430 lines
56 KiB
XML
430 lines
56 KiB
XML
<document ID-DOI="10.11646/zootaxa.5277.2.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="7889695" ID-ZooBank="73ED69F3-3887-460D-86DE-C9F1302C8EC3" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1683099613832" checkinUser="plazi" docAuthor="Kluge, Nikita, Srinivasan, Pandiarajan, Sivaruban, T., Barathy, S. & Isack, Rajasekaran" docDate="2023" docId="03F487D0A3464D7CA2DBFF43FF4BEF42" docLanguage="en" docName="zootaxa.5277.2.1.pdf" docOrigin="Zootaxa 5277 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5277.2.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Baetis (Tenuibaetis) octomaculatus Kluge & Srinivasan & Sivaruban & Barathy & Isack 2023, sp. n." docType="treatment" docVersion="2" lastPageNumber="251" masterDocId="FFCDFFA8A3764D4EA24CFFD4FFABE864" masterDocTitle="Contribution to the knowledge of the subgenus Tenuibaetis Kang & Yang 1994 (Ephemeroptera, Baetidae, Baetis s. l.)" masterLastPageNumber="258" masterPageNumber="201" pageNumber="249" updateTime="1683100342607" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Contribution to the knowledge of the subgenus Tenuibaetis Kang & Yang 1994 (Ephemeroptera, Baetidae, Baetis s. l.)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Kluge, Nikita</mods:namePart>
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<mods:affiliation>Department of Entomology, Saint Petersburg State University, Universitetskaya nab., 7 / 9, Saint Petersburg, 199034, Russia. Website: http: // insecta. bio. spbu. ru</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Srinivasan, Pandiarajan</mods:namePart>
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<mods:affiliation>PG and Research department of Zoology, the American College, Madurai- 625002, India</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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||
</mods:role>
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<mods:namePart>Sivaruban, T.</mods:namePart>
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<mods:nameIdentifier type="ORCID">0000-0001-8997-9355</mods:nameIdentifier>
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<mods:affiliation>PG and Research department of Zoology, the American College, Madurai- 625002, India & sivaruban 270 @ gmail. com; https: // orcid. org / 0000 - 0001 - 8997 - 9355</mods:affiliation>
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<mods:nameIdentifier type="email">sivaruban270@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Barathy, S.</mods:namePart>
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<mods:affiliation>Department of Zoology, Fatima College, Madurai- 625018, India</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Isack, Rajasekaran</mods:namePart>
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<mods:nameIdentifier type="ORCID">0000-0002-9952-4335</mods:nameIdentifier>
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<mods:affiliation>PG and Research department of Zoology, the American College, Madurai- 625002, India & iceisack 143 @ gmail. com; https: // orcid. org / 0000 - 0002 - 9952 - 4335</mods:affiliation>
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<mods:nameIdentifier type="email">iceisack143@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Zootaxa</mods:title>
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||
</mods:titleInfo>
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||
<mods:part>
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||
<mods:date>2023</mods:date>
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<mods:detail type="pubDate">
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||
<mods:number>2023-05-02</mods:number>
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||
</mods:detail>
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||
<mods:detail type="volume">
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||
<mods:number>5277</mods:number>
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||
</mods:detail>
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||
<mods:detail type="issue">
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<mods:number>2</mods:number>
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||
</mods:detail>
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||
<mods:extent unit="page">
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<mods:start>201</mods:start>
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<mods:end>258</mods:end>
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</mods:extent>
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||
</mods:part>
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||
</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.11646/zootaxa.5277.2.1</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.11646/zootaxa.5277.2.1</mods:identifier>
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||
<mods:identifier type="ISSN">1175-5326</mods:identifier>
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||
<mods:identifier type="Zenodo-Dep">7889695</mods:identifier>
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<mods:identifier type="ZooBank">73ED69F3-3887-460D-86DE-C9F1302C8EC3</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.7889727" ID-Zenodo-Dep="7889727" LSID="urn:lsid:plazi:treatment:03F487D0A3464D7CA2DBFF43FF4BEF42" httpUri="http://treatment.plazi.org/id/03F487D0A3464D7CA2DBFF43FF4BEF42" lastPageId="50" lastPageNumber="251" pageId="48" pageNumber="249">
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||
<subSubSection box="[151,678,151,178]" pageId="48" pageNumber="249" type="nomenclature">
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||
<paragraph blockId="48.[151,678,151,214]" box="[151,678,151,178]" pageId="48" pageNumber="249">
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||
<heading bold="true" box="[151,678,151,178]" fontSize="11" level="1" pageId="48" pageNumber="249" reason="1">
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||
<emphasis bold="true" box="[151,678,151,178]" pageId="48" pageNumber="249">
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8.
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<taxonomicName authority="Kluge & Srinivasan & Sivaruban & Barathy & Isack, 2023" authorityName="Kluge & Srinivasan & Sivaruban & Barathy & Isack" authorityYear="2023" box="[181,605,151,178]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="48" pageNumber="249" phylum="Arthropoda" rank="species" species="octomaculatus" status="sp. n." subGenus="Tenuibaetis">
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<emphasis bold="true" box="[181,256,151,177]" italics="true" pageId="48" pageNumber="249">Baetis</emphasis>
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(
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<emphasis bold="true" box="[273,412,151,178]" italics="true" pageId="48" pageNumber="249">Tenuibaetis</emphasis>
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)
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<emphasis bold="true" box="[429,605,151,178]" italics="true" pageId="48" pageNumber="249">octomaculatus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[612,678,151,177]" pageId="48" pageNumber="249" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection box="[151,407,187,214]" pageId="48" pageNumber="249" type="description">
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||
<paragraph blockId="48.[151,678,151,214]" box="[151,407,187,214]" pageId="48" pageNumber="249">
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||
(
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<figureCitation box="[159,288,187,214]" captionStart="FIGURES 13–21" captionStartId="5.[151,264,1840,1864]" captionTargetBox="[151,1436,181,1816]" captionTargetId="figure-19@5.[151,1436,181,1816]" captionTargetPageId="5" captionText="FIGURES 13–21. Imaginal hind wings and larval protoptera (under the same magnification). 13–14, female Baetis (Tenuibaetis) frequentus from India; 15–17, male B. (T.) octomaculatus sp. n.; 18–19, female of same species; 20–21, male B. (T.) bialatus sp. n. (13, 15, 20, metathorax of imago with hind wing; 14, 21, larval exuviae of metanotum; 16, 18, hind wing of imago; 17, 19, cuticle from metathorax of larva ready to molt to subimago, with subimaginal wing inside protopteron: crumpled on Fig. 17, non-crumpled on Fig. 19)." figureDoi="http://doi.org/10.5281/zenodo.7889705" httpUri="https://zenodo.org/record/7889705/files/figure.png" pageId="48" pageNumber="249">Figs 15–19</figureCitation>
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,
