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<document id="C4EAED2A68A9CC24E1B213FF80550CD0" ID-DOI="10.5281/zenodo.13799439" ID-ISSN="0068-547X" ID-Zenodo-Dep="13799439" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722538212676" checkinUser="felipe" docAuthor="Kavanaugh, David H. &amp; Rainio, Johanna" docDate="2016" docId="0A3E26573066FFB1FF8EFE48BDA50C48" docLanguage="en" docName="ProcCalifAcadSci.63.7.201-268.pdf" docOrigin="Proceedings of the California Academy of Sciences 63 (7)" docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.2:ProcCalifAcadSci.2015-.journal_article" docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B" docStyleName="ProcCalifAcadSci.2015-.journal_article" docStyleVersion="2" docTitle="Thysanotus bimaculatoides Kavanaugh and Rainio 2016, sp. nov." docType="treatment" docVersion="2" lastPageNumber="234" masterDocId="F6075E2F3079FF90FFF5FF92BD630D1B" masterDocTitle="Twenty-six New Species of Predaceous Ground Beetles (Coleoptera: Adephaga: Carabidae) from Ranomafana National Park, Madagascar" masterLastPageNumber="268" masterPageNumber="201" pageNumber="232" updateTime="1726771873529" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="D6819875E67CA624B97FDA8E5F65CB94">Twenty-six New Species of Predaceous Ground Beetles (Coleoptera: Adephaga: Carabidae) from Ranomafana National Park, Madagascar</mods:title>
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<mods:name id="89938BE3FF97024642495F6EA2A2A63B" type="personal">
<mods:role id="0946D0E53DD782470AB58FFB37A2078C">
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<mods:namePart id="15052633F69FB62CEE0216A9E0E1543E">Kavanaugh, David H.</mods:namePart>
<mods:affiliation id="B22CD5C63CC79D38A846CD3D6C007D19">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA; Email: dkavanaugh @ calacademy. org; &amp; Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org)</mods:affiliation>
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<mods:namePart id="DE0ACFA19F00F8E42A0328C9434C11C7">Rainio, Johanna</mods:namePart>
<mods:affiliation id="45C833C3DD26E8B21C7E3E733F057706">Department of Ecology and Systematics, University of Helsinki, P. O. Box 65, FIN- 00014 Helsinki, Finland; Email: kjohannarainio @ gmail. com</mods:affiliation>
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<mods:title id="6FF2BB91F9706996C03E4713398325AD">Proceedings of the California Academy of Sciences</mods:title>
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<mods:date id="76038B89BFFB216516E08054AA549AEB">2016</mods:date>
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<mods:number id="7964E66E0733A05C225F7D6295214794">2016-04-29</mods:number>
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<mods:number id="A8350A4AF83BF44EBADC7C971AAF38A5">63</mods:number>
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<treatment id="0A3E26573066FFB1FF8EFE48BDA50C48" LSID="urn:lsid:plazi:treatment:0A3E26573066FFB1FF8EFE48BDA50C48" httpUri="http://treatment.plazi.org/id/0A3E26573066FFB1FF8EFE48BDA50C48" lastPageId="33" lastPageNumber="234" pageId="31" pageNumber="232">
<subSubSection id="CA8DC4CA3066FF8FFF8EFE48BE7F0CE9" box="[123,796,474,498]" pageId="31" pageNumber="232" type="nomenclature">
<paragraph id="822897413066FF8FFF8EFE48BE7F0CE9" blockId="31.[123,796,474,528]" box="[123,796,474,498]" pageId="31" pageNumber="232">
<heading id="D960202D3066FF8FFF8EFE48BE7F0CE9" box="[123,796,474,498]" fontSize="10" level="2" pageId="31" pageNumber="232" reason="4">
<emphasis id="B0E34B533066FF8FFF8EFE48BE7F0CE9" bold="true" box="[123,796,474,498]" pageId="31" pageNumber="232">
<taxonomicName id="4597ECC23066FF8FFF8EFE48BFDB0CE9" ID-CoL="56TVV" authority="Kavanaugh and Rainio" authorityName="Kavanaugh and Rainio" authorityYear="2016" box="[123,696,474,498]" class="Insecta" family="Carabidae" genus="Thysanotus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="232" phylum="Arthropoda" rank="species" species="bimaculatoides" status="sp. nov.">
<emphasis id="B0E34B533066FF8FFF8EFE48BCC40CE9" bold="true" box="[123,423,474,498]" italics="true" pageId="31" pageNumber="232">Thysanotus bimaculatoides</emphasis>
Kavanaugh and Rainio
</taxonomicName>
,
<taxonomicNameLabel id="ABD0F6283066FF8FFD33FE48BE7F0CE9" box="[710,796,474,498]" pageId="31" pageNumber="232" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="CA8DC4CA3066FF8FFF8EFE68BC660F14" box="[123,261,506,528]" pageId="31" pageNumber="232" type="description">
<paragraph id="822897413066FF8FFF8EFE68BC660F14" blockId="31.[123,796,474,528]" box="[123,261,506,528]" pageId="31" pageNumber="232">
<figureCitation id="1AAC8BC43066FF8FFF8EFE68BC660F14" box="[123,261,506,528]" captionStart-0="FIGURE" captionStart-1="FIGURE" captionStartId-0="31.[157,169,1669,1688]" captionStartId-1="32.[155,167,675,694]" captionTargetBox-0="[121,1177,624,1665]" captionTargetBox-1="[140,1163,222,658]" captionTargetId-0="figure-579@31.[121,1177,624,1665]" captionTargetId-1="figure-2339@32.[120,1176,216,672]" captionTargetPageId-0="31" captionTargetPageId-1="32" captionText-0="FIGURE 17. A-D. Digital images of holotype male of Thysanotus bimaculatoides Kavanaugh &amp; Rainio sp. nov. AD. A Habitus, Dorsal aspect; B. Labels. Scale line = 1.0 mm." captionText-1="FIGURE 18. Digital images of aedeagus of male genitalia. A, C. Left lateral aspect; B, D. Dorsal aspect. AB. Holotype male of Thysanotus bimaculatoides Kavanaugh &amp; Rainio sp. nov.; B, D. Thysanotus spinosus Alluaud (from Ranomafana National Park). Scale lines = 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.13799481" figureDoi-1="http://doi.org/10.5281/zenodo.13799483" httpUri-0="https://zenodo.org/record/13799481/files/figure.png" httpUri-1="https://zenodo.org/record/13799483/files/figure.png" pageId="31" pageNumber="232">Figures 1718</figureCitation>
</paragraph>
</subSubSection>
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<paragraph id="822897413066FF8FFF5EFDBCB9F50F5D" blockId="31.[123,1174,558,614]" box="[171,1174,558,582]" pageId="31" pageNumber="232">
<emphasis id="B0E34B533066FF8FFF5EFDBCBC3C0F5F" bold="true" box="[171,351,558,582]" pageId="31" pageNumber="232">
<smallCapsWord id="84CE019D3066FF8FFF5EFDBCBD870F5F" baselines="576,575" box="[171,228,558,582]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Type" pageId="31" pageNumber="232">TYPE</smallCapsWord>
<smallCapsWord id="84CE019D3066FF8FFF07FDA3BC3C0F5F" baselines="575" box="[242,351,561,580]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="material" pageId="31" pageNumber="232">MATERIAL</smallCapsWord>
</emphasis>
.—
<materialsCitation id="32FF9D1C3066FF8FFE73FDBCB9F50F5D" box="[390,1174,558,582]" country="Madagascar" location="MADAGASCAR" pageId="31" pageNumber="232" specimenCount="1" typeStatus="holotype">
<typeStatus id="5D2C29E33066FF8FFE73FDBCBC8A0F5D" box="[390,489,558,582]" pageId="31" pageNumber="232" type="holotype">Holotype</typeStatus>
(
<figureCitation id="1AAC8BC43066FF8FFDF5FDBCBFE10F5E" box="[512,642,558,582]" captionStart="FIGURE" captionStartId="30.[155,167,1246,1265]" captionTargetBox="[120,1176,216,1220]" captionTargetId="figure-760@30.[120,1176,216,1239]" captionTargetPageId="30" captionText="FIGURE 16. Digital images of holotype female of Archicolliuris ranomafanae Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels. Scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13799477" httpUri="https://zenodo.org/record/13799477/files/figure.png" pageId="31" pageNumber="232">Figs.16AB</figureCitation>
), a male, in CAS, labeled: “
<collectingCountry id="FA80D7D13066FF8FFC2DFDBCB9F50F5D" box="[984,1174,558,582]" name="Madagascar" pageId="31" pageNumber="232">MADAGASCAR</collectingCountry>
</materialsCitation>
</paragraph>
<paragraph id="822897413066FF8FFF8EFDDCB9F50F7D" blockId="31.[123,1174,558,614]" box="[123,1174,589,614]" pageId="31" pageNumber="232">
Ranomafana NP. Mangevo plot
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<date id="F629B1813066FF8FFDB5FDDCBFC90F7E" box="[576,682,589,614]" pageId="31" pageNumber="232" value="2005-04-20">20.4.2005</date>
”/ “240” [handwritten label]/ “
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</paragraph>
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<paragraph id="822897413066FF8FFF68F917B9F50B83" blockId="31.[124,1174,1668,1714]" box="[157,1174,1668,1688]" pageId="31" pageNumber="232">
<smallCapsWord id="84CE019D3066FF8FFF68F917BDBA0B8D" baselines="1683,1682" box="[157,217,1669,1688]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="31" pageNumber="232">FIGURE</smallCapsWord>
17. A-D. Digital images of holotype male of
<taxonomicName id="4597ECC23066FF8FFD78F916B9260B8C" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[653,1093,1668,1688]" class="Insecta" family="Carabidae" genus="Thysanotus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="232" phylum="Arthropoda" rank="species" species="bimaculatoides" status="sp. nov.">
<emphasis id="B0E34B533066FF8FFD78F916BE190B8C" box="[653,890,1668,1687]" italics="true" pageId="31" pageNumber="232">Thysanotus bimaculatoides</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F6283066FF8FFBA5F917B9F50B83" box="[1104,1174,1669,1688]" pageId="31" pageNumber="232" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="822897413066FF8FFF89F90CBFC30BA9" blockId="31.[124,1174,1668,1714]" box="[124,672,1694,1714]" pageId="31" pageNumber="232">AD. A Habitus, Dorsal aspect; B. Labels. Scale line = 1.0 mm.</paragraph>
</caption>
<caption id="D6E8C7C93059FFB0FF6EFD31BCDE0FF2" ID-DOI="http://doi.org/10.5281/zenodo.13799483" ID-Zenodo-Dep="13799483" httpUri="https://zenodo.org/record/13799483/files/figure.png" pageId="32" pageNumber="233" startId="32.[155,167,675,694]" targetBox="[140,1163,222,658]" targetPageId="32" targetType="figure">
<paragraph id="822897413059FFB0FF6EFD31BCDE0FF2" blockId="32.[123,1174,675,745]" pageId="32" pageNumber="233">
<smallCapsWord id="84CE019D3059FFB0FF6EFD31BDBB0FAF" baselines="689,688" box="[155,216,675,694]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="32" pageNumber="233">FIGURE</smallCapsWord>
18. Digital images of aedeagus of male genitalia. A, C. Left lateral aspect; B, D. Dorsal aspect. AB. Holotype male of
<taxonomicName id="4597ECC23059FFB0FF37FD2EBF0A0FD4" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[194,617,700,719]" class="Insecta" family="Carabidae" genus="Thysanotus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="233" phylum="Arthropoda" rank="species" species="bimaculatoides" status="sp. nov.">
<emphasis id="B0E34B533059FFB0FF37FD2EBCCF0FD4" box="[194,428,700,719]" italics="true" pageId="32" pageNumber="233">Thysanotus bimaculatoides</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F6283059FFB0FD9AFD2FBFD20FCB" box="[623,689,701,720]" pageId="32" pageNumber="233" rank="species">sp. nov.</taxonomicNameLabel>
; B, D.
<taxonomicName id="4597ECC23059FFB0FD07FD2EBE8E0FD4" authority="Alluaud" authorityName="Alluaud" authorityYear="1936" box="[754,1005,700,719]" class="Insecta" family="Carabidae" genus="Thysanotus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="233" phylum="Arthropoda" rank="species" species="spinosus">
<emphasis id="B0E34B533059FFB0FD07FD2EBEC10FD4" box="[754,930,700,719]" italics="true" pageId="32" pageNumber="233">Thysanotus spinosus</emphasis>
Alluaud
</taxonomicName>
(from Ranomafana National Park). Scale lines = 0.5 mm.
</paragraph>
</caption>
<paragraph id="822897413059FFB0FF8EFC90BE550EC1" blockId="32.[123,1174,770,1725]" pageId="32" pageNumber="233">
<taxonomicName id="4597ECC23059FFB0FF8EFC90BFF30E01" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[123,656,770,794]" class="Insecta" family="Carabidae" genus="Thysanotus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="233" phylum="Arthropoda" rank="species" species="bimaculatoides" status="sp. nov.">
<emphasis id="B0E34B533059FFB0FF8EFC90BCC30E01" box="[123,416,770,794]" italics="true" pageId="32" pageNumber="233">Thysanotus bimaculatoides</emphasis>
<materialsCitation id="32FF9D1C3059FFB0FE5CFC90BFF30E01" box="[425,656,770,794]" collectingDate="2000-11" collectingDateMax="2005-04-22" collectingDateMin="2000-11" collectionCode="MNHN, CAS" collectorName="R. Harin'Hala &amp; M. E. Irwin" country="Madagascar" county="Mangevo" elevation="1600" latitude="-21.827501" location="Foret d'Ambalagoavy Nord" longLatPrecision="21" longitude="47.338886" municipality="Province" pageId="32" pageNumber="233" specimenCount="2" specimenCount-female="2" stateProvince="Fianarantsoa" typeStatus="paratype">Kavanaugh &amp; Rainio</materialsCitation>
</taxonomicName>
<taxonomicNameLabel id="ABD0F6283059FFB0FD6CFC90BFB70E02" box="[665,724,770,793]" pageId="32" pageNumber="233" rank="species">sp. n.</taxonomicNameLabel>
2015” [red label].
<typeStatus id="5D2C29E33059FFB0FC52FC90B96C0E01" box="[935,1039,770,794]" pageId="32" pageNumber="233" type="paratype">Paratypes</typeStatus>
(total of 2):
<specimenCount id="94915CC83059FFB0FF8EFCB0BD810E21" box="[123,226,802,826]" count="1" pageId="32" pageNumber="233" type="female">1 female</specimenCount>
(in
<collectionCode id="E4860F843059FFB0FED7FCB0BC170E22" box="[290,372,802,825]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="32" pageNumber="233">MNHN</collectionCode>
) labeled “
<collectingCountry id="FA80D7D13059FFB0FE0CFCB0BFDB0E21" box="[505,696,802,826]" name="Madagascar" pageId="32" pageNumber="233">MADAGASCAR</collectingCountry>
Ranomafana NP.
<collectingCounty id="6B49EFCD3059FFB0FC6CFCB0BE9C0E21" box="[921,1023,802,826]" pageId="32" pageNumber="233">Mangevo</collectingCounty>
plot
<quantity id="456F3AA43059FFB0FBBEFCB0B9F50E22" box="[1099,1174,802,826]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="32" pageNumber="233" unit="m" value="1600.0">
<elevation id="09BA70723059FFB0FBBEFCB0B9F50E22" box="[1099,1174,802,826]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="32" pageNumber="233" unit="m" value="1600.0">1600m</elevation>
</quantity>
<date id="F629B1813059FFB0FF8EFCD0BD860E41" box="[123,229,834,858]" pageId="32" pageNumber="233" value="2005-04-22">
<collectingDate id="E66D48693059FFB0FF8EFCD0BD860E41" box="[123,229,834,858]" pageId="32" pageNumber="233" value="2005-04-22">22.4.2005</collectingDate>
</date>
”/ “240” [handwritten label];
<specimenCount id="94915CC83059FFB0FDEAFCD0BF1F0E41" box="[543,636,834,858]" count="1" pageId="32" pageNumber="233" type="female">1 female</specimenCount>
(in
<collectionCode id="E4860F843059FFB0FD52FCD0BFB80E41" box="[679,731,834,858]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="32" pageNumber="233" type="Museum">CAS</collectionCode>
) labeled “CASENT 8069538”/ “
<collectingCountry id="FA80D7D13059FFB0FBCEFCD0BD840E61" name="Madagascar" pageId="32" pageNumber="233">MADAGASCAR</collectingCountry>
:
<collectingMunicipality id="624C0D3B3059FFB0FF0FFCF0BC3A0E61" box="[250,345,866,890]" pageId="32" pageNumber="233">Province</collectingMunicipality>
<collectingRegion id="405359A33059FFB0FE91FCF0BC8F0E61" box="[356,492,866,890]" country="Madagascar" name="Fianarantsoa" pageId="32" pageNumber="233">Fianarantsoa</collectingRegion>
,
<location id="8748C19A3059FFB0FE0BFCF0BE4C0E61" LSID="urn:lsid:plazi:treatment:0A3E26573066FFB1FF8EFE48BDA50C48:8748C19A3059FFB0FE0BFCF0BE4C0E61" box="[510,815,866,890]" country="Madagascar" county="Mangevo" latitude="-21.827501" longLatPrecision="21" longitude="47.338886" municipality="Province" name="Foret d'Ambalagoavy Nord" pageId="32" pageNumber="233" stateProvince="Fianarantsoa">Forêt dAmbalagoavy Nord</location>
,
<location id="8748C19A3059FFB0FCB4FCF0BEE80E61" LSID="urn:lsid:plazi:treatment:0A3E26573066FFB1FF8EFE48BDA50C48:8748C19A3059FFB0FCB4FCF0BEE80E61" box="[833,907,866,890]" country="Madagascar" county="Mangevo" latitude="-21.827501" longLatPrecision="21" longitude="47.338886" municipality="Province" name="Ikonga" pageId="32" pageNumber="233" stateProvince="Fianarantsoa">Ikonga</location>
,
<location id="8748C19A3059FFB0FC69FCF0B9680E61" LSID="urn:lsid:plazi:treatment:0A3E26573066FFB1FF8EFE48BDA50C48:8748C19A3059FFB0FC69FCF0B9680E61" box="[924,1035,866,890]" country="Madagascar" county="Mangevo" latitude="-21.827501" longLatPrecision="21" longitude="47.338886" municipality="Province" name="Abatombe" pageId="32" pageNumber="233" stateProvince="Fianarantsoa">Abatombe</location>
<date id="F629B1813059FFB0FBE3FCF1B9E10E61" box="[1046,1154,866,890]" pageId="32" pageNumber="233" value="2000-11">
<collectingDate id="E66D48693059FFB0FBE3FCF1B9E10E61" box="[1046,1154,866,890]" pageId="32" pageNumber="233" value="2000-11">Nov-2000</collectingDate>
</date>
”/ “
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,
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<location id="8748C19A3059FFB0FE62FC10BFF50E81" LSID="urn:lsid:plazi:treatment:0A3E26573066FFB1FF8EFE48BDA50C48:8748C19A3059FFB0FE62FC10BFF50E81" box="[407,662,898,922]" country="Madagascar" county="Mangevo" latitude="-21.827501" longLatPrecision="21" longitude="47.338886" municipality="Province" name="Calif. Acad. of Sciences" pageId="32" pageNumber="233" stateProvince="Fianarantsoa">Calif. Acad. of Sciences</location>
colls:
<collectorName id="2F62F2973059FFB0FD29FC11BE110E81" box="[732,882,898,922]" pageId="32" pageNumber="233">R. HarinHala</collectorName>
&amp;
<collectorName id="2F62F2973059FFB0FC66FC11B96B0E81" box="[915,1032,898,922]" pageId="32" pageNumber="233">M.E. Irwin</collectorName>
,
<collectingMethod id="5BD6EF563059FFB0FBE1FC10B9F60E81" box="[1044,1173,898,922]" pageId="32" pageNumber="233">malaise trap</collectingMethod>
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<elevation id="09BA70723059FFB0FF8EFC30BDDB0EA1" box="[123,184,930,954]" metricMagnitude="2" metricUnit="m" metricValue="6.25" pageId="32" pageNumber="233" unit="m" value="625.0">625m</elevation>
</quantity>
MA-01-12-01”. Both
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also bear the following label: “
<typeStatus id="5D2C29E33059FFB0FC79FC31B9730EA0" box="[908,1040,931,955]" pageId="32" pageNumber="233" type="paratype">PARATYPE</typeStatus>
<taxonomicName id="4597ECC23059FFB0FBE9FC30BF6A0EC0" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" class="Insecta" family="Carabidae" genus="Thysanotus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="233" phylum="Arthropoda" rank="species" species="bimaculatoides" status="sp. nov.">
<emphasis id="B0E34B533059FFB0FBE9FC30BC7D0EC0" italics="true" pageId="32" pageNumber="233">Thysanotus bimaculatoides</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F6283059FFB0FDE5FC51BF290EC1" box="[528,586,963,986]" pageId="32" pageNumber="233" rank="species">sp. n.</taxonomicNameLabel>
2015” [yellow label].
