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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.816.29738" ID-GBIF-Dataset="31bce27d-7094-467b-95ae-009a27e00cf5" ID-PMC="PMC6341056" ID-Pensoft-Pub="1313-2970-816-1" ID-PubMed="30686917" ID-ZBK="51CEEF2E1E1040A8A6731140426ED5A7" ModsDocAuthor="" ModsDocDate="2019" ModsDocID="1313-2970-816-1" ModsDocOrigin="ZooKeys 816" ModsDocTitle="A taxonomic review of the pericaline ground-beetles in Taiwan, with descriptions of new species (Coleoptera, Carabidae, Lebiini)" checkinTime="1547561011487" checkinUser="pensoft" docAuthor="Hunting, Wesley &amp; Yang, Man-Miao" docDate="2019" docId="EFC397CECE22026528BB1F5384241C30" docLanguage="en" docName="ZooKeys 816: 1-164" docOrigin="ZooKeys 816" docSource="http://dx.doi.org/10.3897/zookeys.816.29738" docTitle="Amphimenes absensacidus Hunting &amp; Yang, 2019, sp. n." docType="treatment" docUuid="01478AFD-2061-453B-95EF-D82FF1FC73F1" docUuidSource="ZooBank" docVersion="5" lastPageNumber="16" masterDocId="AA26FF84157BFFAE490E7F133744FF98" masterDocTitle="A taxonomic review of the pericaline ground-beetles in Taiwan, with descriptions of new species (Coleoptera, Carabidae, Lebiini)" masterLastPageNumber="164" masterPageNumber="1" pageNumber="12" updateTime="1668166739332" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A taxonomic review of the pericaline ground-beetles in Taiwan, with descriptions of new species (Coleoptera, Carabidae, Lebiini)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hunting, Wesley</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Yang, Man-Miao</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2019</mods:date>
<mods:detail type="volume">
<mods:number>816</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>164</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.816.29738</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.816.29738</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-816-1</mods:identifier>
<mods:identifier type="ZBK">51CEEF2E1E1040A8A6731140426ED5A7</mods:identifier>
<mods:identifier type="ZooBank">51CEEF2E1E1040A8A6731140426ED5A7</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="154125989" LSID="urn:lsid:zoobank.org:act:01478AFD-2061-453B-95EF-D82FF1FC73F1" httpUri="http://treatment.plazi.org/id/EFC397CECE22026528BB1F5384241C30" lastPageId="15" lastPageNumber="16" pageId="11" pageNumber="12">
<subSubSection pageId="11" pageNumber="12" type="nomenclature">
<paragraph pageId="11" pageNumber="12">
<taxonomicName LSID="http://zoobank.org/01478AFD-2061-453B-95EF-D82FF1FC73F1" class="Insecta" family="Carabidae" genus="Amphimenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphimenes absensacidus" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="absensacidus">Amphimenes absensacidus</taxonomicName>
<taxonomicNameLabel pageId="11" pageNumber="12">sp. n.</taxonomicNameLabel>
Figs 4, 5
<normalizedToken originalValue="AD">A-D</normalizedToken>
, 11B, 12
</paragraph>
</subSubSection>
<subSubSection pageId="12" pageNumber="13" type="specific epithet">
<paragraph pageId="12" pageNumber="13">
<pageBreakToken pageId="12" pageNumber="13" start="start">Specific</pageBreakToken>
epithet.
</paragraph>
<paragraph pageId="12" pageNumber="13">From Latin, absens and acidus, in reference to the lack of cross striations on the intervals of the elytral disc, typical in many species of this genus and all other species known from Taiwan.</paragraph>
</subSubSection>
<subSubSection pageId="12" pageNumber="13" type="type material">
<paragraph pageId="12" pageNumber="13">Types and other material examined.</paragraph>
<paragraph pageId="12" pageNumber="13">
Holotype (male) labeled
<normalizedToken originalValue="“Holotype”">&quot;Holotype&quot;</normalizedToken>
[circular, ringed with red]; &quot;TAIWAN: &quot;TAIWAN: Nantou Co./Huisun Forest Station /Hotel, May 26, 2012/24.0932N, 121.0310E&quot;; &quot;hand coll., veg. nr./Frog Rock trail, night,/~740m, Acc. Ti-170a/Coll. W. M. Hunting&quot;; &quot; NCHU#/100728&quot;. 28 paratypes, 13 males and 15 females. For further details see EH Strickland Virtual Entomology Museum Database.
</paragraph>
</subSubSection>
<subSubSection pageId="12" pageNumber="13" type="type locality">
<paragraph pageId="12" pageNumber="13">Type locality.</paragraph>
<paragraph pageId="12" pageNumber="13">Taiwan. Huisun Forest Station, Nantou county.</paragraph>
</subSubSection>
<subSubSection pageId="12" pageNumber="13" type="diagnosis">
<paragraph pageId="12" pageNumber="13">Diagnosis.</paragraph>
<paragraph pageId="12" pageNumber="13">
Specimens of this species are distinguished from other Taiwanese
<taxonomicName class="Insecta" family="Carabidae" genus="Amphimenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphimenes" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">Amphimenes</taxonomicName>
by lacking cross striations on the elytral disc.
</paragraph>
</subSubSection>
<subSubSection lastPageId="15" lastPageNumber="16" pageId="12" pageNumber="13" type="description">
<paragraph pageId="12" pageNumber="13">Description.</paragraph>
<paragraph pageId="12" pageNumber="13">OBL 5.8 - 6.8 mm. Length (n = ten males, ten females): head 0.58 - 0.70, pronotum 1.08 - 1.24, elytra 3.16 - 3.80, metepisternum 0.72 - 0.84 mm; width: head 1.12 - 1.36, pronotum 1.64 - 2.04, elytra 2.17 - 2.75, metepisternum 0.44 - 0.50 mm.</paragraph>
<paragraph pageId="12" pageNumber="13">Body proportions. HW/HL 1.87 - 2.14; PWM/PL 1.52 - 1.67; EL/EW 1.31 - 1.47; ML/MW 1.50 - 1.82</paragraph>
<paragraph pageId="12" pageNumber="13">Color. Fig. 4. Dorsum of head piceous; clypeus piceous; labrum piceous with brunneo-piceous margins; pronotum piceous with margins markedly pale, rufo-testaceous; proepipleuron rufo-testaceous to darker; antennae rufo-brunneous to brunneous; palpi rufo-brunneous to brunneous; elytral disc piceous, margins rufo-brunneous, translucent; elytral epipleura piceous to brunneo-piceous; thoracic sclerites rufo-brunneous to piceous; abdominal sterna rufo-brunneous medially and brunneo-piceous to piceous at lateral margins; legs with trochanter and femora testaceous to rufo-testaceous, tibia rufo-testaceous to brunneo-piceous.</paragraph>
<caption pageId="12" pageNumber="13">
<paragraph pageId="12" pageNumber="13">
Figure 4. Dorsal habitus and color pattern of
<taxonomicName class="Insecta" family="Carabidae" genus="Amphimenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphimenes absensacidus" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="absensacidus">Amphimenes absensacidus</taxonomicName>
sp. n.. (OBL 6.40 mm).
</paragraph>
</caption>
<paragraph pageId="12" pageNumber="13">
Microsculpture. Dorsum of head and pronotum with distinctive isodiametric mesh pattern easily visible at 50
<normalizedToken originalValue="×">x</normalizedToken>
magnification; elytra with transverse sculpticells visible throughout, moderately elongate; ventral surface of head, prosternum, proepipleuron, mesepisternum and metepisternum with sculpticells forming a moderately distinctive transverse mesh.
</paragraph>
<paragraph pageId="13" pageNumber="14">
<pageBreakToken pageId="13" pageNumber="14" start="start">Macrosculpture</pageBreakToken>
. Pronotum with scattered punctures in lateral margins; elytral striae evenly punctate along length.
</paragraph>
<paragraph pageId="13" pageNumber="14">
Pilosity. Elytra with scattered micro-punctures, each bearing a small seta; not visible at 50
<normalizedToken originalValue="×">x</normalizedToken>
magnification.
</paragraph>
<paragraph pageId="13" pageNumber="14">Luster. Elytra moderately glossy to moderately dull; ventral thoracic sterna and abdominal sterna moderately glossy.</paragraph>
<paragraph pageId="13" pageNumber="14">Head. Labrum more or less quadrate, few with slight indentation along apical margin but not bilobed; eyes distinctly convex, distinctly wider across than width of neck.</paragraph>
<paragraph pageId="13" pageNumber="14">Pronotum. Lateral margins distinctly explanate, broadly rounded; posterio-lateral margins rounded; posterio-lateral fovea relatively deep; basal lobe present.</paragraph>
<paragraph pageId="13" pageNumber="14">Elytra. Humeri broadly rounded; striae moderately impressed; lateral margins nearly parallel; elytral apices truncate.</paragraph>
<paragraph pageId="13" pageNumber="14">Hind wings. Macropterous.</paragraph>
<paragraph pageId="13" pageNumber="14">Legs. Meso-tibia with or without several shallow notches from mid-way to base along ventral surface.</paragraph>
<paragraph pageId="14" pageNumber="15">
<pageBreakToken pageId="14" pageNumber="15" start="start">Abdominal</pageBreakToken>
sterna. Abdominal sterna
<normalizedToken originalValue="IVVI">IV-VI</normalizedToken>
with two to four irregular punctures between fixed apical setae; abdominal sternum VII deeply bilobed.
</paragraph>
<paragraph pageId="14" pageNumber="15">
Male genitalia. Fig. 5
<normalizedToken originalValue="AD">A-D</normalizedToken>
. Length 1.28 - 1.44 mm. Phallus with one carina from base to mid-phallus, centered when viewed ventrally. Left side of shaft distinctively enlarged medially; endophallus with distinctive form, relatively large internal endophallic sclerite (es) near apex and several endophallic lobes (el).
</paragraph>
<caption pageId="14" pageNumber="15">
<paragraph pageId="14" pageNumber="15">
Figure 5. Digital images of male genitalia of
<taxonomicName class="Insecta" family="Carabidae" genus="Amphimenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphimenes absensacidus" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="absensacidus">Amphimenes absensacidus</taxonomicName>
sp. n.. A right lateral aspect, endophallus everted B right lateral aspect C ventral aspect D left lateral aspect. Legend: el endophallic lobes; es endophallic sclerite.
</paragraph>
</caption>
<paragraph pageId="15" pageNumber="16">
<pageBreakToken pageId="15" pageNumber="16" start="start">Female</pageBreakToken>
genitalia. Fig. 11B. Width 0.84 - 0.92mm. Two spermathecae, spermatheca 1 (sp1) and 2 (sp2) with ducts relatively smooth and even throughout length; One spermathecal accessory gland (sg); spermathecal gland duct (sgd) attachment site at base of spermatheca 1 duct, basal to attachment site of spermatheca 2 and associated diverticulum (div); one diverticulum, elongate.
</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="16" type="habitat">
<paragraph pageId="15" pageNumber="16">Habitat, habits, and seasonal occurrence.</paragraph>
<paragraph pageId="15" pageNumber="16">
The known elevational range of
<taxonomicName lsidName="A. absensacidus" pageId="15" pageNumber="16" rank="species" species="absensacidus">A. absensacidus</taxonomicName>
sp. n. is from 700 to 1850 meters. Adults of this species are found in mixed primary and secondary forest of montane areas. Adults are crepuscular or nocturnal with most activity observed on live tree trunks at night. Specimens have been collected from May to August. Methods of collecting include u.v. light and hand collecting at night.
</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="16" type="distribution">
<paragraph pageId="15" pageNumber="16">Geographical distribution.</paragraph>
<paragraph pageId="15" pageNumber="16">
<taxonomicName class="Insecta" family="Carabidae" genus="Amphimenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphimenes absensacidus" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="absensacidus">Amphimenes absensacidus</taxonomicName>
is known only from Taiwan. See Figure 12.
</paragraph>
</subSubSection>
</treatment>
</document>