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<document id="E5672F815844167B308F9D0F0B8BE27F" ID-CLB-Dataset="31368" ID-DOI="10.11646/zootaxa.4351.1.1" ID-GBIF-Dataset="c2a3a65e-90ff-4bba-af4d-91d82594f636" ID-ISSN="1175-5326" ID-Zenodo-Dep="1063746" ID-ZooBank="99BC6304-97CF-4463-B368-D2F102535DD9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1511248295005" checkinUser="plazi" docAuthor="Wang, Lin-Fei, Zhou, Hong-Zhang &amp; Lü, Liang" docDate="2017" docId="B44687B22B59FFD199E2F887FE44FDBB" docLanguage="en" docName="zootaxa.4351.1.1.pdf.imf" docOrigin="Zootaxa 4351 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Anotylus benisculptilis Wang &amp; Zhou &amp; Lü 2017, sp. nov." docType="treatment" docVersion="8" lastPageNumber="18" masterDocId="487FFFCA2B57FFC09975FFABFFDEFFA3" masterDocTitle="Revision of the Anotylus sculpturatus group (Coleoptera: Staphylinidae: Oxytelinae) with descriptions of seven new species from China" masterLastPageNumber="79" masterPageNumber="1" pageNumber="15" updateTime="1698502226059" updateUser="ExternalLinkService">
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<mods:title id="0D9E5DFADC31AF94F03B7DAFED27BBC7">Revision of the Anotylus sculpturatus group (Coleoptera: Staphylinidae: Oxytelinae) with descriptions of seven new species from China</mods:title>
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<mods:namePart id="89040BE2F4CE88A7500BE8DDFDDC6CDA">Wang, Lin-Fei</mods:namePart>
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<mods:namePart id="99782234DA236D21DD13E162D6ACF876">Zhou, Hong-Zhang</mods:namePart>
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<mods:namePart id="A7E77C602B15609E0FB3E019DB144B97">Lü, Liang</mods:namePart>
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<mods:title id="4D005C9C2398BBCCFEE559C7EA6C537D">Zootaxa</mods:title>
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<mods:date id="65D838649EF381A075AE5F790914C8E6">2017</mods:date>
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<mods:number id="309F8A23A1711D85608E1AB9343B99C5">4351</mods:number>
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<treatment id="B44687B22B59FFD199E2F887FE44FDBB" ID-DOI="http://doi.org/10.5281/zenodo.6033711" ID-GBIF-Taxon="137467060" ID-Zenodo-Dep="6033711" LSID="urn:lsid:plazi:treatment:B44687B22B59FFD199E2F887FE44FDBB" httpUri="http://treatment.plazi.org/id/B44687B22B59FFD199E2F887FE44FDBB" lastPageId="17" lastPageNumber="18" pageId="14" pageNumber="15">
<subSubSection id="74F5652F2B59FFCE99E2F887FF2BF8CB" pageId="14" pageNumber="15" type="nomenclature">
<paragraph id="3C5036A42B59FFCE99E2F887FDD0F8E4" blockId="14.[151,526,1836,1896]" box="[151,526,1836,1863]" pageId="14" pageNumber="15">
<heading id="671881C82B59FFCE99E2F887FDD0F8E4" bold="true" box="[151,526,1836,1863]" fontSize="11" level="1" pageId="14" pageNumber="15" reason="1">
<emphasis id="0E9BEAB62B59FFCE99E2F887FDD0F8E4" bold="true" box="[151,526,1836,1863]" pageId="14" pageNumber="15">
<taxonomicName id="FBEF4D272B59FFCE99E2F887FE74F8E5" ID-CoL="5V8F8" authority="Wang &amp; Zhou &amp; Lü, 2017" authorityName="Wang &amp; Zhou &amp; Lü" authorityYear="2017" box="[151,426,1836,1862]" class="Insecta" family="Staphylinidae" genus="Anotylus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="benisculptilis" status="sp. nov.">
<emphasis id="0E9BEAB62B59FFCE99E2F887FE74F8E5" bold="true" box="[151,426,1836,1862]" italics="true" pageId="14" pageNumber="15">Anotylus benisculptilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="15A857CD2B59FFCE98C5F886FDD0F8E4" box="[432,526,1837,1863]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="3C5036A42B59FFCE99E2F8E4FF2BF8CB" blockId="14.[151,526,1836,1896]" box="[151,245,1871,1896]" pageId="14" pageNumber="15">
(
<figureCitation id="A4D42A212B59FFCE99EAF8E4FF33F8CB" box="[159,237,1871,1896]" captionStart="FIGURE 3 4" captionStartId="14.[151,250,1146,1168]" captionTargetBox="[158,1430,194,1115]" captionTargetId="figure@14.[151,1436,193,1125]" captionTargetPageId="14" captionText="FIGURE 3 4. Anotylus varisculptilis sp. nov. different type of variations of forebody. A, type one; B, type two; C, type three; D, type four; E, head and pronotum of type one; F, head and pronotum of type two; G, head and pronotum of type three; H, head and pronotum of type four. Scales = 0.5 mm." httpUri="https://zenodo.org/record/1063758/files/figure.png" pageId="14" pageNumber="15">Figs. 4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="74F5652F2B59FFCE99E2F83EFCDEF855" pageId="14" pageNumber="15" type="materials_examined">
<paragraph id="3C5036A42B59FFCE99E2F83EFCEBF80D" blockId="14.[151,1436,1941,2038]" box="[151,821,1941,1966]" pageId="14" pageNumber="15">
<materialsCitation id="8C873CF92B59FFCE99E2F83EFCEBF80D" ID-GBIF-Occurrence="1701561675" box="[151,821,1941,1966]" country="China" county="Wolong County" location="Wolong County" pageId="14" pageNumber="15" specimenCount="1" stateProvince="Sichuan" typeStatus="holotype">
<emphasis id="0E9BEAB62B59FFCE99E2F83EFEE7F80D" bold="true" box="[151,313,1941,1966]" pageId="14" pageNumber="15">
<typeStatus id="E35488062B59FFCE99E2F83EFF0CF80D" box="[151,210,1941,1966]" pageId="14" pageNumber="15">Type</typeStatus>
locality:
</emphasis>
<collectingCountry id="44F876342B59FFCE9835F83DFE42F80D" box="[320,412,1942,1966]" name="China" pageId="14" pageNumber="15">CHINA</collectingCountry>
,
<collectingRegion id="FE2BF8462B59FFCE98D3F83EFDDDF80D" box="[422,515,1941,1966]" country="China" name="Sichuan" pageId="14" pageNumber="15">Sichuan</collectingRegion>
,
<collectingCounty id="D5314E282B59FFCE9B7AF83EFD1CF80D" box="[527,706,1941,1966]" pageId="14" pageNumber="15">Wolong County</collectingCounty>
, Wulidun.
</materialsCitation>
</paragraph>
<paragraph id="3C5036A42B59FFCE99B2F812FCDEF855" blockId="14.[151,1436,1941,2038]" pageId="14" pageNumber="15">
<materialsCitation id="8C873CF92B59FFCE99B2F812FCDEF855" ID-GBIF-Occurrence="1701561678" collectingDate="2004-05-06" collectingDateMax="2004-05-09" collectingDateMin="2004-05-06" collectorName="Grass &amp; Xiaodong Yu" country="China" county="Wolong County" elevation="2650" location="Wulidun" pageId="14" pageNumber="15" specimenCount="2" stateProvince="Sichuan Province" typeStatus="holotype">
<emphasis id="0E9BEAB62B59FFCE99B2F812FDB7F871" bold="true" box="[199,617,1977,2002]" pageId="14" pageNumber="15">
<typeStatus id="E35488062B59FFCE99B2F812FEDCF871" box="[199,258,1977,2002]" pageId="14" pageNumber="15">Type</typeStatus>
material examined.
<typeStatus id="E35488062B59FFCE9889F812FDB7F871" box="[508,617,1977,2002]" pageId="14" pageNumber="15" type="holotype">Holotype</typeStatus>
</emphasis>
: male,
<emphasis id="0E9BEAB62B59FFCE9BB5F812FC53F871" bold="true" box="[704,909,1977,2002]" pageId="14" pageNumber="15">
<collectingCountry id="44F876342B59FFCE9BB5F812FCC1F871" box="[704,799,1977,2002]" name="China" pageId="14" pageNumber="15">CHINA</collectingCountry>
,
<collectingRegion id="FE2BF8462B59FFCE9A58F812FC53F871" box="[813,909,1977,2002]" country="China" name="Sichuan" pageId="14" pageNumber="15">Sichuan</collectingRegion>
</emphasis>
Province:
<collectingCounty id="D5314E282B59FFCE9D7EF812FB1FF872" box="[1035,1217,1977,2002]" pageId="14" pageNumber="15">Wolong County</collectingCounty>
,
<location id="3930607F2B59FFCE9DB8F812FAEFF871" LSID="urn:lsid:plazi:treatment:B44687B22B59FFD199E2F887FE44FDBB:3930607F2B59FFCE9DB8F812FAEFF871" box="[1229,1329,1977,2002]" country="China" county="Wolong County" name="Wulidun" pageId="14" pageNumber="15" stateProvince="Sichuan Province">Wulidun</location>
,
<elevation id="B7C2D1972B59FFCE9C4BF812FA49F872" box="[1342,1431,1977,2002]" metricMagnitude="3" metricUnit="m" metricValue="2.65" pageId="14" pageNumber="15" unit="m" value="2650.0">
<quantity id="FB179B412B59FFCE9C4BF812FA49F872" box="[1342,1431,1977,2002]" metricMagnitude="3" metricUnit="m" metricValue="2.65" pageId="14" pageNumber="15" unit="m" value="2650.0">2650 m</quantity>
</elevation>
,
<collectorName id="911A53722B59FFCE99E2F875FF09F855" box="[151,215,2014,2038]" pageId="14" pageNumber="15">Grass</collectorName>
land,
<collectingDate id="5815E98C2B59FFCE9869F876FE70F855" box="[284,430,2013,2038]" pageId="14" pageNumber="15" value="2004-05-06" valueMax="2004-05-09" valueMin="2004-05-06">
<date id="485110642B59FFCE9869F876FE70F855" box="[284,430,2013,2038]" pageId="14" pageNumber="15" value="2004-05-06" valueMax="2004-05-09" valueMin="2004-05-06">69. V. 2004</date>
</collectingDate>
. coll.
<collectorName id="911A53722B59FFCE9885F876FD5BF856" box="[496,645,2013,2038]" pageId="14" pageNumber="15">Xiaodong Yu</collectorName>
(IZ-CAS).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="74F5652F2B58FFD199B2FF3CFBC8FF77" lastPageId="17" lastPageNumber="18" pageId="15" pageNumber="16" type="description">
<paragraph id="3C5036A42B58FFCF99B2FF3CFE4DFF77" blockId="15.[151,1437,151,896]" pageId="15" pageNumber="16">
<emphasis id="0E9BEAB62B58FFCF99B2FF3CFE87FF13" bold="true" box="[199,345,151,176]" pageId="15" pageNumber="16">Description.</emphasis>
Body dark brown; elytra and legs brown. Forebody sparsely pubescent, punctate and striate. Body length
<quantity id="FB179B412B58FFCF9852FF17FE50FF77" box="[295,398,188,213]" metricMagnitude="-3" metricUnit="m" metricValue="4.54" pageId="15" pageNumber="16" unit="mm" value="4.54">4.54 mm</quantity>
.
</paragraph>
<paragraph id="3C5036A42B58FFCF99B2FF4BFE7CFE0E" blockId="15.[151,1437,151,896]" pageId="15" pageNumber="16">
<emphasis id="0E9BEAB62B58FFCF99B2FF4BFEDFFF5B" box="[199,257,224,248]" italics="true" pageId="15" pageNumber="16">Male</emphasis>
: Head (
<figureCitation id="A4D42A212B58FFCF9815FF74FDC5FF5B" box="[352,539,223,248]" captionStart="FIGURE 3 4" captionStartId="14.[151,250,1146,1168]" captionTargetBox="[158,1430,194,1115]" captionTargetId="figure@14.[151,1436,193,1125]" captionTargetPageId="14" captionText="FIGURE 3 4. Anotylus varisculptilis sp. nov. different type of variations of forebody. A, type one; B, type two; C, type three; D, type four; E, head and pronotum of type one; F, head and pronotum of type two; G, head and pronotum of type three; H, head and pronotum of type four. Scales = 0.5 mm." httpUri="https://zenodo.org/record/1063758/files/figure.png" pageId="15" pageNumber="16">Fig. 4-1B, 4-2A</figureCitation>
) slightly broader than pronotum. Disc with small punctures and dense striae, vertex extremely convex, mid-longitudinal sulcus deep, slightly fused with deep impression on each side before base. Supra-antennal ridge with margin incrassate. Temples dilated, approximately two times as long as eyes. Clypeus nearly trapezoidal, about 1/3 of total head length, coriaceous, slightly convex, with anterior margin produced. Epistomal suture with lateral portions straight and obliquely inward, running backward to level of anterior margin of eyes.
</paragraph>
<paragraph id="3C5036A42B58FFCF99B2FE1CFC58FE57" blockId="15.[151,1437,151,896]" pageId="15" pageNumber="16">Occipital suture slightly curved forward in middle. Antenna filiform, segment IV globular, VIX short and transverse, with ultimate one stilliform. Neck without smooth area.</paragraph>
<paragraph id="3C5036A42B58FFCF99B2FDABFBBEFC23" blockId="15.[151,1437,151,896]" pageId="15" pageNumber="16">
Pronotum (
<figureCitation id="A4D42A212B58FFCF983EFE54FDD0FDBB" box="[331,526,511,536]" captionStart="FIGURE 3 4" captionStartId="14.[151,250,1146,1168]" captionTargetBox="[158,1430,194,1115]" captionTargetId="figure@14.[151,1436,193,1125]" captionTargetPageId="14" captionText="FIGURE 3 4. Anotylus varisculptilis sp. nov. different type of variations of forebody. A, type one; B, type two; C, type three; D, type four; E, head and pronotum of type one; F, head and pronotum of type two; G, head and pronotum of type three; H, head and pronotum of type four. Scales = 0.5 mm." httpUri="https://zenodo.org/record/1063758/files/figure.png" pageId="15" pageNumber="16">Fig. 4-1B, 4-2A</figureCitation>
) transverse, broadest at near anterior 1/3; disc slightly convex, with sparse longitudinal sculptures and punctures, denser laterally; median longitudinal sulcus deep and broad, with two broader and shallower paramedial sulci; bordered by much sharper longitudinal carina between mid-longitudinal sulcus and paramedial sulcus; with two shallow depressions close to lateral margins. Anterior margin broadly emarginate in middle. Lateral and posterior margins running less continuously, posterior angles obtuse but obvious. Lateral margin crenate on posterior half. Scutellum with crest-shaped impression, with median lobe sharply pointed. Elytra densely sculptured, punctured and pubescent; lateral margin curved and posterior margins truncate. Abdomen coriaceous, covered with punctures and pubescence. Sternite VII (
<figureCitation id="A4D42A212B58FFCF9A9EFD57FB43FCB7" box="[1003,1181,764,789]" captionStart="FIGURE 3 4" captionStartId="14.[151,250,1146,1168]" captionTargetBox="[158,1430,194,1115]" captionTargetId="figure@14.[151,1436,193,1125]" captionTargetPageId="14" captionText="FIGURE 3 4. Anotylus varisculptilis sp. nov. different type of variations of forebody. A, type one; B, type two; C, type three; D, type four; E, head and pronotum of type one; F, head and pronotum of type two; G, head and pronotum of type three; H, head and pronotum of type four. Scales = 0.5 mm." httpUri="https://zenodo.org/record/1063758/files/figure.png" pageId="15" pageNumber="16">Fig. 4-1C, 4-2B</figureCitation>
) with posterior margin slightly emarginate and bearing long setae laterally. Sternite VIII (
<figureCitation id="A4D42A212B58FFCF9AFFFCB4FB9DFC9B" box="[906,1091,799,824]" captionStart="FIGURE 3 4" captionStartId="14.[151,250,1146,1168]" captionTargetBox="[158,1430,194,1115]" captionTargetId="figure@14.[151,1436,193,1125]" captionTargetPageId="14" captionText="FIGURE 3 4. Anotylus varisculptilis sp. nov. different type of variations of forebody. A, type one; B, type two; C, type three; D, type four; E, head and pronotum of type one; F, head and pronotum of type two; G, head and pronotum of type three; H, head and pronotum of type four. Scales = 0.5 mm." httpUri="https://zenodo.org/record/1063758/files/figure.png" pageId="15" pageNumber="16">Fig. 4-1D, 4-2C</figureCitation>
) with posterior margin deeply bi-emarginate and thus forming triangulate plate in middle, with long setae laterally; middle angle rounded obtusely and more angulate in form, with tip slightly beyond lateral angles posteriorly.
</paragraph>
<caption id="6890662C2B58FFCF99E2F887FBC5F823" httpUri="https://zenodo.org/record/1063760/files/figure.png" pageId="15" pageNumber="16" targetBox="[162,1417,952,1814]" targetPageId="15">
<paragraph id="3C5036A42B58FFCF99E2F887FBC5F823" blockId="15.[151,1436,1836,1920]" pageId="15" pageNumber="16">
<emphasis id="0E9BEAB62B58FFCF99E2F887FEF7F8E1" bold="true" box="[151,297,1836,1858]" pageId="15" pageNumber="16">FIGURE 4-1.</emphasis>
<taxonomicName id="FBEF4D272B58FFCF9845F887FDC6F8E1" authority="Wang &amp; Zhou &amp; Lü, 2017" authorityName="Wang &amp; Zhou &amp; Lü" authorityYear="2017" box="[304,536,1836,1858]" class="Insecta" family="Staphylinidae" genus="Anotylus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="benisculptilis" status="sp. nov.">
<emphasis id="0E9BEAB62B58FFCF9845F887FDC6F8E1" box="[304,536,1836,1858]" italics="true" pageId="15" pageNumber="16">Anotylus benisculptilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="15A857CD2B58FFCF9B6AF887FDAFF8E1" box="[543,625,1836,1858]" pageId="15" pageNumber="16" rank="species">
<emphasis id="0E9BEAB62B58FFCF9B6AF887FDAFF8E1" bold="true" box="[543,625,1836,1858]" pageId="15" pageNumber="16">sp. nov.</emphasis>
</taxonomicNameLabel>
A, habitus male; B, head and pronotum male; C, sternite VII male; D, sternite VIII male; E, aedeagus (ventral view); F, median lobe of aedeagus (dorsal view); G, median lobe of aedeagus (lateral view); H, right paramere (in situ) of aedeagus (lateral view). Scales: A - C = 0.5 mm; rest, 0.2 mm.
</paragraph>
</caption>
<caption id="6890662C2B47FFD099E2F883FDA8F8D8" httpUri="https://zenodo.org/record/1063762/files/figure.png" pageId="16" pageNumber="17" targetBox="[220,1384,235,1804]" targetPageId="16">
<paragraph id="3C5036A42B47FFD099E2F883FDA8F8D8" blockId="16.[151,1436,1832,1915]" pageId="16" pageNumber="17">
<emphasis id="0E9BEAB62B47FFD099E2F883FEF6F89D" bold="true" box="[151,296,1832,1854]" pageId="16" pageNumber="17">FIGURE 4-2.</emphasis>
<taxonomicName id="FBEF4D272B47FFD0985BF883FDCAF89D" authority="Wang &amp; Zhou &amp; Lü, 2017" authorityName="Wang &amp; Zhou &amp; Lü" authorityYear="2017" box="[302,532,1832,1854]" class="Insecta" family="Staphylinidae" genus="Anotylus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="benisculptilis" status="sp. nov.">
<emphasis id="0E9BEAB62B47FFD0985BF883FDCAF89D" box="[302,532,1832,1854]" italics="true" pageId="16" pageNumber="17">Anotylus benisculptilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="15A857CD2B47FFD09B6FF883FDB4F89D" box="[538,618,1832,1854]" pageId="16" pageNumber="17" rank="species">
<emphasis id="0E9BEAB62B47FFD09B6FF883FDB4F89D" bold="true" box="[538,618,1832,1854]" pageId="16" pageNumber="17">sp. nov.</emphasis>
</taxonomicNameLabel>
A, head and pronotum male; B, sternite VII male; C, sternite VIII male; D, right paramere (in situ) of aedeagus (lateral view); E aedeagus (ventral view); F, aedeagus (dorsal view); G, median lobe of aedeagus (lateral view). Scale: A= 0.5 mm; rest, 0.2 mm.
</paragraph>
</caption>
<paragraph id="3C5036A42B47FFD099B2F803FA43F846" blockId="16.[151,1437,1960,2021]" pageId="16" pageNumber="17">
Aedeagus (
<figureCitation id="A4D42A212B47FFD09831F803FDE2F863" box="[324,572,1960,1985]" captionStart="FIGURE 3 4" captionStartId="14.[151,250,1146,1168]" captionTargetBox="[158,1430,194,1115]" captionTargetId="figure@14.[151,1436,193,1125]" captionTargetPageId="14" captionText="FIGURE 3 4. Anotylus varisculptilis sp. nov. different type of variations of forebody. A, type one; B, type two; C, type three; D, type four; E, head and pronotum of type one; F, head and pronotum of type two; G, head and pronotum of type three; H, head and pronotum of type four. Scales = 0.5 mm." httpUri="https://zenodo.org/record/1063758/files/figure.png" pageId="16" pageNumber="17">Fig. 4-1EG, 4-2 EG</figureCitation>
) with median lobe elongate, slightly inflated at base and shortly narrowing to apex; apical orifice large and ventral orifice crescentic; internal sac of median lobe with complicated sclerites.
</paragraph>
<paragraph id="3C5036A42B46FFD199E2FF33FBC8FF77" blockId="17.[151,1437,151,536]" pageId="17" pageNumber="18">
Parameres (
<figureCitation id="A4D42A212B46FFD19868FF3CFE04FF13" box="[285,474,151,176]" captionStart="FIGURE 3 4" captionStartId="14.[151,250,1146,1168]" captionTargetBox="[158,1430,194,1115]" captionTargetId="figure@14.[151,1436,193,1125]" captionTargetPageId="14" captionText="FIGURE 3 4. Anotylus varisculptilis sp. nov. different type of variations of forebody. A, type one; B, type two; C, type three; D, type four; E, head and pronotum of type one; F, head and pronotum of type two; G, head and pronotum of type three; H, head and pronotum of type four. Scales = 0.5 mm." httpUri="https://zenodo.org/record/1063758/files/figure.png" pageId="17" pageNumber="18">Fig. 4-1H, 4-2D</figureCitation>
) flattened and enlarged, broader at base, closely situated to each other and twining around median lobe; with thick seta at middle of ventral edge of each paramere.
</paragraph>
</subSubSection>
<subSubSection id="74F5652F2B46FFD199B2FF74FDFBFF5B" box="[199,549,223,248]" pageId="17" pageNumber="18" type="distribution">
<paragraph id="3C5036A42B46FFD199B2FF74FDFBFF5B" blockId="17.[151,1437,151,536]" box="[199,549,223,248]" pageId="17" pageNumber="18">
<emphasis id="0E9BEAB62B46FFD199B2FF74FEBEFF5B" bold="true" box="[199,352,223,248]" pageId="17" pageNumber="18">Distribution.</emphasis>
<collectingCountry id="44F876342B46FFD19812FF74FE74FF5B" box="[359,426,223,248]" name="China" pageId="17" pageNumber="18">China</collectingCountry>
(
<collectingRegion id="FE2BF8462B46FFD198CCFF74FDC4FF5B" box="[441,538,223,248]" country="China" name="Sichuan" pageId="17" pageNumber="18">Sichuan</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="74F5652F2B46FFD199B2FEAFFC38FE9C" pageId="17" pageNumber="18" type="etymology">
<paragraph id="3C5036A42B46FFD199B2FEAFFC38FE9C" blockId="17.[151,1437,151,536]" pageId="17" pageNumber="18">
<emphasis id="0E9BEAB62B46FFD199B2FEAFFE90FEBE" bold="true" box="[199,334,260,285]" pageId="17" pageNumber="18">Etymology.</emphasis>
The species epithet is from the Latin prefix &quot;
<emphasis id="0E9BEAB62B46FFD19A05FEAFFC7EFEBF" box="[880,928,260,284]" italics="true" pageId="17" pageNumber="18">beni</emphasis>
-&quot; and adjective
<emphasis id="0E9BEAB62B46FFD19D10FEAFFB08FEBF" box="[1125,1238,260,284]" italics="true" pageId="17" pageNumber="18">&quot;sculptilis</emphasis>
&quot; (sculptured), to indicate the strong and obvious striae and punctures on head and pronotum.
</paragraph>
</subSubSection>
<subSubSection id="74F5652F2B46FFD199B2FEE7FE44FDBB" pageId="17" pageNumber="18" type="diagnosis">
<paragraph id="3C5036A42B46FFD199B2FEE7FE44FDBB" blockId="17.[151,1437,151,536]" pageId="17" pageNumber="18">
<emphasis id="0E9BEAB62B46FFD199B2FEE7FE0FFEC6" bold="true" box="[199,465,332,357]" pageId="17" pageNumber="18">Differential diagnosis.</emphasis>
This new species is similar to
<taxonomicName id="FBEF4D272B46FFD19A3DFEE7FBBAFEC7" authority="(Sharp, 1874)" authorityName="P. M. Hammond" authorityYear="1874" baseAuthorityName="Sharp" baseAuthorityYear="1874" box="[840,1124,332,357]" class="Insecta" family="Staphylinidae" genus="Anotylus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="vicinus">
<emphasis id="0E9BEAB62B46FFD19A3DFEE7FC64FEC7" box="[840,954,332,356]" italics="true" pageId="17" pageNumber="18">A. vicinus</emphasis>
(
<bibRefCitation id="587E4B552B46FFD19ABBFEE7FB82FEC7" author="Sharp" box="[974,1116,332,357]" pageId="17" pageNumber="18" refString="Sharp, D. S. (1874) The Staphylinidae of Japan. Transactions of the Entomological Society of London, 22 (1), 1 - 103. https: // doi. org / 10.1111 / j. 1365 - 2311.1874. tb 00159. x" type="journal article" year="1874">Sharp, 1874</bibRefCitation>
)
</taxonomicName>
in appearance, but can be distinguished from the latter by having head and pronotum with sculpture deeper and coarser, male sternite VII is slightly emarginate, sternite VIII is bi-emarginate. Whereas
<taxonomicName id="FBEF4D272B46FFD19A35FE3FFB8DFE0F" authority="(Sharp, 1874)" authorityName="P. M. Hammond" authorityYear="1874" baseAuthorityName="Sharp" baseAuthorityYear="1874" box="[832,1107,404,429]" class="Insecta" family="Staphylinidae" genus="Anotylus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="vicinus">
<emphasis id="0E9BEAB62B46FFD19A35FE3FFC6EFE0F" box="[832,944,404,428]" italics="true" pageId="17" pageNumber="18">A. vicinus</emphasis>
(
<bibRefCitation id="587E4B552B46FFD19AB5FE3FFB95FE0F" author="Sharp" box="[960,1099,404,429]" pageId="17" pageNumber="18" refString="Sharp, D. S. (1874) The Staphylinidae of Japan. Transactions of the Entomological Society of London, 22 (1), 1 - 103. https: // doi. org / 10.1111 / j. 1365 - 2311.1874. tb 00159. x" type="journal article" year="1874">Sharp, 1874</bibRefCitation>
)
</taxonomicName>
has male sternite VII almost straight and sternite VIII broadly emarginate. The new species is also similar to
<taxonomicName id="FBEF4D272B46FFD19D31FE13FF06FE57" authority="(Bernhauer, 1938)" baseAuthorityName="Bernhauer" baseAuthorityYear="1938" class="Insecta" family="Staphylinidae" genus="Anotylus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="chinkiangensis">
<emphasis id="0E9BEAB62B46FFD19D31FE13FAD3FE73" box="[1092,1293,440,464]" italics="true" pageId="17" pageNumber="18">A. chinkiangensis</emphasis>
(
<bibRefCitation id="587E4B552B46FFD19C6AFE1CFF0EFE57" author="Bernhauer" pageId="17" pageNumber="18" refString="Bernhauer, M. (1938) Zur Staphylinidenfauna von China u. Japan. Entomologisches Nachrichtenblatt, Troppau, 12, 17 - 39." type="journal article" year="1938">Bernhauer, 1938</bibRefCitation>
)
</taxonomicName>
in the structure of the male genitalia, but can be distinguished from the latter by the structure in the internal sac of the median lobe.
</paragraph>
</subSubSection>
</treatment>
</document>