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<document id="A1580B5CD634CE4216A2834E4E11A48A" ID-DOI="10.11646/zootaxa.3613.3.6" ID-GBIF-Dataset="a8dbc3ea-8d40-4f46-ba18-8ab5a78cb5a1" ID-ISSN="1175-5326" ID-Zenodo-Dep="215765" ID-ZooBank="9FFCFCBC-100D-4CD0-ADAF-BCB51190AF11" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460573934155" checkinUser="plazi" docAuthor="Bitner, Maria Aleksandra, Melnik, Vjacheslav P. &amp; Zezina, Olga N." docDate="2013" docId="F3166866A5359B49FF43F9125A19FB4E" docLanguage="en" docName="zt03613p288.pdf" docOrigin="Zootaxa 3613 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Simpliciforma profunda Bitner &amp; Zezina" docType="treatment" docVersion="9" lastPageNumber="287" masterDocId="0F2F101EA5319B4FFFD4FFB35E2EFF8B" masterDocTitle="New paedomorphic brachiopods from the abyssal zone of the north-eastern Pacific Ocean" masterLastPageNumber="288" masterPageNumber="281" pageNumber="285" updateTime="1698322770872" updateUser="plazi">
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<mods:title id="0E19156C0B1AD66FD027209BB83055B4">New paedomorphic brachiopods from the abyssal zone of the north-eastern Pacific Ocean</mods:title>
</mods:titleInfo>
<mods:name id="329F475DA46804B6199B0D8CE215765D" type="personal">
<mods:role id="C41D3DA08ADAF60B80FACCF8EEAA8378">
<mods:roleTerm id="1BE4BF76FF551E55962486E434C56DAD">Author</mods:roleTerm>
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<mods:namePart id="107C98142865646EAD34ABBC39E7C09C">Bitner, Maria Aleksandra</mods:namePart>
</mods:name>
<mods:name id="C324046FD813A709F25A097C9CB3911A" type="personal">
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<mods:namePart id="4D65E35016E89DB0879BE4564AA001F1">Melnik, Vjacheslav P.</mods:namePart>
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<mods:name id="A8757278B73C49BAB094C4532906523C" type="personal">
<mods:role id="6B13959C6740BF6B2CA2D9F88D731DCF">
<mods:roleTerm id="2226B6C7800F65AAFE75FF29D78F1ED1">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="73D53555FD012E7C86B2BEAE379ECC7D">Zezina, Olga N.</mods:namePart>
</mods:name>
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<mods:title id="7311A9BB902DDF4128D1C283955E67FD">Zootaxa</mods:title>
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<mods:part id="2BE200896ECDF9B46FC1BC5CA747924E">
<mods:date id="233139DBF84BFEB5570479AD77551C4E">2013</mods:date>
<mods:detail id="D13515762489037C2BAEA4FF88F9B21B" type="volume">
<mods:number id="86BFBC8E0FB00AF6B7106D37FAE8EBA7">3613</mods:number>
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<mods:number id="87ADAC12D2CF14F0E475E215FE18A3CC">3</mods:number>
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<mods:classification id="E4ECBEEC771791CB0AFA98D3D558BB26">journal article</mods:classification>
<mods:identifier id="5C5504FE5734326CA0BACE5D5CA4FA7F" type="DOI">10.11646/zootaxa.3613.3.6</mods:identifier>
<mods:identifier id="B37BB9522CAB5109AB494D4824C7D3AD" type="GBIF-Dataset">a8dbc3ea-8d40-4f46-ba18-8ab5a78cb5a1</mods:identifier>
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<mods:identifier id="23412D9392D1AF933E9FB898801FCC44" type="Zenodo-Dep">215765</mods:identifier>
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<treatment id="F3166866A5359B49FF43F9125A19FB4E" ID-DOI="http://doi.org/10.5281/zenodo.5657855" ID-GBIF-Taxon="119559385" ID-Zenodo-Dep="5657855" LSID="urn:lsid:plazi:treatment:F3166866A5359B49FF43F9125A19FB4E" httpUri="http://treatment.plazi.org/id/F3166866A5359B49FF43F9125A19FB4E" lastPageId="6" lastPageNumber="287" pageId="4" pageNumber="285">
<subSubSection id="33A58AFBA5359B4BFF43F9125F39F957" pageId="4" pageNumber="285" type="nomenclature">
<paragraph id="7B00D970A5359B4BFF43F9125D7DF931" blockId="4.[151,851,1696,1756]" box="[151,851,1696,1723]" pageId="4" pageNumber="285">
<heading id="20486E1CA5359B4BFF43F9125D7DF931" bold="true" box="[151,851,1696,1723]" fontSize="11" level="1" pageId="4" pageNumber="285" reason="1">
<emphasis id="49CB0562A5359B4BFF43F9125D7DF931" bold="true" box="[151,851,1696,1723]" pageId="4" pageNumber="285">
<taxonomicName id="BCBFA2F3A5359B4BFF43F9125CA3F931" ID-CoL="6YRHZ" authority="Bitner &amp; Zezina" authorityName="Bitner &amp; Zezina" box="[151,653,1696,1723]" class="Rhynchonellata" genus="Simpliciforma" kingdom="Animalia" order="Terebratulida" pageId="4" pageNumber="285" phylum="Brachiopoda" rank="species" species="profunda">
<emphasis id="49CB0562A5359B4BFF43F9125F92F930" bold="true" box="[151,444,1697,1723]" italics="true" pageId="4" pageNumber="285">Simpliciforma profunda</emphasis>
Bitner &amp; Zezina
</taxonomicName>
gen. et sp. nov.
</emphasis>
</heading>
</paragraph>
<paragraph id="7B00D970A5359B4BFF43F9775F39F957" blockId="4.[151,851,1696,1756]" box="[151,279,1732,1756]" pageId="4" pageNumber="285">
(
<figureCitation id="E384C5F5A5359B4BFF4BF9775ECCF957" box="[159,226,1732,1756]" captionStart="FIGURE 4" captionStartId="5.[151,250,1772,1795]" captionTargetBox="[151,1435,501,1750]" captionTargetId="figure@5.[151,1436,501,1752]" captionTargetPageId="5" captionText="FIGURE 4. Simpliciforma profunda Bitner &amp; Zezina gen. et sp. nov., Clarion-Clipperton Zone, north-eastern Pacific; A, dorsal view of complete specimen, paratype, XI- 52 - 24 / 2, Stn 27, depth 4850 m; B D, dorsal valve, inner view and enlargement of the lophophore (C, D), paratype, XI- 52 - 24 / 3, Stn 185 199, 4580 m; E, F, dorsal view of complete specimen, and enlargement of the umbonal part to show details of the beak, holotype, XI- 52 - 24 / 1, Stn 217, 4680 m; G I, dorsal view of complete specimen, and enlargement of the umbonal part, slightly tilted, to show details of the beak (H) and of the shell surface to show distinct growth lines (I), paratype, XI- 52 - 24 / 4, Stn 185 - 199, 4580 m." httpUri="https://zenodo.org/record/215769/files/figure.png" pageId="4" pageNumber="285">Fig. 4</figureCitation>
AI)
</paragraph>
</subSubSection>
<subSubSection id="33A58AFBA5359B4BFF43F8B85FF6F8AE" box="[151,472,1803,1829]" pageId="4" pageNumber="285" type="diagnosis">
<paragraph id="7B00D970A5359B4BFF43F8B85FF6F8AE" blockId="4.[151,1436,1803,2036]" box="[151,472,1803,1829]" pageId="4" pageNumber="285">
<emphasis id="49CB0562A5359B4BFF43F8B85F3FF8AF" bold="true" box="[151,273,1803,1828]" pageId="4" pageNumber="285">Diagnosis.</emphasis>
<collectingCountry id="03A899E0A5359B4BFECCF8BF5F19F8AF" box="[280,311,1804,1828]" name="American Samoa" pageId="4" pageNumber="285">As</collectingCountry>
for the genus.
</paragraph>
</subSubSection>
<subSubSection id="33A58AFBA5359B4BFF1DF89C5F73F87F" pageId="4" pageNumber="285" type="etymology">
<paragraph id="7B00D970A5359B4BFF1DF89C5F73F87F" blockId="4.[151,1436,1803,2036]" pageId="4" pageNumber="285">
<emphasis id="49CB0562A5359B4BFF1DF89C5F61F8CC" bold="true" box="[201,335,1839,1863]" pageId="4" pageNumber="285">Etymology.</emphasis>
Referring to the great depth at which this brachiopod was found.
<emphasis id="49CB0562A5359B4BFF1DF8E15F4BF8E1" bold="true" box="[201,357,1873,1898]" pageId="4" pageNumber="285">
<typeStatus id="A40467D2A5359B4BFF1DF8E15F2CF8E1" box="[201,258,1874,1898]" pageId="4" pageNumber="285">Type</typeStatus>
locality.
</emphasis>
Clarion-Clipperton Zone, north-eastern Pacific, station 217,
<geoCoordinate id="1E8BBFB7A5359B4BFBC5F8E25A5CF8E2" box="[1041,1138,1873,1897]" direction="north" orientation="latitude" pageId="4" pageNumber="285" precision="555" value="13.91">13.91°N</geoCoordinate>
,
<geoCoordinate id="1E8BBFB7A5359B4BFBAAF8E25AC1F8E2" box="[1150,1263,1873,1897]" direction="west" orientation="longitude" pageId="4" pageNumber="285" precision="555" value="-129.11">129.11°W</geoCoordinate>
, depth
<quantity id="BC477495A5359B4BFA94F8E25BB6F8E2" box="[1344,1432,1873,1897]" metricMagnitude="3" metricUnit="m" metricValue="4.68" pageId="4" pageNumber="285" unit="m" value="4680.0">4680 m</quantity>
.
<emphasis id="49CB0562A5359B4BFF1DF8C75F13F807" bold="true" box="[201,317,1908,1932]" pageId="4" pageNumber="285">
<typeStatus id="A40467D2A5359B4BFF1DF8C75F17F807" box="[201,313,1908,1932]" pageId="4" pageNumber="285" type="holotype">Holotype</typeStatus>
.
</emphasis>
Specimen in
<figureCitation id="E384C5F5A5359B4BFE01F8C75C36F807" box="[469,536,1908,1932]" captionStart="FIGURE 4" captionStartId="5.[151,250,1772,1795]" captionTargetBox="[151,1435,501,1750]" captionTargetId="figure@5.[151,1436,501,1752]" captionTargetPageId="5" captionText="FIGURE 4. Simpliciforma profunda Bitner &amp; Zezina gen. et sp. nov., Clarion-Clipperton Zone, north-eastern Pacific; A, dorsal view of complete specimen, paratype, XI- 52 - 24 / 2, Stn 27, depth 4850 m; B D, dorsal valve, inner view and enlargement of the lophophore (C, D), paratype, XI- 52 - 24 / 3, Stn 185 199, 4580 m; E, F, dorsal view of complete specimen, and enlargement of the umbonal part to show details of the beak, holotype, XI- 52 - 24 / 1, Stn 217, 4680 m; G I, dorsal view of complete specimen, and enlargement of the umbonal part, slightly tilted, to show details of the beak (H) and of the shell surface to show distinct growth lines (I), paratype, XI- 52 - 24 / 4, Stn 185 - 199, 4580 m." httpUri="https://zenodo.org/record/215769/files/figure.png" pageId="4" pageNumber="285">Fig. 4</figureCitation>
E, F (
<date id="0F01FFB0A5359B4BFD81F8C75CEBF807" box="[597,709,1908,1932]" pageId="4" pageNumber="285" value="1952-11-24">XI-52-24</date>
/1), collected in 2003, station 217.
<emphasis id="49CB0562A5359B4BFF1DF8245F66F824" bold="true" box="[201,328,1943,1967]" pageId="4" pageNumber="285">
<typeStatus id="A40467D2A5359B4BFF1DF8245F6DF824" box="[201,323,1943,1967]" pageId="4" pageNumber="285" type="paratype">Paratypes</typeStatus>
.
</emphasis>
Specimen in
<figureCitation id="E384C5F5A5359B4BFE0BF8255C0EF825" box="[479,544,1942,1966]" captionStart="FIGURE 4" captionStartId="5.[151,250,1772,1795]" captionTargetBox="[151,1435,501,1750]" captionTargetId="figure@5.[151,1436,501,1752]" captionTargetPageId="5" captionText="FIGURE 4. Simpliciforma profunda Bitner &amp; Zezina gen. et sp. nov., Clarion-Clipperton Zone, north-eastern Pacific; A, dorsal view of complete specimen, paratype, XI- 52 - 24 / 2, Stn 27, depth 4850 m; B D, dorsal valve, inner view and enlargement of the lophophore (C, D), paratype, XI- 52 - 24 / 3, Stn 185 199, 4580 m; E, F, dorsal view of complete specimen, and enlargement of the umbonal part to show details of the beak, holotype, XI- 52 - 24 / 1, Stn 217, 4680 m; G I, dorsal view of complete specimen, and enlargement of the umbonal part, slightly tilted, to show details of the beak (H) and of the shell surface to show distinct growth lines (I), paratype, XI- 52 - 24 / 4, Stn 185 - 199, 4580 m." httpUri="https://zenodo.org/record/215769/files/figure.png" pageId="4" pageNumber="285">Fig. 4</figureCitation>
A (
<date id="0F01FFB0A5359B4BFD97F8255C81F825" box="[579,687,1942,1966]" pageId="4" pageNumber="285" value="1952-11-24">XI-52-24</date>
/2), collected in 2000, station 27; specimen in
<figureCitation id="E384C5F5A5359B4BFB7BF8255ADEF825" box="[1199,1264,1942,1966]" captionStart="FIGURE 4" captionStartId="5.[151,250,1772,1795]" captionTargetBox="[151,1435,501,1750]" captionTargetId="figure@5.[151,1436,501,1752]" captionTargetPageId="5" captionText="FIGURE 4. Simpliciforma profunda Bitner &amp; Zezina gen. et sp. nov., Clarion-Clipperton Zone, north-eastern Pacific; A, dorsal view of complete specimen, paratype, XI- 52 - 24 / 2, Stn 27, depth 4850 m; B D, dorsal valve, inner view and enlargement of the lophophore (C, D), paratype, XI- 52 - 24 / 3, Stn 185 199, 4580 m; E, F, dorsal view of complete specimen, and enlargement of the umbonal part to show details of the beak, holotype, XI- 52 - 24 / 1, Stn 217, 4680 m; G I, dorsal view of complete specimen, and enlargement of the umbonal part, slightly tilted, to show details of the beak (H) and of the shell surface to show distinct growth lines (I), paratype, XI- 52 - 24 / 4, Stn 185 - 199, 4580 m." httpUri="https://zenodo.org/record/215769/files/figure.png" pageId="4" pageNumber="285">Fig. 4</figureCitation>
GI (
<date id="0F01FFB0A5359B4BFAFEF8255BBDF825" box="[1322,1427,1942,1966]" pageId="4" pageNumber="285" value="1952-11-24">XI-52-24</date>
/ 4), collected in 2003, stations 185199; dorsal valve with preserved lophophore in
<figureCitation id="E384C5F5A5359B4BFBAFF80A5AEDF85A" box="[1147,1219,1977,2001]" captionStart="FIGURE 4" captionStartId="5.[151,250,1772,1795]" captionTargetBox="[151,1435,501,1750]" captionTargetId="figure@5.[151,1436,501,1752]" captionTargetPageId="5" captionText="FIGURE 4. Simpliciforma profunda Bitner &amp; Zezina gen. et sp. nov., Clarion-Clipperton Zone, north-eastern Pacific; A, dorsal view of complete specimen, paratype, XI- 52 - 24 / 2, Stn 27, depth 4850 m; B D, dorsal valve, inner view and enlargement of the lophophore (C, D), paratype, XI- 52 - 24 / 3, Stn 185 199, 4580 m; E, F, dorsal view of complete specimen, and enlargement of the umbonal part to show details of the beak, holotype, XI- 52 - 24 / 1, Stn 217, 4680 m; G I, dorsal view of complete specimen, and enlargement of the umbonal part, slightly tilted, to show details of the beak (H) and of the shell surface to show distinct growth lines (I), paratype, XI- 52 - 24 / 4, Stn 185 - 199, 4580 m." httpUri="https://zenodo.org/record/215769/files/figure.png" pageId="4" pageNumber="285">Fig. 4</figureCitation>
BD (
<date id="0F01FFB0A5359B4BFAD8F80A5B54F85A" box="[1292,1402,1977,2001]" pageId="4" pageNumber="285" value="1952-11-24">XI-52-24</date>
/3), stations 185199.
</paragraph>
</subSubSection>
<paragraph id="7B00D970A5349B4AFF1DFF245FF8FF7C" pageId="5" pageNumber="286">
<table id="09BF2BD0A53464B0FF1DFF245A6EFF7C" box="[201,1088,151,247]" gridcols="6" gridrows="3" pageId="5" pageNumber="286">
<tr id="C58FDB32A53464B0FF1DFF245A6EFF3B" box="[201,1088,151,176]" gridrow="0" pageId="5" pageNumber="286">
<th id="865EB24EA53464B0FF1DFF245C1DFF3B" box="[201,563,151,176]" gridcol="0" gridrow="0" pageId="5" pageNumber="286">
<emphasis id="49CB0562A5349B4AFF1DFF245FD0FF3B" bold="true" box="[201,510,151,176]" pageId="5" pageNumber="286">Other material examined.</emphasis>
One
</th>
<th id="865EB24EA53464B0FDEEFF245C8AFF3B" box="[570,676,151,176]" gridcol="1" gridrow="0" pageId="5" pageNumber="286">specimen</th>
<th id="865EB24EA53464B0FD7FFF245D3EFF3B" box="[683,784,151,176]" gridcol="2" gridrow="0" pageId="5" pageNumber="286">collected</th>
<th id="865EB24EA53464B0FCC3FF245D03FF3B" box="[791,813,151,176]" gridcol="3" gridrow="0" pageId="5" pageNumber="286">in</th>
<th id="865EB24EA53464B0FCE0FF245D5CFF3B" box="[820,882,151,176]" gridcol="4" gridrow="0" pageId="5" pageNumber="286">2003,</th>
<th id="865EB24EA53464B0FCAEFF245A6EFF3B" box="[890,1088,151,176]" gridcol="5" gridrow="0" pageId="5" pageNumber="286">stations 185199.</th>
</tr>
<tr id="C58FDB32A53464B0FF1DFF0F5A6EFF5F" box="[201,1088,188,212]" gridrow="1" pageId="5" pageNumber="286" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<th id="865EB24EA53464B0FF1DFF0F5C1DFF5F" box="[201,563,188,212]" gridcol="0" gridrow="1" pageId="5" pageNumber="286">
<emphasis id="49CB0562A5349B4AFF1DFF0F5F4BFF5F" bold="true" box="[201,357,188,212]" pageId="5" pageNumber="286">Depth range.</emphasis>
45804850 m.
</th>
</tr>
<tr id="C58FDB32A53464B0FF1DFF6C5A6EFF7C" box="[201,1088,223,247]" gridrow="2" pageId="5" pageNumber="286" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<th id="865EB24EA53464B0FF1DFF6C5C1DFF7C" box="[201,563,223,247]" gridcol="0" gridrow="2" pageId="5" pageNumber="286">Measurements (in mm).</th>
</tr>
</table>
</paragraph>
<paragraph id="7B00D970A5349B4AFF4BFEA05A95FE2E" pageId="5" pageNumber="286">
<table id="09BF2BD0A53464B0FF4BFEA05AF7FE2D" box="[159,1241,275,422]" gridcols="3" gridrows="4" pageId="5" pageNumber="286">
<tr id="C58FDB32A53464B0FF4BFEA05AF7FEA2" box="[159,1241,275,297]" gridrow="0" pageId="5" pageNumber="286">
<th id="865EB24EA53464B0FF4BFEA05FDFFEA2" box="[159,497,275,297]" gridcol="0" gridrow="0" pageId="5" pageNumber="286">Station</th>
<th id="865EB24EA53464B0FC5AFEA05DF8FEA2" box="[910,982,275,297]" gridcol="1" gridrow="0" pageId="5" pageNumber="286">Length</th>
<th id="865EB24EA53464B0FB4FFEA05AF7FEA2" box="[1179,1241,275,297]" gridcol="2" gridrow="0" pageId="5" pageNumber="286">Width</th>
</tr>
<tr id="C58FDB32A53464B0FF4BFE8E5AF7FED8" box="[159,1241,317,339]" gridrow="1" pageId="5" pageNumber="286">
<th id="865EB24EA53464B0FF4BFE8E5FDFFED8" box="[159,497,317,339]" gridcol="0" gridrow="1" pageId="5" pageNumber="286">#185199 (paratype, XI-52-24/4)</th>
<td id="865EB24EA53464B0FC5AFE8E5DF8FED8" box="[910,982,317,339]" gridcol="1" gridrow="1" pageId="5" pageNumber="286">1.2</td>
<td id="865EB24EA53464B0FB4FFE8E5AF7FED8" box="[1179,1241,317,339]" gridcol="2" gridrow="1" pageId="5" pageNumber="286">1.0</td>
</tr>
<tr id="C58FDB32A53464B0FF4BFED55AF7FEF7" box="[159,1241,358,380]" gridrow="2" pageId="5" pageNumber="286">
<th id="865EB24EA53464B0FF4BFED55FDFFEF7" box="[159,497,358,380]" gridcol="0" gridrow="2" pageId="5" pageNumber="286">#217 (holotype, (XI-52-24/1)</th>
<td id="865EB24EA53464B0FC5AFED55DF8FEF7" box="[910,982,358,380]" gridcol="1" gridrow="2" pageId="5" pageNumber="286">1.4</td>
<td id="865EB24EA53464B0FB4FFED55AF7FEF7" box="[1179,1241,358,380]" gridcol="2" gridrow="2" pageId="5" pageNumber="286">1.2</td>
</tr>
<tr id="C58FDB32A53464B0FF4BFE235AF7FE2D" box="[159,1241,400,422]" gridrow="3" pageId="5" pageNumber="286">
<th id="865EB24EA53464B0FF4BFE235FDFFE2D" box="[159,497,400,422]" gridcol="0" gridrow="3" pageId="5" pageNumber="286">#27 (paratype, XI-52-24/2)</th>
<td id="865EB24EA53464B0FC5AFE235DF8FE2D" box="[910,982,400,422]" gridcol="1" gridrow="3" pageId="5" pageNumber="286">1.0</td>
<td id="865EB24EA53464B0FB4FFE235AF7FE2D" box="[1179,1241,400,422]" gridcol="2" gridrow="3" pageId="5" pageNumber="286">0.9</td>
</tr>
</table>
</paragraph>
<caption id="2FC089F8A5349B4AFF43F95F5D88F817" httpUri="https://zenodo.org/record/215769/files/figure.png" pageId="5" pageNumber="286" targetBox="[151,1435,501,1750]" targetPageId="5">
<paragraph id="7B00D970A5349B4AFF43F95F5D88F817" blockId="5.[151,1436,1772,1949]" pageId="5" pageNumber="286">
<emphasis id="49CB0562A5349B4AFF43F95F5F3BF889" bold="true" box="[151,277,1772,1795]" pageId="5" pageNumber="286">FIGURE 4.</emphasis>
<taxonomicName id="BCBFA2F3A5349B4AFECBF95E5CE9F888" authority="Bitner &amp; Zezina" authorityName="Bitner &amp; Zezina" box="[287,711,1773,1795]" class="Rhynchonellata" genus="Simpliciforma" kingdom="Animalia" order="Terebratulida" pageId="5" pageNumber="286" phylum="Brachiopoda" rank="species" species="profunda">
<emphasis id="49CB0562A5349B4AFECBF95E5C3AF888" box="[287,532,1773,1795]" italics="true" pageId="5" pageNumber="286">Simpliciforma profunda</emphasis>
Bitner &amp; Zezina
</taxonomicName>
<emphasis id="49CB0562A5349B4AFD05F95E5DAEF889" bold="true" box="[721,896,1773,1794]" pageId="5" pageNumber="286">gen. et sp. nov.</emphasis>
, Clarion-Clipperton Zone, north-eastern Pacific; A, dorsal view of complete specimen, paratype, XI-52-24/2, Stn 27, depth 4850 m; BD, dorsal valve, inner view and enlargement of the lophophore (C, D), paratype, XI-52-24/3, Stn 185199, 4580 m; E, F, dorsal view of complete specimen, and enlargement of the umbonal part to show details of the beak, holotype, XI-52-24/1, Stn 217, 4680 m; GI, dorsal view of complete specimen, and enlargement of the umbonal part, slightly tilted, to show details of the beak (H) and of the shell surface to show distinct growth lines (I), paratype, XI-52-24/4, Stn 185-199, 4580 m.
</paragraph>
</caption>
<subSubSection id="33A58AFBA5379B49FF13FF245CB1FD9C" pageId="6" pageNumber="287" type="description">
<paragraph id="7B00D970A5379B49FF13FF245FFAFEEF" blockId="6.[151,1437,151,1221]" pageId="6" pageNumber="287">
<emphasis id="49CB0562A5379B49FF13FF245F77FF3B" bold="true" box="[199,345,151,176]" pageId="6" pageNumber="287">Description.</emphasis>
Shell very small (maximum observed length
<quantity id="BC477495A5379B49FC5EFF245DC5FF24" box="[906,1003,151,175]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="6" pageNumber="287" unit="mm" value="1.4">1.4 mm</quantity>
), thin, translucent, subpentagonal to subrectangular in outline with greatest width at anterior, biconvex with dorsal valve more convex. Shell surface smooth with numerous, distinct growth lines (
<figureCitation id="E384C5F5A5379B49FD1AFF6C5D35FF7C" box="[718,795,223,247]" captionStart="FIGURE 4" captionStartId="5.[151,250,1772,1795]" captionTargetBox="[151,1435,501,1750]" captionTargetId="figure@5.[151,1436,501,1752]" captionTargetPageId="5" captionText="FIGURE 4. Simpliciforma profunda Bitner &amp; Zezina gen. et sp. nov., Clarion-Clipperton Zone, north-eastern Pacific; A, dorsal view of complete specimen, paratype, XI- 52 - 24 / 2, Stn 27, depth 4850 m; B D, dorsal valve, inner view and enlargement of the lophophore (C, D), paratype, XI- 52 - 24 / 3, Stn 185 199, 4580 m; E, F, dorsal view of complete specimen, and enlargement of the umbonal part to show details of the beak, holotype, XI- 52 - 24 / 1, Stn 217, 4680 m; G I, dorsal view of complete specimen, and enlargement of the umbonal part, slightly tilted, to show details of the beak (H) and of the shell surface to show distinct growth lines (I), paratype, XI- 52 - 24 / 4, Stn 185 - 199, 4580 m." httpUri="https://zenodo.org/record/215769/files/figure.png" pageId="6" pageNumber="287">
Fig.
<date id="0F01FFB0A5379B49FCDFFF6C5D35FF7C" box="[779,795,223,247]" pageId="6" pageNumber="287">4</date>
</figureCitation>
I). Lateral commissures straight, anterior commissure rectimarginate. Hinge line long, straight. Beak short, suberect, beak ridges sharp. Interarea very narrow. Dorsal notothyrium small, ventral delthyrium large, together forming a large, subtriangular foramen (
<figureCitation id="E384C5F5A5379B49FB65FE945ADAFEB4" box="[1201,1268,295,319]" captionStart="FIGURE 4" captionStartId="5.[151,250,1772,1795]" captionTargetBox="[151,1435,501,1750]" captionTargetId="figure@5.[151,1436,501,1752]" captionTargetPageId="5" captionText="FIGURE 4. Simpliciforma profunda Bitner &amp; Zezina gen. et sp. nov., Clarion-Clipperton Zone, north-eastern Pacific; A, dorsal view of complete specimen, paratype, XI- 52 - 24 / 2, Stn 27, depth 4850 m; B D, dorsal valve, inner view and enlargement of the lophophore (C, D), paratype, XI- 52 - 24 / 3, Stn 185 199, 4580 m; E, F, dorsal view of complete specimen, and enlargement of the umbonal part to show details of the beak, holotype, XI- 52 - 24 / 1, Stn 217, 4680 m; G I, dorsal view of complete specimen, and enlargement of the umbonal part, slightly tilted, to show details of the beak (H) and of the shell surface to show distinct growth lines (I), paratype, XI- 52 - 24 / 4, Stn 185 - 199, 4580 m." httpUri="https://zenodo.org/record/215769/files/figure.png" pageId="6" pageNumber="287">Fig. 4</figureCitation>
F, H). Deltidial plates very narrow, disjunct.
</paragraph>
<paragraph id="7B00D970A5379B49FF1DFEDC5CB1FD9C" blockId="6.[151,1437,151,1221]" pageId="6" pageNumber="287">
Ventral valve only slightly larger than dorsal one with large open delthyrium. Dorsal valve interior with very low but thick, widely separated inner socket ridges that do not project beyond posterior margin. Dental sockets rounded and relatively deep. No cardinal process. No hinge plates. No brachial skeleton or median septum. Lophophore trocholophous (
<figureCitation id="E384C5F5A5379B49FE03FE6F5C35FE7F" box="[471,539,476,500]" captionStart="FIGURE 4" captionStartId="5.[151,250,1772,1795]" captionTargetBox="[151,1435,501,1750]" captionTargetId="figure@5.[151,1436,501,1752]" captionTargetPageId="5" captionText="FIGURE 4. Simpliciforma profunda Bitner &amp; Zezina gen. et sp. nov., Clarion-Clipperton Zone, north-eastern Pacific; A, dorsal view of complete specimen, paratype, XI- 52 - 24 / 2, Stn 27, depth 4850 m; B D, dorsal valve, inner view and enlargement of the lophophore (C, D), paratype, XI- 52 - 24 / 3, Stn 185 199, 4580 m; E, F, dorsal view of complete specimen, and enlargement of the umbonal part to show details of the beak, holotype, XI- 52 - 24 / 1, Stn 217, 4680 m; G I, dorsal view of complete specimen, and enlargement of the umbonal part, slightly tilted, to show details of the beak (H) and of the shell surface to show distinct growth lines (I), paratype, XI- 52 - 24 / 4, Stn 185 - 199, 4580 m." httpUri="https://zenodo.org/record/215769/files/figure.png" pageId="6" pageNumber="287">Fig. 4</figureCitation>
BD), consisting of a pair of curving arms projecting horizontally on either side of a mouth and forming a ring; filaments long.
</paragraph>
</subSubSection>
<subSubSection id="33A58AFBA5379B49FF1DFD975A19FB4E" pageId="6" pageNumber="287" type="discussion">
<paragraph id="7B00D970A5379B49FF1DFD975DC3FCBC" blockId="6.[151,1437,151,1221]" pageId="6" pageNumber="287">
<emphasis id="49CB0562A5379B49FF1DFD975F13FDB7" bold="true" box="[201,317,548,572]" pageId="6" pageNumber="287">Remarks.</emphasis>
The investigated specimens, although similar in simple morphological structure and trocholophous lophophore, are easily distinguishable from
<taxonomicName id="BCBFA2F3A5379B49FD5EFDFB5DD5FDD4" authority="Jeffreys, 1859" authorityName="Jeffreys" authorityYear="1859" box="[650,1019,583,608]" class="Rhynchonellata" genus="Gwynia" kingdom="Animalia" order="Terebratulida" pageId="6" pageNumber="287" phylum="Brachiopoda" rank="species" species="capsula">
<emphasis id="49CB0562A5379B49FD5EFDFB5D6FFDEB" box="[650,833,584,608]" italics="true" pageId="6" pageNumber="287">Gwynia capsula</emphasis>
(Jeffreys, 1859)
</taxonomicName>
known from several localities in the eastern Atlantic and Mediterranean (Brunton &amp; Curry 1979; Harper
<emphasis id="49CB0562A5379B49FC6EFDDE5DC1FD0F" box="[954,1007,620,644]" italics="true" pageId="6" pageNumber="287">et al</emphasis>
. 1996; Logan
<emphasis id="49CB0562A5379B49FB4BFDDE5AFBFD0F" box="[1183,1237,620,644]" italics="true" pageId="6" pageNumber="287">et al</emphasis>
. 1997; Simon &amp; Willems 1999).
<taxonomicName id="BCBFA2F3A5379B49FE9EFD235FEEFD23" box="[330,448,656,680]" class="Rhynchonellata" genus="Gwynia" kingdom="Animalia" order="Terebratulida" pageId="6" pageNumber="287" phylum="Brachiopoda" rank="species" species="capsula">
<emphasis id="49CB0562A5379B49FE9EFD235FEEFD23" box="[330,448,656,680]" italics="true" pageId="6" pageNumber="287">G. capsula</emphasis>
</taxonomicName>
is ovate and globose in outline, and its hinge line is slightly curved being shorter than in
<taxonomicName id="BCBFA2F3A5379B49FF43FD075F8FFD47" box="[151,417,692,716]" class="Rhynchonellata" genus="Simpliciforma" kingdom="Animalia" order="Terebratulida" pageId="6" pageNumber="287" phylum="Brachiopoda" rank="species" species="profunda">
<emphasis id="49CB0562A5379B49FF43FD075F8FFD47" box="[151,417,692,716]" italics="true" pageId="6" pageNumber="287">Simpliciforma profunda</emphasis>
</taxonomicName>
. The dorsal valve in
<taxonomicName id="BCBFA2F3A5379B49FD58FD065D2FFD47" box="[652,769,692,716]" class="Rhynchonellata" genus="Gwynia" kingdom="Animalia" order="Terebratulida" pageId="6" pageNumber="287" phylum="Brachiopoda" rank="species" species="capsula">
<emphasis id="49CB0562A5379B49FD58FD065D2FFD47" box="[652,769,692,716]" italics="true" pageId="6" pageNumber="287">G. capsula</emphasis>
</taxonomicName>
is a little larger than the ventral valve. The main difference, however, is visible internally;
<taxonomicName id="BCBFA2F3A5379B49FE2DFD6B5C5DFD7B" box="[505,627,728,752]" class="Rhynchonellata" genus="Gwynia" kingdom="Animalia" order="Terebratulida" pageId="6" pageNumber="287" phylum="Brachiopoda" rank="species" species="capsula">
<emphasis id="49CB0562A5379B49FE2DFD6B5C5DFD7B" box="[505,627,728,752]" italics="true" pageId="6" pageNumber="287">G. capsula</emphasis>
</taxonomicName>
possesses two postero-lateral submarginal ridges (Logan
<emphasis id="49CB0562A5379B49FAC1FD6A5B64FD7B" box="[1301,1354,728,752]" italics="true" pageId="6" pageNumber="287">et al</emphasis>
. 1997; Simon &amp; Willems 1999) that are not observed in
<taxonomicName id="BCBFA2F3A5379B49FD04FD4E5D7AFC9F" box="[720,852,764,788]" class="Rhynchonellata" genus="Simpliciforma" kingdom="Animalia" order="Terebratulida" pageId="6" pageNumber="287" phylum="Brachiopoda" rank="species" species="profunda">
<emphasis id="49CB0562A5379B49FD04FD4E5D7AFC9F" box="[720,852,764,788]" italics="true" pageId="6" pageNumber="287">S. profunda</emphasis>
</taxonomicName>
. Also
<taxonomicName id="BCBFA2F3A5379B49FC75FD4E5A37FC9F" box="[929,1049,764,788]" class="Rhynchonellata" genus="Gwynia" kingdom="Animalia" order="Terebratulida" pageId="6" pageNumber="287" phylum="Brachiopoda" rank="species" species="capsula">
<emphasis id="49CB0562A5379B49FC75FD4E5A37FC9F" box="[929,1049,764,788]" italics="true" pageId="6" pageNumber="287">G. capsula</emphasis>
</taxonomicName>
has a large amphithyrid foramen, while in
<taxonomicName id="BCBFA2F3A5379B49FF2DFC935F55FCB3" box="[249,379,800,824]" class="Rhynchonellata" genus="Simpliciforma" kingdom="Animalia" order="Terebratulida" pageId="6" pageNumber="287" phylum="Brachiopoda" rank="species" species="profunda">
<emphasis id="49CB0562A5379B49FF2DFC935F55FCB3" box="[249,379,800,824]" italics="true" pageId="6" pageNumber="287">S. profunda</emphasis>
</taxonomicName>
the foramen is only slightly marked in the dorsal valve.
</paragraph>
<paragraph id="7B00D970A5379B49FF1DFCF65AF2FB9B" blockId="6.[151,1437,151,1221]" pageId="6" pageNumber="287">
In shell outline, straight hinge line and absence of dorsal submarginal ridges, these specimens are similar to the specimen from the Chatham Rise,
<collectingCountry id="03A899E0A5379B49FDE7FCDA5CFFFCF4" box="[563,721,871,896]" name="New Zealand" pageId="6" pageNumber="287">New Zealand</collectingCountry>
, described by Lüter (2008) as
<taxonomicName id="BCBFA2F3A5379B49FB90FCDB5B6FFC0B" box="[1092,1345,872,896]" class="Rhynchonellata" genus="Gwynia" kingdom="Animalia" order="Terebratulida" pageId="6" pageNumber="287" phylum="Brachiopoda" rank="species" species="macrodentata">
<emphasis id="49CB0562A5379B49FB90FCDB5B6FFC0B" box="[1092,1345,872,896]" italics="true" pageId="6" pageNumber="287">Gwynia macrodentata</emphasis>
</taxonomicName>
, but
<taxonomicName id="BCBFA2F3A5379B49FA50FCDB5F1CFC2F" class="Rhynchonellata" genus="Gwynia" kingdom="Animalia" order="Terebratulida" pageId="6" pageNumber="287" phylum="Brachiopoda" rank="species" species="macrodentata">
<emphasis id="49CB0562A5379B49FA50FCDB5F1CFC2F" italics="true" pageId="6" pageNumber="287">G. macrodentata</emphasis>
</taxonomicName>
differs from
<taxonomicName id="BCBFA2F3A5379B49FE1DFC3E5C62FC2F" box="[457,588,908,932]" class="Rhynchonellata" genus="Simpliciforma" kingdom="Animalia" order="Terebratulida" pageId="6" pageNumber="287" phylum="Brachiopoda" rank="species" species="profunda">
<emphasis id="49CB0562A5379B49FE1DFC3E5C62FC2F" box="[457,588,908,932]" italics="true" pageId="6" pageNumber="287">S. profunda</emphasis>
</taxonomicName>
in having thicker, closely spaced, inner socket ridges (see Lüter 2008, fig. 3B). In addition, dental sockets observed in
<taxonomicName id="BCBFA2F3A5379B49FD4EFC035D31FC43" box="[666,799,944,968]" class="Rhynchonellata" genus="Simpliciforma" kingdom="Animalia" order="Terebratulida" pageId="6" pageNumber="287" phylum="Brachiopoda" rank="species" species="profunda">
<emphasis id="49CB0562A5379B49FD4EFC035D31FC43" box="[666,799,944,968]" italics="true" pageId="6" pageNumber="287">S. profunda</emphasis>
</taxonomicName>
suggest that its teeth are less massive than those in
<taxonomicName id="BCBFA2F3A5379B49FA50FC035F1CFC67" class="Rhynchonellata" genus="Gwynia" kingdom="Animalia" order="Terebratulida" pageId="6" pageNumber="287" phylum="Brachiopoda" rank="species" species="macrodentata">
<emphasis id="49CB0562A5379B49FA50FC035F1CFC67" italics="true" pageId="6" pageNumber="287">G. macrodentata</emphasis>
</taxonomicName>
.
<collectingCountry id="03A899E0A5379B49FE94FC675F71FC67" box="[320,351,980,1004]" name="American Samoa" pageId="6" pageNumber="287">As</collectingCountry>
submarginal ridges in the dorsal valve are considered a diagnostic feature of the genus
<taxonomicName id="BCBFA2F3A5379B49FAEAFC665BBBFC67" box="[1342,1429,981,1004]" class="Rhynchonellata" genus="Gwynia" kingdom="Animalia" order="Terebratulida" pageId="6" pageNumber="287" phylum="Brachiopoda" rank="genus">
<emphasis id="49CB0562A5379B49FAEAFC665BBBFC67" box="[1342,1429,981,1004]" italics="true" pageId="6" pageNumber="287">Gwynia</emphasis>
</taxonomicName>
, but are absent from
<taxonomicName id="BCBFA2F3A5379B49FEA3FC4B5C01FB9B" box="[375,559,1016,1040]" class="Rhynchonellata" genus="Gwynia" kingdom="Animalia" order="Terebratulida" pageId="6" pageNumber="287" phylum="Brachiopoda" rank="species" species="macrodentata">
<emphasis id="49CB0562A5379B49FEA3FC4B5C01FB9B" box="[375,559,1016,1040]" italics="true" pageId="6" pageNumber="287">G. macrodentata</emphasis>
</taxonomicName>
, the latter might better be placed in the genus
<taxonomicName id="BCBFA2F3A5379B49FBE2FC4B5AFBFB9B" box="[1078,1237,1016,1040]" class="Rhynchonellata" genus="Simpliciforma" kingdom="Animalia" order="Terebratulida" pageId="6" pageNumber="287" phylum="Brachiopoda" rank="genus">
<emphasis id="49CB0562A5379B49FBE2FC4B5AFBFB9B" box="[1078,1237,1016,1040]" italics="true" pageId="6" pageNumber="287">Simpliciforma</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="7B00D970A5379B49FF1DFBAF5A19FB4E" blockId="6.[151,1437,151,1221]" pageId="6" pageNumber="287">
Zezina (1976, p. 111) and Simon &amp; Willems (1999, p. 18) have shown that material collected by the Talisman expedition to the eastern Atlantic was erroneously attributed to
<taxonomicName id="BCBFA2F3A5379B49FC56FBF35A15FBD3" box="[898,1083,1088,1112]" class="Rhynchonellata" genus="Gwynia" kingdom="Animalia" order="Terebratulida" pageId="6" pageNumber="287" phylum="Brachiopoda" rank="species" species="capsula">
<emphasis id="49CB0562A5379B49FC56FBF35A15FBD3" box="[898,1083,1088,1112]" italics="true" pageId="6" pageNumber="287">Gwynia capsula</emphasis>
</taxonomicName>
by Fischer &amp; Oehlert (1891). Surprisingly, the specimen illustrated by them displays similarities to
<taxonomicName id="BCBFA2F3A5379B49FC7BFBD65A1FFBF7" box="[943,1073,1124,1148]" class="Rhynchonellata" genus="Simpliciforma" kingdom="Animalia" order="Terebratulida" pageId="6" pageNumber="287" phylum="Brachiopoda" rank="species" species="profunda">
<emphasis id="49CB0562A5379B49FC7BFBD65A1FFBF7" box="[943,1073,1124,1148]" italics="true" pageId="6" pageNumber="287">S. profunda</emphasis>
</taxonomicName>
, particularly in such features as its pointed ventral beak, straight and long hinge line and lack of submarginal ridges, however, their possible conspecifity needs further studies, especially considering a great geographical gap.
</paragraph>
</subSubSection>
</treatment>
</document>