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<document id="D7720190D703370452D6BB39BD5ACBF7" ID-CLB-Dataset="29801" ID-DOI="10.11646/zootaxa.4442.1.4" ID-GBIF-Dataset="0f0f11d9-27bb-487e-93e0-6c33ec5958b6" ID-ISSN="1175-5326" ID-Zenodo-Dep="1301366" ID-ZooBank="937D9736-2646-45D6-8C5D-3625A88FF2FB" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1530293686360" checkinUser="plazi" docAuthor="Grubbs, Scott A. &amp; Dewalt, R. Edward" docDate="2018" docId="451FB544C53BAA70FF6EFF56557C2337" docLanguage="en" docName="zootaxa.4442.1.4.pdf" docOrigin="Zootaxa 4442 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Perlesta xube Stark &amp; Rhodes 1997" docType="treatment" docVersion="8" lastPageNumber="86" masterDocId="B926CD3CC538AA73FFF9FFC1567B2715" masterDocTitle="Perlesta armitagei n. sp. (Plecoptera: Perlidae): More cryptic diversity in darkly pigmented Perlesta from the eastern Nearctic" masterLastPageNumber="100" masterPageNumber="83" pageNumber="86" updateTime="1698551734473" updateUser="ExternalLinkService">
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<mods:title id="41D3E5F80C15C24F78F32D4FD9139508">Perlesta armitagei n. sp. (Plecoptera: Perlidae): More cryptic diversity in darkly pigmented Perlesta from the eastern Nearctic</mods:title>
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<mods:namePart id="2E6EB93EC0BD065FD6C8B6F89482FBA4">Grubbs, Scott A.</mods:namePart>
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<mods:date id="3CCBA6001A9E062EB88C1F21E951ED83">2018</mods:date>
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<treatment id="451FB544C53BAA70FF6EFF56557C2337" ID-DOI="http://doi.org/10.5281/zenodo.4681384" ID-GBIF-Taxon="146043549" ID-Zenodo-Dep="4681384" LSID="urn:lsid:plazi:treatment:451FB544C53BAA70FF6EFF56557C2337" httpUri="http://treatment.plazi.org/id/451FB544C53BAA70FF6EFF56557C2337" lastPageNumber="86" pageId="3" pageNumber="86">
<subSubSection id="85AC57D9C53BAA70FF6EFF56574F27C6" pageId="3" pageNumber="86" type="nomenclature">
<paragraph id="CD090452C53BAA70FF6EFF56543727A7" blockId="3.[151,1023,151,283]" box="[151,588,151,178]" pageId="3" pageNumber="86">
<heading id="9641B33EC53BAA70FF6EFF56543727A7" bold="true" box="[151,588,151,178]" fontSize="11" level="1" pageId="3" pageNumber="86" reason="1">
<taxonomicName id="0AB67FD1C53BAA70FF6EFF56543727A7" ID-CoL="4F6H5" authority="Stark &amp; Rhodes 1997" authorityName="Stark &amp; Rhodes" authorityYear="1997" box="[151,588,151,178]" class="Insecta" family="Perlidae" genus="Perlesta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="86" phylum="Arthropoda" rank="species" species="xube">
<emphasis id="FFC2D840C53BAA70FF6EFF56543727A7" bold="true" box="[151,588,151,178]" pageId="3" pageNumber="86">
<emphasis id="FFC2D840C53BAA70FF6EFF56574127A4" bold="true" box="[151,314,151,177]" italics="true" pageId="3" pageNumber="86">Perlesta xube</emphasis>
<bibRefCitation id="A92779A3C53BAA70FEB8FF59543727A7" box="[321,588,152,178]" pageId="3" pageNumber="86" refId="ref8652" refString="Stark, B. P. &amp; Rhodes, H. A. (1997) Perlesta xube, a new stonefly species from Nebraska (Plecoptera: Perlidae). Entomological News, 108, 92 - 96. Avaliable from: https: // biodiversitylibrary. org / page / 16343581 (Accessed 28 Jun. 2018)" type="journal article">Stark &amp; Rhodes 1997</bibRefCitation>
</emphasis>
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</paragraph>
<paragraph id="CD090452C53BAA70FF6EFF7A574F27C6" blockId="3.[151,1023,151,283]" box="[151,308,187,211]" pageId="3" pageNumber="86">Pawnee Stone</paragraph>
</subSubSection>
<subSubSection id="85AC57D9C53BAA70FF6EFF1F574F260E" pageId="3" pageNumber="86" type="description">
<paragraph id="CD090452C53BAA70FF6EFF1F558427E2" blockId="3.[151,1023,151,283]" box="[151,1023,222,247]" pageId="3" pageNumber="86">
http://lsid.speciesfile.org/urn:lsid:
<taxonomicName id="0AB67FD1C53BAA70FDF5FF1F54F127E2" box="[524,650,222,247]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="86" phylum="Arthropoda" rank="order">Plecoptera</taxonomicName>
.speciesfile.org:TaxonName:1241
</paragraph>
<paragraph id="CD090452C53BAA70FF6EFEC3574F260E" blockId="3.[151,1023,151,283]" box="[151,308,258,283]" pageId="3" pageNumber="86">
(
