343 lines
41 KiB
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343 lines
41 KiB
XML
<document id="8583300FCA6450ACB9E7D990DD2A0029" ID-DOI="10.5252/zoosystema2020v42a6" ID-GBIF-Dataset="69e88778-6ab9-44ad-bce0-bb84eb35391b" ID-ISSN="1638-9387" ID-Zenodo-Dep="3695831" ID-ZooBank="urn:lsid:zoobank.org:pub:C87A10FB-E817-4293-96FD-00C2EF82D371" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1583259932333" checkinUser="felipe" docAuthor="Theil, Emma Palacios & Felder, Darryl L." docDate="2020" docId="DE24878AF655D62D3240C7DB585FF853" docLanguage="en" docName="zoosystema42.6.85-103.pdf.imf" docOrigin="Zoosystema 42 (6)" docStyle="DocumentStyle:5CC576F744449C1860B4E72ABD884631.7:Zoosystema.2020-.journal_article" docStyleId="5CC576F744449C1860B4E72ABD884631" docStyleName="Zoosystema.2020-.journal_article" docStyleVersion="7" docTitle="Scleroplax Rathbun 1894" docType="treatment" docVersion="6" lastPageNumber="98" masterDocId="221DFFF2F65AD6223355C30D5C69FFBE" masterDocTitle="Phylogeny of the genus Pinnixa White, 1846 (Crustacea, Brachyura, Pinnotheridae) and allies inferred from mitochondrial and nuclear molecular markers, with generic reassignment of twenty-one species" masterLastPageNumber="103" masterPageNumber="85" pageNumber="98" updateTime="1698724363688" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="6882A35A33F3E217C4452AA807BB4E8D">Phylogeny of the genus Pinnixa White, 1846 (Crustacea, Brachyura, Pinnotheridae) and allies inferred from mitochondrial and nuclear molecular markers, with generic reassignment of twenty-one species</mods:title>
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<mods:namePart id="1246C2AA38E674E651675916107E6125">Theil, Emma Palacios</mods:namePart>
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<mods:namePart id="56B0F5AED25672ECA7B1ADC34A5F7B82">Felder, Darryl L.</mods:namePart>
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<mods:title id="CD0218500AF96EF1E1EBA4E1D603F769">Zoosystema</mods:title>
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<mods:part id="FB9E69D2CE03A7802E92806C44F7A1C3">
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<mods:date id="1ED995D6220D81DAC0647E5713CC69BB">2020</mods:date>
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<mods:number id="D3E3AFB6347C3ED035F178FD46644999">2020-03-03</mods:number>
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<mods:number id="3BE35DB5D53998CF2737CBCABA448235">42</mods:number>
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<mods:identifier id="76B15D720DAFCD9C640D308BDDA025D1" type="DOI">10.5252/zoosystema2020v42a6</mods:identifier>
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<treatment id="DE24878AF655D62D3240C7DB585FF853" ID-DOI="http://doi.org/10.5281/zenodo.3703644" ID-GBIF-Taxon="162382195" ID-Zenodo-Dep="3703644" LSID="urn:lsid:plazi:treatment:DE24878AF655D62D3240C7DB585FF853" httpUri="http://treatment.plazi.org/id/DE24878AF655D62D3240C7DB585FF853" lastPageNumber="98" pageId="15" pageNumber="98">
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<subSubSection id="1E976517F655D62D3240C7DB5E1FFB4E" box="[277,630,1238,1264]" pageId="15" pageNumber="98" type="nomenclature">
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<paragraph id="5632369CF655D62D3240C7DB5E1FFB4E" blockId="15.[277,630,1238,1264]" box="[277,630,1238,1264]" pageId="15" pageNumber="98">
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<heading id="0D7A81F0F655D62D3240C7DB5E1FFB4E" box="[277,630,1238,1264]" centered="true" fontSize="11" level="2" pageId="15" pageNumber="98" reason="2">
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Genus
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<taxonomicName id="918D4D1FF655D62D3237C7DB5E1FFB4E" ID-CoL="7PZ4B" authority="Rathbun, 1894" authorityName="Rathbun" authorityYear="1894" box="[354,630,1238,1264]" class="Malacostraca" family="Pinnotheridae" genus="Scleroplax" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="98" phylum="Arthropoda" rank="genus">
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<emphasis id="64F9EA8EF655D62D3237C7DB5DA3FB4E" box="[354,458,1238,1264]" italics="true" pageId="15" pageNumber="98">Scleroplax</emphasis>
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<bibRefCitation id="321C4B6DF655D62D3284C7DB5E1FFB4E" author="Rathbun" box="[465,630,1238,1264]" firstAuthor="Rathbun" pageId="15" pageNumber="98" pagination="223 - 260" refId="ref18461" refString="RATHBUN M. J. 1894. - Scientific results of explorations by the U. S. Fish Commission Steamer Albatross. No. XXIV - Descriptions of new genera and species of crabs from the west coast of North America and the Sandwich Islands. Proceedings U. S. National Museum 16: 223 - 260. https: // doi. org / 10.5479 / si. 00963801.933.223" type="journal article" year="1894">Rathbun, 1894</bibRefCitation>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="1E976517F655D62D33D1C6125DDEFA89" box="[132,439,1311,1335]" pageId="15" pageNumber="98" type="reference_group">
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<paragraph id="5632369CF655D62D33D1C6125DDEFA89" blockId="15.[132,439,1311,1335]" box="[132,439,1311,1335]" pageId="15" pageNumber="98">
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<treatmentCitationGroup id="769D11B2F655D62D33D1C6125DDEFA89" box="[132,439,1311,1335]" pageId="15" pageNumber="98">
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<taxonomicName id="918D4D1FF655D62D33D1C6125DD8FA89" ID-CoL="7PZ4B" authority="Rathbun, 1894: 250" authorityName="Rathbun" authorityPageNumber="250" authorityYear="1894" box="[132,433,1311,1335]" class="Malacostraca" family="Pinnotheridae" genus="Scleroplax" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="98" phylum="Arthropoda" rank="genus">
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<emphasis id="64F9EA8EF655D62D33D1C6125C8BFA89" box="[132,226,1311,1335]" italics="true" pageId="15" pageNumber="98">Scleroplax</emphasis>
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<treatmentCitation id="D72C108DF655D62D33BDC6125DD8FA89" author="RATHBUN M. J." box="[232,433,1311,1335]" page="250" pageId="15" pageNumber="98" year="1894">
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<bibRefCitation id="321C4B6DF655D62D33BDC6125DD8FA89" author="Rathbun" box="[232,433,1311,1335]" firstAuthor="Rathbun" pageId="15" pageNumber="98" pagination="223 - 260" refId="ref18461" refString="RATHBUN M. J. 1894. - Scientific results of explorations by the U. S. Fish Commission Steamer Albatross. No. XXIV - Descriptions of new genera and species of crabs from the west coast of North America and the Sandwich Islands. Proceedings U. S. National Museum 16: 223 - 260. https: // doi. org / 10.5479 / si. 00963801.933.223" type="journal article" year="1894">Rathbun, 1894: 250</bibRefCitation>
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</treatmentCitation>
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</taxonomicName>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="1E976517F655D62D33D1C6525E08F939" pageId="15" pageNumber="98" type="materials_examined">
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<paragraph id="5632369CF655D62D33D1C6525D1DFA2C" blockId="15.[132,775,1375,1426]" pageId="15" pageNumber="98">
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<collectionCode id="309CAE59F655D62D33D1C6525D69FACB" box="[132,256,1375,1399]" pageId="15" pageNumber="98">
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<typeStatus id="8936883EF655D62D33D1C6525CDCFACA" box="[132,181,1375,1399]" pageId="15" pageNumber="98">TYPE</typeStatus>
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SPECIES
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</collectionCode>
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. —
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<taxonomicName id="918D4D1FF655D62D327CC6525E17FAC9" authority="Rathbun, 1894" authorityName="Rathbun" authorityYear="1894" box="[297,638,1375,1399]" class="Malacostraca" family="Pinnotheridae" genus="Scleroplax" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="98" phylum="Arthropoda" rank="species" species="granulata">
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<emphasis id="64F9EA8EF655D62D327CC6525D8AFAC9" box="[297,483,1375,1399]" italics="true" pageId="15" pageNumber="98">Scleroplax granulata</emphasis>
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<bibRefCitation id="321C4B6DF655D62D32BDC6525E17FAC9" author="Rathbun" box="[488,638,1375,1399]" firstAuthor="Rathbun" pageId="15" pageNumber="98" pagination="223 - 260" refId="ref18461" refString="RATHBUN M. J. 1894. - Scientific results of explorations by the U. S. Fish Commission Steamer Albatross. No. XXIV - Descriptions of new genera and species of crabs from the west coast of North America and the Sandwich Islands. Proceedings U. S. National Museum 16: 223 - 260. https: // doi. org / 10.5479 / si. 00963801.933.223" type="journal article" year="1894">Rathbun, 1894</bibRefCitation>
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</taxonomicName>
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, by monotypy when genus was erected.
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</paragraph>
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<paragraph id="5632369CF655D62D33D1C6B85E08F939" blockId="15.[132,776,1461,1671]" pageId="15" pageNumber="98">
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<collectionCode id="309CAE59F655D62D33D1C6B85D01FA74" box="[132,360,1461,1485]" pageId="15" pageNumber="98">ORIGINAL DIAGNOSIS BY</collectionCode>
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<bibRefCitation id="321C4B6DF655D62D3238C6B85E7BFA73" author="RATHBUN" box="[365,530,1461,1485]" firstAuthor="RATHBUN" pageId="15" pageNumber="98" pagination="123 - 184" refId="ref18600" refString="RATHBUN M. J. 1918. - The grapsoid crabs of America. Bulletin of the United States National Museum 103: 123 - 184, plates 54 - 66." type="journal article" year="1918">RATHBUN (1918)</bibRefCitation>
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. — “Carapace transverse, subpentagonal, hard, very convex, regions scarcely indicated, lower or true antero-lateral margin curving gradually into postero-lateral margin, not forming an angle with it as in
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<emphasis id="64F9EA8EF655D62D3141C50B5E0FF9A3" box="[532,614,1542,1565]" italics="true" pageId="15" pageNumber="98">
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<taxonomicName id="918D4D1FF655D62D3141C50B5E0BF9A3" authorityName=", Say" authorityYear="1818" box="[532,610,1542,1565]" class="Malacostraca" family="Pinnotheridae" genus="Pinnixa" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="98" phylum="Arthropoda" rank="genus">Pinnixa</taxonomicName>
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.
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</emphasis>
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Ambulatory legs similar, third longest but not unusually long, fourth not noticeably reduced. Ischium of outer maxillipeds rudimentary, merus oblique, palpus three-jointed, the last joint articulating near proximal end of preceding joint. Only a single species known.”
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</paragraph>
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</subSubSection>
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<subSubSection id="1E976517F655D62D33D1C5A75FE6FDFF" pageId="15" pageNumber="98" type="diagnosis">
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<paragraph id="5632369CF655D62D33D1C5A75E9AF8DF" blockId="15.[132,775,1706,1889]" pageId="15" pageNumber="98">
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DIAGNOSIS OF THE GENUS AS MODIFIED BY
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<bibRefCitation id="321C4B6DF655D62D316BC5A75EB7F97C" author="CAMPOS" box="[574,734,1706,1730]" firstAuthor="CAMPOS" pageId="15" pageNumber="98" pagination="33 - 41" refId="ref17258" refString="CAMPOS E. 2006. - Systematics of the genus Scleroplax Rathbun, 1893 (Crustacea: Brachyura: Pinnotheridae). Zootaxa 1344: 33 - 41. https: // doi. org / 10.11646 / zootaxa. 1344.1.3" type="journal article" year="2006">CAMPOS (2006)</bibRefCitation>
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. — “Carapace hard, subheptagonal, highly convex dorsally, anterolateral margins not forming angle with posterolateral margins; MXP3 [= third maxilliped] slightly oblique, covers buccal cavity, ischio-merus subtrapezoidal, propodus extending to end of dactylus, both spoonshaped and larger than carpus. WL1-4 [= walking leg] of similar shape, third pair slightly longer, fourth not noticeably reduced.”
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</paragraph>
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<paragraph id="5632369CF655D62D33D1C4895F83FE61" blockId="15.[132,775,1924,2028]" lastBlockId="15.[813,1456,217,479]" pageId="15" pageNumber="98">
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DIAGNOSIS. — (Modified from
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<bibRefCitation id="321C4B6DF655D62D32E1C4885E2BF823" author="Rathbun" box="[436,578,1925,1949]" firstAuthor="Rathbun" pageId="15" pageNumber="98" pagination="123 - 184" refId="ref18600" refString="RATHBUN M. J. 1918. - The grapsoid crabs of America. Bulletin of the United States National Museum 103: 123 - 184, plates 54 - 66." type="journal article" year="1918">Rathbun 1918</bibRefCitation>
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and
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<bibRefCitation id="321C4B6DF655D62D3125C4885E90F823" author="Campos" box="[624,761,1925,1949]" firstAuthor="Campos" pageId="15" pageNumber="98" pagination="33 - 41" refId="ref17258" refString="CAMPOS E. 2006. - Systematics of the genus Scleroplax Rathbun, 1893 (Crustacea: Brachyura: Pinnotheridae). Zootaxa 1344: 33 - 41. https: // doi. org / 10.11646 / zootaxa. 1344.1.3" type="journal article" year="2006">Campos 2006</bibRefCitation>
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). Carapace transverse, subpentagonal or oblong, hard, very convex, anterolateral margins not forming an acute angle with posterolateral margins; cardiac ridge, if present, not extending entirely across carapace. Third maxilliped slightly oblique, covering buccal cavity, ischiomerus subtrapezoidal; propodus and dactylus elongate, longer than carpus; dactylus inserting near base of propodus, reaching end of propodus or slightly beyond. Male cheliped strong, fixed finger somewhat shortened, straight; female cheliped feeble, fixed finger straight; external palm surface sometimes with longitudinal line of tubercles. Walking pereopods subequal, cylindrical, relative lengths P4> P3 ≥ P2> P5. Male pleon tapering toward end, telson subsemicircular; first pleonal somite lacking gonopodal plate between gonopods.
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</paragraph>
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<paragraph id="5632369CF655D62D3078C1035FE6FDFF" blockId="15.[813,1456,526,683]" pageId="15" pageNumber="98">
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ADDITIONAL SPECIES. —
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<taxonomicName id="918D4D1FF655D62D377FC1025927FD98" authority="(Dana, 1851)" authorityName="Theil & Felder" authorityYear="2020" baseAuthorityName="Dana" baseAuthorityYear="1851" box="[1066,1358,526,551]" class="Malacostraca" family="Pinnotheridae" genus="Scleroplax" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="98" phylum="Arthropoda" rank="species" species="faba" status="comb. nov.">
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<emphasis id="64F9EA8EF655D62D377FC10258D5FD99" box="[1066,1212,527,551]" italics="true" pageId="15" pageNumber="98">Scleroplax faba</emphasis>
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(Dana, 1851)
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</taxonomicName>
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<taxonomicNameLabel id="7FCA57F5F655D62D3600C10259D9FD99" box="[1365,1456,527,551]" pageId="15" pageNumber="98" rank="species">n. comb.</taxonomicNameLabel>
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[
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<taxonomicName id="918D4D1FF655D62D3060C1275FE9FDFF" authorityName=", Say" authorityYear="1818" box="[821,896,554,577]" class="Malacostraca" family="Pinnotheridae" genus="Pinnixa" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="98" phylum="Arthropoda" rank="genus">
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<emphasis id="64F9EA8EF655D62D3060C1275FE9FDFF" box="[821,896,554,577]" italics="true" pageId="15" pageNumber="98">Pinnixa</emphasis>
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</taxonomicName>
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];
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</paragraph>
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</subSubSection>
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<subSubSection id="1E976517F655D62D3078C14E592DFD14" pageId="15" pageNumber="98" type="reference_group">
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<paragraph id="5632369CF655D62D3078C14E592FFDCB" blockId="15.[813,1456,526,683]" pageId="15" pageNumber="98">
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<taxonomicName id="918D4D1FF655D62D3078C14E58CCFDE2" ID-CoL="84H9T" authority="(Rathbun, 1918)" authorityName="Theil & Felder" authorityYear="2020" baseAuthorityName="Rathbun" baseAuthorityYear="1918" box="[813,1189,579,604]" class="Malacostraca" family="Pinnotheridae" genus="Scleroplax" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="98" phylum="Arthropoda" rank="species" species="franciscana" status="comb. nov.">
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<emphasis id="64F9EA8EF655D62D3078C14E5F90FDE2" box="[813,1017,579,604]" italics="true" pageId="15" pageNumber="98">Scleroplax franciscana</emphasis>
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(
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<bibRefCitation id="321C4B6DF655D62D3753C14E58F6FDE2" author="Rathbun" box="[1030,1183,579,604]" firstAuthor="Rathbun" pageId="15" pageNumber="98" pagination="123 - 184" refId="ref18600" refString="RATHBUN M. J. 1918. - The grapsoid crabs of America. Bulletin of the United States National Museum 103: 123 - 184, plates 54 - 66." type="journal article" year="1918">Rathbun, 1918</bibRefCitation>
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)
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</taxonomicName>
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<taxonomicNameLabel id="7FCA57F5F655D62D37FEC149596AFDE5" box="[1195,1283,579,603]" pageId="15" pageNumber="98" rank="species">n. comb.</taxonomicNameLabel>
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[
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<taxonomicName id="918D4D1FF655D62D3644C1495935FDE5" ID-CoL="7PS58" authorityName=", Say" authorityYear="1818" box="[1297,1372,580,603]" class="Malacostraca" family="Pinnotheridae" genus="Pinnixa" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="98" phylum="Arthropoda" rank="genus">
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<emphasis id="64F9EA8EF655D62D3644C1495935FDE5" box="[1297,1372,580,603]" italics="true" pageId="15" pageNumber="98">Pinnixa</emphasis>
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</taxonomicName>
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];
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<taxonomicName id="918D4D1FF655D62D3078C15358E9FDC8" ID-CoL="84H9V" authority="(Holmes, 1894)" authorityName="Theil & Felder" authorityYear="2020" baseAuthorityName="Holmes" baseAuthorityYear="1894" box="[813,1152,606,630]" class="Malacostraca" family="Pinnotheridae" genus="Scleroplax" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="98" phylum="Arthropoda" rank="species" species="littoralis" status="comb. nov.">
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<emphasis id="64F9EA8EF655D62D3078C1535FB7FDC8" box="[813,990,606,630]" italics="true" pageId="15" pageNumber="98">Scleroplax littoralis</emphasis>
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(Holmes, 1894)
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</taxonomicName>
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<taxonomicNameLabel id="7FCA57F5F655D62D37D2C15358B6FDC8" box="[1159,1247,606,630]" pageId="15" pageNumber="98" rank="species">n. comb.</taxonomicNameLabel>
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[
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<taxonomicName id="918D4D1FF655D62D37B9C152595EFDC8" ID-CoL="7PS58" authorityName=", Say" authorityYear="1818" box="[1260,1335,607,630]" class="Malacostraca" family="Pinnotheridae" genus="Pinnixa" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="98" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="64F9EA8EF655D62D37B9C152595EFDC8" box="[1260,1335,607,630]" italics="true" pageId="15" pageNumber="98">Pinnixa</emphasis>
|
|
</taxonomicName>
|
|
];
|
|
</paragraph>
|
|
<paragraph id="5632369CF655D62D3078C175592DFD14" blockId="15.[813,1456,526,683]" pageId="15" pageNumber="98">
|
|
<taxonomicName id="918D4D1FF655D62D3078C17558EEFD2F" ID-CoL="84H9W" authority="(Rathbun, 1918)" authorityName="Theil & Felder" authorityYear="2020" baseAuthorityName="Rathbun" baseAuthorityYear="1918" box="[813,1159,632,657]" class="Malacostraca" family="Pinnotheridae" genus="Scleroplax" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="98" phylum="Arthropoda" rank="species" species="schmitti" status="comb. nov.">
|
|
<emphasis id="64F9EA8EF655D62D3078C1755FB2FD2E" box="[813,987,632,656]" italics="true" pageId="15" pageNumber="98">Scleroplax schmitti</emphasis>
|
|
(
|
|
<bibRefCitation id="321C4B6DF655D62D30BDC17558E9FD2F" author="Rathbun" box="[1000,1152,632,657]" firstAuthor="Rathbun" pageId="15" pageNumber="98" pagination="123 - 184" refId="ref18600" refString="RATHBUN M. J. 1918. - The grapsoid crabs of America. Bulletin of the United States National Museum 103: 123 - 184, plates 54 - 66." type="journal article" year="1918">Rathbun, 1918</bibRefCitation>
|
|
)
|
|
</taxonomicName>
|
|
<taxonomicNameLabel id="7FCA57F5F655D62D37D8C174588CFD2E" box="[1165,1253,632,656]" pageId="15" pageNumber="98" rank="species">n. comb.</taxonomicNameLabel>
|
|
[
|
|
<taxonomicName id="918D4D1FF655D62D37A6C1745957FD2E" ID-CoL="7PS58" authorityName=", Say" authorityYear="1818" box="[1267,1342,633,656]" class="Malacostraca" family="Pinnotheridae" genus="Pinnixa" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="98" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="64F9EA8EF655D62D37A6C1745957FD2E" box="[1267,1342,633,656]" italics="true" pageId="15" pageNumber="98">Pinnixa</emphasis>
|
|
</taxonomicName>
|
|
];
|
|
<taxonomicName id="918D4D1FF655D62D3078C19E5817FD15" ID-CoL="84H9X" authority="(Holmes, 1894)" authorityName="Theil & Felder" authorityYear="2020" baseAuthorityName="Holmes" baseAuthorityYear="1894" box="[813,1150,659,683]" class="Malacostraca" family="Pinnotheridae" genus="Scleroplax" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="98" phylum="Arthropoda" rank="species" species="tubicola" status="comb. nov.">
|
|
<emphasis id="64F9EA8EF655D62D3078C19E5FB5FD15" box="[813,988,659,683]" italics="true" pageId="15" pageNumber="98">Scleroplax tubicola</emphasis>
|
|
(Holmes, 1894)
|
|
</taxonomicName>
|
|
<taxonomicNameLabel id="7FCA57F5F655D62D37D1C19E58B5FD15" box="[1156,1244,659,683]" pageId="15" pageNumber="98" rank="species">n. comb.</taxonomicNameLabel>
|
|
[
|
|
<taxonomicName id="918D4D1FF655D62D37BFC199595CFD15" ID-CoL="7PS58" authorityName=", Say" authorityYear="1818" box="[1258,1333,660,683]" class="Malacostraca" family="Pinnotheridae" genus="Pinnixa" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="98" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="64F9EA8EF655D62D37BFC199595CFD15" box="[1258,1333,660,683]" italics="true" pageId="15" pageNumber="98">Pinnixa</emphasis>
|
|
</taxonomicName>
|
|
].
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="1E976517F655D62D3078C1D75F93FCE3" pageId="15" pageNumber="98" type="materials_examined">
|
|
<paragraph id="5632369CF655D62D3078C1D75840FC96" blockId="15.[813,1456,730,1046]" pageId="15" pageNumber="98">
|
|
MATERIAL EXAMINED. — In addition to the material included in the phylogenetic analyses (
|
|
<tableCitation id="1B0F0327F655D62D3769C1F858E0FCB3" box="[1084,1161,757,781]" captionStart="TABLE" captionStartId="5.[132,143,221,239]" captionText="TABLE 1. — Pinnotherid specimens used in phylogenetic analyses of the genus Pinnixa White, 1846 s.l. and allies. Names shown are in accord with Pinnixa s.l. prior to revisions in present paper. See “Material and Methods” for museum abbreviations." httpUri="http://table.plazi.org/id/02F26614F65FD62733D1C3D05F1CFEB8" pageId="15" pageNumber="98" tableUuid="02F26614F65FD62733D1C3D05F1CFEB8">Table 1</tableCitation>
|
|
) the following samples were available for examination:
|
|
</paragraph>
|
|
<paragraph id="5632369CF655D62D3078C0275F93FCE3" blockId="15.[813,1456,730,1046]" pageId="15" pageNumber="98">
|
|
<materialsCitation id="E6E53CC1F655D62D3078C0275F9FFCE3" ID-GBIF-Occurrence="2573304558" collectingLocationText="or DreamTaqTM Green DNA Polymerase, Fermentas, currently ThermoFisher Scientific, Waltham, MA, USA) in a Stratagene® Robocycler® Gradient 96 (Santa Clara, CA, USA). Three fragments were targeted: 1) a mitochondrial complex of about 830 bp consisting of part of the 16S rRNA gene, the tRNA-Leu, and part of the gene for NADH dehydrogenase subunit 1 (NADH1), 2) part of the mitochondrial 12S rRNA gene, and 3) part of the nuclear histone subunit 3 gene. These genes represent a diverse group of mitochondrial protein- or RNA-coding genes, as well as a nuclear protein-coding gene. They have been previously shown to be informative and have an adequate resolution to solve phylogenetic relationships between and among species and genera within Pinnotheridae (Palacios Theil et al. 2016; Tsang et al. 2018). In addition, they have proved to be obtained with relative ease using the primers indicated in Table 2, which shows also the length of the fragments obtained with each primer combination. PCR products were either sent to be purified and sequenced by Beckman Coulter Genomics (Danvers, MA, USA) or purified using SureClean (Bioline, Taunton, MA, USA), resuspended in water, sequenced with the ABI BigDye terminator mix, sequencing products being run on a 3100 Applied Biosystems (Foster City, CA, USA) automated sequencer." collectionCode="CA, USA" country="United States of America" pageId="15" pageNumber="89" specimenCode="ULLZ 5625, ULLZ 5626" specimenCount="1">
|
|
<taxonomicName id="918D4D1FF655D62D3078C0275F90FCFC" authorityName="Theil & Felder" authorityYear="2020" baseAuthorityName="Rathbun" baseAuthorityYear="1918" box="[813,1017,810,834]" class="Malacostraca" family="Pinnotheridae" genus="Scleroplax" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="98" phylum="Arthropoda" rank="species" species="franciscana" status="comb. nov.">
|
|
<emphasis id="64F9EA8EF655D62D3078C0275F90FCFC" box="[813,1017,810,834]" italics="true" pageId="15" pageNumber="98">Scleroplax franciscana</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicNameLabel id="7FCA57F5F655D62D3755C0265830FCFC" box="[1024,1113,810,834]" pageId="15" pageNumber="98" rank="species">n. comb.</taxonomicNameLabel>
|
|
<emphasis id="64F9EA8EF655D62D370BC026581EFCFC" box="[1118,1143,811,834]" italics="true" pageId="15" pageNumber="98">—</emphasis>
|
|
<specimenCode id="062B9EE7F655D62D372BC0265890FCFC" box="[1150,1273,810,834]" pageId="15" pageNumber="98">ULLZ 5625</specimenCode>
|
|
,
|
|
<specimenCode id="062B9EE7F655D62D3651C0265914FCFC" box="[1284,1405,810,834]" pageId="15" pageNumber="98">ULLZ 5626</specimenCode>
|
|
(Bodega Bay,
|
|
<collectionCode id="309CAE59F655D62D30DBC0485FDBFCE3" box="[910,946,837,861]" pageId="15" pageNumber="98">CA</collectionCode>
|
|
,
|
|
<collectionCode id="309CAE59F655D62D30E9C0485F86FCE3" box="[956,1007,837,861]" pageId="15" pageNumber="98">USA</collectionCode>
|
|
)
|
|
</materialsCitation>
|
|
;
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="1E976517F655D62D3078C052597BFBA8" pageId="15" pageNumber="98" type="description">
|
|
<paragraph id="5632369CF655D62D3078C0525973FC2C" blockId="15.[813,1456,730,1046]" pageId="15" pageNumber="98">
|
|
<materialsCitation id="E6E53CC1F655D62D3078C0525F03FC2C" ID-GBIF-Occurrence="2573304564" collectionCode="WA, USA" country="United States of America" pageId="15" pageNumber="98" specimenCode="ULLZ 8505" specimenCount="1">
|
|
<taxonomicName id="918D4D1FF655D62D3078C0525F88FCC9" authorityName="Theil & Felder" authorityYear="2020" baseAuthorityName="Holmes" baseAuthorityYear="1894" box="[813,993,863,887]" class="Malacostraca" family="Pinnotheridae" genus="Scleroplax" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="98" phylum="Arthropoda" rank="species" species="littoralis" status="comb. nov.">
|
|
<emphasis id="64F9EA8EF655D62D3078C0525F88FCC9" box="[813,993,863,887]" italics="true" pageId="15" pageNumber="98">Scleroplax littoralis</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicNameLabel id="7FCA57F5F655D62D30BDC052582AFCC9" box="[1000,1091,863,887]" pageId="15" pageNumber="98" rank="species">n. comb.</taxonomicNameLabel>
|
|
<emphasis id="64F9EA8EF655D62D371CC06D580BFCC9" box="[1097,1122,864,887]" italics="true" pageId="15" pageNumber="98">—</emphasis>
|
|
<specimenCode id="062B9EE7F655D62D373EC052588CFCC9" box="[1131,1253,863,887]" pageId="15" pageNumber="98">ULLZ 8505</specimenCode>
|
|
(10) (Poulsbo,
|
|
<collectionCode id="309CAE59F655D62D36D6C05259C5FCC9" box="[1411,1452,863,887]" pageId="15" pageNumber="98">WA</collectionCode>
|
|
,
|
|
<collectionCode id="309CAE59F655D62D3078C0775F09FC2C" box="[813,864,890,914]" pageId="15" pageNumber="98">USA</collectionCode>
|
|
),
|
|
</materialsCitation>
|
|
<materialsCitation id="E6E53CC1F655D62D3025C077597EFC2C" ID-GBIF-Occurrence="2573304535" box="[880,1303,889,914]" collectionCode="WA, USA" country="United States of America" pageId="15" pageNumber="98" specimenCode="ULLZ 14072" specimenCount="1">
|
|
<specimenCode id="062B9EE7F655D62D3025C0775F9CFC2C" box="[880,1013,890,914]" pageId="15" pageNumber="98">ULLZ 14072</specimenCode>
|
|
(4) (Gamble Bay,
|
|
<collectionCode id="309CAE59F655D62D37FFC07758BAFC2C" box="[1194,1235,890,914]" pageId="15" pageNumber="98">WA</collectionCode>
|
|
,
|
|
<collectionCode id="309CAE59F655D62D3788C0775979FC2C" box="[1245,1296,890,914]" pageId="15" pageNumber="98">USA</collectionCode>
|
|
)
|
|
</materialsCitation>
|
|
;
|
|
</paragraph>
|
|
<paragraph id="5632369CF655D62D3078C0995940FC5F" blockId="15.[813,1456,730,1046]" pageId="15" pageNumber="98">
|
|
<materialsCitation id="E6E53CC1F655D62D3078C099585AFC78" ID-GBIF-Occurrence="2573304550" collectionCode="AK, USA" country="Iceland" location="Island" pageId="15" pageNumber="98" specimenCode="ULLZ 14036, ULLZ 14842" specimenCount="1">
|
|
<taxonomicName id="918D4D1FF655D62D3078C0995FB6FC12" authorityName="Theil & Felder" authorityYear="2020" baseAuthorityName="Rathbun" baseAuthorityYear="1918" box="[813,991,916,940]" class="Malacostraca" family="Pinnotheridae" genus="Scleroplax" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="98" phylum="Arthropoda" rank="species" species="schmitti" status="comb. nov.">
|
|
<emphasis id="64F9EA8EF655D62D3078C0995FB6FC12" box="[813,991,916,940]" italics="true" pageId="15" pageNumber="98">Scleroplax schmitti</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicNameLabel id="7FCA57F5F655D62D30B2C099582BFC12" box="[999,1090,916,940]" pageId="15" pageNumber="98" rank="species">n. comb.</taxonomicNameLabel>
|
|
<emphasis id="64F9EA8EF655D62D371CC098580AFC12" box="[1097,1123,917,940]" italics="true" pageId="15" pageNumber="98">—</emphasis>
|
|
<specimenCode id="062B9EE7F655D62D373EC099589FFC12" box="[1131,1270,916,940]" pageId="15" pageNumber="98">ULLZ 14036</specimenCode>
|
|
,
|
|
<specimenCode id="062B9EE7F655D62D3657C09959E3FC12" box="[1282,1418,916,940]" pageId="15" pageNumber="98">ULLZ 14842</specimenCode>
|
|
(8) (Baranof
|
|
<collectingCountry id="2E9A760CF655D62D30D0C0A35FABFC78" box="[901,962,942,966]" name="Iceland" pageId="15" pageNumber="98" value="Iceland">Island</collectingCountry>
|
|
,
|
|
<collectionCode id="309CAE59F655D62D309EC0A25F87FC79" box="[971,1006,943,967]" pageId="15" pageNumber="98">AK</collectionCode>
|
|
,
|
|
<collectionCode id="309CAE59F655D62D30A3C0A35840FC78" box="[1014,1065,942,966]" pageId="15" pageNumber="98">USA</collectionCode>
|
|
),
|
|
</materialsCitation>
|
|
<materialsCitation id="E6E53CC1F655D62D376DC0A2594FFC5F" ID-GBIF-Occurrence="2573304565" collectionCode="AK, USA" country="Iceland" location="Island" pageId="15" pageNumber="98" specimenCode="ULLZ 14117, ULLZ 14119" specimenCount="1">
|
|
<specimenCode id="062B9EE7F655D62D376DC0A258D5FC78" box="[1080,1212,942,966]" pageId="15" pageNumber="98">ULLZ 14117</specimenCode>
|
|
, MNHN-IU-2017-9369 = former
|
|
<specimenCode id="062B9EE7F655D62D30DDC0C45864FC5F" box="[904,1037,969,993]" pageId="15" pageNumber="98">ULLZ 14119</specimenCode>
|
|
(Japonski
|
|
<collectingCountry id="2E9A760CF655D62D3723C0C458DDFC5F" box="[1142,1204,969,993]" name="Iceland" pageId="15" pageNumber="98" value="Iceland">Island</collectingCountry>
|
|
,
|
|
<collectionCode id="309CAE59F655D62D37EBC0C4588BFC5F" box="[1214,1250,969,993]" pageId="15" pageNumber="98">AK</collectionCode>
|
|
,
|
|
<collectionCode id="309CAE59F655D62D37B9C0C45976FC5F" box="[1260,1311,969,993]" pageId="15" pageNumber="98">USA</collectionCode>
|
|
)
|
|
</materialsCitation>
|
|
;
|
|
</paragraph>
|
|
<paragraph id="5632369CF655D62D3078C0EE597BFBA8" blockId="15.[813,1456,730,1046]" pageId="15" pageNumber="98">
|
|
<materialsCitation id="E6E53CC1F655D62D3078C0EE5F03FBA8" ID-GBIF-Occurrence="2573304531" collectionCode="AK, USA" country="Iceland" location="Island" pageId="15" pageNumber="98" specimenCode="ULLZ 14116" specimenCount="1">
|
|
<taxonomicName id="918D4D1FF655D62D3078C0EE5FB5FC45" authorityName="Theil & Felder" authorityYear="2020" baseAuthorityName="Holmes" baseAuthorityYear="1894" box="[813,988,995,1019]" class="Malacostraca" family="Pinnotheridae" genus="Scleroplax" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="98" phylum="Arthropoda" rank="species" species="tubicola" status="comb. nov.">
|
|
<emphasis id="64F9EA8EF655D62D3078C0EE5FB5FC45" box="[813,988,995,1019]" italics="true" pageId="15" pageNumber="98">Scleroplax tubicola</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicNameLabel id="7FCA57F5F655D62D30B7C0E95853FC45" box="[994,1082,995,1019]" pageId="15" pageNumber="98" rank="species">n. comb.</taxonomicNameLabel>
|
|
<emphasis id="64F9EA8EF655D62D376AC0E95831FC45" box="[1087,1112,996,1019]" italics="true" pageId="15" pageNumber="98">—</emphasis>
|
|
<specimenCode id="062B9EE7F655D62D370AC0E9588DFC45" box="[1119,1252,995,1019]" pageId="15" pageNumber="98">ULLZ 14116</specimenCode>
|
|
(Middle
|
|
<collectingCountry id="2E9A760CF655D62D3615C0EE5917FC45" box="[1344,1406,995,1019]" name="Iceland" pageId="15" pageNumber="98" value="Iceland">Island</collectingCountry>
|
|
,
|
|
<collectionCode id="309CAE59F655D62D36DDC0EE59C5FC45" box="[1416,1452,995,1019]" pageId="15" pageNumber="98">AK</collectionCode>
|
|
,
|
|
<collectionCode id="309CAE59F655D62D3078C0F35F09FBA8" box="[813,864,1022,1046]" pageId="15" pageNumber="98">USA</collectionCode>
|
|
),
|
|
</materialsCitation>
|
|
<materialsCitation id="E6E53CC1F655D62D3025C0F35967FBA8" ID-GBIF-Occurrence="2573304542" box="[880,1294,1022,1046]" collectionCode="AK, USA" country="Iceland" location="Island" pageId="15" pageNumber="98" specimenCode="ULLZ 14118" specimenCount="1">
|
|
<specimenCode id="062B9EE7F655D62D3025C0F35F9CFBA8" box="[880,1013,1022,1046]" pageId="15" pageNumber="98">ULLZ 14118</specimenCode>
|
|
(Japonski
|
|
<collectingCountry id="2E9A760CF655D62D370BC0F358F5FBA8" box="[1118,1180,1022,1046]" name="Iceland" pageId="15" pageNumber="98" value="Iceland">Island</collectingCountry>
|
|
,
|
|
<collectionCode id="309CAE59F655D62D37F3C0F358A3FBA8" box="[1190,1226,1022,1046]" pageId="15" pageNumber="98">AK</collectionCode>
|
|
,
|
|
<collectionCode id="309CAE59F655D62D3781C0F35961FBA8" box="[1236,1288,1022,1046]" pageId="15" pageNumber="98">USA</collectionCode>
|
|
)
|
|
</materialsCitation>
|
|
.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="1E976517F655D62D3078C758585FF853" pageId="15" pageNumber="98" type="discussion">
|
|
<paragraph id="5632369CF655D62D3078C7585FF8FBD0" blockId="15.[813,1456,1109,2030]" box="[813,913,1109,1135]" pageId="15" pageNumber="98">REMARKS</paragraph>
|
|
<paragraph id="5632369CF655D62D3078C77B5875F910" blockId="15.[813,1456,1109,2030]" pageId="15" pageNumber="98">
|
|
Genetic distances and the morphological differences observed among some of the species in this group are similar to those shown among conspecific populations in other pinnotherid genera, for instance
|
|
<taxonomicName id="918D4D1FF655D62D3751C7DB5800FB4E" authorityName="Heard & Manning" authorityYear="1997" box="[1028,1129,1238,1264]" class="Malacostraca" family="Pinnotheridae" genus="Austinixa" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="98" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="64F9EA8EF655D62D3751C7DB5800FB4E" box="[1028,1129,1238,1264]" italics="true" pageId="15" pageNumber="98">Austinixa</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="918D4D1FF655D62D3720C7DB59C2FB4E" authority="Campos, 1989" authorityName="Campos" authorityYear="1989" box="[1141,1451,1238,1264]" class="Malacostraca" family="Pinnotheridae" genus="Tumidotheres" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="98" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="64F9EA8EF655D62D3720C7DB596BFB4E" box="[1141,1282,1238,1264]" italics="true" pageId="15" pageNumber="98">Tumidotheres</emphasis>
|
|
Campos, 1989
|
|
</taxonomicName>
|
|
, or
|
|
<taxonomicName id="918D4D1FF655D62D301FC7F85813FAB1" authority="Campos, 1996" authorityName="Campos" authorityYear="1996" box="[842,1146,1269,1296]" class="Malacostraca" family="Pinnotheridae" genus="Tunicotheres" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="98" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="64F9EA8EF655D62D301FC7F85FA7FAB1" box="[842,974,1269,1295]" italics="true" pageId="15" pageNumber="98">Tunicotheres</emphasis>
|
|
Campos, 1996
|
|
</taxonomicName>
|
|
. Furthermore, for some species there seems to be striking variation in key characters between juveniles and adults. For example, juveniles of
|
|
<taxonomicName id="918D4D1FF655D62D3078C65B5FC9FAD1" authorityName="Theil & Felder" authorityYear="2020" baseAuthorityName="Holmes" baseAuthorityYear="1894" box="[813,928,1365,1391]" class="Malacostraca" family="Pinnotheridae" genus="Scleroplax" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="98" phylum="Arthropoda" rank="species" species="littoralis">
|
|
<emphasis id="64F9EA8EF655D62D3078C65B5FC9FAD1" box="[813,928,1365,1391]" italics="true" pageId="15" pageNumber="98">P. littoralis</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="918D4D1FF655D62D308EC65B584CFAD1" authorityName="Theil & Felder" authorityYear="2020" baseAuthorityName="Dana" baseAuthorityYear="1851" box="[987,1061,1365,1391]" class="Malacostraca" family="Pinnotheridae" genus="Scleroplax" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="98" phylum="Arthropoda" rank="species" species="faba">
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<emphasis id="64F9EA8EF655D62D308EC65B584CFAD1" box="[987,1061,1365,1391]" italics="true" pageId="15" pageNumber="98">P. faba</emphasis>
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</taxonomicName>
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appear to be extremely difficult to discriminate (
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<bibRefCitation id="321C4B6DF655D62D309FC6785802FA31" author="Zmarzly" box="[970,1131,1397,1423]" firstAuthor="Zmarzly" pageId="15" pageNumber="98" pagination="677 - 713" refId="ref19522" refString="ZMARZLY D. L. 1992. - Taxonomic review of pea crabs in the genus Pinnixa (Decapoda: Brachyura: Pinnotheridae) occurring on the California shelf, with descriptions of two new species. Journal of Crustacean Biology 12 (4): 677 - 713. https: // doi. org / 10.1163 / 193724092 X 00166" type="journal article" year="1992">Zmarzly 1992</bibRefCitation>
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). A more detailed investigation with larger sample sizes and markers appropriate to determine variability between populations of these species is required to clarify phylogenetic relationships within and among them. In addition, knowledge of host associations is required to accompany samples, as these taxa might represent species complexes of separate, but morphologically similar, populations that have adapted to different hosts, which may also be reflected in variations between inshore and offshore samples.
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</paragraph>
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<paragraph id="5632369CF655D62D3016C5B9585FF853" blockId="15.[813,1456,1109,2030]" pageId="15" pageNumber="98">
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The only specimen of
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<taxonomicName id="918D4D1FF655D62D377AC5B858A0F970" authorityName="Martin & Zmarzly" authorityYear="1994" box="[1071,1225,1717,1742]" class="Malacostraca" family="Pinnotheridae" genus="Pinnixa" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="98" phylum="Arthropoda" rank="species" species="scamit">
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<emphasis id="64F9EA8EF655D62D377AC5B858A0F970" box="[1071,1225,1717,1742]" italics="true" pageId="15" pageNumber="98">Pinnixa scamit</emphasis>
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</taxonomicName>
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available for molecular analyses and morphological examination was a juvenile (UF 11969), and had therefore been identified provisionally. Genetically it was closely allied to
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<taxonomicName id="918D4D1FF655D62D37CBC419596FF890" authorityName="Rathbun" authorityYear="1894" box="[1182,1286,1812,1838]" class="Malacostraca" family="Pinnotheridae" genus="Scleroplax" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="98" phylum="Arthropoda" rank="genus">
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<emphasis id="64F9EA8EF655D62D37CBC419596FF890" box="[1182,1286,1812,1838]" italics="true" pageId="15" pageNumber="98">Scleroplax</emphasis>
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</taxonomicName>
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, however, morphologically it showed characters similar to those in
|
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<taxonomicName id="918D4D1FF655D62D360CC43E5F32F8D3" authority="Theil & Felder, 2020" authorityName="Theil & Felder" authorityYear="2020" class="Malacostraca" family="Pinnotheridae" genus="Rathbunixa" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="98" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="64F9EA8EF655D62D360CC43E5F32F8D3" italics="true" pageId="15" pageNumber="98">Rathbunixa</emphasis>
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</taxonomicName>
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|
<taxonomicNameLabel id="7FCA57F5F655D62D3034C4595FC5F8D0" box="[865,940,1876,1902]" pageId="15" pageNumber="98" rank="genus">n. gen.</taxonomicNameLabel>
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It had long slender legs, somewhat compressed, and a sculpted carapace. Despite the results of the molecular analysis, we choose not to transfer
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<taxonomicName id="918D4D1FF655D62D37C6C4995945F813" authorityName="Martin & Zmarzly" authorityYear="1994" box="[1171,1324,1940,1965]" class="Malacostraca" family="Pinnotheridae" genus="Pinnixa" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="98" phylum="Arthropoda" rank="species" species="scamit">
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<emphasis id="64F9EA8EF655D62D37C6C4995945F813" box="[1171,1324,1940,1965]" italics="true" pageId="15" pageNumber="98">Pinnixa scamit</emphasis>
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</taxonomicName>
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to the genus
|
|
<taxonomicName id="918D4D1FF655D62D3078C4BE5FFAF873" authorityName="Rathbun" authorityYear="1894" box="[813,915,1971,1997]" class="Malacostraca" family="Pinnotheridae" genus="Scleroplax" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="98" phylum="Arthropoda" rank="genus">
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<emphasis id="64F9EA8EF655D62D3078C4BE5FFAF873" box="[813,915,1971,1997]" italics="true" pageId="15" pageNumber="98">Scleroplax</emphasis>
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</taxonomicName>
|
|
, until specimens definitively identifiable as
|
|
<taxonomicName id="918D4D1FF655D62D360CC4B959C6F873" authorityName="Martin & Zmarzly" authorityYear="1994" box="[1369,1455,1972,1997]" class="Malacostraca" family="Pinnotheridae" genus="Pinnixa" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="98" phylum="Arthropoda" rank="species" species="scamit">
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<emphasis id="64F9EA8EF655D62D360CC4B959C6F873" box="[1369,1455,1972,1997]" italics="true" pageId="15" pageNumber="98">P. scamit</emphasis>
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|
</taxonomicName>
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are available for analysis.
|
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |