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<mods:title id="43ED6CCDC2ED9D7D2113991441B0C704">A taxonomic review of the Australian genus Eurylychnus Bates (Coleoptera: Carabidae: Broscini)</mods:title>
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<subSubSection id="42AA655EFB6EFFDEFF3BFF790DB8FF6F" box="[151,455,223,250]" pageId="1" pageNumber="248" type="nomenclature">
<paragraph id="0A0F36D5FB6EFFDEFF3BFF790DB8FF6F" blockId="1.[151,455,223,285]" box="[151,455,223,250]" pageId="1" pageNumber="248">
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<taxonomicName id="CDB04D56FB6EFFDEFF3BFF790DB8FF6F" ID-CoL="8KWXM" authority="Bates, 1891" authorityName="Bates" authorityYear="1891" box="[151,455,223,250]" class="Insecta" family="Carabidae" genus="Eurylychnus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="248" phylum="Arthropoda" rank="genus">
<emphasis id="38C4EAC7FB6EFFDEFF3BFF790DB8FF6F" bold="true" box="[151,455,223,250]" pageId="1" pageNumber="248">
<emphasis id="38C4EAC7FB6EFFDEFF3BFF790D4DFF6C" bold="true" box="[151,306,223,250]" italics="true" pageId="1" pageNumber="248">Eurylychnus</emphasis>
<bibRefCitation id="6E214B24FB6EFFDEFE95FF790DB8FF6F" author="Bates, H. W." box="[313,455,223,249]" pageId="1" pageNumber="248" pagination="285 - 287" refId="ref13266" refString="Bates, H. W. (1891) Note on three Australian Carabidae. Entomologists Monthly Magazine, 2 (2), 285 - 287." type="journal article" year="1891">Bates, 1891</bibRefCitation>
</emphasis>
</taxonomicName>
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<subSubSection id="42AA655EFB6EFFDEFF3BFEA5089AFE0B" pageId="1" pageNumber="248" type="description">
<paragraph id="0A0F36D5FB6EFFDEFF3BFEA50D6DFE8B" blockId="1.[151,455,223,285]" box="[151,274,259,285]" pageId="1" pageNumber="248">
(
<figureCitation id="928B2A50FB6EFFDEFF33FEA50D76FE8B" box="[159,265,259,285]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStart-6="FIGURE 7" captionStartId-0="3.[152,255,1888,1912]" captionStartId-1="4.[152,255,1943,1967]" captionStartId-2="8.[152,255,1943,1967]" captionStartId-3="9.[152,255,988,1012]" captionStartId-4="10.[152,255,1886,1910]" captionStartId-5="11.[152,255,1081,1105]" captionStartId-6="16.[152,255,688,712]" captionTargetBox-0="[163,1424,345,1852]" captionTargetBox-1="[304,1283,185,1915]" captionTargetBox-2="[298,1290,186,1915]" captionTargetBox-3="[154,1433,184,961]" captionTargetBox-4="[154,1433,1068,1858]" captionTargetBox-5="[151,1435,181,1057]" captionTargetBox-6="[151,1435,181,664]" captionTargetId-0="figure-96@3.[151,1436,345,1864]" captionTargetId-1="figure-25@4.[299,1287,181,1919]" captionTargetId-2="figure-25@8.[294,1294,181,1919]" captionTargetId-3="figure-15@9.[151,1435,181,964]" captionTargetId-4="figure-426@10.[151,1435,1065,1862]" captionTargetId-5="figure-15@11.[151,1435,181,1057]" captionTargetId-6="figure-25@16.[151,1435,181,664]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="8" captionTargetPageId-3="9" captionTargetPageId-4="10" captionTargetPageId-5="11" captionTargetPageId-6="16" captionText-0="FIGURE 1. Ventral surface of male protarsi of A, Eurylychnus dyschirioides; B, Eurylychnus cylindricus." captionText-1="FIGURE 2. Dorsal habitus of A, Eurylychnus blagravei; B, Eurylychnus cylindricus; C, Eurylychnus regularis; D, Eurylychnus victoriae. Scalebars are 2mm." captionText-2="FIGURE 3. Dorsal habitus of A, Eurylychnus dyschirioides; B, Eurylychnus ovipennis; C, Eurylychnus kershawi; D, Eurylychnus femoralis. Scalebars are 2mm." captionText-3="FIGURE 4. Male genitalia of Eurylychnus blagravei (A–C): A, left paramere, B, right paramere, C, median lobe left lateral; E. cylindricus (D–F): D, left paramere, E, right paramere, F, median lobe left lateral; E. regularis (G–I): G, left paramere, H, right paramere, I, median lobe left lateral; E. victoriae (J–L): J, left paramere, K, right paramere, L, median lobe left lateral. Scale bars are 0.5mm." captionText-4="FIGURE 5. Male genitalia of Eurylychnus dyschirioides (A–C):A, left paramere, B, right paramere, C, median lobe left lateral; E. ovipennis (D–F): D, left paramere, E, right paramere, F, median lobe left lateral; E. kershawi (G–I): G, left paramere, H, right paramere, I, median lobe left lateral; E. femoralis (J–L): J, left paramere, K, right paramere, L, median lobe left lateral. Scale bars are 0.5mm." captionText-5="FIGURE 6. Pronotum of Eurylychnus dyschirioides. PC = pronotal carina; PS = pronotal setae; MI = median impression; PF = pronotal fovea; PT = pronotal tubercle. Scalebar is 1mm." captionText-6="FIGURE 7. A, Eurylychnus blagravei, with parascuttelar striole isolated from striae 1; B, Eurylychnus victoriae, with parascuttelar striole joined to apical portion of striae 1." figureDoi-0="http://doi.org/10.5281/zenodo.8249354" figureDoi-1="http://doi.org/10.5281/zenodo.8249356" figureDoi-2="http://doi.org/10.5281/zenodo.8249358" figureDoi-3="http://doi.org/10.5281/zenodo.8249360" figureDoi-4="http://doi.org/10.5281/zenodo.8249362" figureDoi-5="http://doi.org/10.5281/zenodo.8249364" figureDoi-6="http://doi.org/10.5281/zenodo.8249366" httpUri-0="https://zenodo.org/record/8249354/files/figure.png" httpUri-1="https://zenodo.org/record/8249356/files/figure.png" httpUri-2="https://zenodo.org/record/8249358/files/figure.png" httpUri-3="https://zenodo.org/record/8249360/files/figure.png" httpUri-4="https://zenodo.org/record/8249362/files/figure.png" httpUri-5="https://zenodo.org/record/8249364/files/figure.png" httpUri-6="https://zenodo.org/record/8249366/files/figure.png" pageId="1" pageNumber="248">Figs. 1–7</figureCitation>
)
</paragraph>
<paragraph id="0A0F36D5FB6EFFDEFF3BFEEE08DAFEE8" blockId="1.[151,1437,327,413]" pageId="1" pageNumber="248">
<bibRefCitation id="6E214B24FB6EFFDEFF3BFEEE0D35FEC9" author="Bates, H. W." box="[151,330,327,352]" pageId="1" pageNumber="248" pagination="285 - 287" refId="ref13266" refString="Bates, H. W. (1891) Note on three Australian Carabidae. Entomologists Monthly Magazine, 2 (2), 285 - 287." type="journal article" year="1891">Bates 1891: 285</bibRefCitation>
.
<bibRefCitation id="6E214B24FB6EFFDEFEF5FEEE0E77FEC9" author="Sloane, T. G." box="[345,520,327,352]" pageId="1" pageNumber="248" pagination="45 - 64" refId="ref13969" refString="Sloane, T. G. (1892) Studies in Australian Entomology. No. V. Notes on the subfamily Broscini (Carabidae), with descriptions of new species. Proceedings of the Linnean Society of New South Wales, 7 (2), 45 - 64. https: // doi. org / 10.5962 / bhl. part. 26042" type="journal article" year="1892">Sloane 1892: 50</bibRefCitation>
(key to species).
<bibRefCitation id="6E214B24FB6EFFDEFD63FEEE087FFEF6" author="Moore, B. P. &amp; Weir, T. A. &amp; Pyke, J. E." box="[719,1024,328,352]" pageId="1" pageNumber="248" refId="ref13723" refString="Moore, B. P., Weir, T. A. &amp; Pyke, J. E. (1987) Zoological catalogue of Australia. Vol. 4. Coleoptera: Rhysodidae and Carabidae. Australian Government Publishing Service, Canberra, 444 pp." type="book" year="1987">
Moore
<emphasis id="38C4EAC7FB6EFFDEFCB3FEEE0F28FEF6" box="[799,855,328,352]" italics="true" pageId="1" pageNumber="248">et al.</emphasis>
1987: 120–121
</bibRefCitation>
(catalogue).
<bibRefCitation id="6E214B24FB6EFFDEFB3CFEEE09E2FEC9" author="Roig-Junent, S." box="[1168,1437,327,352]" pageId="1" pageNumber="248" refId="ref13836" refString="Roig-Junent, S. (2000) The subtribes and genera of the tribe Broscini (Coleoptera: Carabidae): cladistic analysis, taxonomic treatment, and biogeographical considerations. Bulletin of the American Museum of Natural History, 255, 01 - 90. https: // doi. org / 10.1206 / 0003 - 0090 (2000) 255 &lt;0001: TSAGOT&gt; 2.0. CO; 2" type="book" year="2000">Roig-Juñent 2000: 36–38</bibRefCitation>
(description).
<bibRefCitation id="6E214B24FB6EFFDEFEF9FEC00E24FEE8" author="Hackel, M. &amp; Farkac, J. &amp; Wrase, D. W." box="[341,603,358,382]" pageId="1" pageNumber="248" pagination="43 - 84" refId="ref13364" refString="Hackel, M., Farkac, J. &amp; Wrase, D. W. (2010) A check-list of the tribe Broscini Hope, 1838 of the World (Coleoptera: Carabidae). Studies and Reports, Taxonomical Series, 6 (1 - 2), 43 - 84." type="journal article" year="2010">
Häckel
<emphasis id="38C4EAC7FB6EFFDEFE0FFEC10DA8FEE8" box="[419,471,358,382]" italics="true" pageId="1" pageNumber="248">et al.</emphasis>
2010: 65–66
</bibRefCitation>
(catalogue).
<bibRefCitation id="6E214B24FB6EFFDEFD4DFEC00F8DFEE8" author="Baehr, M. &amp; Will, K." box="[737,1010,358,382]" pageId="1" pageNumber="248" pagination="1802" refId="ref13189" refString="Baehr, M. &amp; Will, K. (2019) 12. Carabidae Latreille, 1802. In: Slipinski, A. &amp; Lawrence, J. (Eds.), Australian Beetles. Vol. 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, Victoria, pp. 61 - 217. https: // doi. org / 10.1071 / 9780643097315" type="book chapter" year="2019">Baehr &amp; Will 2019: 90–91</bibRefCitation>
; 93 (description).
</paragraph>
<paragraph id="0A0F36D5FB6EFFDEFF3BFE23089AFE0B" blockId="1.[151,1437,327,413]" box="[151,1253,389,413]" pageId="1" pageNumber="248">
<typeStatus id="D50B8877FB6EFFDEFF3BFE230CB6FE0B" box="[151,201,389,413]" pageId="1" pageNumber="248">Type</typeStatus>
species, by monotypy:
<taxonomicName id="CDB04D56FB6EFFDEFE15FE230E8BFE0B" authority="Bates, 1891" authorityName="Bates" authorityYear="1891" box="[441,756,389,413]" class="Insecta" family="Carabidae" genus="Eurylychnus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="248" phylum="Arthropoda" rank="species" species="olliffi">
<emphasis id="38C4EAC7FB6EFFDEFE15FE230E0AFE0B" box="[441,629,389,413]" italics="true" pageId="1" pageNumber="248">Eurylychnus olliffi</emphasis>
<bibRefCitation id="6E214B24FB6EFFDEFDD0FE230E8BFE0B" author="Bates, H. W." box="[636,756,389,413]" pageId="1" pageNumber="248" pagination="285 - 287" refId="ref13266" refString="Bates, H. W. (1891) Note on three Australian Carabidae. Entomologists Monthly Magazine, 2 (2), 285 - 287." type="journal article" year="1891">Bates, 1891</bibRefCitation>
</taxonomicName>
[=
<taxonomicName id="CDB04D56FB6EFFDEFCBBFE23089FFE0B" authority="(Castelnau, 1867)" baseAuthorityName="Castelnau" baseAuthorityYear="1867" box="[791,1248,389,413]" class="Insecta" family="Carabidae" genus="Eurylychnus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="248" phylum="Arthropoda" rank="species" species="dyschirioides">
<emphasis id="38C4EAC7FB6EFFDEFCBBFE23085CFE0B" box="[791,1059,389,413]" italics="true" pageId="1" pageNumber="248">Eurylychnus dyschirioides</emphasis>
(
<bibRefCitation id="6E214B24FB6EFFDEFB9CFE2308A7FE0B" author="Castelnau, F. L." box="[1072,1240,389,413]" pageId="1" pageNumber="248" refId="ref13340" refString="Castelnau, F. L. (1867) Notes on Australian Coleoptera. Royal Society of Victoria, Melbourne, 139 pp." type="book" year="1867">Castelnau, 1867</bibRefCitation>
)
</taxonomicName>
]
</paragraph>
</subSubSection>
<subSubSection id="42AA655EFB6EFFDEFF3BFE6D0863FD28" pageId="1" pageNumber="248" type="diagnosis">
<paragraph id="0A0F36D5FB6EFFDEFF3BFE6D0863FD28" blockId="1.[151,1437,459,1928]" pageId="1" pageNumber="248">
<emphasis id="38C4EAC7FB6EFFDEFF3BFE6D0D75FE73" bold="true" box="[151,266,459,485]" pageId="1" pageNumber="248">Diagnosis</emphasis>
:
<taxonomicName id="CDB04D56FB6EFFDEFEB7FE6D0DD8FE73" authorityName="Bates" authorityYear="1891" box="[283,423,459,485]" class="Insecta" family="Carabidae" genus="Eurylychnus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="248" phylum="Arthropoda" rank="genus">
<emphasis id="38C4EAC7FB6EFFDEFEB7FE6D0DD8FE73" box="[283,423,459,485]" italics="true" pageId="1" pageNumber="248">Eurylychnus</emphasis>
</taxonomicName>
can be separated from all other genera of Australian
<taxonomicName id="CDB04D56FB6EFFDEFBA1FE6D08CDFE73" authorityName="Roig-Junent" authorityYear="2000" box="[1037,1202,459,485]" pageId="1" pageNumber="247" rank="subTribe" subTribe="Nothobroscina">Nothobroscina</taxonomicName>
by having all of the following characters: a seta in the scrobe of the mandibles, a transverse impression behind the eyes and a single supraorbital seta above each eye.
<taxonomicName id="CDB04D56FB6EFFDEFDA7FDB50E18FDBB" authorityName="Sloane" authorityYear="1920" box="[523,615,531,557]" class="Insecta" family="Carabidae" genus="Chylnus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="248" phylum="Arthropoda" rank="genus">
<emphasis id="38C4EAC7FB6EFFDEFDA7FDB50E18FDBB" box="[523,615,531,557]" italics="true" pageId="1" pageNumber="248">Chylnus</emphasis>
</taxonomicName>
resembles
<taxonomicName id="CDB04D56FB6EFFDEFD48FDB50F0FFDBB" authorityName="Bates" authorityYear="1891" box="[740,880,531,557]" class="Insecta" family="Carabidae" genus="Eurylychnus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="248" phylum="Arthropoda" rank="genus">
<emphasis id="38C4EAC7FB6EFFDEFD48FDB50F0FFDBB" box="[740,880,531,557]" italics="true" pageId="1" pageNumber="248">Eurylychnus</emphasis>
</taxonomicName>
in having a transverse impression behind the eyes and a single supraorbital seta above each eye, but
<taxonomicName id="CDB04D56FB6EFFDEFD7EFD910F51FDC7" authorityName="Sloane" authorityYear="1920" box="[722,814,567,593]" class="Insecta" family="Carabidae" genus="Chylnus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="248" phylum="Arthropoda" rank="genus">
<emphasis id="38C4EAC7FB6EFFDEFD7EFD910F51FDC7" box="[722,814,567,593]" italics="true" pageId="1" pageNumber="248">Chylnus</emphasis>
</taxonomicName>
lack a seta in the scrobe of the mandibles.
<taxonomicName id="CDB04D56FB6EFFDEFAB7FD910D1EFDE3" authority="Sloane" authorityName="Sloane" authorityYear="1892" class="Insecta" family="Carabidae" genus="Percolestus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="248" phylum="Arthropoda" rank="species" species="blackburni">
<emphasis id="38C4EAC7FB6EFFDEFAB7FD910D6FFDE3" italics="true" pageId="1" pageNumber="248">Percolestus blackburni</emphasis>
Sloane
</taxonomicName>
is sympatric with
<taxonomicName id="CDB04D56FB6EFFDEFD89FDFA0F5CFDE3" authority="Castelnau" authorityName="Castelnau" box="[549,803,603,629]" class="Insecta" family="Carabidae" genus="Eurylychnus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="248" phylum="Arthropoda" rank="species" species="blagravei">
<emphasis id="38C4EAC7FB6EFFDEFD89FDFA0ED2FDE3" box="[549,685,603,629]" italics="true" pageId="1" pageNumber="248">E. blagravei</emphasis>
Castelnau
</taxonomicName>
, however
<taxonomicName id="CDB04D56FB6EFFDEFC3CFDFA085DFDE3" authorityName="Sloane" authorityYear="1892" box="[912,1058,603,629]" class="Insecta" family="Carabidae" genus="Percolestus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="248" phylum="Arthropoda" rank="species" species="blackburni">
<emphasis id="38C4EAC7FB6EFFDEFC3CFDFA085DFDE3" box="[912,1058,603,629]" italics="true" pageId="1" pageNumber="248">P. blackburni</emphasis>
</taxonomicName>
lacks both a transverse impression behind the eyes and a seta in the scrobe of the mandibles.
<taxonomicName id="CDB04D56FB6EFFDEFCBEFD260FF0FD0F" authorityName="Schaum" authorityYear="1858" box="[786,911,640,665]" class="Insecta" family="Carabidae" genus="Percosoma" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="248" phylum="Arthropoda" rank="genus">
<emphasis id="38C4EAC7FB6EFFDEFCBEFD260FF0FD0F" box="[786,911,640,665]" italics="true" pageId="1" pageNumber="248">Percosoma</emphasis>
</taxonomicName>
lack the transverse impression behind the eyes, have no seta in the scrobe of the mandibles and have multiple supraorbital setae.
</paragraph>
</subSubSection>
<subSubSection id="42AA655EFB6EFFDEFF6BFD610861FBFB" pageId="1" pageNumber="248" type="description">
<paragraph id="0A0F36D5FB6EFFDEFF6BFD610861FBFB" blockId="1.[151,1437,459,1928]" pageId="1" pageNumber="248">
<emphasis id="38C4EAC7FB6EFFDEFF6BFD610D24FD77" bold="true" box="[199,347,711,737]" pageId="1" pageNumber="248">Description:</emphasis>
Body length
<quantity id="CD489B30FB6EFFDEFE57FD610EF2FD77" box="[507,653,711,737]" metricMagnitude="-2" metricUnit="m" metricValue="1.66" metricValueMax="2.33" metricValueMin="0.99" pageId="1" pageNumber="248" unit="mm" value="16.6" valueMax="23.3" valueMin="9.9">9.9–23.3mm</quantity>
. Body black, legs red to black, mouthparts and antennae piceous to black. Head with a transverse impression behind the eyes; frontal impression begins at the eyes, curving inwards, joining the frontoclypeal suture.Antennae with antennomeres 1 and 2 glabrous, 3 with an apical ring of setae, 4 with pubescence from apical half, all remaining antennomeres pubescent. Eyes with a single supraorbital seta. Mandibles with a seta in the scrobe. Mentum with paramedian foveae; tooth bifid. Submentum with 2–6 setae. Pronotum with 1–5 setae on lateral margins, with posterior angles constricted, rounded or straight. Males with or without squamose setae under protarsi 1–3. Elytra with parascutellar striole isolated or joined to the apical portion of stria 1, with setae present only on the lateral margins in the umbilical series. Abdominal ventrites 3–5 with or without paramedian setae. Hind wings absent. Aedeagus left paramere broad, asetose, with a denticle at the apex; right paramere longer and narrower than the left paramere, with setae on apical half of the ventral margin, sometimes extending onto the dorsal margin and middle of the paramere at the apex. Aedeagus median lobe weakly to strongly curved ventrally, deflected to the right; apex rounded or pointed, with or without a tooth ventrally.
</paragraph>
</subSubSection>
<subSubSection id="42AA655EFB6EFFDEFF6BFBD10FC0F813" pageId="1" pageNumber="248" type="discussion">
<paragraph id="0A0F36D5FB6EFFDEFF6BFBD10F30F9D7" blockId="1.[151,1437,459,1928]" pageId="1" pageNumber="248">
<emphasis id="38C4EAC7FB6EFFDEFF6BFBD10D42FB07" bold="true" box="[199,317,1143,1169]" pageId="1" pageNumber="248">Remarks:</emphasis>
In a cladistic analysis by
<bibRefCitation id="6E214B24FB6EFFDEFDF5FBD10F4AFB07" author="Roig-Junent, S." box="[601,821,1143,1169]" pageId="1" pageNumber="248" refId="ref13836" refString="Roig-Junent, S. (2000) The subtribes and genera of the tribe Broscini (Coleoptera: Carabidae): cladistic analysis, taxonomic treatment, and biogeographical considerations. Bulletin of the American Museum of Natural History, 255, 01 - 90. https: // doi. org / 10.1206 / 0003 - 0090 (2000) 255 &lt;0001: TSAGOT&gt; 2.0. CO; 2" type="book" year="2000">Roig-Juñent (2000)</bibRefCitation>
, 2 species groups were identified based on morphology (group 1:
<taxonomicName id="CDB04D56FB6EFFDEFEA8FB3A0E7BFB23" authority="Castelnau" authorityName="Castelnau" box="[260,516,1179,1205]" class="Insecta" family="Carabidae" genus="Eurylychnus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="248" phylum="Arthropoda" rank="species" species="blagravei">
<emphasis id="38C4EAC7FB6EFFDEFEA8FB3A0DF2FB23" box="[260,397,1179,1205]" italics="true" pageId="1" pageNumber="248">E. blagravei</emphasis>
Castelnau
</taxonomicName>
,
<taxonomicName id="CDB04D56FB6EFFDEFDA3FB3A0E81FB23" authority="Sloane" authorityName="Sl. Id. By T. G. Sloane" authorityYear="1916" box="[527,766,1179,1205]" class="Insecta" family="Carabidae" genus="Eurylychnus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="248" phylum="Arthropoda" rank="species" species="cylindricus">
<emphasis id="38C4EAC7FB6EFFDEFDA3FB3A0ED6FB23" box="[527,681,1179,1205]" italics="true" pageId="1" pageNumber="248">E. cylindricus</emphasis>
Sloane
</taxonomicName>
,
<taxonomicName id="CDB04D56FB6EFFDEFCA4FB3A0F9DFB23" authority="Sloane" authorityName="Sloane" box="[776,994,1179,1205]" class="Insecta" family="Carabidae" genus="Eurylychnus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="248" phylum="Arthropoda" rank="species" species="regularis">
<emphasis id="38C4EAC7FB6EFFDEFCA4FB3A0FF3FB23" box="[776,908,1179,1205]" italics="true" pageId="1" pageNumber="248">E. regularis</emphasis>
Sloane
</taxonomicName>
; group 2:
<taxonomicName id="CDB04D56FB6EFFDEFBE3FB3A0905FB23" authority="Castelnau" authorityName="Castelnau" box="[1103,1402,1179,1205]" class="Insecta" family="Carabidae" genus="Eurylychnus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="248" phylum="Arthropoda" rank="species" species="dyschirioides">
<emphasis id="38C4EAC7FB6EFFDEFBE3FB3A097DFB23" box="[1103,1282,1179,1205]" italics="true" pageId="1" pageNumber="248">E. dyschirioides</emphasis>
Castelnau
</taxonomicName>
,
<taxonomicName id="CDB04D56FB6EFFDEFA28FB3A0D29FB4F" authority="Sloane" authorityName="Sloane" authorityYear="1915" class="Insecta" family="Carabidae" genus="Eurylychnus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="248" phylum="Arthropoda" rank="species" species="femoralis">
<emphasis id="38C4EAC7FB6EFFDEFA28FB3A0D7EFB4F" italics="true" pageId="1" pageNumber="248">E. femoralis</emphasis>
Sloane
</taxonomicName>
,
<taxonomicName id="CDB04D56FB6EFFDEFECCFB660E40FB4F" authority="Sloane" authorityName="Sloane" box="[352,575,1215,1241]" class="Insecta" family="Carabidae" genus="Eurylychnus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="248" phylum="Arthropoda" rank="species" species="ovipennis">
<emphasis id="38C4EAC7FB6EFFDEFECCFB660D95FB4F" box="[352,490,1216,1241]" italics="true" pageId="1" pageNumber="248">E. ovipennis</emphasis>
Sloane
</taxonomicName>
,
<taxonomicName id="CDB04D56FB6EFFDEFDE6FB660F60FB4F" authority="Sloane" authorityName="Sloane" authorityYear="1892" box="[586,799,1215,1241]" class="Insecta" family="Carabidae" genus="Eurylychnus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="248" phylum="Arthropoda" rank="species" species="victoriae">
<emphasis id="38C4EAC7FB6EFFDEFDE6FB660EB3FB4F" box="[586,716,1216,1241]" italics="true" pageId="1" pageNumber="248">E. victoriae</emphasis>
Sloane
</taxonomicName>
) and it was found that the two groups were monophyletic and most closely related to
<taxonomicName id="CDB04D56FB6EFFDEFE67FB450F36FB6B" authority="Roig-Junent &amp; Ball" authorityName="Roig-Junent &amp; Ball" authorityYear="1995" box="[459,841,1251,1277]" class="Insecta" family="Carabidae" genus="Nothobroscus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="248" phylum="Arthropoda" rank="genus">
<emphasis id="38C4EAC7FB6EFFDEFE67FB450E19FB6B" box="[459,614,1251,1277]" italics="true" pageId="1" pageNumber="248">Nothobroscus</emphasis>
Roig-Juñent &amp; Ball
</taxonomicName>
from
<collectingCountry id="72A77645FB6EFFDEFC21FB450FB2FB6B" box="[909,973,1251,1277]" name="Chile" pageId="1" pageNumber="248">Chile</collectingCountry>
. However, not all species of
<taxonomicName id="CDB04D56FB6EFFDEFABCFB4509E3FB6B" authorityName="Bates" authorityYear="1891" box="[1296,1436,1251,1277]" class="Insecta" family="Carabidae" genus="Eurylychnus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="248" phylum="Arthropoda" rank="genus">
<emphasis id="38C4EAC7FB6EFFDEFABCFB4509E3FB6B" box="[1296,1436,1251,1277]" italics="true" pageId="1" pageNumber="248">Eurylychnus</emphasis>
</taxonomicName>
were examined, and the present study has found some of the characters used to be tenuous. In a key to
<taxonomicName id="CDB04D56FB6EFFDEFB5BFAA109E3FAB7" authorityName="Roig-Junent" authorityYear="2000" box="[1271,1436,1287,1313]" pageId="1" pageNumber="247" rank="subTribe" subTribe="Nothobroscina">Nothobroscina</taxonomicName>
(
<bibRefCitation id="6E214B24FB6EFFDEFF33FA8D0DC5FAD3" author="Seldon, D. S. &amp; Holwell, G. I." box="[159,442,1323,1349]" pageId="1" pageNumber="248" pagination="724 - 728" refId="ref13908" refString="Seldon, D. S. &amp; Holwell, G. I. (2019) Monteremita gen. nov. (Coleoptera: Carabidae), a new genus of ground beetle from New Caledonia. Austral Entomology, 58 (4), 724 - 728. https: // doi. org / 10.1111 / aen. 12422" type="journal article" year="2019">Seldon &amp; Holwell 2019</bibRefCitation>
),
<taxonomicName id="CDB04D56FB6EFFDEFE7DFA8D0E22FAD3" authorityName="Bates" authorityYear="1891" box="[465,605,1323,1349]" class="Insecta" family="Carabidae" genus="Eurylychnus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="248" phylum="Arthropoda" rank="genus">
<emphasis id="38C4EAC7FB6EFFDEFE7DFA8D0E22FAD3" box="[465,605,1323,1349]" italics="true" pageId="1" pageNumber="248">Eurylychnus</emphasis>
</taxonomicName>
are reported as having a vertexal groove without punctures, abdominal ventrites 3–5 without paramedian setae and protarsi with squamose setae. However, this study has found exceptions in these characters: the vertexal grove is punctate in
<taxonomicName id="CDB04D56FB6EFFDEFD57FAD20FCDFA1B" box="[763,946,1395,1421]" class="Insecta" family="Carabidae" genus="Eurylychnus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="248" phylum="Arthropoda" rank="species" species="dyschirioides">
<emphasis id="38C4EAC7FB6EFFDEFD57FAD20FCDFA1B" box="[763,946,1395,1421]" italics="true" pageId="1" pageNumber="248">E. dyschirioides</emphasis>
</taxonomicName>
; abdominal ventrites 3–5 lack paramedian setae only in
<taxonomicName id="CDB04D56FB6EFFDEFE9DFA3E0DC2FA27" box="[305,445,1431,1457]" class="Insecta" family="Carabidae" genus="Eurylychnus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="248" phylum="Arthropoda" rank="species" species="blagravei">
<emphasis id="38C4EAC7FB6EFFDEFE9DFA3E0DC2FA27" box="[305,445,1431,1457]" italics="true" pageId="1" pageNumber="248">E. blagravei</emphasis>
</taxonomicName>
,
<taxonomicName id="CDB04D56FB6EFFDEFE61FA3E0E15FA27" authorityName="Sl. Id. By T. G. Sloane" authorityYear="1916" box="[461,618,1431,1457]" class="Insecta" family="Carabidae" genus="Eurylychnus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="248" phylum="Arthropoda" rank="species" species="cylindricus">
<emphasis id="38C4EAC7FB6EFFDEFE61FA3E0E15FA27" box="[461,618,1431,1457]" italics="true" pageId="1" pageNumber="248">E. cylindricus</emphasis>
</taxonomicName>
,
<taxonomicName id="CDB04D56FB6EFFDEFDD6FA3E0F7EFA27" box="[634,769,1431,1457]" class="Insecta" family="Carabidae" genus="Eurylychnus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="248" phylum="Arthropoda" rank="species" species="regularis">
<emphasis id="38C4EAC7FB6EFFDEFDD6FA3E0F7EFA27" box="[634,769,1431,1457]" italics="true" pageId="1" pageNumber="248">E. regularis</emphasis>
</taxonomicName>
and sometimes in
<emphasis id="38C4EAC7FB6EFFDEFC76FA3E0810FA27" box="[986,1135,1432,1457]" italics="true" pageId="1" pageNumber="248">
<taxonomicName id="CDB04D56FB6EFFDEFC76FA3E0816FA27" box="[986,1129,1432,1457]" class="Insecta" family="Carabidae" genus="Eurylychnus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="248" phylum="Arthropoda" rank="species" species="ovipennis">E. ovipennis</taxonomicName>
;
</emphasis>
squamose setae is present under protarsomeres 1–3 only in males of
<taxonomicName id="CDB04D56FB6EFFDEFD24FA1A0F40FA43" box="[648,831,1467,1493]" class="Insecta" family="Carabidae" genus="Eurylychnus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="248" phylum="Arthropoda" rank="species" species="dyschirioides">
<emphasis id="38C4EAC7FB6EFFDEFD24FA1A0F40FA43" box="[648,831,1467,1493]" italics="true" pageId="1" pageNumber="248">E. dyschirioides</emphasis>
</taxonomicName>
and
<emphasis id="38C4EAC7FB6EFFDEFCD0FA1A086DFA43" box="[892,1042,1468,1493]" italics="true" pageId="1" pageNumber="248">
<taxonomicName id="CDB04D56FB6EFFDEFCD0FA1A0871FA43" box="[892,1038,1468,1493]" class="Insecta" family="Carabidae" genus="Eurylychnus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="248" phylum="Arthropoda" rank="species" species="ovipennis">E. ovipennis</taxonomicName>
.
</emphasis>
<bibRefCitation id="6E214B24FB6EFFDEFBB7FA1D0971FA43" author="Baehr, M. &amp; Will, K." box="[1051,1294,1467,1493]" pageId="1" pageNumber="248" pagination="1802" refId="ref13189" refString="Baehr, M. &amp; Will, K. (2019) 12. Carabidae Latreille, 1802. In: Slipinski, A. &amp; Lawrence, J. (Eds.), Australian Beetles. Vol. 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, Victoria, pp. 61 - 217. https: // doi. org / 10.1071 / 9780643097315" type="book chapter" year="2019">Baehr &amp; Will (2019)</bibRefCitation>
erroneously reported that squamose setae under male protarsus did not occur in
<taxonomicName id="CDB04D56FB6EFFDEFC08FA79084FFA6F" authorityName="Bates" authorityYear="1891" box="[932,1072,1503,1529]" class="Insecta" family="Carabidae" genus="Eurylychnus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="248" phylum="Arthropoda" rank="genus">
<emphasis id="38C4EAC7FB6EFFDEFC08FA79084FFA6F" box="[932,1072,1503,1529]" italics="true" pageId="1" pageNumber="248">Eurylychnus</emphasis>
</taxonomicName>
.
<bibRefCitation id="6E214B24FB6EFFDEFBECFA46095CFA6F" author="Moore, B. P. &amp; Weir, T. A. &amp; Pyke, J. E." box="[1088,1315,1503,1530]" pageId="1" pageNumber="248" refId="ref13723" refString="Moore, B. P., Weir, T. A. &amp; Pyke, J. E. (1987) Zoological catalogue of Australia. Vol. 4. Coleoptera: Rhysodidae and Carabidae. Australian Government Publishing Service, Canberra, 444 pp." type="book" year="1987">
Moore
<emphasis id="38C4EAC7FB6EFFDEFB38FA4608B0FA6F" box="[1172,1231,1503,1529]" italics="true" pageId="1" pageNumber="248">et al.</emphasis>
(1987)
</bibRefCitation>
noted this genus as occurring in
<collectingCountry id="72A77645FB6EFFDEFE3EF9A20E3CF98B" box="[402,579,1539,1566]" name="New Caledonia" pageId="1" pageNumber="248">New Caledonia</collectingCountry>
, however the specimens collected from there have since been assigned to a new monotypic genus
<taxonomicName id="CDB04D56FB6EFFDEFE38F98E0F37F9D7" authority="Seldon &amp; Holwell (2019)" authorityName="Seldon &amp; Holwell" authorityYear="2019" box="[404,840,1575,1601]" class="Insecta" family="Carabidae" genus="Monteremita" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="248" phylum="Arthropoda" rank="genus">
<emphasis id="38C4EAC7FB6EFFDEFE38F98E0E5BF9D7" box="[404,548,1576,1601]" italics="true" pageId="1" pageNumber="248">Monteremita</emphasis>
<bibRefCitation id="6E214B24FB6EFFDEFD87F9810F37F9D7" author="Seldon, D. S. &amp; Holwell, G. I." box="[555,840,1575,1601]" pageId="1" pageNumber="248" pagination="724 - 728" refId="ref13908" refString="Seldon, D. S. &amp; Holwell, G. I. (2019) Monteremita gen. nov. (Coleoptera: Carabidae), a new genus of ground beetle from New Caledonia. Austral Entomology, 58 (4), 724 - 728. https: // doi. org / 10.1111 / aen. 12422" type="journal article" year="2019">Seldon &amp; Holwell (2019)</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="0A0F36D5FB6EFFDEFF6BF9ED0FC0F813" blockId="1.[151,1437,459,1928]" pageId="1" pageNumber="248">
Details of the life history and biology are not well documented beyond being flightless nocturnal predators (
<bibRefCitation id="6E214B24FB6EFFDEFF33F9C90D0EF91F" author="Moore, B. P. &amp; Weir, T. A. &amp; Pyke, J. E." box="[159,369,1647,1673]" pageId="1" pageNumber="248" refId="ref13723" refString="Moore, B. P., Weir, T. A. &amp; Pyke, J. E. (1987) Zoological catalogue of Australia. Vol. 4. Coleoptera: Rhysodidae and Carabidae. Australian Government Publishing Service, Canberra, 444 pp." type="book" year="1987">
Moore
<emphasis id="38C4EAC7FB6EFFDEFF58F9D60D51F91F" box="[244,302,1647,1673]" italics="true" pageId="1" pageNumber="248">et al.</emphasis>
1987
</bibRefCitation>
;
<bibRefCitation id="6E214B24FB6EFFDEFED2F9C90E21F91F" author="Baehr, M. &amp; Will, K." box="[382,606,1647,1673]" pageId="1" pageNumber="248" pagination="1802" refId="ref13189" refString="Baehr, M. &amp; Will, K. (2019) 12. Carabidae Latreille, 1802. In: Slipinski, A. &amp; Lawrence, J. (Eds.), Australian Beetles. Vol. 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, Victoria, pp. 61 - 217. https: // doi. org / 10.1071 / 9780643097315" type="book chapter" year="2019">Baehr &amp; Will 2019</bibRefCitation>
). While collecting material for this study, the author collected specimens under fallen timber in cool temperate rainforest (
<taxonomicName id="CDB04D56FB6EFFDEFD78F9320F0DF93B" authorityName="Sl. Id. By T. G. Sloane" authorityYear="1916" box="[724,882,1683,1709]" class="Insecta" family="Carabidae" genus="Eurylychnus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="248" phylum="Arthropoda" rank="species" species="cylindricus">
<emphasis id="38C4EAC7FB6EFFDEFD78F9320F0DF93B" box="[724,882,1683,1709]" italics="true" pageId="1" pageNumber="248">E. cylindricus</emphasis>
</taxonomicName>
,
<taxonomicName id="CDB04D56FB6EFFDEFC28F9320872F93B" box="[900,1037,1683,1709]" class="Insecta" family="Carabidae" genus="Eurylychnus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="248" phylum="Arthropoda" rank="species" species="regularis">
<emphasis id="38C4EAC7FB6EFFDEFC28F9320872F93B" box="[900,1037,1683,1709]" italics="true" pageId="1" pageNumber="248">E. regularis</emphasis>
</taxonomicName>
,
<taxonomicName id="CDB04D56FB6EFFDEFBB3F93208D3F93B" box="[1055,1196,1684,1709]" class="Insecta" family="Carabidae" genus="Eurylychnus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="248" phylum="Arthropoda" rank="species" species="ovipennis">
<emphasis id="38C4EAC7FB6EFFDEFBB3F93208D3F93B" box="[1055,1196,1684,1709]" italics="true" pageId="1" pageNumber="248">E. ovipennis</emphasis>
</taxonomicName>
,
<taxonomicName id="CDB04D56FB6EFFDEFB12F9320939F93B" authorityName="Sloane" authorityYear="1915" box="[1214,1350,1683,1709]" class="Insecta" family="Carabidae" genus="Eurylychnus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="248" phylum="Arthropoda" rank="species" species="kershawi">
<emphasis id="38C4EAC7FB6EFFDEFB12F9320939F93B" box="[1214,1350,1683,1709]" italics="true" pageId="1" pageNumber="248">E. kershawi</emphasis>
</taxonomicName>
and
<taxonomicName id="CDB04D56FB6EFFDEFA28F9320C84F947" authorityName="Sloane" authorityYear="1892" class="Insecta" family="Carabidae" genus="Eurylychnus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="248" phylum="Arthropoda" rank="species" species="victoriae">
<emphasis id="38C4EAC7FB6EFFDEFA28F9320C84F947" italics="true" pageId="1" pageNumber="248">E. victoriae</emphasis>
</taxonomicName>
) and Eucalypt Forest (
<emphasis id="38C4EAC7FB6EFFDEFDAFF91E0FFDF947" box="[515,898,1719,1745]" italics="true" pageId="1" pageNumber="248">
<taxonomicName id="CDB04D56FB6EFFDEFDAFF91E0EEFF947" box="[515,656,1719,1745]" class="Insecta" family="Carabidae" genus="Eurylychnus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="248" phylum="Arthropoda" rank="species" species="blagravei">E. blagravei</taxonomicName>
and
<taxonomicName id="CDB04D56FB6EFFDEFD60F91E0FFDF947" box="[716,898,1719,1745]" class="Insecta" family="Carabidae" genus="Eurylychnus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="248" phylum="Arthropoda" rank="species" species="dyschirioides">E. dyschirioides</taxonomicName>
</emphasis>
) and, with the exception of
<taxonomicName id="CDB04D56FB6EFFDEFB65F91E092AF947" box="[1225,1365,1720,1745]" class="Insecta" family="Carabidae" genus="Eurylychnus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="248" phylum="Arthropoda" rank="species" species="ovipennis">
<emphasis id="38C4EAC7FB6EFFDEFB65F91E092AF947" box="[1225,1365,1720,1745]" italics="true" pageId="1" pageNumber="248">E. ovipennis</emphasis>
</taxonomicName>
, were quite abundant. Specimens of
<taxonomicName id="CDB04D56FB6EFFDEFE4AF97A0EFEF963" authorityName="Sl. Id. By T. G. Sloane" authorityYear="1916" box="[486,641,1755,1781]" class="Insecta" family="Carabidae" genus="Eurylychnus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="248" phylum="Arthropoda" rank="species" species="cylindricus">
<emphasis id="38C4EAC7FB6EFFDEFE4AF97A0EFEF963" box="[486,641,1755,1781]" italics="true" pageId="1" pageNumber="248">E. cylindricus</emphasis>
</taxonomicName>
were kept in captivity where they survived for 6 months from February to August and readily fed on mincemeat, suggesting scavenging could form part of the wild diet. The defensive secretions of
<taxonomicName id="CDB04D56FB6EFFDEFE86F8820DCCF8AB" box="[298,435,1827,1853]" class="Insecta" family="Carabidae" genus="Eurylychnus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="248" phylum="Arthropoda" rank="species" species="blagravei">
<emphasis id="38C4EAC7FB6EFFDEFE86F8820DCCF8AB" box="[298,435,1827,1853]" italics="true" pageId="1" pageNumber="248">E. blagravei</emphasis>
</taxonomicName>
and
<taxonomicName id="CDB04D56FB6EFFDEFE44F8820EE5F8AB" box="[488,666,1827,1853]" class="Insecta" family="Carabidae" genus="Eurylychnus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="248" phylum="Arthropoda" rank="species" species="dyschirioides">
<emphasis id="38C4EAC7FB6EFFDEFE44F8820EE5F8AB" box="[488,666,1827,1853]" italics="true" pageId="1" pageNumber="248">E. dyschirioides</emphasis>
</taxonomicName>
were examined by
<bibRefCitation id="6E214B24FB6EFFDEFCDDF88208E7F8AB" author="Moore, B. P. &amp; Wallbank, B. E." box="[881,1176,1827,1854]" pageId="1" pageNumber="248" pagination="62 - 72" refId="ref13648" refString="Moore, B. P. &amp; Wallbank, B. E. (1968) Chemical composition of the defensive secretion in carabid beetles and its importance as a taxonomic character. Proceedings of the Royal Entomological Society of London (B), 37 (5 - 6), 62 - 72. https: // doi. org / 10.1111 / j. 1365 - 3113.1968. tb 00199. x" type="journal article" year="1968">Moore &amp; Wallbank (1968)</bibRefCitation>
who detected saturated and unsaturated acids. The larvae of
<taxonomicName id="CDB04D56FB6EFFDEFD98F8E10EBFF8F7" authorityName="Bates" authorityYear="1891" box="[564,704,1863,1889]" class="Insecta" family="Carabidae" genus="Eurylychnus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="248" phylum="Arthropoda" rank="genus">
<emphasis id="38C4EAC7FB6EFFDEFD98F8E10EBFF8F7" box="[564,704,1863,1889]" italics="true" pageId="1" pageNumber="248">Eurylychnus</emphasis>
</taxonomicName>
are poorly documented, although the larvae of
<taxonomicName id="CDB04D56FB6EFFDEFB79F8EE091FF8F7" box="[1237,1376,1863,1889]" class="Insecta" family="Carabidae" genus="Eurylychnus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="248" phylum="Arthropoda" rank="species" species="blagravei">
<emphasis id="38C4EAC7FB6EFFDEFB79F8EE091FF8F7" box="[1237,1376,1863,1889]" italics="true" pageId="1" pageNumber="248">E. blagravei</emphasis>
</taxonomicName>
have been described (
<bibRefCitation id="6E214B24FB6EFFDEFEE1F8CD0DA6F813" author="Moore, B. P." box="[333,473,1899,1925]" pageId="1" pageNumber="248" pagination="242 - 246" refId="ref13612" refString="Moore, B. P. (1964) Australian larval Carabidae of the subfamilies Broscinae, Psydrinae and Pseudomorphinae (Coleoptera). Pacific Insects, 6 (2), 242 - 246." type="journal article" year="1964">Moore 1964</bibRefCitation>
) and figured (
<bibRefCitation id="6E214B24FB6EFFDEFDDBF8CD0FCCF813" author="Lawrence, J. F. &amp; Slipinski, A." box="[631,947,1899,1928]" pageId="1" pageNumber="248" refId="ref13419" refString="Lawrence, J. F. &amp; Slipinski, A. (2013) Australian Beetles. Vol. 1. Morphology, Classification and Keys. CSIRO Publishing, Collingwood, Victoria, 576 pp. https: // doi. org / 10.1071 / 9780643097292" type="book" year="2013">Lawrence &amp; Ślipiński 2013</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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