334 lines
43 KiB
XML
334 lines
43 KiB
XML
<document id="58E2DD6B54F225E6FBB39C02716B76F8" ID-DOI="10.11646/phytotaxa.622.5.2" ID-ISSN="1179-3163" ID-Zenodo-Dep="10150899" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1700250772755" checkinUser="plazi" docAuthor="Londoño-Echeverri, Yeison & Trujillo-López, Ana María" docDate="2023" docId="03FC87ECFFC9FFE11C87F852FD7DFE51" docLanguage="en" docName="phytotaxa.622.5.2.pdf" docOrigin="Phytotaxa 622 (5)" docSource="https://phytotaxa.mapress.com/pt/article/download/phytotaxa.622.5.2/51262" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.7:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="7" docTitle="Aenigmanu chococola Londono-Ech. & A. M. Trujillo 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="286" masterDocId="FFC5FF94FFC8FFE41C0FFFC6FFC4FFAC" masterDocTitle="Taxonomic studies in Colombian Picramniaceae (Picramniales): new species of Aenigmanu, Nothotalisia and Picramnia from the Pacific and Magdalena Valley regions" masterLastPageNumber="297" masterPageNumber="281" pageNumber="282" updateTime="1700251273136" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="BBE1D844EB5A6F20F66A320F75188D5D">Taxonomic studies in Colombian Picramniaceae (Picramniales): new species of Aenigmanu, Nothotalisia and Picramnia from the Pacific and Magdalena Valley regions</mods:title>
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<mods:roleTerm id="ACE9549C2F66D847BB154EAFEAB878BC">Author</mods:roleTerm>
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<mods:namePart id="5D64A0760C5497717B74C90F7934AF3F">Londoño-Echeverri, Yeison</mods:namePart>
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<mods:affiliation id="0133C2E633274BFDC67014517793D2D2">Herbario Universidad de Antioquia (HUA) y Grupo de Estudios Botánicos (GEOBOTA), Instituto de Biología, Facultad de Ciencias Exactas y Naturales, Universidad de Antioquia, A. A, 1226, Medellín, Colombia & Herbario Gabriel Gutiérrez Villegas (MEDEL) y Semillero en estudios taxonómicos de plantas de Colombia, Facultad de Ciencias, Universidad Nacional de Colombia, A. A, 3840, Medellín, Colombia & ylondono @ unal. edu. co; https: // orcid. org / 0000 - 0001 - 5169 - 598 X</mods:affiliation>
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<mods:nameIdentifier id="D3A3B51FFE0165200A486016B9894DAB" type="email">ylondono@unal.edu.co</mods:nameIdentifier>
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<mods:namePart id="3CC8C9D69881F7FCF4CD7CD110703524">Trujillo-López, Ana María</mods:namePart>
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<mods:affiliation id="BAFFFA2AE1B8B62DE0CD8E1ECFBB86DE">Herbario Universidad de Antioquia (HUA) y Grupo de Estudios Botánicos (GEOBOTA), Instituto de Biología, Facultad de Ciencias Exactas y Naturales, Universidad de Antioquia, A. A, 1226, Medellín, Colombia & Herbario Gabriel Gutiérrez Villegas (MEDEL) y Semillero en estudios taxonómicos de plantas de Colombia, Facultad de Ciencias, Universidad Nacional de Colombia, A. A, 3840, Medellín, Colombia & amtrujillol @ unal. edu. co; https: // orcid. org / 0000 - 0002 - 0898 - 1646</mods:affiliation>
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<mods:nameIdentifier id="504C27A7843968CAF39804CC7B224496" type="email">amtrujillol@unal.edu.co</mods:nameIdentifier>
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<mods:title id="6314D923B88879505CAFCC8D14C527F2">Phytotaxa</mods:title>
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<mods:part id="2396FD4EECEF25FC478B28B47595C9E2">
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<mods:date id="7889697C5DB570D665C1414C12806ABF">2023</mods:date>
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<mods:number id="CB6AB4255572F0A17BE68EABE08D9762">2023-11-03</mods:number>
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<mods:number id="61633CE6620A41F68C6F34B23F122B63">622</mods:number>
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<subSubSection id="C34F6571FFC9FFE51C87F852FC07F802" box="[136,963,1940,1967]" pageId="1" pageNumber="282" type="nomenclature">
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<heading id="D0A28196FFC9FFE51C87F852FC07F802" box="[136,963,1940,1967]" fontSize="11" level="2" pageId="1" pageNumber="282" reason="3">
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<taxonomicName id="4C554D79FFC9FFE51C87F852FD2CF802" authority="Londono-Ech. & A.M.Trujillo" authorityName="Londono-Ech. & A. M. Trujillo" authorityYear="2023" box="[136,744,1940,1967]" class="Magnoliopsida" family="Picramniaceae" genus="Aenigmanu" kingdom="Plantae" order="Picramniales" pageId="1" pageNumber="282" phylum="Tracheophyta" rank="species" species="chococola" status="sp. nov.">
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<emphasis id="B921EAE8FFC9FFE51C87F852FE4FF803" bold="true" box="[136,395,1940,1967]" italics="true" pageId="1" pageNumber="282">Aenigmanu chococola</emphasis>
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Londoño-Ech. & A.M.Trujillo
|
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</taxonomicName>
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,
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<emphasis id="B921EAE8FFC9FFE51EFCF853FC8CF802" box="[755,840,1941,1966]" italics="true" pageId="1" pageNumber="282">
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<taxonomicNameLabel id="A2125793FFC9FFE51EFCF853FC8CF802" box="[755,840,1941,1966]" pageId="1" pageNumber="282" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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(
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<figureCitation id="136E2A7FFFC9FFE51F59F852FC59F802" box="[854,925,1940,1966]" captionStart="FIGURE 1" captionStartId="3.[136,229,1980,2002]" captionTargetBox="[165,1422,187,1953]" captionTargetId="figure-17@3.[165,1422,187,1953]" captionTargetPageId="3" captionText="FIGURE 1. Holotype of Aenigmanu chococola." figureDoi="http://doi.org/10.5281/zenodo.10150901" httpUri="https://zenodo.org/record/10150901/files/figure.png" pageId="1" pageNumber="282">Figs 1</figureCitation>
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,
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<figureCitation id="136E2A7FFFC9FFE51FA4F852FC7EF802" box="[939,954,1940,1966]" captionStart="FIGURE 2" captionStartId="4.[136,229,1883,1905]" captionTargetBox="[154,1432,193,1773]" captionTargetId="figure-27@4.[151,1435,190,1859]" captionTargetPageId="4" captionText="FIGURE 2. Aenigmanu chococola. A. Flowering branchlet of pistillate plant. B. Stipules. C. Detail of leaves blade at very tip, abaxial view. D. Pistillate inflorescence. E. Pistillate flower beginning to develop fruit. (A–E from the holotype; illustration by Ana Cristina Pareja, HUA)." figureDoi="http://doi.org/10.5281/zenodo.10150903" httpUri="https://zenodo.org/record/10150903/files/figure.png" pageId="1" pageNumber="282">2</figureCitation>
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)
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</heading>
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</paragraph>
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<subSubSection id="C34F6571FFC9FFE61C87F826FD37FB42" lastPageId="2" lastPageNumber="283" pageId="1" pageNumber="282" type="materials_examined">
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<paragraph id="8BEA36FAFFC9FFE51C87F826FBE8F7B5" blockId="1.[136,1452,2016,2074]" pageId="1" pageNumber="282">
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<materialsCitation id="3B3D3CA7FFC9FFE51C87F826FBE3F7B5" collectingDate="1994-09-08" collectionCode="SO, HUA" country="Colombia" county="Pijiba" elevation="28" location="Type" municipality="Nuqui" pageId="1" pageNumber="282" specimenCount="1" typeStatus="holotype">
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:
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<emphasis id="B921EAE8FFC9FFE51CB1F826FF12F859" bold="true" box="[190,214,2016,2037]" pageId="1" pageNumber="282">—</emphasis>
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<collectingCountry id="F342766AFFC9FFE51CD9F826FE9DF859" box="[214,345,2016,2037]" name="Colombia" pageId="1" pageNumber="282">COLOMBIA</collectingCountry>
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.
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<location id="8E8A6021FFC9FFE51D6DF826FE60F85A" LSID="urn:lsid:plazi:treatment:03FC87ECFFC9FFE11C87F852FD7DFE51:8E8A6021FFC9FFE51D6DF826FE60F85A" box="[354,420,2016,2038]" country="Colombia" county="Pijiba" municipality="Nuqui" name="Choco" pageId="1" pageNumber="282">Chocó</location>
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: Mun.
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<location id="8E8A6021FFC9FFE51DE8F826FDE1F859" LSID="urn:lsid:plazi:treatment:03FC87ECFFC9FFE11C87F852FD7DFE51:8E8A6021FFC9FFE51DE8F826FDE1F859" box="[487,549,2016,2037]" country="Colombia" county="Pijiba" municipality="Nuqui" name="Nuqui" pageId="1" pageNumber="282">Nuquí</location>
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, costa del
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<location id="8E8A6021FFC9FFE51E85F826FD1FF85A" LSID="urn:lsid:plazi:treatment:03FC87ECFFC9FFE11C87F852FD7DFE51:8E8A6021FFC9FFE51E85F826FD1FF85A" box="[650,731,2016,2038]" country="Colombia" county="Pijiba" municipality="Nuqui" name="Pacifico" pageId="1" pageNumber="282">Pacífico</location>
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, correg.
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<location id="8E8A6021FFC9FFE51F22F826FC43F85A" LSID="urn:lsid:plazi:treatment:03FC87ECFFC9FFE11C87F852FD7DFE51:8E8A6021FFC9FFE51F22F826FC43F85A" box="[813,903,2016,2038]" country="Colombia" county="Pijiba" municipality="Nuqui" name="Termales" pageId="1" pageNumber="282">Termales</location>
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, quebrada
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<location id="8E8A6021FFC9FFE51FE0F826FBB0F85A" LSID="urn:lsid:plazi:treatment:03FC87ECFFC9FFE11C87F852FD7DFE51:8E8A6021FFC9FFE51FE0F826FBB0F85A" box="[1007,1140,2016,2038]" country="Colombia" county="Pijiba" municipality="Nuqui" name="Piedra-Piedra" pageId="1" pageNumber="282">Piedra-Piedra</location>
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, 1 hr.
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<collectionCode id="ED44AE3FFFC9FFE518A1F826FB09F859" box="[1198,1229,2016,2037]" country="Bulgaria" lsid="urn:lsid:biocol.org:col:14079" name="Sofia University" pageId="1" pageNumber="282" type="Herbarium">SO</collectionCode>
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de
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<collectingMunicipality id="6B8EAC80FFC9FFE518E0F826FAE9F859" box="[1263,1325,2016,2037]" pageId="1" pageNumber="282">Nuquí</collectingMunicipality>
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, hotel
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<collectingCounty id="628B4E76FFC9FFE51962F826FA6CF85A" box="[1389,1448,2016,2038]" pageId="1" pageNumber="282">Pijibá</collectingCounty>
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,
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, 5°37ʹ0ʹʹN, 77°15ʹ0ʹʹW,
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<date id="FFEB103AFFC9FFE51DE0F7C2FD9FF7B5" box="[495,603,2052,2073]" pageId="1" pageNumber="282" value="1994-09-08">
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</date>
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(im fr),
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<emphasis id="B921EAE8FFC9FFE51EA3F7C2FCB6F7B5" box="[684,882,2052,2073]" italics="true" pageId="1" pageNumber="282">Callejas et al. 11230</emphasis>
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(
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<typeStatus id="54EE8858FFC9FFE51F70F7C2FC1CF7B6" box="[895,984,2052,2074]" pageId="1" pageNumber="282" type="holotype">holotype</typeStatus>
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:
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<collectionCode id="ED44AE3FFFC9FFE51FEEF7C2FBD9F7B5" box="[993,1053,2052,2073]" country="Colombia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15677" lsid="urn:lsid:biocol.org:col:15677" name="Universidad de Antioquia" pageId="1" pageNumber="282" type="Herbarium">HUA</collectionCode>
|
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!)
|
||
</materialsCitation>
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.
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</paragraph>
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<paragraph id="8BEA36FAFFCAFFE61C87FF59FF36FE8C" blockId="2.[136,1451,159,288]" pageId="2" pageNumber="283">
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<taxonomicName id="4C554D79FFCAFFE61C87FF59FE9BFF18" authorityName="Londono-Ech. & A. M. Trujillo" authorityYear="2023" box="[136,351,159,180]" class="Magnoliopsida" family="Picramniaceae" genus="Aenigmanu" kingdom="Plantae" order="Picramniales" pageId="2" pageNumber="283" phylum="Tracheophyta" rank="species" species="chococola">
|
||
<emphasis id="B921EAE8FFCAFFE61C87FF59FE9BFF18" box="[136,351,159,180]" italics="true" pageId="2" pageNumber="283">Aenigmanu chococola</emphasis>
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||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C554D79FFCAFFE61DEFFF59FD9BFF18" box="[480,607,159,180]" class="Magnoliopsida" family="Picramniaceae" genus="Aenigmanu" kingdom="Plantae" order="Picramniales" pageId="2" pageNumber="283" phylum="Tracheophyta" rank="species" species="alvareziae">
|
||
<emphasis id="B921EAE8FFCAFFE61DEFFF59FD9BFF18" box="[480,607,159,180]" italics="true" pageId="2" pageNumber="283">A. alvareziae</emphasis>
|
||
</taxonomicName>
|
||
by its longer stipules 8.2–11.3 ×
|
||
<quantity id="4CAD9B1FFFCAFFE61FAAFF59FBDDFF19" box="[933,1049,159,181]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" metricValueMax="1.2" metricValueMin="0.8" pageId="2" pageNumber="283" unit="mm" value="1.0" valueMax="1.2" valueMin="0.8">0.8–1.2 mm</quantity>
|
||
(vs.
|
||
<emphasis id="B921EAE8FFCAFFE61845FF66FBA3FF19" box="[1098,1127,160,181]" italics="true" pageId="2" pageNumber="283">ca.</emphasis>
|
||
6.5 ×
|
||
<quantity id="4CAD9B1FFFCAFFE618A8FF59FB31FF18" box="[1191,1269,159,181]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="283" unit="mm" value="1.5">1.5 mm</quantity>
|
||
), wider leaf blades (7.0–)
|
||
<quantity id="4CAD9B1FFFCAFFE61CF3FF05FEB6FF75" box="[252,370,195,217]" metricMagnitude="-2" metricUnit="m" metricValue="9.45" metricValueMax="10.6" metricValueMin="8.3" pageId="2" pageNumber="283" unit="cm" value="9.45" valueMax="10.6" valueMin="8.3">8.3–10.6 cm</quantity>
|
||
wide (vs. 3.4–5.5 [–7.5] cm) that are basally convex (vs. cuneate to decurrent) and commonly apically emarginate at the very tip (vs. commonly mucronate) and bracts of pistillate inflorescence longer 1.5–2.3 (–3.7) mm (vs. equal or less than
|
||
<quantity id="4CAD9B1FFFCAFFE61990FF21FF22FE8C" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="283" unit="mm" value="1.0">1 mm</quantity>
|
||
).
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</paragraph>
|
||
<paragraph id="8BEA36FAFFCAFFE61C87FE89FD37FB42" blockId="2.[136,1452,335,2090]" pageId="2" pageNumber="283">
|
||
Shrubs
|
||
<quantity id="4CAD9B1FFFCAFFE61CD2FE96FEE0FEC6" box="[221,292,336,362]" metricMagnitude="0" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="2" pageNumber="283" unit="m" value="3.5" valueMax="4.0" valueMin="3.0">3–4 m</quantity>
|
||
tall, indumentum of simple trichomes, buds and young branchlets densely shortly strigose, glabrescent, stipules 8.2–11.3 ×
|
||
<quantity id="4CAD9B1FFFCAFFE61D62FEB2FE3DFE22" box="[365,505,372,398]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" metricValueMax="1.2" metricValueMin="0.8" pageId="2" pageNumber="283" unit="mm" value="1.0" valueMax="1.2" valueMin="0.8">0.8–1.2 mm</quantity>
|
||
, linear to narrowly oblanceolate, apically acute and cuneate to shortly acuminate, shortly strigose on both surfaces, more densely abaxially; primary vein raised, and occasionally secondary veins slightly raised abaxially; caducous, leaving transversal scars up to
|
||
<quantity id="4CAD9B1FFFCAFFE61F82FE7AFC15FE7A" box="[909,977,444,470]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="283" unit="mm" value="3.0">3 mm</quantity>
|
||
long on fully expanded branchlet nodes. Leaves simple (possibly unifoliolate), alternate, petiole
|
||
<quantity id="4CAD9B1FFFCAFFE61EFBFE19FC95FE56" box="[756,849,479,506]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="2" pageNumber="283" unit="mm" value="6.0" valueMax="7.0" valueMin="5.0">5–7 mm</quantity>
|
||
long, flat adaxially, shortly strigose; blades 18.5–23.6 × (7.0–)
|
||
<quantity id="4CAD9B1FFFCAFFE61CE6FDC5FEB2FDB2" box="[233,374,515,542]" metricMagnitude="-2" metricUnit="m" metricValue="9.45" metricValueMax="10.6" metricValueMin="8.3" pageId="2" pageNumber="283" unit="cm" value="9.45" valueMax="10.6" valueMin="8.3">8.3–10.6 cm</quantity>
|
||
, broadly obovate, basally obtuse and convex, apically acute to obtuse and acuminate, commonly emarginate at the very tip; shortly strigose abaxially, nearly glabrous on the blade, sparsely so toward the primary and secondary veins; puberulous adaxially, nearly glabrous on the blade, densely so toward the primary and secondary veins; primary vein raised abaxially, flat to slightly raised adaxially, secondary veins 10–13 on each side, raised abaxially, slightly impressed adaxially, brochidodromous, loops consistently of 2° gauge and with course monopodial to slightly sympodial, tertiary veins raised abaxially, slightly raised to flat adaxially, percurrent, vein framework visible up to 6° gauge; margin entire. Staminate inflorescences and flowers not seen. Pistillate inflorescences forming cymules, congested as scaly short-shoots, 1–2-forked, leaf opposed, 7.0–
|
||
<quantity id="4CAD9B1FFFCAFFE61F54FD39FC7BFCB6" box="[859,959,767,794]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" pageId="2" pageNumber="283" unit="mm" value="14.5">14.5 mm</quantity>
|
||
long (including a common penduncle
|
||
<quantity id="4CAD9B1FFFCAFFE61975FCC6FF25FC92" metricMagnitude="-3" metricUnit="m" metricValue="3.3499999999999996" metricValueMax="3.9" metricValueMin="2.8" pageId="2" pageNumber="283" unit="mm" value="3.3499999999999996" valueMax="3.9" valueMin="2.8">2.8– 3.9 mm</quantity>
|
||
long), bearing 1–2 flowers at a time, axes shortly strigose; bracts 1.5–2.3 (–3.7) ×
|
||
<quantity id="4CAD9B1FFFCAFFE618BCFCE2FAFBFC92" box="[1203,1343,804,830]" metricMagnitude="-3" metricUnit="m" metricValue="1.0499999999999998" metricValueMax="1.4" metricValueMin="0.7" pageId="2" pageNumber="283" unit="mm" value="1.0499999999999998" valueMax="1.4" valueMin="0.7">0.7–1.4 mm</quantity>
|
||
, ovate to lanceolate, apically acute and cuneate to narrowly long-acuminate, shortly strigose throughout and apically densely puberulous on both surfaces, persistent; pedicel ca.
|
||
<quantity id="4CAD9B1FFFCAFFE61EC6FCADFCE5FC2A" box="[713,801,875,902]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="283" unit="mm" value="1.5">1.5 mm</quantity>
|
||
, ca.
|
||
<quantity id="4CAD9B1FFFCAFFE61F5CFCAAFC6DFC2A" box="[851,937,876,902]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="283" unit="mm" value="0.8">0.8 mm</quantity>
|
||
diam., densely shortly strigose to puberulous, bracteoles not seen. Pistillate flowers pentamerous (the specimen label indicate “flowers white” but colour is not preserved, the description here provided is from flowers beginning to develop fruits), calyx 3.5–4.0 mm long, connate basally, reflexed but becoming inflexed toward the lobes, the basal connate portion ca.
|
||
<quantity id="4CAD9B1FFFCAFFE61851FC1EFB71FC5E" box="[1118,1205,984,1010]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="2" pageNumber="283" unit="mm" value="0.7">0.7 mm</quantity>
|
||
long; lobes 2.8–3.3 ×
|
||
<quantity id="4CAD9B1FFFCAFFE61C87FC3AFED6FBBA" box="[136,274,1020,1046]" metricMagnitude="-3" metricUnit="m" metricValue="1.5499999999999998" metricValueMax="1.7" metricValueMin="1.4" pageId="2" pageNumber="283" unit="mm" value="1.5499999999999998" valueMax="1.7" valueMin="1.4">1.4–1.7 mm</quantity>
|
||
, ovate to lanceolate, apically acute and cuneate, densely shortly strigose to puberulous abaxially, mainly glabrous and only apically puberulous adaxially, margin entire; petals not seen, their scars alternating with the sepals; staminodes not seen; disc ca.
|
||
<quantity id="4CAD9B1FFFCAFFE61DDDFB85FDE3FBF2" box="[466,551,1091,1118]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="2" pageNumber="283" unit="mm" value="0.6">0.6 mm</quantity>
|
||
tall, ca.
|
||
<quantity id="4CAD9B1FFFCAFFE61E89FB82FD02FBF2" box="[646,710,1092,1118]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="283" unit="mm" value="2.0">2 mm</quantity>
|
||
diam., intrastaminal, like a gynophore (i.e., not annular, but rather subtending the ovary), glabrous; ovary syncarpous, 7-carpellate, 3.0–
|
||
<quantity id="4CAD9B1FFFCAFFE61F93FBAEFC37FB2E" box="[924,1011,1128,1154]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="2" pageNumber="283" unit="mm" value="3.4">3.4 mm</quantity>
|
||
tall, 4.0–
|
||
<quantity id="4CAD9B1FFFCAFFE6186FFBA1FB7CFB2E" box="[1120,1208,1127,1154]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="2" pageNumber="283" unit="mm" value="4.6">4.6 mm</quantity>
|
||
in diam., globose, 7- lobed, densely strigose to puberulous, transversally ridged; common style ca.
|
||
<quantity id="4CAD9B1FFFCAFFE61FEAFB4AFBFEFB0A" box="[997,1082,1164,1190]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="2" pageNumber="283" unit="mm" value="0.4">0.4 mm</quantity>
|
||
long, ca.
|
||
<quantity id="4CAD9B1FFFCAFFE618AAFB4DFB3EFB0A" box="[1189,1274,1163,1190]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="2" pageNumber="283" unit="mm" value="0.6">0.6 mm</quantity>
|
||
diam. (obsolete by fruit expansion), densely strigose to puberulous, stigma 7-lobed, each ca. 0.8 ×
|
||
<quantity id="4CAD9B1FFFCAFFE61837FB76FB55FB66" box="[1080,1169,1200,1226]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="283" unit="mm" value="0.2">0.2 mm</quantity>
|
||
, patent to erect, sparsely shortly strigose. Mature fruits not seen. Seeds not seen.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34F6571FFCAFFE61CB2FB31FA41FAD1" pageId="2" pageNumber="283" type="distribution">
|
||
<paragraph id="8BEA36FAFFCAFFE61CB2FB31FAAFFAF5" blockId="2.[136,1452,335,2090]" pageId="2" pageNumber="283">
|
||
<emphasis id="B921EAE8FFCAFFE61CB2FB31FDCCFABD" bold="true" box="[189,520,1271,1298]" pageId="2" pageNumber="283">Distribution and habitat:—</emphasis>
|
||
Endemic to western
|
||
<collectingCountry id="F342766AFFCAFFE61EFDFB31FCA7FABD" box="[754,867,1271,1297]" name="Colombia" pageId="2" pageNumber="283">Colombia</collectingCountry>
|
||
, where it has been recorded on nearly coastal areas from the Nuquí Municipality in the
|
||
<collectingRegion id="4991F818FFCAFFE61E1FFADDFD26FA9A" box="[528,738,1307,1334]" country="Colombia" name="Choco" pageId="2" pageNumber="283">Chocó Department</collectingRegion>
|
||
(
|
||
<figureCitation id="136E2A7FFFCAFFE61EE0FADDFCEFFA9A" box="[751,811,1307,1334]" captionStart="FIGURE 3" captionStartId="5.[136,229,1981,2003]" captionTargetBox="[161,1433,546,1954]" captionTargetId="figure-185@5.[151,1436,533,1956]" captionTargetPageId="5" captionText="FIGURE 3. Distribution of Aenigmanu chococola, Nothotalisia karsticola and Picramnia vinacea." figureDoi="http://doi.org/10.5281/zenodo.10150905" httpUri="https://zenodo.org/record/10150905/files/figure.png" pageId="2" pageNumber="283">Fig 3</figureCitation>
|
||
), Pacific Biogeographical Region, at
|
||
<quantity id="4CAD9B1FFFCAFFE618C7FADDFADFFA9A" box="[1224,1307,1307,1334]" metricMagnitude="1" metricUnit="m" metricValue="2.5" metricValueMax="5.0" metricValueMin="0.0" pageId="2" pageNumber="283" unit="m" value="25.0" valueMax="50.0" valueMin="0.0">0–50 m</quantity>
|
||
in equatorial rainforest climate
|
||
<typeStatus id="54EE8858FFCAFFE61D5BFA86FE40FAF6" box="[340,388,1344,1370]" pageId="2" pageNumber="283">type</typeStatus>
|
||
(Af), a region globally considered one of the wettest and most biodiverse (
|
||
<bibRefCitation id="EFC44B0BFFCAFFE618C3FA86FA9AFAF5" author="Gentry, A." box="[1228,1374,1343,1370]" pageId="2" pageNumber="283" pagination="71 - 92" refId="ref8564" refString="Gentry, A. (1986) Species richness and floristic composition of Choco region plant communities. Caldasia 15 (71 / 75): 71 - 92. https: // www. jstor. org / stable / 43406071" type="journal article" year="1986">Gentry 1986</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BEA36FAFFCAFFE61CB2FAA2FA41FAD1" blockId="2.[136,1452,335,2090]" box="[189,1413,1379,1406]" pageId="2" pageNumber="283">
|
||
<emphasis id="B921EAE8FFCAFFE61CB2FAA2FE9BFAD2" bold="true" box="[189,351,1380,1406]" pageId="2" pageNumber="283">Phenology:—</emphasis>
|
||
Only immature fruits known in September; flowering on the same date (no flowers preserved).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34F6571FFCAFFE11CB2FA4EFD7DFE51" lastPageId="5" lastPageNumber="286" pageId="2" pageNumber="283" type="etymology">
|
||
<paragraph id="8BEA36FAFFCAFFE61CB2FA4EFF15FA69" blockId="2.[136,1452,335,2090]" pageId="2" pageNumber="283">
|
||
<emphasis id="B921EAE8FFCAFFE61CB2FA4EFEA0FA0E" bold="true" box="[189,356,1416,1442]" pageId="2" pageNumber="283">Etymology:—</emphasis>
|
||
Referring to the
|
||
<collectingRegion id="4991F818FFCAFFE61E10FA41FDACFA0D" box="[543,616,1415,1441]" country="Panama" name="Embera-Wounaan" pageId="2" pageNumber="283">Choco</collectingRegion>
|
||
Biogeographical Region (= Pacific Biogeographical Region
|
||
<emphasis id="B921EAE8FFCAFFE61914FA4EFA9DFA0D" box="[1307,1369,1416,1441]" italics="true" pageId="2" pageNumber="283">sensu</emphasis>
|
||
<bibRefCitation id="EFC44B0BFFCAFFE6196EFA41FF01FA69" author="Bernal, R." pageId="2" pageNumber="283" pagination="19 - 32" refId="ref8231" refString="Bernal, R. (2016) Geografia de Colombia. In: Bernal, R., Gradstein, S. R. & Celis, M. (Eds.) Catalogo de plantas y liquenes de Colombia. Instituto de Ciencias Naturales, Universidad Nacional de Colombia, Bogota, pp. 19 - 32." type="book chapter" year="2016">Bernal 2016</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BEA36FAFFCAFFE61CB2FA09FD5CF96D" blockId="2.[136,1452,335,2090]" pageId="2" pageNumber="283">
|
||
<emphasis id="B921EAE8FFCAFFE61CB2FA09FE0BFA46" bold="true" box="[189,463,1487,1514]" pageId="2" pageNumber="283">Conservation status:—</emphasis>
|
||
Known only from a subpopulation found outside protected areas, with AOO =
|
||
<quantity id="4CAD9B1FFFCAFFE61931FA16FAB3FA45" box="[1342,1399,1487,1514]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="2" pageNumber="283" unit="km" value="4.0">4 km</quantity>
|
||
<superScript id="7C209BB2FFCAFFE61978FA09FABBFA71" attach="left" box="[1399,1407,1487,1501]" fontSize="6" pageId="2" pageNumber="283">2</superScript>
|
||
and less than
|
||
<quantity id="4CAD9B1FFFCAFFE61CFBFA35FEF9F9A1" box="[244,317,1523,1550]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="283" unit="km" value="15.0">15 km</quantity>
|
||
from a populated centre. The forests of the Pacific Biogeographical Region in
|
||
<collectingCountry id="F342766AFFCAFFE618C4FA35FAF8F9A1" box="[1227,1340,1523,1549]" name="Colombia" pageId="2" pageNumber="283">Colombia</collectingCountry>
|
||
, although apparently continuous, exhibit high levels of habitat degradation due to selective logging and mining that, according to
|
||
<bibRefCitation id="EFC44B0BFFCAFFE61C87F9FAFDE4F9FA" author="Moreno, G. R. & Ledezma-Renteria, E." box="[136,544,1595,1622]" pageId="2" pageNumber="283" pagination="58 - 65" refId="ref8899" refString="Moreno, G. R. & Ledezma-Renteria, E. (2007) Efectos de las actividades socio-economicas (mineria y explotacion maderera) sobre los bosques del departamento del Choco. Revista institucional universidad tecnologica del Choco 26: 58 - 65." type="journal article" year="2007">Moreno & Ledezma-Rentería (2007)</bibRefCitation>
|
||
, are the main economic activities of the region. Rangel-Ch. (2004) also emphasised the high threats and habitat degradation in this region, suggesting a loss of forest cover at 25–36%. Because to its small distribution outside protected areas and the threats to its habitat,
|
||
<taxonomicName id="4C554D79FFCAFFE61F92F942FBEBF931" authorityName="Londono-Ech. & A. M. Trujillo" authorityYear="2023" box="[925,1071,1667,1694]" class="Magnoliopsida" family="Picramniaceae" genus="Aenigmanu" kingdom="Plantae" order="Picramniales" pageId="2" pageNumber="283" phylum="Tracheophyta" rank="species" species="chococola">
|
||
<emphasis id="B921EAE8FFCAFFE61F92F942FBEBF931" box="[925,1071,1667,1694]" italics="true" pageId="2" pageNumber="283">A. chococola</emphasis>
|
||
</taxonomicName>
|
||
should be considered as critically endangered, CR according to criteria B2ab(iii).
|
||
</paragraph>
|
||
<paragraph id="8BEA36FAFFCAFFE61CB2F90AFDC1F8FD" blockId="2.[136,1452,335,2090]" pageId="2" pageNumber="283">
|
||
<emphasis id="B921EAE8FFCAFFE61CB2F90AFEE2F94A" bold="true" box="[189,294,1740,1766]" pageId="2" pageNumber="283">Notes:—</emphasis>
|
||
<taxonomicName id="4C554D79FFCAFFE61D29F90AFDE7F949" authorityName="Londono-Ech. & A. M. Trujillo" authorityYear="2023" box="[294,547,1739,1766]" class="Magnoliopsida" family="Picramniaceae" genus="Aenigmanu" kingdom="Plantae" order="Picramniales" pageId="2" pageNumber="283" phylum="Tracheophyta" rank="species" species="chococola">
|
||
<emphasis id="B921EAE8FFCAFFE61D29F90AFDE7F949" box="[294,547,1739,1766]" italics="true" pageId="2" pageNumber="283">Aenigmanu chococola</emphasis>
|
||
</taxonomicName>
|
||
is known only by a pistillate specimen collected by Ricardo Callejas, Pedro Acevedo and Steven Paul Churchill in 1994, at which time it would has been almost impossible to consider that it belonged to
|
||
<taxonomicName id="4C554D79FFCAFFE61D17F8D5FE7BF881" box="[280,447,1811,1837]" class="Magnoliopsida" family="Picramniaceae" kingdom="Plantae" order="Picramniales" pageId="2" pageNumber="283" phylum="Tracheophyta" rank="family">Picramniaceae</taxonomicName>
|
||
. Despite extensive searches for other specimens in Colombian and foreign herbaria, no other specimens have been found.
|
||
</paragraph>
|
||
<paragraph id="8BEA36FAFFCAFFE11CB2F89AFADBFF71" blockId="2.[136,1452,335,2090]" lastBlockId="5.[136,1452,158,509]" lastPageId="5" lastPageNumber="286" pageId="2" pageNumber="283">
|
||
Because of the absence of additional specimens, the diagnosis is mainly based on vegetative features. Otherwise, only the longer bracts of pistillate inflorescences are useful in recognising
|
||
<taxonomicName id="4C554D79FFCAFFE61FD9F846FBADF835" authorityName="Londono-Ech. & A. M. Trujillo" authorityYear="2023" box="[982,1129,1919,1946]" class="Magnoliopsida" family="Picramniaceae" genus="Aenigmanu" kingdom="Plantae" order="Picramniales" pageId="2" pageNumber="283" phylum="Tracheophyta" rank="species" species="chococola">
|
||
<emphasis id="B921EAE8FFCAFFE61FD9F846FBADF835" box="[982,1129,1919,1946]" italics="true" pageId="2" pageNumber="283">A. chococola</emphasis>
|
||
</taxonomicName>
|
||
in the phenological stage of the
|
||
<typeStatus id="54EE8858FFCAFFE61CBCF862FF27F812" box="[179,227,1956,1982]" pageId="2" pageNumber="283">type</typeStatus>
|
||
specimen. Although the floral and fruit morphology is distinctive among
|
||
<taxonomicName id="4C554D79FFCAFFE61839F865FB1DF811" box="[1078,1241,1955,1981]" class="Magnoliopsida" family="Picramniaceae" kingdom="Plantae" order="Picramniales" pageId="2" pageNumber="283" phylum="Tracheophyta" rank="family">Picramniaceae</taxonomicName>
|
||
genera, it is often uniform within each genus, especially in apparently related species, resulting in the wide use of vegetative features for recognition at the species level,
|
||
<emphasis id="B921EAE8FFCAFFE61E2AF82AFD89F7A9" box="[549,589,2028,2053]" italics="true" pageId="2" pageNumber="283">e.g.</emphasis>
|
||
, taxonomic keys of
|
||
<bibRefCitation id="EFC44B0BFFCAFFE61F35F82DFC26F7AA" author="Thomas, W. W." box="[826,994,2027,2054]" pageId="2" pageNumber="283" pagination="89 - 105" refId="ref9405" refString="Thomas, W. W. (1988) A conspectus of Mexican and Central American Picramnia (Simaroubaceae). Brittonia 40: 89 - 105. https: // doi. org / 10.2307 / 2806882" type="journal article" year="1988">Thomas (1988</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFC44B0BFFCAFFE61FE0F82AFBEEF7AA" author="Thomas, W. W." box="[1007,1066,2028,2054]" pageId="2" pageNumber="283" pagination="171 - 174" refId="ref9443" refString="Thomas, W. W. (1990) A new species of Picramnia (Simaroubaceae) from Amazonian Peru. Brittonia 42: 171 - 174. https: // doi. org / 10.2307 / 2807207" type="journal article" year="1990">1990</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFC44B0BFFCAFFE61837F82AFBB5F7AA" author="Thomas, W. W." box="[1080,1137,2028,2054]" pageId="2" pageNumber="283" pagination="51 - 61" refId="ref9481" refString="Thomas, W. W. (2011) Nothotalisia, a new genus of Picramniaceae from tropical America. Brittonia 63: 51 - 61. https: // doi. org / 10.1007 / s 12228 - 010 - 9130 - 8" type="journal article" year="2011">2011</bibRefCitation>
|
||
) and
|
||
<bibRefCitation id="EFC44B0BFFCAFFE618BDF82DFA63F7AA" author="Thomas, W. W. & Hensold, N. & Foster, R. & Ree, R. H. & Soares Neto, R. L." box="[1202,1447,2027,2054]" pageId="2" pageNumber="283" pagination="1239 - 1247" refId="ref9525" refString="Thomas, W. W., Hensold, N., Foster, R., Ree, R. H. & Soares Neto, R. L. (2021) Aenigmanu, a new genus of Picramniaceae from western Amazonia. Taxon 70: 1239 - 1247. https: // doi. org / 10.1002 / tax. 12588" type="journal article" year="2021">
|
||
Thomas
|
||
<emphasis id="B921EAE8FFCAFFE61919F82AFA95F7A9" box="[1302,1361,2027,2053]" italics="true" pageId="2" pageNumber="283">et al.</emphasis>
|
||
(2021)
|
||
</bibRefCitation>
|
||
. Further studies should supplement the morphological description of
|
||
<taxonomicName id="4C554D79FFCAFFE61F9BF7D6FBECF785" authorityName="Londono-Ech. & A. M. Trujillo" authorityYear="2023" box="[916,1064,2063,2090]" class="Magnoliopsida" family="Picramniaceae" genus="Aenigmanu" kingdom="Plantae" order="Picramniales" pageId="2" pageNumber="283" phylum="Tracheophyta" rank="species" species="chococola">
|
||
<emphasis id="B921EAE8FFCAFFE61F9BF7D6FBECF785" box="[916,1064,2063,2090]" italics="true" pageId="2" pageNumber="283">A. chococola</emphasis>
|
||
</taxonomicName>
|
||
provided here, permitting the use of additional diagnostic floral features. Nevertheless, the diagnosis here proposed was verified using all available specimens of
|
||
<taxonomicName id="4C554D79FFCDFFE11D2DFF05FE71FF70" box="[290,437,194,221]" class="Magnoliopsida" family="Picramniaceae" genus="Aenigmanu" kingdom="Plantae" order="Picramniales" pageId="5" pageNumber="286" phylum="Tracheophyta" rank="species" species="alvareziae">
|
||
<emphasis id="B921EAE8FFCDFFE11D2DFF05FE71FF70" box="[290,437,194,221]" italics="true" pageId="5" pageNumber="286">A. alvareziae</emphasis>
|
||
</taxonomicName>
|
||
and distinguishing
|
||
<taxonomicName id="4C554D79FFCDFFE11E9EFF05FCE7FF70" authorityName="Londono-Ech. & A. M. Trujillo" authorityYear="2023" box="[657,803,194,221]" class="Magnoliopsida" family="Picramniaceae" genus="Aenigmanu" kingdom="Plantae" order="Picramniales" pageId="5" pageNumber="286" phylum="Tracheophyta" rank="species" species="chococola">
|
||
<emphasis id="B921EAE8FFCDFFE11E9EFF05FCE7FF70" box="[657,803,194,221]" italics="true" pageId="5" pageNumber="286">A. chococola</emphasis>
|
||
</taxonomicName>
|
||
from any Amazonian specimen of the genus.
