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<document id="4E67D8F7322AAD4A16E14CF7CE6ABF57" ID-DOI="10.5281/zenodo.157029" ID-GBIF-Dataset="aba28cae-a8bc-4fb7-98bd-5bd284ccc89d" ID-ISSN="1175­5326" ID-Zenodo-Dep="157029" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459625440537" checkinUser="plazi" docAuthor="Johanson, Kjell Arne" docDate="2003" docId="736E87EFFFEAFFD62F5DE5FBFAE0836F" docLanguage="en" docName="zt00283.pdf" docOrigin="Zootaxa 283" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Helicopsychidae" docType="key" docVersion="9" lastPageNumber="9" masterDocId="8F57FF97FFEDFFDE2E55E307FFCB8739" masterDocTitle="The Helicopsyche (Feropsyche) (Insecta, Trichoptera, Helicopsychidae) from Barro Colorado Island, Panama" masterLastPageNumber="12" masterPageNumber="1" pageNumber="8" updateTime="1698203957936" updateUser="plazi">
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<mods:title id="F50FA90B7A7466664587E775BF2A1D98">The Helicopsyche (Feropsyche) (Insecta, Trichoptera, Helicopsychidae) from Barro Colorado Island, Panama</mods:title>
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<emphasis id="C9B3EAEBFFEAFFD92F5DE5FBFBF9802F" bold="true" box="[264,1074,1788,1814]" pageId="7" pageNumber="8">
Key to the males of
<taxonomicName id="3CC74D7AFFEAFFD92FAAE5FBFD01802F" ID-CoL="6252B" box="[511,714,1788,1814]" class="Insecta" family="Helicopsychidae" kingdom="Animalia" order="Trichoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="family">Helicopsychidae</taxonomicName>
from Barro Colorado Island
</emphasis>
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<paragraph id="FB7836F9FFEAFFD92F5DE43EFAE0806A" blockId="7.[264,1323,1849,1875]" box="[264,1323,1849,1875]" pageId="7" pageNumber="8">
<keyLead id="403695CCFFEAFFD92F5DE43EFAE0806A" box="[264,1323,1849,1875]" pageId="7" pageNumber="8">1 Postantennal setal warts strongly enlarged and larger than the cephalic setal warts in</keyLead>
</paragraph>
<paragraph id="FB7836F9FFE5FFD62F6DE21BFAE68697" blockId="8.[264,1325,284,1110]" pageId="8" pageNumber="9">
lateral view (fig. 42A in
<bibRefCitation id="9F564B08FFE5FFD62C36E21BFCD2860F" author="Johanson" box="[611,793,284,310]" pageId="8" pageNumber="12" refString="Johanson, K. A. (2002) Systematic revision of American Helicopsyche of the subgenus Feropsyche (Trichoptera: Helicopsychidae). Insect Systematics and Evolution Supplement, 60, 1 - 147." type="journal article" year="2002">Johanson 2002</bibRefCitation>
); cephalic setal warts anteriorly swollen and separate from head capsule (fig. 42A in
<bibRefCitation id="9F564B08FFE5FFD62D5BE243FC0B8667" author="Johanson" box="[782,960,324,350]" pageId="8" pageNumber="12" refString="Johanson, K. A. (2002) Systematic revision of American Helicopsyche of the subgenus Feropsyche (Trichoptera: Helicopsychidae). Insect Systematics and Evolution Supplement, 60, 1 - 147." type="journal article" year="2002">Johanson 2002</bibRefCitation>
)
<emphasis id="C9B3EAEBFFE5FFD62D9BE243FB1C8664" box="[974,1239,324,349]" italics="true" pageId="8" pageNumber="9">
................
<taxonomicName id="3CC74D7AFFE5FFD62A13E243FB1C8664" ID-CoL="7VCBY" authority="Bueno" authorityName="Bueno" box="[1094,1239,324,349]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="tuxtlensis">H. tuxtlensis</taxonomicName>
</emphasis>
Bueno
<keyLead id="403695CCFFE5FFD62F5DE26BFAE68697" pageId="8" pageNumber="9">­ Postantennal setal warts much smaller than the cephalic setal warts; cephalic setal warts not swollen anteriorly and fused with head capsule along its length .................. 2</keyLead>
</paragraph>
</keyStep>
<keyStep id="40332E5CFFE5FFD62F5DE2BBFAE085A7" pageId="8" pageNumber="9">
<paragraph id="FB7836F9FFE5FFD62F5DE2BBFAE0851F" blockId="8.[264,1325,284,1110]" pageId="8" pageNumber="9">
<keyLead id="403695CCFFE5FFD62F5DE2BBFAE0851F" pageId="8" pageNumber="9">
2 Xth tergum with row of megasetae starting proximally on the segment; gonocoxite strongly sickle­shaped; basimesal lobe tuboid; phallus uniformly curving ventrally along its length .................................................................................
<taxonomicName id="3CC74D7AFFE5FFD62A07E10BFAE0851F" ID-CoL="7VCC3" authority="Ross" authorityName="Ross" box="[1106,1323,524,550]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="vergelana">
<emphasis id="C9B3EAEBFFE5FFD62A07E10BFB21851C" box="[1106,1258,524,549]" italics="true" pageId="8" pageNumber="9">H. vergelana</emphasis>
Ross
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</paragraph>
<paragraph id="FB7836F9FFE5FFD62F5DE133FAE085A7" blockId="8.[264,1325,284,1110]" pageId="8" pageNumber="9">
<keyLead id="403695CCFFE5FFD62F5DE133FAE085A7" pageId="8" pageNumber="9">
­ Xth tergum with megasetae present only on mid and distal part of the segment (
<figureCitation id="63FC2A7CFFE5FFD62A8CE133FAEE8577" box="[1241,1317,564,590]" captionStart="FIGURES 1 ­ 8" captionStartId="2.[264,382,1813,1837]" captionTargetBox="[266,1307,465,1791]" captionTargetId="figure@2.[249,1324,449,1803]" captionTargetPageId="2" captionText="FIGURES 1 ­ 8. Helicopsyche woldai sp. n., male holotype. 1, right wings; 2, abdominal segment VI, lateral view; 3, abdominal VIth sternal process, ventral view; 4, genitalia, lateral view; 5, genitalia, dorsal view; 6, genitalia, ventral view; 7, phallus, lateral view; 8, phallus, ventral view." httpUri="https://zenodo.org/record/157030/files/figure.png" pageId="8" pageNumber="9">Figs 5</figureCitation>
,
<figureCitation id="63FC2A7CFFE5FFD62F6DE15BFE93854F" box="[312,344,604,630]" captionStart="FIGURES 9 ­ 16" captionStartId="5.[264,382,1617,1641]" captionTargetBox="[272,1322,318,1589]" captionTargetId="figure@5.[268,1343,302,1602]" captionTargetPageId="5" captionText="FIGURES 9 ­ 16. Helicopsyche fridae sp. n., male holotype. 9, right wings; 10, abdominal segment VI, lateral view; 11, abdominal VIth sternal process, ventral view; 12, genitalia, lateral view; 13, genitalia, dorsal view; 14, genitalia, ventral view; 15, phallus, lateral view; 16, phallus, ventral view." httpUri="https://zenodo.org/record/157031/files/figure.png" pageId="8" pageNumber="9">13</figureCitation>
); gonocoxite shape different (
<figureCitation id="63FC2A7CFFE5FFD62CE3E15BFCC2854F" box="[694,777,604,630]" captionStart="FIGURES 1 ­ 8" captionStartId="2.[264,382,1813,1837]" captionTargetBox="[266,1307,465,1791]" captionTargetId="figure@2.[249,1324,449,1803]" captionTargetPageId="2" captionText="FIGURES 1 ­ 8. Helicopsyche woldai sp. n., male holotype. 1, right wings; 2, abdominal segment VI, lateral view; 3, abdominal VIth sternal process, ventral view; 4, genitalia, lateral view; 5, genitalia, dorsal view; 6, genitalia, ventral view; 7, phallus, lateral view; 8, phallus, ventral view." httpUri="https://zenodo.org/record/157030/files/figure.png" pageId="8" pageNumber="9">Figs. 4</figureCitation>
,
<figureCitation id="63FC2A7CFFE5FFD62D43E15BFCFD854F" box="[790,822,604,630]" captionStart="FIGURES 9 ­ 16" captionStartId="5.[264,382,1617,1641]" captionTargetBox="[272,1322,318,1589]" captionTargetId="figure@5.[268,1343,302,1602]" captionTargetPageId="5" captionText="FIGURES 9 ­ 16. Helicopsyche fridae sp. n., male holotype. 9, right wings; 10, abdominal segment VI, lateral view; 11, abdominal VIth sternal process, ventral view; 12, genitalia, lateral view; 13, genitalia, dorsal view; 14, genitalia, ventral view; 15, phallus, lateral view; 16, phallus, ventral view." httpUri="https://zenodo.org/record/157031/files/figure.png" pageId="8" pageNumber="9">12</figureCitation>
); basimesal lobe cone­shaped (
<figureCitation id="63FC2A7CFFE5FFD62AF6E15BFB24854F" box="[1187,1263,604,630]" captionStart="FIGURES 1 ­ 8" captionStartId="2.[264,382,1813,1837]" captionTargetBox="[266,1307,465,1791]" captionTargetId="figure@2.[249,1324,449,1803]" captionTargetPageId="2" captionText="FIGURES 1 ­ 8. Helicopsyche woldai sp. n., male holotype. 1, right wings; 2, abdominal segment VI, lateral view; 3, abdominal VIth sternal process, ventral view; 4, genitalia, lateral view; 5, genitalia, dorsal view; 6, genitalia, ventral view; 7, phallus, lateral view; 8, phallus, ventral view." httpUri="https://zenodo.org/record/157030/files/figure.png" pageId="8" pageNumber="9">Figs 6</figureCitation>
,
<figureCitation id="63FC2A7CFFE5FFD62AAEE15BFAD0854F" box="[1275,1307,604,630]" captionStart="FIGURES 9 ­ 16" captionStartId="5.[264,382,1617,1641]" captionTargetBox="[272,1322,318,1589]" captionTargetId="figure@5.[268,1343,302,1602]" captionTargetPageId="5" captionText="FIGURES 9 ­ 16. Helicopsyche fridae sp. n., male holotype. 9, right wings; 10, abdominal segment VI, lateral view; 11, abdominal VIth sternal process, ventral view; 12, genitalia, lateral view; 13, genitalia, dorsal view; 14, genitalia, ventral view; 15, phallus, lateral view; 16, phallus, ventral view." httpUri="https://zenodo.org/record/157031/files/figure.png" pageId="8" pageNumber="9">14</figureCitation>
); phallus strongly curving ventrally at midlength (
<figureCitation id="63FC2A7CFFE5FFD62D38E183FC7185A7" box="[877,954,644,670]" captionStart="FIGURES 1 ­ 8" captionStartId="2.[264,382,1813,1837]" captionTargetBox="[266,1307,465,1791]" captionTargetId="figure@2.[249,1324,449,1803]" captionTargetPageId="2" captionText="FIGURES 1 ­ 8. Helicopsyche woldai sp. n., male holotype. 1, right wings; 2, abdominal segment VI, lateral view; 3, abdominal VIth sternal process, ventral view; 4, genitalia, lateral view; 5, genitalia, dorsal view; 6, genitalia, ventral view; 7, phallus, lateral view; 8, phallus, ventral view." httpUri="https://zenodo.org/record/157030/files/figure.png" pageId="8" pageNumber="9">Figs 7</figureCitation>
,
<figureCitation id="63FC2A7CFFE5FFD62D93E183FC2F85A7" box="[966,996,644,670]" captionStart="FIGURES 9 ­ 16" captionStartId="5.[264,382,1617,1641]" captionTargetBox="[272,1322,318,1589]" captionTargetId="figure@5.[268,1343,302,1602]" captionTargetPageId="5" captionText="FIGURES 9 ­ 16. Helicopsyche fridae sp. n., male holotype. 9, right wings; 10, abdominal segment VI, lateral view; 11, abdominal VIth sternal process, ventral view; 12, genitalia, lateral view; 13, genitalia, dorsal view; 14, genitalia, ventral view; 15, phallus, lateral view; 16, phallus, ventral view." httpUri="https://zenodo.org/record/157031/files/figure.png" pageId="8" pageNumber="9">15</figureCitation>
) ....................................... 3
</keyLead>
</paragraph>
</keyStep>
<keyStep id="40332E5CFFE5FFD62F5DE1ABFAE6845F" pageId="8" pageNumber="9">
<paragraph id="FB7836F9FFE5FFD62F5DE1ABFAE1842F" blockId="8.[264,1325,284,1110]" pageId="8" pageNumber="9">
<keyLead id="403695CCFFE5FFD62F5DE1ABFAE1842F" pageId="8" pageNumber="9">
3 Gonocoxite sharply incised toward basimesal lobe (
<figureCitation id="63FC2A7CFFE5FFD62DCCE1ABFC3A85FF" box="[921,1009,684,710]" captionStart="FIGURES 9 ­ 16" captionStartId="5.[264,382,1617,1641]" captionTargetBox="[272,1322,318,1589]" captionTargetId="figure@5.[268,1343,302,1602]" captionTargetPageId="5" captionText="FIGURES 9 ­ 16. Helicopsyche fridae sp. n., male holotype. 9, right wings; 10, abdominal segment VI, lateral view; 11, abdominal VIth sternal process, ventral view; 12, genitalia, lateral view; 13, genitalia, dorsal view; 14, genitalia, ventral view; 15, phallus, lateral view; 16, phallus, ventral view." httpUri="https://zenodo.org/record/157031/files/figure.png" pageId="8" pageNumber="9">Fig. 12</figureCitation>
); gonocoxite with basimesal lobe very long (
<figureCitation id="63FC2A7CFFE5FFD62C48E1D3FDBE85D7" box="[541,629,724,750]" captionStart="FIGURES 9 ­ 16" captionStartId="5.[264,382,1617,1641]" captionTargetBox="[272,1322,318,1589]" captionTargetId="figure@5.[268,1343,302,1602]" captionTargetPageId="5" captionText="FIGURES 9 ­ 16. Helicopsyche fridae sp. n., male holotype. 9, right wings; 10, abdominal segment VI, lateral view; 11, abdominal VIth sternal process, ventral view; 12, genitalia, lateral view; 13, genitalia, dorsal view; 14, genitalia, ventral view; 15, phallus, lateral view; 16, phallus, ventral view." httpUri="https://zenodo.org/record/157031/files/figure.png" pageId="8" pageNumber="9">Fig. 12</figureCitation>
); gonocoxite sharply triangular (
<figureCitation id="63FC2A7CFFE5FFD62DADE1D3FB9B85D7" box="[1016,1104,724,750]" captionStart="FIGURES 9 ­ 16" captionStartId="5.[264,382,1617,1641]" captionTargetBox="[272,1322,318,1589]" captionTargetId="figure@5.[268,1343,302,1602]" captionTargetPageId="5" captionText="FIGURES 9 ­ 16. Helicopsyche fridae sp. n., male holotype. 9, right wings; 10, abdominal segment VI, lateral view; 11, abdominal VIth sternal process, ventral view; 12, genitalia, lateral view; 13, genitalia, dorsal view; 14, genitalia, ventral view; 15, phallus, lateral view; 16, phallus, ventral view." httpUri="https://zenodo.org/record/157031/files/figure.png" pageId="8" pageNumber="9">Fig. 12</figureCitation>
); posterior margin of gonocoxite with prominent processes (
<figureCitation id="63FC2A7CFFE5FFD62D49E1FBFCBA842F" box="[796,881,764,790]" captionStart="FIGURES 9 ­ 16" captionStartId="5.[264,382,1617,1641]" captionTargetBox="[272,1322,318,1589]" captionTargetId="figure@5.[268,1343,302,1602]" captionTargetPageId="5" captionText="FIGURES 9 ­ 16. Helicopsyche fridae sp. n., male holotype. 9, right wings; 10, abdominal segment VI, lateral view; 11, abdominal VIth sternal process, ventral view; 12, genitalia, lateral view; 13, genitalia, dorsal view; 14, genitalia, ventral view; 15, phallus, lateral view; 16, phallus, ventral view." httpUri="https://zenodo.org/record/157031/files/figure.png" pageId="8" pageNumber="9">Fig. 12</figureCitation>
) .................................
<taxonomicName id="3CC74D7AFFE5FFD62AD7E1FBFB27842C" box="[1154,1260,764,789]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="fridae" status="sp. nov.">
<emphasis id="C9B3EAEBFFE5FFD62AD7E1FBFB27842C" box="[1154,1260,764,789]" italics="true" pageId="8" pageNumber="9">H. fridae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D2805790FFE5FFD62AA6E1FBFAE1842F" box="[1267,1322,764,790]" pageId="8" pageNumber="9" rank="species">sp.n.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="FB7836F9FFE5FFD62F5DE023FAE6845F" blockId="8.[264,1325,284,1110]" pageId="8" pageNumber="9">
<keyLead id="403695CCFFE5FFD62F5DE023FAE6845F" pageId="8" pageNumber="9">
­ Gonocoxite widely incised toward basimesal lobe (
<figureCitation id="63FC2A7CFFE5FFD62DC2E023FC298407" box="[919,994,804,830]" captionStart="FIGURES 1 ­ 8" captionStartId="2.[264,382,1813,1837]" captionTargetBox="[266,1307,465,1791]" captionTargetId="figure@2.[249,1324,449,1803]" captionTargetPageId="2" captionText="FIGURES 1 ­ 8. Helicopsyche woldai sp. n., male holotype. 1, right wings; 2, abdominal segment VI, lateral view; 3, abdominal VIth sternal process, ventral view; 4, genitalia, lateral view; 5, genitalia, dorsal view; 6, genitalia, ventral view; 7, phallus, lateral view; 8, phallus, ventral view." httpUri="https://zenodo.org/record/157030/files/figure.png" pageId="8" pageNumber="9">Fig. 4</figureCitation>
); posterior margin of gonocoxite with row of prominent processes (
<figureCitation id="63FC2A7CFFE5FFD62D42E04BFC94845F" box="[791,863,844,870]" captionStart="FIGURES 1 ­ 8" captionStartId="2.[264,382,1813,1837]" captionTargetBox="[266,1307,465,1791]" captionTargetId="figure@2.[249,1324,449,1803]" captionTargetPageId="2" captionText="FIGURES 1 ­ 8. Helicopsyche woldai sp. n., male holotype. 1, right wings; 2, abdominal segment VI, lateral view; 3, abdominal VIth sternal process, ventral view; 4, genitalia, lateral view; 5, genitalia, dorsal view; 6, genitalia, ventral view; 7, phallus, lateral view; 8, phallus, ventral view." httpUri="https://zenodo.org/record/157030/files/figure.png" pageId="8" pageNumber="9">Fig. 4</figureCitation>
) or processes are lacking ................. 4
</keyLead>
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<keyStep id="40332E5CFFE5FFD62F5DE073FAE0836F" pageId="8" pageNumber="9">
<paragraph id="FB7836F9FFE5FFD62F5DE073FAE184E7" blockId="8.[264,1325,284,1110]" pageId="8" pageNumber="9">
<keyLead id="403695CCFFE5FFD62F5DE073FAE184E7" pageId="8" pageNumber="9">
4 Gonocoxite (
<figureCitation id="63FC2A7CFFE5FFD62F85E073FDDD84B7" box="[464,534,884,910]" captionStart="FIGURES 1 ­ 8" captionStartId="2.[264,382,1813,1837]" captionTargetBox="[266,1307,465,1791]" captionTargetId="figure@2.[249,1324,449,1803]" captionTargetPageId="2" captionText="FIGURES 1 ­ 8. Helicopsyche woldai sp. n., male holotype. 1, right wings; 2, abdominal segment VI, lateral view; 3, abdominal VIth sternal process, ventral view; 4, genitalia, lateral view; 5, genitalia, dorsal view; 6, genitalia, ventral view; 7, phallus, lateral view; 8, phallus, ventral view." httpUri="https://zenodo.org/record/157030/files/figure.png" pageId="8" pageNumber="9">Fig. 4</figureCitation>
) parallelogram­shaped, and with row of processes at posteroventral margin; basimesal lobe (
<figureCitation id="63FC2A7CFFE5FFD62C0EE09BFD69848F" box="[603,674,924,950]" captionStart="FIGURES 1 ­ 8" captionStartId="2.[264,382,1813,1837]" captionTargetBox="[266,1307,465,1791]" captionTargetId="figure@2.[249,1324,449,1803]" captionTargetPageId="2" captionText="FIGURES 1 ­ 8. Helicopsyche woldai sp. n., male holotype. 1, right wings; 2, abdominal segment VI, lateral view; 3, abdominal VIth sternal process, ventral view; 4, genitalia, lateral view; 5, genitalia, dorsal view; 6, genitalia, ventral view; 7, phallus, lateral view; 8, phallus, ventral view." httpUri="https://zenodo.org/record/157030/files/figure.png" pageId="8" pageNumber="9">Fig. 6</figureCitation>
) with smooth lateral and median margins, and bearing about three stout megasetae
<emphasis id="C9B3EAEBFFE5FFD62CD1E0C3FB2784E4" box="[644,1260,964,989]" italics="true" pageId="8" pageNumber="9">
....................................................................
<taxonomicName id="3CC74D7AFFE5FFD62A2CE0C3FB2784E4" box="[1145,1260,964,989]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="woldai" status="sp. nov.">H. woldai</taxonomicName>
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</paragraph>
<paragraph id="FB7836F9FFE5FFD62F5DE0EBFAE0836F" blockId="8.[264,1325,284,1110]" pageId="8" pageNumber="9">
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­ Gonocoxite widens slightly distally; posteroventral margin without processes; basimesal lobe with irregular median and lateral margins associated with numerous setae and no stout megasetae .................................................................................
<taxonomicName id="3CC74D7AFFE5FFD62AD5E73BFAE0836F" ID-CoL="7VC9S" authority="Ross" authorityName="Ross" box="[1152,1323,1084,1110]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="incisa">
<emphasis id="C9B3EAEBFFE5FFD62AD5E73BFB21836C" box="[1152,1258,1084,1109]" italics="true" pageId="8" pageNumber="9">H. incisa</emphasis>
Ross
</taxonomicName>
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