175 lines
24 KiB
XML
175 lines
24 KiB
XML
<document id="B95CA6C8AABBCE4B6D54275BB9819248" ID-DOI="10.5281/zenodo.175414" ID-GBIF-Dataset="f485f2e0-6053-49c6-9863-4c71023258b3" ID-ISSN="11755326" ID-Zenodo-Dep="175414" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459808583420" checkinUser="plazi" docAuthor="Thomas, Richard & Hedges, Blair" docDate="2007" docId="004FE949FFA6FFDFFF3FFBDCFCDD2350" docLanguage="en" docName="zt01400p026.pdf" docOrigin="Zootaxa 1400" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Typhlops arator Thomas & Hedges, 2007, new species" docType="treatment" docVersion="10" lastPageNumber="20" masterDocId="FC769131FFB4FFCCFFA8FFC5FF942112" masterDocTitle="Eleven new species of snakes of the genus Typhlops (Serpentes: Typhlopidae) from Hispaniola and Cuba" masterLastPageNumber="26" masterPageNumber="1" pageNumber="19" updateTime="1698220755238" updateUser="plazi">
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<mods:mods id="7EE87FDD41E488F901C72FDF8611ED1D" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="41D441169724B36731F1F7DDBFA398BA">
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<mods:title id="CB42ADC2541195953BD610653E61CB9C">Eleven new species of snakes of the genus Typhlops (Serpentes: Typhlopidae) from Hispaniola and Cuba</mods:title>
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<mods:name id="38ADF12E437DED51DDA2ED0D200B6B9F" type="personal">
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<mods:namePart id="1CFFDFE7A0882B435FB895B8860014F6">Thomas, Richard</mods:namePart>
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<mods:name id="79A57A53FA55DA99194147F781F294A8" type="personal">
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<mods:namePart id="02D6EC41829A2B81173C5C555A4E45F6">Hedges, Blair</mods:namePart>
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<mods:typeOfResource id="E23C253A341F5C023F401C887C4F52AE">text</mods:typeOfResource>
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<mods:title id="990F9299EF03E41AF39048310585C4E4">Zootaxa</mods:title>
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<mods:part id="EE23FFD0E494852E40433ECFED58F12F">
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<mods:date id="9E134FF64D9AD4E9A7AE7ADB04B54BD7">2007</mods:date>
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<mods:number id="FE88CC1F226875C7868BBD2D24FD3500">1400</mods:number>
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<mods:classification id="158AC4313EBD165A1727130B733878C5">journal article</mods:classification>
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<mods:identifier id="EED023CC75C4BF5D6F226706D8A34292" type="DOI">10.5281/zenodo.175414</mods:identifier>
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<treatment id="004FE949FFA6FFDFFF3FFBDCFCDD2350" ID-DOI="http://doi.org/10.5281/zenodo.5677331" ID-GBIF-Taxon="119353301" ID-Zenodo-Dep="5677331" LSID="urn:lsid:plazi:treatment:004FE949FFA6FFDFFF3FFBDCFCDD2350" httpUri="http://treatment.plazi.org/id/004FE949FFA6FFDFFF3FFBDCFCDD2350" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
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<subSubSection id="C0FC0BD4FFA6FFDEFF3FFBDCFD1B2521" box="[151,655,1049,1075]" pageId="18" pageNumber="19" type="nomenclature">
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<paragraph id="8859585FFFA6FFDEFF3FFBDCFD1B2521" blockId="18.[151,655,1049,1075]" box="[151,655,1049,1075]" pageId="18" pageNumber="19">
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<heading id="D311EF33FFA6FFDEFF3FFBDCFD1B2521" bold="true" box="[151,655,1049,1075]" fontSize="11" level="1" pageId="18" pageNumber="19" reason="1">
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<emphasis id="BA92844DFFA6FFDEFF3FFBDCFD1B2521" bold="true" box="[151,655,1049,1075]" pageId="18" pageNumber="19">
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<taxonomicName id="4FE623DCFFA6FFDEFF3FFBDCFEC32521" box="[151,343,1049,1075]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="arator" status="sp. nov.">
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<emphasis id="BA92844DFFA6FFDEFF3FFBDCFEC32521" bold="true" box="[151,343,1049,1075]" italics="true" pageId="18" pageNumber="19">Typhlops arator</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A1A13936FFA6FFDEFEF6FBDCFE792521" box="[350,493,1049,1075]" pageId="18" pageNumber="19" rank="species">new species</taxonomicNameLabel>
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(
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<figureCitation id="10DD44DAFFA6FFDEFE55FBDCFDD02521" box="[509,580,1049,1075]" captionStart="FIGURE 8" captionStartId="13.[151,255,1722,1746]" captionTargetBox="[273,1319,349,1708]" captionTargetId="figure@13.[273,1320,349,1709]" captionTargetPageId="13" captionText="FIGURE 8. Head scalation in the Typhlops biminiensis Group, including seven new species from Cuba. (A) T. biminiensis (ASFS V 11655), from Elbow Cay, Cay Sal Bank, Bahamas. (B) T. paradoxus (ASFS V 12067), from Great Inagua, Bahamas. (C) T. epactius (ASFS V 11673), from Cayman Brac, Cayman Islands. (D) T. caymanensis (ASFS V 17112), from Grand Cayman, Cayman Islands. (E) T. perimychus (USNM 564800), holotype, from Guantanamo Bay Naval Station, Cuba. (F) T. arator (MNHNCu 191), holotype, from El Narigon, La Habana, Province, Cuba). (G) T. anousius (MNHNCu 4550), holotype, from 3.5 km E Tortuguilla, Guantánamo Province, Cuba. (H) T. notorachius (MNHNCu 4551), holotype, from 9.4 km W Imias, Guantánamo Province, Cuba. (I) T. contorhinus (MNHNCu 4552), holotype, from 4.9 km S La Tinta, Guantánamo Province, Cuba. (J) T. anchaurus (MNHNCu 4553), holotype, from within 1 km of Cueva de Agua (2.5 km airline NW Maisí), Guantánamo Province, Cuba. (K) T. satelles (SMF 61303), holotype, from Rancho Luna, about 12 km airline S Cienfuegos, Cienfuegos Province, Cuba." httpUri="https://zenodo.org/record/175421/files/figure.png" pageId="18" pageNumber="19">Fig. 8</figureCitation>
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F, 9B)
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C0FC0BD4FFA6FFDEFF3FFBA1FE1525E4" pageId="18" pageNumber="19" type="materials_examined">
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<paragraph id="8859585FFFA6FFDEFF3FFBA1FB1725B4" blockId="18.[151,1439,1124,2030]" pageId="18" pageNumber="19">
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<emphasis id="BA92844DFFA6FFDEFF3FFBA1FE98256F" box="[151,268,1124,1149]" italics="true" pageId="18" pageNumber="19">
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<typeStatus id="575DE6FDFFA6FFDEFF3FFBA1FE92256F" box="[151,262,1124,1149]" pageId="18" pageNumber="19" type="holotype">Holotype</typeStatus>
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:
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</emphasis>
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MNHNCu 191, a female, collected at “El Narigon and
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<collectingCountry id="F0F118CFFFA6FFDEFC65FBA1FBAE256C" box="[973,1082,1124,1150]" name="India" pageId="18" pageNumber="19">La India</collectingCountry>
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,” near Puerto Escondido, La Habana, Province,
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<collectingCountry id="F0F118CFFFA6FFDEFEDEFB49FE2325B4" box="[374,439,1164,1190]" name="Cuba" pageId="18" pageNumber="19">Cuba</collectingCountry>
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, by Ruben Regalado and Carlos Wotzkow, on
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<date id="FC587E9FFFA6FFDEFC7FFB49FBEB25B4" box="[983,1151,1164,1190]" pageId="18" pageNumber="19" value="1988-04-20">20 April 1988</date>
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.
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</paragraph>
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<paragraph id="8859585FFFA6FFDEFF6EFB71FE1525E4" blockId="18.[151,1439,1124,2030]" pageId="18" pageNumber="19">
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<emphasis id="BA92844DFFA6FFDEFF6EFB71FEA825DF" box="[198,316,1204,1229]" italics="true" pageId="18" pageNumber="19">
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<typeStatus id="575DE6FDFFA6FFDEFF6EFB71FEA125DF" box="[198,309,1204,1229]" pageId="18" pageNumber="19" type="paratype">Paratype</typeStatus>
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:
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</emphasis>
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<collectionCode id="EEF7C09AFFA6FFDEFEEBFB71FE0C25DC" LSID="urn:lsid:biocol.org:col:34871" box="[323,408,1204,1230]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="18" pageNumber="19">USNM</collectionCode>
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564784, same locality as
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<typeStatus id="575DE6FDFFA6FFDEFD61FB71FCA625DC" box="[713,818,1204,1230]" pageId="18" pageNumber="19" type="holotype">holotype</typeStatus>
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, collected by Richard Thomas and Daniel McCallister on
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<date id="FC587E9FFFA6FFDEFF49FB19FEE825E4" box="[225,380,1244,1270]" pageId="18" pageNumber="19" value="1990-07-10">10 July 1990</date>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C0FC0BD4FFA6FFDEFF6EFAC1FAF924F4" pageId="18" pageNumber="19" type="diagnosis">
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<paragraph id="8859585FFFA6FFDEFF6EFAC1FAF924F4" blockId="18.[151,1439,1124,2030]" pageId="18" pageNumber="19">
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<emphasis id="BA92844DFFA6FFDEFF6EFAC1FED3240F" box="[198,327,1284,1309]" italics="true" pageId="18" pageNumber="19">Diagnosis:</emphasis>
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A large species of the
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<taxonomicName id="4FE623DCFFA6FFDEFDC0FAC1FCF6240F" box="[616,866,1284,1309]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="biminiensis">
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<emphasis id="BA92844DFFA6FFDEFDC0FAC1FCF6240F" box="[616,866,1284,1309]" italics="true" pageId="18" pageNumber="19">Typhlops biminiensis</emphasis>
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</taxonomicName>
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Group that has a greater number of middorsal scales than any other West Indian species (578–579). Also, it has 26 scale rows anteriorly, more than any other West Indian species. It also has a uniquely shaped rostral (
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<figureCitation id="10DD44DAFFA6FFDEFCEBFA91FC1E247C" box="[835,906,1364,1390]" captionStart="FIGURE 8" captionStartId="13.[151,255,1722,1746]" captionTargetBox="[273,1319,349,1708]" captionTargetId="figure@13.[273,1320,349,1709]" captionTargetPageId="13" captionText="FIGURE 8. Head scalation in the Typhlops biminiensis Group, including seven new species from Cuba. (A) T. biminiensis (ASFS V 11655), from Elbow Cay, Cay Sal Bank, Bahamas. (B) T. paradoxus (ASFS V 12067), from Great Inagua, Bahamas. (C) T. epactius (ASFS V 11673), from Cayman Brac, Cayman Islands. (D) T. caymanensis (ASFS V 17112), from Grand Cayman, Cayman Islands. (E) T. perimychus (USNM 564800), holotype, from Guantanamo Bay Naval Station, Cuba. (F) T. arator (MNHNCu 191), holotype, from El Narigon, La Habana, Province, Cuba). (G) T. anousius (MNHNCu 4550), holotype, from 3.5 km E Tortuguilla, Guantánamo Province, Cuba. (H) T. notorachius (MNHNCu 4551), holotype, from 9.4 km W Imias, Guantánamo Province, Cuba. (I) T. contorhinus (MNHNCu 4552), holotype, from 4.9 km S La Tinta, Guantánamo Province, Cuba. (J) T. anchaurus (MNHNCu 4553), holotype, from within 1 km of Cueva de Agua (2.5 km airline NW Maisí), Guantánamo Province, Cuba. (K) T. satelles (SMF 61303), holotype, from Rancho Luna, about 12 km airline S Cienfuegos, Cienfuegos Province, Cuba." httpUri="https://zenodo.org/record/175421/files/figure.png" pageId="18" pageNumber="19">Fig. 8</figureCitation>
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), which differs from
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<taxonomicName id="4FE623DCFFA6FFDEFB2CFA91FAB2247F" box="[1156,1318,1364,1389]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="biminiensis">
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<emphasis id="BA92844DFFA6FFDEFB2CFA91FAB2247F" box="[1156,1318,1364,1389]" italics="true" pageId="18" pageNumber="19">T. biminiensis</emphasis>
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</taxonomicName>
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in having an acuminate posterior edge (as in other Cuban species), not broadly rounded (
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<figureCitation id="10DD44DAFFA6FFDEFB9DFAB9FBEF2484" box="[1077,1147,1404,1430]" captionStart="FIGURE 8" captionStartId="13.[151,255,1722,1746]" captionTargetBox="[273,1319,349,1708]" captionTargetId="figure@13.[273,1320,349,1709]" captionTargetPageId="13" captionText="FIGURE 8. Head scalation in the Typhlops biminiensis Group, including seven new species from Cuba. (A) T. biminiensis (ASFS V 11655), from Elbow Cay, Cay Sal Bank, Bahamas. (B) T. paradoxus (ASFS V 12067), from Great Inagua, Bahamas. (C) T. epactius (ASFS V 11673), from Cayman Brac, Cayman Islands. (D) T. caymanensis (ASFS V 17112), from Grand Cayman, Cayman Islands. (E) T. perimychus (USNM 564800), holotype, from Guantanamo Bay Naval Station, Cuba. (F) T. arator (MNHNCu 191), holotype, from El Narigon, La Habana, Province, Cuba). (G) T. anousius (MNHNCu 4550), holotype, from 3.5 km E Tortuguilla, Guantánamo Province, Cuba. (H) T. notorachius (MNHNCu 4551), holotype, from 9.4 km W Imias, Guantánamo Province, Cuba. (I) T. contorhinus (MNHNCu 4552), holotype, from 4.9 km S La Tinta, Guantánamo Province, Cuba. (J) T. anchaurus (MNHNCu 4553), holotype, from within 1 km of Cueva de Agua (2.5 km airline NW Maisí), Guantánamo Province, Cuba. (K) T. satelles (SMF 61303), holotype, from Rancho Luna, about 12 km airline S Cienfuegos, Cienfuegos Province, Cuba." httpUri="https://zenodo.org/record/175421/files/figure.png" pageId="18" pageNumber="19">Fig. 8</figureCitation>
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F). The rostral scale also rounds the tip of the snout without narrowing, which is unique among Cuban species in the group, reflected in a high RW2 (
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<figureCitation id="10DD44DAFFA6FFDEFE9BFA09FE1D24F4" box="[307,393,1484,1510]" captionStart="FIGURE 10" captionStartId="17.[151,255,1319,1343]" captionTargetBox="[293,1229,214,1276]" captionTargetId="figure@17.[279,1308,194,1295]" captionTargetPageId="17" captionText="FIGURE 10. Variation in measurements of Typhlops biminiensis and Cuban species of the Typhlops biminiensis Group. For all graphs (A – F), the following species are indicated by symbols: T. anchaurus (open diamond), T. anousius (closed square), T. arator (closed diamond), T. biminiensis (open circle), T. contorhinus (open triangle), T. notorachius (closed triangle), T. paradoxus (open square), T. perimychus (closed circle), T. satelles (“ X ”). Trendlines are indicated on some graphs for T. biminiensis and T. perimychus. See text for definition of abbreviations." httpUri="https://zenodo.org/record/175423/files/figure.png" pageId="18" pageNumber="19">Fig. 10</figureCitation>
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D).
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<taxonomicName id="4FE623DCFFA6FFDEFE1CFA09FDE424F7" box="[436,624,1484,1509]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="arator">
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<emphasis id="BA92844DFFA6FFDEFE1CFA09FDE424F7" box="[436,624,1484,1509]" italics="true" pageId="18" pageNumber="19">Typhlops arator</emphasis>
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</taxonomicName>
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additionally has a large protrusion (umbo) of the snout (
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<figureCitation id="10DD44DAFFA6FFDEFAAFFA09FAD824F4" box="[1287,1356,1484,1510]" captionStart="FIGURE 8" captionStartId="13.[151,255,1722,1746]" captionTargetBox="[273,1319,349,1708]" captionTargetId="figure@13.[273,1320,349,1709]" captionTargetPageId="13" captionText="FIGURE 8. Head scalation in the Typhlops biminiensis Group, including seven new species from Cuba. (A) T. biminiensis (ASFS V 11655), from Elbow Cay, Cay Sal Bank, Bahamas. (B) T. paradoxus (ASFS V 12067), from Great Inagua, Bahamas. (C) T. epactius (ASFS V 11673), from Cayman Brac, Cayman Islands. (D) T. caymanensis (ASFS V 17112), from Grand Cayman, Cayman Islands. (E) T. perimychus (USNM 564800), holotype, from Guantanamo Bay Naval Station, Cuba. (F) T. arator (MNHNCu 191), holotype, from El Narigon, La Habana, Province, Cuba). (G) T. anousius (MNHNCu 4550), holotype, from 3.5 km E Tortuguilla, Guantánamo Province, Cuba. (H) T. notorachius (MNHNCu 4551), holotype, from 9.4 km W Imias, Guantánamo Province, Cuba. (I) T. contorhinus (MNHNCu 4552), holotype, from 4.9 km S La Tinta, Guantánamo Province, Cuba. (J) T. anchaurus (MNHNCu 4553), holotype, from within 1 km of Cueva de Agua (2.5 km airline NW Maisí), Guantánamo Province, Cuba. (K) T. satelles (SMF 61303), holotype, from Rancho Luna, about 12 km airline S Cienfuegos, Cienfuegos Province, Cuba." httpUri="https://zenodo.org/record/175421/files/figure.png" pageId="18" pageNumber="19">Fig. 8</figureCitation>
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F).
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</paragraph>
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</subSubSection>
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<subSubSection id="C0FC0BD4FFA6FFDEFF6EFA31FF2D268C" pageId="18" pageNumber="19" type="description">
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<paragraph id="8859585FFFA6FFDEFF6EFA31FF2D268C" blockId="18.[151,1439,1124,2030]" pageId="18" pageNumber="19">
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<emphasis id="BA92844DFFA6FFDEFF6EFA31FECF271F" box="[198,347,1524,1549]" italics="true" pageId="18" pageNumber="19">Description:</emphasis>
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Snout narrow and ogival in outline, lobed, tapering from maximum width of head. Rostral broad and nearly parallelsided, curving gently mesiad to apex (RW1/RL1 0.97), distinctly flared on apex and with a prominent apical (not ventral) umbo; no labial flare. Anterior nasal width as a proportion of RW1 0.35. Preocular angle 110°, apex rounded; lower portion contacting labials 2 and 3 of upper labial series. OL 1/2 height, OS 0.19. Rostronasal pattern strongly divergent. Postocular about twice as high as long. First parietal wide and winglike, spanning two scale rows. Second parietal present on left and equal in size to the first, absent on right. TL
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<quantity id="4F1EF5BAFFA6FFDEFED7F921FE7027EC" box="[383,484,1764,1790]" metricMagnitude="-1" metricUnit="m" metricValue="4.6" pageId="18" pageNumber="19" unit="mm" value="460.0">460 mm</quantity>
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. TL/TA 46–48. TL/MBD 51–55. Middorsal scales 578–579. Scale rows 26 with posterior reduction to 24 rows occurring at 34–41% of TL. Coloration bicolor with dorsal pigmentation (very pale brown) ending abruptly along a midlateral or dorsolateral line by dropping out of pigmentation on individual scales; the pigmentation is so light that this is difficult to discern; pigmented scale rows at midbody 9– 11.
