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<document id="B3A37892F002ED2FF14227E344DC6188" ID-DOI="10.11646/zootaxa.3920.3.2" ID-GBIF-Dataset="f491c6f8-0266-4c7e-8819-1e5c32c34f2e" ID-ISSN="1175-5326" ID-Zenodo-Dep="238674" ID-ZooBank="99ABD0BA-173B-418F-A87F-647867BD68A1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460997415643" checkinUser="plazi" docAuthor="Bolzern, Angelo &amp; Jäger, Peter" docDate="2015" docId="031F87E1FFF9011DFF5BFF7053717967" docLanguage="en" docName="zt03920p442.pdf" docOrigin="Zootaxa 3920 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Eratigena Bolzern, Burckhardt &amp; Hanggi 2013" docType="treatment" docVersion="11" lastPageNumber="434" masterDocId="FF26FF99FFFA0119FFCCFFE752647803" masterDocTitle="Unexpected occurrence of the genus Eratigena in Laos with description of a new species (Araneae: Agelenidae)" masterLastPageNumber="442" masterPageNumber="431" pageNumber="434" updateTime="1698563223356" updateUser="plazi">
<mods:mods id="BF8C04B5A61AA0C1E6DCEE7A091918CF" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="F6A16EBAFA20D37E05B5F86C779AF2AC">Unexpected occurrence of the genus Eratigena in Laos with description of a new species (Araneae: Agelenidae)</mods:title>
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<mods:namePart id="F14520A695D2F3C7DD470E29F46271B3">Bolzern, Angelo</mods:namePart>
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<mods:namePart id="C42419471A943355E2AF6C3D8FCAFEBC">Jäger, Peter</mods:namePart>
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<mods:title id="548BEC79574DFFE6EFE8CCF3525A2EE6">Zootaxa</mods:title>
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<mods:date id="F6EEAB3168D80E48815CEA59ACFC1316">2015</mods:date>
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<mods:number id="95681A198DAB0510178B4558FF4A02FB">3920</mods:number>
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<mods:classification id="6B8F632BFAE817E5221AC25AF624F553">journal article</mods:classification>
<mods:identifier id="D59538517C571E15CD08D41BC178E491" type="DOI">10.11646/zootaxa.3920.3.2</mods:identifier>
<mods:identifier id="39E329D59FB4996C38A4F1CD201D4FCA" type="GBIF-Dataset">f491c6f8-0266-4c7e-8819-1e5c32c34f2e</mods:identifier>
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<treatment id="031F87E1FFF9011DFF5BFF7053717967" ID-DOI="http://doi.org/10.5281/zenodo.5296389" ID-GBIF-Taxon="119597532" ID-Zenodo-Dep="5296389" LSID="urn:lsid:plazi:treatment:031F87E1FFF9011DFF5BFF7053717967" httpUri="http://treatment.plazi.org/id/031F87E1FFF9011DFF5BFF7053717967" lastPageId="4" lastPageNumber="434" pageId="3" pageNumber="434">
<subSubSection id="C3AC657CFFF9011AFF5BFF70508278B1" box="[151,742,151,178]" pageId="3" pageNumber="434" type="nomenclature">
<paragraph id="8B0936F7FFF9011AFF5BFF70508278B1" blockId="3.[151,742,151,178]" box="[151,742,151,178]" pageId="3" pageNumber="434">
<heading id="D041819BFFF9011AFF5BFF70508278B1" bold="true" box="[151,742,151,178]" fontSize="11" level="1" pageId="3" pageNumber="434" reason="1">
<taxonomicName id="4CB64D74FFF9011AFF5BFF70508278B1" ID-CoL="633YC" ID-ENA="1522684" authority="Bolzern, Burckhardt &amp; Hanggi, 2013" authorityName="Bolzern, Burckhardt &amp; Hanggi" authorityYear="2013" box="[151,742,151,178]" class="Arachnida" family="Agelenidae" genus="Eratigena" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="434" phylum="Arthropoda" rank="genus">
<emphasis id="B9C2EAE5FFF9011AFF5BFF70508278B1" bold="true" box="[151,742,151,178]" pageId="3" pageNumber="434">
<emphasis id="B9C2EAE5FFF9011AFF5BFF70537478B2" bold="true" box="[151,272,151,177]" italics="true" pageId="3" pageNumber="434">Eratigena</emphasis>
<bibRefCitation id="EF274B06FFF9011AFEDAFF7F508278B1" author="Bolzern" box="[278,742,152,178]" pageId="3" pageNumber="434" refString="Bolzern, A., Burckhardt, D. &amp; Hanggi, A. (2013) Phylogeny and taxonomy of European funnel-web spiders of the Tegenaria- Malthonica complex (Araneae: Agelenidae) based upon morphological and molecular data. Zoological Journal of the Linnean Society, 168, 723 - 848. http: // dx. doi. org / 10.1111 / zoj. 12040" type="journal article" year="2013">Bolzern, Burckhardt &amp; Hänggi, 2013</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3AC657CFFF9011AFF5BFF07568478F7" box="[151,1248,222,245]" pageId="3" pageNumber="434" type="reference_group">
<paragraph id="8B0936F7FFF9011AFF5BFF07568478F7" blockId="3.[151,1248,222,245]" box="[151,1248,222,245]" pageId="3" pageNumber="434">
<treatmentCitationGroup id="ABA611D9FFF9011AFF5BFF07568478F7" box="[151,1248,222,245]" pageId="3" pageNumber="434">
<taxonomicName id="4CB64D74FFF9011AFF5BFF07539E78F7" ID-CoL="633YC" authority="Bolzern et al. 2013: 738" authorityName="Bolzern et al." authorityPageNumber="738" authorityYear="2013" box="[151,506,222,245]" class="Arachnida" family="Agelenidae" genus="Eratigena" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="434" phylum="Arthropoda" rank="genus">
<emphasis id="B9C2EAE5FFF9011AFF5BFF07529878F6" box="[151,252,224,245]" italics="true" pageId="3" pageNumber="434">Eratigena</emphasis>
<treatmentCitation id="0A1710E6FFF9011AFECEFF39539E78F7" ID-DOI="http://doi.org/10.5281/zenodo.6984000" ID-Zenodo-Dep="6984000" LSID="urn:lsid:zoobank.org:act:4E508ABD-A527-4D70-8BE6-075F3A5948F4" author="Bolzern" box="[258,506,222,245]" httpUri="http://treatment.plazi.org/id/BD701413E232B65657FBFB1FC09E113D" page="738" pageId="3" pageNumber="434" year="2013">
<bibRefCitation id="EF274B06FFF9011AFECEFF3953AC78F7" author="Bolzern" box="[258,456,222,245]" pageId="3" pageNumber="434" refString="Bolzern, A., Burckhardt, D. &amp; Hanggi, A. (2013) Phylogeny and taxonomy of European funnel-web spiders of the Tegenaria- Malthonica complex (Araneae: Agelenidae) based upon morphological and molecular data. Zoological Journal of the Linnean Society, 168, 723 - 848. http: // dx. doi. org / 10.1111 / zoj. 12040" type="journal article" year="2013">
Bolzern
<emphasis id="B9C2EAE5FFF9011AFE95FF0753E878F7" box="[345,396,222,245]" italics="true" pageId="3" pageNumber="434">et al.</emphasis>
2013
</bibRefCitation>
: 738
</treatmentCitation>
</taxonomicName>
.
<typeStatus id="540D8855FFF9011AFDC8FF38505278F7" box="[516,566,223,244]" pageId="3" pageNumber="434">Type</typeStatus>
species:
<taxonomicName id="4CB64D74FFF9011AFD5FFF07566178F7" ID-CoL="6GCBJ" authority="C. L. Koch, 1843" authorityName="C. L. Koch" authorityYear="1843" box="[659,1029,222,245]" class="Arachnida" family="Agelenidae" genus="Eratigena" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="434" phylum="Arthropoda" rank="species" species="atrica">
<emphasis id="B9C2EAE5FFF9011AFD5FFF07515F78F6" box="[659,827,224,245]" italics="true" pageId="3" pageNumber="434">Eratigena atrica</emphasis>
(C. L. Koch, 1843)
</taxonomicName>
, original designation.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3AC657CFFF9011AFF5BFEC6538B79A5" pageId="3" pageNumber="434" type="discussion">
<paragraph id="8B0936F7FFF9011AFF5BFEC6538B79A5" blockId="3.[151,1436,289,422]" pageId="3" pageNumber="434">
<emphasis id="B9C2EAE5FFF9011AFF5BFEC6536F7939" bold="true" box="[151,267,289,314]" pageId="3" pageNumber="434">Remarks.</emphasis>
In this recently designated genus, 18 species have been placed so far (Word Spider Catalog 2014), 16 of which are mainly limited to
<collectingCountry id="F3A17667FFF9011AFE3AFEA2504E795D" box="[502,554,325,350]" name="Italy" pageId="3" pageNumber="434">Italy</collectingCountry>
,
<collectingCountry id="F3A17667FFF9011AFDF0FEA150EF795E" box="[572,651,326,349]" name="France" pageId="3" pageNumber="434">France</collectingCountry>
, and the Iberian Peninsula. Only two species,
<taxonomicName id="4CB64D74FFF9011AFB0AFEA153157982" authority="Walckenaer, 1802" authorityName="Walckenaer" authorityYear="1802" class="Arachnida" family="Agelenidae" genus="Eratigena" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="434" phylum="Arthropoda" rank="species" species="agrestis">
<emphasis id="B9C2EAE5FFF9011AFB0AFEA157F8795E" box="[1222,1436,326,349]" italics="true" pageId="3" pageNumber="434">Eratigena agrestis</emphasis>
(Walckenaer, 1802)
</taxonomicName>
and
<taxonomicName id="4CB64D74FFF9011AFE6BFE8D506D7982" box="[423,521,362,385]" class="Arachnida" family="Agelenidae" genus="Eratigena" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="434" phylum="Arthropoda" rank="species" species="atrica">
<emphasis id="B9C2EAE5FFF9011AFE6BFE8D506D7982" box="[423,521,362,385]" italics="true" pageId="3" pageNumber="434">E. atrica</emphasis>
</taxonomicName>
, also occur in Central Europe and may have been introduced to the
<collectingCountry id="F3A17667FFF9011AFB31FE8D57417982" box="[1277,1317,362,385]" name="United Kingdom" pageId="3" pageNumber="434">UK</collectingCountry>
and North
<collectingCountry id="F3A17667FFF9011AFF5BFE6A529379A5" box="[151,247,397,422]" name="United States of America" pageId="3" pageNumber="434">America</collectingCountry>
(
<bibRefCitation id="EF274B06FFF9011AFECAFE6A538779A5" author="Bolzern" box="[262,483,397,422]" pageId="3" pageNumber="434" refString="Bolzern, A., Burckhardt, D. &amp; Hanggi, A. (2013) Phylogeny and taxonomy of European funnel-web spiders of the Tegenaria- Malthonica complex (Araneae: Agelenidae) based upon morphological and molecular data. Zoological Journal of the Linnean Society, 168, 723 - 848. http: // dx. doi. org / 10.1111 / zoj. 12040" type="journal article" year="2013">
Bolzern
<emphasis id="B9C2EAE5FFF9011AFEABFE6953FB79A5" box="[359,415,397,422]" italics="true" pageId="3" pageNumber="434">et al.</emphasis>
2013
</bibRefCitation>
).
</paragraph>
</subSubSection>
<caption id="DFC9667FFFF9011AFF5BF9EA50EF7E41" httpUri="https://zenodo.org/record/238675/files/figure.png" pageId="3" pageNumber="434" targetBox="[150,1430,487,1506]" targetPageId="3">
<paragraph id="8B0936F7FFF9011AFF5BF9EA50EF7E41" blockId="3.[151,1436,1549,1602]" pageId="3" pageNumber="434">
<emphasis id="B9C2EAE5FFF9011AFF5BF9EA53587E20" bold="true" box="[151,316,1549,1572]" pageId="3" pageNumber="434">FIGURES 1–4.</emphasis>
<taxonomicName id="4CB64D74FFF9011AFE88F9E853977E27" box="[324,499,1550,1572]" class="Arachnida" family="Agelenidae" genus="Eratigena" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="434" phylum="Arthropoda" rank="species" species="laksao" status="sp. nov.">
<emphasis id="B9C2EAE5FFF9011AFE88F9E853977E27" box="[324,499,1550,1572]" italics="true" pageId="3" pageNumber="434">Eratigena laksao</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2F1579EFFF9011AFE30F9E950537E20" box="[508,567,1550,1571]" pageId="3" pageNumber="434" rank="species">
<emphasis id="B9C2EAE5FFF9011AFE30F9E950537E20" bold="true" box="[508,567,1550,1571]" pageId="3" pageNumber="434">sp. n.</emphasis>
</taxonomicNameLabel>
, male paratype habitus of living specimen (1 lateral view; 2 anterior view; 3 dorsal view; 4 same specimen as subadult, dorsal view).
</paragraph>
</caption>
<subSubSection id="C3AC657CFFF9011DFF0BF98D53717967" lastPageId="4" lastPageNumber="435" pageId="3" pageNumber="434" type="diagnosis">
<paragraph id="8B0936F7FFF9011DFF0BF98D53717967" blockId="3.[151,1436,1642,2027]" lastBlockId="4.[151,1436,151,357]" lastPageId="4" lastPageNumber="435" pageId="3" pageNumber="434">
<emphasis id="B9C2EAE5FFF9011AFF0BF98D53257E80" bold="true" box="[199,321,1642,1667]" pageId="3" pageNumber="434">Diagnosis.</emphasis>
Medium to large sized (carapace length between 2 and
<quantity id="4C4E9B12FFF9011AFC17F98C56457E81" box="[987,1057,1642,1666]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="3" pageNumber="434" unit="mm" value="7.0">7 mm</quantity>
)
<taxonomicName id="4CB64D74FFF9011AFBF8F98D56D07E80" box="[1076,1204,1642,1667]" class="Arachnida" family="Agelenidae" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="434" phylum="Arthropoda" rank="family">Agelenidae</taxonomicName>
with plumose hairs present (absent in
<taxonomicName id="4CB64D74FFF9011AFEA8F96A50177EA5" authority="Lucas, 1846" authorityName="Lucas" authorityYear="1846" box="[356,627,1677,1702]" class="Arachnida" family="Agelenidae" genus="Lycosoides" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="434" phylum="Arthropoda" rank="genus">
<emphasis id="B9C2EAE5FFF9011AFEA8F96A53847EA5" box="[356,480,1677,1702]" italics="true" pageId="3" pageNumber="434">Lycosoides</emphasis>
Lucas, 1846
</taxonomicName>
,
<taxonomicName id="4CB64D74FFF9011AFDB2F96851F97EA5" authority="Lehtinen, 1967" authorityName="Lehtinen" authorityYear="1967" box="[638,925,1677,1702]" class="Arachnida" family="Agelenidae" genus="Maimuna" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="434" phylum="Arthropoda" rank="genus">
<emphasis id="B9C2EAE5FFF9011AFDB2F968508D7EA5" box="[638,745,1679,1702]" italics="true" pageId="3" pageNumber="434">Maimuna</emphasis>
Lehtinen, 1967
</taxonomicName>
, and
<taxonomicName id="4CB64D74FFF9011AFC14F96856817EA5" authority="Sundevall, 1833" authorityName="Sundevall" authorityYear="1833" box="[984,1253,1677,1702]" class="Arachnida" family="Agelenidae" genus="Textrix" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="434" phylum="Arthropoda" rank="genus">
<emphasis id="B9C2EAE5FFF9011AFC14F96856477EA5" box="[984,1059,1679,1702]" italics="true" pageId="3" pageNumber="434">Textrix</emphasis>
Sundevall, 1833
</taxonomicName>
), AER and PER straight or only slightly curved in dorsal view (
<figureCitation id="138D2A72FFF9011AFD6AF95551687EC8" box="[678,780,1714,1739]" captionStart-0="FIGURES 1 – 4" captionStart-1="FIGURES 5 – 13" captionStartId-0="3.[151,264,1549,1572]" captionStartId-1="4.[151,264,1667,1690]" captionTargetBox-0="[150,1430,487,1506]" captionTargetBox-1="[165,1419,403,1636]" captionTargetId-0="figure@3.[145,939,487,1105]" captionTargetId-1="figure@4.[529,1025,816,1312]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURES 1 – 4. Eratigena laksao sp. n., male paratype habitus of living specimen (1 lateral view; 2 anterior view; 3 dorsal view; 4 same specimen as subadult, dorsal view)." captionText-1="FIGURES 5 – 13. Eratigena laksao sp. n., male holotype. 5. Carapace, dorsal view. 6. Abdomen, dorsal view. 7. Carapace and chelicerae, anterior view. 8. Sternum, ventral view. 9. Spinnerets, ventral view. 10. Chelicerae, posteroventral view. 11. Mouthpart, ventral view. 12. Coxa with trochanter and basal femora, ventral view. 13. Claws of leg 1, lateral view." httpUri-0="https://zenodo.org/record/238675/files/figure.png" httpUri-1="https://zenodo.org/record/238676/files/figure.png" pageId="3" pageNumber="434">Figs 3–5</figureCitation>
; both rows recurved in
<taxonomicName id="4CB64D74FFF9011AFBD8F95556F47EC8" box="[1044,1168,1714,1739]" class="Arachnida" family="Agelenidae" genus="Lycosoides" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="434" phylum="Arthropoda" rank="genus">
<emphasis id="B9C2EAE5FFF9011AFBD8F95556F47EC8" box="[1044,1168,1714,1739]" italics="true" pageId="3" pageNumber="434">Lycosoides</emphasis>
</taxonomicName>
,
<taxonomicName id="4CB64D74FFF9011AFB52F954576D7EC9" box="[1182,1289,1715,1738]" class="Arachnida" family="Agelenidae" genus="Maimuna" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="434" phylum="Arthropoda" rank="genus">
<emphasis id="B9C2EAE5FFF9011AFB52F954576D7EC9" box="[1182,1289,1715,1738]" italics="true" pageId="3" pageNumber="434">Maimuna</emphasis>
</taxonomicName>
, and
<taxonomicName id="4CB64D74FFF9011AFA8BF95457F67EC9" box="[1351,1426,1715,1738]" class="Arachnida" family="Agelenidae" genus="Textrix" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="434" phylum="Arthropoda" rank="genus">
<emphasis id="B9C2EAE5FFF9011AFA8BF95457F67EC9" box="[1351,1426,1715,1738]" italics="true" pageId="3" pageNumber="434">Textrix</emphasis>
</taxonomicName>
; both rows procurved in
<taxonomicName id="4CB64D74FFF9011AFE6FF93250AA7EED" authority="Walckenaer, 1805" authorityName="Walckenaer" authorityYear="1805" box="[419,718,1749,1774]" class="Arachnida" family="Agelenidae" genus="Agelena" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="434" phylum="Arthropoda" rank="genus">
<emphasis id="B9C2EAE5FFF9011AFE6FF932539B7EED" box="[419,511,1749,1774]" italics="true" pageId="3" pageNumber="434">Agelena</emphasis>
Walckenaer, 1805
</taxonomicName>
,
<taxonomicName id="4CB64D74FFF9011AFD10F93251847EED" authority="Levy, 1996" authorityName="Levy" authorityYear="1996" box="[732,992,1749,1775]" class="Arachnida" family="Agelenidae" genus="Agelescape" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="434" phylum="Arthropoda" rank="genus">
<emphasis id="B9C2EAE5FFF9011AFD10F93251387EED" box="[732,860,1749,1774]" italics="true" pageId="3" pageNumber="434">Agelescape</emphasis>
Levy, 1996
</taxonomicName>
,
<taxonomicName id="4CB64D74FFF9011AFC22F93256957EED" authority="Zhang, Zhu" authorityName="Zhang, Zhu" box="[1006,1265,1749,1774]" class="Arachnida" family="Agelenidae" genus="Allagelena" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="434" phylum="Arthropoda" rank="genus">
<emphasis id="B9C2EAE5FFF9011AFC22F93256037EED" box="[1006,1127,1749,1774]" italics="true" pageId="3" pageNumber="434">Allagelena</emphasis>
Zhang, Zhu
</taxonomicName>
&amp; Song, 2006, and
<taxonomicName id="4CB64D74FFF9011AFF05F91C53847F10" authority="Lehtinen, 1967" authorityName="Lehtinen" authorityYear="1967" box="[201,480,1786,1811]" class="Arachnida" family="Agelenidae" genus="Benoitia" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="434" phylum="Arthropoda" rank="genus">
<emphasis id="B9C2EAE5FFF9011AFF05F91C534C7F11" box="[201,296,1787,1810]" italics="true" pageId="3" pageNumber="434">Benoitia</emphasis>
Lehtinen, 1967
</taxonomicName>
) and moderately procurved in frontal view (
<figureCitation id="138D2A72FFF9011AFC2BF91D56577F11" box="[999,1075,1786,1811]" captionStart="FIGURES 1 – 4" captionStartId="3.[151,264,1549,1572]" captionTargetBox="[150,1430,487,1506]" captionTargetId="figure@3.[145,939,487,1105]" captionTargetPageId="3" captionText="FIGURES 1 – 4. Eratigena laksao sp. n., male paratype habitus of living specimen (1 lateral view; 2 anterior view; 3 dorsal view; 4 same specimen as subadult, dorsal view)." httpUri="https://zenodo.org/record/238675/files/figure.png" pageId="3" pageNumber="434">Figs 2</figureCitation>
,
<figureCitation id="138D2A72FFF9011AFB8EF91C56367F11" box="[1090,1106,1787,1810]" captionStart="FIGURES 5 – 13" captionStartId="4.[151,264,1667,1690]" captionTargetBox="[165,1419,403,1636]" captionTargetId="figure@4.[529,1025,816,1312]" captionTargetPageId="4" captionText="FIGURES 5 – 13. Eratigena laksao sp. n., male holotype. 5. Carapace, dorsal view. 6. Abdomen, dorsal view. 7. Carapace and chelicerae, anterior view. 8. Sternum, ventral view. 9. Spinnerets, ventral view. 10. Chelicerae, posteroventral view. 11. Mouthpart, ventral view. 12. Coxa with trochanter and basal femora, ventral view. 13. Claws of leg 1, lateral view." httpUri="https://zenodo.org/record/238676/files/figure.png" pageId="3" pageNumber="434">7</figureCitation>
; AER strongly procurved in
<taxonomicName id="4CB64D74FFF9011AFF5BF8FA52977F35" box="[151,243,1821,1846]" class="Arachnida" family="Agelenidae" genus="Agelena" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="434" phylum="Arthropoda" rank="genus">
<emphasis id="B9C2EAE5FFF9011AFF5BF8FA52977F35" box="[151,243,1821,1846]" italics="true" pageId="3" pageNumber="434">Agelena</emphasis>
</taxonomicName>
,
<taxonomicName id="4CB64D74FFF9011AFECDF8FA53E47F35" box="[257,384,1821,1846]" class="Arachnida" family="Agelenidae" genus="Agelescape" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="434" phylum="Arthropoda" rank="genus">
<emphasis id="B9C2EAE5FFF9011AFECDF8FA53E47F35" box="[257,384,1821,1846]" italics="true" pageId="3" pageNumber="434">Agelescape</emphasis>
</taxonomicName>
,
<taxonomicName id="4CB64D74FFF9011AFE43F8FA506C7F35" box="[399,520,1821,1846]" class="Arachnida" family="Agelenidae" genus="Allagelena" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="434" phylum="Arthropoda" rank="genus">
<emphasis id="B9C2EAE5FFF9011AFE43F8FA506C7F35" box="[399,520,1821,1846]" italics="true" pageId="3" pageNumber="434">Allagelena</emphasis>
</taxonomicName>
,
<taxonomicName id="4CB64D74FFF9011AFDDAF8F850117F35" box="[534,629,1823,1846]" class="Arachnida" family="Agelenidae" genus="Benoitia" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="434" phylum="Arthropoda" rank="genus">
<emphasis id="B9C2EAE5FFF9011AFDDAF8F850117F35" box="[534,629,1823,1846]" italics="true" pageId="3" pageNumber="434">Benoitia</emphasis>
</taxonomicName>
, and
<taxonomicName id="4CB64D74FFF9011AFD7FF8FA51AA7F35" authority="Simon, 1898" authorityName="Simon" authorityYear="1898" box="[691,974,1821,1846]" class="Arachnida" family="Agelenidae" genus="Malthonica" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="434" phylum="Arthropoda" rank="genus">
<emphasis id="B9C2EAE5FFF9011AFD7FF8FA51507F35" box="[691,820,1821,1846]" italics="true" pageId="3" pageNumber="434">Malthonica</emphasis>
Simon, 1898
</taxonomicName>
; AER recurved in
<taxonomicName id="4CB64D74FFF9011AFB6CF8FA57787F35" box="[1184,1308,1821,1846]" class="Arachnida" family="Agelenidae" genus="Lycosoides" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="434" phylum="Arthropoda" rank="genus">
<emphasis id="B9C2EAE5FFF9011AFB6CF8FA57787F35" box="[1184,1308,1821,1846]" italics="true" pageId="3" pageNumber="434">Lycosoides</emphasis>
</taxonomicName>
,
<taxonomicName id="4CB64D74FFF9011AFAE6F8F857F17F35" box="[1322,1429,1823,1846]" class="Arachnida" family="Agelenidae" genus="Maimuna" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="434" phylum="Arthropoda" rank="genus">
<emphasis id="B9C2EAE5FFF9011AFAE6F8F857F17F35" box="[1322,1429,1823,1846]" italics="true" pageId="3" pageNumber="434">Maimuna</emphasis>
</taxonomicName>
, and
<taxonomicName id="4CB64D74FFF9011AFF04F8A453777F59" box="[200,275,1859,1882]" class="Arachnida" family="Agelenidae" genus="Textrix" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="434" phylum="Arthropoda" rank="genus">
<emphasis id="B9C2EAE5FFF9011AFF04F8A453777F59" box="[200,275,1859,1882]" italics="true" pageId="3" pageNumber="434">Textrix</emphasis>
</taxonomicName>
), cheliceral retromargin with six or more teeth (
<figureCitation id="138D2A72FFF9011AFCF2F8A551F77F59" box="[830,915,1858,1883]" captionStart="FIGURES 5 – 13" captionStartId="4.[151,264,1667,1690]" captionTargetBox="[165,1419,403,1636]" captionTargetId="figure@4.[529,1025,816,1312]" captionTargetPageId="4" captionText="FIGURES 5 – 13. Eratigena laksao sp. n., male holotype. 5. Carapace, dorsal view. 6. Abdomen, dorsal view. 7. Carapace and chelicerae, anterior view. 8. Sternum, ventral view. 9. Spinnerets, ventral view. 10. Chelicerae, posteroventral view. 11. Mouthpart, ventral view. 12. Coxa with trochanter and basal femora, ventral view. 13. Claws of leg 1, lateral view." httpUri="https://zenodo.org/record/238676/files/figure.png" pageId="3" pageNumber="434">Fig. 10</figureCitation>
; up to six teeth in
<taxonomicName id="4CB64D74FFF9011AFBA0F8A457F47F59" authority="Latreille, 1804" authorityName="Latreille" authorityYear="1804" box="[1132,1424,1858,1883]" class="Arachnida" family="Agelenidae" genus="Tegenaria" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="434" phylum="Arthropoda" rank="genus">
<emphasis id="B9C2EAE5FFF9011AFBA0F8A456B97F59" box="[1132,1245,1859,1882]" italics="true" pageId="3" pageNumber="434">Tegenaria</emphasis>
Latreille, 1804
</taxonomicName>
), straight or slightly curved trochanters (
<figureCitation id="138D2A72FFF9011AFD85F88250F87F7D" box="[585,668,1893,1918]" captionStart="FIGURES 5 – 13" captionStartId="4.[151,264,1667,1690]" captionTargetBox="[165,1419,403,1636]" captionTargetId="figure@4.[529,1025,816,1312]" captionTargetPageId="4" captionText="FIGURES 5 – 13. Eratigena laksao sp. n., male holotype. 5. Carapace, dorsal view. 6. Abdomen, dorsal view. 7. Carapace and chelicerae, anterior view. 8. Sternum, ventral view. 9. Spinnerets, ventral view. 10. Chelicerae, posteroventral view. 11. Mouthpart, ventral view. 12. Coxa with trochanter and basal femora, ventral view. 13. Claws of leg 1, lateral view." httpUri="https://zenodo.org/record/238676/files/figure.png" pageId="3" pageNumber="434">Fig. 12</figureCitation>
; notched in
<taxonomicName id="4CB64D74FFF9011AFCEFF88156137F7D" authority="Bolzern et al., 2010" authorityName="Bolzern et al." authorityYear="2010" box="[803,1143,1893,1918]" class="Arachnida" family="Agelenidae" genus="Aterigena" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="434" phylum="Arthropoda" rank="genus">
<emphasis id="B9C2EAE5FFF9011AFCEFF88151F57F7E" box="[803,913,1894,1917]" italics="true" pageId="3" pageNumber="434">Aterigena</emphasis>
<bibRefCitation id="EF274B06FFF9011AFC54F88256137F7D" author="Bolzern" box="[920,1143,1893,1918]" pageId="3" pageNumber="434" refString="Bolzern, A., Hanggi, A. &amp; Burckhardt, D. (2010) Aterigena, a new genus of funnel-web spider, shedding some light on the Tegenaria-Malthonica problem (Araneae: Agelenidae). The Journal of Arachnology, 28, 162 - 182. http: // dx. doi. org / 10.1636 / A 09 - 78.1" type="journal article" year="2010">Bolzern et al., 2010</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4CB64D74FFF9011AFB4DF88057FC7F7D" authority="Thorell, 1869" authorityName="Thorell" authorityYear="1869" box="[1153,1432,1893,1918]" class="Arachnida" family="Agelenidae" genus="Histopona" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="434" phylum="Arthropoda" rank="genus">
<emphasis id="B9C2EAE5FFF9011AFB4DF88056927F7D" box="[1153,1270,1895,1918]" italics="true" pageId="3" pageNumber="434">Histopona</emphasis>
Thorell, 1869
</taxonomicName>
, and
<taxonomicName id="4CB64D74FFF9011AFF0AF86D53237FA0" box="[198,327,1930,1955]" class="Arachnida" family="Agelenidae" genus="Malthonica" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="434" phylum="Arthropoda" rank="genus">
<emphasis id="B9C2EAE5FFF9011AFF0AF86D53237FA0" box="[198,327,1930,1955]" italics="true" pageId="3" pageNumber="434">Malthonica</emphasis>
</taxonomicName>
), lateral spines at patellae absent (as in
<taxonomicName id="4CB64D74FFF9011AFCCBF86C511F7FA1" box="[775,891,1931,1954]" class="Arachnida" family="Agelenidae" genus="Histopona" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="434" phylum="Arthropoda" rank="genus">
<emphasis id="B9C2EAE5FFF9011AFCCBF86C511F7FA1" box="[775,891,1931,1954]" italics="true" pageId="3" pageNumber="434">Histopona</emphasis>
</taxonomicName>
,
<taxonomicName id="4CB64D74FFF9011AFC46F86D566F7FA0" box="[906,1035,1930,1955]" class="Arachnida" family="Agelenidae" genus="Malthonica" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="434" phylum="Arthropoda" rank="genus">
<emphasis id="B9C2EAE5FFF9011AFC46F86D566F7FA0" box="[906,1035,1930,1955]" italics="true" pageId="3" pageNumber="434">Malthonica</emphasis>
</taxonomicName>
, and T
<emphasis id="B9C2EAE5FFF9011AFB9BF86C56DF7FA1" box="[1111,1211,1931,1954]" italics="true" pageId="3" pageNumber="434">egenaria</emphasis>
; present in all other European genera), and colulus forming a rectangular or trapezoidal plate with distal margin straight or W-shaped (
<figureCitation id="138D2A72FFF9011AFF53F83552877FE9" box="[159,227,2002,2027]" captionStart="FIGURES 5 – 13" captionStartId="4.[151,264,1667,1690]" captionTargetBox="[165,1419,403,1636]" captionTargetId="figure@4.[529,1025,816,1312]" captionTargetPageId="4" captionText="FIGURES 5 – 13. Eratigena laksao sp. n., male holotype. 5. Carapace, dorsal view. 6. Abdomen, dorsal view. 7. Carapace and chelicerae, anterior view. 8. Sternum, ventral view. 9. Spinnerets, ventral view. 10. Chelicerae, posteroventral view. 11. Mouthpart, ventral view. 12. Coxa with trochanter and basal femora, ventral view. 13. Claws of leg 1, lateral view." httpUri="https://zenodo.org/record/238676/files/figure.png" pageId="3" pageNumber="434">Fig. 9</figureCitation>
; similar in
<taxonomicName id="4CB64D74FFF9011AFEACF83453B47FE9" box="[352,464,2003,2026]" class="Arachnida" family="Agelenidae" genus="Tegenaria" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="434" phylum="Arthropoda" rank="genus">
<emphasis id="B9C2EAE5FFF9011AFEACF83453B47FE9" box="[352,464,2003,2026]" italics="true" pageId="3" pageNumber="434">Tegenaria</emphasis>
</taxonomicName>
, strongly reduced in
<taxonomicName id="4CB64D74FFF9011AFD77F83551BD7FE8" authority="Keyserling, 1862" authorityName="Keyserling" authorityYear="1862" box="[699,985,2002,2027]" class="Arachnida" family="Agelenidae" genus="Hadites" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="434" phylum="Arthropoda" rank="genus">
<emphasis id="B9C2EAE5FFF9011AFD77F83551767FE8" box="[699,786,2002,2027]" italics="true" pageId="3" pageNumber="434">Hadites</emphasis>
Keyserling, 1862
</taxonomicName>
and
<taxonomicName id="4CB64D74FFF9011AFBC2F83556EB7FE8" box="[1038,1167,2002,2027]" class="Arachnida" family="Agelenidae" genus="Malthonica" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="434" phylum="Arthropoda" rank="genus">
<emphasis id="B9C2EAE5FFF9011AFBC2F83556EB7FE8" box="[1038,1167,2002,2027]" italics="true" pageId="3" pageNumber="434">Malthonica</emphasis>
</taxonomicName>
; two separated plates in all other European agelenids). Male genitalia are similar to those of
<taxonomicName id="4CB64D74FFFE011DFC6CFF7E567578B3" box="[928,1041,153,176]" class="Arachnida" family="Agelenidae" genus="Tegenaria" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="435" phylum="Arthropoda" rank="genus">
<emphasis id="B9C2EAE5FFFE011DFC6CFF7E567578B3" box="[928,1041,153,176]" italics="true" pageId="4" pageNumber="435">Tegenaria</emphasis>
</taxonomicName>
specimens, but differ in having an RTA with one or two branches only (
<figureCitation id="138D2A72FFFE011DFDF9FF5B50D278D6" box="[565,694,188,213]" captionStart="FIGURES 14 – 15" captionStartId="5.[151,264,1922,1945]" captionTargetBox="[161,1423,519,1898]" captionTargetId="figure@5.[161,922,519,1898]" captionTargetPageId="5" captionText="FIGURES 14 – 15. Eratigena laksao sp. n. 14. Left palp in ventral view (male holotype). 15. Same, retrolateral view. Abbreviations: Br = bristles; C = conductor; dRTA = dorsal branch of RTA; E = embolus; MA = median apophysis; R = membranous ridge; RTA = retrolateral tibial apophysis; vRTA = ventral branch of RTA." httpUri="https://zenodo.org/record/238677/files/figure.png" pageId="4" pageNumber="435">Figs 14–15</figureCitation>
,
<figureCitation id="138D2A72FFFE011DFD0EFF5B515178D7" box="[706,821,188,212]" captionStart="FIGURES 16 – 21" captionStartId="6.[151,264,1374,1397]" captionTargetBox="[150,1434,159,1338]" captionTargetId="figure@6.[602,1024,159,783]" captionTargetPageId="6" captionText="FIGURES 16 – 21. Eratigena laksao sp. n., male holotype. 16. Left palp, prolateral view. 17. Same, ventral view. 18. Same, retrolateral view. 19. Bulb of left palp, ventral view. 20. Same, retrolateral view. 21. Palpal tibia, dorsal view." httpUri="https://zenodo.org/record/238678/files/figure.png" pageId="4" pageNumber="435">17–18, 21</figureCitation>
; rather than more complex in most
<taxonomicName id="4CB64D74FFFE011DFB0FFF5A575578D7" box="[1219,1329,189,212]" class="Arachnida" family="Agelenidae" genus="Tegenaria" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="435" phylum="Arthropoda" rank="genus">
<emphasis id="B9C2EAE5FFFE011DFB0FFF5A575578D7" box="[1219,1329,189,212]" italics="true" pageId="4" pageNumber="435">Tegenaria</emphasis>
</taxonomicName>
species), a typically massive conductor with a membranous transverse ridge (
<figureCitation id="138D2A72FFFE011DFC19FF38563F78FB" box="[981,1115,223,248]" captionStart="FIGURES 14 – 15" captionStartId="5.[151,264,1922,1945]" captionTargetBox="[161,1423,519,1898]" captionTargetId="figure@5.[161,922,519,1898]" captionTargetPageId="5" captionText="FIGURES 14 – 15. Eratigena laksao sp. n. 14. Left palp in ventral view (male holotype). 15. Same, retrolateral view. Abbreviations: Br = bristles; C = conductor; dRTA = dorsal branch of RTA; E = embolus; MA = median apophysis; R = membranous ridge; RTA = retrolateral tibial apophysis; vRTA = ventral branch of RTA." httpUri="https://zenodo.org/record/238677/files/figure.png" pageId="4" pageNumber="435">Figs 14–15</figureCitation>
(R), 17–20), and an only moderately elongated median apophysis with distal plate-like sclerite or without sclerite (
<figureCitation id="138D2A72FFFE011DFBB0FEE3569E791E" box="[1148,1274,260,285]" captionStart="FIGURES 14 – 15" captionStartId="5.[151,264,1922,1945]" captionTargetBox="[161,1423,519,1898]" captionTargetId="figure@5.[161,922,519,1898]" captionTargetPageId="5" captionText="FIGURES 14 – 15. Eratigena laksao sp. n. 14. Left palp in ventral view (male holotype). 15. Same, retrolateral view. Abbreviations: Br = bristles; C = conductor; dRTA = dorsal branch of RTA; E = embolus; MA = median apophysis; R = membranous ridge; RTA = retrolateral tibial apophysis; vRTA = ventral branch of RTA." httpUri="https://zenodo.org/record/238677/files/figure.png" pageId="4" pageNumber="435">Figs 14–15</figureCitation>
(MA), 17–20; sclerite also absent in
<taxonomicName id="4CB64D74FFFE011DFE5EFEC0538A7943" box="[402,494,295,320]" class="Arachnida" family="Agelenidae" genus="Agelena" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="435" phylum="Arthropoda" rank="genus">
<emphasis id="B9C2EAE5FFFE011DFE5EFEC0538A7943" box="[402,494,295,320]" italics="true" pageId="4" pageNumber="435">Agelena</emphasis>
</taxonomicName>
,
<taxonomicName id="4CB64D74FFFE011DFE30FEC0501F7943" box="[508,635,295,320]" class="Arachnida" family="Agelenidae" genus="Agelescape" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="435" phylum="Arthropoda" rank="genus">
<emphasis id="B9C2EAE5FFFE011DFE30FEC0501F7943" box="[508,635,295,320]" italics="true" pageId="4" pageNumber="435">Agelescape</emphasis>
</taxonomicName>
, and
<taxonomicName id="4CB64D74FFFE011DFD77FECE517E7943" box="[699,794,297,320]" class="Arachnida" family="Agelenidae" genus="Benoitia" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="435" phylum="Arthropoda" rank="genus">
<emphasis id="B9C2EAE5FFFE011DFD77FECE517E7943" box="[699,794,297,320]" italics="true" pageId="4" pageNumber="435">Benoitia</emphasis>
</taxonomicName>
; MA absent in
<taxonomicName id="4CB64D74FFFE011DFC00FECE56257943" box="[972,1089,297,320]" class="Arachnida" family="Agelenidae" genus="Histopona" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="435" phylum="Arthropoda" rank="genus">
<emphasis id="B9C2EAE5FFFE011DFC00FECE56257943" box="[972,1089,297,320]" italics="true" pageId="4" pageNumber="435">Histopona</emphasis>
</taxonomicName>
and
<taxonomicName id="4CB64D74FFFE011DFBB5FECE56A17943" box="[1145,1221,297,320]" class="Arachnida" family="Agelenidae" genus="Textrix" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="435" phylum="Arthropoda" rank="genus">
<emphasis id="B9C2EAE5FFFE011DFBB5FECE56A17943" box="[1145,1221,297,320]" italics="true" pageId="4" pageNumber="435">Textrix</emphasis>
</taxonomicName>
). See also
<bibRefCitation id="EF274B06FFFE011DFA8FFEC053767967" author="Bolzern" pageId="4" pageNumber="435" refString="Bolzern, A., Burckhardt, D. &amp; Hanggi, A. (2013) Phylogeny and taxonomy of European funnel-web spiders of the Tegenaria- Malthonica complex (Araneae: Agelenidae) based upon morphological and molecular data. Zoological Journal of the Linnean Society, 168, 723 - 848. http: // dx. doi. org / 10.1111 / zoj. 12040" type="journal article" year="2013">
Bolzern
<emphasis id="B9C2EAE5FFFE011DFF5BFEAA52AB7966" box="[151,207,332,357]" italics="true" pageId="4" pageNumber="435">et al.</emphasis>
2013
</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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