<figureCitation id="558D18D7C53BAA70FF66FEC35697260E" box="[159,236,258,283]" captionStart="FIGURE 2" captionStartId="4.[151,250,1876,1898]" captionTargetBox="[179,1408,393,1788]" captionTargetId="figure@4.[151,1436,367,1819]" captionTargetPageId="4" captionText="FIGURE 2. Perlesta xube, a–b, Illinois, Little Saline River; c–d, Nebraska, Cherry Creek (paratypes); 11, Ohio, Pee Pee Creek. a, c, e, male, head + pronotum, dorsal profile; b, d, female, head + pronotum, dorsal profile." httpUri="https://zenodo.org/record/1301370/files/figure.png" pageId="3" pageNumber="86">Figs. 2</figureCitation>
&amp;
<figureCitation id="558D18D7C53BAA70FEE9FEC25750260E" box="[272,299,259,283]" captionStart="FIGURE 11" captionStartId="14.[151,250,1938,1960]" captionTargetBox="[205,1400,405,1888]" captionTargetId="figure@14.[167,1431,373,1925]" captionTargetPageId="14" captionText="FIGURE 11. Distribution maps of Perlesta based on material examined during this study. a, P. adena, P. browni, P. cinctipes, and P. xube; b, P. armitagei sp. nov. The record from northern West Virginia (red circle; Tarter &amp; Nelson 2006) is conditional since that material was not available for study. The type locality is represented by the purple circle." httpUri="https://zenodo.org/record/1301388/files/figure.png" pageId="3" pageNumber="86">11</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="85AC57D9C53BAA70FF6EFE8557C5266D" pageId="3" pageNumber="86" type="reference_group">
<paragraph id="CD090452C53BAA70FF6EFE85529B264F" blockId="3.[151,1248,324,376]" box="[151,1248,324,346]" pageId="3" pageNumber="86">
<materialsCitation id="7DDE0E0FC53BAA70FF6EFE85529B264F" ID-GBIF-Occurrence="1852083302" box="[151,1248,324,346]" collectionCode="USNM" county="Cherry Co." location="Merriman" municipality="Dry Creek" pageId="3" pageNumber="86" specimenCount="1" specimenCount-male="1" stateProvince="Nebraska" typeStatus="holotype">
<treatmentCitationGroup id="EDA6237CC53BAA70FF6EFE85529B264F" box="[151,1248,324,346]" pageId="3" pageNumber="86">
<taxonomicName id="0AB67FD1C53BAA70FF6EFE85545E264F" ID-CoL="4F6H5" authority="Stark &amp; Rhodes 1997: 92" authorityName="Stark &amp; Rhodes" authorityPageNumber="92" authorityYear="1997" box="[151,549,324,346]" class="Insecta" family="Perlidae" genus="Perlesta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="86" phylum="Arthropoda" rank="species" species="xube">
<emphasis id="FFC2D840C53BAA70FF6EFE855759264F" box="[151,290,324,346]" italics="true" pageId="3" pageNumber="86">Perlesta xube</emphasis>
<treatmentCitation id="4C172243C53BAA70FED1FE85545E264F" author="" box="[296,549,324,346]" page="92" pageId="3" pageNumber="86">
<bibRefCitation id="A92779A3C53BAA70FED1FE85547D264F" box="[296,518,324,346]" pageId="3" pageNumber="86" refId="ref8652" refString="Stark, B. P. &amp; Rhodes, H. A. (1997) Perlesta xube, a new stonefly species from Nebraska (Plecoptera: Perlidae). Entomological News, 108, 92 - 96. Avaliable from: https: // biodiversitylibrary. org / page / 16343581 (Accessed 28 Jun. 2018)" type="journal article">Stark &amp; Rhodes 1997</bibRefCitation>
:92
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</taxonomicName>
.
<typeStatus id="120DBAF0C53BAA70FDD6FE8554F6264F" box="[559,653,324,346]" pageId="3" pageNumber="86" type="holotype">Holotype</typeStatus>
<specimenCount id="DBB0CFDBC53BAA70FD6AFE8454DD264F" box="[659,678,325,346]" pageId="3" pageNumber="86" type="male">♂</specimenCount>
(
<collectionCode id="ABA79C97C53BAA70FD4DFE855579264F" box="[692,770,324,346]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="3" pageNumber="86">USNM</collectionCode>
),
<collectingMunicipality id="2D6D9E28C53BAA70FCEAFE8455FB264F" box="[787,896,324,346]" pageId="3" pageNumber="86">Dry Creek</collectingMunicipality>
,
<location id="C8695289C53BAA70FC73FE855589264F" LSID="urn:lsid:plazi:treatment:451FB544C53BAA70FF6EFF56557C2337:C8695289C53BAA70FC73FE855589264F" box="[906,1010,324,346]" county="Cherry Co." municipality="Dry Creek" name="Merriman" pageId="3" pageNumber="86" stateProvince="Nebraska">Merriman</location>
,
<collectingCounty id="24687CDEC53BAA70FC05FE855214264F" box="[1020,1135,324,346]" pageId="3" pageNumber="86">Cherry Co.</collectingCounty>
,
<collectingRegion id="0F72CAB0C53BAA70FB80FE8552A7264F" box="[1145,1244,324,346]" country="United States of America" name="Nebraska" pageId="3" pageNumber="86">Nebraska</collectingRegion>
.
</treatmentCitationGroup>
</materialsCitation>
</paragraph>
<paragraph id="CD090452C53BAA70FF6EFEA357C5266D" blockId="3.[151,1248,324,376]" box="[151,446,354,376]" pageId="3" pageNumber="86">
<treatmentCitationGroup id="EDA6237CC53BAA70FF6EFEA357C5266D" box="[151,446,354,376]" pageId="3" pageNumber="86">
<taxonomicName id="0AB67FD1C53BAA70FF6EFEA357C5266D" ID-CoL="4F6H5" authority=": Stark 2004: 95" authorityName=": Stark" authorityPageNumber="95" authorityYear="2004" box="[151,446,354,376]" class="Insecta" family="Perlidae" genus="Perlesta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="86" phylum="Arthropoda" rank="species" species="xube">
<emphasis id="FFC2D840C53BAA70FF6EFEA35759266D" box="[151,290,354,376]" italics="true" pageId="3" pageNumber="86">Perlesta xube</emphasis>
:
<treatmentCitation id="4C172243C53BAA70FED6FEA357C5266D" author="" box="[303,446,354,376]" page="95" pageId="3" pageNumber="86">
<bibRefCitation id="A92779A3C53BAA70FED6FEA357DA266D" box="[303,417,354,376]" pageId="3" pageNumber="86" refId="ref8501" refString="Stark, B. P. (2004) Perlidae (The Stones). In: Stark, B. P. &amp; Armitage, B. J. (Eds.), The stoneflies (Plecoptera) of eastern North America. Vol. II. Chloroperlidae, Perlidae, and Perlodidae (Perlodinae). Bulletin of the Ohio Biological Survey. New Series. Uol. 14. Ohio Biological Society, Columbus, Ohio, pp. 61 - 148." type="journal article">Stark 2004</bibRefCitation>
:95
</treatmentCitation>
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</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="85AC57D9C53BAA70FF6EFE64523726AB" box="[151,1100,421,446]" pageId="3" pageNumber="86" type="distribution">
<paragraph id="CD090452C53BAA70FF6EFE64523726AB" blockId="3.[151,1437,421,1058]" box="[151,1100,421,446]" pageId="3" pageNumber="86">
<emphasis id="FFC2D840C53BAA70FF6EFE64574B26AB" bold="true" box="[151,304,421,446]" pageId="3" pageNumber="86">Distribution.</emphasis>
<collectingCountry id="B5A144C2C53BAA70FECEFE67570826AB" box="[311,371,422,446]" name="United States of America" pageId="3" pageNumber="86">USA</collectingCountry>
: IA, IL, ND, NE, OH (
<bibRefCitation id="A92779A3C53BAA70FD8BFE64553026AB" box="[626,843,421,446]" pageId="3" pageNumber="86" refId="ref6880" refString="DeWalt, R. E., Maehr, M. D., Neu-Becker, U. &amp; Stueber, G. (2018) Plecoptera Species File Online. Version 5.0 / 5.0. Available from: http: // Plecoptera. SpeciesFile. org (accessed 15 March 2018)" type="journal article">
DeWalt
<emphasis id="FFC2D840C53BAA70FD29FE67557326AB" box="[720,776,421,446]" italics="true" pageId="3" pageNumber="86">et al.</emphasis>
2018
</bibRefCitation>
), IN (new state record)
</paragraph>
</subSubSection>
<subSubSection id="85AC57D9C53BAA70FF3EFE08557C2337" pageId="3" pageNumber="86" type="discussion">
<paragraph id="CD090452C53BAA70FF3EFE085550245F" blockId="3.[151,1437,421,1058]" pageId="3" pageNumber="86">
<emphasis id="FFC2D840C53BAA70FF3EFE08574026F7" bold="true" box="[199,315,457,482]" pageId="3" pageNumber="86">Remarks.</emphasis>
The head masks of males and females of
<taxonomicName id="0AB67FD1C53BAA70FCD1FE0B550C26F7" authorityName=": Stark" authorityYear="2004" box="[808,887,457,482]" class="Insecta" family="Perlidae" genus="Perlesta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="86" phylum="Arthropoda" rank="species" species="xube">
<emphasis id="FFC2D840C53BAA70FCD1FE0B550C26F7" box="[808,887,457,482]" italics="true" pageId="3" pageNumber="86">P. xube</emphasis>
</taxonomicName>
(
<figureCitation id="558D18D7C53BAA70FC74FE08526826F4" box="[909,1043,457,482]" captionStart="FIGURE 2" captionStartId="4.[151,250,1876,1898]" captionTargetBox="[179,1408,393,1788]" captionTargetId="figure@4.[151,1436,367,1819]" captionTargetPageId="4" captionText="FIGURE 2. Perlesta xube, a–b, Illinois, Little Saline River; c–d, Nebraska, Cherry Creek (paratypes); 11, Ohio, Pee Pee Creek. a, c, e, male, head + pronotum, dorsal profile; b, d, female, head + pronotum, dorsal profile." httpUri="https://zenodo.org/record/1301370/files/figure.png" pageId="3" pageNumber="86">Figs. 2a‒2e</figureCitation>
) are readily distinguished from
<taxonomicName id="0AB67FD1C53BAA70FA7EFE0B577F2510" authority="Grubbs &amp; Dewalt, 2018" authorityName="Grubbs &amp; Dewalt" authorityYear="2018" class="Insecta" family="Perlidae" genus="Perlesta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="86" phylum="Arthropoda" rank="species" species="armitagei" status="sp. nov.">
<emphasis id="FFC2D840C53BAA70FA7EFE0B577F2510" italics="true" pageId="3" pageNumber="86">P. armitagei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E4F1653BC53BAA70FEE9FE2C57142513" box="[272,367,493,518]" pageId="3" pageNumber="86" rank="species">
<emphasis id="FFC2D840C53BAA70FEE9FE2C57142513" bold="true" box="[272,367,493,518]" pageId="3" pageNumber="86">sp. nov.</emphasis>
</taxonomicNameLabel>
(
<figureCitation id="558D18D7C53BAA70FE7DFE2C54742513" box="[388,527,493,518]" captionStart="FIGURE 6" captionStartId="9.[151,250,1983,2005]" captionTargetBox="[224,1362,218,1928]" captionTargetId="figure@9.[188,1398,193,1962]" captionTargetPageId="9" captionText="FIGURE 6. Perlesta armitagei sp. nov. a–b, Ohio, Deer Creek; c–d, Indiana, Blue River; e–f, Kentucky, Russell Creek; g–h, Pennsylvania, French Creek; a, c, e, g, male, head + pronotum, dorsal profile; b, d, f, h. female, head + pronotum, dorsal profile." httpUri="https://zenodo.org/record/1301378/files/figure.png" pageId="3" pageNumber="86">Figs. 6a‒6h</figureCitation>
) and
<taxonomicName id="0AB67FD1C53BAA70FDA1FE2F54A22510" baseAuthorityName="Banks" baseAuthorityYear="1905" box="[600,729,494,517]" class="Insecta" family="Perlidae" genus="Perlesta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="86" phylum="Arthropoda" rank="species" species="cinctipes">
<emphasis id="FFC2D840C53BAA70FDA1FE2F54A22510" box="[600,729,494,517]" italics="true" pageId="3" pageNumber="86">P. cinctipes</emphasis>
</taxonomicName>
(
<figureCitation id="558D18D7C53BAA70FD08FE2C55052513" box="[753,894,493,518]" captionStart="FIGURE 5" captionStartId="7.[151,250,1839,1861]" captionTargetBox="[175,1408,392,1789]" captionTargetId="figure@7.[151,1436,375,1820]" captionTargetPageId="7" captionText="FIGURE 5. Perlesta cinctipes. a–b, Arkansas, unknown locality; c–f, Missouri, South Moreau Creek; a, c, male, head + pronotum, dorsal profile; b, d, female, head + pronotum, dorsal profile; e, right paraproct, lateral view, 180X; f, right paraproct, lateral view, 330X." httpUri="https://zenodo.org/record/1301376/files/figure.png" pageId="3" pageNumber="86">Figs. 5a‒5d</figureCitation>
). The specimens (all males) determined as
<taxonomicName id="0AB67FD1C53BAA70FA7EFE2F57D6253C" authority="in Stark (2004)" authorityName="in Stark" authorityYear="2004" class="Insecta" family="Perlidae" genus="Perlesta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="86" phylum="Arthropoda" rank="species" species="cinctipes">
<emphasis id="FFC2D840C53BAA70FA7EFE2F5680253C" italics="true" pageId="3" pageNumber="86">P. cinctipes</emphasis>
in
<bibRefCitation id="A92779A3C53BAA70FED9FDD057D6253C" box="[288,429,529,554]" pageId="3" pageNumber="86" refId="ref8501" refString="Stark, B. P. (2004) Perlidae (The Stones). In: Stark, B. P. &amp; Armitage, B. J. (Eds.), The stoneflies (Plecoptera) of eastern North America. Vol. II. Chloroperlidae, Perlidae, and Perlodidae (Perlodinae). Bulletin of the Ohio Biological Survey. New Series. Uol. 14. Ohio Biological Society, Columbus, Ohio, pp. 61 - 148." type="journal article">Stark (2004)</bibRefCitation>
</taxonomicName>
were not of
<taxonomicName id="0AB67FD1C53BAA70FDC4FDD354BD253C" authority="Grubbs &amp; Dewalt, 2018" authorityName="Grubbs &amp; Dewalt" authorityYear="2018" box="[573,710,530,553]" class="Insecta" family="Perlidae" genus="Perlesta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="86" phylum="Arthropoda" rank="species" species="armitagei" status="sp. nov.">
<emphasis id="FFC2D840C53BAA70FDC4FDD354BD253C" box="[573,710,530,553]" italics="true" pageId="3" pageNumber="86">P. armitagei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E4F1653BC53BAA70FD37FDD05553253F" box="[718,808,529,554]" pageId="3" pageNumber="86" rank="species">
<emphasis id="FFC2D840C53BAA70FD37FDD05553253F" bold="true" box="[718,808,529,554]" pageId="3" pageNumber="86">sp. nov.</emphasis>
</taxonomicNameLabel>
but of
<taxonomicName id="0AB67FD1C53BAA70FC83FDD355B2253F" authorityName=": Stark" authorityYear="2004" box="[890,969,529,554]" class="Insecta" family="Perlidae" genus="Perlesta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="86" phylum="Arthropoda" rank="species" species="xube">
<emphasis id="FFC2D840C53BAA70FC83FDD355B2253F" box="[890,969,529,554]" italics="true" pageId="3" pageNumber="86">P. xube</emphasis>
</taxonomicName>
. Males of
<taxonomicName id="0AB67FD1C53BAA70FBBCFDD352EC253F" authorityName=": Stark" authorityYear="2004" box="[1093,1175,529,554]" class="Insecta" family="Perlidae" genus="Perlesta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="86" phylum="Arthropoda" rank="species" species="xube">
<emphasis id="FFC2D840C53BAA70FBBCFDD352EC253F" box="[1093,1175,529,554]" italics="true" pageId="3" pageNumber="86">P. xube</emphasis>
</taxonomicName>
and
<taxonomicName id="0AB67FD1C53BAA70FB37FDD35330253C" authorityName="in Stark" authorityYear="2004" box="[1230,1355,530,553]" class="Insecta" family="Perlidae" genus="Perlesta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="86" phylum="Arthropoda" rank="species" species="cinctipes">
<emphasis id="FFC2D840C53BAA70FB37FDD35330253C" box="[1230,1355,530,553]" italics="true" pageId="3" pageNumber="86">P. cinctipes</emphasis>
</taxonomicName>
key to couplet
<quantity id="0A4EA9B7C53BAA70FF09FDF45760255B" box="[240,283,565,590]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="3" pageNumber="86" unit="in" value="6.0">6 in</quantity>
<bibRefCitation id="A92779A3C53BAA70FED8FDF457CA255B" box="[289,433,565,590]" pageId="3" pageNumber="86" refId="ref8501" refString="Stark, B. P. (2004) Perlidae (The Stones). In: Stark, B. P. &amp; Armitage, B. J. (Eds.), The stoneflies (Plecoptera) of eastern North America. Vol. II. Chloroperlidae, Perlidae, and Perlodidae (Perlodinae). Bulletin of the Ohio Biological Survey. New Series. Uol. 14. Ohio Biological Society, Columbus, Ohio, pp. 61 - 148." type="journal article">Stark (2004)</bibRefCitation>
. This couplet focuses on whether the dorsal patch is “wider at apex than at base” (
<taxonomicName id="0AB67FD1C53BAA70FABCFDF7576F2567" authority=", Stark 2004" authorityName=", Stark" authorityYear="2004" class="Insecta" family="Perlidae" genus="Perlesta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="86" phylum="Arthropoda" rank="species" species="xube">
<emphasis id="FFC2D840C53BAA70FABCFDF753EA255B" box="[1349,1425,565,590]" italics="true" pageId="3" pageNumber="86">P. xube</emphasis>
,
<bibRefCitation id="A92779A3C53BAA70FF6EFD98576F2567" box="[151,276,601,626]" pageId="3" pageNumber="86" refId="ref8501" refString="Stark, B. P. (2004) Perlidae (The Stones). In: Stark, B. P. &amp; Armitage, B. J. (Eds.), The stoneflies (Plecoptera) of eastern North America. Vol. II. Chloroperlidae, Perlidae, and Perlodidae (Perlodinae). Bulletin of the Ohio Biological Survey. New Series. Uol. 14. Ohio Biological Society, Columbus, Ohio, pp. 61 - 148." type="journal article">Stark 2004</bibRefCitation>
</taxonomicName>
, his Fig. 7.370‒7.371) or “narrow throughout most of length” (
<taxonomicName id="0AB67FD1C53BAA70FC22FD9B529D2567" authority=", Stark 2004" authorityName=", Stark" authorityYear="2004" box="[987,1254,601,626]" class="Insecta" family="Perlidae" genus="Perlesta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="86" phylum="Arthropoda" rank="species" species="cinctipes">
<emphasis id="FFC2D840C53BAA70FC22FD9B52202564" box="[987,1115,602,625]" italics="true" pageId="3" pageNumber="86">P. cinctipes</emphasis>
,
<bibRefCitation id="A92779A3C53BAA70FB90FD98529D2567" box="[1129,1254,601,626]" pageId="3" pageNumber="86" refId="ref8501" refString="Stark, B. P. (2004) Perlidae (The Stones). In: Stark, B. P. &amp; Armitage, B. J. (Eds.), The stoneflies (Plecoptera) of eastern North America. Vol. II. Chloroperlidae, Perlidae, and Perlodidae (Perlodinae). Bulletin of the Ohio Biological Survey. New Series. Uol. 14. Ohio Biological Society, Columbus, Ohio, pp. 61 - 148." type="journal article">Stark 2004</bibRefCitation>
</taxonomicName>
, his Fig. 7.300).
<taxonomicName id="0AB67FD1C53BAA70FF6EFDBC571B2580" authorityName=", Stark" authorityYear="2004" box="[151,352,637,662]" class="Insecta" family="Perlidae" genus="Perlesta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="86" phylum="Arthropoda" rank="species" species="cinctipes">
<emphasis id="FFC2D840C53BAA70FF6EFDBC571B2580" box="[151,352,637,662]" italics="true" pageId="3" pageNumber="86">Perlesta cinctipes</emphasis>
</taxonomicName>
is one of few
<taxonomicName id="0AB67FD1C53BAA70FDFFFDBC541F2583" authorityName="Banks" authorityYear="1906" box="[518,612,637,662]" class="Insecta" family="Perlidae" genus="Perlesta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="86" phylum="Arthropoda" rank="genus">
<emphasis id="FFC2D840C53BAA70FDFFFDBC541F2583" box="[518,612,637,662]" italics="true" pageId="3" pageNumber="86">Perlesta</emphasis>
</taxonomicName>
species where the fully-extruded male aedeagus has yet to be illustrated with a line drawing or SEM. The length of the sac is not fully understood, whether in relation to the length of the tube or length and shape of the caecum. An additional aedeagal characteristic is useful in differentiating between these two species. The dorsal patch of
<taxonomicName id="0AB67FD1C53BAA70FDA1FD2B54D02417" authorityName=", Stark" authorityYear="2004" box="[600,683,745,770]" class="Insecta" family="Perlidae" genus="Perlesta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="86" phylum="Arthropoda" rank="species" species="xube">
<emphasis id="FFC2D840C53BAA70FDA1FD2B54D02417" box="[600,683,745,770]" italics="true" pageId="3" pageNumber="86">P. xube</emphasis>
</taxonomicName>
is expanded at the base and body of the caecum but not apically (
<bibRefCitation id="A92779A3C53BAA70FF67FCCC57EC2433" box="[158,407,781,806]" pageId="3" pageNumber="86" refId="ref8652" refString="Stark, B. P. &amp; Rhodes, H. A. (1997) Perlesta xube, a new stonefly species from Nebraska (Plecoptera: Perlidae). Entomological News, 108, 92 - 96. Avaliable from: https: // biodiversitylibrary. org / page / 16343581 (Accessed 28 Jun. 2018)" type="journal article">Stark &amp; Rhodes 1997</bibRefCitation>
, their
<figureCitation id="558D18D7C53BAA70FE27FCCC54592433" box="[478,546,781,806]" captionStart="FIGURE 6" captionStartId="9.[151,250,1983,2005]" captionTargetBox="[224,1362,218,1928]" captionTargetId="figure@9.[188,1398,193,1962]" captionTargetPageId="9" captionText="FIGURE 6. Perlesta armitagei sp. nov. a–b, Ohio, Deer Creek; c–d, Indiana, Blue River; e–f, Kentucky, Russell Creek; g–h, Pennsylvania, French Creek; a, c, e, g, male, head + pronotum, dorsal profile; b, d, f, h. female, head + pronotum, dorsal profile." httpUri="https://zenodo.org/record/1301378/files/figure.png" pageId="3" pageNumber="86">Fig. 6</figureCitation>
;
<bibRefCitation id="A92779A3C53BAA70FDD6FCCC54D62433" box="[559,685,781,806]" pageId="3" pageNumber="86" refId="ref8501" refString="Stark, B. P. (2004) Perlidae (The Stones). In: Stark, B. P. &amp; Armitage, B. J. (Eds.), The stoneflies (Plecoptera) of eastern North America. Vol. II. Chloroperlidae, Perlidae, and Perlodidae (Perlodinae). Bulletin of the Ohio Biological Survey. New Series. Uol. 14. Ohio Biological Society, Columbus, Ohio, pp. 61 - 148." type="journal article">Stark 2004</bibRefCitation>
, his Figs. 7.370‒7.371). The dorsal patch of
<taxonomicName id="0AB67FD1C53BAA70FB5EFCCF535D2430" authorityName=", Stark" authorityYear="2004" box="[1191,1318,782,805]" class="Insecta" family="Perlidae" genus="Perlesta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="86" phylum="Arthropoda" rank="species" species="cinctipes">
<emphasis id="FFC2D840C53BAA70FB5EFCCF535D2430" box="[1191,1318,782,805]" italics="true" pageId="3" pageNumber="86">P. cinctipes</emphasis>
</taxonomicName>
, however, expands well onto the caecum (
<bibRefCitation id="A92779A3C53BAA70FE0EFCF05564245F" box="[503,799,817,842]" pageId="3" pageNumber="86" refId="ref8501" refString="Stark, B. P. (2004) Perlidae (The Stones). In: Stark, B. P. &amp; Armitage, B. J. (Eds.), The stoneflies (Plecoptera) of eastern North America. Vol. II. Chloroperlidae, Perlidae, and Perlodidae (Perlodinae). Bulletin of the Ohio Biological Survey. New Series. Uol. 14. Ohio Biological Society, Columbus, Ohio, pp. 61 - 148." type="journal article">Stark 2004, his Fig. 7.299</bibRefCitation>
).
</paragraph>
<paragraph id="CD090452C53BAA70FF3EFC94545F24CF" blockId="3.[151,1437,421,1058]" pageId="3" pageNumber="86">
Females of
<taxonomicName id="0AB67FD1C53BAA70FEB2FC9757E2247B" authorityName=", Stark" authorityYear="2004" box="[331,409,853,878]" class="Insecta" family="Perlidae" genus="Perlesta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="86" phylum="Arthropoda" rank="species" species="xube">
<emphasis id="FFC2D840C53BAA70FEB2FC9757E2247B" box="[331,409,853,878]" italics="true" pageId="3" pageNumber="86">P. xube</emphasis>
</taxonomicName>
can be easily differentiated from
<taxonomicName id="0AB67FD1C53BAA70FCE3FC9755ED2478" authorityName=", Stark" authorityYear="2004" box="[794,918,854,877]" class="Insecta" family="Perlidae" genus="Perlesta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="86" phylum="Arthropoda" rank="species" species="cinctipes">
<emphasis id="FFC2D840C53BAA70FCE3FC9755ED2478" box="[794,918,854,877]" italics="true" pageId="3" pageNumber="86">P. cinctipes</emphasis>
</taxonomicName>
using
<bibRefCitation id="A92779A3C53BAA70FC1FFC945208247B" box="[998,1139,853,878]" pageId="3" pageNumber="86" refId="ref8501" refString="Stark, B. P. (2004) Perlidae (The Stones). In: Stark, B. P. &amp; Armitage, B. J. (Eds.), The stoneflies (Plecoptera) of eastern North America. Vol. II. Chloroperlidae, Perlidae, and Perlodidae (Perlodinae). Bulletin of the Ohio Biological Survey. New Series. Uol. 14. Ohio Biological Society, Columbus, Ohio, pp. 61 - 148." type="journal article">Stark (2004)</bibRefCitation>
and from
<taxonomicName id="0AB67FD1C53BAA70FB10FC9753092478" authority="Grubbs &amp; Dewalt, 2018" authorityName="Grubbs &amp; Dewalt" authorityYear="2018" box="[1257,1394,854,877]" class="Insecta" family="Perlidae" genus="Perlesta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="86" phylum="Arthropoda" rank="species" species="armitagei" status="sp. nov.">
<emphasis id="FFC2D840C53BAA70FB10FC9753092478" box="[1257,1394,854,877]" italics="true" pageId="3" pageNumber="86">P. armitagei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E4F1653BC53BAA70FA83FC9456B32487" pageId="3" pageNumber="86" rank="species">
<emphasis id="FFC2D840C53BAA70FA83FC9456B32487" bold="true" pageId="3" pageNumber="86">sp. nov.</emphasis>
</taxonomicNameLabel>
here with subgenital plate characteristics. Eggs of
<taxonomicName id="0AB67FD1C53BAA70FCFCFCBB552D2487" authorityName=", Stark" authorityYear="2004" box="[773,854,889,914]" class="Insecta" family="Perlidae" genus="Perlesta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="86" phylum="Arthropoda" rank="species" species="xube">
<emphasis id="FFC2D840C53BAA70FCFCFCBB552D2487" box="[773,854,889,914]" italics="true" pageId="3" pageNumber="86">P. xube</emphasis>
</taxonomicName>
(
<bibRefCitation id="A92779A3C53BAA70FC9CFCB852272487" box="[869,1116,889,914]" pageId="3" pageNumber="86" refId="ref8652" refString="Stark, B. P. &amp; Rhodes, H. A. (1997) Perlesta xube, a new stonefly species from Nebraska (Plecoptera: Perlidae). Entomological News, 108, 92 - 96. Avaliable from: https: // biodiversitylibrary. org / page / 16343581 (Accessed 28 Jun. 2018)" type="journal article">Stark &amp; Rhodes 1997</bibRefCitation>
, their
<figureCitation id="558D18D7C53BAA70FB5BFCB8529E2487" box="[1186,1253,889,914]" captionStart="FIGURE 8" captionStartId="11.[151,250,1811,1833]" captionTargetBox="[225,1364,464,1787]" captionTargetId="figure@11.[188,1398,445,1790]" captionTargetPageId="11" captionText="FIGURE 8. Perlesta armitagei sp. nov., male, aedeagus. a, Ohio, Deer Creek, lateral; b, Indiana, Blue River, lateral, 95X; c, Ohio, Deer Creek, lateral, 200X; d, Ohio Deer Creek, dorsal, 220X; e, Ohio Deer Creek, dorsal; f, Pennsylvania, French Creek, details, of caecum, 950X." httpUri="https://zenodo.org/record/1301382/files/figure.png" pageId="3" pageNumber="86">Fig. 8</figureCitation>
,
<bibRefCitation id="A92779A3C53BAA70FB0AFCB8530B2487" box="[1267,1392,889,914]" pageId="3" pageNumber="86" refId="ref8501" refString="Stark, B. P. (2004) Perlidae (The Stones). In: Stark, B. P. &amp; Armitage, B. J. (Eds.), The stoneflies (Plecoptera) of eastern North America. Vol. II. Chloroperlidae, Perlidae, and Perlodidae (Perlodinae). Bulletin of the Ohio Biological Survey. New Series. Uol. 14. Ohio Biological Society, Columbus, Ohio, pp. 61 - 148." type="journal article">Stark 2004</bibRefCitation>
, his Fig. 7.404) are also distinct from both
<taxonomicName id="0AB67FD1C53BAA70FDBAFC5F54C524A0" authorityName=", Stark" authorityYear="2004" box="[579,702,926,949]" class="Insecta" family="Perlidae" genus="Perlesta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="86" phylum="Arthropoda" rank="species" species="cinctipes">
<emphasis id="FFC2D840C53BAA70FDBAFC5F54C524A0" box="[579,702,926,949]" italics="true" pageId="3" pageNumber="86">P. cinctipes</emphasis>
</taxonomicName>
(
<bibRefCitation id="A92779A3C53BAA70FD29FC5C553524A3" box="[720,846,925,950]" pageId="3" pageNumber="86" refId="ref8452" refString="Stark, B. P. (1989) Perlesta placida (Hagen), an eastern Nearctic species complex (Plecoptera: Perlidae). Insect Systematics and Evolution, 20, 263 - 286. https: // doi. org / 10.1163 / 187631289 X 00339" type="journal article">Stark 1989</bibRefCitation>
, his Fig. 17,
<bibRefCitation id="A92779A3C53BAA70FC25FC5C522324A3" box="[988,1112,925,950]" pageId="3" pageNumber="86" refId="ref8501" refString="Stark, B. P. (2004) Perlidae (The Stones). In: Stark, B. P. &amp; Armitage, B. J. (Eds.), The stoneflies (Plecoptera) of eastern North America. Vol. II. Chloroperlidae, Perlidae, and Perlodidae (Perlodinae). Bulletin of the Ohio Biological Survey. New Series. Uol. 14. Ohio Biological Society, Columbus, Ohio, pp. 61 - 148." type="journal article">Stark 2004</bibRefCitation>
, his Fig. 7.394‒7.395) and
<taxonomicName id="0AB67FD1C53BAA70FA7EFC5F577F24CC" authority="Grubbs &amp; Dewalt, 2018" authorityName="Grubbs &amp; Dewalt" authorityYear="2018" class="Insecta" family="Perlidae" genus="Perlesta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="86" phylum="Arthropoda" rank="species" species="armitagei" status="sp. nov.">
<emphasis id="FFC2D840C53BAA70FA7EFC5F577F24CC" italics="true" pageId="3" pageNumber="86">P. armitagei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E4F1653BC53BAA70FEF2FC00571F24CF" box="[267,356,961,986]" pageId="3" pageNumber="86" rank="species">
<emphasis id="FFC2D840C53BAA70FEF2FC00571F24CF" bold="true" box="[267,356,961,986]" pageId="3" pageNumber="86">sp. nov.</emphasis>
</taxonomicNameLabel>
(
<figureCitation id="558D18D7C53BAA70FE8DFC00546C24CF" box="[372,535,961,986]" captionStart="FIGURE 10" captionStartId="13.[151,250,1112,1134]" captionTargetBox="[236,1344,225,1054]" captionTargetId="figure@13.[206,1377,193,1099]" captionTargetPageId="13" captionText="FIGURE 10. Perlesta armitagei sp. nov., egg. a, Kentucky, Russell Creek, entire egg, 270X; b, Kentucky, Russell Creek, anterior pole and collar, 1600X; c, Ohio, Deer Creek, details of collar, 2300X; d, Kentucky, Russell Creek, posterior pole and micropyles, 500X." httpUri="https://zenodo.org/record/1301386/files/figure.png" pageId="3" pageNumber="86">Figs. 10a‒10d</figureCitation>
).
</paragraph>
<paragraph id="CD090452C53BAA70FF3EFC24557C2337" blockId="3.[151,1437,421,1058]" pageId="3" pageNumber="86">
<taxonomicName id="0AB67FD1C53BAA70FF3EFC24571B24EB" authorityName=", Stark" authorityYear="2004" box="[199,352,997,1022]" class="Insecta" family="Perlidae" genus="Perlesta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="86" phylum="Arthropoda" rank="species" species="xube">
<emphasis id="FFC2D840C53BAA70FF3EFC24571B24EB" box="[199,352,997,1022]" italics="true" pageId="3" pageNumber="86">Perlesta xube</emphasis>
</taxonomicName>
overlaps in distribution with
<taxonomicName id="0AB67FD1C53BAA70FD5EFC27555424E8" authority="Grubbs &amp; Dewalt, 2018" authorityName="Grubbs &amp; Dewalt" authorityYear="2018" box="[679,815,998,1021]" class="Insecta" family="Perlidae" genus="Perlesta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="86" phylum="Arthropoda" rank="species" species="armitagei" status="sp. nov.">
<emphasis id="FFC2D840C53BAA70FD5EFC27555424E8" box="[679,815,998,1021]" italics="true" pageId="3" pageNumber="86">P. armitagei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E4F1653BC53BAA70FCCFFC2455F424EB" box="[822,911,997,1022]" pageId="3" pageNumber="86" rank="species">
<emphasis id="FFC2D840C53BAA70FCCFFC2455F424EB" bold="true" box="[822,911,997,1022]" pageId="3" pageNumber="86">sp. nov.</emphasis>
</taxonomicNameLabel>
from southern
<collectingRegion id="0F72CAB0C53BAA70FBC0FC2452F624EB" box="[1081,1165,997,1022]" country="United States of America" name="Indiana" pageId="3" pageNumber="86">Indiana</collectingRegion>
east to
<collectingRegion id="0F72CAB0C53BAA70FB1BFC24536124EB" box="[1250,1306,997,1022]" country="United States of America" name="Ohio" pageId="3" pageNumber="86">Ohio</collectingRegion>
and with
<taxonomicName id="0AB67FD1C53BAA70FA7EFC2756802334" authorityName=", Stark" authorityYear="2004" class="Insecta" family="Perlidae" genus="Perlesta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="86" phylum="Arthropoda" rank="species" species="cinctipes">
<emphasis id="FFC2D840C53BAA70FA7EFC2756802334" italics="true" pageId="3" pageNumber="86">P. cinctipes</emphasis>
</taxonomicName>
to date only in southern
<collectingRegion id="0F72CAB0C53BAA70FDE8FBCB54322334" box="[529,585,1034,1057]" country="United States of America" name="Iowa" pageId="3" pageNumber="86">Iowa</collectingRegion>
(
<figureCitation id="558D18D7C53BAA70FDA1FBC854812337" box="[600,762,1033,1058]" captionStart="FIGURE 11" captionStartId="14.[151,250,1938,1960]" captionTargetBox="[205,1400,405,1888]" captionTargetId="figure@14.[167,1431,373,1925]" captionTargetPageId="14" captionText="FIGURE 11. Distribution maps of Perlesta based on material examined during this study. a, P. adena, P. browni, P. cinctipes, and P. xube; b, P. armitagei sp. nov. The record from northern West Virginia (red circle; Tarter &amp; Nelson 2006) is conditional since that material was not available for study. The type locality is represented by the purple circle." httpUri="https://zenodo.org/record/1301388/files/figure.png" pageId="3" pageNumber="86">Figs. 11a‒11b</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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