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<document id="6DE9A403EE6FFDA4DB75603489715F14" ID-DOI="10.11646/zootaxa.1161.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="5058330" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1625190197324" checkinUser="felipe" docAuthor="Hippa, Heikki" docDate="2006" docId="5D4C87852866FF8C5C1A0860FEB3FAA1" docLanguage="en" docName="zt01161p049.pdf" docOrigin="Zootaxa 1161 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.1161.1.1" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Manota curvata Hippa 2006, sp. n." docType="treatment" docVersion="4" lastPageNumber="36" masterDocId="A175FFFD2844FFAF5D120E76FFC5FFE7" masterDocTitle="Diversity of Manota Williston (Diptera: Mycetophilidae) in a Malaysian rainforest: description of twenty-seven new sympatric species" masterLastPageNumber="49" masterPageNumber="1" pageNumber="35" updateTime="1698983105622" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="9305223AC2E6292F162D782A7BDF8F56" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="11FE7C38EA4FDF537B4D557EDDB4A22D">Diversity of Manota Williston (Diptera: Mycetophilidae) in a Malaysian rainforest: description of twenty-seven new sympatric species</mods:title>
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<mods:namePart id="D7A133DE51B848EB7833A28E6CEB7ED7">Hippa, Heikki</mods:namePart>
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<mods:title id="D19C697265E4E7091A18A823B8CEFC00">Zootaxa</mods:title>
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<mods:date id="9622D191B9DFF7292B578D7C502DE734">2006</mods:date>
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<mods:number id="6BBBF8DF6DC6346DAD02271834362056">2006-03-30</mods:number>
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<mods:number id="74D621CECD911EF1788A9975E32D15C5">1161</mods:number>
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<mods:url id="35C9B876D08B056724C27E2EE72AA2E6">https://biotaxa.org/Zootaxa/article/view/zootaxa.1161.1.1</mods:url>
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<mods:classification id="985F1DE7D915B39856C73ED7E17A8AB3">journal article</mods:classification>
<mods:identifier id="D0448AEB60689600AFC04B736285BDE7" type="DOI">10.11646/zootaxa.1161.1.1</mods:identifier>
<mods:identifier id="59F7301F54540675026DD4A9921B97AC" type="ISSN">1175­5334</mods:identifier>
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<treatment id="5D4C87852866FF8C5C1A0860FEB3FAA1" ID-DOI="http://doi.org/10.5281/zenodo.5059324" ID-Zenodo-Dep="5059324" LSID="urn:lsid:plazi:treatment:5D4C87852866FF8C5C1A0860FEB3FAA1" httpUri="http://treatment.plazi.org/id/5D4C87852866FF8C5C1A0860FEB3FAA1" lastPageId="35" lastPageNumber="36" pageId="34" pageNumber="35">
<subSubSection id="9DFF65182866FF8D5C1A0860FDD7F9D7" box="[264,530,1558,1584]" pageId="34" pageNumber="35" type="nomenclature">
<paragraph id="D55A36932866FF8D5C1A0860FDD7F9D7" blockId="34.[264,530,1558,1619]" box="[264,530,1558,1584]" pageId="34" pageNumber="35">
<heading id="8E1281FF2866FF8D5C1A0860FDD7F9D7" bold="true" box="[264,530,1558,1584]" fontSize="11" level="1" pageId="34" pageNumber="35" reason="1">
<emphasis id="E791EA812866FF8D5C1A0860FDD7F9D7" bold="true" box="[264,530,1558,1584]" pageId="34" pageNumber="35">
<taxonomicName id="12E54D102866FF8D5C1A0860FE0DF9D7" ID-CoL="3XVVP" authority="Hippa, 2006" authorityName="Hippa" authorityYear="2006" box="[264,456,1558,1584]" class="Insecta" family="Mycetophilidae" genus="Manota" kingdom="Animalia" order="Diptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="curvata" status="sp. nov.">
<emphasis id="E791EA812866FF8D5C1A0860FE0DF9D7" bold="true" box="[264,456,1558,1584]" italics="true" pageId="34" pageNumber="35">Manota curvata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FCA257FA2866FF8D5CC20860FDD7F9D7" box="[464,530,1558,1584]" pageId="34" pageNumber="35" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="9DFF65182866FF8C5C1A084FFEB3FAA1" lastPageId="35" lastPageNumber="36" pageId="34" pageNumber="35" type="description">
<paragraph id="D55A36932866FF8D5C1A084FFE14F9B4" blockId="34.[264,530,1558,1619]" box="[264,465,1593,1619]" pageId="34" pageNumber="35">
(
<figureCitation id="4DDE2A162866FF8D5C03084FFE0CF9B4" box="[273,457,1593,1619]" captionStart="FIGURE 13" captionStartId="36.[264,367,1753,1777]" captionTargetBox="[291,1293,332,1691]" captionTargetId="figure-66@36.[263,1322,297,1747]" captionTargetPageId="36" captionText="FIGURE 13. Antennal flagellomere 4, lateral view (A, C); hypopygium, dorsal (B, E) and ventral view (D). A, B. Manota globigera sp. n. (holotype). C, D, E. Manota curvata sp. n. (D holotype, E paratype). Scale for A and C 0.05 mm; for B, D, and E 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.5058370" httpUri="https://zenodo.org/record/5058370/files/figure.png" pageId="34" pageNumber="35">Figs. 13C, D, E</figureCitation>
)
</paragraph>
<paragraph id="D55A36932866FF8D5C1A08FFFDD6F944" blockId="34.[264,1322,1673,1899]" box="[264,531,1673,1699]" pageId="34" pageNumber="35">
A small­sized
<taxonomicName id="12E54D102866FF8D5CA308FFFDC9F945" authorityName="Williston" authorityYear="1896" box="[433,524,1673,1698]" class="Insecta" family="Mycetophilidae" genus="Manota" kingdom="Animalia" order="Diptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="E791EA812866FF8D5CA308FFFDC9F945" box="[433,524,1673,1698]" italics="true" pageId="34" pageNumber="35">Manota</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="D55A36932866FF8C5C2A08C7FEA3FD21" blockId="34.[264,1322,1673,1899]" lastBlockId="35.[264,1322,283,750]" lastPageId="35" lastPageNumber="36" pageId="34" pageNumber="35">
Male.
<emphasis id="E791EA812866FF8D5C9408C7FE15F92C" bold="true" box="[390,464,1713,1739]" pageId="34" pageNumber="35">Head.</emphasis>
Flagellomere 4,
<figureCitation id="4DDE2A162866FF8D5FB308C7FCCBF92C" box="[673,782,1713,1739]" captionStart="FIGURE 13" captionStartId="36.[264,367,1753,1777]" captionTargetBox="[291,1293,332,1691]" captionTargetId="figure-66@36.[263,1322,297,1747]" captionTargetPageId="36" captionText="FIGURE 13. Antennal flagellomere 4, lateral view (A, C); hypopygium, dorsal (B, E) and ventral view (D). A, B. Manota globigera sp. n. (holotype). C, D, E. Manota curvata sp. n. (D holotype, E paratype). Scale for A and C 0.05 mm; for B, D, and E 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.5058370" httpUri="https://zenodo.org/record/5058370/files/figure.png" pageId="34" pageNumber="35">Fig. 13C</figureCitation>
. Maxillary palpus with palpomere 3 bearing 5–6 apically expanded curved sensilla; palpomere 4 with parasegment.
<emphasis id="E791EA812866FF8D59D408AFFAEFF914" bold="true" box="[1222,1322,1753,1779]" pageId="34" pageNumber="35">Thorax.</emphasis>
Anepisternum setose. Anterior basalare non­setose. Preepisternum 2 setose. Laterotergite non­setose. Episternum 3 setose.
<emphasis id="E791EA812866FF8D5FA5095FFD3EF8A4" bold="true" box="[695,763,1833,1859]" pageId="34" pageNumber="35">Wing</emphasis>
. Length
<quantity id="121D9B762866FF8D5E77095FFC33F8A4" box="[869,1014,1833,1859]" metricMagnitude="-3" metricUnit="m" metricValue="1.5499999999999998" metricValueMax="1.7" metricValueMin="1.4" pageId="34" pageNumber="35" unit="mm" value="1.5499999999999998" valueMax="1.7" valueMin="1.4">1.4–1.7 mm</quantity>
. Wing membrane dorsally with few setae at posterior margin. Sc distally of h non­setose. A1 indicated by few setae.
<emphasis id="E791EA812867FF8C5C1A0F6DFE6FFED2" bold="true" box="[264,426,283,309]" pageId="35" pageNumber="36">Hypopygium</emphasis>
(
<figureCitation id="4DDE2A162867FF8C5CAF0F6AFD9EFED1" box="[445,603,284,310]" captionStart="FIGURE 13" captionStartId="36.[264,367,1753,1777]" captionTargetBox="[291,1293,332,1691]" captionTargetId="figure-66@36.[263,1322,297,1747]" captionTargetPageId="36" captionText="FIGURE 13. Antennal flagellomere 4, lateral view (A, C); hypopygium, dorsal (B, E) and ventral view (D). A, B. Manota globigera sp. n. (holotype). C, D, E. Manota curvata sp. n. (D holotype, E paratype). Scale for A and C 0.05 mm; for B, D, and E 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.5058370" httpUri="https://zenodo.org/record/5058370/files/figure.png" pageId="35" pageNumber="36">Figs. 13D, E</figureCitation>
). Sternite 9 large, extending posteriorly over half of ventral length of gonocoxa, laterally and posteriorly sharply delimited, almost round, setae similar to adjacent ventral setae of gonocoxa except at posterior margin where they are twice as large. The dorsomesial margin of gonocoxa simple, in position III with 2 curved or sigmoid simple megasetae arising from common rather low basal body; gonocoxa apicolaterally with tapering, mesially setose, subtriangular lobe, some setae conspicuously flattened. Gonostylus long, narrow, strongly curved in outline, apicodorsally with crest with strong setae. Parastylar lobe present, broad, without distinct apex, with 2 or 3 setae directed transversely mesiad. Tegmen rather short and broad, subtriangular, with distinct shoulders. Setae on sternite
<quantity id="121D9B762867FF8C5F770CF2FD63FD79" box="[613,678,644,670]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="35" pageNumber="36" unit="in" value="10.0">10 in</quantity>
3 rows on each half, mostly posteriorly of apex of tegmen.
</paragraph>
<paragraph id="D55A36932867FF8C5C2A0CA2FCDBFD09" blockId="35.[264,1322,283,750]" box="[312,798,724,750]" pageId="35" pageNumber="36">Female and preimaginal stages unknown.</paragraph>
<paragraph id="D55A36932867FF8C5C1A0D52FE4DFCDA" blockId="35.[264,1323,804,1070]" box="[264,392,804,829]" pageId="35" pageNumber="36">
<emphasis id="E791EA812867FF8C5C1A0D52FE4DFCDA" box="[264,392,804,829]" italics="true" pageId="35" pageNumber="36">Discussion</emphasis>
</paragraph>
<paragraph id="D55A36932867FF8C5C2A0D3AFE35FBC9" blockId="35.[264,1323,804,1070]" pageId="35" pageNumber="36">
<taxonomicName id="12E54D102867FF8C5C2A0D3AFE30FC82" authorityName="Hippa" authorityYear="2006" box="[312,501,844,869]" class="Insecta" family="Mycetophilidae" genus="Manota" kingdom="Animalia" order="Diptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="curvata">
<emphasis id="E791EA812867FF8C5C2A0D3AFE30FC82" box="[312,501,844,869]" italics="true" pageId="35" pageNumber="36">Manota curvata</emphasis>
</taxonomicName>
is not especially similar to any of the other species. Its hypopygium resembles that of many of the species that have an apicolateral subtriangular lobe on the gonocoxa, especially those that also have the dorsal apicomesial corner of the gonocoxa unmodified and rounded (e.g.,
<taxonomicName id="12E54D102867FF8C5F690DB2FD33FC3A" authorityName="Hippa" authorityYear="2006" box="[635,758,964,989]" class="Insecta" family="Mycetophilidae" genus="Manota" kingdom="Animalia" order="Diptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="ferrata">
<emphasis id="E791EA812867FF8C5F690DB2FD33FC3A" box="[635,758,964,989]" italics="true" pageId="35" pageNumber="36">M. ferrata</emphasis>
</taxonomicName>
).
<taxonomicName id="12E54D102867FF8C5E020DB2FC0BFC3A" authorityName="Hippa" authorityYear="2006" box="[784,974,964,989]" class="Insecta" family="Mycetophilidae" genus="Manota" kingdom="Animalia" order="Diptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="curvata">
<emphasis id="E791EA812867FF8C5E020DB2FC0BFC3A" box="[784,974,964,989]" italics="true" pageId="35" pageNumber="36">Manota curvata</emphasis>
</taxonomicName>
differs from all such species by its narrow, curved gonostylus and by having the setae on the apicolateral lobe of the gonocoxa flattened.
</paragraph>
<paragraph id="D55A36932867FF8C5C1A0A12FE8FFB9A" blockId="35.[264,1323,1124,1350]" box="[264,330,1124,1149]" pageId="35" pageNumber="36">
<emphasis id="E791EA812867FF8C5C1A0A12FE8FFB9A" box="[264,330,1124,1149]" italics="true" pageId="35" pageNumber="36">Types</emphasis>
</paragraph>
<paragraph id="D55A36932867FF8C5C2A0AFAFEB3FAA1" blockId="35.[264,1323,1124,1350]" pageId="35" pageNumber="36">
<materialsCitation id="658D3CCE2867FF8C5C2A0AFAFE7BFB11" collectingDate="1997-02-24" collectingDateMax="1997-03-21" collectingDateMin="1997-02-24" collectorName="H. Hippa &amp; M. Jaschhof &amp; B. Viklund" country="Malaysia" county="Male" elevation="244" location="Ulu Gombak" municipality="West" pageId="35" pageNumber="36" specimenCount="1" stateProvince="Selangor" typeStatus="holotype">
<emphasis id="E791EA812867FF8C5C2A0AFAFE66FB42" box="[312,419,1164,1189]" italics="true" pageId="35" pageNumber="36">
<typeStatus id="0A5E88312867FF8C5C2A0AFAFE66FB42" box="[312,419,1164,1189]" pageId="35" pageNumber="36" type="holotype">Holotype</typeStatus>
</emphasis>
.
<collectingCounty id="3C3B4E1F2867FF8C5CA60AFAFE36FB41" box="[436,499,1164,1190]" pageId="35" pageNumber="36">Male</collectingCounty>
,
<collectingMunicipality id="353EACE92867FF8C5F100AFAFDF9FB41" box="[514,572,1164,1190]" pageId="35" pageNumber="36">West</collectingMunicipality>
<collectingCountry id="ADF276032867FF8C5F570AFAFD76FB41" box="[581,691,1164,1190]" name="Malaysia" pageId="35" pageNumber="36">Malaysia</collectingCountry>
,
<collectingRegion id="1721F8712867FF8C5FD30AFAFCEEFB41" box="[705,811,1164,1190]" country="Malaysia" name="Selangor" pageId="35" pageNumber="36">Selangor</collectingRegion>
,
<location id="D03A60482867FF8C5E280AFAFC12FB41" LSID="urn:lsid:plazi:treatment:5D4C87852866FF8C5C1A0860FEB3FAA1:D03A60482867FF8C5E280AFAFC12FB41" box="[826,983,1164,1190]" country="Malaysia" county="Male" municipality="West" name="Ulu Gombak" pageId="35" pageNumber="36" stateProvince="Selangor">Ulu Gombak</location>
,
<location id="D03A60482867FF8C5EF70AFAFE60FB29" LSID="urn:lsid:plazi:treatment:5D4C87852866FF8C5C1A0860FEB3FAA1:D03A60482867FF8C5EF70AFAFE60FB29" country="Malaysia" county="Male" municipality="West" name="University of Malaya Field Study Centre" pageId="35" pageNumber="36" stateProvince="Selangor">University of Malaya Field Study Centre</location>
,
<quantity id="121D9B762867FF8C5CA10AC2FE39FB29" box="[435,508,1204,1230]" metricMagnitude="2" metricUnit="m" metricValue="2.4384" pageId="35" pageNumber="36" unit="ft" value="800.0">
<elevation id="5EC8D1A02867FF8C5CA10AC2FE39FB29" box="[435,508,1204,1230]" metricMagnitude="2" metricUnit="m" metricValue="2.4384" pageId="35" pageNumber="36" unit="ft" value="800.0">800 ft</elevation>
</quantity>
, Malaise in jungle,
<date id="A15B10532867FF8C5FFA0AC2FBCAFB29" box="[744,1039,1204,1230]" pageId="35" pageNumber="36" value="1997-02-24" valueMax="1997-03-21" valueMin="1997-02-24">
<collectingDate id="B11FE9BB2867FF8C5FFA0AC2FBCAFB29" box="[744,1039,1204,1230]" pageId="35" pageNumber="36" value="1997-02-24" valueMax="1997-03-21" valueMin="1997-02-24">24 Feb.–21. March 1997</collectingDate>
</date>
,
<collectorName id="781053452867FF8C590F0AC2FB4BFB29" box="[1053,1166,1204,1230]" pageId="35" pageNumber="36">H. Hippa</collectorName>
,
<collectorName id="781053452867FF8C59880AC2FAEFFB29" box="[1178,1322,1204,1230]" pageId="35" pageNumber="36">M. Jaschhof</collectorName>
and
<collectorName id="781053452867FF8C5C280AAAFE7BFB11" box="[314,446,1244,1270]" pageId="35" pageNumber="36">B. Viklund</collectorName>
</materialsCitation>
.
<materialsCitation id="658D3CCE2867FF8C5CDB0AAAFC29FB11" box="[457,1004,1244,1270]" collectingDate="1997-02-24" collectingDateMax="1997-03-21" collectingDateMin="1997-02-24" collectorName="H. Hippa &amp; M. Jaschhof &amp; B. Viklund" country="Malaysia" county="Male" elevation="244" location="Ulu Gombak" municipality="West" pageId="35" pageNumber="36" specimenCount="7" specimenCount-male="7" stateProvince="Selangor" typeStatus="paratype">
<typeStatus id="0A5E88312867FF8C5CDB0AAAFDFAFB11" box="[457,575,1244,1270]" pageId="35" pageNumber="36" type="paratype">Paratypes</typeStatus>
.
<specimenCount id="C3E3FD1A2867FF8C5F580AAAFD61FB11" box="[586,676,1244,1270]" pageId="35" pageNumber="36" type="male">7 males</specimenCount>
with same data as holotype
</materialsCitation>
;
<materialsCitation id="658D3CCE2867FF8C5EEA0AAAFEABFAF9" collectingDate="1997-02-24" collectingDateMax="1997-03-21" collectingDateMin="1997-02-24" collectorName="H. Hippa &amp; M. Jaschhof &amp; B. Viklund" country="Malaysia" county="Male" elevation="244" location="Ulu Gombak" municipality="West" pageId="35" pageNumber="36" specimenCount="6" specimenCount-male="6" stateProvince="Selangor" typeStatus="paratype">
<specimenCount id="C3E3FD1A2867FF8C5EEA0AAAFB97FB11" box="[1016,1106,1244,1270]" pageId="35" pageNumber="36" type="male">6 males</specimenCount>
with same data as holotype
</materialsCitation>
except for 22. Feb.–21. March.
<materialsCitation id="658D3CCE2867FF8C5E030B72FEB7FAA1" collectingDate="1997-02-24" collectingDateMax="1997-03-21" collectingDateMin="1997-02-24" collectorName="H. Hippa &amp; M. Jaschhof &amp; B. Viklund" country="Malaysia" county="Male" elevation="244" location="Ulu Gombak" municipality="West" pageId="35" pageNumber="36" specimenCount="2" specimenCount-male="2" stateProvince="Selangor">
Other material.
<specimenCount id="C3E3FD1A2867FF8C5EC50B72FBFDFAF9" box="[983,1080,1284,1310]" pageId="35" pageNumber="36" type="male">2 males</specimenCount>
with same data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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