153 lines
20 KiB
XML
153 lines
20 KiB
XML
<document id="8AD5E26A866F7D7DD123543D39F66E38" ID-DOI="10.11646/zootaxa.4098.3.6" ID-GBIF-Dataset="687e2f7e-8268-4707-9cd7-bf46f9753f3b" ID-ISSN="1175-5326" ID-Zenodo-Dep="257723" ID-ZooBank="A005DB6C-7389-4506-B2D5-392531F832EB" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460012672336" checkinUser="plazi" docAuthor="Phan, Quoc Toan & Kompier, Tom" docDate="2016" docId="8A1EB74F897DFF94C7F36717FC3690D6" docLanguage="en" docName="zootaxa.4098.3.6.pdf.imf" docOrigin="Zootaxa 4098 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Protosticta caroli" docType="treatment" docVersion="7" lastPageNumber="532" masterDocId="7627CF37897FFF97C7646277FF99917B" masterDocTitle="A study of the genus Protosticta Selys, 1855, with descriptions of four new species from Vietnam (Odonata: Platystictidae)" masterLastPageNumber="544" masterPageNumber="529" pageNumber="531" updateTime="1698233463814" updateUser="plazi">
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<mods:titleInfo id="66F3574A98216F7C4EF0DD84050EA8EE">
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<mods:title id="AFD9954CAE6F9DA1DCFEC93A6C2677C8">A study of the genus Protosticta Selys, 1855, with descriptions of four new species from Vietnam (Odonata: Platystictidae)</mods:title>
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<mods:namePart id="FFB87AB9E673DBD8709E2F0B599A6C1D">Phan, Quoc Toan</mods:namePart>
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<mods:roleTerm id="4C3DA8FB1CAE3F3907979FED71DC138C">Author</mods:roleTerm>
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<mods:namePart id="15D280421E1AC02BCD2B127A72F3C40E">Kompier, Tom</mods:namePart>
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<mods:typeOfResource id="8629952D1BD3CCF51C584B330635C766">text</mods:typeOfResource>
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<mods:title id="674D46DF75CA54D51F70A930065E143E">Zootaxa</mods:title>
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<mods:part id="1A9237F7A063E2CF96DC35171A8FB983">
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<mods:date id="3E2E87C8F1AF70F644D44705E0CC4975">2016</mods:date>
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<mods:number id="73AF9C699017514D5C0324BBDA88B1CF">4098</mods:number>
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<mods:identifier id="099907945E3FA6FE544D3E29E89B36C2" type="DOI">10.11646/zootaxa.4098.3.6</mods:identifier>
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<treatment id="8A1EB74F897DFF94C7F36717FC3690D6" ID-DOI="http://doi.org/10.5281/zenodo.6056373" ID-GBIF-Taxon="127620791" ID-Zenodo-Dep="6056373" LSID="urn:lsid:plazi:treatment:8A1EB74F897DFF94C7F36717FC3690D6" httpUri="http://treatment.plazi.org/id/8A1EB74F897DFF94C7F36717FC3690D6" lastPageId="3" lastPageNumber="532" pageId="2" pageNumber="531">
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<subSubSection id="4AAD55D2897DFF95C7F36717FDA89401" box="[151,561,1375,1402]" pageId="2" pageNumber="531" type="nomenclature">
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<paragraph id="02080659897DFF95C7F36717FDA89401" blockId="2.[151,561,1375,1435]" box="[151,561,1375,1402]" pageId="2" pageNumber="531">
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<heading id="5940B135897DFF95C7F36717FDA89401" bold="true" box="[151,561,1375,1402]" fontSize="11" level="1" pageId="2" pageNumber="531" reason="1">
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<emphasis id="30C3DA4B897DFF95C7F36717FDA89401" bold="true" box="[151,561,1375,1402]" pageId="2" pageNumber="531">
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2.
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<taxonomicName id="C5B77DDA897DFF95C7D06728FE1A9402" ID-CoL="4N782" box="[180,387,1375,1401]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="531" phylum="Arthropoda" rank="species" species="caroli">
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<emphasis id="30C3DA4B897DFF95C7D06728FE1A9402" bold="true" box="[180,387,1375,1401]" italics="true" pageId="2" pageNumber="531">Protosticta caroli</emphasis>
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</taxonomicName>
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van
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<bibRefCitation id="66267BA8897DFF95C6DB6717FDA89401" author="Tol" box="[447,561,1376,1402]" pageId="2" pageNumber="531" refString="van Tol, J. (2008) Notes on some species of the genus Protosticta from Vietnam (Odonata, Platystictidae). Zoologische Mededelingen Leiden, 82, 217 - 234." type="journal article" year="2008">Tol, 2008</bibRefCitation>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="4AAD55D2897DFF95C7F367F5FEAD94E0" box="[151,308,1410,1435]" pageId="2" pageNumber="531" type="description">
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<paragraph id="02080659897DFF95C7F367F5FEAD94E0" blockId="2.[151,561,1375,1435]" box="[151,308,1410,1435]" pageId="2" pageNumber="531">
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(
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<figureCitation id="9A8C1ADC897DFF95C7FA67F5FF6494E0" box="[158,253,1410,1435]" captionStart="FIGURE 2" captionStartId="3.[151,250,1687,1709]" captionTargetBox="[159,1432,476,1674]" captionTargetId="figure@3.[151,1437,469,1675]" captionTargetPageId="3" captionText="FIGURE 2. Protosticta caroli, Ba Na. [A – G] male— (A) head & thorax in lateral view; (B) prothorax in dorsal view; anal appendages in (C) dorsal & (D) lateral view; (E) right cercus and paraproct in oblique-dorsal view; (F) male paraprocts in ventral view; (G) penile organ in dorsal view; [H – J] female— (H) head & thorax in lateral view; (I) prothorax in dorsal view; (J) tip abdomen in lateral view." httpUri="https://zenodo.org/record/257725/files/figure.png" pageId="2" pageNumber="531">Figure 2</figureCitation>
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A–J)
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</paragraph>
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</subSubSection>
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<subSubSection id="4AAD55D2897DFF95C7F367BCFE0D961F" pageId="2" pageNumber="531" type="reference_group">
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<paragraph id="02080659897DFF95C7F367BCFC3E9484" blockId="2.[151,1436,1482,1535]" pageId="2" pageNumber="531">
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<taxonomicName id="C5B77DDA897DFF95C7F367BCFED0949B" ID-CoL="4N782" box="[151,329,1482,1504]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="531" phylum="Arthropoda" rank="species" species="caroli">
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<emphasis id="30C3DA4B897DFF95C7F367BCFED0949B" box="[151,329,1482,1504]" italics="true" pageId="2" pageNumber="531">Protosticta caroli</emphasis>
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</taxonomicName>
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: van
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<bibRefCitation id="66267BA8897DFF95C6E067BDFE6A949B" author="Tol" box="[388,499,1482,1504]" pageId="2" pageNumber="531" refString="van Tol, J. (2008) Notes on some species of the genus Protosticta from Vietnam (Odonata, Platystictidae). Zoologische Mededelingen Leiden, 82, 217 - 234." type="journal article" year="2008">Tol (2008)</bibRefCitation>
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, pp. 227–229, Figs. 19–22.
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<typeStatus id="DD0CB8FB897DFF95C47067BCFCDF949B" box="[788,838,1483,1504]" pageId="2" pageNumber="531">Type</typeStatus>
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locality: Chu Yang Sin National Park, Dak Lak Province, southern
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<collectingCountry id="7AA046C9897DFF95C640679EFEE29484" box="[292,379,1513,1535]" name="Vietnam" pageId="2" pageNumber="531">Vietnam</collectingCountry>
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.—
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<bibRefCitation id="66267BA8897DFF95C6FC679EFD4C9484" author="Yokoi" box="[408,725,1513,1535]" pageId="2" pageNumber="531" refString="Yokoi, N. & Souphanthong, V. (2014) A list of Lao Dragonflies. Kyoei Printing Co. Ltd. Koriyama, 110 pp, 28 pls." type="book" year="2014">Yokoi & Souphanthong (2014)</bibRefCitation>
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, p. 25, Pl. 2, Fig. 11.
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</paragraph>
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<paragraph id="02080659897DFF95C7F3645BFA08973E" blockId="2.[151,1437,1580,2037]" box="[151,1425,1580,1605]" pageId="2" pageNumber="531">
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<emphasis id="30C3DA4B897DFF95C7F3645BFE16973E" bold="true" box="[151,399,1580,1605]" pageId="2" pageNumber="531">Specimens examined</emphasis>
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.
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<specimenCount id="14B1CDD0897DFF95C6F9645BFE6F973E" box="[413,502,1580,1605]" pageId="2" pageNumber="531" type="male">9 males</specimenCount>
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,
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<specimenCount id="14B1CDD0897DFF95C565645BFDE9973E" box="[513,624,1580,1605]" pageId="2" pageNumber="531" type="female">2 females</specimenCount>
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, Ba Na Nature Reserve, Da Nang city,
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<date id="76092099897DFF95C342645BFB02973E" box="[1062,1179,1580,1605]" pageId="2" pageNumber="531" value="2015-05-25">25.V.2015</date>
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, Quoc Toan Phan leg.
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</paragraph>
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<paragraph id="02080659897DFF95C7A36438FE0D961F" blockId="2.[151,1437,1580,2037]" pageId="2" pageNumber="531">
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<emphasis id="30C3DA4B897DFF95C7A36438FE939713" bold="true" box="[199,266,1615,1640]" pageId="2" pageNumber="531">Notes</emphasis>
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. Van
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<bibRefCitation id="66267BA8897DFF95C62A6438FE5E9713" author="Tol" box="[334,455,1615,1640]" pageId="2" pageNumber="531" refString="van Tol, J. (2008) Notes on some species of the genus Protosticta from Vietnam (Odonata, Platystictidae). Zoologische Mededelingen Leiden, 82, 217 - 234." type="journal article" year="2008">Tol (2008)</bibRefCitation>
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described
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<emphasis id="30C3DA4B897DFF95C5206426FD069713" box="[580,671,1615,1640]" italics="true" pageId="2" pageNumber="531">P. c a ro l i</emphasis>
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from southern
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<collectingCountry id="7AA046C9897DFF95C4306438FC2A9713" box="[852,947,1615,1640]" name="Vietnam" pageId="2" pageNumber="531">Vietnam</collectingCountry>
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based on male specimens. The body color pattern, anal appendages and penile structure of our specimens (
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<figureCitation id="9A8C1ADC897DFF95C4166403FC2697F7" box="[882,959,1652,1677]" captionStart="FIGURE 2" captionStartId="3.[151,250,1687,1709]" captionTargetBox="[159,1432,476,1674]" captionTargetId="figure@3.[151,1437,469,1675]" captionTargetPageId="3" captionText="FIGURE 2. Protosticta caroli, Ba Na. [A – G] male— (A) head & thorax in lateral view; (B) prothorax in dorsal view; anal appendages in (C) dorsal & (D) lateral view; (E) right cercus and paraproct in oblique-dorsal view; (F) male paraprocts in ventral view; (G) penile organ in dorsal view; [H – J] female— (H) head & thorax in lateral view; (I) prothorax in dorsal view; (J) tip abdomen in lateral view." httpUri="https://zenodo.org/record/257725/files/figure.png" pageId="2" pageNumber="531">Figs. 2</figureCitation>
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A–G) match the original description well, allowing identification as
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<emphasis id="30C3DA4B897DFF95C6DD64EEFD8B97CB" box="[441,530,1687,1712]" italics="true" pageId="2" pageNumber="531">P. c aro l i</emphasis>
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. However, males from central
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<collectingCountry id="7AA046C9897DFF95C40A64E0FC5497CB" box="[878,973,1687,1712]" name="Vietnam" pageId="2" pageNumber="531">Vietnam</collectingCountry>
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are much larger than southern specimens (hind wing
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<quantity id="C54FABBC897DFF95C64664CBFEE197AF" box="[290,376,1724,1749]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="2" pageNumber="531" unit="mm" value="25.0">25 mm</quantity>
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, abdomen + appendages
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<quantity id="C54FABBC897DFF95C5C164CBFCBC97AF" box="[677,805,1724,1749]" metricMagnitude="-2" metricUnit="m" metricValue="5.05" metricValueMax="5.2" metricValueMin="4.9" pageId="2" pageNumber="531" unit="mm" value="50.5" valueMax="52.0" valueMin="49.0">49–52 mm</quantity>
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, but
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<quantity id="C54FABBC897DFF95C40264CBFC5797AF" box="[870,974,1724,1749]" metricMagnitude="-2" metricUnit="m" metricValue="2.05" pageId="2" pageNumber="531" unit="mm" value="20.5">20.5 mm</quantity>
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and
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<quantity id="C54FABBC897DFF95C36A64CBFBFB97AF" box="[1038,1122,1724,1748]" metricMagnitude="-2" metricUnit="m" metricValue="3.9" pageId="2" pageNumber="531" unit="mm" value="39.0">39 mm</quantity>
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in
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<typeStatus id="DD0CB8FB897DFF95C3F464CBFB6C97AE" box="[1168,1269,1724,1749]" pageId="2" pageNumber="531" type="holotype">holotype</typeStatus>
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, respectively). Moreover, van
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<bibRefCitation id="66267BA8897DFF95C62564A8FE689783" author="Tol" box="[321,497,1759,1784]" pageId="2" pageNumber="531" refString="van Tol, J. (2008) Notes on some species of the genus Protosticta from Vietnam (Odonata, Platystictidae). Zoologische Mededelingen Leiden, 82, 217 - 234." type="journal article" year="2008">Tol (2008: 229)</bibRefCitation>
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described the paraprocts of the
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<typeStatus id="DD0CB8FB897DFF95C43264A8FC219783" box="[854,952,1759,1784]" pageId="2" pageNumber="531" type="holotype">holotype</typeStatus>
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as “in ventral view forming a heart-shaped figure, (.) in posterior view a long and stout terminal spine, which is bent abaxiad, visible”, but in our specimens the left and right paraproct are somewhat separated, not forming a heart shape; the spine can be seen easily in ventral view (
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<figureCitation id="9A8C1ADC897DFF95C655653BFEEC961F" box="[305,373,1868,1893]" captionStart="FIGURE 2" captionStartId="3.[151,250,1687,1709]" captionTargetBox="[159,1432,476,1674]" captionTargetId="figure@3.[151,1437,469,1675]" captionTargetPageId="3" captionText="FIGURE 2. Protosticta caroli, Ba Na. [A – G] male— (A) head & thorax in lateral view; (B) prothorax in dorsal view; anal appendages in (C) dorsal & (D) lateral view; (E) right cercus and paraproct in oblique-dorsal view; (F) male paraprocts in ventral view; (G) penile organ in dorsal view; [H – J] female— (H) head & thorax in lateral view; (I) prothorax in dorsal view; (J) tip abdomen in lateral view." httpUri="https://zenodo.org/record/257725/files/figure.png" pageId="2" pageNumber="531">Fig. 2</figureCitation>
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F).
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</paragraph>
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</subSubSection>
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<subSubSection id="4AAD55D2897DFF94C7A36518FE2C901F" lastPageId="3" lastPageNumber="532" pageId="2" pageNumber="531" type="description">
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<paragraph id="02080659897DFF94C7A36518FA059067" blockId="2.[151,1437,1580,2037]" lastBlockId="3.[151,1437,151,429]" lastPageId="3" lastPageNumber="532" pageId="2" pageNumber="531">
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<emphasis id="30C3DA4B897DFF95C7A36518FE6D96F3" bold="true" box="[199,500,1903,1928]" pageId="2" pageNumber="531">Description of the female</emphasis>
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. Body color pattern as male. Head (
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<figureCitation id="9A8C1ADC897DFF95C4E96518FC4996F3" box="[909,976,1903,1928]" captionStart="FIGURE 2" captionStartId="3.[151,250,1687,1709]" captionTargetBox="[159,1432,476,1674]" captionTargetId="figure@3.[151,1437,469,1675]" captionTargetPageId="3" captionText="FIGURE 2. Protosticta caroli, Ba Na. [A – G] male— (A) head & thorax in lateral view; (B) prothorax in dorsal view; anal appendages in (C) dorsal & (D) lateral view; (E) right cercus and paraproct in oblique-dorsal view; (F) male paraprocts in ventral view; (G) penile organ in dorsal view; [H – J] female— (H) head & thorax in lateral view; (I) prothorax in dorsal view; (J) tip abdomen in lateral view." httpUri="https://zenodo.org/record/257725/files/figure.png" pageId="2" pageNumber="531">Fig. 2</figureCitation>
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H) black, with half of labrum, genae and entire anteclypeus bluish yellow; inferior margin of labrum black, forming a small dot at the middle that is absent in the male. Antennae black, but basal half of second segment pale yellow. Thoracic markings (
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<figureCitation id="9A8C1ADC897DFF95C3AE65CFFA9596AB" box="[1226,1292,1976,2001]" captionStart="FIGURE 2" captionStartId="3.[151,250,1687,1709]" captionTargetBox="[159,1432,476,1674]" captionTargetId="figure@3.[151,1437,469,1675]" captionTargetPageId="3" captionText="FIGURE 2. Protosticta caroli, Ba Na. [A – G] male— (A) head & thorax in lateral view; (B) prothorax in dorsal view; anal appendages in (C) dorsal & (D) lateral view; (E) right cercus and paraproct in oblique-dorsal view; (F) male paraprocts in ventral view; (G) penile organ in dorsal view; [H – J] female— (H) head & thorax in lateral view; (I) prothorax in dorsal view; (J) tip abdomen in lateral view." httpUri="https://zenodo.org/record/257725/files/figure.png" pageId="2" pageNumber="531">Fig. 2</figureCitation>
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H) similar to male; prothorax pale yellow, with a black marking on middle of anterior lobe and an all-black posterior lobe, black extending onto distal side of lateral lobes. Wings hyaline, 16 postnodal veins in forewing and
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<quantity id="C54FABBC897CFF94C3A462E0FABD91CB" box="[1216,1316,151,176]" metricMagnitude="-1" metricUnit="m" metricValue="4.191" metricValueMax="4.318" metricValueMin="4.064" pageId="3" pageNumber="532" unit="in" value="16.5" valueMax="17.0" valueMin="16.0">16–17 in</quantity>
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hindwing. Pterostigma black. Abdomen black, with yellow markings as follows: S1 with an anteroventral oval spot; in S2, this marking extends to middle of segment; S3–5 with ventral spots at either end of segment; S6–7 with a yellow ventral marking; S8 black with an anteroventral yellow mark as
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<figureCitation id="9A8C1ADC897CFF94C4046373FC3B9067" box="[864,930,260,285]" captionStart="FIGURE 2" captionStartId="3.[151,250,1687,1709]" captionTargetBox="[159,1432,476,1674]" captionTargetId="figure@3.[151,1437,469,1675]" captionTargetPageId="3" captionText="FIGURE 2. Protosticta caroli, Ba Na. [A – G] male— (A) head & thorax in lateral view; (B) prothorax in dorsal view; anal appendages in (C) dorsal & (D) lateral view; (E) right cercus and paraproct in oblique-dorsal view; (F) male paraprocts in ventral view; (G) penile organ in dorsal view; [H – J] female— (H) head & thorax in lateral view; (I) prothorax in dorsal view; (J) tip abdomen in lateral view." httpUri="https://zenodo.org/record/257725/files/figure.png" pageId="3" pageNumber="532">Fig. 2</figureCitation>
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J; S9–10 entirely black. Ovipositor as
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<figureCitation id="9A8C1ADC897CFF94C22E6373FA159067" box="[1354,1420,260,285]" captionStart="FIGURE 2" captionStartId="3.[151,250,1687,1709]" captionTargetBox="[159,1432,476,1674]" captionTargetId="figure@3.[151,1437,469,1675]" captionTargetPageId="3" captionText="FIGURE 2. Protosticta caroli, Ba Na. [A – G] male— (A) head & thorax in lateral view; (B) prothorax in dorsal view; anal appendages in (C) dorsal & (D) lateral view; (E) right cercus and paraproct in oblique-dorsal view; (F) male paraprocts in ventral view; (G) penile organ in dorsal view; [H – J] female— (H) head & thorax in lateral view; (I) prothorax in dorsal view; (J) tip abdomen in lateral view." httpUri="https://zenodo.org/record/257725/files/figure.png" pageId="3" pageNumber="532">Fig. 2</figureCitation>
|
||
J.
|
||
</paragraph>
|
||
<paragraph id="02080659897CFF94C7A36350FE2C901F" blockId="3.[151,1437,151,429]" pageId="3" pageNumber="532">
|
||
<emphasis id="30C3DA4B897CFF94C7A36350FEEF903B" bold="true" box="[199,374,295,320]" pageId="3" pageNumber="532">Measurements</emphasis>
|
||
. Male. Hindwing 25; abdomen including appendages 49–52. Female. Hindwing 24, abdomen including appendages 40.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4AAD55D2897CFF94C7A36318FC3690D6" pageId="3" pageNumber="532" type="distribution">
|
||
<paragraph id="02080659897CFF94C7A36318FC3690D6" blockId="3.[151,1437,151,429]" pageId="3" pageNumber="532">
|
||
<emphasis id="30C3DA4B897CFF94C7A36318FEC090F3" bold="true" box="[199,345,367,392]" pageId="3" pageNumber="532">Distribution</emphasis>
|
||
.
|
||
<collectingCountry id="7AA046C9897CFF94C6036318FE5F90F3" box="[359,454,367,392]" name="Vietnam" pageId="3" pageNumber="532">Vietnam</collectingCountry>
|
||
(Ba Na Nature Reserve, Da Nang city; Chu Yang Sin National Park, Dak Lak Province; Bao Loc, Lam Dong Province), Lao PDR (Luangnamtha, Muangsing).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |