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<document id="EDBB2B8E2B0E107E7464954EEEC8F0AB" ID-CLB-Dataset="22806" ID-DOI="10.11646/zootaxa.4407.1.2" ID-GBIF-Dataset="8e0aac3a-6556-4b96-a67e-fce9f0db0ba9" ID-ISSN="1175-5326" ID-Zenodo-Dep="3766290" ID-ZooBank="B45E9ABD-CB52-4EE7-91D6-783E8E958AC5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1587827908598" checkinUser="plazi" docAuthor="Short, Andrew Edward Z. &amp; Girón, Jennifer C." docDate="2018" docId="676A87E0FFA0B14F2392F996473A0A5A" docLanguage="en" docName="zootaxa.4407.1.2.pdf.imf" docOrigin="Zootaxa 4407 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Helochares (Hydrobaticus) MacLeay 1871" docType="treatment" docVersion="6" lastPageNumber="34" masterDocId="9B53FF98FFA4B14A2305FFE3420C0E4A" masterDocTitle="Review of the Helochares (Hydrobaticus) MacLeay of the New World (Coleoptera: Hydrophilidae: Acidocerinae)" masterLastPageNumber="50" masterPageNumber="29" pageNumber="33" updateTime="1698835862916" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="EED187DCF97910B0B2B46DBE752A42D4">Review of the Helochares (Hydrobaticus) MacLeay of the New World (Coleoptera: Hydrophilidae: Acidocerinae)</mods:title>
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<mods:namePart id="B1816F9572670382E1A05E332D2FAC44">Short, Andrew Edward Z.</mods:namePart>
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<mods:namePart id="6EFB2A637BC3DCD07DBC08E5EBFBD872">Girón, Jennifer C.</mods:namePart>
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<mods:title id="2D8B4F0C819DB06599F87A3307DBD7DD">Zootaxa</mods:title>
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<treatment id="676A87E0FFA0B14F2392F996473A0A5A" ID-DOI="http://doi.org/10.5281/zenodo.3803364" ID-GBIF-Taxon="163743978" ID-Zenodo-Dep="3803364" LSID="urn:lsid:plazi:treatment:676A87E0FFA0B14F2392F996473A0A5A" httpUri="http://treatment.plazi.org/id/676A87E0FFA0B14F2392F996473A0A5A" lastPageId="5" lastPageNumber="34" pageId="4" pageNumber="33">
<subSubSection id="A7D9657DFFA0B14E2392F996404E08C5" box="[151,578,1653,1679]" pageId="4" pageNumber="33" type="nomenclature">
<paragraph id="EF7C36F6FFA0B14E2392F996404E08C5" blockId="4.[151,578,1653,1679]" box="[151,578,1653,1679]" pageId="4" pageNumber="33">
<heading id="B434819AFFA0B14E2392F996404E08C5" bold="true" box="[151,578,1653,1679]" fontSize="11" level="1" pageId="4" pageNumber="33" reason="1">
<treatmentCitationGroup id="CFD311D8FFA0B14E2392F996404E08C5" box="[151,578,1653,1679]" pageId="4" pageNumber="33">
<emphasis id="DDB7EAE4FFA0B14E2392F996404E08C5" bold="true" box="[151,578,1653,1679]" pageId="4" pageNumber="33">
<treatmentCitation id="6E6210E7FFA0B14E2392F996403108C5" author="MacLeay, W. S." box="[151,573,1653,1679]" page="131" pageId="4" pageNumber="33" year="1871">
<taxonomicName id="28C34D75FFA0B14E2392F996403108C5" authority="MacLeay, 1871: 131" authorityName="MacLeay" authorityYear="1871" box="[151,573,1653,1679]" class="Insecta" family="Hydrophilidae" genus="Helochares" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="29" phylum="Arthropoda" rank="subGenus" subGenus="Hydrobaticus">
<emphasis id="DDB7EAE4FFA0B14E2392F996433608C5" bold="true" box="[151,314,1653,1679]" italics="true" pageId="4" pageNumber="33">Hydrobaticus</emphasis>
<bibRefCitation id="8B524B07FFA0B14E2244F996403108C5" author="MacLeay, W. S." box="[321,573,1653,1679]" pageId="4" pageNumber="33" pagination="79 - 205" refId="ref17101" refString="MacLeay, W. S. (1871) Notes on a collection of insects from Gayndah. Transactions of the Entomological Society of New South Wales, 2, 79 - 205. https: // doi. org / 10.5962 / bhl. title. 9427" type="journal article" year="1871">MacLeay, 1871: 131</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</emphasis>
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</heading>
</paragraph>
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<subSubSection id="A7D9657DFFA0B14E2392F95F46D009E7" pageId="4" pageNumber="33" type="diagnosis">
<paragraph id="EF7C36F6FFA0B14E2392F95F46D009E7" blockId="4.[151,1436,1724,2037]" pageId="4" pageNumber="33">
<emphasis id="DDB7EAE4FFA0B14E2392F95F431D089F" bold="true" box="[151,273,1724,1749]" pageId="4" pageNumber="33">Diagnosis.</emphasis>
Among New World
<taxonomicName id="28C34D75FFA0B14E2115F95F40A4089F" baseAuthorityName="Zaitzev" baseAuthorityYear="1908" box="[528,680,1724,1749]" class="Insecta" family="Hydrophilidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="33" phylum="Arthropoda" rank="subFamily" subFamily="Acidocerinae">Acidocerinae</taxonomicName>
, species of
<taxonomicName id="28C34D75FFA0B14E203CF95F41DC089E" authorityName="MacLeay" authorityYear="1871" box="[825,976,1724,1748]" class="Insecta" family="Hydrophilidae" genus="Helochares" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="48" phylum="Arthropoda" rank="subGenus" subGenus="Hydrobaticus">
<emphasis id="DDB7EAE4FFA0B14E203CF95F41DC089E" box="[825,976,1724,1748]" italics="true" pageId="4" pageNumber="33">Hydrobaticus</emphasis>
</taxonomicName>
can be distinguished by the following combination of characters: (1) antennae with 9 antennomeres, (2) fifth ventrite with a distinct apicomedial emargination lined with coarse setae, (3) mesoventrite without a distinctly elevated keel or protuberance, (4) elytra with ten rows of serial punctures (except
<emphasis id="DDB7EAE4FFA0B14E2166F8CA417E090A" box="[611,882,1832,1856]" italics="true" pageId="4" pageNumber="33">
<taxonomicName id="28C34D75FFA0B14E2166F8CA40F1090A" authorityName="Sharp" authorityYear="1882" box="[611,765,1832,1856]" class="Insecta" family="Hydrophilidae" genus="Helochares" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="33" phylum="Arthropoda" rank="species" species="championi">H. championi</taxonomicName>
,
<taxonomicName id="28C34D75FFA0B14E200DF8CA4162090A" authorityName="Short &amp; Girón" authorityYear="2018" box="[776,878,1832,1856]" class="Insecta" family="Hydrophilidae" genus="Helochares" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="33" phylum="Arthropoda" rank="species" species="laevis">H. laevis</taxonomicName>
,
</emphasis>
and
<taxonomicName id="28C34D75FFA0B14E20ADF8CA461A090A" authorityName="Short &amp; Girón" authorityYear="2018" box="[936,1046,1832,1856]" class="Insecta" family="Hydrophilidae" genus="Helochares" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="33" phylum="Arthropoda" rank="species" species="politus">
<emphasis id="DDB7EAE4FFA0B14E20ADF8CA461A090A" box="[936,1046,1832,1856]" italics="true" pageId="4" pageNumber="33">H. politus</emphasis>
</taxonomicName>
), and (5) aedeagus with parameres simple and fused, forming a tube-like structure, with median lobe appearing as a flat, tongue-like strap (
<figureCitation id="77F82A73FFA0B14E262DF8AF477E092E" box="[1320,1394,1868,1893]" captionStart="FIGURE 7" captionStartId="13.[151,250,1200,1222]" captionTargetBox="[347,1229,206,1166]" captionTargetId="figure@13.[340,1247,193,1179]" captionTargetPageId="13" captionText="FIGURE 7. Aedeagi of Helochares (Hydrobaticus) spp. (A) H. trujillo n. sp., (B) H. zamora n. sp., (C) H. maculicollis (Florida), (D) H. maculicollis (Kansas), (E) H. maculicollis (Pennsylvania), (F) H. nexus n. sp. (Venezuela), (G) H. nexus n. sp. (Panama), (H) H. nexus n. sp. (Ecuador), (I) H. politus n. sp., (J) H. laevis n. sp., (K) H. championi." figureDoi="http://doi.org/10.5281/zenodo.3766304" httpUri="https://zenodo.org/record/3766304/files/figure.png" pageId="4" pageNumber="33">Figs 7</figureCitation>
,
<figureCitation id="77F82A73FFA0B14E267BF8AF4782092E" box="[1406,1422,1868,1892]" captionStart="FIGURE 8" captionStartId="15.[151,250,1934,1956]" captionTargetBox="[360,1216,1351,1885]" captionTargetId="figure@15.[340,1247,1345,1913]" captionTargetPageId="15" captionText="FIGURE 8. Aedeagi of Helochares (Hydrobaticus) normatus. (A) Oregon, (B) California, (C) Arizona, (D) Mexico, (E) Costa Rica." figureDoi="http://doi.org/10.5281/zenodo.3766306" httpUri="https://zenodo.org/record/3766306/files/figure.png" pageId="4" pageNumber="33">8</figureCitation>
). It should be emphasized that these characters will only separate species of the New World species, and are not sufficient to distinguish
<taxonomicName id="28C34D75FFA0B14E22A3F877403109E6" authorityName="MacLeay" authorityYear="1871" box="[422,573,1940,1964]" class="Insecta" family="Hydrophilidae" genus="Helochares" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="48" phylum="Arthropoda" rank="subGenus" subGenus="Hydrobaticus">
<emphasis id="DDB7EAE4FFA0B14E22A3F877403109E6" box="[422,573,1940,1964]" italics="true" pageId="4" pageNumber="33">Hydrobaticus</emphasis>
</taxonomicName>
from species of other subgenera in other parts of the world.
</paragraph>
</subSubSection>
<subSubSection id="A7D9657DFFA0B14F23C2F85440A20DEF" lastPageId="5" lastPageNumber="34" pageId="4" pageNumber="33" type="description">
<paragraph id="EF7C36F6FFA0B14F23C2F85440A20DEF" blockId="4.[151,1436,1724,2037]" lastBlockId="5.[151,1437,151,1040]" lastPageId="5" lastPageNumber="34" pageId="4" pageNumber="33">
<emphasis id="DDB7EAE4FFA0B14E23C2F8544355099A" bold="true" box="[199,345,1975,2000]" pageId="4" pageNumber="33">Description.</emphasis>
Body size: 4.2–7.0 mm.
<emphasis id="DDB7EAE4FFA0B14E2186F85440C6099A" bold="true" box="[643,714,1975,2000]" pageId="4" pageNumber="33">Head.</emphasis>
Antennae with nine antennomeres, including three-segmented pubescent club. Maxillary palps curved inward and moderate in length, generally subequal to slightly longer in length to the width of head just anterior to eyes; inner face of palpomere 2 slightly curved; apical palpomere subequal in length to slightly shorter than penultimate. Labial palps short, distinctly shorter than mentum width. Mentum slightly concave centrally, strongly emarginated anteromedially; surface moderately to strongly sculptured; appearing mostly glabrous to possessing a few scattered short setae. Frons with series of irregular systematic, setae-bearing punctures anterior to each eye. Systematic punctures also present on clypeus and labrum but often blend with ground punctation, though occasionally appearing distinct. Eyes not or only slightly bulging, continuous with outline of the head or nearly so; not emarginated anteriorly by the frons.
<emphasis id="DDB7EAE4FFA1B14F2789FE8C46E00FC2" bold="true" box="[1164,1260,367,392]" pageId="5" pageNumber="34">Thorax.</emphasis>
Pronotum with systematic punctation in lateral thirds, each puncture usually bearing a short seta. Prosternum moderately wide, tectiform medially and usually with a narrow, fine longitudinal carina; very slightly elevated in anterior third. Mesoventrite with a low, weakly elevated bulge, never with an elevated keel, lamina, or tubercle. Metaventrite without a glabrous patch posteromedially, or at most with two small glabrous spots situated adjacent to midline in posterior quarter. Elytra with ten rows of very coarse serial punctures (except
<emphasis id="DDB7EAE4FFA1B14F272DFDC6474F0C76" box="[1064,1347,548,572]" italics="true" pageId="5" pageNumber="34">
<taxonomicName id="28C34D75FFA1B14F272DFDC646CB0C76" authorityName="Sharp" authorityYear="1882" box="[1064,1223,548,572]" class="Insecta" family="Hydrophilidae" genus="Helochares" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="34" phylum="Arthropoda" rank="species" species="championi">H. championi</taxonomicName>
,
<taxonomicName id="28C34D75FFA1B14F27D0FDC647330C76" authorityName="Short &amp; Girón" authorityYear="2018" box="[1237,1343,548,572]" class="Insecta" family="Hydrophilidae" genus="Helochares" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="34" phylum="Arthropoda" rank="species" species="laevis">H. laevis</taxonomicName>
,
</emphasis>
and
<taxonomicName id="28C34D75FFA1B14F2684FDC642EF0C2A" authorityName="Short &amp; Girón" authorityYear="2018" class="Insecta" family="Hydrophilidae" genus="Helochares" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="34" phylum="Arthropoda" rank="species" species="politus">
<emphasis id="DDB7EAE4FFA1B14F2684FDC642EF0C2A" italics="true" pageId="5" pageNumber="34">H. politus</emphasis>
</taxonomicName>
) which may or may not be depressed into grooves on at least the posterior half of the elytra. Procoxae set with sparse setae, but without thickened spines. Hind femora densely pubescent except on distal fifth–sixth or less. All tarsi with five tarsomeres.
<emphasis id="DDB7EAE4FFA1B14F22F2FD6C407F0CE2" bold="true" box="[503,627,655,680]" pageId="5" pageNumber="34">Abdomen.</emphasis>
With five pubescent ventrites, density of setae ranging from sparse to very dense. Fifth ventrite distinctly emarginated apicomedially, emargination set with thickened setae. Aedeagus (
<figureCitation id="77F82A73FFA1B14F239AFD3443010CBA" box="[159,269,727,752]" captionStart-0="FIGURE 7" captionStart-1="FIGURE 8" captionStartId-0="13.[151,250,1200,1222]" captionStartId-1="15.[151,250,1934,1956]" captionTargetBox-0="[347,1229,206,1166]" captionTargetBox-1="[360,1216,1351,1885]" captionTargetId-0="figure@13.[340,1247,193,1179]" captionTargetId-1="figure@15.[340,1247,1345,1913]" captionTargetPageId-0="13" captionTargetPageId-1="15" captionText-0="FIGURE 7. Aedeagi of Helochares (Hydrobaticus) spp. (A) H. trujillo n. sp., (B) H. zamora n. sp., (C) H. maculicollis (Florida), (D) H. maculicollis (Kansas), (E) H. maculicollis (Pennsylvania), (F) H. nexus n. sp. (Venezuela), (G) H. nexus n. sp. (Panama), (H) H. nexus n. sp. (Ecuador), (I) H. politus n. sp., (J) H. laevis n. sp., (K) H. championi." captionText-1="FIGURE 8. Aedeagi of Helochares (Hydrobaticus) normatus. (A) Oregon, (B) California, (C) Arizona, (D) Mexico, (E) Costa Rica." figureDoi-0="http://doi.org/10.5281/zenodo.3766304" figureDoi-1="http://doi.org/10.5281/zenodo.3766306" httpUri-0="https://zenodo.org/record/3766304/files/figure.png" httpUri-1="https://zenodo.org/record/3766306/files/figure.png" pageId="5" pageNumber="34">Figs. 7–8</figureCitation>
) 2.0 to 2.8-times longer than its widest point, with basal piece short, nearly one third the length of parameres; parameres basally fused together into a dorsoventrally flattened tube with basal margin rounded to mesally notched; lateral outer margins of parameres slightly sinuate, parallel along basal half or third, continuing nearly parallel to apically diverging; widest point of aedeagus usually near apex of parameres; widest point of each paramere narrower than median lobe; median lobe with lateral basal apodemes, nearly as long as median lobe; gonopore situated on distal half of median lobe.
</paragraph>
</subSubSection>
<subSubSection id="A7D9657DFFA1B14F23C2FC4C473A0A5A" pageId="5" pageNumber="34" type="discussion">
<paragraph id="EF7C36F6FFA1B14F23C2FC4C473A0A5A" blockId="5.[151,1437,151,1040]" pageId="5" pageNumber="34">
<emphasis id="DDB7EAE4FFA1B14F23C2FC4C43350D82" bold="true" box="[199,313,943,968]" pageId="5" pageNumber="34">Remarks.</emphasis>
As previously mentioned, characters used to define the subgenus as a whole do not seem to demarcate natural groups of species. The diagnosis that is provided here should be used to separate Nearctic and Neotropical members of
<taxonomicName id="28C34D75FFA1B14F2225FC1B43B80A5A" authorityName="MacLeay" authorityYear="1871" box="[288,436,1016,1040]" class="Insecta" family="Hydrophilidae" genus="Helochares" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="48" phylum="Arthropoda" rank="subGenus" subGenus="Hydrobaticus">
<emphasis id="DDB7EAE4FFA1B14F2225FC1B43B80A5A" box="[288,436,1016,1040]" italics="true" pageId="5" pageNumber="34">Hydrobaticus</emphasis>
</taxonomicName>
only, though some of these features are common to the subgenus in other regions.
</paragraph>
</subSubSection>
</treatment>
</document>
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