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</paragraph>
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</subSubSection>
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<subSubSection id="C0FC0BD4FFA6FFDEFF6EF869FD2D26FC" pageId="18" pageNumber="19" type="distribution">
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<paragraph id="8859585FFFA6FFDEFF6EF869FD2D26FC" blockId="18.[151,1439,1124,2030]" pageId="18" pageNumber="19">
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<emphasis id="BA92844DFFA6FFDEFF6EF869FEF426D7" box="[198,352,1964,1989]" italics="true" pageId="18" pageNumber="19">Distribution:</emphasis>
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Known only from Narigon (Habana Prov.), although possibly more widely distributed in western
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<collectingCountry id="F0F118CFFFA6FFDEFF51F811FEA326FC" box="[249,311,2004,2030]" name="Cuba" pageId="18" pageNumber="19">Cuba</collectingCountry>
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(see comments below) (
|
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<figureCitation id="10DD44DAFFA6FFDEFDFEF811FD3F26FC" box="[598,683,2004,2030]" captionStart="FIGURE 11" captionStartId="18.[151,255,865,889]" captionTargetBox="[266,1321,437,838]" captionTargetId="figure@18.[266,1321,437,839]" captionTargetPageId="18" captionText="FIGURE 11. Geographic distribution of seven species of blindsnakes (Typhlops) in Cuba. Closed triangles = T. perimychus, closed hexagons = T. arator, open circle = T. anousius, closed circle = T. notorachius, open square = T. contorhinus, open triangle = T. anchaurus, closed square = T. satelles." httpUri="https://zenodo.org/record/175424/files/figure.png" pageId="18" pageNumber="19">Fig. 11</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C0FC0BD4FFA7FFDFFF6EFF5DFB2D21A0" box="[198,1209,152,178]" pageId="19" pageNumber="20" type="etymology">
|
||
<paragraph id="8859585FFFA7FFDFFF6EFF5DFB2D21A0" blockId="19.[151,1437,152,578]" box="[198,1209,152,178]" pageId="19" pageNumber="20">
|
||
<emphasis id="BA92844DFFA7FFDFFF6EFF5DFED921A3" box="[198,333,152,177]" italics="true" pageId="19" pageNumber="20">Etymology:</emphasis>
|
||
From the Greek,
|
||
<taxonomicName id="4FE623DCFFA7FFDFFDB5FF5DFDF321A3" box="[541,615,152,177]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="arator">
|
||
<emphasis id="BA92844DFFA7FFDFFDB5FF5DFDF321A3" box="[541,615,152,177]" italics="true" pageId="19" pageNumber="20">arator</emphasis>
|
||
</taxonomicName>
|
||
, a plow, in allusion to the prominent rostral umbo.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C0FC0BD4FFA7FFDFFF6EFF05FCDD2350" pageId="19" pageNumber="20" type="discussion">
|
||
<paragraph id="8859585FFFA7FFDFFF6EFF05FEB920E0" blockId="19.[151,1437,152,578]" pageId="19" pageNumber="20">
|
||
<emphasis id="BA92844DFFA7FFDFFF6EFF05FED821CB" box="[198,332,192,217]" italics="true" pageId="19" pageNumber="20">Comments:</emphasis>
|
||
In addition to the
|
||
<typeStatus id="575DE6FDFFA7FFDFFD8DFF05FD1F21C8" box="[549,651,192,218]" pageId="19" pageNumber="20" type="holotype">holotype</typeStatus>
|
||
and
|
||
<typeStatus id="575DE6FDFFA7FFDFFD6DFF05FCB921C8" box="[709,813,192,218]" pageId="19" pageNumber="20" type="paratype">paratype</typeStatus>
|
||
, both from the same locality in La Habana Province, we are aware of a very large specimen collected by Julio Novo at Cueva de San Miguel near Viñales (Pinar del Rio Prov.) in Western
|
||
<collectingCountry id="F0F118CFFFA7FFDFFE34FED5FE4E2038" box="[412,474,272,298]" name="Cuba" pageId="19" pageNumber="20">Cuba</collectingCountry>
|
||
during the 1980s. It was even larger than the
|
||
<typeStatus id="575DE6FDFFA7FFDFFC54FED5FBF72038" box="[1020,1123,272,298]" pageId="19" pageNumber="20" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4FE623DCFFA7FFDFFB24FED5FB67203B" box="[1164,1267,272,297]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="arator">
|
||
<emphasis id="BA92844DFFA7FFDFFB24FED5FB67203B" box="[1164,1267,272,297]" italics="true" pageId="19" pageNumber="20">T. arator</emphasis>
|
||
</taxonomicName>
|
||
, and probably one of the largest scolecophidians known from the Western Hemisphere. Although that specimen apparently is lost (L. Diaz, pers. comm.), DNA sequences (S. B. Hedges, unpublished) indicate that it is most closely related to
|
||
<taxonomicName id="4FE623DCFFA7FFDFFEA3FE4DFEE620B3" box="[267,370,392,417]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="arator">
|
||
<emphasis id="BA92844DFFA7FFDFFEA3FE4DFEE620B3" box="[267,370,392,417]" italics="true" pageId="19" pageNumber="20">T. arator</emphasis>
|
||
</taxonomicName>
|
||
, suggesting that it is either a closely related species or that
|
||
<taxonomicName id="4FE623DCFFA7FFDFFB8CFE4DFB1820B3" box="[1060,1164,392,417]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="arator">
|
||
<emphasis id="BA92844DFFA7FFDFFB8CFE4DFB1820B3" box="[1060,1164,392,417]" italics="true" pageId="19" pageNumber="20">T. arator</emphasis>
|
||
</taxonomicName>
|
||
is more widely distributed in western
|
||
<collectingCountry id="F0F118CFFFA7FFDFFEF8FE75FE0620D8" box="[336,402,432,458]" name="Cuba" pageId="19" pageNumber="20">Cuba</collectingCountry>
|
||
. Until that specimen is located, or additional material is collected, we are unable to speculate further.
|
||
</paragraph>
|
||
<paragraph id="8859585FFFA7FFDFFF6EFDC5FCDD2350" blockId="19.[151,1437,152,578]" pageId="19" pageNumber="20">
|
||
The next new species having affinities with
|
||
<taxonomicName id="4FE623DCFFA7FFDFFD7CFDC5FCE3230B" box="[724,887,512,537]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="biminiensis">
|
||
<emphasis id="BA92844DFFA7FFDFFD7CFDC5FCE3230B" box="[724,887,512,537]" italics="true" pageId="19" pageNumber="20">T. biminiensis</emphasis>
|
||
</taxonomicName>
|
||
occurs on the southern coast of eastern
|
||
<collectingCountry id="F0F118CFFFA7FFDFFAFFFDC5FA0C2308" box="[1367,1432,512,538]" name="Cuba" pageId="19" pageNumber="20">Cuba</collectingCountry>
|
||
, east of the region of Guantánamo Bay. It may be known as
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |