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<document id="2663EFC844157AACCFBBF089ED2EEC57" ID-CLB-Dataset="264215" ID-DOI="10.1080/00222933.2023.2226375" ID-GBIF-Dataset="79c6cc16-cda1-4ea6-9ffb-05855d9cbfcf" ID-ISSN="1464-5262" ID-Zenodo-Dep="8270799" ID-ZooBank="FFDF1435-92D6-4C19-9B51-3AB61E1BD7DA" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1692610082376" checkinUser="plazi" docAuthor="Gustafsson, Daniel R., Grossi, Alexandra A., Ren, Mengjiao &amp; Zou, Fasheng" docDate="2023" docId="03A25902FFA0CF3B3E55FDCFFE85FA02" docLanguage="en" docName="JNATHIST.57.17-20.996-1048.pdf" docOrigin="Journal of Natural History 57 (17 - 20)" docSource="http://dx.doi.org/10.1080/00222933.2023.2226375" docStyle="DocumentStyle:3C762F9783B768A90048B82ACD135A62.3:JNatHist.2017-.journal_article.0cover" docStyleId="3C762F9783B768A90048B82ACD135A62" docStyleName="JNatHist.2017-.journal_article.0cover" docStyleVersion="3" docTitle="Goniocotes rectangulatus Nitzsch 1866" docType="treatment" docVersion="6" lastPageNumber="1022" masterDocId="FF9B217AFFB4CF213F37FFE1FFE9FFA8" masterDocTitle="The Goniodidae (Phthiraptera: Ischnocera) of peafowl (Aves: Galliformes: Pavo), with description of a new genus" masterLastPageNumber="1048" masterPageNumber="996" pageNumber="1016" updateTime="1699594595493" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="849D7467CBAEE16AB9F12381DE7D91EC">The Goniodidae (Phthiraptera: Ischnocera) of peafowl (Aves: Galliformes: Pavo), with description of a new genus</mods:title>
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<mods:namePart id="5A423F3A5EA65889D449F1FE52D43CD1">Gustafsson, Daniel R.</mods:namePart>
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<paragraph id="8BB4E814FFA0CF353E55FDCFFC29FDE0" blockId="20.[354,960,558,584]" box="[354,960,558,584]" pageId="20" pageNumber="1016">
<heading id="D0FC5F78FFA0CF353E55FDCFFC29FDE0" box="[354,960,558,584]" centered="true" fontSize="11" level="2" pageId="20" pageNumber="1016" reason="5">
<taxonomicName id="4C0B9397FFA0CF353E55FDCFFD11FDE0" ID-CoL="3GWGX" authority="Nitzsch" authorityName="Nitzsch" authorityYear="1866" box="[354,760,558,584]" class="Insecta" family="Philopteridae" genus="Goniocotes" kingdom="Animalia" order="Psocodea" pageId="20" pageNumber="1016" phylum="Arthropoda" rank="species" species="rectangulatus">
<emphasis id="B97F3406FFA0CF353E55FDCFFD75FDE0" bold="true" box="[354,668,558,584]" italics="true" pageId="20" pageNumber="1016">Goniocotes rectangulatus</emphasis>
Nitzsch
</taxonomicName>
[in Giebel], 1866
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<subSubSection id="C311BB9FFFA0CF353F97FD82FC1DFCAF" pageId="20" pageNumber="1016" type="reference_group">
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<treatmentCitation id="0AAACE05FFA0CF353F97FD82FD9AFDD4" author="Nitzsch CL" box="[160,627,611,636]" page="294" pageId="20" pageNumber="1016" year="1818">
<taxonomicName id="4C0B9397FFA0CF353F97FD82FD9AFDD4" authority="Nitzsch, 1818: 294" authorityName="Nitzsch" authorityPageNumber="294" authorityYear="1818" box="[160,627,611,636]" class="Insecta" family="Philopteridae" genus="Goniodes" kingdom="Animalia" order="Psocodea" pageId="20" pageNumber="1016" phylum="Arthropoda" rank="species" species="rectangulatus">
<emphasis id="B97F3406FFA0CF353F97FD82FE48FDD4" box="[160,417,611,636]" italics="true" pageId="20" pageNumber="1016">Goniodes rectangulatus</emphasis>
<bibRefCitation id="EF9A95E5FFA0CF353E9EFD82FD9AFDD4" author="Nitzsch CL" box="[425,627,611,636]" pageId="20" pageNumber="1016" pagination="261 - 316" refId="ref20470" refString="Nitzsch CL. 1818. Die Familien und Gattungen der Thierinsekten (insecta epizoica); als Prodromus einer Naturgeschechte derselben. Magazin der Entomologie. 3: 261 - 316." type="journal article" year="1818">Nitzsch, 1818: 294</bibRefCitation>
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<emphasis id="B97F3406FFA0CF353D4CFD82FCF5FDD4" box="[635,796,611,636]" italics="true" pageId="20" pageNumber="1016">nomen nudum</emphasis>
.
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</paragraph>
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<treatmentCitationGroup id="AB1BCF3AFFA0CF353F97FD67FEABFD69" pageId="20" pageNumber="1016">
<taxonomicName id="4C0B9397FFA0CF353F97FD67FD54FD37" authority="(Nitzsch)" baseAuthorityName="Nitzsch" box="[160,701,646,671]" class="Insecta" family="Philopteridae" genus="Goniodes" kingdom="Animalia" order="Psocodea" pageId="20" pageNumber="1016" phylum="Arthropoda" rank="species" species="rectangulus" subGenus="Philopedon">
<emphasis id="B97F3406FFA0CF353F97FD67FEE1FD37" box="[160,264,646,671]" italics="true" pageId="20" pageNumber="1016">Goniodes</emphasis>
(
<emphasis id="B97F3406FFA0CF353E13FD67FE4BFD37" box="[292,418,646,671]" italics="true" pageId="20" pageNumber="1016">Philopedon</emphasis>
)
<emphasis id="B97F3406FFA0CF353EF7FD67FDABFD37" box="[448,578,646,671]" italics="true" pageId="20" pageNumber="1016">rectangulus</emphasis>
(Nitzsch)
</taxonomicName>
;
<treatmentCitation id="0AAACE05FFA0CF353DE1FD67FC3BFD37" author="Stephens JF" box="[726,978,646,671]" page="333" pageId="20" pageNumber="1016" year="1829">
<bibRefCitation id="EF9A95E5FFA0CF353DE1FD67FC3BFD37" author="Stephens JF" box="[726,978,646,671]" pageId="20" pageNumber="1016" refId="ref20866" refString="Stephens JF. 1829. A systematic catalogue of British insects: being an attempt to arrange all the hitherto discovered indigenous insects in accordance with their natural affinities. London (UK): Baldwin &amp; Cradock; xxxiv + 388 pp." type="book" year="1829">Stephens 1829: 333</bibRefCitation>
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[misprint for
<taxonomicName id="4C0B9397FFA0CF353F97FD49FEDDFD69" authorityName="Nitzsch" authorityYear="1818" box="[160,308,680,705]" class="Insecta" family="Philopteridae" genus="Goniodes" kingdom="Animalia" order="Psocodea" pageId="20" pageNumber="1016" phylum="Arthropoda" rank="species" species="rectangulatus">
<emphasis id="B97F3406FFA0CF353F97FD49FEDDFD69" box="[160,308,680,705]" italics="true" pageId="20" pageNumber="1016">rectangulatus</emphasis>
</taxonomicName>
].
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<taxonomicName id="4C0B9397FFA0CF353F97FD2AFDE4FD4C" ID-CoL="3GWGX" authority="Nitzsch" authorityName="Nitzsch" authorityYear="1866" box="[160,525,715,740]" class="Insecta" family="Philopteridae" genus="Goniocotes" kingdom="Animalia" order="Psocodea" pageId="20" pageNumber="1016" phylum="Arthropoda" rank="species" species="rectangulatus">
<emphasis id="B97F3406FFA0CF353F97FD2AFE5CFD4C" box="[160,437,715,740]" italics="true" pageId="20" pageNumber="1016">Goniocotes rectangulatus</emphasis>
Nitzsch
</taxonomicName>
[in Giebel], 1866: 389.
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<paragraph id="8BB4E814FFA0CF353F97FD0FFC1DFCAF" blockId="20.[160,1156,611,809]" box="[160,1012,750,775]" pageId="20" pageNumber="1016">
<treatmentCitationGroup id="AB1BCF3AFFA0CF353F97FD0FFC1DFCAF" box="[160,1012,750,775]" pageId="20" pageNumber="1016">
<treatmentCitation id="0AAACE05FFA0CF353F97FD0FFD8EFCAF" author="Giebel CG" box="[160,615,750,775]" page="185" pageId="20" pageNumber="1016" year="1874">
<taxonomicName id="4C0B9397FFA0CF353F97FD0FFD8EFCAF" ID-CoL="3GWGY" authority="Giebel, 1874: 185" authorityName="Giebel" authorityPageNumber="185" authorityYear="1874" box="[160,615,750,775]" class="Insecta" family="Philopteridae" genus="Goniocotes" kingdom="Animalia" order="Psocodea" pageId="20" pageNumber="1016" phylum="Arthropoda" rank="species" species="rectangulus">
<emphasis id="B97F3406FFA0CF353F97FD0FFE77FCAF" box="[160,414,750,775]" italics="true" pageId="20" pageNumber="1016">Goniocotes rectangulus</emphasis>
<bibRefCitation id="EF9A95E5FFA0CF353E91FD0FFD8EFCAF" author="Giebel CG" box="[422,615,750,775]" pageId="20" pageNumber="1016" refId="ref18933" refString="Giebel CG. 1874. Insecta epizoa. Die auf Saugetieren und Vogeln schmarotzenden Insecten nach Chr. L. Nitzsch ̾ s Nachlass bearbeitet. Leipzig: Otto Wigand; xiv + 308 pp., 20 plates." type="book" year="1874">Giebel, 1874: 185</bibRefCitation>
</taxonomicName>
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<emphasis id="B97F3406FFA0CF353D58FD0FFCE7FCAF" box="[623,782,750,775]" italics="true" pageId="20" pageNumber="1016">nomen novum</emphasis>
for
<taxonomicName id="4C0B9397FFA0CF353C0BFD0FFC06FCAF" ID-CoL="3GWGX" authorityName="Nitzsch" authorityYear="1866" box="[828,1007,750,775]" class="Insecta" family="Philopteridae" genus="Goniocotes" kingdom="Animalia" order="Psocodea" pageId="20" pageNumber="1016" phylum="Arthropoda" rank="species" species="rectangulatus">
<emphasis id="B97F3406FFA0CF353C0BFD0FFC06FCAF" box="[828,1007,750,775]" italics="true" pageId="20" pageNumber="1016">G. rectangulatus</emphasis>
</taxonomicName>
.
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<subSubSection id="C311BB9FFFA0CF353F97FCF1FD58FC31" pageId="20" pageNumber="1016" type="description">
<paragraph id="8BB4E814FFA0CF353F97FCF1FEA4FC81" blockId="20.[160,1156,611,809]" box="[160,333,784,809]" pageId="20" pageNumber="1016">
(
<figureCitation id="1330F491FFA0CF353F9FFCF1FEACFC81" box="[168,325,784,809]" captionStart-0="Figure 18" captionStart-1="Figure 19" captionStart-2="Figures 2022" captionStart-3="Figure 23" captionStartId-0="21.[160,225,1746,1769]" captionStartId-1="22.[160,225,1746,1769]" captionStartId-2="23.[160,235,1631,1654]" captionStartId-3="24.[160,225,1574,1597]" captionTargetBox-0="[195,1122,143,1711]" captionTargetBox-1="[216,1097,143,1709]" captionTargetBox-2="[164,1152,143,1598]" captionTargetBox-3="[163,1152,142,1528]" captionTargetId-0="figure-1@21.[192,1123,142,1714]" captionTargetId-1="figure-1@22.[215,1100,142,1714]" captionTargetId-2="figure-1@23.[163,1153,142,1599]" captionTargetId-3="figure-3@24.[163,1153,142,1543]" captionTargetPageId-0="21" captionTargetPageId-1="22" captionTargetPageId-2="23" captionTargetPageId-3="24" captionText-0="Figure 18. Goniocotes rectangulatus Nitzsch [in Giebel], 1866, male habitus, dorsal and ventral views. Paratergal plates not clearly demarcated in specimen, and illustrated approximately." captionText-1="Figure 19. Goniocotes rectangulatus Nitzsch [in Giebel], 1866, female habitus, dorsal and ventral views. Paratergal plates not clearly demarcated in specimen, and illustrated approximately." captionText-2="Figures 2022. Goniocotes rectangulatus Nitzsch [in Giebel], 1866. 20, male head, dorsal and ventral views. 21, male genitalia, dorsal view. 22, female genitalia and abdominal segments VIIIXI, ventral view." captionText-3="Figure 23. Photographs of the remaining specimens (top), the slide labels (middle), and the slide itself (bottom) of Nitzsch̍s Goniocotes rectangulatus Nitzsch [in Giebel], 1866, at the Zentralmagazin Naturwissenschaftlischer Sammlungen, Martin-Luther-Universität, Halle-Wittenberg, provided by Dr Hendrik Müller at the same institute. These specimens constitute the syntype series of G. reticulatus, and are the same as those on which von Kéler (1940) based his illustrations." figureDoi-0="http://doi.org/10.5281/zenodo.8270825" figureDoi-1="http://doi.org/10.5281/zenodo.8270827" figureDoi-2="http://doi.org/10.5281/zenodo.8270831" figureDoi-3="http://doi.org/10.5281/zenodo.8270833" httpUri-0="https://zenodo.org/record/8270825/files/figure.png" httpUri-1="https://zenodo.org/record/8270827/files/figure.png" httpUri-2="https://zenodo.org/record/8270831/files/figure.png" httpUri-3="https://zenodo.org/record/8270833/files/figure.png" pageId="20" pageNumber="1016">Figures 1823</figureCitation>
)
</paragraph>
<paragraph id="8BB4E814FFA0CF353F97FCBDFEFAFCDE" blockId="20.[160,689,860,921]" box="[160,275,860,886]" pageId="20" pageNumber="1016">
<emphasis id="B97F3406FFA0CF353F97FCBDFEFAFCDE" bold="true" box="[160,275,860,886]" italics="true" pageId="20" pageNumber="1016">
<typeStatus id="54B056B6FFA0CF353F97FCBDFF31FCDE" box="[160,216,860,886]" pageId="20" pageNumber="1016">Type</typeStatus>
host
</emphasis>
</paragraph>
<paragraph id="8BB4E814FFA0CF353F97FC61FD58FC31" blockId="20.[160,689,860,921]" box="[160,689,896,921]" pageId="20" pageNumber="1016">
<taxonomicName id="4C0B9397FFA0CF353F97FC61FE0EFC31" authority="Linnaeus, 1758" authorityName="Linnaeus" authorityYear="1758" box="[160,487,896,921]" class="Aves" family="Phasianidae" genus="Pavo" kingdom="Animalia" order="Galliformes" pageId="20" pageNumber="1016" phylum="Chordata" rank="species" species="cristatus">
<emphasis id="B97F3406FFA0CF353F97FC61FEDEFC31" box="[160,311,896,921]" italics="true" pageId="20" pageNumber="1016">Pavo cristatus</emphasis>
<bibRefCitation id="EF9A95E5FFA0CF353E08FC61FE0EFC31" author="Linnaeus CV" box="[319,487,896,921]" pageId="20" pageNumber="1016" refId="ref20038" refString="Linnaeus CV. 1758. Systema naturae per Regna Tria Naturae, Secundum Classes, Ordines, Genera, Species, cum Characteribus, Differentiis, Synonymis, Locis. 10 th ed. Stockholm (Sweden): Salvius; p. iv + 824." type="book" year="1758">Linnaeus, 1758</bibRefCitation>
</taxonomicName>
Indian peafowl.
</paragraph>
</subSubSection>
<subSubSection id="C311BB9FFFA0CF353F97FC2DFE35FBA0" pageId="20" pageNumber="1016" type="materials_examined">
<paragraph id="8BB4E814FFA0CF353F97FC2DFEDFFC4E" blockId="20.[160,476,972,1032]" box="[160,310,972,998]" pageId="20" pageNumber="1016">
<emphasis id="B97F3406FFA0CF353F97FC2DFEDFFC4E" bold="true" box="[160,310,972,998]" italics="true" pageId="20" pageNumber="1016">
<typeStatus id="54B056B6FFA0CF353F97FC2DFF31FC4E" box="[160,216,972,998]" pageId="20" pageNumber="1016">Type</typeStatus>
locality
</emphasis>
</paragraph>
<paragraph id="8BB4E814FFA0CF353F97FC0EFE35FBA0" blockId="20.[160,476,972,1032]" box="[160,476,1007,1032]" pageId="20" pageNumber="1016">
None given; likely
<collectingCountry id="F31CA884FFA0CF353E47FC0EFE30FBA0" box="[368,473,1007,1032]" name="Germany" pageId="20" pageNumber="1016">Germany</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C311BB9FFFA0CF383F97FBDDFCB8FE32" lastPageId="25" lastPageNumber="1021" pageId="20" pageNumber="1016" type="description">
<paragraph id="8BB4E814FFA0CF353F97FBDDFECFFBFE" blockId="20.[160,1156,1084,1547]" box="[160,294,1084,1110]" pageId="20" pageNumber="1016">
<emphasis id="B97F3406FFA0CF353F97FBDDFECFFBFE" bold="true" box="[160,294,1084,1110]" italics="true" pageId="20" pageNumber="1016">Description</emphasis>
</paragraph>
<paragraph id="8BB4E814FFA0CF353F97FBBFFD87F9A3" blockId="20.[160,1156,1084,1547]" pageId="20" pageNumber="1016">
<emphasis id="B97F3406FFA0CF353F97FBBFFECDFBD0" bold="true" box="[160,292,1118,1144]" italics="true" pageId="20" pageNumber="1016">Both sexes.</emphasis>
Head shape as in
<figureCitation id="1330F491FFA0CF353D35FBBEFD86FBD0" box="[514,623,1119,1144]" captionStart="Figures 1216" captionStartId="14.[160,235,1597,1620]" captionTargetBox="[163,1151,144,1564]" captionTargetId="figure-1@14.[163,1153,142,1565]" captionTargetPageId="14" captionText="Figures 1216. Goniodes meinertzhageni Clay, 1940. 12, male head, dorsal and ventral views. 13, female antenna and conus, ventral view. 14, male genitalia, dorsal view. 15, male genitalia, ventral view. 16, female genitalia and abdominal segments VIIXI, ventral view." figureDoi="http://doi.org/10.5281/zenodo.8270819" httpUri="https://zenodo.org/record/8270819/files/figure.png" pageId="20" pageNumber="1016">Figure 12</figureCitation>
; temples not flaring significantly, and without distinct corner at
<emphasis id="B97F3406FFA0CF353E5DFB60FE49FB32" box="[362,416,1153,1178]" italics="true" pageId="20" pageNumber="1016">mts1</emphasis>
. Marginal carina broad, widening near frons, with irregular inner margin and clear attendant canals to at least some preantennal setae. Preantennal nodi large, connected to marginal carina by narrow neck. Coni broad, blunt, with a tendency in some specimens to curve slightly medianly (
<figureCitation id="1330F491FFA0CF353DE6FB04FCA9FB56" box="[721,832,1253,1278]" captionStart="Figure 19" captionStartId="22.[160,225,1746,1769]" captionTargetBox="[216,1097,143,1709]" captionTargetId="figure-1@22.[215,1100,142,1714]" captionTargetPageId="22" captionText="Figure 19. Goniocotes rectangulatus Nitzsch [in Giebel], 1866, female habitus, dorsal and ventral views. Paratergal plates not clearly demarcated in specimen, and illustrated approximately." figureDoi="http://doi.org/10.5281/zenodo.8270827" httpUri="https://zenodo.org/record/8270827/files/figure.png" pageId="20" pageNumber="1016">Figure 19</figureCitation>
). Antennae sexually monomorphic. Head chaetotaxy as in
<figureCitation id="1330F491FFA0CF353D3DFAE6FD9CFA88" box="[522,629,1287,1312]" captionStart="Figures 2022" captionStartId="23.[160,235,1631,1654]" captionTargetBox="[164,1152,143,1598]" captionTargetId="figure-1@23.[163,1153,142,1599]" captionTargetPageId="23" captionText="Figures 2022. Goniocotes rectangulatus Nitzsch [in Giebel], 1866. 20, male head, dorsal and ventral views. 21, male genitalia, dorsal view. 22, female genitalia and abdominal segments VIIIXI, ventral view." figureDoi="http://doi.org/10.5281/zenodo.8270831" httpUri="https://zenodo.org/record/8270831/files/figure.png" pageId="20" pageNumber="1016">Figure 20</figureCitation>
;
<emphasis id="B97F3406FFA0CF353DB7FAE6FD5BFA88" box="[640,690,1287,1312]" italics="true" pageId="20" pageNumber="1016">
s
<figureCitation id="1330F491FFA0CF353DBAFAE6FD5BFA88" box="[653,690,1287,1312]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figures 37" captionStartId-0="4.[160,225,1746,1769]" captionStartId-1="7.[160,225,1746,1769]" captionStartId-2="8.[160,235,1571,1594]" captionTargetBox-0="[168,1146,142,1711]" captionTargetBox-1="[227,1087,143,1713]" captionTargetBox-2="[169,1145,149,1533]" captionTargetId-0="figure-1@4.[167,1148,142,1714]" captionTargetId-1="figure-1@7.[226,1090,142,1714]" captionTargetId-2="figure-1@8.[163,1153,142,1539]" captionTargetPageId-0="4" captionTargetPageId-1="7" captionTargetPageId-2="8" captionText-0="Figure 1. Goniodes pavonis (Linnaeus, 1758), male habitus, dorsal and ventral views." captionText-1="Figure 2. Goniodes pavonis (Linnaeus, 1758), female habitus, dorsal and ventral views." captionText-2="Figures 37. Goniodes pavonis (Linnaeus, 1758). 3, male head, dorsal and ventral views. 4, female antenna and conus, ventral view. 5, male genitalia, dorsal view. 6, male genitalia, ventral view. 7, female genitalia and abdominal segments VIIXI, ventral view." figureDoi-0="http://doi.org/10.5281/zenodo.8270803" figureDoi-1="http://doi.org/10.5281/zenodo.8270807" figureDoi-2="http://doi.org/10.5281/zenodo.8270809" httpUri-0="https://zenodo.org/record/8270803/files/figure.png" httpUri-1="https://zenodo.org/record/8270807/files/figure.png" httpUri-2="https://zenodo.org/record/8270809/files/figure.png" pageId="20" pageNumber="1016">16</figureCitation>
</emphasis>
present in male, but only
<emphasis id="B97F3406FFA0CF353CEEFAE6FBE2FA88" box="[985,1035,1287,1312]" italics="true" pageId="20" pageNumber="1016">s13</emphasis>
present in female;
<emphasis id="B97F3406FFA0CF353FCEFAC8FEF8FAEA" box="[249,273,1321,1346]" italics="true" pageId="20" pageNumber="1016">os</emphasis>
sexually monomorphic;
<emphasis id="B97F3406FFA0CF353D11FAC8FDB2FAEA" box="[550,603,1321,1346]" italics="true" pageId="20" pageNumber="1016">mts5</emphasis>
microseta in both sexes. Thoracic segments as in
<figureCitation id="1330F491FFA0CF353F97FAABFEA8FACB" box="[160,321,1354,1379]" captionStart-0="Figure 18" captionStart-1="Figure 19" captionStartId-0="21.[160,225,1746,1769]" captionStartId-1="22.[160,225,1746,1769]" captionTargetBox-0="[195,1122,143,1711]" captionTargetBox-1="[216,1097,143,1709]" captionTargetId-0="figure-1@21.[192,1123,142,1714]" captionTargetId-1="figure-1@22.[215,1100,142,1714]" captionTargetPageId-0="21" captionTargetPageId-1="22" captionText-0="Figure 18. Goniocotes rectangulatus Nitzsch [in Giebel], 1866, male habitus, dorsal and ventral views. Paratergal plates not clearly demarcated in specimen, and illustrated approximately." captionText-1="Figure 19. Goniocotes rectangulatus Nitzsch [in Giebel], 1866, female habitus, dorsal and ventral views. Paratergal plates not clearly demarcated in specimen, and illustrated approximately." figureDoi-0="http://doi.org/10.5281/zenodo.8270825" figureDoi-1="http://doi.org/10.5281/zenodo.8270827" httpUri-0="https://zenodo.org/record/8270825/files/figure.png" httpUri-1="https://zenodo.org/record/8270827/files/figure.png" pageId="20" pageNumber="1016">Figures 1819</figureCitation>
. Proepimera fused medianly, mesofurcal pit sublateral. One sternal seta on each side of both meso- and metathorax, no thoracic sternal plates. Pterothoracic setae sparse;
<emphasis id="B97F3406FFA0CF353E01FA6FFEB4FA0F" box="[310,349,1422,1447]" italics="true" pageId="20" pageNumber="1016">lpts</emphasis>
and
<emphasis id="B97F3406FFA0CF353EA3FA6FFE52FA0F" box="[404,443,1422,1447]" italics="true" pageId="20" pageNumber="1016">ipts</emphasis>
widely separated,
<emphasis id="B97F3406FFA0CF353DB9FA6FFD2FFA0F" box="[654,710,1422,1447]" italics="true" pageId="20" pageNumber="1016">smns</emphasis>
and
<emphasis id="B97F3406FFA0CF353DCAFA6FFCDDFA0F" box="[765,820,1422,1447]" italics="true" pageId="20" pageNumber="1016">mpts</emphasis>
absent. Abdominal segments and chaetotaxy as in
<figureCitation id="1330F491FFA0CF353EA7FA4EFDC7FA60" box="[400,558,1455,1480]" captionStart-0="Figure 18" captionStart-1="Figure 19" captionStartId-0="21.[160,225,1746,1769]" captionStartId-1="22.[160,225,1746,1769]" captionTargetBox-0="[195,1122,143,1711]" captionTargetBox-1="[216,1097,143,1709]" captionTargetId-0="figure-1@21.[192,1123,142,1714]" captionTargetId-1="figure-1@22.[215,1100,142,1714]" captionTargetPageId-0="21" captionTargetPageId-1="22" captionText-0="Figure 18. Goniocotes rectangulatus Nitzsch [in Giebel], 1866, male habitus, dorsal and ventral views. Paratergal plates not clearly demarcated in specimen, and illustrated approximately." captionText-1="Figure 19. Goniocotes rectangulatus Nitzsch [in Giebel], 1866, female habitus, dorsal and ventral views. Paratergal plates not clearly demarcated in specimen, and illustrated approximately." figureDoi-0="http://doi.org/10.5281/zenodo.8270825" figureDoi-1="http://doi.org/10.5281/zenodo.8270827" httpUri-0="https://zenodo.org/record/8270825/files/figure.png" httpUri-1="https://zenodo.org/record/8270827/files/figure.png" pageId="20" pageNumber="1016">Figures 1819</figureCitation>
; tergopleurites sexually dimorphic. Paratergal plates poorly demarcated in both sexes, and illustrated approximately. Sternal plates absent in both sexes. Measurements as in
<tableCitation id="C689DDAFFFA0CF353D22FA13FD80F9A3" box="[533,617,1522,1547]" captionStart="Table 1" captionStartId="10.[160,216,142,165]" captionTargetPageId="10" captionText="Table 1. Measurements (in millimetres) of the species treated here. Abbreviations: TL = total length (along midline); HL = head length (along midline); HW = head width (at temples); PRW = prothoracic width; PTW = pterothoracic width; AW = abdominal width (at segment V, for consistency, AW is measured at segment V even in Pavoniocotes, but the abdomen is wider at the posterior end of segment IV in some specimens of this genus). One specimen of Pavoniocotes parviceps (Piaget, 1880) which appears to be either a tenereal male or an exuvium is not measured." httpUri="http://table.plazi.org/id/DF74B89CFFBECF2B3F97FF6FFC66FE90" pageId="20" pageNumber="1016" tableUuid="DF74B89CFFBECF2B3F97FF6FFC66FE90">Table 1</tableCitation>
.
</paragraph>
<paragraph id="8BB4E814FFA0CF383F97F9DFFB98FF62" blockId="20.[160,1156,1598,1728]" lastBlockId="25.[160,1156,142,202]" lastPageId="25" lastPageNumber="1021" pageId="20" pageNumber="1016">
<emphasis id="B97F3406FFA0CF353F97F9DFFF08F9F0" bold="true" box="[160,225,1598,1624]" italics="true" pageId="20" pageNumber="1016">Male.</emphasis>
Tergopleurites IIVIII reduced, widely separated medianly (
<figureCitation id="1330F491FFA0CF353CBFF9DEFC1BF9F0" box="[904,1010,1599,1624]" captionStart="Figure 18" captionStartId="21.[160,225,1746,1769]" captionTargetBox="[195,1122,143,1711]" captionTargetId="figure-1@21.[192,1123,142,1714]" captionTargetPageId="21" captionText="Figure 18. Goniocotes rectangulatus Nitzsch [in Giebel], 1866, male habitus, dorsal and ventral views. Paratergal plates not clearly demarcated in specimen, and illustrated approximately." figureDoi="http://doi.org/10.5281/zenodo.8270825" httpUri="https://zenodo.org/record/8270825/files/figure.png" pageId="20" pageNumber="1016">Figure 18</figureCitation>
). Abdominal chaetotaxy sparse, without
<emphasis id="B97F3406FFA0CF353EE2F983FE1CF9D3" box="[469,501,1634,1659]" italics="true" pageId="20" pageNumber="1016">tps</emphasis>
or setal rows on all segments. Tergopleurite IX elongated, with median hook-like extension, not overlapping central tergopleurite X. Subgenital plate not clearly demarcated in examined specimen, and not illustrated; presumably similar in shape and size to those of other species in the genus. Male genitalia simple, solenoid, with faint wing-like extensions on distal ends which may be part of genitalia.
</paragraph>
<caption id="DF74B89CFFA1CF343F97F933FC0CF8AE" ID-DOI="http://doi.org/10.5281/zenodo.8270825" ID-Zenodo-Dep="8270825" httpUri="https://zenodo.org/record/8270825/files/figure.png" pageId="21" pageNumber="1017" startId="21.[160,225,1746,1769]" targetBox="[195,1122,143,1711]" targetPageId="21" targetType="figure">
<paragraph id="8BB4E814FFA1CF343F97F933FC0CF8AE" blockId="21.[160,1157,1746,1798]" pageId="21" pageNumber="1017">
<emphasis id="B97F3406FFA1CF343F97F933FEEEF941" bold="true" box="[160,263,1746,1769]" pageId="21" pageNumber="1017">Figure 18.</emphasis>
<taxonomicName id="4C0B9397FFA1CF343E39F933FDA7F941" authority="Nitzsch" authorityName="Nitzsch" authorityYear="1866" box="[270,590,1746,1769]" class="Insecta" family="Philopteridae" genus="Goniocotes" kingdom="Animalia" order="Psocodea" pageId="21" pageNumber="1017" phylum="Arthropoda" rank="species" species="rectangulatus">
<emphasis id="B97F3406FFA1CF343E39F933FDE9F941" box="[270,512,1746,1769]" italics="true" pageId="21" pageNumber="1017">Goniocotes rectangulatus</emphasis>
Nitzsch
</taxonomicName>
[in Giebel], 1866, male habitus, dorsal and ventral views. Paratergal plates not clearly demarcated in specimen, and illustrated approximately.
</paragraph>
</caption>
<caption id="DF74B89CFFA2CF373F97F933FC0CF8AE" ID-DOI="http://doi.org/10.5281/zenodo.8270827" ID-Zenodo-Dep="8270827" httpUri="https://zenodo.org/record/8270827/files/figure.png" pageId="22" pageNumber="1018" startId="22.[160,225,1746,1769]" targetBox="[216,1097,143,1709]" targetPageId="22" targetType="figure">
<paragraph id="8BB4E814FFA2CF373F97F933FC0CF8AE" blockId="22.[160,1157,1746,1798]" pageId="22" pageNumber="1018">
<emphasis id="B97F3406FFA2CF373F97F933FEEFF941" bold="true" box="[160,262,1746,1769]" pageId="22" pageNumber="1018">Figure 19.</emphasis>
<taxonomicName id="4C0B9397FFA2CF373E3CF933FDA1F941" authority="Nitzsch" authorityName="Nitzsch" authorityYear="1866" box="[267,584,1746,1769]" class="Insecta" family="Philopteridae" genus="Goniocotes" kingdom="Animalia" order="Psocodea" pageId="22" pageNumber="1018" phylum="Arthropoda" rank="species" species="rectangulatus">
<emphasis id="B97F3406FFA2CF373E3CF933FE12F941" box="[267,507,1746,1769]" italics="true" pageId="22" pageNumber="1018">Goniocotes rectangulatus</emphasis>
Nitzsch
</taxonomicName>
[in Giebel], 1866, female habitus, dorsal and ventral views. Paratergal plates not clearly demarcated in specimen, and illustrated approximately.
</paragraph>
</caption>
<caption id="DF74B89CFFA3CF363F97F9BEFF3DF919" ID-DOI="http://doi.org/10.5281/zenodo.8270831" ID-Zenodo-Dep="8270831" httpUri="https://zenodo.org/record/8270831/files/figure.png" pageId="23" pageNumber="1019" startId="23.[160,235,1631,1654]" targetBox="[164,1152,143,1598]" targetPageId="23" targetType="figure">
<paragraph id="8BB4E814FFA3CF363F97F9BEFF3DF919" blockId="23.[160,1155,1631,1713]" pageId="23" pageNumber="1019">
<emphasis id="B97F3406FFA3CF363F97F9BEFEDEF9DE" bold="true" box="[160,311,1631,1654]" pageId="23" pageNumber="1019">Figures 2022.</emphasis>
<taxonomicName id="4C0B9397FFA3CF363E09F9BEFD96F9DE" authority="Nitzsch" authorityName="Nitzsch" authorityYear="1866" box="[318,639,1631,1654]" class="Insecta" family="Philopteridae" genus="Goniocotes" kingdom="Animalia" order="Psocodea" pageId="23" pageNumber="1019" phylum="Arthropoda" rank="species" species="rectangulatus">
<emphasis id="B97F3406FFA3CF363E09F9BEFDD9F9DE" box="[318,560,1631,1654]" italics="true" pageId="23" pageNumber="1019">Goniocotes rectangulatus</emphasis>
Nitzsch
</taxonomicName>
[in Giebel], 1866.
<emphasis id="B97F3406FFA3CF363C02F9BEFCA6F9DE" bold="true" box="[821,847,1631,1654]" pageId="23" pageNumber="1019">20</emphasis>
, male head, dorsal and ventral views.
<emphasis id="B97F3406FFA3CF363FD3F99DFF17F93B" bold="true" box="[228,254,1660,1683]" pageId="23" pageNumber="1019">21</emphasis>
, male genitalia, dorsal view.
<emphasis id="B97F3406FFA3CF363D15F99DFDD5F93B" bold="true" box="[546,572,1660,1683]" pageId="23" pageNumber="1019">22</emphasis>
, female genitalia and abdominal segments VIIIXI, ventral view.
</paragraph>
</caption>
<caption id="DF74B89CFFACCF393F97F9C7FBC9F91B" ID-DOI="http://doi.org/10.5281/zenodo.8270833" ID-Zenodo-Dep="8270833" httpUri="https://zenodo.org/record/8270833/files/figure.png" pageId="24" pageNumber="1020" startId="24.[160,225,1574,1597]" targetBox="[163,1152,142,1528]" targetPageId="24" targetType="figure">
<paragraph id="8BB4E814FFACCF393F97F9C7FBC9F91B" blockId="24.[160,1156,1574,1715]" pageId="24" pageNumber="1020">
<emphasis id="B97F3406FFACCF393F97F9C7FEEFF995" bold="true" box="[160,262,1574,1597]" pageId="24" pageNumber="1020">Figure 23.</emphasis>
Photographs of the remaining specimens (top), the slide labels (middle), and the slide itself (bottom) of Nitzsch̍s
<taxonomicName id="4C0B9397FFACCF393EB2F9A5FD3DF9F3" authority="Nitzsch" authorityName="Nitzsch" authorityYear="1866" box="[389,724,1604,1627]" class="Insecta" family="Philopteridae" genus="Goniocotes" kingdom="Animalia" order="Psocodea" pageId="24" pageNumber="1020" phylum="Arthropoda" rank="species" species="rectangulatus">
<emphasis id="B97F3406FFACCF393EB2F9A5FD68F9F3" box="[389,641,1604,1627]" italics="true" pageId="24" pageNumber="1020">Goniocotes rectangulatus</emphasis>
Nitzsch
</taxonomicName>
[in Giebel], 1866, at the Zentralmagazin Naturwissenschaftlischer Sammlungen, Martin-Luther-Universität, Halle-Wittenberg, provided by Dr Hendrik Müller at the same institute. These specimens constitute the syntype series of
<taxonomicName id="4C0B9397FFACCF393F97F97DFEF7F91B" authorityName="von Keler" authorityYear="1939" box="[160,286,1692,1715]" class="Insecta" family="Philopteridae" genus="Goniocotes" kingdom="Animalia" order="Psocodea" pageId="24" pageNumber="1020" phylum="Arthropoda" rank="species" species="reticulatus">
<emphasis id="B97F3406FFACCF393F97F97DFEF7F91B" box="[160,286,1692,1715]" italics="true" pageId="24" pageNumber="1020">G. reticulatus</emphasis>
</taxonomicName>
, and are the same as those on which
<bibRefCitation id="EF9A95E5FFACCF393DADF97DFCD5F91B" author="von Keler S" box="[666,828,1692,1715]" pageId="24" pageNumber="1020" pagination="1 - 354" refId="ref21322" refString="von Keler S. 1940. Baustoffe zu einer Monographie der mallophagen. II. Teil: Uberfamilie der Nirmoidea. Nova Acta Leopoldiana Abhandlungen der Kaiserlich Leopoldinisch-Carolinisch Deutschen Akademie der Naturforscher Neue Folge. 8: 1 - 354 + 4 plates." type="book chapter" year="1940">von Kéler (1940)</bibRefCitation>
based his illustrations.
</paragraph>
</caption>
<paragraph id="8BB4E814FFADCF383F97FF17FCB8FE32" blockId="25.[160,1155,246,410]" pageId="25" pageNumber="1021">
<emphasis id="B97F3406FFADCF383F97FF17FF14FEB8" bold="true" box="[160,253,246,272]" italics="true" pageId="25" pageNumber="1021">Female.</emphasis>
Tergopleurites IIVIII elongated (
<figureCitation id="1330F491FFADCF383D40FF17FD36FEA7" box="[631,735,246,271]" captionStart="Figure 19" captionStartId="22.[160,225,1746,1769]" captionTargetBox="[216,1097,143,1709]" captionTargetId="figure-1@22.[215,1100,142,1714]" captionTargetPageId="22" captionText="Figure 19. Goniocotes rectangulatus Nitzsch [in Giebel], 1866, female habitus, dorsal and ventral views. Paratergal plates not clearly demarcated in specimen, and illustrated approximately." figureDoi="http://doi.org/10.5281/zenodo.8270827" httpUri="https://zenodo.org/record/8270827/files/figure.png" pageId="25" pageNumber="1021">Figure 19</figureCitation>
). Abdominal chaetotaxy sparse, without
<emphasis id="B97F3406FFADCF383FE4FEF8FF1DFE9A" box="[211,244,281,306]" italics="true" pageId="25" pageNumber="1021">tps</emphasis>
or setal rows on all segments. Tergopleurites IXXI fused, proximal margin indented medianly. Subgenital plate poorly sclerotised, seemingly as broad, medianly continuous sclerite at vulval margin (
<figureCitation id="1330F491FFADCF383D76FEBFFD42FEDF" box="[577,683,350,375]" captionStart="Figures 2022" captionStartId="23.[160,235,1631,1654]" captionTargetBox="[164,1152,143,1598]" captionTargetId="figure-1@23.[163,1153,142,1599]" captionTargetPageId="23" captionText="Figures 2022. Goniocotes rectangulatus Nitzsch [in Giebel], 1866. 20, male head, dorsal and ventral views. 21, male genitalia, dorsal view. 22, female genitalia and abdominal segments VIIIXI, ventral view." figureDoi="http://doi.org/10.5281/zenodo.8270831" httpUri="https://zenodo.org/record/8270831/files/figure.png" pageId="25" pageNumber="1021">Figure 22</figureCitation>
). Vulval margin rounded, with chaetotaxy as in
<figureCitation id="1330F491FFADCF383FECFE60FEACFE32" box="[219,325,385,410]" captionStart="Figures 2022" captionStartId="23.[160,235,1631,1654]" captionTargetBox="[164,1152,143,1598]" captionTargetId="figure-1@23.[163,1153,142,1599]" captionTargetPageId="23" captionText="Figures 2022. Goniocotes rectangulatus Nitzsch [in Giebel], 1866. 20, male head, dorsal and ventral views. 21, male genitalia, dorsal view. 22, female genitalia and abdominal segments VIIIXI, ventral view." figureDoi="http://doi.org/10.5281/zenodo.8270831" httpUri="https://zenodo.org/record/8270831/files/figure.png" pageId="25" pageNumber="1021">Figure 22</figureCitation>
. Subvulval plates absent or poorly sclerotised.
</paragraph>
</subSubSection>
<subSubSection id="C311BB9FFFADCF383F97FE27FD14FD8D" pageId="25" pageNumber="1021" type="materials_examined">
<paragraph id="8BB4E814FFADCF383F97FE27FE96FE48" blockId="25.[160,1156,454,549]" box="[160,383,454,480]" pageId="25" pageNumber="1021">
<emphasis id="B97F3406FFADCF383F97FE27FE96FE48" bold="true" box="[160,383,454,480]" italics="true" pageId="25" pageNumber="1021">Material examined</emphasis>
</paragraph>
<paragraph id="8BB4E814FFADCF383F97FE0BFD14FD8D" blockId="25.[160,1156,454,549]" pageId="25" pageNumber="1021">
Non-types (erroneously marked ̍Neoparatypes̾).
<materialsCitation id="3B63E249FFADCF383C37FE0BFD10FD8D" ID-GBIF-Occurrence="4398940313" collectingDate="1975-05-03" collectionCode="NHML" country="Poland" location="Wroclaw Zoo" pageId="25" pageNumber="1021" specimenCode="BM1993-8, NHMUK010575871-2" specimenCount="1" specimenCount-female="1">
1♂,
<specimenCount id="9D0D239DFFADCF383C0DFE0BFCB5FDA9" box="[826,860,490,515]" pageId="25" pageNumber="1021" type="female">1♀</specimenCount>
,
<location id="8ED4BECFFFADCF383C43FE0BFBF0FDAB" LSID="urn:lsid:plazi:treatment:03A25902FFA0CF3B3E55FDCFFE85FA02:8ED4BECFFFADCF383C43FE0BFBF0FDAB" box="[884,1049,490,515]" country="Poland" name="Wroclaw Zoo" pageId="25" pageNumber="1021">Wroclaw Zoo</location>
,
<collectingCountry id="F31CA884FFADCF383B19FE0BFB68FDAB" box="[1070,1153,490,515]" name="Poland" pageId="25" pageNumber="1021">Poland</collectingCountry>
,
<date id="FFB5CED4FFADCF383F97FDEDFECCFD8D" box="[160,293,524,549]" pageId="25" pageNumber="1021" value="1975-05-03">
<collectingDate id="EFF1373CFFADCF383F97FDEDFECCFD8D" box="[160,293,524,549]" pageId="25" pageNumber="1021" value="1975-05-03">3 May 1975</collectingDate>
</date>
,
<specimenCode id="DBAD406FFFADCF383E18FDEDFE4BFD8D" box="[303,418,524,549]" collectionCode="BM" country="United Kingdom" name="Bristol Museum" pageId="25" pageNumber="1021">BM1993-8</specimenCode>
,
<specimenCode id="DBAD406FFFADCF383E9AFDEDFD72FD8D" box="[429,667,524,549]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="25" pageNumber="1021" type="Museum">NHMUK0105758712</specimenCode>
(
<collectionCode id="ED1A70D1FFADCF383D9FFDEDFD1AFD8D" box="[680,755,524,549]" country="Libya" httpUri="http://grbio.org/cool/49nc-28y0" name="Natural History Museum, Tripoli" pageId="25" pageNumber="1021">NHML</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C311BB9FFFADCF3B3F97FDB0FE85FA02" lastPageId="26" lastPageNumber="1022" pageId="25" pageNumber="1021" type="discussion">
<paragraph id="8BB4E814FFADCF383F97FDB0FEEFFDC3" blockId="25.[160,1156,593,1728]" box="[160,262,593,619]" pageId="25" pageNumber="1021">
<emphasis id="B97F3406FFADCF383F97FDB0FEEFFDC3" bold="true" box="[160,262,593,619]" italics="true" pageId="25" pageNumber="1021">Remarks</emphasis>
</paragraph>
<paragraph id="8BB4E814FFADCF383F97FD94FEF6FC28" blockId="25.[160,1156,593,1728]" pageId="25" pageNumber="1021">
<taxonomicName id="4C0B9397FFADCF383F97FD94FE5AFD26" authorityName="Nitzsch" authorityYear="1866" box="[160,435,629,654]" class="Insecta" family="Philopteridae" genus="Goniocotes" kingdom="Animalia" order="Psocodea" pageId="25" pageNumber="1021" phylum="Arthropoda" rank="species" species="rectangulatus">
<emphasis id="B97F3406FFADCF383F97FD94FE5AFD26" box="[160,435,629,654]" italics="true" pageId="25" pageNumber="1021">Goniocotes rectangulatus</emphasis>
</taxonomicName>
has been reported rarely but regularly over the last 100 years (eg Roman-Bolelli 1947;
<bibRefCitation id="EF9A95E5FFADCF383EBBFD76FDFAFD18" author="Rekasi J" box="[396,531,663,688]" pageId="25" pageNumber="1021" pagination="71 - 93" refId="ref20648" refString="Rekasi J. 1993. Bird lice (Mallophaga) parasiting the birds of Hungary. Aquila. 100: 71 - 93." type="journal article" year="1993">Rékási 1993</bibRefCitation>
), but no description or illustration of this species has been published since
<bibRefCitation id="EF9A95E5FFADCF383EA4FD5BFDA0FD7B" author="von Keler S" box="[403,585,698,723]" pageId="25" pageNumber="1021" pagination="1 - 354" refId="ref21322" refString="von Keler S. 1940. Baustoffe zu einer Monographie der mallophagen. II. Teil: Uberfamilie der Nirmoidea. Nova Acta Leopoldiana Abhandlungen der Kaiserlich Leopoldinisch-Carolinisch Deutschen Akademie der Naturforscher Neue Folge. 8: 1 - 354 + 4 plates." type="book chapter" year="1940">von Kéler (1940)</bibRefCitation>
. As
<bibRefCitation id="EF9A95E5FFADCF383D43FD5BFCC0FD7B" author="von Keler S" box="[628,809,698,723]" pageId="25" pageNumber="1021" pagination="1 - 354" refId="ref21322" refString="von Keler S. 1940. Baustoffe zu einer Monographie der mallophagen. II. Teil: Uberfamilie der Nirmoidea. Nova Acta Leopoldiana Abhandlungen der Kaiserlich Leopoldinisch-Carolinisch Deutschen Akademie der Naturforscher Neue Folge. 8: 1 - 354 + 4 plates." type="book chapter" year="1940">von Kéler (1940)</bibRefCitation>
only illustrated the female, it is possible that some published records refer to either of the species of
<taxonomicName id="4C0B9397FFADCF383CC5FD3DFB6DFD5D" authorityName="Gustafsson &amp; Grossi &amp; Ren &amp; Zou" authorityYear="2023" box="[1010,1156,732,757]" class="Insecta" family="Goniodidae" genus="Pavoniocotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Psocodea" pageId="25" pageNumber="1021" phylum="Arthropoda" rank="genus">
<emphasis id="B97F3406FFADCF383CC5FD3DFB6DFD5D" box="[1010,1156,732,757]" italics="true" pageId="25" pageNumber="1021">Pavoniocotes</emphasis>
</taxonomicName>
described above. As discussed in more detail below, Clay confused
<taxonomicName id="4C0B9397FFADCF383CA9FD1EFBB8FCB0" authorityName="Nitzsch" authorityYear="1866" box="[926,1105,767,792]" class="Insecta" family="Philopteridae" genus="Goniocotes" kingdom="Animalia" order="Psocodea" pageId="25" pageNumber="1021" phylum="Arthropoda" rank="species" species="rectangulatus">
<emphasis id="B97F3406FFADCF383CA9FD1EFBB8FCB0" box="[926,1105,767,792]" italics="true" pageId="25" pageNumber="1021">G. rectangulatus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0B9397FFADCF383F97FCC3FEE4FC93" authorityName="Lakshminarayana and Emerson" authorityYear="1971" box="[160,269,802,827]" class="Insecta" family="Goniodidae" genus="Pavoniocotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Psocodea" pageId="25" pageNumber="1021" phylum="Arthropoda" rank="species" species="mayuri">
<emphasis id="B97F3406FFADCF383F97FCC3FEE4FC93" box="[160,269,802,827]" italics="true" pageId="25" pageNumber="1021">G. mayuri</emphasis>
</taxonomicName>
, and this confusion may have been more widespread. However, the lack of illustrations or photos in most reports preclude an assessment of whether these reports are correct.
</paragraph>
<paragraph id="8BB4E814FFADCF383FF7FC6BFCCEFA88" blockId="25.[160,1156,593,1728]" pageId="25" pageNumber="1021">
The identity of
<taxonomicName id="4C0B9397FFADCF383E58FC6BFDCAFC0B" authorityName="Nitzsch" authorityYear="1866" box="[367,547,906,931]" class="Insecta" family="Philopteridae" genus="Goniocotes" kingdom="Animalia" order="Psocodea" pageId="25" pageNumber="1021" phylum="Arthropoda" rank="species" species="rectangulatus">
<emphasis id="B97F3406FFADCF383E58FC6BFDCAFC0B" box="[367,547,906,931]" italics="true" pageId="25" pageNumber="1021">G. rectangulatus</emphasis>
</taxonomicName>
is not straightforward.
<bibRefCitation id="EF9A95E5FFADCF383C18FC6BFC26FC0B" author="Nitzsch CL" box="[815,975,906,931]" pageId="25" pageNumber="1021" pagination="261 - 316" refId="ref20470" refString="Nitzsch CL. 1818. Die Familien und Gattungen der Thierinsekten (insecta epizoica); als Prodromus einer Naturgeschechte derselben. Magazin der Entomologie. 3: 261 - 316." type="journal article" year="1818">Nitzsch (1818)</bibRefCitation>
and
<bibRefCitation id="EF9A95E5FFADCF383B3DFC6BFF0FFC6E" author="Burmeister H" pageId="25" pageNumber="1021" pagination="418 - 443" refId="ref17783" refString="Burmeister H. 1838. Mallophaga Nitzsch. In: Handbuch der Entomologie. Zweite Abteilung. Besondere Entomologie. Zweite Abteilunf. Lauskerfe. Gymnognatha. (Zweiter Halfte; vulgo Neuroptera), Vol. 2. Theod. Chr. Fried Enslin, Berlin; p. 418 - 443." type="book chapter" year="1838">Burmeister (1838)</bibRefCitation>
only mentioned the name, without making any attempt to describe it. The first description is therefore in
<bibRefCitation id="EF9A95E5FFADCF383EF8FC2EFD8EFC40" author="Giebel CG" box="[463,615,975,1000]" pageId="25" pageNumber="1021" pagination="353 - 397" refId="ref18899" refString="Giebel CG. 1866. Die im zoologischen Museum der Universitat Halle aufgestellten Epizoen nebst Beobachtungen uber dieselben. Zeitschrift fur die Gesammten Naturwissenschaften (Halle). 28: 353 - 397." type="journal article" year="1866">Giebel (1866)</bibRefCitation>
, who published Nitzsch̾s Latin manuscript; the author is thus Nitzsch.
<bibRefCitation id="EF9A95E5FFADCF383E94FC13FDD0FBA3" author="Giebel CG" box="[419,569,1010,1035]" pageId="25" pageNumber="1021" refId="ref18933" refString="Giebel CG. 1874. Insecta epizoa. Die auf Saugetieren und Vogeln schmarotzenden Insecten nach Chr. L. Nitzsch ̾ s Nachlass bearbeitet. Leipzig: Otto Wigand; xiv + 308 pp., 20 plates." type="book" year="1874">Giebel (1874)</bibRefCitation>
published a German description of the species, but the combination of characters given in these two descriptions do not fit well with either
<taxonomicName id="4C0B9397FFADCF383F97FBD6FEF1FBF8" authorityName="Burmeister" authorityYear="1838" box="[160,280,1079,1104]" class="Insecta" family="Philopteridae" genus="Goniocotes" kingdom="Animalia" order="Psocodea" pageId="25" pageNumber="1021" phylum="Arthropoda" rank="genus">
<emphasis id="B97F3406FFADCF383F97FBD6FEF1FBF8" box="[160,280,1079,1104]" italics="true" pageId="25" pageNumber="1021">Goniocotes</emphasis>
</taxonomicName>
found on peafowls or the group here described as
<taxonomicName id="4C0B9397FFADCF383C49FBD6FBE4FBF8" authority="Gustafsson &amp; Grossi &amp; Ren &amp; Zou, 2023" authorityName="Gustafsson &amp; Grossi &amp; Ren &amp; Zou" authorityYear="2023" box="[894,1037,1079,1104]" class="Insecta" family="Goniodidae" genus="Pavoniocotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Psocodea" pageId="25" pageNumber="1021" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B97F3406FFADCF383C49FBD6FBE4FBF8" box="[894,1037,1079,1104]" italics="true" pageId="25" pageNumber="1021">Pavoniocotes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24C897DFFADCF383B2FFBD6FB6DFBF8" box="[1048,1156,1079,1104]" pageId="25" pageNumber="1021" rank="genus">gen. nov.</taxonomicNameLabel>
Nitzsch [in Giebel] (1866) described the temples as ̍[
<emphasis id="B97F3406FFADCF383DCBFBBBFCE3FBDB" box="[764,778,1114,1139]" italics="true" pageId="25" pageNumber="1021">a</emphasis>
]
<emphasis id="B97F3406FFADCF383C26FBBBFCB8FBDB" box="[785,849,1114,1139]" italics="true" pageId="25" pageNumber="1021">ngulis</emphasis>
[...]
<emphasis id="B97F3406FFADCF383CB9FBBBFBF1FBDB" box="[910,1048,1114,1139]" italics="true" pageId="25" pageNumber="1021">exactis rectis</emphasis>
̾ [= exact right angles], and
<bibRefCitation id="EF9A95E5FFADCF383E51FB9CFE13FB3E" author="Giebel CG" box="[358,506,1149,1174]" pageId="25" pageNumber="1021" refId="ref18933" refString="Giebel CG. 1874. Insecta epizoa. Die auf Saugetieren und Vogeln schmarotzenden Insecten nach Chr. L. Nitzsch ̾ s Nachlass bearbeitet. Leipzig: Otto Wigand; xiv + 308 pp., 20 plates." type="book" year="1874">Giebel (1874)</bibRefCitation>
wrote that the temples are ̍rechtwinklig, aber nicht scharf̾ [= right angles, but not sharp]. This fits with males of
<taxonomicName id="4C0B9397FFADCF383C37FB7EFC14FB10" authorityName="Linnaeus" authorityYear="1758" box="[768,1021,1183,1208]" class="Insecta" family="Goniodidae" genus="Pavoniocotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Psocodea" pageId="25" pageNumber="1021" phylum="Arthropoda" rank="species" species="parviceps">
<emphasis id="B97F3406FFADCF383C37FB7EFC14FB10" box="[768,1021,1183,1208]" italics="true" pageId="25" pageNumber="1021">Pavoniocotes parviceps</emphasis>
</taxonomicName>
(
<figureCitation id="1330F491FFADCF383B3BFB7EFB9FFB10" box="[1036,1142,1183,1208]" captionStart="Figures 2630" captionStartId="30.[160,235,1580,1603]" captionTargetBox="[165,1150,143,1545]" captionTargetId="figure-1@30.[163,1153,142,1548]" captionTargetPageId="30" captionText="Figures 2630. Pavoniocotes parviceps (Piaget, 1880) ex Pavo cristatus Linnaeus, 1758. 26, male head, dorsal and ventral views. 27, female antenna and conus, ventral view. 28, male genitalia, dorsal view. 29, male genitalia, ventral view. 30, female genitalia and abdominal segments VIIXI, ventral view." figureDoi="http://doi.org/10.5281/zenodo.8270841" httpUri="https://zenodo.org/record/8270841/files/figure.png" pageId="25" pageNumber="1021">Figure 26</figureCitation>
), but not with
<taxonomicName id="4C0B9397FFADCF383E03FB23FE0EFB73" authorityName="Nitzsch" authorityYear="1866" box="[308,487,1218,1243]" class="Insecta" family="Philopteridae" genus="Goniocotes" kingdom="Animalia" order="Psocodea" pageId="25" pageNumber="1021" phylum="Arthropoda" rank="species" species="rectangulatus">
<emphasis id="B97F3406FFADCF383E03FB23FE0EFB73" box="[308,487,1218,1243]" italics="true" pageId="25" pageNumber="1021">G. rectangulatus</emphasis>
</taxonomicName>
(
<figureCitation id="1330F491FFADCF383EC2FB23FDB7FB73" box="[501,606,1218,1243]" captionStart="Figures 2022" captionStartId="23.[160,235,1631,1654]" captionTargetBox="[164,1152,143,1598]" captionTargetId="figure-1@23.[163,1153,142,1599]" captionTargetPageId="23" captionText="Figures 2022. Goniocotes rectangulatus Nitzsch [in Giebel], 1866. 20, male head, dorsal and ventral views. 21, male genitalia, dorsal view. 22, female genitalia and abdominal segments VIIIXI, ventral view." figureDoi="http://doi.org/10.5281/zenodo.8270831" httpUri="https://zenodo.org/record/8270831/files/figure.png" pageId="25" pageNumber="1021">Figure 20</figureCitation>
). However,
<bibRefCitation id="EF9A95E5FFADCF383DD3FB23FC90FB73" author="Giebel CG" box="[740,889,1218,1243]" pageId="25" pageNumber="1021" pagination="353 - 397" refId="ref18899" refString="Giebel CG. 1866. Die im zoologischen Museum der Universitat Halle aufgestellten Epizoen nebst Beobachtungen uber dieselben. Zeitschrift fur die Gesammten Naturwissenschaften (Halle). 28: 353 - 397." type="journal article" year="1866">Giebel (1866)</bibRefCitation>
and
<bibRefCitation id="EF9A95E5FFADCF383C85FB23FBAEFB73" author="Giebel CG" box="[946,1095,1218,1243]" pageId="25" pageNumber="1021" refId="ref18933" refString="Giebel CG. 1874. Insecta epizoa. Die auf Saugetieren und Vogeln schmarotzenden Insecten nach Chr. L. Nitzsch ̾ s Nachlass bearbeitet. Leipzig: Otto Wigand; xiv + 308 pp., 20 plates." type="book" year="1874">Giebel (1874)</bibRefCitation>
both write that the antennae are sexually monomorphic, which fits with
<taxonomicName id="4C0B9397FFADCF383CFDFB04FB6DFB56" authorityName="Nitzsch" authorityYear="1866" box="[970,1156,1253,1278]" class="Insecta" family="Philopteridae" genus="Goniocotes" kingdom="Animalia" order="Psocodea" pageId="25" pageNumber="1021" phylum="Arthropoda" rank="species" species="rectangulatus">
<emphasis id="B97F3406FFADCF383CFDFB04FB6DFB56" box="[970,1156,1253,1278]" italics="true" pageId="25" pageNumber="1021">G. rectangulatus</emphasis>
</taxonomicName>
(
<figureCitation id="1330F491FFADCF383F9FFAE6FEACFA88" box="[168,325,1287,1312]" captionStart-0="Figure 18" captionStart-1="Figure 19" captionStartId-0="21.[160,225,1746,1769]" captionStartId-1="22.[160,225,1746,1769]" captionTargetBox-0="[195,1122,143,1711]" captionTargetBox-1="[216,1097,143,1709]" captionTargetId-0="figure-1@21.[192,1123,142,1714]" captionTargetId-1="figure-1@22.[215,1100,142,1714]" captionTargetPageId-0="21" captionTargetPageId-1="22" captionText-0="Figure 18. Goniocotes rectangulatus Nitzsch [in Giebel], 1866, male habitus, dorsal and ventral views. Paratergal plates not clearly demarcated in specimen, and illustrated approximately." captionText-1="Figure 19. Goniocotes rectangulatus Nitzsch [in Giebel], 1866, female habitus, dorsal and ventral views. Paratergal plates not clearly demarcated in specimen, and illustrated approximately." figureDoi-0="http://doi.org/10.5281/zenodo.8270825" figureDoi-1="http://doi.org/10.5281/zenodo.8270827" httpUri-0="https://zenodo.org/record/8270825/files/figure.png" httpUri-1="https://zenodo.org/record/8270827/files/figure.png" pageId="25" pageNumber="1021">Figures 1819</figureCitation>
), but not with
<taxonomicName id="4C0B9397FFADCF383ED9FAE6FD86FA88" authorityName="Linnaeus" authorityYear="1758" box="[494,623,1287,1312]" class="Insecta" family="Goniodidae" genus="Pavoniocotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Psocodea" pageId="25" pageNumber="1021" phylum="Arthropoda" rank="species" species="parviceps">
<emphasis id="B97F3406FFADCF383ED9FAE6FD86FA88" box="[494,623,1287,1312]" italics="true" pageId="25" pageNumber="1021">P. parviceps</emphasis>
</taxonomicName>
(
<figureCitation id="1330F491FFADCF383D49FAE6FCF3FA88" box="[638,794,1287,1312]" captionStart="Figures 2630" captionStartId="30.[160,235,1580,1603]" captionTargetBox="[165,1150,143,1545]" captionTargetId="figure-1@30.[163,1153,142,1548]" captionTargetPageId="30" captionText="Figures 2630. Pavoniocotes parviceps (Piaget, 1880) ex Pavo cristatus Linnaeus, 1758. 26, male head, dorsal and ventral views. 27, female antenna and conus, ventral view. 28, male genitalia, dorsal view. 29, male genitalia, ventral view. 30, female genitalia and abdominal segments VIIXI, ventral view." figureDoi="http://doi.org/10.5281/zenodo.8270841" httpUri="https://zenodo.org/record/8270841/files/figure.png" pageId="25" pageNumber="1021">Figures 2627</figureCitation>
).
</paragraph>
<paragraph id="8BB4E814FFADCF383FF7FACBFBC0F99E" blockId="25.[160,1156,593,1728]" pageId="25" pageNumber="1021">
The species illustrated by
<bibRefCitation id="EF9A95E5FFADCF383ED5FACBFD70FAEB" author="von Keler S" box="[482,665,1322,1347]" pageId="25" pageNumber="1021" pagination="1 - 354" refId="ref21322" refString="von Keler S. 1940. Baustoffe zu einer Monographie der mallophagen. II. Teil: Uberfamilie der Nirmoidea. Nova Acta Leopoldiana Abhandlungen der Kaiserlich Leopoldinisch-Carolinisch Deutschen Akademie der Naturforscher Neue Folge. 8: 1 - 354 + 4 plates." type="book chapter" year="1940">von Kéler (1940)</bibRefCitation>
does not have temples that are ̍exact right angles̾, but only females were in Nitzsch̾s collection at the time Kéler examined it, and Kéler does not comment on the dimorphism of the antennae; however, Kéler̾s placement of this species in ̍
<taxonomicName id="4C0B9397FFADCF383E53FA73FDE9FA03" authorityName="von Kéler" authorityYear="1963" box="[356,512,1426,1451]" class="Insecta" family="Goniodidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Psocodea" pageId="25" pageNumber="1015" phylum="Arthropoda" rank="subFamily" subFamily="Goniocotinae">Goniocotinae</taxonomicName>
̾ implies that the antennae are sexually monomorphic, as this is a character of this subfamily. Conceivably, Nitzsch may have had access to additional specimens, which were lost by the time Kéler examined the collection. Nitzsch̾s collection may even have included some males of
<taxonomicName id="4C0B9397FFADCF383DD1FA1BFC86F9BB" box="[742,879,1530,1555]" class="Insecta" family="Goniodidae" genus="Pavoniocotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Psocodea" pageId="25" pageNumber="1021" phylum="Arthropoda" rank="species" species="parvifrons">
<emphasis id="B97F3406FFADCF383DD1FA1BFC86F9BB" box="[742,879,1530,1555]" italics="true" pageId="25" pageNumber="1021">P. parvifrons</emphasis>
</taxonomicName>
, on which he may have based the description of the temples and, presumably, the name ̍
<taxonomicName id="4C0B9397FFADCF383CBEF9FCFBF7F99E" authorityName="Nitzsch" authorityYear="1866" box="[905,1054,1565,1590]" class="Insecta" family="Philopteridae" genus="Goniocotes" kingdom="Animalia" order="Psocodea" pageId="25" pageNumber="1021" phylum="Arthropoda" rank="species" species="rectangulatus">
<emphasis id="B97F3406FFADCF383CBEF9FCFBF7F99E" box="[905,1054,1565,1590]" italics="true" pageId="25" pageNumber="1021">rectangulatus</emphasis>
</taxonomicName>
̾.
</paragraph>
<paragraph id="8BB4E814FFADCF3B3FF7F9DEFD6BFF62" blockId="25.[160,1156,593,1728]" lastBlockId="26.[160,1156,142,1450]" lastPageId="26" lastPageNumber="1022" pageId="25" pageNumber="1021">
Nevertheless, Kéler̾s description and illustration is based on one of the specimens identified by Nitzsch and Giebel as
<taxonomicName id="4C0B9397FFADCF383D1FF983FD30F9D3" authorityName="Nitzsch" authorityYear="1866" box="[552,729,1634,1659]" class="Insecta" family="Philopteridae" genus="Goniocotes" kingdom="Animalia" order="Psocodea" pageId="25" pageNumber="1021" phylum="Arthropoda" rank="species" species="rectangulatus">
<emphasis id="B97F3406FFADCF383D1FF983FD30F9D3" box="[552,729,1634,1659]" italics="true" pageId="25" pageNumber="1021">G. rectangulatus</emphasis>
</taxonomicName>
, and this constitutes the only detailed published illustration of this species to date. This is thus presumably the species that has been identified by researchers as
<taxonomicName id="4C0B9397FFADCF383D1EF946FD37F968" authorityName="Nitzsch" authorityYear="1866" box="[553,734,1703,1728]" class="Insecta" family="Philopteridae" genus="Goniocotes" kingdom="Animalia" order="Psocodea" pageId="25" pageNumber="1021" phylum="Arthropoda" rank="species" species="rectangulatus">
<emphasis id="B97F3406FFADCF383D1EF946FD37F968" box="[553,734,1703,1728]" italics="true" pageId="25" pageNumber="1021">G. rectangulatus</emphasis>
</taxonomicName>
since at least 1940. Neither Nitzsch, Giebel, nor Kéler designated any
<typeStatus id="54B056B6FFAECF3B3D3AFF6FFD98FF0F" box="[525,625,142,167]" pageId="26" pageNumber="1022" type="holotype">holotype</typeStatus>
or
<typeStatus id="54B056B6FFAECF3B3DA3FF6FFCE9FF0F" box="[660,768,142,167]" pageId="26" pageNumber="1022" type="lectotype">lectotype</typeStatus>
, but
<bibRefCitation id="EF9A95E5FFAECF3B3C03FF6FFC01FF0F" author="von Keler S" box="[820,1000,142,167]" pageId="26" pageNumber="1022" pagination="1 - 354" refId="ref21322" refString="von Keler S. 1940. Baustoffe zu einer Monographie der mallophagen. II. Teil: Uberfamilie der Nirmoidea. Nova Acta Leopoldiana Abhandlungen der Kaiserlich Leopoldinisch-Carolinisch Deutschen Akademie der Naturforscher Neue Folge. 8: 1 - 354 + 4 plates." type="book chapter" year="1940">von Kéler (1940)</bibRefCitation>
designated ̍2 complete and 1 damaged
<specimenCount id="9D0D239DFFAECF3B3EFDFF50FE37FF60" box="[458,478,177,200]" pageId="26" pageNumber="1022" type="female"></specimenCount>
̾ as (syn)types.
</paragraph>
<paragraph id="8BB4E814FFAECF3B3FF7FF35FD55FD25" blockId="26.[160,1156,142,1450]" pageId="26" pageNumber="1022">
<bibRefCitation id="EF9A95E5FFAECF3B3FF7FF35FD99FF45" author="Lakshminarayana KV &amp; Emerson KC" box="[192,624,212,237]" pageId="26" pageNumber="1022" pagination="95 - 102" refId="ref19904" refString="Lakshminarayana KV, Emerson KC. 1971. Mallophaga Indica. VI. Notes on Goniocotes (Mallophaga: Philopteridae) found on Pavo cristatus, with description of a new species. Orient Insects. 5: 95 - 102. doi: 10.1080 / 00305316.1971.10433994." type="journal article" year="1971">Lakshminarayana and Emerson (1971)</bibRefCitation>
and
<bibRefCitation id="EF9A95E5FFAECF3B3D9AFF35FC6FFF45" author="Nasser M &amp; Al-Ahmed A &amp; Shobrak M &amp; Aldryhim Y" box="[685,902,212,237]" pageId="26" pageNumber="1022" pagination="88 - 94" refId="ref20323" refString="Nasser M, Al-Ahmed A, Shobrak M, Aldryhim Y. 2015. Identification key for chewing lice (Phthiraptera: Amblycera, Ischnocera) infesting the Indian Peacock (Pavo cristatus) with one new country record and a new host record for Saudi Arabia. Turk J Zool. 38: 88 - 94. doi: 10.3906 / zoo- 1312 - 44." type="journal article" year="2015">
Nasser
<emphasis id="B97F3406FFAECF3B3C37FF35FCD8FF45" box="[768,817,212,237]" italics="true" pageId="26" pageNumber="1022">et al</emphasis>
. (2015)
</bibRefCitation>
claimed that the
<typeStatus id="54B056B6FFAECF3B3B62FF35FF35FEA7" pageId="26" pageNumber="1022" type="syntype">syntypes</typeStatus>
were lost, and the specimens we have examined are labelled ̍Neoparatypes̾. It is unclear where this designation originated. The collection locality (Wroclaw,
<collectingCountry id="F31CA884FFAECF3B3CD8FEF8FBA8FE9A" box="[1007,1089,281,306]" name="Poland" pageId="26" pageNumber="1022">Poland</collectingCountry>
), date and host are written in J. Złotorzycka̾s handwriting and style. However, the identification label is from the ZSI, and signed by K.V. Lakshminarayana. The lice were collected in 1975, but integrated into the collection of the NHMUK in 1993, where they are now the only specimens of
<taxonomicName id="4C0B9397FFAECF3B3E0BFE45FE07FE15" authorityName="Nitzsch" authorityYear="1866" box="[316,494,420,445]" class="Insecta" family="Philopteridae" genus="Goniocotes" kingdom="Animalia" order="Psocodea" pageId="26" pageNumber="1022" phylum="Arthropoda" rank="species" species="rectangulatus">
<emphasis id="B97F3406FFAECF3B3E0BFE45FE07FE15" box="[316,494,420,445]" italics="true" pageId="26" pageNumber="1022">G. rectangulatus</emphasis>
</taxonomicName>
[see
<bibRefCitation id="EF9A95E5FFAECF3B3D1DFE45FC3CFE15" author="Lakshminarayana KV &amp; Emerson KC" box="[554,981,420,445]" pageId="26" pageNumber="1022" pagination="95 - 102" refId="ref19904" refString="Lakshminarayana KV, Emerson KC. 1971. Mallophaga Indica. VI. Notes on Goniocotes (Mallophaga: Philopteridae) found on Pavo cristatus, with description of a new species. Orient Insects. 5: 95 - 102. doi: 10.1080 / 00305316.1971.10433994." type="journal article" year="1971">Lakshminarayana and Emerson (1971)</bibRefCitation>
on the identity of specimens with this name from the Piaget collection]. Notably, neither Złotorzycka̾s collection at MNHW nor the collection at the ZSI contains any
<typeStatus id="54B056B6FFAECF3B3C46FE08FC31FDAA" box="[881,984,489,514]" pageId="26" pageNumber="1022" type="neotype">neotypes</typeStatus>
of this species (
<bibRefCitation id="EF9A95E5FFAECF3B3F91FDEDFE4EFD8D" author="Lakshminarayana KV" box="[166,423,524,549]" pageId="26" pageNumber="1022" pagination="1 - 37" refId="ref19849" refString="Lakshminarayana KV. 1982. Notes on the type collection of the chewing-lice (Phthiraptera, Mallophaga s. l.: Insecta) in the Zoological Survey of India. Records of the Zoological Survey of India, Miscellaneous Publication, Occasional Paper No. 33, 1 - 37 + 7 plates." type="book chapter" year="1982">Lakshminarayana 1982</bibRefCitation>
;
<bibRefCitation id="EF9A95E5FFAECF3B3E85FDEDFD4AFD8D" author="Jaloszynski P &amp; Gustafsson DR &amp; Wanat A &amp; Wanat M" box="[434,675,524,549]" pageId="26" pageNumber="1022" pagination="645 - 661" refId="ref19453" refString="Jaloszynski P, Gustafsson DR, Wanat A, Wanat M. 2014. Type specimens of Phthiraptera in the collection of Jadwiga Zlotorzycka preserved in the Museum of Natural History, University of Wroclaw. Genus. 25: 645 - 661." type="journal article" year="2014">
Jałoszynski
<emphasis id="B97F3406FFAECF3B3D05FDEDFD89FD8D" box="[562,608,524,549]" italics="true" pageId="26" pageNumber="1022">et al</emphasis>
. 2014
</bibRefCitation>
;
<bibRefCitation id="EF9A95E5FFAECF3B3D99FDEDFC87FD8D" author="Sheela S &amp; Venkataraman K &amp; Das D" box="[686,878,524,549]" pageId="26" pageNumber="1022" pagination="1 - 43" refId="ref20806" refString="Sheela S, Venkataraman K, Das D. 2015. The type specimens in the National Zoological Collection. Phthiraptera. Type Catalogue Series. 2: 1 - 43." type="journal article" year="2015">
Sheela
<emphasis id="B97F3406FFAECF3B3DCAFDEDFCC5FD8D" box="[765,812,524,549]" italics="true" pageId="26" pageNumber="1022">et al</emphasis>
. 2015
</bibRefCitation>
), and we have not found any publication in which either Lakshminarayana or Złotorzycka designated
<typeStatus id="54B056B6FFAECF3B3CC9FDCFFB8CFDEF" box="[1022,1125,558,583]" pageId="26" pageNumber="1022" type="neotype">neotypes</typeStatus>
of
<taxonomicName id="4C0B9397FFAECF3B3F97FDB0FE8AFDC2" authorityName="Nitzsch" authorityYear="1866" box="[160,355,593,618]" class="Insecta" family="Philopteridae" genus="Goniocotes" kingdom="Animalia" order="Psocodea" pageId="26" pageNumber="1022" phylum="Arthropoda" rank="species" species="rectangulatus">
<emphasis id="B97F3406FFAECF3B3F97FDB0FE8AFDC2" box="[160,355,593,618]" italics="true" pageId="26" pageNumber="1022">G. rectangulatus</emphasis>
</taxonomicName>
. Moreover,
<bibRefCitation id="EF9A95E5FFAECF3B3D32FDB0FC35FDC2" author="Zlotorzycka J &amp; Modrzejewska M" box="[517,988,593,618]" pageId="26" pageNumber="1022" refId="ref21499" refString="Zlotorzycka J, Modrzejewska M. 1988. Katalog Fauny Polski. Czesc XIX, zeszyt 1. Wszoly. Mallophaga. Warszawa (Poland): Panstwowe Wydawnictwo Naukowe; 332 pp." type="book" year="1988">Złotorzycka and Modrzejewska (1988)</bibRefCitation>
did not list
<taxonomicName id="4C0B9397FFAECF3B3F97FD95FEBAFD25" authorityName="Nitzsch" authorityYear="1866" box="[160,339,628,653]" class="Insecta" family="Philopteridae" genus="Goniocotes" kingdom="Animalia" order="Psocodea" pageId="26" pageNumber="1022" phylum="Arthropoda" rank="species" species="rectangulatus">
<emphasis id="B97F3406FFAECF3B3F97FD95FEBAFD25" box="[160,339,628,653]" italics="true" pageId="26" pageNumber="1022">G. rectangulatus</emphasis>
</taxonomicName>
in their catalogue of Polish lice.
</paragraph>
<paragraph id="8BB4E814FFAECF3B3FF7FD77FC09FBE7" blockId="26.[160,1156,142,1450]" pageId="26" pageNumber="1022">
Regardless of where this
<typeStatus id="54B056B6FFAECF3B3EEAFD77FDD3FD07" box="[477,570,662,687]" pageId="26" pageNumber="1022" type="neotype">neotype</typeStatus>
designation was published, it was superfluous. The Halle collection, where most of the specimens examined by Nitzsch and Giebel were stored, was largely destroyed during World War II (eg
<bibRefCitation id="EF9A95E5FFAECF3B3C36FD3DFC87FD5D" author="Clay T" box="[769,878,732,757]" pageId="26" pageNumber="1022" pagination="1 - 25" refId="ref18090" refString="Clay T. 1949. Species of the genus Saemundssonia (Mallophaga) from the Sterninae. Am Mus Novit. 1409: 1 - 25." type="journal article" year="1949">Clay 1949</bibRefCitation>
; Hopkins and
<bibRefCitation id="EF9A95E5FFAECF3B3B27FD3DFB97FD5D" author="Clay T &amp; Hopkins GHE" box="[1040,1150,732,757]" pageId="26" pageNumber="1022" pagination="49 - 73" refId="ref18168" refString="Clay T, Hopkins GHE. 1955. Notes on the Rudow Collection of Mallophaga at Hamburg. Mitteilungen aus dem Hamburg Zoologischen Museum Institut. 53: 49 - 73." type="journal article" year="1955">Clay 1955</bibRefCitation>
;
<bibRefCitation id="EF9A95E5FFAECF3B3F97FD1FFE42FCBF" author="Palma RL &amp; Pilgrim RLC" box="[160,427,766,791]" pageId="26" pageNumber="1022" pagination="145 - 166" refId="ref20502" refString="Palma RL, Pilgrim RLC. 1984. A revision of the genus Harrisoniella (Mallophaga: Philopteridae). N Z J Zool. 11: 145 - 166. doi: 10.1080 / 03014223.1984.10423754." type="journal article" year="1984">Palma and Pilgrim 1984</bibRefCitation>
), but some specimens survived as they were not at the museum during the bombing. This supposedly includes the specimens listed by
<bibRefCitation id="EF9A95E5FFAECF3B3CF7FCC0FF0FFCF5" author="Clay T &amp; Hopkins GHE" pageId="26" pageNumber="1022" pagination="49 - 73" refId="ref18168" refString="Clay T, Hopkins GHE. 1955. Notes on the Rudow Collection of Mallophaga at Hamburg. Mitteilungen aus dem Hamburg Zoologischen Museum Institut. 53: 49 - 73." type="journal article" year="1955">Clay and Hopkins (1955)</bibRefCitation>
as well as those listed by
<bibRefCitation id="EF9A95E5FFAECF3B3D13FCA5FD46FCF5" author="von Keler S" box="[548,687,836,861]" pageId="26" pageNumber="1022" pagination="124 - 136" refId="ref21366" refString="von Keler S. 1941. Systematisches Verzeichnis der von Chr. L. Nitzsch begrundeten und Ch. G. A. von Giebel und O. Taschenberg fortgefuhrten und bereicherten Sammlung von Mallophagen des Zoologischen Instituts der Universitat Halle. Zeitschrift fur Naturwissenschaften. 95: 124 - 136." type="journal article" year="1941">Kéler (1941)</bibRefCitation>
; the latter list includes
<specimenCount id="9D0D239DFFAECF3B3CF1FCA5FB88FCF5" box="[966,1121,836,861]" pageId="26" pageNumber="1022" type="female">three females</specimenCount>
of
<taxonomicName id="4C0B9397FFAECF3B3F97FC87FEB8FCD7" authorityName="Nitzsch" authorityYear="1866" box="[160,337,870,895]" class="Insecta" family="Philopteridae" genus="Goniocotes" kingdom="Animalia" order="Psocodea" pageId="26" pageNumber="1022" phylum="Arthropoda" rank="species" species="rectangulatus">
<emphasis id="B97F3406FFAECF3B3F97FC87FEB8FCD7" box="[160,337,870,895]" italics="true" pageId="26" pageNumber="1022">G. rectangulatus</emphasis>
</taxonomicName>
that Kéler indicate are the types. Dr Hendrik Müller (
<emphasis id="B97F3406FFAECF3B3CAFFC87FC26FCD7" box="[920,975,870,895]" italics="true" pageId="26" pageNumber="1022">in litt</emphasis>
.) at the ZNS has confirmed that the specimens are still in that collection and provided some photos of the three remaining specimens (
<figureCitation id="1330F491FFAECF3B3EE9FC4DFDAFFC6D" box="[478,582,940,965]" captionStart="Figure 23" captionStartId="24.[160,225,1574,1597]" captionTargetBox="[163,1152,142,1528]" captionTargetId="figure-3@24.[163,1153,142,1543]" captionTargetPageId="24" captionText="Figure 23. Photographs of the remaining specimens (top), the slide labels (middle), and the slide itself (bottom) of Nitzsch̍s Goniocotes rectangulatus Nitzsch [in Giebel], 1866, at the Zentralmagazin Naturwissenschaftlischer Sammlungen, Martin-Luther-Universität, Halle-Wittenberg, provided by Dr Hendrik Müller at the same institute. These specimens constitute the syntype series of G. reticulatus, and are the same as those on which von Kéler (1940) based his illustrations." figureDoi="http://doi.org/10.5281/zenodo.8270833" httpUri="https://zenodo.org/record/8270833/files/figure.png" pageId="26" pageNumber="1022">Figure 23</figureCitation>
). These are the same specimens as those examined by
<bibRefCitation id="EF9A95E5FFAECF3B3FF4FC2FFE92FC4F" author="von Keler S" box="[195,379,974,999]" pageId="26" pageNumber="1022" pagination="1 - 354" refId="ref21322" refString="von Keler S. 1940. Baustoffe zu einer Monographie der mallophagen. II. Teil: Uberfamilie der Nirmoidea. Nova Acta Leopoldiana Abhandlungen der Kaiserlich Leopoldinisch-Carolinisch Deutschen Akademie der Naturforscher Neue Folge. 8: 1 - 354 + 4 plates." type="book chapter" year="1940">von Kéler (1940)</bibRefCitation>
, and thus constitute the only remaining
<typeStatus id="54B056B6FFAECF3B3C73FC2FFC4FFC4F" box="[836,934,974,999]" pageId="26" pageNumber="1022" type="syntype">syntypes</typeStatus>
of
<taxonomicName id="4C0B9397FFAECF3B3CFBFC2FFB97FC4F" authorityName="Nitzsch" authorityYear="1866" box="[972,1150,974,999]" class="Insecta" family="Philopteridae" genus="Goniocotes" kingdom="Animalia" order="Psocodea" pageId="26" pageNumber="1022" phylum="Arthropoda" rank="species" species="rectangulatus">
<emphasis id="B97F3406FFAECF3B3CFBFC2FFB97FC4F" box="[972,1150,974,999]" italics="true" pageId="26" pageNumber="1022">G. rectangulatus</emphasis>
</taxonomicName>
. These specimens appear to represent a typical species of
<taxonomicName id="4C0B9397FFAECF3B3C00FC10FC46FBA2" authorityName="Burmeister" authorityYear="1838" box="[823,943,1009,1034]" class="Insecta" family="Philopteridae" genus="Goniocotes" kingdom="Animalia" order="Psocodea" pageId="26" pageNumber="1022" phylum="Arthropoda" rank="genus">
<emphasis id="B97F3406FFAECF3B3C00FC10FC46FBA2" box="[823,943,1009,1034]" italics="true" pageId="26" pageNumber="1022">Goniocotes</emphasis>
</taxonomicName>
, indicating that at least since
<bibRefCitation id="EF9A95E5FFAECF3B3E16FBF5FE37FB85" author="von Keler S" box="[289,478,1044,1069]" pageId="26" pageNumber="1022" pagination="1 - 354" refId="ref21322" refString="von Keler S. 1940. Baustoffe zu einer Monographie der mallophagen. II. Teil: Uberfamilie der Nirmoidea. Nova Acta Leopoldiana Abhandlungen der Kaiserlich Leopoldinisch-Carolinisch Deutschen Akademie der Naturforscher Neue Folge. 8: 1 - 354 + 4 plates." type="book chapter" year="1940">von Kéler (1940)</bibRefCitation>
examined these specimens, there has been no mixture of species in the type series; there is therefore no need to select a
<typeStatus id="54B056B6FFAECF3B3C46FBD7FC34FBE7" box="[881,989,1078,1103]" pageId="26" pageNumber="1022" type="lectotype">lectotype</typeStatus>
.
</paragraph>
<paragraph id="8BB4E814FFAECF3B3FF7FBB8FE85FA02" blockId="26.[160,1156,142,1450]" pageId="26" pageNumber="1022">
<taxonomicName id="4C0B9397FFAECF3B3FF7FBB8FE3AFBDA" authorityName="Nitzsch" authorityYear="1866" box="[192,467,1113,1138]" class="Insecta" family="Philopteridae" genus="Goniocotes" kingdom="Animalia" order="Psocodea" pageId="26" pageNumber="1022" phylum="Arthropoda" rank="species" species="rectangulatus">
<emphasis id="B97F3406FFAECF3B3FF7FBB8FE3AFBDA" box="[192,467,1113,1138]" italics="true" pageId="26" pageNumber="1022">Goniocotes rectangulatus</emphasis>
</taxonomicName>
is the only member of
<taxonomicName id="4C0B9397FFAECF3B3DE5FBB8FCA3FBDA" authorityName="Burmeister" authorityYear="1838" box="[722,842,1113,1138]" class="Insecta" family="Philopteridae" genus="Goniocotes" kingdom="Animalia" order="Psocodea" pageId="26" pageNumber="1022" phylum="Arthropoda" rank="genus">
<emphasis id="B97F3406FFAECF3B3DE5FBB8FCA3FBDA" box="[722,842,1113,1138]" italics="true" pageId="26" pageNumber="1022">Goniocotes</emphasis>
</taxonomicName>
known from any host in the genus
<taxonomicName id="4C0B9397FFAECF3B3FDCFB9DFE0CFB3D" authority="(Price et al. 2003)" baseAuthorityName="Price" baseAuthorityYear="2003" box="[235,485,1148,1173]" class="Aves" family="Phasianidae" genus="Pavo" kingdom="Animalia" order="Galliformes" pageId="26" pageNumber="1022" phylum="Chordata" rank="genus">
<emphasis id="B97F3406FFAECF3B3FDCFB9DFEC9FB3D" box="[235,288,1148,1173]" italics="true" pageId="26" pageNumber="1022">Pavo</emphasis>
(
<bibRefCitation id="EF9A95E5FFAECF3B3E1BFB9DFE35FB3D" author="Price RD &amp; Hellenthal RA &amp; Palma RL" box="[300,476,1148,1173]" pageId="26" pageNumber="1022" pagination="501" refId="ref20568" refString="Price RD, Hellenthal RA, Palma RL. 2003. World checklist of chewing lice with host associations and keys to families and genera. In: Price RD, Hellenthal RA, Palma RL, Johnson KP, Clayton DH, editors. The chewing lice: world checklist and biological overview. Illinois Natural History Survey Special Publications 24, Illinois Natural History Survey. Champaign (IL USA); p. x + 501." type="book chapter" year="2003">
Price
<emphasis id="B97F3406FFAECF3B3E5CFB9DFE70FB3D" box="[363,409,1148,1173]" italics="true" pageId="26" pageNumber="1022">et al</emphasis>
. 2003
</bibRefCitation>
)
</taxonomicName>
. Two species of
<taxonomicName id="4C0B9397FFAECF3B3DAEFB9DFCF8FB3D" authorityName="Burmeister" authorityYear="1838" box="[665,785,1148,1173]" class="Insecta" family="Philopteridae" genus="Goniocotes" kingdom="Animalia" order="Psocodea" pageId="26" pageNumber="1022" phylum="Arthropoda" rank="genus" sensu="lato">
<emphasis id="B97F3406FFAECF3B3DAEFB9DFCF8FB3D" box="[665,785,1148,1173]" italics="true" pageId="26" pageNumber="1022">Goniocotes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24C897DFFAECF3B3C2FFB9DFCB9FB3D" box="[792,848,1148,1173]" pageId="26" pageNumber="1022" sensu="lato">
<emphasis id="B97F3406FFAECF3B3C2FFB9DFCA3FB3D" box="[792,842,1148,1173]" italics="true" pageId="26" pageNumber="1022">s. lat</emphasis>
.
</taxonomicNameLabel>
are known from the
<collectingCountry id="F31CA884FFAECF3B3B0EFB9DFB6AFB3D" box="[1081,1155,1148,1173]" name="Democratic Republic of the Congo" pageId="26" pageNumber="1022">Congo</collectingCountry>
peafowl (
<taxonomicName id="4C0B9397FFAECF3B3E3DFB7FFE09FB1F" authorityName="Chapin" authorityYear="1936" box="[266,480,1182,1207]" class="Aves" family="Phasianidae" genus="Afropavo" kingdom="Animalia" order="Galliformes" pageId="26" pageNumber="1022" phylum="Chordata" rank="species" species="congensis">
<emphasis id="B97F3406FFAECF3B3E3DFB7FFE09FB1F" box="[266,480,1182,1207]" italics="true" pageId="26" pageNumber="1022">Afropavo congensis</emphasis>
</taxonomicName>
), which are the closest relatives of
<taxonomicName id="4C0B9397FFAECF3B3C40FB7FFC45FB1F" box="[887,940,1182,1207]" class="Aves" family="Phasianidae" genus="Pavo" kingdom="Animalia" order="Galliformes" pageId="26" pageNumber="1022" phylum="Chordata" rank="genus">
<emphasis id="B97F3406FFAECF3B3C40FB7FFC45FB1F" box="[887,940,1182,1207]" italics="true" pageId="26" pageNumber="1022">Pavo</emphasis>
</taxonomicName>
(
<bibRefCitation id="EF9A95E5FFAECF3B3C8CFB7FFB97FB1F" author="Wang N &amp; Kimball RT &amp; Braun EL &amp; Liang B &amp; Zhang Z" box="[955,1150,1182,1207]" pageId="26" pageNumber="1022" refId="ref21418" refString="Wang N, Kimball RT, Braun EL, Liang B, Zhang Z. 2014. Assessing phylogenetic relationships among Galliformes: a multigene phylogeny with expanded taxon sampling in Phasianidae. PLoS One. 8: e 64312 [12 pp.]. doi: 10.1371 / journal. pone. 0064312." type="book" year="2014">
Wang
<emphasis id="B97F3406FFAECF3B3B30FB7FFBD1FB1F" box="[1031,1080,1182,1207]" italics="true" pageId="26" pageNumber="1022">et al</emphasis>
. 2014
</bibRefCitation>
;
<bibRefCitation id="EF9A95E5FFAECF3B3F97FB20FEBFFB72" author="Chen D &amp; Hosner PA &amp; Dittmann DL &amp; Neill JP &amp; Birks SM &amp; Braun EL &amp; Kimball RT" box="[160,342,1217,1242]" pageId="26" pageNumber="1022" pagination="209" refId="ref17975" refString="Chen D, Hosner PA, Dittmann DL, O ̾ Neill JP, Birks SM, Braun EL, Kimball RT. 2021. Divergence time estimation of Galliformes based on the best gene shopping scheme of ultraconserved elements. BMC Ecol Evol. 21: 209. doi: 10.1186 / s 12862 - 021 - 01935 - 1" type="journal article" year="2021">
Chen
<emphasis id="B97F3406FFAECF3B3FD5FB20FEFBFB72" box="[226,274,1217,1242]" italics="true" pageId="26" pageNumber="1022">et al</emphasis>
. 2021
</bibRefCitation>
;
<bibRefCitation id="EF9A95E5FFAECF3B3E54FB20FDD8FB72" author="Kimball RT &amp; Hosner PA &amp; Braun EL" box="[355,561,1217,1242]" pageId="26" pageNumber="1022" refId="ref19718" refString="Kimball RT, Hosner PA, Braun EL. 2021. A phylogenetic supermatrix of Galliformes (Landfowl) reveals biased branch lengths. Mol Phylogenet Evol. 158: 107091 [10 pp.]. doi: 10.1016 / j. ympev. 2021. 107091." type="book" year="2021">
Kimball
<emphasis id="B97F3406FFAECF3B3E89FB20FE04FB72" box="[446,493,1217,1242]" italics="true" pageId="26" pageNumber="1022">et al</emphasis>
. 2021
</bibRefCitation>
). However, the lice of
<collectingCountry id="F31CA884FFAECF3B3C19FB20FC91FB72" box="[814,888,1217,1242]" name="Democratic Republic of the Congo" pageId="26" pageNumber="1022">Congo</collectingCountry>
peafowl are morphologically distinct (
<bibRefCitation id="EF9A95E5FFAECF3B3E67FB05FE28FB55" author="Clay T" box="[336,449,1252,1277]" pageId="26" pageNumber="1022" pagination="1 - 11" refId="ref18036" refString="Clay T. 1938. New species of Mallophaga from Afropavo congensis Chapin. Am Mus Novit. 1008: 1 - 11." type="journal article" year="1938">Clay 1938</bibRefCitation>
), and likely do not belong in
<taxonomicName id="4C0B9397FFAECF3B3C22FB05FC64FB55" authorityName="Burmeister" authorityYear="1838" box="[789,909,1252,1277]" class="Insecta" family="Philopteridae" genus="Goniocotes" kingdom="Animalia" order="Psocodea" pageId="26" pageNumber="1022" phylum="Arthropoda" rank="genus">
<emphasis id="B97F3406FFAECF3B3C22FB05FC64FB55" box="[789,909,1252,1277]" italics="true" pageId="26" pageNumber="1022">Goniocotes</emphasis>
</taxonomicName>
. Given the paucity of verified records of
<taxonomicName id="4C0B9397FFAECF3B3E44FAE7FE02FAB7" authorityName="Burmeister" authorityYear="1838" box="[371,491,1286,1311]" class="Insecta" family="Philopteridae" genus="Goniocotes" kingdom="Animalia" order="Psocodea" pageId="26" pageNumber="1022" phylum="Arthropoda" rank="genus">
<emphasis id="B97F3406FFAECF3B3E44FAE7FE02FAB7" box="[371,491,1286,1311]" italics="true" pageId="26" pageNumber="1022">Goniocotes</emphasis>
</taxonomicName>
on peafowls,
<taxonomicName id="4C0B9397FFAECF3B3DBEFAE7FCD5FAB7" authorityName="Nitzsch" authorityYear="1866" box="[649,828,1286,1311]" class="Insecta" family="Philopteridae" genus="Goniocotes" kingdom="Animalia" order="Psocodea" pageId="26" pageNumber="1022" phylum="Arthropoda" rank="species" species="rectangulatus">
<emphasis id="B97F3406FFAECF3B3DBEFAE7FCD5FAB7" box="[649,828,1286,1311]" italics="true" pageId="26" pageNumber="1022">G. rectangulatus</emphasis>
</taxonomicName>
may represent a straggler to
<taxonomicName id="4C0B9397FFAECF3B3F97FAC8FEF3FAEA" authorityName="Linnaeus" authorityYear="1758" box="[160,282,1321,1346]" class="Aves" family="Phasianidae" genus="Pavo" kingdom="Animalia" order="Galliformes" pageId="26" pageNumber="1022" phylum="Chordata" rank="species" species="cristatus">
<emphasis id="B97F3406FFAECF3B3F97FAC8FEF3FAEA" box="[160,282,1321,1346]" italics="true" pageId="26" pageNumber="1022">P. cristatus</emphasis>
</taxonomicName>
from an unknown host; among the other species of
<taxonomicName id="4C0B9397FFAECF3B3CACFAC8FBFDFAEA" authorityName="Burmeister" authorityYear="1838" box="[923,1044,1321,1346]" class="Insecta" family="Philopteridae" genus="Goniocotes" kingdom="Animalia" order="Psocodea" pageId="26" pageNumber="1022" phylum="Arthropoda" rank="genus">
<emphasis id="B97F3406FFAECF3B3CACFAC8FBFDFAEA" box="[923,1044,1321,1346]" italics="true" pageId="26" pageNumber="1022">Goniocotes</emphasis>
</taxonomicName>
we have examined (DRG, in prep.), none are particularly close to
<taxonomicName id="4C0B9397FFAECF3B3C3DFAADFC52FACD" authorityName="Nitzsch" authorityYear="1866" box="[778,955,1356,1381]" class="Insecta" family="Philopteridae" genus="Goniocotes" kingdom="Animalia" order="Psocodea" pageId="26" pageNumber="1022" phylum="Arthropoda" rank="species" species="rectangulatus">
<emphasis id="B97F3406FFAECF3B3C3DFAADFC52FACD" box="[778,955,1356,1381]" italics="true" pageId="26" pageNumber="1022">G. rectangulatus</emphasis>
</taxonomicName>
, but most species in the genus have not been re-examined. A general revision of
<taxonomicName id="4C0B9397FFAECF3B3C43FA8FFC05FA2F" authorityName="Burmeister" authorityYear="1838" box="[884,1004,1390,1415]" class="Insecta" family="Philopteridae" genus="Goniocotes" kingdom="Animalia" order="Psocodea" pageId="26" pageNumber="1022" phylum="Arthropoda" rank="genus">
<emphasis id="B97F3406FFAECF3B3C43FA8FFC05FA2F" box="[884,1004,1390,1415]" italics="true" pageId="26" pageNumber="1022">Goniocotes</emphasis>
</taxonomicName>
is needed to address this issue.
</paragraph>
</subSubSection>
</treatment>
</document>