" italics="true" pageId="45" pageNumber="246">T.</emphasis>
)
<emphasis box="[927,1017,259,285]" italics="true" pageId="45" pageNumber="246">bialatus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1023,1086,259,285]" pageId="45" pageNumber="246" rank="species">
<emphasis bold="true" box="[1023,1079,259,285]" pageId="45" pageNumber="246">sp. n</emphasis>
.
</taxonomicNameLabel>
Here
<emphasis box="[1154,1171,259,284]" italics="true" pageId="45" pageNumber="246">B</emphasis>
.
<taxonomicName authorityName="Kumar, Sundar &amp; Sivaramakrishnan" authorityYear="2012" box="[1185,1436,259,285]" class="Insecta" family="Baetidae" genus="Indobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="45" pageNumber="246" phylum="Arthropoda" rank="species" species="michaelohubbardi" subGenus="Tenuibaetis">
(
<emphasis box="[1194,1210,259,284]" italics="true" pageId="45" pageNumber="246">T</emphasis>
.)
<emphasis box="[1232,1436,259,285]" italics="true" pageId="45" pageNumber="246">michaelohubbardi</emphasis>
</taxonomicName>
is synonymized with
<taxonomicName authorityName="Muller-Liebenau &amp; Hubbard" authorityYear="1985" box="[390,584,295,321]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="45" pageNumber="246" phylum="Arthropoda" rank="species" species="frequentus" subGenus="Tenuibaetis">
<emphasis box="[390,414,295,320]" italics="true" pageId="45" pageNumber="246">B.</emphasis>
(
<emphasis box="[430,452,295,320]" italics="true" pageId="45" pageNumber="246">T.</emphasis>
)
<emphasis box="[467,584,295,321]" italics="true" pageId="45" pageNumber="246">frequentus</emphasis>
</taxonomicName>
(see above).
</paragraph>
</subSubSection>
</treatment>
</document>
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