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<figureCitation box="[300,399,187,213]" captionStart-0="FIGURES 242–248" captionStart-1="FIGURES 249–252" captionStart-2="FIGURES 253–256" captionStart-3="FIGURES 257–264" captionStart-4="FIGURES 265–274" captionStart-5="FIGURES 275–278" captionStartId-0="49.[151,264,1844,1868]" captionStartId-1="50.[151,264,1481,1505]" captionStartId-2="51.[151,264,1450,1474]" captionStartId-3="52.[151,264,1879,1903]" captionStartId-4="53.[151,264,1843,1867]" captionStartId-5="54.[151,264,1584,1608]" captionTargetBox-0="[151,1436,181,1820]" captionTargetBox-1="[151,1436,181,1457]" captionTargetBox-2="[151,1436,181,1426]" captionTargetBox-3="[151,1435,181,1855]" captionTargetBox-4="[151,1435,181,1819]" captionTargetBox-5="[151,1436,181,1560]" captionTargetId-0="figure-19@49.[151,1436,181,1820]" captionTargetId-1="figure-36@50.[151,1436,181,1457]" captionTargetId-2="figure-19@51.[151,1436,181,1426]" captionTargetId-3="figure-36@52.[151,1435,181,1855]" captionTargetId-4="figure-19@53.[151,1435,181,1819]" captionTargetId-5="figure-36@54.[151,1436,181,1560]" captionTargetPageId-0="49" captionTargetPageId-1="50" captionTargetPageId-2="51" captionTargetPageId-3="52" captionTargetPageId-4="53" captionTargetPageId-5="54" captionText-0="FIGURES 242–248. Baetis (Tenuibaetis) octomaculatus sp. n. 242–247, exuviae of female larvae: 242, 245, half of pronotum and mesonotum; 243, 246, head; 244, 247, abdomen; 248, young larva to show hypodermal coloration (242–244, holotype)." captionText-1="FIGURES 249–252. Baetis (Tenuibaetis) octomaculatus sp. n. 249, labrum (holotype); 250–251, left and right mandibles; 252, maxilla and labium." captionText-2="FIGURES 253–256. Baetis (Tenuibaetis) octomaculatus sp. n. (holotype). 253–255, larval exuviae of fore, middle and hind legs; 256, base of femur with villopore." captionText-3="FIGURES 257–264. Baetis (Tenuibaetis) octomaculatus sp. n. 257–258, fore and hind femora; 259–260, fore and middle tibiae; 261–263, abdominal terga I, VII and X; 264, abdominal sternum IX and paraprocts (257–260, holotype)." captionText-4="FIGURES 265–274. Baetis (Tenuibaetis) octomaculatus sp. n. 265, subimaginal exuviae of half of mesonotum; 266, male imago; 267, female imago; 268–269, fore legs of two female imagines (268, holotype); 270, female imago; 271, male subimago; 272, abdomen of male imago; 273, fore leg of male imago; 274, its subimaginal exuviae." captionText-5="FIGURES 275–278. Baetis (Tenuibaetis) octomaculatus sp. n. 275, subimaginal gonostyli crumpled under larval cuticle; 276, imaginal genitalia; 277, imaginal genitalia of another individual, with focus on ventral side; 278, the same, with focus on gonovectes. Abbreviation: s.s, sterno-styligeral muscle." figureDoi-0="http://doi.org/10.5281/zenodo.7889803" figureDoi-1="http://doi.org/10.5281/zenodo.7889805" figureDoi-2="http://doi.org/10.5281/zenodo.7889807" figureDoi-3="http://doi.org/10.5281/zenodo.7889809" figureDoi-4="http://doi.org/10.5281/zenodo.7889811" figureDoi-5="http://doi.org/10.5281/zenodo.7889813" httpUri-0="https://zenodo.org/record/7889803/files/figure.png" httpUri-1="https://zenodo.org/record/7889805/files/figure.png" httpUri-2="https://zenodo.org/record/7889807/files/figure.png" httpUri-3="https://zenodo.org/record/7889809/files/figure.png" httpUri-4="https://zenodo.org/record/7889811/files/figure.png" httpUri-5="https://zenodo.org/record/7889813/files/figure.png" pageId="48" pageNumber="249">242–278</figureCitation>
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||
)
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection pageId="48" pageNumber="249" type="etymology">
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||
<paragraph blockId="48.[151,1437,259,2014]" pageId="48" pageNumber="249">
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||
<emphasis bold="true" box="[151,286,259,285]" pageId="48" pageNumber="249">Etymology.</emphasis>
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||
Allusion to eight contrasting dark brown maculae on abdominal terga in all stages of both sexes (
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||
<figureCitation captionStart="FIGURES 242–248" captionStartId="49.[151,264,1844,1868]" captionTargetBox="[151,1436,181,1820]" captionTargetId="figure-19@49.[151,1436,181,1820]" captionTargetPageId="49" captionText="FIGURES 242–248. Baetis (Tenuibaetis) octomaculatus sp. n. 242–247, exuviae of female larvae: 242, 245, half of pronotum and mesonotum; 243, 246, head; 244, 247, abdomen; 248, young larva to show hypodermal coloration (242–244, holotype)." figureDoi="http://doi.org/10.5281/zenodo.7889803" httpUri="https://zenodo.org/record/7889803/files/figure.png" pageId="48" pageNumber="249">Figs 248</figureCitation>
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,
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<figureCitation box="[207,421,295,321]" captionStart="FIGURES 265–274" captionStartId="53.[151,264,1843,1867]" captionTargetBox="[151,1435,181,1819]" captionTargetId="figure-19@53.[151,1435,181,1819]" captionTargetPageId="53" captionText="FIGURES 265–274. Baetis (Tenuibaetis) octomaculatus sp. n. 265, subimaginal exuviae of half of mesonotum; 266, male imago; 267, female imago; 268–269, fore legs of two female imagines (268, holotype); 270, female imago; 271, male subimago; 272, abdomen of male imago; 273, fore leg of male imago; 274, its subimaginal exuviae." figureDoi="http://doi.org/10.5281/zenodo.7889811" httpUri="https://zenodo.org/record/7889811/files/figure.png" pageId="48" pageNumber="249">266, 267, 270, 271</figureCitation>
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).
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection lastPageId="50" lastPageNumber="251" pageId="48" pageNumber="249" type="materials_examined">
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||
<paragraph blockId="48.[151,1437,259,2014]" pageId="48" pageNumber="249">
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||
<emphasis bold="true" box="[199,431,331,357]" pageId="48" pageNumber="249">Material examined.</emphasis>
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||
<materialsCitation collectingDate="2015-02-14" collectorName="N. Kluge & L. Sheiko." country="Thailand" location="Pai" municipality="river Moei" pageId="48" pageNumber="249" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
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||
<emphasis bold="true" box="[436,554,331,357]" pageId="48" pageNumber="249">
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||
<typeStatus box="[436,549,331,357]" pageId="48" pageNumber="249">Holotype</typeStatus>
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||
:
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</emphasis>
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L-S-I
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<specimenCount box="[628,642,331,357]" pageId="48" pageNumber="249" type="female">♀</specimenCount>
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{specimen [XVIII](2) 2015},
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||
<collectingCountry box="[978,1126,331,357]" name="Thailand" pageId="48" pageNumber="249">THAILAND</collectingCountry>
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||
,
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||
<location LSID="urn:lsid:plazi:treatment:03F487D0A3464D7CA2DBFF43FF4BEF42:8E82601DA3464D7EA623FE9FFB3EE901" box="[1135,1173,331,357]" country="Thailand" municipality="river Moei" name="Pai" pageId="48" pageNumber="249">Pai</location>
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,
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||
<collectingMunicipality box="[1183,1303,331,357]" pageId="48" pageNumber="249">river Moei</collectingMunicipality>
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,
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||
<date box="[1313,1432,331,358]" pageId="48" pageNumber="249" value="2015-02-14">
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<collectingDate box="[1313,1432,331,358]" pageId="48" pageNumber="249" value="2015-02-14">14.II.2015</collectingDate>
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||
</date>
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||
, coll.
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||
<collectorName box="[211,317,367,394]" pageId="48" pageNumber="249">N. Kluge</collectorName>
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||
&
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||
<collectorName box="[360,480,367,394]" pageId="48" pageNumber="249">L. Sheiko.</collectorName>
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||
</materialsCitation>
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||
<emphasis bold="true" box="[490,619,367,393]" pageId="48" pageNumber="249">
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||
<typeStatus box="[490,614,367,393]" pageId="48" pageNumber="249">Paratypes</typeStatus>
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||
:
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||
</emphasis>
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||
the same place, date and collectors: 2 L-S-I
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||
<specimenCount box="[1150,1164,367,393]" pageId="48" pageNumber="249" type="female">♀</specimenCount>
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||
, 1 S♁,
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||
<specimenCount box="[1258,1369,367,393]" pageId="48" pageNumber="249" type="larva">30 larvae</specimenCount>
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||
;
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||
<materialsCitation collectingDate="1998-01-11" collectorName="Grigorenko" country="Thailand" location="river Pai" pageId="48" pageNumber="249" specimenCount="1" typeStatus="paratype">
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||
<location LSID="urn:lsid:plazi:treatment:03F487D0A3464D7CA2DBFF43FF4BEF42:8E82601DA3464D7EA72BFEBBFF16E9C9" country="Thailand" name="river Pai" pageId="48" pageNumber="249">river Pai</location>
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||
,
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||
<date box="[202,310,403,429]" pageId="48" pageNumber="249" value="1998-01-11">
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||
<collectingDate box="[202,310,403,429]" pageId="48" pageNumber="249" value="1998-01-11">11.I.1998</collectingDate>
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||
</date>
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||
, coll.
