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<document id="755D5B375819DA86A93E2CDD553AAF5E" ID-DOI="10.5281/zenodo.157800" ID-GBIF-Dataset="94266e3f-2fa2-45bc-8344-98e7b4f0c559" ID-ISSN="1175­5326" ID-Zenodo-Dep="157800" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461018378539" checkinUser="plazi" docAuthor="Valerio, A. A., Deans, A. R. &amp; Whitfield, J. B." docDate="2004" docId="03FD293BFF8C9462137954FBA9968B63" docLanguage="en" docName="zt00526.pdf" docOrigin="Zootaxa 526" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Exoryza schoenobii Wilkinson" docType="treatment" docVersion="8" lastPageNumber="8" masterDocId="FFC45143FF89946512715607AB53885D" masterDocTitle="Review of the microgastrine braconid wasp genus Exoryza, with description of a new species, E. monocavus, from Central America" masterLastPageNumber="11" masterPageNumber="1" pageNumber="6" updateTime="1698569481161" updateUser="plazi">
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<mods:title id="9AD288F91BAD2705435FFEE75C8CF900">Review of the microgastrine braconid wasp genus Exoryza, with description of a new species, E. monocavus, from Central America</mods:title>
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<mods:namePart id="FEE2648A73808196B09EC798BE5CB0D4">Valerio, A. A.</mods:namePart>
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<mods:namePart id="B48411695306C754EF97D75102A4255A">Deans, A. R.</mods:namePart>
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<mods:namePart id="334F72F6F193B1255C52D3138784F430">Whitfield, J. B.</mods:namePart>
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<mods:date id="2003214D989C172EFD14D8BE01384536">2004</mods:date>
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<treatment id="03FD293BFF8C9462137954FBA9968B63" ID-DOI="http://doi.org/10.5281/zenodo.6272669" ID-GBIF-Taxon="119602026" ID-Zenodo-Dep="6272669" LSID="urn:lsid:plazi:treatment:03FD293BFF8C9462137954FBA9968B63" httpUri="http://treatment.plazi.org/id/03FD293BFF8C9462137954FBA9968B63" lastPageId="7" lastPageNumber="8" pageId="5" pageNumber="6">
<subSubSection id="C34ECBA6FF8C9460137954FBAAF58B64" pageId="5" pageNumber="6" type="nomenclature">
<paragraph id="8BEB982DFF8C9460137954FBA9C38B4B" blockId="5.[264,656,764,825]" box="[264,656,764,790]" pageId="5" pageNumber="6">
<heading id="D0A32F41FF8C9460137954FBA9C38B4B" bold="true" box="[264,656,764,790]" fontSize="11" level="1" pageId="5" pageNumber="6" reason="1">
<taxonomicName id="4C54E3AEFF8C9460137954FBA9C38B4B" authority="Wilkinson" authorityName="Wilkinson" box="[264,656,764,790]" class="Insecta" family="Braconidae" genus="Exoryza" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="schoenobii">
<emphasis id="B920443FFF8C9460137954FBA9C38B4B" bold="true" box="[264,656,764,790]" pageId="5" pageNumber="6">
<emphasis id="B920443FFF8C9460137954FBAAA68B4B" bold="true" box="[264,501,764,790]" italics="true" pageId="5" pageNumber="6">Exoryza schoenobii</emphasis>
(Wilkinson)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BEB982DFF8C946013795518AAF58B64" blockId="5.[264,656,764,825]" box="[264,422,799,825]" pageId="5" pageNumber="6">
(
<figureCitation id="136F84A8FF8C946013605518AAD38B64" box="[273,384,799,825]" captionStart="FIGURES 1 5. 1" captionStartId="2.[264,383,1652,1676]" captionTargetBox="[331,1267,471,1639]" captionTargetId="figure@2.[306,1314,454,1644]" captionTargetPageId="2" captionText="FIGURES 1 5. 1: Exoryza monocavus, female, lateral habitus; 2,3: Forewing 1 ­ CU of E. minnesota and E. schoenobii, respectively; 4,5: Mesopleura of E. minnesota and E. schoenobii, respectively. Scale = 0.5 mm." httpUri="https://zenodo.org/record/157801/files/figure.png" pageId="5" pageNumber="6">Figs. 3, 5</figureCitation>
,
<figureCitation id="136F84A8FF8C946013FC5518AACF8B64" box="[397,412,799,825]" captionStart="FIGURE 6 8" captionStartId="4.[264,368,1102,1126]" captionTargetBox="[513,1046,337,1071]" captionTargetId="figure@4.[505,1081,329,1076]" captionTargetPageId="4" captionText="FIGURE 6 8. Metasomal tergites I III of: 6, Exoryza minnesota, 7, E. schoenobii, 8, E. monocavus. Scale = 0.5 mm." httpUri="https://zenodo.org/record/157802/files/figure.png" pageId="5" pageNumber="6">7</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34ECBA6FF8C946013795568AFCA8BF8" pageId="5" pageNumber="6" type="reference_group">
<paragraph id="8BEB982DFF8C946013795568A8198BD8" blockId="5.[264,1177,878,934]" box="[264,842,878,902]" pageId="5" pageNumber="6">
<taxonomicName id="4C54E3AEFF8C946013795568A9AA8BD8" ID-CoL="F7Y3" authority="Wilkinson, 1932 Stylops" authorityName="Wilkinson, 1932 Stylops" box="[264,761,878,902]" class="Insecta" family="Braconidae" genus="Apanteles" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="schoenobii">
<emphasis id="B920443FFF8C946013795568AABF8BDB" box="[264,492,879,902]" italics="true" pageId="5" pageNumber="6">Apanteles schoenobii</emphasis>
<bibRefCitation id="EFC5E5DCFF8C946013835569A9F18BD8" author="Wilkinson" box="[498,674,878,901]" pageId="5" pageNumber="11" refString="Wilkinson, D. S. (1932) Four new Apanteles (Hym. Brac.) Stylops, 1, 139 - 144." type="journal article" year="1932">Wilkinson, 1932</bibRefCitation>
Stylops
</taxonomicName>
1: 142.
</paragraph>
<paragraph id="8BEB982DFF8C946013795588AFCA8BF8" blockId="5.[264,1177,878,934]" box="[264,1177,910,934]" pageId="5" pageNumber="6">
<taxonomicName id="4C54E3AEFF8C946013795588A9388BF8" authority="Wiilkinson" authorityName="Wiilkinson" box="[264,619,910,934]" class="Insecta" family="Braconidae" genus="Exoryza" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="schoenobii">
<emphasis id="B920443FFF8C946013795588AA8A8BFB" box="[264,473,911,934]" italics="true" pageId="5" pageNumber="6">Exoryza schoenobii</emphasis>
(Wiilkinson)
</taxonomicName>
:
<bibRefCitation id="EFC5E5DCFF8C946010045589A9AE8BF8" author="Mason" box="[629,765,910,933]" pageId="5" pageNumber="10" refString="Mason, W. R. M. (1981) The polyphyletic nature of Apanteles Foerster (Hymenoptera: Braconidae): A phylogeny and reclassification of Microgastrinae. Memoirs of the Entomological Society of Canada, 115, 1 - 147." type="journal article" year="1981">Mason,1981</bibRefCitation>
, Mem. Entomol. Soc.
<collectingCountry id="F343D8BDFF8C9460119D5589AF6F8BF8" box="[1004,1084,910,933]" name="Canada" pageId="5" pageNumber="6">Canada</collectingCountry>
115: 40.
</paragraph>
</subSubSection>
<subSubSection id="C34ECBA6FF8C9463137955D1A9C38E84" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6" type="description">
<paragraph id="8BEB982DFF8C9460137955D1A9D08BAC" blockId="5.[264,1324,982,1894]" box="[264,643,982,1009]" pageId="5" pageNumber="6">
<emphasis id="B920443FFF8C9460137955D1AA308BAD" bold="true" box="[264,355,982,1008]" pageId="5" pageNumber="6">Female</emphasis>
. Body length =
<quantity id="4CAC35C8FF8C9460105055D0A92D8BAC" box="[545,638,983,1009]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="5" pageNumber="6" unit="mm" value="3.7">3.7 mm</quantity>
.
</paragraph>
<paragraph id="8BEB982DFF8C9460134955F9AFE38D53" blockId="5.[264,1324,982,1894]" pageId="5" pageNumber="6">
<emphasis id="B920443FFF8C9460134955F9AAEC8C45" bold="true" box="[312,447,1022,1048]" pageId="5" pageNumber="6">Body color</emphasis>
. Antenna brownish yellow as are hind coxae, palpus, distal edge of metasomal 3rd to 6th tergites, ovipositor and lateral areas of metasoma; hind femur distal tip dark brown as are tarsomeres and distal portion of hind tibia; hypopygium yellow as are most of legs; hind tibial spurs whitish yellow; compound eyes silver; ocelli yellow; mandible (except distal tips dark brown) and labrum light brown; palpus yellow as are fore leg telotarsus, distal edge of 3rd to 6th metasomal terga. Wings hyaline; forewing veins brownish except base of M+CU and basal 1/3 of 11A; hind wing veins brownish yellow.
</paragraph>
<paragraph id="8BEB982DFF8C94601349531BA81B8E13" blockId="5.[264,1324,982,1894]" pageId="5" pageNumber="6">
<emphasis id="B920443FFF8C94601349531BAA288D6B" bold="true" box="[312,379,1308,1334]" pageId="5" pageNumber="6">Head</emphasis>
. Head height/width = 1.2; compound eye height/width = 1.7; intertentorial pit distance/distance from tentorial pit to compound eye = 3.2; width face at dorsal clypeal edge =
<quantity id="4CAC35C8FF8C94601310536BAA9C8DDB" box="[353,463,1388,1414]" metricMagnitude="-4" metricUnit="m" metricValue="4.4" pageId="5" pageNumber="6" unit="mm" value="0.44">0.44 mm</quantity>
; clypeus width/height = 3; vertex width/distance between anterior ocelli and edge of torulus = 2.6; first flagellomere length/width = 1.0; length of first flagellomere/length second flagellomere = 1.0; length of first flagellomere/length of third flagellomere = 1.1; distal flagellomere length/subdistal flagellomere length = 0.14; distal flagellomere length/width = 0.25; malar space height/basal width of mandible = 0.46; ocell­ocular distance/lateral ocelli distance = 1.8.
</paragraph>
<paragraph id="8BEB982DFF8C94601349505BAA208EB3" blockId="5.[264,1324,982,1894]" pageId="5" pageNumber="6">Face and clypeus with dense and fine punctate sculpturing, less evident and less dense on clypeus than on face, remainder of head with same sculpturing; ocell­ocular area with more widely spaced and less evident punctate sculpturing; frons with most of scrobal areas punctate.</paragraph>
<paragraph id="8BEB982DFF8C9463134950FBAFFF8C23" blockId="5.[264,1324,982,1894]" lastBlockId="6.[264,1325,284,1833]" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
<emphasis id="B920443FFF8C9460134950FBAAEF8F4B" bold="true" box="[312,444,1788,1814]" pageId="5" pageNumber="6">Mesosoma</emphasis>
. Mesosoma length/width = 1.6. Propleuron anteriorly with confused foveate sculpturing, distally with less dense and larger punctations than remainder of propleuron, dorsal area of medial half with confused punctate sculpturing; pronotum anteriorly with sparse large transverse ridges, lateroventral groove with well­defined but miniature scrobiculate sculpturing, upper groove appearing as a well­defined nitid depression, area between lateral grooves with sparse confused punctate sculpturing; mesonotum with punctate sculpturing which becomes less dense towards scutellar groove; scutellar groove with 10 well defined costulae of approximately same width, medial ones slightly longer than lateral ones; scutellum with tiny shallow punctations, lateral areas of scutellum with narrow transverse costulae distal tip less sculptured than rest of scutellum; axilla through mesonotum with transverse costulate sculpturing on anterior edge of lunulae, remainder nitid; mesopleuron almost totally covered with punctate sculpturing except sternaulus with thick transverse sculpturing, sternaulus appearing as an elongated depression that runs at least 1/3 of mesopleuron length, dorsal edge with well defined and dense sculpturing on medial 1/3, but upper part distally almost completely nitid, anterior 1/3 of dorsal edge with confused and shallow scrobiculate sculpturing; metanotum with anterior 1/3 nitid as on distal edge, remainder with two transverse ridges and conspicuously depressed; metapleuron with anterior 1/3 with punctate sculpturing, medial pit present, distal 2/3 with fine and confused rugulose sculpturing mixed with shallow punctate sculpturing, upper and distal edge with coarsely spaced rugulose sculpturing (distal edge with less sculpturing), remain­ der of metapleuron nitid; anterolateral area of propodeum with rugose aciculate sculpturing, spiracular carina cristate and well­developed, spiracular area with fine rugulose sculpturing, areola deeper than anterolateral areas (appearing sunken into the propodeum), transverse carinae confused with few shallow punctate sculpturing at areolar area, posterolateral areas with carinate sculpturing mixed with shallow punctate sculpturing.
</paragraph>
<paragraph id="8BEB982DFF8F94631349528BAAFB8C93" blockId="6.[264,1325,284,1833]" pageId="6" pageNumber="7">
<emphasis id="B920443FFF8F94631349528BAA208CFB" bold="true" box="[312,371,1164,1190]" pageId="6" pageNumber="7">Legs</emphasis>
.
<collectingCountry id="F343D8BDFF8F946313F0528BAAEF8CFB" box="[385,444,1164,1190]" name="India" pageId="6" pageNumber="7">Hind</collectingCountry>
femur length/width = 3.1; hind tibia length/hind femur length = 1.4. Tarsal claws simple.
</paragraph>
<paragraph id="8BEB982DFF8F9463134952DBAF218D33" blockId="6.[264,1325,284,1833]" pageId="6" pageNumber="7">
<emphasis id="B920443FFF8F9463134952DBAAD48CAB" bold="true" box="[312,391,1244,1270]" pageId="6" pageNumber="7">Wings</emphasis>
. Forewing length =
<quantity id="4CAC35C8FF8F9463100552DBA9838CAB" box="[628,720,1244,1270]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="6" pageNumber="7" unit="mm" value="3.5">3.5 mm</quantity>
; 1CUa length/2Cub length = 0.62; length RS+Ma =
<quantity id="4CAC35C8FF8F946313795303AA248D43" box="[264,375,1284,1310]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" pageId="6" pageNumber="7" unit="mm" value="0.48">0.48 mm</quantity>
; length M+CU =
<quantity id="4CAC35C8FF8F946310275303A9E58D43" box="[598,694,1284,1310]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="6" pageNumber="7" unit="mm" value="1.3">1.3 mm</quantity>
; 1M length/ m­cu length = 1.5; pterostigma length/ height = 1.9. Hindwing: 1M length =
<quantity id="4CAC35C8FF8F946310AF532BA86D8D1B" box="[734,830,1324,1350]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="6" pageNumber="7" unit="mm" value="0.7">0.7 mm</quantity>
; 1M length/2M length = 2.1; 1M length/ M+CU length = 2.1; length r­m/length Cua = 1.1; 1
<collectingCountry id="F343D8BDFF8F9463112E5353A8C68D33" box="[863,917,1364,1390]" name="South Africa" pageId="6" pageNumber="7">RSa</collectingCountry>
length/2r­m = 1.7.
</paragraph>
<paragraph id="8BEB982DFF8F94631349537BAF238DE3" blockId="6.[264,1325,284,1833]" pageId="6" pageNumber="7">
<emphasis id="B920443FFF8F94631349537BAAE98DCB" bold="true" box="[312,442,1404,1430]" pageId="6" pageNumber="7">Metasoma</emphasis>
. First tergum length/distal width = 1; second tergum length/distal width = 2.2; third tergum length/distal width = 0.3; Hypopygium length =
<quantity id="4CAC35C8FF8F9463167E53A3AF3F8DE3" box="[1039,1132,1444,1470]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="6" pageNumber="7" unit="mm" value="3.2">3.2 mm</quantity>
;
</paragraph>
<paragraph id="8BEB982DFF8F9463137953CBA9C38E84" blockId="6.[264,1325,284,1833]" pageId="6" pageNumber="7">First and second metasomal terga with rugulose aciculate sculpturing; mediolongitudinal area of first metasomal tergum depressed but not clearly defined; 3rd metasomal tergum with shallow punctate sculpturing on anterior edge, becoming less evident towards posterior edge; metasomal terga 4 to 7 mainly nitid; hypopygium with longitudinal desclerotized pleats present medially; ovipositor long and thin throughout its length; ovipositor sheaths squarely truncate distally and slightly wider distally than basally with fine and dense setae throughout its length.</paragraph>
</subSubSection>
<subSubSection id="C34ECBA6FF8F9462134950E0AFFC8903" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7" type="materials_examined">
<paragraph id="8BEB982DFF8F9463134950E0AF368F74" blockId="6.[264,1325,284,1833]" pageId="6" pageNumber="7">
<emphasis id="B920443FFF8F9463134950E0A9758F5C" bold="true" box="[312,550,1767,1793]" pageId="6" pageNumber="7">Material examined</emphasis>
. Female, “
<collectingCountry id="F343D8BDFF8F946310D950E0A8618F5C" box="[680,818,1767,1793]" name="Philippines" pageId="6" pageNumber="7">Philippines</collectingCountry>
, Pangasinan, San Manuel,
<date id="FFEABEEDFF8F9463160450E0AFB18F5C" box="[1141,1250,1767,1793]" pageId="6" pageNumber="7" value="1953-02-04">4/ii/1953</date>
, H. &amp; M. Townes.” Deposited in
<collectionCode id="ED4500E8FF8F9463103A5108A9D48F74" box="[587,647,1807,1833]" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="6" pageNumber="7">CNC</collectionCode>
under the name “
<taxonomicName id="4C54E3AEFF8F946311265108AF028F75" box="[855,1105,1807,1832]" class="Insecta" family="Braconidae" genus="Apanteles" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="schoenobii">
<emphasis id="B920443FFF8F946311265108AF028F75" box="[855,1105,1807,1832]" italics="true" pageId="6" pageNumber="7">Apanteles schoenobii</emphasis>
</taxonomicName>
.”
</paragraph>
<paragraph id="8BEB982DFF8E94621349571BAFFC8903" blockId="7.[264,1323,284,830]" pageId="7" pageNumber="8">
The
<typeStatus id="54EF268FFF8E9462131E571BAA86896B" box="[367,469,284,310]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
is in the Natural History Museum, London and was not examined but our material was compared and matched with the
<typeStatus id="54EF268FFF8E946211205743A8EB8903" box="[849,952,324,350]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
by W. R. M. Mason.
</paragraph>
</subSubSection>
<subSubSection id="C34ECBA6FF8E94621349576CA9968B63" pageId="7" pageNumber="8" type="discussion">
<paragraph id="8BEB982DFF8E94621349576CA98589A3" blockId="7.[264,1323,284,830]" pageId="7" pageNumber="8">
<emphasis id="B920443FFF8E94621349576CAAEC89D8" bold="true" box="[312,447,363,389]" pageId="7" pageNumber="8">Comments</emphasis>
. This species is easily recognized by the forewing vein 1CUa which is contrastingly shorter than vein 1Cub (approximately 0.6 X) and the metapleuron being mainly covered in punctate sculpturing. Neither of the two remaining
<taxonomicName id="4C54E3AEFF8E9462161B57BBAF998988" box="[1130,1226,444,469]" class="Insecta" family="Braconidae" genus="Exoryza" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B920443FFF8E9462161B57BBAF998988" box="[1130,1226,444,469]" italics="true" pageId="7" pageNumber="8">Exoryza</emphasis>
</taxonomicName>
species exhibits this combination of characters.
</paragraph>
<paragraph id="8BEB982DFF8E94621349540BAFE18AB3" blockId="7.[264,1323,284,830]" pageId="7" pageNumber="8">
The original description of Wilkinson departs from the one presented here and the one presented by
<bibRefCitation id="EFC5E5DCFF8E946213DC5433A9048A13" author="Mason" box="[429,599,564,590]" pageId="7" pageNumber="10" refString="Mason, W. R. M. (1981) The polyphyletic nature of Apanteles Foerster (Hymenoptera: Braconidae): A phylogeny and reclassification of Microgastrinae. Memoirs of the Entomological Society of Canada, 115, 1 - 147." type="journal article" year="1981">Mason (1981)</bibRefCitation>
in the dark coloration of the mouthparts as well as of the clypeus, the yellow color of the legs in combination with the brownish tarsomeres; the wings not being distinctly infumate, the presence of more than 6 costulae in the scutellar groove; the absence of minute rugose sculpturing on the anterior edge of the mesopleuron. The remainder of Wilkinsons description agrees with our material in all details.
</paragraph>
<paragraph id="8BEB982DFF8E9462134954FBA9968B63" blockId="7.[264,1323,284,830]" pageId="7" pageNumber="8">
<emphasis id="B920443FFF8E9462134954FBA9518B4B" bold="true" box="[312,514,764,790]" pageId="7" pageNumber="8">Rearing records</emphasis>
. Reared from stem­boring Pyraloidea larvae on rice plants according to Wilkinsons and Masons accounts.
</paragraph>
</subSubSection>
</treatment>
</document>