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<document ID-DOI="10.37520/fi.2022.016" ID-ISSN="2533-4069" ID-Zenodo-Dep="7522801" approvalRequired="105" approvalRequired_for_taxonomicNames="25" approvalRequired_for_textStreams="59" approvalRequired_for_treatments="21" checkinTime="1673382488781" checkinUser="felipe" docAuthor="Friis, Else Marie, Crane, Peter R., Pedersen, Kaj Raunsgaard, Mendes, Mário Miguel &amp; Kvaček, Jiří" docDate="2022" docId="03FD87F2FFD5FFC6FCCFFD81C77EF91D" docLanguage="en" docName="FossilImprint.78.2.341-424.pdf" docOrigin="Fossil Imprint 78 (2)" docSource="http://dx.doi.org/10.37520/fi.2022.016" docStyle="DocumentStyle:0EC54BF35E227CA4627F894F6BA30634.2:FossilImprint.2016-.journal_article" docStyleId="0EC54BF35E227CA4627F894F6BA30634" docStyleName="FossilImprint.2016-.journal_article" docStyleVersion="2" docTitle="Endressistemon cateficensis E. M. FRIIS, P. R. CRANE, K. R. PEDERSEN, M. M. MENDES et J. KVACEK 2022, sp. nov." docType="treatment" docVersion="2" lastPageNumber="381" masterDocId="FFC4FF8AFFFDFFEEFFBFFFFFC378FFA6" masterDocTitle="The Early Cretaceous Mesofossil Flora Of Catefica, Portugal: Angiosperms" masterLastPageNumber="424" masterPageNumber="341" pageNumber="381" updateTime="1673383361075" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:title>The Early Cretaceous Mesofossil Flora Of Catefica, Portugal: Angiosperms</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Friis, Else Marie</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Crane, Peter R.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Pedersen, Kaj Raunsgaard</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Mendes, Mário Miguel</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kvaček, Jiří</mods:namePart>
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<mods:title>Fossil Imprint</mods:title>
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<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-12-20</mods:number>
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<mods:detail type="volume">
<mods:number>78</mods:number>
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<mods:number>2</mods:number>
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<mods:start>341</mods:start>
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<mods:url>http://dx.doi.org/10.37520/fi.2022.016</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.37520/fi.2022.016</mods:identifier>
<mods:identifier type="ISSN">2533-4069</mods:identifier>
<mods:identifier type="Zenodo-Dep">7522801</mods:identifier>
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<treatment LSID="urn:lsid:plazi:treatment:03FD87F2FFD5FFC6FCCFFD81C77EF91D" httpUri="http://treatment.plazi.org/id/03FD87F2FFD5FFC6FCCFFD81C77EF91D" lastPageNumber="381" pageId="40" pageNumber="381">
<subSubSection pageId="40" pageNumber="381" type="nomenclature">
<paragraph blockId="40.[880,1444,638,695]" pageId="40" pageNumber="381">
<heading centered="true" fontSize="8" level="2" pageId="40" pageNumber="381" reason="2">
<emphasis bold="true" pageId="40" pageNumber="381">
<taxonomicName authority="E.M.FRIIS, P.R.CRANE, K.R.PEDERSEN, M.M.MENDES et J.KVACEK" authorityName="E. M. FRIIS, P. R. CRANE, K. R. PEDERSEN, M. M. MENDES et J. KVACEK" authorityYear="2022" class="Magnoliopsida" family="Platanaceae" genus="Endressistemon" kingdom="Plantae" order="Proteales" pageId="40" pageNumber="381" phylum="Tracheophyta" rank="species" species="cateficensis" status="sp. nov.">
<emphasis bold="true" box="[880,1188,638,662]" italics="true" pageId="40" pageNumber="381">Endressistemon cateficensis</emphasis>
E.M.FRIIS, P.R.CRANE, K.R.PEDERSEN, M.M.MENDES et J.KVAČEK
</taxonomicName>
<taxonomicNameLabel box="[1354,1439,671,695]" pageId="40" pageNumber="381" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph blockId="40.[1081,1244,704,728]" box="[1081,1244,704,728]" pageId="40" pageNumber="381">
<figureCitation box="[1081,1244,704,728]" captionStart="Text-fig" captionStartId="45.[125,206,1434,1455]" captionTargetBox="[125,1440,151,1415]" captionTargetId="figure-130@45.[122,1443,148,1421]" captionTargetPageId="45" captionText="Text-fig. 28. Scanning electron microscope (SEM) images of stamens and pollen of Endressistemon cateficensis gen. et sp. nov.; Catefica locality, Portugal. a, b) Staminal structure with two lateral stamens and one median structure seen in ventral and dorsal view (orientation unknown) showing that each stamen has a prominent apical extension and two pairs of pollen sacs separated by a narrow connective; both stamens are borne on a common base together with the median structure and their anthers are sessile on the common stalk; note the apical projection of the median structure (asterisk) between the two stamens and the ribs over probable vascular bundles that extend from the common base into the apical projections of both stamens and the median structure (arrows); c) Staminal structure showing two stamens with prominent apical projections and median axis-like structure between the two stamens (arrow); d) Staminal structure showing two stamens with prominent apical projections borne on a common base; e) Staminal structure in (a) and (b), from the same orientation as (b), showing the ribs over probable vascular bundles (yellow) that extend into the apical projections; f) Detail of staminal structure in (a) and (b) showing the fused or strongly adhering apical projections of the two lateral stamens and the median structure (asterisk); g) Monocolpate, reticulate pollen grains from the pollen sacs of stamen in (d). Specimens, Catefica 49-S107778 (holotype, a, b, e, f), Catefica 49-S107769 (c), Catefica 49-S107751 (d, g). Scale bars = 600 Μm (ad), 300 Μm (e), 100 Μm (f), 6 Μm (g)." figureDoi="http://doi.org/10.5281/zenodo.7522859" httpUri="https://zenodo.org/record/7522859/files/figure.png" pageId="40" pageNumber="381">Text-fig. 28ag</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="40" pageNumber="381" type="description">
<paragraph blockId="40.[843,1482,751,808]" box="[881,1482,751,776]" pageId="40" pageNumber="381">H o l o t y p e. S107778 (Catefica sample 49; figured</paragraph>
<paragraph blockId="40.[843,1482,751,808]" box="[843,1069,784,808]" pageId="40" pageNumber="381">
<figureCitation box="[843,1056,784,808]" captionStart="Text-fig" captionStartId="45.[125,206,1434,1455]" captionTargetBox="[125,1440,151,1415]" captionTargetId="figure-130@45.[122,1443,148,1421]" captionTargetPageId="45" captionText="Text-fig. 28. Scanning electron microscope (SEM) images of stamens and pollen of Endressistemon cateficensis gen. et sp. nov.; Catefica locality, Portugal. a, b) Staminal structure with two lateral stamens and one median structure seen in ventral and dorsal view (orientation unknown) showing that each stamen has a prominent apical extension and two pairs of pollen sacs separated by a narrow connective; both stamens are borne on a common base together with the median structure and their anthers are sessile on the common stalk; note the apical projection of the median structure (asterisk) between the two stamens and the ribs over probable vascular bundles that extend from the common base into the apical projections of both stamens and the median structure (arrows); c) Staminal structure showing two stamens with prominent apical projections and median axis-like structure between the two stamens (arrow); d) Staminal structure showing two stamens with prominent apical projections borne on a common base; e) Staminal structure in (a) and (b), from the same orientation as (b), showing the ribs over probable vascular bundles (yellow) that extend into the apical projections; f) Detail of staminal structure in (a) and (b) showing the fused or strongly adhering apical projections of the two lateral stamens and the median structure (asterisk); g) Monocolpate, reticulate pollen grains from the pollen sacs of stamen in (d). Specimens, Catefica 49-S107778 (holotype, a, b, e, f), Catefica 49-S107769 (c), Catefica 49-S107751 (d, g). Scale bars = 600 Μm (ad), 300 Μm (e), 100 Μm (f), 6 Μm (g)." figureDoi="http://doi.org/10.5281/zenodo.7522859" httpUri="https://zenodo.org/record/7522859/files/figure.png" pageId="40" pageNumber="381">Text-fig. 28a, b, e, f</figureCitation>
).
</paragraph>
<paragraph blockId="40.[843,1482,831,888]" box="[881,1482,831,856]" pageId="40" pageNumber="381">P l a n t F o s s i l N a m e s R e g i s t r y N u m b e r.</paragraph>
<paragraph blockId="40.[843,1482,831,888]" box="[843,1169,864,888]" pageId="40" pageNumber="381">PFN002797 (for new species).</paragraph>
<paragraph blockId="40.[843,1481,911,968]" box="[881,1481,911,936]" pageId="40" pageNumber="381">P a r a t y p e s. S107751, S107768, S107769, S266022</paragraph>
<paragraph blockId="40.[843,1481,911,968]" box="[843,1071,944,968]" pageId="40" pageNumber="381">(Catefica sample 49).</paragraph>
<paragraph blockId="40.[843,1482,992,1081]" pageId="40" pageNumber="381">
R e p o s i t o r y. Palaeobotanical Collections, Department of Palaeobiology, Swedish Museum of Natural History,
<collectingRegion box="[843,961,1057,1081]" country="Sweden" name="Stockholm" pageId="40" pageNumber="381">Stockholm</collectingRegion>
,
<collectingCountry box="[972,1056,1057,1081]" name="Sweden" pageId="40" pageNumber="381">Sweden</collectingCountry>
(S).
</paragraph>
<paragraph blockId="40.[843,1482,1104,1161]" pageId="40" pageNumber="381">E t y m o l o g y. From the Catefica locality where the fossils were collected.</paragraph>
<paragraph blockId="40.[843,1481,1183,1273]" pageId="40" pageNumber="381">
<materialsCitation country="Portugal" county="Torres Vedras" latitude="39.058334" location="Catefica" longLatPrecision="19" longitude="-9.241667" municipality="Mugideira" pageId="40" pageNumber="381" specimenCount="1">
Ty p e l o c a l i t y. Catefica (
<geoCoordinate box="[1184,1322,1183,1208]" degrees="39" direction="north" minutes="03" orientation="latitude" pageId="40" pageNumber="381" precision="15" seconds="30" value="39.058334">39° 03ʹ30ʺ N</geoCoordinate>
;
<geoCoordinate box="[1330,1469,1183,1208]" degrees="09" direction="west" minutes="14" orientation="longitude" pageId="40" pageNumber="381" precision="15" seconds="30" value="-9.241667">09°14ʹ 30ʺ W</geoCoordinate>
), between the villages of
<location LSID="urn:lsid:plazi:treatment:03FD87F2FFD5FFC6FCCFFD81C77EF91D:8E8B603FFFD5FFC6FBF6FB3EC7D8FB7F" box="[1097,1184,1217,1241]" country="Portugal" county="Torres Vedras" latitude="39.058334" longLatPrecision="19" longitude="-9.241667" municipality="Mugideira" name="Catefica" pageId="40" pageNumber="381">Catefica</location>
and
<collectingMunicipality box="[1236,1350,1217,1241]" pageId="40" pageNumber="381">Mugideira</collectingMunicipality>
, about
<quantity box="[1427,1481,1217,1241]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="40" pageNumber="381" unit="km" value="4.0">4 km</quantity>
south of
<collectingCounty box="[936,1085,1249,1273]" pageId="40" pageNumber="381">Torres Vedras</collectingCounty>
,
<collectingCountry box="[1095,1189,1249,1273]" name="Portugal" pageId="40" pageNumber="381">Portugal</collectingCountry>
</materialsCitation>
.
</paragraph>
<paragraph blockId="40.[843,1482,1297,1353]" box="[881,1482,1297,1321]" pageId="40" pageNumber="381">T y p e s t r a t u m a n d a g e. Almargem Formation,</paragraph>
<paragraph blockId="40.[843,1482,1297,1353]" box="[843,1270,1329,1353]" pageId="40" pageNumber="381">Early Cretaceous (Aptian-early Albian).</paragraph>
<paragraph blockId="40.[881,1366,1377,1401]" box="[881,1366,1377,1401]" pageId="40" pageNumber="381">S p e c i f i c d i a g n o s i s.As for the genus.</paragraph>
<paragraph blockId="40.[843,1482,1425,1481]" box="[881,1482,1425,1449]" pageId="40" pageNumber="381">
D i m e n s i o n s. Stamens up to about
<quantity box="[1300,1381,1425,1449]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="40" pageNumber="381" unit="mm" value="0.9">0.9 mm</quantity>
long and
</paragraph>
<paragraph blockId="40.[843,1482,1425,1481]" box="[843,1027,1457,1481]" pageId="40" pageNumber="381">
<quantity box="[843,924,1457,1481]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="40" pageNumber="381" unit="mm" value="0.2">0.2 mm</quantity>
in broad.
</paragraph>
<paragraph blockId="40.[843,1482,1505,1919]" pageId="40" pageNumber="381">
D e s c r i p t i o n a n d r e m a r k s. The material includes isolated staminal structures each consisting of two lateral stamens and in some specimens a median axislike structure. The two stamens and axis-like structure are borne on a short, common stalk (
<figureCitation box="[1294,1470,1635,1659]" captionStart="Text-fig" captionStartId="45.[125,206,1434,1455]" captionTargetBox="[125,1440,151,1415]" captionTargetId="figure-130@45.[122,1443,148,1421]" captionTargetPageId="45" captionText="Text-fig. 28. Scanning electron microscope (SEM) images of stamens and pollen of Endressistemon cateficensis gen. et sp. nov.; Catefica locality, Portugal. a, b) Staminal structure with two lateral stamens and one median structure seen in ventral and dorsal view (orientation unknown) showing that each stamen has a prominent apical extension and two pairs of pollen sacs separated by a narrow connective; both stamens are borne on a common base together with the median structure and their anthers are sessile on the common stalk; note the apical projection of the median structure (asterisk) between the two stamens and the ribs over probable vascular bundles that extend from the common base into the apical projections of both stamens and the median structure (arrows); c) Staminal structure showing two stamens with prominent apical projections and median axis-like structure between the two stamens (arrow); d) Staminal structure showing two stamens with prominent apical projections borne on a common base; e) Staminal structure in (a) and (b), from the same orientation as (b), showing the ribs over probable vascular bundles (yellow) that extend into the apical projections; f) Detail of staminal structure in (a) and (b) showing the fused or strongly adhering apical projections of the two lateral stamens and the median structure (asterisk); g) Monocolpate, reticulate pollen grains from the pollen sacs of stamen in (d). Specimens, Catefica 49-S107778 (holotype, a, b, e, f), Catefica 49-S107769 (c), Catefica 49-S107751 (d, g). Scale bars = 600 Μm (ad), 300 Μm (e), 100 Μm (f), 6 Μm (g)." figureDoi="http://doi.org/10.5281/zenodo.7522859" httpUri="https://zenodo.org/record/7522859/files/figure.png" pageId="40" pageNumber="381">Text-fig. 28ae</figureCitation>
). The anthers are basifixed, tetrasporangiate and dithecate (
<figureCitation box="[850,1017,1699,1724]" captionStart="Text-fig" captionStartId="45.[125,206,1434,1455]" captionTargetBox="[125,1440,151,1415]" captionTargetId="figure-130@45.[122,1443,148,1421]" captionTargetPageId="45" captionText="Text-fig. 28. Scanning electron microscope (SEM) images of stamens and pollen of Endressistemon cateficensis gen. et sp. nov.; Catefica locality, Portugal. a, b) Staminal structure with two lateral stamens and one median structure seen in ventral and dorsal view (orientation unknown) showing that each stamen has a prominent apical extension and two pairs of pollen sacs separated by a narrow connective; both stamens are borne on a common base together with the median structure and their anthers are sessile on the common stalk; note the apical projection of the median structure (asterisk) between the two stamens and the ribs over probable vascular bundles that extend from the common base into the apical projections of both stamens and the median structure (arrows); c) Staminal structure showing two stamens with prominent apical projections and median axis-like structure between the two stamens (arrow); d) Staminal structure showing two stamens with prominent apical projections borne on a common base; e) Staminal structure in (a) and (b), from the same orientation as (b), showing the ribs over probable vascular bundles (yellow) that extend into the apical projections; f) Detail of staminal structure in (a) and (b) showing the fused or strongly adhering apical projections of the two lateral stamens and the median structure (asterisk); g) Monocolpate, reticulate pollen grains from the pollen sacs of stamen in (d). Specimens, Catefica 49-S107778 (holotype, a, b, e, f), Catefica 49-S107769 (c), Catefica 49-S107751 (d, g). Scale bars = 600 Μm (ad), 300 Μm (e), 100 Μm (f), 6 Μm (g)." figureDoi="http://doi.org/10.5281/zenodo.7522859" httpUri="https://zenodo.org/record/7522859/files/figure.png" pageId="40" pageNumber="381">Text-fig. 28a, b</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>