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<document id="ABFFD9E01347682A2FD6EFD068A6D130" ID="10.11646/zootaxa.4273.4.2SLASH11258" ID-CLB-Dataset="32892" ID-DOI="10.11646/zootaxa.4273.4.2" ID-GBIF-Dataset="316e1968-2a32-462e-a554-99165078f1fc" ID-ISSN="1175-5326" ID-Zenodo-Dep="803688" ID-ZooBank="77091F8E-2A3C-4A9B-81C2-656B5BF72C22" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1496820183743" checkinUser="plazi" docAuthor="Majnon-Jahromi, Bahareh, Gheibi, Mehdi, Fallahzadeh, Majid, Kehlmaier, Christian &amp; Hesami, Shahram" docDate="2017" docId="03F487A3B074FF90B8D3FD92B76765F0" docLanguage="en" docName="zootaxa.4273.4.2SLASH11258.pdf" docOrigin="Zootaxa 4273 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tomosvaryella angulata Kehlmaier &amp; Majnon-Jahromi, sp. nov." docType="treatment" docVersion="7" lastPageNumber="493" masterDocId="FFCDFFDBB071FF95B844FC24B20A632E" masterDocTitle="Pipunculidae (Diptera) from southern Iran, including two new species of the genus Tomosvaryella Aczél" masterLastPageNumber="500" masterPageNumber="488" pageNumber="493" updateTime="1698440239418" updateUser="ExternalLinkService">
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<mods:title id="82AF53A7A4EC535ECE8DF65FE0AEDFA9">Pipunculidae (Diptera) from southern Iran, including two new species of the genus Tomosvaryella Aczél</mods:title>
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<mods:namePart id="276F4AFF01A501F7DBA0DDA90EFD60D8">Majnon-Jahromi, Bahareh</mods:namePart>
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<mods:namePart id="47559B93655A79832AEDB9610B9B6D83">Gheibi, Mehdi</mods:namePart>
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<mods:namePart id="60C6616D5A034EF3B3233BF657B81F1C">Fallahzadeh, Majid</mods:namePart>
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<mods:namePart id="0E3A52421E8931D78FDDBF143CA9FB9E">Kehlmaier, Christian</mods:namePart>
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<mods:date id="1F8882CEED7BDA0E906981FF5698BBF4">2017</mods:date>
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<treatment id="03F487A3B074FF90B8D3FD92B76765F0" ID-DOI="http://doi.org/10.5281/zenodo.6020265" ID-GBIF-Taxon="131812481" ID-Zenodo-Dep="6020265" LSID="urn:lsid:plazi:treatment:03F487A3B074FF90B8D3FD92B76765F0" httpUri="http://treatment.plazi.org/id/03F487A3B074FF90B8D3FD92B76765F0" lastPageNumber="493" pageId="5" pageNumber="493">
<subSubSection id="C347653EB074FF90B8D3FD92B2F362DF" pageId="5" pageNumber="493" type="nomenclature">
<paragraph id="8BE236B5B074FF90B8D3FD92B1AD62FE" blockId="5.[151,935,438,498]" box="[151,935,438,464]" pageId="5" pageNumber="493">
<heading id="D0AA81D9B074FF90B8D3FD92B1AD62FE" bold="true" box="[151,935,438,464]" fontSize="11" level="1" pageId="5" pageNumber="493" reason="1">
<emphasis id="B929EAA7B074FF90B8D3FD92B1AD62FE" bold="true" box="[151,935,438,464]" pageId="5" pageNumber="493">
<taxonomicName id="4C5D4D36B074FF90B8D3FD92B14862FE" ID-CoL="9HKN3" authority="Kehlmaier &amp; Majnon-Jahromi" authorityName="Kehlmaier &amp; Majnon-Jahromi" box="[151,834,438,464]" class="Insecta" family="Pipunculidae" genus="Tomosvaryella" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="493" phylum="Arthropoda" rank="species" species="angulata" status="sp. nov.">
<emphasis id="B929EAA7B074FF90B8D3FD92B3B262FE" bold="true" box="[151,440,438,464]" italics="true" pageId="5" pageNumber="493">Tomosvaryella angulata</emphasis>
Kehlmaier &amp; Majnon-Jahromi
</taxonomicName>
<taxonomicNameLabel id="A21A57DCB074FF90BB0DFD92B1AD62FE" box="[841,935,438,464]" pageId="5" pageNumber="493" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BE236B5B074FF90B8D3FDFDB2F362DF" blockId="5.[151,935,438,498]" box="[151,249,473,498]" pageId="5" pageNumber="493">
<figureCitation id="13662A30B074FF90B8D3FDFDB2F362DF" box="[151,249,473,498]" captionStart="FIGURES 3 8" captionStartId="6.[151,264,1729,1752]" captionTargetBox="[188,1398,267,1707]" captionTargetId="figure@6.[188,1398,267,1707]" captionTargetPageId="6" captionText="FIGURES 3 8. Tomosvaryella angulata Kehlmaier &amp; Majnon-Jahromi sp. nov. — 3 6: male paratype (DNA CK 831); 7 8: female paratype (DNA CK 804). 3 — right surstylus and phallus, lateral view; 4 — syntergosternite 8, epandrium and surstyli, dorsal view; 5 — left surstylus, lateral view; 6 — phallus, lateral view; 7 — ovipositor, dorsal view; 8 — ovipositor, lateral view. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/803694/files/figure.png" pageId="5" pageNumber="493">Figs 38</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C347653EB074FF90B8D3FE05B2F561AC" pageId="5" pageNumber="493" type="materials_examined">
<paragraph id="8BE236B5B074FF90B8D3FE05B2F561AC" blockId="5.[151,1436,545,1758]" pageId="5" pageNumber="493">
<emphasis id="B929EAA7B074FF90B8D3FE05B34E6114" bold="true" box="[151,324,545,570]" pageId="5" pageNumber="493">Type material.</emphasis>
HOLOTYPE: 1♂, loc. 7,
<date id="FFE31075B074FF90BA2EFE05B0FE6114" box="[618,756,545,570]" pageId="5" pageNumber="493" value="2014-05-09" valueMax="2014-05-16" valueMin="2014-05-09">916.v.2014</date>
(SMTD); PARATYPES: 1♂ (DNA CK831,
<accessionNumber id="940EAB56B074FF90BCACFE05B76A6114" box="[1256,1376,545,570]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/LT671787" pageId="5" pageNumber="493">LT671787</accessionNumber>
), loc. 7,
<date id="FFE31075B074FF90B8F0FE61B3346170" box="[180,318,581,606]" pageId="5" pageNumber="493" value="2014-05-09" valueMax="2014-05-16" valueMin="2014-05-09">916.v.2014</date>
(SMTD); 1♂, loc. 4,
<date id="FFE31075B074FF90BA72FE61B0D26170" box="[566,728,581,606]" pageId="5" pageNumber="493" value="2015-06-18" valueMax="2015-06-28" valueMin="2015-06-18">1828.vi.2015</date>
(SHIAU); 1♀ (DNA CK804,
<accessionNumber id="940EAB56B074FF90BC74FE61B6A36170" box="[1072,1193,581,606]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/LT671786" pageId="5" pageNumber="493">LT671786</accessionNumber>
), loc. 3,
<date id="FFE31075B074FF90BD4EFE61B7966170" box="[1290,1436,581,606]" pageId="5" pageNumber="493" value="2012-04-05" valueMax="2012-04-15" valueMin="2012-04-05">515.iv.2012</date>
(SMTD).