</paragraph>
<paragraph id="822897413059FFB0FF5EFC71B9500EE1" blockId="32.[123,1174,770,1725]" box="[171,1075,995,1019]" pageId="32" pageNumber="233">
<materialsCitation id="32FF9D1C3059FFB0FF5EFC71B94F0EE0" box="[171,1068,995,1019]" country="Madagascar" location="Ranomafana National Park" pageId="32" pageNumber="233" specimenCount="1" stateProvince="Fianarantsoa" typeStatus="holotype">
<emphasis id="B0E34B533059FFB0FF5EFC71BC340EE2" bold="true" box="[171,343,995,1019]" pageId="32" pageNumber="233">
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<typeStatus id="5D2C29E33059FFB0FF5EFC71BD870EE2" box="[171,228,995,1019]" pageId="32" pageNumber="233">TYPE</typeStatus>
</smallCapsWord>
<smallCapsWord id="84CE019D3059FFB0FF19FC74BC340EE2" baselines="1012" box="[236,343,998,1017]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="locality" pageId="32" pageNumber="233">LOCALITY</smallCapsWord>
</emphasis>
.—
<collectingCountry id="FA80D7D13059FFB0FE8BFC71BC9C0EE0" box="[382,511,995,1019]" name="Madagascar" pageId="32" pageNumber="233">Madagascar</collectingCountry>
,
<collectingRegion id="405359A33059FFB0FDF8FC71BF9F0EE0" box="[525,764,995,1019]" country="Madagascar" name="Fianarantsoa" pageId="32" pageNumber="233">Fianarantsoa Province</collectingRegion>
,
<location id="8748C19A3059FFB0FCFFFC71B94F0EE0" LSID="urn:lsid:plazi:treatment:0A3E26573066FFB1FF8EFE48BDA50C48:8748C19A3059FFB0FCFFFC71B94F0EE0" box="[778,1068,995,1019]" country="Madagascar" name="Ranomafana National Park" pageId="32" pageNumber="233" stateProvince="Fianarantsoa">Ranomafana National Park</location>
</materialsCitation>
.
</paragraph>
<paragraph id="822897413059FFB0FF5EFB91BF5D0960" blockId="32.[123,1174,770,1725]" pageId="32" pageNumber="233">
<emphasis id="B0E34B533059FFB0FF5EFB91BC8C0902" bold="true" box="[171,495,1027,1051]" pageId="32" pageNumber="233">
<smallCapsWord id="84CE019D3059FFB0FF5EFB91BC4D0902" baselines="1045,1044" box="[171,302,1027,1051]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Derivation" pageId="32" pageNumber="233">DERIVATION</smallCapsWord>
<smallCapsWord id="84CE019D3059FFB0FEC0FB94BC320902" baselines="1044" box="[309,337,1030,1049]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="of" pageId="32" pageNumber="233">OF</smallCapsWord>
<smallCapsWord id="84CE019D3059FFB0FEA2FB94BCC90902" baselines="1044" box="[343,426,1030,1049]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="species" pageId="32" pageNumber="233">SPECIES</smallCapsWord>
<smallCapsWord id="84CE019D3059FFB0FE47FB94BC8C0902" baselines="1044" box="[434,495,1030,1049]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="name" pageId="32" pageNumber="233">NAME</smallCapsWord>
</emphasis>
.— The species epithet,
<taxonomicName id="4597ECC23059FFB0FD06FB91BEF50900" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[755,918,1027,1051]" class="Insecta" family="Carabidae" genus="Thysanotus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="233" phylum="Arthropoda" rank="species" species="bimaculatoides">
<emphasis id="B0E34B533059FFB0FD06FB91BEF50900" box="[755,918,1027,1051]" italics="true" pageId="32" pageNumber="233">bimaculatoides</emphasis>
</taxonomicName>
, is an adjective derived from the Latin words,
<emphasis id="B0E34B533059FFB0FE9CFBB1BCEB0920" box="[361,392,1059,1083]" italics="true" pageId="32" pageNumber="233">bis</emphasis>
, meaning two, and
<emphasis id="B0E34B533059FFB0FDAFFBB1BFCA0920" box="[602,681,1059,1083]" italics="true" pageId="32" pageNumber="233">macula</emphasis>
, meaning spot or spotted, and the Greek word
<emphasis id="B0E34B533059FFB0FF8EFBD1BDD00940" box="[123,179,1091,1115]" italics="true" pageId="32" pageNumber="233">eidos</emphasis>
, meaning like. The name refers to the similarity between members of this species and those of
<taxonomicName id="4597ECC23059FFB0FF6DFBF1BF540960" authority="Deuve (2010)" authorityName="Deuve" authorityYear="2010" box="[152,567,1123,1147]" class="Insecta" family="Carabidae" genus="Thysanotus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="233" phylum="Arthropoda" rank="species" species="bimaculatus">
<emphasis id="B0E34B533059FFB0FF6DFBF1BCF80960" box="[152,411,1123,1147]" italics="true" pageId="32" pageNumber="233">Thysanotus bimaculatus</emphasis>
<bibRefCitation id="E606EAB03059FFB0FE56FBF6BF540960" author="DEUVE, T." box="[419,567,1123,1147]" pageId="32" pageNumber="233" pagination="165 - 176" refId="ref30046" refString="DEUVE, T. 2010. Note sur le genre Thysanotus Chaudoir et quelques autres Lebiinae Pericalini de Madagascar [Coleoptera, Caraboidea, Harpalidae]. Revue Francaise d'Entomologie (N. S.) 32: 165 - 176." type="journal article" year="2010">Deuve (2010)</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="822897413059FFB0FF5EFB11BEC10B66" blockId="32.[123,1174,770,1725]" pageId="32" pageNumber="233">
<smallCapsWord id="84CE019D3059FFB0FF5EFB11BC210981" baselines="1173,1173" box="[171,322,1155,1179]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Recognition" pageId="32" pageNumber="233">
<emphasis id="B0E34B533059FFB0FF5EFB11BC210981" bold="true" box="[171,322,1155,1179]" pageId="32" pageNumber="233">RECOGNITION</emphasis>
</smallCapsWord>
.— Size slightly less than average for genus, SBL of male =
<quantity id="456F3AA43059FFB0FC32FB11B97B0980" box="[967,1048,1155,1179]" metricMagnitude="-3" metricUnit="m" metricValue="5.7" pageId="32" pageNumber="233" unit="mm" value="5.7">5.7 mm</quantity>
, of females =
<quantity id="456F3AA43059FFB0FF65FB31BD8709A0" box="[144,228,1187,1212]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="32" pageNumber="233" unit="mm" value="5.3">5.3 mm</quantity>
. Members of this species (
<figureCitation id="1AAC8BC43059FFB0FDE4FB31BF1109A7" box="[529,626,1187,1212]" captionStart="FIGURE" captionStartId="31.[157,169,1669,1688]" captionTargetBox="[121,1177,624,1665]" captionTargetId="figure-579@31.[121,1177,624,1665]" captionTargetPageId="31" captionText="FIGURE 17. A-D. Digital images of holotype male of Thysanotus bimaculatoides Kavanaugh &amp; Rainio sp. nov. AD. A Habitus, Dorsal aspect; B. Labels. Scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13799481" httpUri="https://zenodo.org/record/13799481/files/figure.png" pageId="32" pageNumber="233">Fig. 17A</figureCitation>
) key to
<taxonomicName id="4597ECC23059FFB0FD26FB31B91409A7" authority="Deuve (2010)" authorityName="Deuve" authorityYear="2010" box="[723,1143,1187,1212]" class="Insecta" family="Carabidae" genus="Thysanotus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="233" phylum="Arthropoda" rank="species" species="bimaculatus">
<emphasis id="B0E34B533059FFB0FD26FB31BEBB09A0" box="[723,984,1187,1211]" italics="true" pageId="32" pageNumber="233">Thysanotus bimaculatus</emphasis>
<bibRefCitation id="E606EAB03059FFB0FC14FB36B91409A7" author="DEUVE, T." box="[993,1143,1188,1212]" pageId="32" pageNumber="233" pagination="165 - 176" refId="ref30046" refString="DEUVE, T. 2010. Note sur le genre Thysanotus Chaudoir et quelques autres Lebiinae Pericalini de Madagascar [Coleoptera, Caraboidea, Harpalidae]. Revue Francaise d'Entomologie (N. S.) 32: 165 - 176." type="journal article" year="2010">Deuve (2010)</bibRefCitation>
</taxonomicName>
in Deuves key based on the following combination of features: elytra shiny, intervals smooth, elytral microsculpture indistinct (comprised of faintly impressed microlines and markedly transverse sculpticells, each elytron with a large, pale macula near the base on intervals 2 to 4, also extended partially onto interval 1 and to scutellum at basomedial part of macula, elytral apex bluntly round- ed, not toothed. Members of
<taxonomicName id="4597ECC23059FFB0FE40FAD7BF100847" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[437,627,1348,1372]" class="Insecta" family="Carabidae" genus="Thysanotus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="233" phylum="Arthropoda" rank="species" species="bimaculatoides">
<emphasis id="B0E34B533059FFB0FE40FAD7BF100847" box="[437,627,1348,1372]" italics="true" pageId="32" pageNumber="233">T. bimaculatoides</emphasis>
</taxonomicName>
differ from the
<typeStatus id="5D2C29E33059FFB0FCD4FAD6BE1D0847" box="[801,894,1348,1372]" pageId="32" pageNumber="233" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4597ECC23059FFB0FC51FAD7B9220847" authorityName="Deuve" authorityYear="2010" box="[932,1089,1348,1372]" class="Insecta" family="Carabidae" genus="Thysanotus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="233" phylum="Arthropoda" rank="species" species="bimaculatus">
<emphasis id="B0E34B533059FFB0FC51FAD7B9220847" box="[932,1089,1348,1372]" italics="true" pageId="32" pageNumber="233">T. bimaculatus</emphasis>
</taxonomicName>
in having slightly larger body size (SBL of
<typeStatus id="5D2C29E33059FFB0FDD1FAF6BFE20867" box="[548,641,1380,1404]" pageId="32" pageNumber="233" type="holotype">holotype</typeStatus>
female of
<taxonomicName id="4597ECC23059FFB0FCF5FAF7BEC20867" authorityName="Deuve" authorityYear="2010" box="[768,929,1380,1404]" class="Insecta" family="Carabidae" genus="Thysanotus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="233" phylum="Arthropoda" rank="species" species="bimaculatus">
<emphasis id="B0E34B533059FFB0FCF5FAF7BEC20867" box="[768,929,1380,1404]" italics="true" pageId="32" pageNumber="233">T. bimaculatus</emphasis>
</taxonomicName>
=
<quantity id="456F3AA43059FFB0FC4BFAF6B9700867" box="[958,1043,1380,1404]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="32" pageNumber="233" unit="mm" value="4.9">4.9 mm</quantity>
), pronotum slightly narrower, ratio of PW/PL = 1.1 (ratio of PW/PL =
<quantity id="456F3AA43059FFB0FD01FA16BE510887" box="[756,818,1412,1436]" metricMagnitude="-2" metricUnit="m" metricValue="3.048" pageId="32" pageNumber="233" unit="in" value="1.2">1.2 in</quantity>
<taxonomicName id="4597ECC23059FFB0FCCCFA17BEB50887" authorityName="Deuve" authorityYear="2010" box="[825,982,1412,1436]" class="Insecta" family="Carabidae" genus="Thysanotus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="233" phylum="Arthropoda" rank="species" species="bimaculatus">
<emphasis id="B0E34B533059FFB0FCCCFA17BEB50887" box="[825,982,1412,1436]" italics="true" pageId="32" pageNumber="233">T. bimaculatus</emphasis>
</taxonomicName>
), narrower basally with subbasal sinuation of the lateral margin slightly longer and deeper and hind angles sharp- er, rectangular to distinctly acute (compare with
<bibRefCitation id="E606EAB03059FFB0FD76FA57BE6608C6" author="DEUVE, T." box="[643,773,1477,1501]" pageId="32" pageNumber="233" pagination="165 - 176" refId="ref30046" refString="DEUVE, T. 2010. Note sur le genre Thysanotus Chaudoir et quelques autres Lebiinae Pericalini de Madagascar [Coleoptera, Caraboidea, Harpalidae]. Revue Francaise d'Entomologie (N. S.) 32: 165 - 176." type="journal article" year="2010">Deuve 2010</bibRefCitation>
,
<figureCitation id="1AAC8BC43059FFB0FCE6FA57BE3C08C6" box="[787,863,1477,1501]" captionStart="FIGURE" captionStartId="31.[157,169,1669,1688]" captionTargetBox="[121,1177,624,1665]" captionTargetId="figure-579@31.[121,1177,624,1665]" captionTargetPageId="31" captionText="FIGURE 17. A-D. Digital images of holotype male of Thysanotus bimaculatoides Kavanaugh &amp; Rainio sp. nov. AD. A Habitus, Dorsal aspect; B. Labels. Scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13799481" httpUri="https://zenodo.org/record/13799481/files/figure.png" pageId="32" pageNumber="233">Fig. 17</figureCitation>
). Deuve did not have a male specimen of
<taxonomicName id="4597ECC23059FFB0FEF0FA77BCC108E6" authorityName="Deuve" authorityYear="2010" box="[261,418,1509,1533]" class="Insecta" family="Carabidae" genus="Thysanotus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="233" phylum="Arthropoda" rank="species" species="bimaculatus">
<emphasis id="B0E34B533059FFB0FEF0FA77BCC108E6" box="[261,418,1509,1533]" italics="true" pageId="32" pageNumber="233">T. bimaculatus</emphasis>
</taxonomicName>
, so we cannot compare male genitalia of these two species. However, for future comparative purposes, we provide digital photographs of the genitalia (
<figureCitation id="1AAC8BC43059FFB0FC19F997B90C0B06" box="[1004,1135,1541,1565]" captionStart="FIGURE" captionStartId="32.[155,167,675,694]" captionTargetBox="[140,1163,222,658]" captionTargetId="figure-2339@32.[120,1176,216,672]" captionTargetPageId="32" captionText="FIGURE 18. Digital images of aedeagus of male genitalia. A, C. Left lateral aspect; B, D. Dorsal aspect. AB. Holotype male of Thysanotus bimaculatoides Kavanaugh &amp; Rainio sp. nov.; B, D. Thysanotus spinosus Alluaud (from Ranomafana National Park). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13799483" httpUri="https://zenodo.org/record/13799483/files/figure.png" pageId="32" pageNumber="233">Figs. 18A,C</figureCitation>
) of the
<typeStatus id="5D2C29E33059FFB0FF53F9B7BC600B26" box="[166,259,1573,1597]" pageId="32" pageNumber="233" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4597ECC23059FFB0FEDBF9B4BC8C0B26" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[302,495,1573,1597]" class="Insecta" family="Carabidae" genus="Thysanotus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="233" phylum="Arthropoda" rank="species" species="bimaculatoides">
<emphasis id="B0E34B533059FFB0FEDBF9B4BC8C0B26" box="[302,495,1573,1597]" italics="true" pageId="32" pageNumber="233">T. bimaculatoides</emphasis>
</taxonomicName>
as well as those of a male of
<taxonomicName id="4597ECC23059FFB0FCB9F9B7BDA60B46" authority="(Alluaud 1936 c)" baseAuthorityName="Alluaud" baseAuthorityYear="1936" class="Insecta" family="Carabidae" genus="Thysanotus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="233" phylum="Arthropoda" rank="species" species="spinosus">
<emphasis id="B0E34B533059FFB0FCB9F9B7B94F0B26" box="[844,1068,1573,1597]" italics="true" pageId="32" pageNumber="233">Thysanotus spinosus</emphasis>
(
<bibRefCitation id="E606EAB03059FFB0FBCAF9B7BDDF0B46" author="ALLUAUD, C." pageId="32" pageNumber="233" pagination="1 - 10" refId="ref29048" refString="ALLUAUD, C. 1936 c. Carabiques nouveaux de Madagascar. Afra, Cahiers d'Entomologie 12: 1 - 10." type="journal article" year="1936">Alluaud 1936c</bibRefCitation>
)
</taxonomicName>
(
<figureCitation id="1AAC8BC43059FFB0FF20F9D7BC340B46" box="[213,343,1605,1629]" captionStart="FIGURE" captionStartId="32.[155,167,675,694]" captionTargetBox="[140,1163,222,658]" captionTargetId="figure-2339@32.[120,1176,216,672]" captionTargetPageId="32" captionText="FIGURE 18. Digital images of aedeagus of male genitalia. A, C. Left lateral aspect; B, D. Dorsal aspect. AB. Holotype male of Thysanotus bimaculatoides Kavanaugh &amp; Rainio sp. nov.; B, D. Thysanotus spinosus Alluaud (from Ranomafana National Park). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13799483" httpUri="https://zenodo.org/record/13799483/files/figure.png" pageId="32" pageNumber="233">Figs. 18B,D</figureCitation>
) [males of this species previously unknown], to complement the illustrations of other
<taxonomicName id="4597ECC23059FFB0FF23F9F7BCC70B66" box="[214,420,1637,1661]" pageId="32" pageNumber="233">
<emphasis id="B0E34B533059FFB0FF23F9F7BC2C0B66" box="[214,335,1637,1661]" italics="true" pageId="32" pageNumber="233">Thysanotus</emphasis>
species
</taxonomicName>
provided by Jeannel (1949) and
<bibRefCitation id="E606EAB03059FFB0FCF2F9F4BEF80B66" author="DEUVE, T." box="[775,923,1637,1661]" pageId="32" pageNumber="233" pagination="165 - 176" refId="ref30046" refString="DEUVE, T. 2010. Note sur le genre Thysanotus Chaudoir et quelques autres Lebiinae Pericalini de Madagascar [Coleoptera, Caraboidea, Harpalidae]. Revue Francaise d'Entomologie (N. S.) 32: 165 - 176." type="journal article" year="2010">Deuve (2010)</bibRefCitation>
.
</paragraph>
<paragraph id="822897413059FFB0FF5EF917B9030BA6" blockId="32.[123,1174,770,1725]" pageId="32" pageNumber="233">
<emphasis id="B0E34B533059FFB0FF5EF917BC930B87" bold="true" box="[171,496,1669,1693]" pageId="32" pageNumber="233">
<smallCapsWord id="84CE019D3059FFB0FF5EF917BC350B87" baselines="1687,1687" box="[171,342,1669,1693]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Geographical" pageId="32" pageNumber="233">GEOGRAPHICAL</smallCapsWord>
<smallCapsWord id="84CE019D3059FFB0FEA9F91BBC930B87" baselines="1687" box="[348,496,1673,1692]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="32" pageNumber="233">DISTRIBUTION</smallCapsWord>
</emphasis>
.— At present, known only from two localities,
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apart, in central
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. Both localities are on the eastern slope off the central plateau.
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.—
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and
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were collected by hand in the
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, about
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distance in from the edge of montane rainforest, at an elevation of
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.
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other
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was collected in a
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in montane rainforest at an elevation of
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.
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</paragraph>
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