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||
<collectorName box="[382,516,403,429]" pageId="48" pageNumber="249">Grigorenko</collectorName>
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||
: 1 L/S
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||
</materialsCitation>
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||
♁;
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||
<materialsCitation box="[627,1338,403,430]" collectingDate="2015-02-15" collectingDateMax="2015-02-19" collectingDateMin="2015-02-15" collectionCode="II" collectorName="N. Kluge & L. Sheiko" country="Thailand" location="Mae-Sae" pageId="48" pageNumber="249" specimenCount="2" specimenCount-larva="2" typeStatus="paratype">
|
||
<location LSID="urn:lsid:plazi:treatment:03F487D0A3464D7CA2DBFF43FF4BEF42:8E82601DA3464D7EA03FFE47FD72E9C9" box="[627,729,403,429]" country="Thailand" name="Mae-Sae" pageId="48" pageNumber="249">Mae-Sae</location>
|
||
,
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||
<date box="[742,903,403,430]" pageId="48" pageNumber="249" value="2015-02-15" valueMax="2015-02-19" valueMin="2015-02-15">
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||
<collectingDate box="[742,903,403,430]" pageId="48" pageNumber="249" value="2015-02-15" valueMax="2015-02-19" valueMin="2015-02-15">
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||
15–19.
|
||
<collectionCode box="[817,837,403,430]" pageId="48" pageNumber="249">II</collectionCode>
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||
.2015
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||
</collectingDate>
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||
</date>
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||
, coll.
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||
<collectorName box="[974,1078,403,430]" pageId="48" pageNumber="249">N. Kluge</collectorName>
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||
&
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||
<collectorName box="[1117,1231,403,430]" pageId="48" pageNumber="249">L. Sheiko</collectorName>
|
||
:
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||
<specimenCount box="[1244,1338,403,429]" pageId="48" pageNumber="249" type="larva">2 larvae</specimenCount>
|
||
</materialsCitation>
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||
;
|
||
<materialsCitation collectingDate="2015-02-02" collectingDateMax="2015-02-04" collectingDateMin="2015-02-02" collectionCode="II" collectorName="N. Kluge & L. Sheiko" country="Thailand" location="Erawan National Park" pageId="48" pageNumber="249" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
|
||
<location LSID="urn:lsid:plazi:treatment:03F487D0A3464D7CA2DBFF43FF4BEF42:8E82601DA3464D7EA70BFE40FE9EE9B5" country="Thailand" name="Erawan National Park" pageId="48" pageNumber="249">Erawan National Park</location>
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||
,
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||
<date box="[320,453,439,466]" pageId="48" pageNumber="249" value="2015-02-02" valueMax="2015-02-04" valueMin="2015-02-02">
|
||
<collectingDate box="[320,453,439,466]" pageId="48" pageNumber="249" value="2015-02-02" valueMax="2015-02-04" valueMin="2015-02-02">
|
||
2–4.
|
||
<collectionCode box="[368,388,439,466]" pageId="48" pageNumber="249">II</collectionCode>
|
||
.2015
|
||
</collectingDate>
|
||
</date>
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||
, coll.
|
||
<collectorName box="[521,623,439,466]" pageId="48" pageNumber="249">N. Kluge</collectorName>
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||
&
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||
<collectorName box="[659,772,439,466]" pageId="48" pageNumber="249">L. Sheiko</collectorName>
|
||
: 1 S-I♁, 1 S-I
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||
<specimenCount box="[935,951,439,465]" pageId="48" pageNumber="249" type="female">♀</specimenCount>
|
||
</materialsCitation>
|
||
, 9 I♁,
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||
<specimenCount box="[1028,1121,439,465]" pageId="48" pageNumber="249" type="larva">2 larvae</specimenCount>
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||
.
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</paragraph>
|
||
<paragraph blockId="48.[151,1437,259,2014]" box="[199,348,475,501]" pageId="48" pageNumber="249">
|
||
<emphasis bold="true" box="[199,348,475,501]" pageId="48" pageNumber="249">Descriptions</emphasis>
|
||
</paragraph>
|
||
<paragraph blockId="48.[151,1437,259,2014]" pageId="48" pageNumber="249">
|
||
<emphasis bold="true" box="[199,275,511,537]" pageId="48" pageNumber="249">
|
||
<emphasis bold="true" box="[199,267,511,537]" italics="true" pageId="48" pageNumber="249">Larva</emphasis>
|
||
.
|
||
</emphasis>
|
||
CUTICULAR COLORATION: Head brown (
|
||
<figureCitation box="[824,987,511,537]" captionStart="FIGURES 242–248" captionStartId="49.[151,264,1844,1868]" captionTargetBox="[151,1436,181,1820]" captionTargetId="figure-19@49.[151,1436,181,1820]" captionTargetPageId="49" captionText="FIGURES 242–248. Baetis (Tenuibaetis) octomaculatus sp. n. 242–247, exuviae of female larvae: 242, 245, half of pronotum and mesonotum; 243, 246, head; 244, 247, abdomen; 248, young larva to show hypodermal coloration (242–244, holotype)." figureDoi="http://doi.org/10.5281/zenodo.7889803" httpUri="https://zenodo.org/record/7889803/files/figure.png" pageId="48" pageNumber="249">Figs 243, 246</figureCitation>
|
||
). Pronotum brown, with sigilla slightly darker than background; mesonotum and fore protoptera brown, with diffusive blanks (
|
||
<figureCitation box="[1156,1315,547,574]" captionStart="FIGURES 242–248" captionStartId="49.[151,264,1844,1868]" captionTargetBox="[151,1436,181,1820]" captionTargetId="figure-19@49.[151,1436,181,1820]" captionTargetPageId="49" captionText="FIGURES 242–248. Baetis (Tenuibaetis) octomaculatus sp. n. 242–247, exuviae of female larvae: 242, 245, half of pronotum and mesonotum; 243, 246, head; 244, 247, abdomen; 248, young larva to show hypodermal coloration (242–244, holotype)." figureDoi="http://doi.org/10.5281/zenodo.7889803" httpUri="https://zenodo.org/record/7889803/files/figure.png" pageId="48" pageNumber="249">Fig. 242, 245</figureCitation>
|
||
). Thoracic pleura brownish, sterna colorless. Legs with darker and lighter brown areas; fore femur with wide oval proximal blank (
|
||
<figureCitation box="[229,390,619,646]" captionStart="FIGURES 253–256" captionStartId="51.[151,264,1450,1474]" captionTargetBox="[151,1436,181,1426]" captionTargetId="figure-19@51.[151,1436,181,1426]" captionTargetPageId="51" captionText="FIGURES 253–256. Baetis (Tenuibaetis) octomaculatus sp. n. (holotype). 253–255, larval exuviae of fore, middle and hind legs; 256, base of femur with villopore." figureDoi="http://doi.org/10.5281/zenodo.7889807" httpUri="https://zenodo.org/record/7889807/files/figure.png" pageId="48" pageNumber="249">Figs 253–255</figureCitation>
|
||
). Abdominal terga vary from nearly uniformly brown to more or less contrasting; tergum
|
||
<collectionCode box="[1407,1437,620,646]" pageId="48" pageNumber="249">IV</collectionCode>
|
||
either nearly as dark as others (
|
||
<figureCitation box="[500,598,655,681]" captionStart="FIGURES 242–248" captionStartId="49.[151,264,1844,1868]" captionTargetBox="[151,1436,181,1820]" captionTargetId="figure-19@49.[151,1436,181,1820]" captionTargetPageId="49" captionText="FIGURES 242–248. Baetis (Tenuibaetis) octomaculatus sp. n. 242–247, exuviae of female larvae: 242, 245, half of pronotum and mesonotum; 243, 246, head; 244, 247, abdomen; 248, young larva to show hypodermal coloration (242–244, holotype)." figureDoi="http://doi.org/10.5281/zenodo.7889803" httpUri="https://zenodo.org/record/7889803/files/figure.png" pageId="48" pageNumber="249">Fig. 247</figureCitation>
|
||
), or colorless laterally (
|
||
<figureCitation box="[862,961,655,681]" captionStart="FIGURES 242–248" captionStartId="49.[151,264,1844,1868]" captionTargetBox="[151,1436,181,1820]" captionTargetId="figure-19@49.[151,1436,181,1820]" captionTargetPageId="49" captionText="FIGURES 242–248. Baetis (Tenuibaetis) octomaculatus sp. n. 242–247, exuviae of female larvae: 242, 245, half of pronotum and mesonotum; 243, 246, head; 244, 247, abdomen; 248, young larva to show hypodermal coloration (242–244, holotype)." figureDoi="http://doi.org/10.5281/zenodo.7889803" httpUri="https://zenodo.org/record/7889803/files/figure.png" pageId="48" pageNumber="249">Fig. 244</figureCitation>
|
||
). Caudalii uniformly brown.
|
||
</paragraph>
|
||
<paragraph blockId="48.[151,1437,259,2014]" pageId="48" pageNumber="249">
|
||
HYPODERMAL COLORATION: In young and mature larvae of both sexes, abdominal terga
|
||
<collectionCode box="[1324,1342,692,718]" pageId="48" pageNumber="249">II</collectionCode>
|
||
–
|
||
<collectionCode box="[1357,1385,692,718]" pageId="48" pageNumber="249">III</collectionCode>
|
||
and
|
||
<collectionCode box="[151,180,728,754]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="48" pageNumber="249">VI</collectionCode>
|
||
–
|
||
<collectionCode box="[195,234,728,754]" pageId="48" pageNumber="249">VII</collectionCode>
|
||
with pair of contrasting dark brown maculae; these and other abdominal terga with dark brown line on posterior margin (
|
||
<figureCitation box="[352,450,763,789]" captionStart="FIGURES 242–248" captionStartId="49.[151,264,1844,1868]" captionTargetBox="[151,1436,181,1820]" captionTargetId="figure-19@49.[151,1436,181,1820]" captionTargetPageId="49" captionText="FIGURES 242–248. Baetis (Tenuibaetis) octomaculatus sp. n. 242–247, exuviae of female larvae: 242, 245, half of pronotum and mesonotum; 243, 246, head; 244, 247, abdomen; 248, young larva to show hypodermal coloration (242–244, holotype)." figureDoi="http://doi.org/10.5281/zenodo.7889803" httpUri="https://zenodo.org/record/7889803/files/figure.png" pageId="48" pageNumber="249">Fig. 248</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="48.[151,1437,259,2014]" pageId="48" pageNumber="249">
|
||
<collectionCode box="[200,512,799,825]" pageId="48" pageNumber="249">SHAPE AND SETATION</collectionCode>
|
||
: Labrum of medium proportions (
|
||
<figureCitation box="[909,1009,799,825]" captionStart="FIGURES 249–252" captionStartId="50.[151,264,1481,1505]" captionTargetBox="[151,1436,181,1457]" captionTargetId="figure-36@50.[151,1436,181,1457]" captionTargetPageId="50" captionText="FIGURES 249–252. Baetis (Tenuibaetis) octomaculatus sp. n. 249, labrum (holotype); 250–251, left and right mandibles; 252, maxilla and labium." figureDoi="http://doi.org/10.5281/zenodo.7889805" httpUri="https://zenodo.org/record/7889805/files/figure.png" pageId="48" pageNumber="249">Fig. 249</figureCitation>
|
||
). Mandibles—
|
||
<figureCitation box="[1178,1337,799,826]" captionStart="FIGURES 249–252" captionStartId="50.[151,264,1481,1505]" captionTargetBox="[151,1436,181,1457]" captionTargetId="figure-36@50.[151,1436,181,1457]" captionTargetPageId="50" captionText="FIGURES 249–252. Baetis (Tenuibaetis) octomaculatus sp. n. 249, labrum (holotype); 250–251, left and right mandibles; 252, maxilla and labium." figureDoi="http://doi.org/10.5281/zenodo.7889805" httpUri="https://zenodo.org/record/7889805/files/figure.png" pageId="48" pageNumber="249">Figs 250–251</figureCitation>
|
||
. Maxilla and labium—
|
||
<figureCitation box="[308,402,835,862]" captionStart="FIGURES 249–252" captionStartId="50.[151,264,1481,1505]" captionTargetBox="[151,1436,181,1457]" captionTargetId="figure-36@50.[151,1436,181,1457]" captionTargetPageId="50" captionText="FIGURES 249–252. Baetis (Tenuibaetis) octomaculatus sp. n. 249, labrum (holotype); 250–251, left and right mandibles; 252, maxilla and labium." figureDoi="http://doi.org/10.5281/zenodo.7889805" httpUri="https://zenodo.org/record/7889805/files/figure.png" pageId="48" pageNumber="249">Fig. 252</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph blockId="48.[151,1437,259,2014]" box="[199,623,871,898]" pageId="48" pageNumber="249">
|
||
Hind protoptera present (
|
||
<figureCitation box="[480,611,871,897]" captionStart="FIGURES 13–21" captionStartId="5.[151,264,1840,1864]" captionTargetBox="[151,1436,181,1816]" captionTargetId="figure-19@5.[151,1436,181,1816]" captionTargetPageId="5" captionText="FIGURES 13–21. Imaginal hind wings and larval protoptera (under the same magnification). 13–14, female Baetis (Tenuibaetis) frequentus from India; 15–17, male B. (T.) octomaculatus sp. n.; 18–19, female of same species; 20–21, male B. (T.) bialatus sp. n. (13, 15, 20, metathorax of imago with hind wing; 14, 21, larval exuviae of metanotum; 16, 18, hind wing of imago; 17, 19, cuticle from metathorax of larva ready to molt to subimago, with subimaginal wing inside protopteron: crumpled on Fig. 17, non-crumpled on Fig. 19)." figureDoi="http://doi.org/10.5281/zenodo.7889705" httpUri="https://zenodo.org/record/7889705/files/figure.png" pageId="48" pageNumber="249">Figs 17–19</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="48.[151,1437,259,2014]" pageId="48" pageNumber="249">
|
||
Stout two-channel setae forming row along outer side of femur (including two subapical setae) elongate, blunt, distally widened and flattened (
|
||
<figureCitation box="[503,663,943,970]" captionStart="FIGURES 253–256" captionStartId="51.[151,264,1450,1474]" captionTargetBox="[151,1436,181,1426]" captionTargetId="figure-19@51.[151,1436,181,1426]" captionTargetPageId="51" captionText="FIGURES 253–256. Baetis (Tenuibaetis) octomaculatus sp. n. (holotype). 253–255, larval exuviae of fore, middle and hind legs; 256, base of femur with villopore." figureDoi="http://doi.org/10.5281/zenodo.7889807" httpUri="https://zenodo.org/record/7889807/files/figure.png" pageId="48" pageNumber="249">Figs 253–255</figureCitation>
|
||
). Anterior-apical margin of femur with small, pseudo-bifurcate setae, as on tibiae (
|
||
<figureCitation box="[292,390,979,1006]" captionStart="FIGURES 257–264" captionStartId="52.[151,264,1879,1903]" captionTargetBox="[151,1435,181,1855]" captionTargetId="figure-36@52.[151,1435,181,1855]" captionTargetPageId="52" captionText="FIGURES 257–264. Baetis (Tenuibaetis) octomaculatus sp. n. 257–258, fore and hind femora; 259–260, fore and middle tibiae; 261–263, abdominal terga I, VII and X; 264, abdominal sternum IX and paraprocts (257–260, holotype)." figureDoi="http://doi.org/10.5281/zenodo.7889809" httpUri="https://zenodo.org/record/7889809/files/figure.png" pageId="48" pageNumber="249">Fig. 257</figureCitation>
|
||
). Stout setae on anterior side of femur mostly elongate, distally widened and rounded, varying from large to small (
|
||
<figureCitation box="[379,537,1015,1042]" captionStart="FIGURES 253–256" captionStartId="51.[151,264,1450,1474]" captionTargetBox="[151,1436,181,1426]" captionTargetId="figure-19@51.[151,1436,181,1426]" captionTargetPageId="51" captionText="FIGURES 253–256. Baetis (Tenuibaetis) octomaculatus sp. n. (holotype). 253–255, larval exuviae of fore, middle and hind legs; 256, base of femur with villopore." figureDoi="http://doi.org/10.5281/zenodo.7889807" httpUri="https://zenodo.org/record/7889807/files/figure.png" pageId="48" pageNumber="249">Figs 253–255</figureCitation>
|
||
,
|
||
<figureCitation box="[548,651,1015,1042]" captionStart="FIGURES 257–264" captionStartId="52.[151,264,1879,1903]" captionTargetBox="[151,1435,181,1855]" captionTargetId="figure-36@52.[151,1435,181,1855]" captionTargetPageId="52" captionText="FIGURES 257–264. Baetis (Tenuibaetis) octomaculatus sp. n. 257–258, fore and hind femora; 259–260, fore and middle tibiae; 261–263, abdominal terga I, VII and X; 264, abdominal sternum IX and paraprocts (257–260, holotype)." figureDoi="http://doi.org/10.5281/zenodo.7889809" httpUri="https://zenodo.org/record/7889809/files/figure.png" pageId="48" pageNumber="249">257–258</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="48.[151,1437,259,2014]" pageId="48" pageNumber="249">
|
||
Tibia of each leg with pseudo-bifurcate setae on outer side, smaller and sparser on fore leg (
|
||
<figureCitation box="[1251,1351,1051,1078]" captionStart="FIGURES 257–264" captionStartId="52.[151,264,1879,1903]" captionTargetBox="[151,1435,181,1855]" captionTargetId="figure-36@52.[151,1435,181,1855]" captionTargetPageId="52" captionText="FIGURES 257–264. Baetis (Tenuibaetis) octomaculatus sp. n. 257–258, fore and hind femora; 259–260, fore and middle tibiae; 261–263, abdominal terga I, VII and X; 264, abdominal sternum IX and paraprocts (257–260, holotype)." figureDoi="http://doi.org/10.5281/zenodo.7889809" httpUri="https://zenodo.org/record/7889809/files/figure.png" pageId="48" pageNumber="249">Fig. 259</figureCitation>
|
||
), larger and denser on middle and hind legs (
|
||
<figureCitation box="[575,677,1087,1113]" captionStart="FIGURES 257–264" captionStartId="52.[151,264,1879,1903]" captionTargetBox="[151,1435,181,1855]" captionTargetId="figure-36@52.[151,1435,181,1855]" captionTargetPageId="52" captionText="FIGURES 257–264. Baetis (Tenuibaetis) octomaculatus sp. n. 257–258, fore and hind femora; 259–260, fore and middle tibiae; 261–263, abdominal terga I, VII and X; 264, abdominal sternum IX and paraprocts (257–260, holotype)." figureDoi="http://doi.org/10.5281/zenodo.7889809" httpUri="https://zenodo.org/record/7889809/files/figure.png" pageId="48" pageNumber="249">Fig. 260</figureCitation>
|
||
). Stout setae along patella-tibial suture two-channel, short, distally widened and rounded (
|
||
<figureCitation box="[407,567,1123,1150]" captionStart="FIGURES 257–264" captionStartId="52.[151,264,1879,1903]" captionTargetBox="[151,1435,181,1855]" captionTargetId="figure-36@52.[151,1435,181,1855]" captionTargetPageId="52" captionText="FIGURES 257–264. Baetis (Tenuibaetis) octomaculatus sp. n. 257–258, fore and hind femora; 259–260, fore and middle tibiae; 261–263, abdominal terga I, VII and X; 264, abdominal sternum IX and paraprocts (257–260, holotype)." figureDoi="http://doi.org/10.5281/zenodo.7889809" httpUri="https://zenodo.org/record/7889809/files/figure.png" pageId="48" pageNumber="249">Figs 259–260</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="48.[151,1437,259,2014]" pageId="48" pageNumber="249">
|
||
Denticles on posterior margins of all abdominal terga I–X well-developed, short and blunt (
|
||
<figureCitation box="[1262,1425,1159,1185]" captionStart="FIGURES 257–264" captionStartId="52.[151,264,1879,1903]" captionTargetBox="[151,1435,181,1855]" captionTargetId="figure-36@52.[151,1435,181,1855]" captionTargetPageId="52" captionText="FIGURES 257–264. Baetis (Tenuibaetis) octomaculatus sp. n. 257–258, fore and hind femora; 259–260, fore and middle tibiae; 261–263, abdominal terga I, VII and X; 264, abdominal sternum IX and paraprocts (257–260, holotype)." figureDoi="http://doi.org/10.5281/zenodo.7889809" httpUri="https://zenodo.org/record/7889809/files/figure.png" pageId="48" pageNumber="249">Figs 261–263</figureCitation>
|
||
). Denticles on posterior margin of sternum triangular, present on sternum
|
||
<collectionCode box="[976,1006,1196,1222]" pageId="48" pageNumber="249">IX</collectionCode>
|
||
(
|
||
<figureCitation box="[1022,1122,1195,1221]" captionStart="FIGURES 257–264" captionStartId="52.[151,264,1879,1903]" captionTargetBox="[151,1435,181,1855]" captionTargetId="figure-36@52.[151,1435,181,1855]" captionTargetPageId="52" captionText="FIGURES 257–264. Baetis (Tenuibaetis) octomaculatus sp. n. 257–258, fore and hind femora; 259–260, fore and middle tibiae; 261–263, abdominal terga I, VII and X; 264, abdominal sternum IX and paraprocts (257–260, holotype)." figureDoi="http://doi.org/10.5281/zenodo.7889809" httpUri="https://zenodo.org/record/7889809/files/figure.png" pageId="48" pageNumber="249">Fig. 264</figureCitation>
|
||
), fewer on sternum
|
||
<collectionCode box="[1350,1402,1196,1222]" pageId="48" pageNumber="249">VIII</collectionCode>
|
||
; in male row of denticles not interrupted by protogonostyli. Paraproct with pointed denticles (
|
||
<figureCitation box="[1192,1294,1231,1257]" captionStart="FIGURES 257–264" captionStartId="52.[151,264,1879,1903]" captionTargetBox="[151,1435,181,1855]" captionTargetId="figure-36@52.[151,1435,181,1855]" captionTargetPageId="52" captionText="FIGURES 257–264. Baetis (Tenuibaetis) octomaculatus sp. n. 257–258, fore and hind femora; 259–260, fore and middle tibiae; 261–263, abdominal terga I, VII and X; 264, abdominal sternum IX and paraprocts (257–260, holotype)." figureDoi="http://doi.org/10.5281/zenodo.7889809" httpUri="https://zenodo.org/record/7889809/files/figure.png" pageId="48" pageNumber="249">Fig. 264</figureCitation>
|
||
). Tergalius I much smaller than others (
|
||
<figureCitation box="[448,607,1267,1293]" captionStart="FIGURES 242–248" captionStartId="49.[151,264,1844,1868]" captionTargetBox="[151,1436,181,1820]" captionTargetId="figure-19@49.[151,1436,181,1820]" captionTargetPageId="49" captionText="FIGURES 242–248. Baetis (Tenuibaetis) octomaculatus sp. n. 242–247, exuviae of female larvae: 242, 245, half of pronotum and mesonotum; 243, 246, head; 244, 247, abdomen; 248, young larva to show hypodermal coloration (242–244, holotype)." figureDoi="http://doi.org/10.5281/zenodo.7889803" httpUri="https://zenodo.org/record/7889803/files/figure.png" pageId="48" pageNumber="249">Figs 244, 247</figureCitation>
|
||
). Paracercus significantly smaller than cerci (
|
||
<figureCitation box="[1116,1214,1267,1293]" captionStart="FIGURES 242–248" captionStartId="49.[151,264,1844,1868]" captionTargetBox="[151,1436,181,1820]" captionTargetId="figure-19@49.[151,1436,181,1820]" captionTargetPageId="49" captionText="FIGURES 242–248. Baetis (Tenuibaetis) octomaculatus sp. n. 242–247, exuviae of female larvae: 242, 245, half of pronotum and mesonotum; 243, 246, head; 244, 247, abdomen; 248, young larva to show hypodermal coloration (242–244, holotype)." figureDoi="http://doi.org/10.5281/zenodo.7889803" httpUri="https://zenodo.org/record/7889803/files/figure.png" pageId="48" pageNumber="249">Fig. 247</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="48.[151,1437,259,2014]" pageId="48" pageNumber="249">
|
||
<emphasis bold="true" box="[199,323,1303,1330]" pageId="48" pageNumber="249">
|
||
<emphasis bold="true" box="[199,316,1303,1330]" italics="true" pageId="48" pageNumber="249">Subimago</emphasis>
|
||
.
|
||
</emphasis>
|
||
CUTICULAR COLORATION: Head colorless, antennae brown. Pronotum with light brown and colorless areas. Mesonotum mostly light brownish (
|
||
<figureCitation box="[738,837,1339,1366]" captionStart="FIGURES 265–274" captionStartId="53.[151,264,1843,1867]" captionTargetBox="[151,1435,181,1819]" captionTargetId="figure-19@53.[151,1435,181,1819]" captionTargetPageId="53" captionText="FIGURES 265–274. Baetis (Tenuibaetis) octomaculatus sp. n. 265, subimaginal exuviae of half of mesonotum; 266, male imago; 267, female imago; 268–269, fore legs of two female imagines (268, holotype); 270, female imago; 271, male subimago; 272, abdomen of male imago; 273, fore leg of male imago; 274, its subimaginal exuviae." figureDoi="http://doi.org/10.5281/zenodo.7889811" httpUri="https://zenodo.org/record/7889811/files/figure.png" pageId="48" pageNumber="249">Fig. 265</figureCitation>
|
||
). Thoracic pleura and sterna with light brownish and dark brown areas (as in
|
||
<figureCitation box="[419,517,1375,1402]" captionStart="FIGURES 215–224" captionStartId="43.[151,264,1770,1794]" captionTargetBox="[151,1435,181,1746]" captionTargetId="figure-19@43.[151,1435,181,1746]" captionTargetPageId="43" captionText="FIGURES 215–224. Baetis (Tenuibaetis) bialatus sp. n. 215, subimaginal exuviae of mesopleuron; 216, subimaginal exuviae of half of mesonotum; 217–218, exuviae of fore and hind legs of male subimago; 219–220, fore and middle legs of male imago; 221, male imago; 222, female imago; 223, subimaginal exuviae of abdominal segments VIII–IX; 224, egg (215–220, holotype)." figureDoi="http://doi.org/10.5281/zenodo.7889793" httpUri="https://zenodo.org/record/7889793/files/figure.png" pageId="48" pageNumber="249">Fig. 215</figureCitation>
|
||
). Legs brownish (
|
||
<figureCitation box="[717,816,1375,1401]" captionStart="FIGURES 265–274" captionStartId="53.[151,264,1843,1867]" captionTargetBox="[151,1435,181,1819]" captionTargetId="figure-19@53.[151,1435,181,1819]" captionTargetPageId="53" captionText="FIGURES 265–274. Baetis (Tenuibaetis) octomaculatus sp. n. 265, subimaginal exuviae of half of mesonotum; 266, male imago; 267, female imago; 268–269, fore legs of two female imagines (268, holotype); 270, female imago; 271, male subimago; 272, abdomen of male imago; 273, fore leg of male imago; 274, its subimaginal exuviae." figureDoi="http://doi.org/10.5281/zenodo.7889811" httpUri="https://zenodo.org/record/7889811/files/figure.png" pageId="48" pageNumber="249">Fig. 274</figureCitation>
|
||
). Abdominal terga and sterna light brownish.
|
||
</paragraph>
|
||
<paragraph blockId="48.[151,1437,259,2014]" pageId="48" pageNumber="249">
|
||
HYPODERMAL COLORATION: In both sexes, abdominal terga mostly ochre; terga
|
||
<collectionCode box="[1183,1201,1412,1438]" pageId="48" pageNumber="249">II</collectionCode>
|
||
–
|
||
<collectionCode box="[1216,1244,1412,1438]" pageId="48" pageNumber="249">III</collectionCode>
|
||
and
|
||
<collectionCode box="[1297,1326,1412,1438]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="48" pageNumber="249">VI</collectionCode>
|
||
–
|
||
<collectionCode box="[1341,1380,1412,1438]" pageId="48" pageNumber="249">VII</collectionCode>
|
||
with pair of contrasting dark brown maculae; these and other abdominal terga with dark brown line on posterior margin (
|
||
<figureCitation box="[159,258,1483,1509]" captionStart="FIGURES 265–274" captionStartId="53.[151,264,1843,1867]" captionTargetBox="[151,1435,181,1819]" captionTargetId="figure-19@53.[151,1435,181,1819]" captionTargetPageId="53" captionText="FIGURES 265–274. Baetis (Tenuibaetis) octomaculatus sp. n. 265, subimaginal exuviae of half of mesonotum; 266, male imago; 267, female imago; 268–269, fore legs of two female imagines (268, holotype); 270, female imago; 271, male subimago; 272, abdomen of male imago; 273, fore leg of male imago; 274, its subimaginal exuviae." figureDoi="http://doi.org/10.5281/zenodo.7889811" httpUri="https://zenodo.org/record/7889811/files/figure.png" pageId="48" pageNumber="249">Fig. 271</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="48.[151,1437,259,2014]" pageId="48" pageNumber="249">
|
||
<emphasis bold="true" box="[199,346,1519,1546]" italics="true" pageId="48" pageNumber="249">Imago, male</emphasis>
|
||
(
|
||
<figureCitation box="[362,526,1519,1545]" captionStart="FIGURES 265–274" captionStartId="53.[151,264,1843,1867]" captionTargetBox="[151,1435,181,1819]" captionTargetId="figure-19@53.[151,1435,181,1819]" captionTargetPageId="53" captionText="FIGURES 265–274. Baetis (Tenuibaetis) octomaculatus sp. n. 265, subimaginal exuviae of half of mesonotum; 266, male imago; 267, female imago; 268–269, fore legs of two female imagines (268, holotype); 270, female imago; 271, male subimago; 272, abdomen of male imago; 273, fore leg of male imago; 274, its subimaginal exuviae." figureDoi="http://doi.org/10.5281/zenodo.7889811" httpUri="https://zenodo.org/record/7889811/files/figure.png" pageId="48" pageNumber="249">Figs 266, 272</figureCitation>
|
||
). Head light brown. Antennae light brown. Turbinate eyes with facetted surface orange, stem ochre. Thorax brown. Forewing with membrane entirely colorless; veins ochre or brown; pterostigma with few oblique cross veins and disconnected vein fragments. Hind wing diminished, colorless, with short and non-hooked costal projection, with 2 longitudinal veins (
|
||
<figureCitation box="[745,878,1627,1654]" captionStart="FIGURES 13–21" captionStartId="5.[151,264,1840,1864]" captionTargetBox="[151,1436,181,1816]" captionTargetId="figure-19@5.[151,1436,181,1816]" captionTargetPageId="5" captionText="FIGURES 13–21. Imaginal hind wings and larval protoptera (under the same magnification). 13–14, female Baetis (Tenuibaetis) frequentus from India; 15–17, male B. (T.) octomaculatus sp. n.; 18–19, female of same species; 20–21, male B. (T.) bialatus sp. n. (13, 15, 20, metathorax of imago with hind wing; 14, 21, larval exuviae of metanotum; 16, 18, hind wing of imago; 17, 19, cuticle from metathorax of larva ready to molt to subimago, with subimaginal wing inside protopteron: crumpled on Fig. 17, non-crumpled on Fig. 19)." figureDoi="http://doi.org/10.5281/zenodo.7889705" httpUri="https://zenodo.org/record/7889705/files/figure.png" pageId="48" pageNumber="249">Figs 15–16</figureCitation>
|
||
). Legs either uniformly ochre (
|
||
<figureCitation box="[1234,1334,1627,1653]" captionStart="FIGURES 265–274" captionStartId="53.[151,264,1843,1867]" captionTargetBox="[151,1435,181,1819]" captionTargetId="figure-19@53.[151,1435,181,1819]" captionTargetPageId="53" captionText="FIGURES 265–274. Baetis (Tenuibaetis) octomaculatus sp. n. 265, subimaginal exuviae of half of mesonotum; 266, male imago; 267, female imago; 268–269, fore legs of two female imagines (268, holotype); 270, female imago; 271, male subimago; 272, abdomen of male imago; 273, fore leg of male imago; 274, its subimaginal exuviae." figureDoi="http://doi.org/10.5281/zenodo.7889811" httpUri="https://zenodo.org/record/7889811/files/figure.png" pageId="48" pageNumber="249">Fig. 273</figureCitation>
|
||
), or with diffusive reddish markings. Middle and hind tarsi with 2 apical spines (on 1st+2nd and 3rd segments). Abdominal terga
|
||
<collectionCode box="[214,232,1700,1726]" pageId="48" pageNumber="249">II</collectionCode>
|
||
–
|
||
<collectionCode box="[247,275,1700,1726]" pageId="48" pageNumber="249">III</collectionCode>
|
||
and
|
||
<collectionCode box="[328,357,1700,1726]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="48" pageNumber="249">VI</collectionCode>
|
||
–
|
||
<collectionCode box="[372,411,1700,1726]" pageId="48" pageNumber="249">VII</collectionCode>
|
||
with pair of contrasting dark brown maculae and brownish-ochre area between them; these and other abdominal terga with dark brown line on posterior margin; terga
|
||
<collectionCode box="[1007,1037,1736,1762]" pageId="48" pageNumber="249">IV</collectionCode>
|
||
–
|
||
<collectionCode box="[1051,1071,1736,1762]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="48" pageNumber="249" type="Museum">V</collectionCode>
|
||
mostly light ochre, tergum
|
||
<collectionCode box="[1388,1436,1736,1762]" pageId="48" pageNumber="249">VIII</collectionCode>
|
||
mostly brownish-ochre, terga
|
||
<collectionCode box="[485,514,1772,1798]" pageId="48" pageNumber="249">IX</collectionCode>
|
||
–X lighter ochre (
|
||
<figureCitation box="[711,868,1771,1797]" captionStart="FIGURES 265–274" captionStartId="53.[151,264,1843,1867]" captionTargetBox="[151,1435,181,1819]" captionTargetId="figure-19@53.[151,1435,181,1819]" captionTargetPageId="53" captionText="FIGURES 265–274. Baetis (Tenuibaetis) octomaculatus sp. n. 265, subimaginal exuviae of half of mesonotum; 266, male imago; 267, female imago; 268–269, fore legs of two female imagines (268, holotype); 270, female imago; 271, male subimago; 272, abdomen of male imago; 273, fore leg of male imago; 274, its subimaginal exuviae." figureDoi="http://doi.org/10.5281/zenodo.7889811" httpUri="https://zenodo.org/record/7889811/files/figure.png" pageId="48" pageNumber="249">Figs 266, 272</figureCitation>
|
||
. Cerci ochre. Gonostyli ochre; 1st segment slightly narrowing toward apex; 3rd (terminal) segment short, often truncate (
|
||
<figureCitation box="[966,1069,1807,1833]" captionStart="FIGURES 275–278" captionStartId="54.[151,264,1584,1608]" captionTargetBox="[151,1436,181,1560]" captionTargetId="figure-36@54.[151,1436,181,1560]" captionTargetPageId="54" captionText="FIGURES 275–278. Baetis (Tenuibaetis) octomaculatus sp. n. 275, subimaginal gonostyli crumpled under larval cuticle; 276, imaginal genitalia; 277, imaginal genitalia of another individual, with focus on ventral side; 278, the same, with focus on gonovectes. Abbreviation: s.s, sterno-styligeral muscle." figureDoi="http://doi.org/10.5281/zenodo.7889813" httpUri="https://zenodo.org/record/7889813/files/figure.png" pageId="48" pageNumber="249">Fig. 277</figureCitation>
|
||
), sometimes clavate (
|
||
<figureCitation box="[1321,1424,1807,1833]" captionStart="FIGURES 275–278" captionStartId="54.[151,264,1584,1608]" captionTargetBox="[151,1436,181,1560]" captionTargetId="figure-36@54.[151,1436,181,1560]" captionTargetPageId="54" captionText="FIGURES 275–278. Baetis (Tenuibaetis) octomaculatus sp. n. 275, subimaginal gonostyli crumpled under larval cuticle; 276, imaginal genitalia; 277, imaginal genitalia of another individual, with focus on ventral side; 278, the same, with focus on gonovectes. Abbreviation: s.s, sterno-styligeral muscle." figureDoi="http://doi.org/10.5281/zenodo.7889813" httpUri="https://zenodo.org/record/7889813/files/figure.png" pageId="48" pageNumber="249">Fig. 276</figureCitation>
|
||
). Gonovectes sharply bent, light brownish. Sterno-styligeral muscle vestigial (
|
||
<figureCitation box="[1009,1107,1843,1869]" captionStart="FIGURES 275–278" captionStartId="54.[151,264,1584,1608]" captionTargetBox="[151,1436,181,1560]" captionTargetId="figure-36@54.[151,1436,181,1560]" captionTargetPageId="54" captionText="FIGURES 275–278. Baetis (Tenuibaetis) octomaculatus sp. n. 275, subimaginal gonostyli crumpled under larval cuticle; 276, imaginal genitalia; 277, imaginal genitalia of another individual, with focus on ventral side; 278, the same, with focus on gonovectes. Abbreviation: s.s, sterno-styligeral muscle." figureDoi="http://doi.org/10.5281/zenodo.7889813" httpUri="https://zenodo.org/record/7889813/files/figure.png" pageId="48" pageNumber="249">Fig. 277</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="48.[151,1437,259,2014]" pageId="48" pageNumber="249">
|
||
<emphasis bold="true" box="[199,366,1879,1906]" italics="true" pageId="48" pageNumber="249">Imago, female</emphasis>
|
||
(
|
||
<figureCitation box="[380,539,1879,1905]" captionStart="FIGURES 265–274" captionStartId="53.[151,264,1843,1867]" captionTargetBox="[151,1435,181,1819]" captionTargetId="figure-19@53.[151,1435,181,1819]" captionTargetPageId="53" captionText="FIGURES 265–274. Baetis (Tenuibaetis) octomaculatus sp. n. 265, subimaginal exuviae of half of mesonotum; 266, male imago; 267, female imago; 268–269, fore legs of two female imagines (268, holotype); 270, female imago; 271, male subimago; 272, abdomen of male imago; 273, fore leg of male imago; 274, its subimaginal exuviae." figureDoi="http://doi.org/10.5281/zenodo.7889811" httpUri="https://zenodo.org/record/7889811/files/figure.png" pageId="48" pageNumber="249">Figs 267, 270</figureCitation>
|
||
). Head ochre. Thorax ochre. Hind wing much smaller than in male (
|
||
<figureCitation box="[1299,1382,1879,1905]" captionStart="FIGURES 13–21" captionStartId="5.[151,264,1840,1864]" captionTargetBox="[151,1436,181,1816]" captionTargetId="figure-19@5.[151,1436,181,1816]" captionTargetPageId="5" captionText="FIGURES 13–21. Imaginal hind wings and larval protoptera (under the same magnification). 13–14, female Baetis (Tenuibaetis) frequentus from India; 15–17, male B. (T.) octomaculatus sp. n.; 18–19, female of same species; 20–21, male B. (T.) bialatus sp. n. (13, 15, 20, metathorax of imago with hind wing; 14, 21, larval exuviae of metanotum; 16, 18, hind wing of imago; 17, 19, cuticle from metathorax of larva ready to molt to subimago, with subimaginal wing inside protopteron: crumpled on Fig. 17, non-crumpled on Fig. 19)." figureDoi="http://doi.org/10.5281/zenodo.7889705" httpUri="https://zenodo.org/record/7889705/files/figure.png" pageId="48" pageNumber="249">Fig. 18</figureCitation>
|
||
). All tarsi with 2 apical spines (fore tarsus on 2nd and 3rd segments, middle and hind tarsi on 1st+2nd and 3rd segments). Abdominal terga mostly ochre; terga
|
||
<collectionCode box="[564,582,1952,1978]" pageId="48" pageNumber="249">II</collectionCode>
|
||
–
|
||
<collectionCode box="[597,625,1952,1978]" pageId="48" pageNumber="249">III</collectionCode>
|
||
and
|
||
<collectionCode box="[676,705,1952,1978]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="48" pageNumber="249">VI</collectionCode>
|
||
–
|
||
<collectionCode box="[720,759,1952,1978]" pageId="48" pageNumber="249">VII</collectionCode>
|
||
with pair of contrasting dark brown maculae; these and other abdominal terga with dark brown line on posterior margin (
|
||
<figureCitation box="[815,975,1987,2013]" captionStart="FIGURES 265–274" captionStartId="53.[151,264,1843,1867]" captionTargetBox="[151,1435,181,1819]" captionTargetId="figure-19@53.[151,1435,181,1819]" captionTargetPageId="53" captionText="FIGURES 265–274. Baetis (Tenuibaetis) octomaculatus sp. n. 265, subimaginal exuviae of half of mesonotum; 266, male imago; 267, female imago; 268–269, fore legs of two female imagines (268, holotype); 270, female imago; 271, male subimago; 272, abdomen of male imago; 273, fore leg of male imago; 274, its subimaginal exuviae." figureDoi="http://doi.org/10.5281/zenodo.7889811" httpUri="https://zenodo.org/record/7889811/files/figure.png" pageId="48" pageNumber="249">Figs 267, 270</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7889803" ID-Zenodo-Dep="7889803" httpUri="https://zenodo.org/record/7889803/files/figure.png" pageId="49" pageNumber="250" startId="49.[151,264,1844,1868]" targetBox="[151,1436,181,1820]" targetPageId="49" targetType="figure">
|
||
<paragraph blockId="49.[151,1437,1844,1905]" pageId="49" pageNumber="250">
|
||
<emphasis bold="true" box="[151,365,1844,1868]" pageId="49" pageNumber="250">FIGURES 242–248.</emphasis>
|
||
<taxonomicName authority="Kluge & Srinivasan & Sivaruban & Barathy & Isack, 2023" authorityName="Kluge & Srinivasan & Sivaruban & Barathy & Isack" authorityYear="2023" box="[372,729,1844,1868]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="49" pageNumber="250" phylum="Arthropoda" rank="species" species="octomaculatus" status="sp. n." subGenus="Tenuibaetis">
|
||
<emphasis box="[372,435,1845,1868]" italics="true" pageId="49" pageNumber="250">Baetis</emphasis>
|
||
(
|
||
<emphasis box="[450,566,1844,1868]" italics="true" pageId="49" pageNumber="250">Tenuibaetis</emphasis>
|
||
)
|
||
<emphasis box="[581,729,1844,1868]" italics="true" pageId="49" pageNumber="250">octomaculatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[735,791,1844,1868]" pageId="49" pageNumber="250" rank="species">
|
||
<emphasis bold="true" box="[735,785,1844,1868]" pageId="49" pageNumber="250">sp. n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
242–247, exuviae of female larvae: 242, 245, half of pronotum and mesonotum; 243, 246, head; 244, 247, abdomen; 248, young larva to show hypodermal coloration (242–244, holotype).
|
||
</paragraph>
|
||
</caption>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7889805" ID-Zenodo-Dep="7889805" httpUri="https://zenodo.org/record/7889805/files/figure.png" pageId="50" pageNumber="251" startId="50.[151,264,1481,1505]" targetBox="[151,1436,181,1457]" targetPageId="50" targetType="figure">
|
||
<paragraph blockId="50.[151,1436,1481,1541]" pageId="50" pageNumber="251">
|
||
<emphasis bold="true" box="[151,364,1481,1505]" pageId="50" pageNumber="251">FIGURES 249–252.</emphasis>
|
||
<taxonomicName authority="Kluge & Srinivasan & Sivaruban & Barathy & Isack, 2023" authorityName="Kluge & Srinivasan & Sivaruban & Barathy & Isack" authorityYear="2023" box="[371,726,1481,1506]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="50" pageNumber="251" phylum="Arthropoda" rank="species" species="octomaculatus" status="sp. n." subGenus="Tenuibaetis">
|
||
<emphasis box="[371,434,1482,1505]" italics="true" pageId="50" pageNumber="251">Baetis</emphasis>
|
||
(
|
||
<emphasis box="[448,564,1481,1505]" italics="true" pageId="50" pageNumber="251">Tenuibaetis</emphasis>
|
||
)
|
||
<emphasis box="[578,726,1481,1505]" italics="true" pageId="50" pageNumber="251">octomaculatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[732,786,1482,1505]" pageId="50" pageNumber="251" rank="species">sp. n.</taxonomicNameLabel>
|
||
249, labrum (holotype); 250–251, left and right mandibles; 252, maxilla and labium.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="50.[151,1437,1587,1830]" box="[199,604,1587,1614]" pageId="50" pageNumber="251">
|
||
<emphasis bold="true" box="[199,334,1587,1613]" pageId="50" pageNumber="251">Dimension.</emphasis>
|
||
Forewing length
|
||
<quantity box="[532,600,1588,1614]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="50" pageNumber="251" unit="mm" value="4.0">4 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="50" pageNumber="251" type="distribution">
|
||
<paragraph blockId="50.[151,1437,1587,1830]" box="[199,466,1623,1650]" pageId="50" pageNumber="251">
|
||
<emphasis bold="true" box="[199,353,1623,1649]" pageId="50" pageNumber="251">Distribution.</emphasis>
|
||
<collectingCountry box="[359,462,1624,1650]" name="Thailand" pageId="50" pageNumber="251">Thailand</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph blockId="50.[151,1437,1587,1830]" pageId="50" pageNumber="251">
|
||
<emphasis bold="true" box="[199,354,1659,1685]" pageId="50" pageNumber="251">Comparison.</emphasis>
|
||
The new species differs from other known species by peculiar hypodermal coloration which is well-expressed in all stages of the both sexes, i.e. with eight dark brown maculae, by a pair on the abdominal terga II, III, VI and VII. By greatly diminished hind wings of female, the new species resembles
|
||
<emphasis box="[1193,1210,1732,1757]" italics="true" pageId="50" pageNumber="251">B</emphasis>
|
||
.
|
||
<taxonomicName authorityName="Kluge & Srinivasan & Sivaruban & Barathy & Isack" authorityYear="2023" baseAuthorityName="Kluge & Srinivasan & Sivaruban & Barathy & Isack" baseAuthorityYear="2011" box="[1226,1404,1732,1758]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="50" pageNumber="251" phylum="Arthropoda" rank="species" species="parvipterus" subGenus="Tenuibaetis">
|
||
(
|
||
<emphasis box="[1235,1251,1732,1757]" italics="true" pageId="50" pageNumber="251">T</emphasis>
|
||
.)
|
||
<emphasis box="[1275,1404,1732,1757]" italics="true" pageId="50" pageNumber="251">parvipterus</emphasis>
|
||
</taxonomicName>
|
||
; it differs from
|
||
<emphasis box="[293,310,1768,1793]" italics="true" pageId="50" pageNumber="251">B</emphasis>
|
||
.
|
||
<taxonomicName authorityName="Kluge & Srinivasan & Sivaruban & Barathy & Isack" authorityYear="2023" baseAuthorityName="Kluge & Srinivasan & Sivaruban & Barathy & Isack" baseAuthorityYear="2011" box="[325,503,1768,1794]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="50" pageNumber="251" phylum="Arthropoda" rank="species" species="parvipterus" subGenus="Tenuibaetis">
|
||
(
|
||
<emphasis box="[334,350,1768,1793]" italics="true" pageId="50" pageNumber="251">T</emphasis>
|
||
.)
|
||
<emphasis box="[374,503,1768,1793]" italics="true" pageId="50" pageNumber="251">parvipterus</emphasis>
|
||
</taxonomicName>
|
||
, besides the hypodermal coloration, by very long stout setae on anterior surface of femur.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |