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<mods:title id="959607900CF401F2E4D65F55FB172045">Extraordinary majoid crabs: the genus Esopus A. Milne-Edwards, 1875 in the new subfamily Esopinae subfam. nov., and erection of Paulitinae subfam. nov. (Crustacea, Decapoda, Brachyura, Majoidea, Inachoididae Dana, 1851)</mods:title>
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[
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Salacini [
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Salaciinae 
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Salaciini 
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genus
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<emphasis id="DCE4EAC2FFB9A61FB743FD6DFE71FD0A" box="[300,436,664,690]" italics="true" pageId="16" pageNumber="117">Leurocyclus</emphasis>
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, 
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H. 
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being preoccupied by 
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<emphasis id="DCE4EAC2FFB9A61FB327FD6DFA59FD0A" box="[1352,1436,664,690]" italics="true" pageId="16" pageNumber="117">Salacia</emphasis>
Lamouroux, 1816
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, a hydrozoan, and replaced by 
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<emphasis id="DCE4EAC2FFB9A61FB4A6FD49FC94FD6E" box="[713,849,700,726]" italics="true" pageId="16" pageNumber="117">Leurocyclus</emphasis>
(
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[(
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species: 
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<emphasis id="DCE4EAC2FFB9A61FB2ADFD48FAAAFD6E" box="[1218,1391,700,726]" italics="true" pageId="16" pageNumber="117">L. tuberculosus</emphasis>
(H. 
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)
</taxonomicName>
]. See also 
<bibRefCitation id="8A014B21FFB9A61FB419FD15FCC8FD43" author="Miers" box="[630,781,736,763]" firstAuthor="Miers" pageId="16" pageNumber="117" pagination="634 - 673" refId="ref23493" refString="Miers, E. J. (1879) On the classification of the maioid Crustacea or Oxyrhyncha, with a synopsis of the families, subfamilies, and genera. Journal of the Linnean Society, London, 14, 634 - 673, pls. 12, 13. https: // doi. org / 10.1111 / j. 1096 - 3642.1879. tb 02457. x" type="book chapter" year="1879">Miers (1879)</bibRefCitation>
. Although 
<taxonomicName id="29904D53FFB9A61FB5E7FD15FC19FD42" ID-CoL="7PG52" authorityName="Lamouroux" authorityYear="1816" box="[904,988,736,762]" class="Malacostraca" family="Inachoididae" genus="Salacia" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="DCE4EAC2FFB9A61FB5E7FD15FC19FD42" box="[904,988,736,762]" italics="true" pageId="16" pageNumber="117">Salacia</emphasis>
</taxonomicName>
is not a valid nomen, the subfamily 
<taxonomicName id="29904D53FFB9A61FB318FD14FEB2FCA7" authority="Dana, 1851" authorityName="Dana" authorityYear="1851" class="Malacostraca" family="Inachoididae" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="117" phylum="Arthropoda" rank="subFamily" subFamily="Salaciinae">
Salaciinae 
<bibRefCitation id="8A014B21FFB9A61FB69AFCF0FEB2FCA7" author="Dana" box="[245,375,772,799]" firstAuthor="Dana" pageId="16" pageNumber="117" pagination="425 - 434" refId="ref20420" refString="Dana, J. D. (1851) On the classification of the Maioid Crustacea or Oxyrhyncha. American Journal of Science and Arts, Series 2, 11, 425 - 434." type="journal article" year="1851">Dana, 1851</bibRefCitation>
</taxonomicName>
(
<taxonomicName id="29904D53FFB9A61FB7EAFCF1FE37FCA6" authorityName="Dana" authorityYear="1851" box="[389,498,772,798]" class="Malacostraca" family="Inachoididae" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="117" phylum="Arthropoda" rank="subFamily" subFamily="Salacinae">Salacinae</taxonomicName>
is an incorrect spelling of the nomen) is an available nomen that was used by 
<bibRefCitation id="8A014B21FFB9A61FB30EFCF0FEDBFCFB" author="Dana" firstAuthor="Dana" pageId="16" pageNumber="117" pagination="425 - 434" refId="ref20420" refString="Dana, J. D. (1851) On the classification of the Maioid Crustacea or Oxyrhyncha. American Journal of Science and Arts, Series 2, 11, 425 - 434." type="journal article" year="1851">Dana (1851: 430)</bibRefCitation>
, with 
<taxonomicName id="29904D53FFB9A61FB70AFCDDFE28FCFA" ID-CoL="7PMRD" baseAuthorityName="Rathbun" baseAuthorityYear="1897" box="[357,493,808,834]" class="Malacostraca" family="Inachoididae" genus="Leurocyclus" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="DCE4EAC2FFB9A61FB70AFCDDFE28FCFA" box="[357,493,808,834]" italics="true" pageId="16" pageNumber="117">Leurocyclus</emphasis>
</taxonomicName>
as 
<typeStatus id="312B8872FFB9A61FB47BFCDCFD81FCFB" box="[532,580,809,835]" pageId="16" pageNumber="117">type</typeStatus>
genus (see 
<bibRefCitation id="8A014B21FFB9A61FB4A4FCDDFC0DFCFB" author="Drach &amp; Guinot" box="[715,968,808,835]" firstAuthor="Drach" pageId="16" pageNumber="117" pagination="37 - 42" refId="ref21056" refString="Drach, P. &amp; Guinot, D. (1983) Les Inachoididae Dana, famille de Majoidea caracterisee par des connexions morphologiques d'un type nouveau entre carapace, pleurites, sternites et pleon (Crustacea Decapoda). Comptes rendus hebdomadaires des Seances de l'Academie des Sciences, Serie 3, 297, 37 - 42." type="journal article" year="1983">Drach &amp; Guinot 1983</bibRefCitation>
: figs. 6–9; 
<bibRefCitation id="8A014B21FFB9A61FB229FCDDFB1DFCFB" author="Guinot" box="[1094,1240,808,835]" firstAuthor="Guinot" pageId="16" pageNumber="117" pagination="377 - 395" refId="ref21724" refString="Guinot, D. (1984) Le genre Leurocyclus Rathbun, 1897 (Crustacea Decapoda Brachyura). Bulletin du Museum national d'Histoire naturelle, ser. 4, 6, sect. A (2), 377 - 395." type="book chapter" year="1984">Guinot 1984</bibRefCitation>
: fig. 1; 
<bibRefCitation id="8A014B21FFB9A61FB35FFCDDFE52FCDF" author="Guinot &amp; Richer de Forges" firstAuthor="Guinot" pageId="16" pageNumber="117" pagination="453 - 502" refId="ref22156" refString="Guinot, D. &amp; Richer de Forges, B. (1997) Affinites entre les Hymenosomatidae MacLeay, 1838 et les Inachoididae Dana, 1851 (Crustacea Decapoda Brachyura). Zoosystema, 19 (2 &amp; 3), 453 - 502." type="journal article" year="1997">Guinot &amp; Richer de Forges 1997</bibRefCitation>
: fig. 12B; 
<bibRefCitation id="8A014B21FFB9A61FB464FCB9FCF0FCDF" author="Guinot &amp; Cleva" box="[523,821,844,871]" firstAuthor="Guinot" pageId="16" pageNumber="117" pagination="35 - 43" refId="ref22037" refString="Guinot, D. &amp; Cleva, R. (2002 a) D'Orbigny et les Crustaces. In: Taquet, P. (Ed.), Alcide d'Orbigny. Du Nouveau Monde ... au Passe du Monde. Nathan / VUEF et Museum national d'Histoire naturelle, Paris, pp. 35 - 43." type="book chapter" year="2002" yearSuffix="a">Guinot &amp; Cleva 2002a: 40</bibRefCitation>
; 
<bibRefCitation id="8A014B21FFB9A61FB52EFCB9FC07FCDF" author="Guinot &amp; Cleva" box="[833,962,844,871]" firstAuthor="Guinot" pageId="16" pageNumber="117" pagination="499 - 515" refId="ref22095" refString="Guinot, D. &amp; Cleva, R. (2002 b) Les Crustaces recoltes par d'Orbigny en Amerique du Sud et deposes au Museum national d'Histoire naturelle, Paris. Palevol, Paris, 1, 499 - 515. https: // doi. org / 10.1016 / S 1631 - 0683 (02) 00055 - 6" type="journal article" year="2002" yearSuffix="b">2002b: 508</bibRefCitation>
; 
<bibRefCitation id="8A014B21FFB9A61FB5A2FCB8FBADFCDF" author="Santana, W. R. A." box="[973,1128,845,871]" pageId="16" pageNumber="117" refId="ref24839" refString="Santana, W. R. A. (2008) Revisao taxonomica e relacoes filogeneticas em Inachoididae Dana, 1851 (Crustacea, Brachyura, Majoidea). Unpublished doctoral thesis, Universidade de Sao Paulo, Sao Paulo, 244 pp." type="book" year="2008">Santana 2008</bibRefCitation>
: figs. 45C, D, 47B, C, 48A; 
<bibRefCitation id="8A014B21FFB9A61FB6F8FC85FEA3FC33" author="Guinot" box="[151,358,880,907]" etAl="et al." firstAuthor="Guinot" pageId="16" pageNumber="117" pagination="1 - 414" refId="ref22407" refString="Guinot, D., Tavares, M. &amp; Castro, P. (2013) Significance of the sexual openings and supplementary structures on the phylogeny of brachyuran crabs (Crustacea, Decapoda, Brachyura), with new nomina for higher-ranked podotreme taxa. Zootaxa, 3665 (1), 1 - 414. https: // doi. org / 10.11646 / zootaxa. 3665.1.1" type="journal article" year="2013">
Guinot 
<emphasis id="DCE4EAC2FFB9A61FB683FC84FEE1FC32" box="[236,292,880,906]" italics="true" pageId="16" pageNumber="117">et al.</emphasis>
2013
</bibRefCitation>
: fig. 47I). An unfortunate typographical mistake in (2012) Guinot’s abstract attributed the authorhip of the family to H. 
<bibRefCitation id="8A014B21FFB9A61FB7CDFC61FD38FC17" author="Milne Edwards &amp; Lucas" box="[418,765,916,943]" firstAuthor="Milne Edwards" pageId="16" pageNumber="117" refId="ref23942" refString="Milne Edwards, H. &amp; Lucas, H. (1842 - 1844) Crustaces. In: d'Orbigny, A. (Ed.), Voyage dans l'Amerique meridionale (le Bresil, la Republique orientale de l'Uruguay, la Republique Argentine, la Patagonie, la Republique du Chili, la Republique de Bolivia, la Republique du Perou), execute pendant les annees 1826, 1827, 1828, 1829, 1830, 1831, 1832 et 1833. Tome sixieme. Premiere partie. P. Bertrand, Paris and Vve Levrault, Strasbourg, pls. 1 + 3 (1842: 62); pls. 4 + 6 + 13 (67); pls. 2 + 10 (68); pp. 1 - 8, pls. 5 + 7 + 11 (1843: 69); pp. 9 - 16, pl. 8 bis (71); pp. 17 - 24, pl. 8 (1844: 72); pp. 25 - 32, pl. 7 bis (73); 33 - 39, pl. 9 (74); pls. 14 - 16 (75); pl. 17 (77)] https: // doi. org / 10.5962 / bhl. title. 110540" type="journal volume" year="1842">Milne Edwards &amp; Lucas, 1842</bibRefCitation>
, instead of 
<bibRefCitation id="8A014B21FFB9A61FB512FC60FBC4FC17" author="Dana" box="[893,1025,916,943]" firstAuthor="Dana" pageId="16" pageNumber="117" pagination="425 - 434" refId="ref20420" refString="Dana, J. D. (1851) On the classification of the Maioid Crustacea or Oxyrhyncha. American Journal of Science and Arts, Series 2, 11, 425 - 434." type="journal article" year="1851">Dana, 1851</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A68A655BFFB9A61FB6A8FC4DFAB3F85E" pageId="16" pageNumber="117" type="description">
<paragraph id="EE2F36D0FFB9A61FB6A8FC4DFAB3F85E" blockId="16.[151,1437,664,2023]" pageId="16" pageNumber="117">
<emphasis id="DCE4EAC2FFB9A61FB6A8FC4DFE94FC6B" bold="true" box="[199,337,952,979]" pageId="16" pageNumber="117">Description</emphasis>
. Carapace much broader than long, suborbicular, depressed, dilated posterolaterally (H. 
<bibRefCitation id="8A014B21FFB9A61FB336FC4DFE61FC4F" author="Milne Edwards &amp; Lucas" firstAuthor="Milne Edwards" pageId="16" pageNumber="117" refId="ref23942" refString="Milne Edwards, H. &amp; Lucas, H. (1842 - 1844) Crustaces. In: d'Orbigny, A. (Ed.), Voyage dans l'Amerique meridionale (le Bresil, la Republique orientale de l'Uruguay, la Republique Argentine, la Patagonie, la Republique du Chili, la Republique de Bolivia, la Republique du Perou), execute pendant les annees 1826, 1827, 1828, 1829, 1830, 1831, 1832 et 1833. Tome sixieme. Premiere partie. P. Bertrand, Paris and Vve Levrault, Strasbourg, pls. 1 + 3 (1842: 62); pls. 4 + 6 + 13 (67); pls. 2 + 10 (68); pp. 1 - 8, pls. 5 + 7 + 11 (1843: 69); pp. 9 - 16, pl. 8 bis (71); pp. 17 - 24, pl. 8 (1844: 72); pp. 25 - 32, pl. 7 bis (73); 33 - 39, pl. 9 (74); pls. 14 - 16 (75); pl. 17 (77)] https: // doi. org / 10.5962 / bhl. title. 110540" type="journal volume" year="1842">Milne Edwards &amp; Lucas 1842</bibRefCitation>
: pl. 2, fig. 1, as 
<taxonomicName id="29904D53FFB9A61FB436FC29FD68FC4E" authorityName="Lamouroux" authorityYear="1816" box="[601,685,988,1014]" class="Malacostraca" family="Inachoididae" genus="Salacia" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="DCE4EAC2FFB9A61FB436FC29FD68FC4E" box="[601,685,988,1014]" italics="true" pageId="16" pageNumber="117">Salacia</emphasis>
</taxonomicName>
; 
<bibRefCitation id="8A014B21FFB9A61FB4D2FC29FC9AFC4F" author="Rathbun" box="[701,863,988,1015]" firstAuthor="Rathbun" pageId="16" pageNumber="117" pagination="1 - 613" refId="ref24756" refString="Rathbun, M. J. (1925) The spider crabs of America. Bulletin of the U. S. National Museum, 129, 1 - 613. https: // doi. org / 10.5479 / si. 03629236.129. i" type="journal article" year="1925">Rathbun 1925</bibRefCitation>
: pl. 233, fig. 1; 
<bibRefCitation id="8A014B21FFB9A61FB27DFC29FAC8FC4F" author="Drach &amp; Guinot" box="[1042,1293,988,1015]" firstAuthor="Drach" pageId="16" pageNumber="117" pagination="37 - 42" refId="ref21056" refString="Drach, P. &amp; Guinot, D. (1983) Les Inachoididae Dana, famille de Majoidea caracterisee par des connexions morphologiques d'un type nouveau entre carapace, pleurites, sternites et pleon (Crustacea Decapoda). Comptes rendus hebdomadaires des Seances de l'Academie des Sciences, Serie 3, 297, 37 - 42." type="journal article" year="1983">Drach &amp; Guinot 1983</bibRefCitation>
: pl. 1, fig. 6; 
<bibRefCitation id="8A014B21FFB9A61FB6F8FBF5FEE2FBA3" author="Guinot" box="[151,295,1024,1051]" firstAuthor="Guinot" pageId="16" pageNumber="117" pagination="377 - 395" refId="ref21724" refString="Guinot, D. (1984) Le genre Leurocyclus Rathbun, 1897 (Crustacea Decapoda Brachyura). Bulletin du Museum national d'Histoire naturelle, ser. 4, 6, sect. A (2), 377 - 395." type="book chapter" year="1984">Guinot 1984</bibRefCitation>
: fig. 1; 
<bibRefCitation id="8A014B21FFB9A61FB715FBF5FE3DFBA2" author="Melo" box="[378,504,1024,1050]" firstAuthor="Melo" pageId="16" pageNumber="117" refId="ref23457" refString="Melo, G. A. S. de (1996) Manual de Identificacao dos Brachyura (Caranguejos e Siris) do Litoral Brasileiro. Pleiade / FAPESP, Sao Paulo, 604 pp." type="book" year="1996">Melo 1996</bibRefCitation>
: fig. p. 207; 
<bibRefCitation id="8A014B21FFB9A61FB4EDFBF5FC37FBA3" author="Guinot &amp; Richer de Forges" box="[642,1010,1024,1051]" firstAuthor="Guinot" pageId="16" pageNumber="117" pagination="453 - 502" refId="ref22156" refString="Guinot, D. &amp; Richer de Forges, B. (1997) Affinites entre les Hymenosomatidae MacLeay, 1838 et les Inachoididae Dana, 1851 (Crustacea Decapoda Brachyura). Zoosystema, 19 (2 &amp; 3), 453 - 502." type="journal article" year="1997">Guinot &amp; Richer de Forges 1997</bibRefCitation>
: fig. 12B). Carapace resting on setting gutter. Dorsal surface with longitudinal carina marked by granules; regions well marked by deep grooves and ornamented (
<bibRefCitation id="8A014B21FFB9A61FB698FBBCFE56FBDB" author="Santana, W. R. A." box="[247,403,1097,1123]" pageId="16" pageNumber="117" refId="ref24839" refString="Santana, W. R. A. (2008) Revisao taxonomica e relacoes filogeneticas em Inachoididae Dana, 1851 (Crustacea, Brachyura, Majoidea). Unpublished doctoral thesis, Universidade de Sao Paulo, Sao Paulo, 244 pp." type="book" year="2008">Santana 2008</bibRefCitation>
: figs. 45C, D, 47C). Rostrum small, narrow, subtriangular. Proepistome laterally compressed, reduced to recessed, concealed portion, not joining anterior process (a narrow septum) of rudimentary epistome (
<bibRefCitation id="8A014B21FFB9A61FB6F0FB65FE87FB13" author="Rathbun" box="[159,322,1168,1195]" firstAuthor="Rathbun" pageId="16" pageNumber="117" pagination="1 - 613" refId="ref24756" refString="Rathbun, M. J. (1925) The spider crabs of America. Bulletin of the U. S. National Museum, 129, 1 - 613. https: // doi. org / 10.5479 / si. 03629236.129. i" type="journal article" year="1925">Rathbun 1925</bibRefCitation>
: pl. 232, fig. 6). Antennulae: both vertically folded in two fossae, without visible separation, the latter being deeply recessed. Antenna: urinary article and basal article adjacent, seemingly fused; basal article (2+3) very wide, concave, without longitudinal carina, with lateroexternal lobe (H. 
<bibRefCitation id="8A014B21FFB9A61FB590FB2DFA92FB4B" author="Milne Edwards &amp; Lucas" box="[1023,1367,1240,1267]" firstAuthor="Milne Edwards" pageId="16" pageNumber="117" refId="ref23942" refString="Milne Edwards, H. &amp; Lucas, H. (1842 - 1844) Crustaces. In: d'Orbigny, A. (Ed.), Voyage dans l'Amerique meridionale (le Bresil, la Republique orientale de l'Uruguay, la Republique Argentine, la Patagonie, la Republique du Chili, la Republique de Bolivia, la Republique du Perou), execute pendant les annees 1826, 1827, 1828, 1829, 1830, 1831, 1832 et 1833. Tome sixieme. Premiere partie. P. Bertrand, Paris and Vve Levrault, Strasbourg, pls. 1 + 3 (1842: 62); pls. 4 + 6 + 13 (67); pls. 2 + 10 (68); pp. 1 - 8, pls. 5 + 7 + 11 (1843: 69); pp. 9 - 16, pl. 8 bis (71); pp. 17 - 24, pl. 8 (1844: 72); pp. 25 - 32, pl. 7 bis (73); 33 - 39, pl. 9 (74); pls. 14 - 16 (75); pl. 17 (77)] https: // doi. org / 10.5962 / bhl. title. 110540" type="journal volume" year="1842">Milne Edwards &amp; Lucas 1842</bibRefCitation>
: pl. 2, fig. 2, as 
<taxonomicName id="29904D53FFB9A61FB76DFB09FE93FAAE" authorityName="Lamouroux" authorityYear="1816" box="[258,342,1276,1302]" class="Malacostraca" family="Inachoididae" genus="Salacia" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="DCE4EAC2FFB9A61FB76DFB09FE93FAAE" box="[258,342,1276,1302]" italics="true" pageId="16" pageNumber="117">Salacia</emphasis>
</taxonomicName>
; 
<bibRefCitation id="8A014B21FFB9A61FB709FB09FDCCFAAF" author="Rathbun" box="[358,521,1276,1303]" firstAuthor="Rathbun" pageId="16" pageNumber="117" pagination="1 - 613" refId="ref24756" refString="Rathbun, M. J. (1925) The spider crabs of America. Bulletin of the U. S. National Museum, 129, 1 - 613. https: // doi. org / 10.5479 / si. 03629236.129. i" type="journal article" year="1925">Rathbun 1925</bibRefCitation>
: pl. 232, fig. 11; 
<bibRefCitation id="8A014B21FFB9A61FB4A3FB09FC9BFAAF" author="Guinot" box="[716,862,1276,1303]" firstAuthor="Guinot" pageId="16" pageNumber="117" pagination="377 - 395" refId="ref21724" refString="Guinot, D. (1984) Le genre Leurocyclus Rathbun, 1897 (Crustacea Decapoda Brachyura). Bulletin du Museum national d'Histoire naturelle, ser. 4, 6, sect. A (2), 377 - 395." type="book chapter" year="1984">Guinot 1984</bibRefCitation>
: fig. 2; 
<bibRefCitation id="8A014B21FFB9A61FB5D9FB08FB96FAAF" author="Santana, W. R. A." box="[950,1107,1277,1303]" pageId="16" pageNumber="117" refId="ref24839" refString="Santana, W. R. A. (2008) Revisao taxonomica e relacoes filogeneticas em Inachoididae Dana, 1851 (Crustacea, Brachyura, Majoidea). Unpublished doctoral thesis, Universidade de Sao Paulo, Sao Paulo, 244 pp." type="book" year="2008">Santana 2008</bibRefCitation>
: fig. 48A); following articles (antennal movable portion) entirely visible in dorsal view at side of rostrum. Orbits rather large, directed forward, close to each other; preorbital tooth absent; postorbital tooth large forming postocular cup (supraocular eave) accommodating eyestalk in resting position (
<bibRefCitation id="8A014B21FFB9A61FB411FA9CFCDBFA3B" author="Santana, W. R. A." box="[638,798,1385,1411]" pageId="16" pageNumber="117" refId="ref24839" refString="Santana, W. R. A. (2008) Revisao taxonomica e relacoes filogeneticas em Inachoididae Dana, 1851 (Crustacea, Brachyura, Majoidea). Unpublished doctoral thesis, Universidade de Sao Paulo, Sao Paulo, 244 pp." type="book" year="2008">Santana 2008</bibRefCitation>
: fig. 48A). Mxp3: ischium broad, with developed crista dentata formed by strong spines; merus cordiform (H. 
<bibRefCitation id="8A014B21FFB9A61FB498FA79FB8EFA1F" author="Milne Edwards &amp; Lucas" box="[759,1099,1420,1447]" firstAuthor="Milne Edwards" pageId="16" pageNumber="117" refId="ref23942" refString="Milne Edwards, H. &amp; Lucas, H. (1842 - 1844) Crustaces. In: d'Orbigny, A. (Ed.), Voyage dans l'Amerique meridionale (le Bresil, la Republique orientale de l'Uruguay, la Republique Argentine, la Patagonie, la Republique du Chili, la Republique de Bolivia, la Republique du Perou), execute pendant les annees 1826, 1827, 1828, 1829, 1830, 1831, 1832 et 1833. Tome sixieme. Premiere partie. P. Bertrand, Paris and Vve Levrault, Strasbourg, pls. 1 + 3 (1842: 62); pls. 4 + 6 + 13 (67); pls. 2 + 10 (68); pp. 1 - 8, pls. 5 + 7 + 11 (1843: 69); pp. 9 - 16, pl. 8 bis (71); pp. 17 - 24, pl. 8 (1844: 72); pp. 25 - 32, pl. 7 bis (73); 33 - 39, pl. 9 (74); pls. 14 - 16 (75); pl. 17 (77)] https: // doi. org / 10.5962 / bhl. title. 110540" type="journal volume" year="1842">Milne Edwards &amp; Lucas 1842</bibRefCitation>
: pl. 2, fig. 5, as 
<taxonomicName id="29904D53FFB9A61FB293FA79FA95FA1E" authorityName="Lamouroux" authorityYear="1816" box="[1276,1360,1420,1446]" class="Malacostraca" family="Inachoididae" genus="Salacia" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="DCE4EAC2FFB9A61FB293FA79FA95FA1E" box="[1276,1360,1420,1446]" italics="true" pageId="16" pageNumber="117">Salacia</emphasis>
</taxonomicName>
; 
<bibRefCitation id="8A014B21FFB9A61FB331FA79FEC0FA73" author="Rathbun" firstAuthor="Rathbun" pageId="16" pageNumber="117" pagination="1 - 613" refId="ref24756" refString="Rathbun, M. J. (1925) The spider crabs of America. Bulletin of the U. S. National Museum, 129, 1 - 613. https: // doi. org / 10.5479 / si. 03629236.129. i" type="journal article" year="1925">Rathbun 1925</bibRefCitation>
: pl. 232, figs. 8, 10). Pleurites regularly connecting medially and fused to carapace by pillars (
<bibRefCitation id="8A014B21FFB9A61FB357FA45FEE2FA57" author="Drach &amp; Guinot" firstAuthor="Drach" pageId="16" pageNumber="117" pagination="37 - 42" refId="ref21056" refString="Drach, P. &amp; Guinot, D. (1983) Les Inachoididae Dana, famille de Majoidea caracterisee par des connexions morphologiques d'un type nouveau entre carapace, pleurites, sternites et pleon (Crustacea Decapoda). Comptes rendus hebdomadaires des Seances de l'Academie des Sciences, Serie 3, 297, 37 - 42." type="journal article" year="1983">Drach &amp; Guinot 1983</bibRefCitation>
: pl. 1, figs. 7, 8). Thoracic sternum: lateral suture 3/4 short, ending as perforation of the sternal surface (
<bibRefCitation id="8A014B21FFB9A61FB6F0FA0DFE84F9AB" author="Rathbun" box="[159,321,1528,1555]" firstAuthor="Rathbun" pageId="16" pageNumber="117" pagination="1 - 613" refId="ref24756" refString="Rathbun, M. J. (1925) The spider crabs of America. Bulletin of the U. S. National Museum, 129, 1 - 613. https: // doi. org / 10.5479 / si. 03629236.129. i" type="journal article" year="1925">Rathbun 1925</bibRefCitation>
: pl. 233, fig. 2); sutures 4/5–7/8 interrupted; a bridge at base of sternite 8, then median line extending on sternite 8. Sternite 3 forming a narrow, short extension; no sternum/pterygostome junction at level of sternite 4 (H. 
<bibRefCitation id="8A014B21FFB9A61FB6ACF9B5FDDDF9E3" author="Milne Edwards &amp; Lucas" box="[195,536,1600,1627]" firstAuthor="Milne Edwards" pageId="16" pageNumber="117" refId="ref23942" refString="Milne Edwards, H. &amp; Lucas, H. (1842 - 1844) Crustaces. In: d'Orbigny, A. (Ed.), Voyage dans l'Amerique meridionale (le Bresil, la Republique orientale de l'Uruguay, la Republique Argentine, la Patagonie, la Republique du Chili, la Republique de Bolivia, la Republique du Perou), execute pendant les annees 1826, 1827, 1828, 1829, 1830, 1831, 1832 et 1833. Tome sixieme. Premiere partie. P. Bertrand, Paris and Vve Levrault, Strasbourg, pls. 1 + 3 (1842: 62); pls. 4 + 6 + 13 (67); pls. 2 + 10 (68); pp. 1 - 8, pls. 5 + 7 + 11 (1843: 69); pp. 9 - 16, pl. 8 bis (71); pp. 17 - 24, pl. 8 (1844: 72); pp. 25 - 32, pl. 7 bis (73); 33 - 39, pl. 9 (74); pls. 14 - 16 (75); pl. 17 (77)] https: // doi. org / 10.5962 / bhl. title. 110540" type="journal volume" year="1842">Milne Edwards &amp; Lucas 1842</bibRefCitation>
: pl. 2, fig. 8, as 
<taxonomicName id="29904D53FFB9A61FB4A4F9B5FCDAF9E2" authorityName="Lamouroux" authorityYear="1816" box="[715,799,1600,1626]" class="Malacostraca" family="Inachoididae" genus="Salacia" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="DCE4EAC2FFB9A61FB4A4F9B5FCDAF9E2" box="[715,799,1600,1626]" italics="true" pageId="16" pageNumber="117">Salacia</emphasis>
</taxonomicName>
; Guinot 1979: pl. 16, figs. 5, 6; 
<bibRefCitation id="8A014B21FFB9A61FB2E7F9B4FAE1F9E3" author="Santana, W. R. A." box="[1160,1316,1601,1627]" pageId="16" pageNumber="117" refId="ref24839" refString="Santana, W. R. A. (2008) Revisao taxonomica e relacoes filogeneticas em Inachoididae Dana, 1851 (Crustacea, Brachyura, Majoidea). Unpublished doctoral thesis, Universidade de Sao Paulo, Sao Paulo, 244 pp." type="book" year="2008">Santana 2008</bibRefCitation>
: fig. 47D). Milne-Edwards openings not separated from chelipeds. Sternal extensions joining exposed pleurites (sternum/pleurites connections) between pereiopods. Pleurites 5–8 exposed but proportionally small and narrow, each pointed at each end (thus collar with festooned appearance), ornamented (
<bibRefCitation id="8A014B21FFB9A61FB53BF959FB89F97F" author="Drach &amp; Guinot" box="[852,1100,1708,1735]" firstAuthor="Drach" pageId="16" pageNumber="117" pagination="37 - 42" refId="ref21056" refString="Drach, P. &amp; Guinot, D. (1983) Les Inachoididae Dana, famille de Majoidea caracterisee par des connexions morphologiques d'un type nouveau entre carapace, pleurites, sternites et pleon (Crustacea Decapoda). Comptes rendus hebdomadaires des Seances de l'Academie des Sciences, Serie 3, 297, 37 - 42." type="journal article" year="1983">Drach &amp; Guinot 1983</bibRefCitation>
: pl. 1, fig. 6; 
<bibRefCitation id="8A014B21FFB9A61FB2B3F959FAA9F97F" author="Guinot" box="[1244,1388,1708,1735]" firstAuthor="Guinot" pageId="16" pageNumber="117" pagination="377 - 395" refId="ref21724" refString="Guinot, D. (1984) Le genre Leurocyclus Rathbun, 1897 (Crustacea Decapoda Brachyura). Bulletin du Museum national d'Histoire naturelle, ser. 4, 6, sect. A (2), 377 - 395." type="book chapter" year="1984">Guinot 1984</bibRefCitation>
: fig. 1; 
<bibRefCitation id="8A014B21FFB9A61FB6DCF924FE8BF953" author="Santana, W. R. A." box="[179,334,1745,1771]" pageId="16" pageNumber="117" refId="ref24839" refString="Santana, W. R. A. (2008) Revisao taxonomica e relacoes filogeneticas em Inachoididae Dana, 1851 (Crustacea, Brachyura, Majoidea). Unpublished doctoral thesis, Universidade de Sao Paulo, Sao Paulo, 244 pp." type="book" year="2008">Santana 2008</bibRefCitation>
: fig. 47B, C). Male chelipeds very short; palm swollen; fingers long, slightly curved, inner margins denticulate (
<bibRefCitation id="8A014B21FFB9A61FB74FF901FE06F8B7" author="Rathbun" box="[288,451,1780,1807]" firstAuthor="Rathbun" pageId="16" pageNumber="117" pagination="1 - 613" refId="ref24756" refString="Rathbun, M. J. (1925) The spider crabs of America. Bulletin of the U. S. National Museum, 129, 1 - 613. https: // doi. org / 10.5479 / si. 03629236.129. i" type="journal article" year="1925">Rathbun 1925</bibRefCitation>
: pl. 232, fig. 9; 
<bibRefCitation id="8A014B21FFB9A61FB41FF900FCCEF8B7" author="Santana, W. R. A." box="[624,779,1781,1807]" pageId="16" pageNumber="117" refId="ref24839" refString="Santana, W. R. A. (2008) Revisao taxonomica e relacoes filogeneticas em Inachoididae Dana, 1851 (Crustacea, Brachyura, Majoidea). Unpublished doctoral thesis, Universidade de Sao Paulo, Sao Paulo, 244 pp." type="book" year="2008">Santana 2008</bibRefCitation>
: fig. 47A, B). Ambulatory legs very long, spinulous; merus subcylindrical; propodus compressed; dactylus long, slender, compressed, slightly curved (
<bibRefCitation id="8A014B21FFB9A61FB2F6F8EDFAFBF88B" author="Rathbun" box="[1177,1342,1816,1843]" firstAuthor="Rathbun" pageId="16" pageNumber="117" pagination="1 - 613" refId="ref24756" refString="Rathbun, M. J. (1925) The spider crabs of America. Bulletin of the U. S. National Museum, 129, 1 - 613. https: // doi. org / 10.5479 / si. 03629236.129. i" type="journal article" year="1925">Rathbun 1925</bibRefCitation>
: pls. 82, 83, pl. 233, fig. 1; 
<bibRefCitation id="8A014B21FFB9A61FB708F8C9FE32F8EF" author="Guinot" box="[359,503,1852,1879]" firstAuthor="Guinot" pageId="16" pageNumber="117" pagination="377 - 395" refId="ref21724" refString="Guinot, D. (1984) Le genre Leurocyclus Rathbun, 1897 (Crustacea Decapoda Brachyura). Bulletin du Museum national d'Histoire naturelle, ser. 4, 6, sect. A (2), 377 - 395." type="book chapter" year="1984">Guinot 1984</bibRefCitation>
: figs. 3, 4; 
<bibRefCitation id="8A014B21FFB9A61FB41CF8C8FCCBF8EF" author="Santana, W. R. A." box="[627,782,1853,1879]" pageId="16" pageNumber="117" refId="ref24839" refString="Santana, W. R. A. (2008) Revisao taxonomica e relacoes filogeneticas em Inachoididae Dana, 1851 (Crustacea, Brachyura, Majoidea). Unpublished doctoral thesis, Universidade de Sao Paulo, Sao Paulo, 244 pp." type="book" year="2008">Santana 2008</bibRefCitation>
: fig. 47B). Pleon: only first somite dorsal, well developed. Male pleon with somites 1–5 free, somite 6 being fused to telson (pleotelson); female pleon with somites 1–4 free, somites 5, 6 being fused to telson (pleotelson). Male gonopore partially condylar, partially coxal (
<bibRefCitation id="8A014B21FFB9A61FB2A4F871FF06F87B" author="Guinot" etAl="et al." firstAuthor="Guinot" pageId="16" pageNumber="117" pagination="1 - 414" refId="ref22407" refString="Guinot, D., Tavares, M. &amp; Castro, P. (2013) Significance of the sexual openings and supplementary structures on the phylogeny of brachyuran crabs (Crustacea, Decapoda, Brachyura), with new nomina for higher-ranked podotreme taxa. Zootaxa, 3665 (1), 1 - 414. https: // doi. org / 10.11646 / zootaxa. 3665.1.1" type="journal article" year="2013">
Guinot 
<emphasis id="DCE4EAC2FFB9A61FB34FF870FA95F826" box="[1312,1360,1924,1950]" italics="true" pageId="16" pageNumber="117">et al</emphasis>
. 2013: 127
</bibRefCitation>
, fig. 31C). G1 as straight, simple plate, with apical aperture (Garth 1968: pl. T, fig. 9). Vulvae located on anteriorly displaced on sternite 6, not projecting on sternite 5. Penis emerging from coxo-sternal condyle’s extremity.
</paragraph>
</subSubSection>
<subSubSection id="A68A655BFFB8A61EB6A8FF6DFAB4FD12" pageId="17" pageNumber="118" type="discussion">
<paragraph id="EE2F36D0FFB8A61EB6A8FF6DFDE3FF0B" blockId="17.[151,1437,152,683]" box="[199,550,152,179]" pageId="17" pageNumber="118">
<emphasis id="DCE4EAC2FFB8A61EB6A8FF6DFE11FF0B" bold="true" box="[199,468,152,179]" pageId="17" pageNumber="118">Other included genera</emphasis>
. None.
</paragraph>
<paragraph id="EE2F36D0FFB8A61EB6A8FF48FC99FEFA" blockId="17.[151,1437,152,683]" pageId="17" pageNumber="118">
<emphasis id="DCE4EAC2FFB8A61EB6A8FF48FEF1FF6F" bold="true" box="[199,308,189,215]" pageId="17" pageNumber="118">Remarks</emphasis>
. H. 
<bibRefCitation id="8A014B21FFB8A61EB70BFF49FD01FF6F" author="Milne Edwards &amp; Lucas" box="[356,708,188,215]" firstAuthor="Milne Edwards" pageId="17" pageNumber="118" refId="ref23942" refString="Milne Edwards, H. &amp; Lucas, H. (1842 - 1844) Crustaces. In: d'Orbigny, A. (Ed.), Voyage dans l'Amerique meridionale (le Bresil, la Republique orientale de l'Uruguay, la Republique Argentine, la Patagonie, la Republique du Chili, la Republique de Bolivia, la Republique du Perou), execute pendant les annees 1826, 1827, 1828, 1829, 1830, 1831, 1832 et 1833. Tome sixieme. Premiere partie. P. Bertrand, Paris and Vve Levrault, Strasbourg, pls. 1 + 3 (1842: 62); pls. 4 + 6 + 13 (67); pls. 2 + 10 (68); pp. 1 - 8, pls. 5 + 7 + 11 (1843: 69); pp. 9 - 16, pl. 8 bis (71); pp. 17 - 24, pl. 8 (1844: 72); pp. 25 - 32, pl. 7 bis (73); 33 - 39, pl. 9 (74); pls. 14 - 16 (75); pl. 17 (77)] https: // doi. org / 10.5962 / bhl. title. 110540" type="journal volume" year="1842">Milne Edwards &amp; Lucas (1842</bibRefCitation>
: pl. 2, figs. 1–8; 1843: 12, as 
<taxonomicName id="29904D53FFB8A61EB260FF49FB28FF6E" authorityName="H.Milne Edwards &amp; Lucas" authorityYear="1842" box="[1039,1261,188,214]" class="Malacostraca" family="Inachoididae" genus="Salacia" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="118" phylum="Arthropoda" rank="species" species="tuberculosa">
<emphasis id="DCE4EAC2FFB8A61EB260FF49FB28FF6E" box="[1039,1261,188,214]" italics="true" pageId="17" pageNumber="118">Salacia tuberculosa</emphasis>
</taxonomicName>
) based the species on three specimens collected by A. d’Orbigny (see 
<bibRefCitation id="8A014B21FFB8A61EB56EFF15FBC4FF43" author="Guinot &amp; Cleva" box="[769,1025,224,251]" firstAuthor="Guinot" pageId="17" pageNumber="118" pagination="35 - 43" refId="ref22037" refString="Guinot, D. &amp; Cleva, R. (2002 a) D'Orbigny et les Crustaces. In: Taquet, P. (Ed.), Alcide d'Orbigny. Du Nouveau Monde ... au Passe du Monde. Nathan / VUEF et Museum national d'Histoire naturelle, Paris, pp. 35 - 43." type="book chapter" year="2002" yearSuffix="a">Guinot &amp; Cleva 2002a</bibRefCitation>
, 
<bibRefCitation id="8A014B21FFB8A61EB264FF15FB96FF43" author="Guinot &amp; Cleva" box="[1035,1107,224,251]" firstAuthor="Guinot" pageId="17" pageNumber="118" pagination="499 - 515" refId="ref22095" refString="Guinot, D. &amp; Cleva, R. (2002 b) Les Crustaces recoltes par d'Orbigny en Amerique du Sud et deposes au Museum national d'Histoire naturelle, Paris. Palevol, Paris, 1, 499 - 515. https: // doi. org / 10.1016 / S 1631 - 0683 (02) 00055 - 6" type="journal article" year="2002" yearSuffix="b">2002b</bibRefCitation>
) and diagnosed it by a practically circular carapace and a short, narrow rostrum. Both nomina 
<taxonomicName id="29904D53FFB8A61EB51EFEF1FC00FEA6" authorityName="Lamouroux" authorityYear="1816" box="[881,965,260,286]" class="Malacostraca" family="Inachoididae" genus="Salacia" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="118" phylum="Arthropoda" rank="genus">
<emphasis id="DCE4EAC2FFB8A61EB51EFEF1FC00FEA6" box="[881,965,260,286]" italics="true" pageId="17" pageNumber="118">Salacia</emphasis>
</taxonomicName>
and 
<taxonomicName id="29904D53FFB8A61EB595FEF1FBA0FEA6" authorityName="Dana" authorityYear="1851" box="[1018,1125,260,286]" class="Malacostraca" family="Inachoididae" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="118" phylum="Arthropoda" rank="subFamily" subFamily="Salacinae">Salacinae</taxonomicName>
were used by 
<bibRefCitation id="8A014B21FFB8A61EB36AFEF1FED1FEFA" author="Brito Capello" firstAuthor="Brito Capello" pageId="17" pageNumber="118" pagination="262 - 265" refId="ref19798" refString="Brito Capello, F. de (1871) Descripcao de algumas especies novas de crustaceos. Jornal de Sciencias mathematicas, physicas e naturales, Lisboa, 12, 262 - 265 (1 - 4)." type="journal article" year="1871">Brito Capello (1871: 263</bibRefCitation>
, 264, pl. 3, fig. 3, 3a), then not used for a long time.
</paragraph>
<paragraph id="EE2F36D0FFB8A61EB6A8FEB8FB99FDA3" blockId="17.[151,1437,152,683]" pageId="17" pageNumber="118">
An unconstrained analysis of larval characters of 
<taxonomicName id="29904D53FFB8A61EB482FEB9FBCFFEDE" baseAuthorityName="H. Milne Edwards &amp; Lucas" baseAuthorityYear="1842" box="[749,1034,332,358]" class="Malacostraca" family="Inachoididae" genus="Leurocyclus" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="118" phylum="Arthropoda" rank="species" species="tuberculosus">
<emphasis id="DCE4EAC2FFB8A61EB482FEB9FBCFFEDE" box="[749,1034,332,358]" italics="true" pageId="17" pageNumber="118">Leurocyclus tuberculosus</emphasis>
</taxonomicName>
suggested that it was the most basal taxon of 
<taxonomicName id="29904D53FFB8A61EB693FE85FEACFE32" authorityName="MacLeay" authorityYear="1838" box="[252,361,368,394]" class="Malacostraca" family="Inachidae" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="118" phylum="Arthropoda" rank="family">Inachidae</taxonomicName>
+ 
<taxonomicName id="29904D53FFB8A61EB7E8FE85FDDDFE32" authorityName="Drach &amp; Guinot" authorityYear="1983" box="[391,536,368,394]" class="Malacostraca" family="Inachoididae" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="118" phylum="Arthropoda" rank="family">Inachoididae</taxonomicName>
(
<bibRefCitation id="8A014B21FFB8A61EB448FE84FC98FE32" author="Marques &amp; Pohle" box="[551,861,368,395]" firstAuthor="Marques" pageId="17" pageNumber="118" pagination="71 - 82" refId="ref23399" refString="Marques, F. P. L. &amp; Pohle, G. (2003) Searching for larval support for majid subfamilies (Crustacea: Brachyura) with particular reference to Inachoidinae Dana, 1851. Invertebrate Reproduction and Development, 43, 71 - 82. https: // doi. org / 10.1080 / 07924259.2003.9652523" type="journal article" year="2003">
<collectingRegion id="2C54F832FFB8A61EB448FE84FD4EFE33" box="[551,651,369,395]" country="Italy" name="Marche" pageId="17" pageNumber="118">Marques</collectingRegion>
&amp; Pohle 2003: 76
</bibRefCitation>
, figs. 1A, 2). The larval morphology of 
<taxonomicName id="29904D53FFB8A61EB34FFE84FF14FE16" baseAuthorityName="H. Milne Edwards &amp; Lucas" baseAuthorityYear="1842" class="Malacostraca" family="Inachoididae" genus="Leurocyclus" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="118" phylum="Arthropoda" rank="species" species="tuberculosus">
<emphasis id="DCE4EAC2FFB8A61EB34FFE84FF14FE16" italics="true" pageId="17" pageNumber="118">L. tuberculosus</emphasis>
</taxonomicName>
conforms to the general pattern found in 
<taxonomicName id="29904D53FFB8A61EB4C0FE61FCD9FE16" authorityName="Samouelle" authorityYear="1819" box="[687,796,404,430]" class="Malacostraca" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="103" phylum="Arthropoda" rank="superFamily" superFamily="Majoidea">Majoidea</taxonomicName>
, but the two zoeal stages and the megalopa differs from those of all known inachoidids by a series of characters (
<bibRefCitation id="8A014B21FFB8A61EB54DFE4CFBF4FE6B" author="Marques &amp; Pohle" box="[802,1073,440,467]" firstAuthor="Marques" pageId="17" pageNumber="118" pagination="71 - 82" refId="ref23399" refString="Marques, F. P. L. &amp; Pohle, G. (2003) Searching for larval support for majid subfamilies (Crustacea: Brachyura) with particular reference to Inachoidinae Dana, 1851. Invertebrate Reproduction and Development, 43, 71 - 82. https: // doi. org / 10.1080 / 07924259.2003.9652523" type="journal article" year="2003">
<collectingRegion id="2C54F832FFB8A61EB54DFE4CFC43FE6B" box="[802,902,441,467]" country="Italy" name="Marche" pageId="17" pageNumber="118">Marques</collectingRegion>
&amp; Pohle 2003
</bibRefCitation>
; 
<bibRefCitation id="8A014B21FFB8A61EB251FE4CFAA3FE6B" author="Santana &amp; Marques" box="[1086,1382,441,467]" firstAuthor="Santana" pageId="17" pageNumber="118" pagination="49 - 58" refId="ref24883" refString="Santana, W. R. &amp; Marques, F. P. L. (2009) Larval morphology of the spider crab Leurocyclus tuberculosus (Decapoda: Majoidea: Inachoididae). Nauplius, 17, 49 - 58." type="journal article" year="2009">
Santana &amp; 
<collectingRegion id="2C54F832FFB8A61EB2D1FE4CFAE5FE6B" box="[1214,1312,441,467]" country="Italy" name="Marche" pageId="17" pageNumber="118">Marques</collectingRegion>
2009
</bibRefCitation>
). All this supports the separation of the genus in its own subfamily. 
<taxonomicName id="29904D53FFB8A61EB536FE29FBB2FE4E" baseAuthorityName="H. Milne Edwards &amp; Lucas" baseAuthorityYear="1842" box="[857,1143,476,502]" class="Malacostraca" family="Inachoididae" genus="Leurocyclus" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="118" phylum="Arthropoda" rank="species" species="tuberculosus">
<emphasis id="DCE4EAC2FFB8A61EB536FE29FBB2FE4E" box="[857,1143,476,502]" italics="true" pageId="17" pageNumber="118">Leurocyclus tuberculosus</emphasis>
</taxonomicName>
, as other inachoidids, is a determinate growth crab, but growth patterns differ beween the sexes (
<bibRefCitation id="8A014B21FFB8A61EB5C2FDF4FB95FDA3" author="MacLay" box="[941,1104,512,539]" firstAuthor="MacLay" pageId="17" pageNumber="118" pagination="245 - 316" refId="ref23113" refString="MacLay, C. L. (2015) Chapter 71 - 5. Moulting and growth. In: Castro, P., Davie, P. J. F., Guinot, D., Schram, F. R. &amp; von Vaupel Klein, J. C. (Eds.), Treatise on Zoology-Anatomy, Taxonomy, Biology. The Crustacea. Vol. 9. Part C-I. Brill, Leiden and Boston, pp. 245 - 316." type="book chapter" year="2015">MacLay 2015</bibRefCitation>
).
</paragraph>
<paragraph id="EE2F36D0FFB8A61EB6A8FDD1FAB4FD12" blockId="17.[151,1437,152,683]" pageId="17" pageNumber="118">
The biology of 
<taxonomicName id="29904D53FFB8A61EB71CFDD1FDB8FD86" box="[371,637,548,574]" class="Malacostraca" family="Inachoididae" genus="Leurocyclus" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="118" phylum="Arthropoda" rank="species" species="granulosus">
<emphasis id="DCE4EAC2FFB8A61EB71CFDD1FDB8FD86" box="[371,637,548,574]" italics="true" pageId="17" pageNumber="118">Leurocyclus granulosus</emphasis>
</taxonomicName>
is well studied (
<bibRefCitation id="8A014B21FFB8A61EB55DFDD0FC32FD87" author="Baron" box="[818,1015,548,575]" etAl="et al." firstAuthor="Baron" pageId="17" pageNumber="118" pagination="267 - 273" refId="ref19474" refString="Baron, P. J., Quiroga, P., Leal, G. &amp; Gonzalez-Pisani, X. (2009) Morphological maturity of the spider crab Leurocyclus tuberculosus (H. Milne Edwards and Lucas, 1842) (Majidae, Brachyura) in the northern Patagonian gulfs. Crustaceana, 82 (3), 267 - 273. https: // doi. org / 10.1163 / 156854008 X 390407" type="journal article" year="2009">
Barón 
<emphasis id="DCE4EAC2FFB8A61EB511FDD0FC6AFD86" box="[894,943,548,574]" italics="true" pageId="17" pageNumber="118">et al</emphasis>
. 2009
</bibRefCitation>
; 
<bibRefCitation id="8A014B21FFB8A61EB26DFDD1FB1FFD87" author="Stauffer" box="[1026,1242,548,575]" etAl="et al." firstAuthor="Stauffer" pageId="17" pageNumber="118" pagination="55 - 61" refId="ref25276" refString="Stauffer, T., Ostrovski, M. C., Goncalves da Silva-Ferreira, T. C. &amp; Costa, T. (2011) Biology of the crab Leurocyclus tuberculosus (H. Milne Edwards &amp; Lucas, 1843) by catch from pink shrimp trawl fishery in the coast of Rio de Janeiro, Brazil. Nauplius, 19 (1), 55 - 61. https: // doi. org / 10.1590 / S 0104 - 64972011000100006" type="journal article" year="2011">
Stauffer 
<emphasis id="DCE4EAC2FFB8A61EB20DFDD0FB5FFD86" box="[1122,1178,548,574]" italics="true" pageId="17" pageNumber="118">et al.</emphasis>
2011
</bibRefCitation>
; 
<bibRefCitation id="8A014B21FFB8A61EB28AFDD1FED6FDDB" author="Gonzalez-Pisani" etAl="et al." firstAuthor="Gonzalez-Pisani" pageId="17" pageNumber="118" pagination="61 - 74" refId="ref21590" refString="Gonzalez-Pisani, X., Baron, P. J. &amp; Lopez Greco, L. S. (2011) Functional anatomy of the female reproductive systems of two spider crabs (Decapoda, Majoidea). Invertebrate Biology, 131 (1), 61 - 74. https: // doi. org / 10.1111 / j. 1744 - 7410.2011.00252. x" type="journal article" year="2011">
González-Pisani 
<emphasis id="DCE4EAC2FFB8A61EB6F8FDBCFF0FFDDA" box="[151,202,584,610]" italics="true" pageId="17" pageNumber="118">et al</emphasis>
. 2011
</bibRefCitation>
, 
<bibRefCitation id="8A014B21FFB8A61EB770FDBCFE99FDDB" author="Gonzalez-Pisani" box="[287,348,585,611]" firstAuthor="Gonzalez-Pisani" pageId="17" pageNumber="118" pagination="473 - 483" refId="ref21657" refString="Gonzalez-Pisani, X., Baron, P. J. &amp; Lopez Greco, L. S. (2017) Integrated analysis of sexual maturation through successive growth instars in the spider crab Leurocyclus tuberculosus (Decapoda: Majoidea). Canadian Journal of Zoology, 95, 473 - 483. https: // doi. org / 10.1139 / cjz- 2016 - 0034" type="journal article" year="2017">2017</bibRefCitation>
). According to 
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González-Pisani 
<emphasis id="DCE4EAC2FFB8A61EB4A5FDBCFCC6FDDA" box="[714,771,584,610]" italics="true" pageId="17" pageNumber="118">et al.</emphasis>
(2017)
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, the integrated analysis of size at maturity and size frequency distributions showed that in both sexes molt to gonadal, morphological, morphometric, and functional sexual maturity occurred in advance of the terminal molt, in contrast with patterns observed in other 
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.
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