</paragraph>
</subSubSection>
<subSubSection id="C347653EB074FF90B883FEA9B1D86504" pageId="5" pageNumber="493" type="description">
<paragraph id="8BE236B5B074FF90B883FEA9B1D86504" blockId="5.[151,1436,545,1758]" pageId="5" pageNumber="493">
<emphasis id="B929EAA7B074FF90B883FEA9B3BE6188" bold="true" box="[199,436,653,678]" pageId="5" pageNumber="493">Description. MALE</emphasis>
. Body length
<quantity id="4CA59B50B074FF90BA10FEAAB0D4618B" box="[596,734,654,678]" metricMagnitude="-3" metricUnit="m" metricValue="3.0999999999999996" metricValueMax="3.3" metricValueMin="2.9" pageId="5" pageNumber="493" unit="mm" value="3.0999999999999996" valueMax="3.3" valueMin="2.9">2.93.3 mm</quantity>
; wing length
<quantity id="4CA59B50B074FF90BB33FEA9B60B618B" box="[887,1025,653,678]" metricMagnitude="-3" metricUnit="m" metricValue="3.65" metricValueMax="3.8" metricValueMin="3.5" pageId="5" pageNumber="493" unit="mm" value="3.65" valueMax="3.8" valueMin="3.5">3.53.8 mm</quantity>
; eyes meeting for a distance slightly more than twice the length of ocellar triangle, i.e., for approximately six to nine ommatidia; pedicellus with two minute bristles dorsally and one ventrally; flagellum pointed, tawny, tip may be paler; thorax dark brown except postpronotal lobe yellow; dorsal bristles hardly discernible, black; wing membrane with a faint brownish tinge; wing membrane covered in microtrichia except for the base, including the anal lobe basally; third costal segment 1/ 3 the length of fourth costal segment; crossvein r-m reaching cell dm at or shortly after the middle; crossvein bm-cu interrupted right before reaching cell br on both wings; halter with dark base, whitish stem and light brown knob; leg dark brown except for the narrowly yellow apex of femora, basal 1/3 and apex of tibia and tarsal segments except distitarsus; hind trochanter ventrally smooth, without spines or projection; fore femora with posteroventral row of minute black spines in apical half; mid femora with antero- and posteroventral row of minute black spines for more than apical half; hind femora with anteroventral row of minute black spines in apical half; pulvilli slightly shorter than distitarsus; abdomen dark brown; tergite 1 with lateral patch of about ten short black setae on each side, slightly shorter than width of tarsal segments; dorsal setae of abdomen black, hardly discernible; sclerites dark brown, apparently evenly sclerotised; genitalia, in dorsal view, with short epandrium, broader than long, slightly less sclerotised in the central part; surstyli almost symmetrical, gradually tapering towards apices, which are distinctly knobbed; left surstylus slightly protruding the right one; membrane of syntergosternite 8 large, almost reaching epandrium, caudally occupying 1/3 of syntergosternite 8; in lateral view, apical half of surstyli bent towards venter by 90° (see inner margin); phallus trifid, with two longer and one shorter upright and wavy ejaculatory ducts, the longest one carrying about six scale-like projections at its base (see at high magnification, i.e., 100x); phallic guide small and inconspicuous, tip pointing towards epandrium.
<emphasis id="B929EAA7B074FF90BC1FF91DB6D9667C" bold="true" box="[1115,1235,1337,1362]" pageId="5" pageNumber="493">FEMALE</emphasis>
. Body length
<quantity id="4CA59B50B074FF90BD3DF91EB2CC665B" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="5" pageNumber="493" unit="mm" value="2.7">2.7 mm</quantity>
; wing length
<quantity id="4CA59B50B074FF90B922F97AB3CB665B" box="[358,449,1374,1398]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="5" pageNumber="493" unit="mm" value="3.2">3.2 mm</quantity>
, differing from male by dichoptic eyes; frontal ommatidial facets distinctly enlarged; frons widened in its middle, twice the width of the ocellar triangle or equal to the length of four enlarged ommatidial facets; flagellum with paler apical half; crossvein r-m reaching cell dm shortly after the middle; pulvilli of fore and mid legs as long as distitarsi; ovipositor with dark base and yellow piercer; in lateral view, base of ovipositor with a distinct ventral protuberance; piercer gently undulating; in dorsal view, base of ovipositor rather trapezoid and piercer with weak lateral flanges in the middle of its length.
</paragraph>
</subSubSection>
<subSubSection id="C347653EB074FF90B883FA11B04F6597" pageId="5" pageNumber="493" type="diagnosis">
<paragraph id="8BE236B5B074FF90B883FA11B04F6597" blockId="5.[151,1436,545,1758]" pageId="5" pageNumber="493">
<emphasis id="B929EAA7B074FF90B883FA11B3DB6560" bold="true" box="[199,465,1589,1614]" pageId="5" pageNumber="493">Differential diagnosis.</emphasis>
To the best of our knowledge,
<taxonomicName id="4C5D4D36B074FF90BB03FA12B1C06560" box="[839,970,1589,1614]" class="Insecta" family="Pipunculidae" genus="Tomosvaryella" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="493" phylum="Arthropoda" rank="species" species="angulata" status="sp. nov.">
<emphasis id="B929EAA7B074FF90BB03FA12B1C06560" box="[839,970,1589,1614]" italics="true" pageId="5" pageNumber="493">T. angulata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21A57DCB074FF90BB90FA11B63B6563" box="[980,1073,1589,1614]" pageId="5" pageNumber="493" rank="species">
<emphasis id="B929EAA7B074FF90BB90FA11B1E46560" bold="true" box="[980,1006,1589,1614]" pageId="5" pageNumber="493">sp</emphasis>
.
<emphasis id="B929EAA7B074FF90BC44FA11B6206560" bold="true" box="[1024,1066,1589,1614]" pageId="5" pageNumber="493">nov</emphasis>
.
</taxonomicNameLabel>
is a unique species among the Palaearctic
<taxonomicName id="4C5D4D36B074FF90B95EFA7DB3A5655C" box="[282,431,1625,1650]" class="Insecta" family="Pipunculidae" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="493" phylum="Arthropoda" rank="family">Pipunculidae</taxonomicName>
, due to its distinctly apically-knobbed and laterally-bent surstyli in the male, and due to the weakly undulating ovipositor that bears a distinct ventral protuberance in the female (
<figureCitation id="13662A30B074FF90BCE4FA59B70365B8" box="[1184,1289,1661,1686]" captionStart="FIGURES 3 8" captionStartId="6.[151,264,1729,1752]" captionTargetBox="[188,1398,267,1707]" captionTargetId="figure@6.[188,1398,267,1707]" captionTargetPageId="6" captionText="FIGURES 3 8. Tomosvaryella angulata Kehlmaier &amp; Majnon-Jahromi sp. nov. — 3 6: male paratype (DNA CK 831); 7 8: female paratype (DNA CK 804). 3 — right surstylus and phallus, lateral view; 4 — syntergosternite 8, epandrium and surstyli, dorsal view; 5 — left surstylus, lateral view; 6 — phallus, lateral view; 7 — ovipositor, dorsal view; 8 — ovipositor, lateral view. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/803694/files/figure.png" pageId="5" pageNumber="493">Figs 38</figureCitation>
). For further remarks, see under
<taxonomicName id="4C5D4D36B074FF90B934FA86B3E96597" box="[368,483,1698,1721]" class="Insecta" family="Pipunculidae" genus="Tomosvaryella" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="493" phylum="Arthropoda" rank="species" species="pistacia" status="sp. nov.">
<emphasis id="B929EAA7B074FF90B934FA86B3E96597" box="[368,483,1698,1721]" italics="true" pageId="5" pageNumber="493">T. pistacia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21A57DCB074FF90B9A8FA85B04F6597" box="[492,581,1697,1722]" pageId="5" pageNumber="493" rank="species">
<emphasis id="B929EAA7B074FF90B9A8FA85B00C6594" bold="true" box="[492,518,1697,1722]" pageId="5" pageNumber="493">sp</emphasis>
.
<emphasis id="B929EAA7B074FF90BA50FA85B0346594" bold="true" box="[532,574,1697,1722]" pageId="5" pageNumber="493">nov</emphasis>
.
</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C347653EB074FF90B883FAE1B76765F0" box="[199,1389,1733,1758]" pageId="5" pageNumber="493" type="etymology">
<paragraph id="8BE236B5B074FF90B883FAE1B76765F0" blockId="5.[151,1436,545,1758]" box="[199,1389,1733,1758]" pageId="5" pageNumber="493">
<emphasis id="B929EAA7B074FF90B883FAE1B34465F0" bold="true" box="[199,334,1733,1758]" pageId="5" pageNumber="493">Etymology.</emphasis>
The Latin adjective
<taxonomicName id="4C5D4D36B074FF90BA71FAE1B09365F0" box="[565,665,1733,1758]" class="Insecta" family="Pipunculidae" genus="Tomosvaryella" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="493" phylum="Arthropoda" rank="species" species="angulata">
<emphasis id="B929EAA7B074FF90BA71FAE1B09365F0" box="[565,665,1733,1758]" italics="true" pageId="5" pageNumber="493">angulata</emphasis>
</taxonomicName>
(English: angular) refers to the angled surstyli of male genitalia.
</paragraph>
</subSubSection>
</treatment>
</document>