|
||
</paragraph>
|
||
<caption id="DF2A6672FFCBFFE71C87F87AFD9DF87E" ID-DOI="http://doi.org/10.5281/zenodo.10150901" ID-Zenodo-Dep="10150901" box="[136,601,1980,2002]" httpUri="https://zenodo.org/record/10150901/files/figure.png" pageId="3" pageNumber="284" startId="3.[136,229,1980,2002]" targetBox="[165,1422,187,1953]" targetPageId="3" targetType="figure">
|
||
<paragraph id="8BEA36FAFFCBFFE71C87F87AFD9DF87E" blockId="3.[136,601,1980,2002]" box="[136,601,1980,2002]" pageId="3" pageNumber="284">
|
||
<emphasis id="B921EAE8FFCBFFE71C87F87AFF3AF87D" bold="true" box="[136,254,1980,2002]" pageId="3" pageNumber="284">FIGURE 1.</emphasis>
|
||
Holotype of
|
||
<taxonomicName id="4C554D79FFCBFFE71D72F87BFD97F87D" authorityName="Londono-Ech. & A. M. Trujillo" authorityYear="2023" box="[381,595,1980,2002]" class="Magnoliopsida" family="Picramniaceae" genus="Aenigmanu" kingdom="Plantae" order="Picramniales" pageId="3" pageNumber="284" phylum="Tracheophyta" rank="species" species="chococola">
|
||
<emphasis id="B921EAE8FFCBFFE71D72F87BFD97F87D" box="[381,595,1980,2002]" italics="true" pageId="3" pageNumber="284">Aenigmanu chococola</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF2A6672FFCCFFE01C87F89DFED7F814" ID-DOI="http://doi.org/10.5281/zenodo.10150903" ID-Zenodo-Dep="10150903" httpUri="https://zenodo.org/record/10150903/files/figure.png" pageId="4" pageNumber="285" startId="4.[136,229,1883,1905]" targetBox="[154,1432,193,1773]" targetPageId="4" targetType="figure">
|
||
<paragraph id="8BEA36FAFFCCFFE01C87F89DFED7F814" blockId="4.[136,1451,1883,1977]" pageId="4" pageNumber="285">
|
||
<emphasis id="B921EAE8FFCCFFE01C87F89DFF3BF8DC" bold="true" box="[136,255,1883,1905]" pageId="4" pageNumber="285">FIGURE 2.</emphasis>
|
||
<taxonomicName id="4C554D79FFCCFFE01D09F89DFE19F8DC" authorityName="Londono-Ech. & A. M. Trujillo" authorityYear="2023" box="[262,477,1883,1904]" class="Magnoliopsida" family="Picramniaceae" genus="Aenigmanu" kingdom="Plantae" order="Picramniales" pageId="4" pageNumber="285" phylum="Tracheophyta" rank="species" species="chococola">
|
||
<emphasis id="B921EAE8FFCCFFE01D09F89DFE19F8DC" box="[262,477,1883,1904]" italics="true" pageId="4" pageNumber="285">Aenigmanu chococola</emphasis>
|
||
</taxonomicName>
|
||
. A. Flowering branchlet of pistillate plant. B. Stipules. C. Detail of leaves blade at very tip, abaxial view. D. Pistillate inflorescence. E. Pistillate flower beginning to develop fruit. (A–E from the holotype; illustration by Ana
|
||
<taxonomicName id="4C554D79FFCCFFE01950F8B9FECCF814" authority="Pareja, HUA" authorityName="Pareja, HUA" class="Arachnida" family="Phalangiidae" genus="Cristina" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="285" phylum="Arthropoda" rank="genus">Cristina Pareja, HUA</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BEA36FAFFCDFFE11CB2FF21FAB5FE3C" blockId="5.[136,1452,158,509]" pageId="5" pageNumber="286">
|
||
Two additional features in foliar architecture of
|
||
<taxonomicName id="4C554D79FFCDFFE11EECFF21FC1AFEAC" authorityName="Londono-Ech. & A. M. Trujillo" authorityYear="2023" box="[739,990,230,257]" class="Magnoliopsida" family="Picramniaceae" genus="Aenigmanu" kingdom="Plantae" order="Picramniales" pageId="5" pageNumber="286" phylum="Tracheophyta" rank="species" species="chococola">
|
||
<emphasis id="B921EAE8FFCDFFE11EECFF21FC1AFEAC" box="[739,990,230,257]" italics="true" pageId="5" pageNumber="286">Aenigmanu chococola</emphasis>
|
||
</taxonomicName>
|
||
can be useful, although their taxonomic relevance depends on further study with more specimens of both species: loops between secondary veins consistently of 2° gauge and monopodial to slightly sympodial (vs. loops noticeably reduced in gauge towards the superjacent secondary vein with course strongly sympodial in
|
||
<taxonomicName id="4C554D79FFCDFFE11EBCFE95FC81FEC0" box="[691,837,338,365]" class="Magnoliopsida" family="Picramniaceae" genus="Aenigmanu" kingdom="Plantae" order="Picramniales" pageId="5" pageNumber="286" phylum="Tracheophyta" rank="species" species="alvareziae">
|
||
<emphasis id="B921EAE8FFCDFFE11EBCFE95FC81FEC0" box="[691,837,338,365]" italics="true" pageId="5" pageNumber="286">A. alvareziae</emphasis>
|
||
</taxonomicName>
|
||
), and veins framework abaxially visible up to 6° giving them a more reticulate appearance (vs. veins abaxially visible up to 5° giving them a less reticulate appearance).
|
||
</paragraph>
|
||
<paragraph id="8BEA36FAFFCDFFE11CB2FE5CFD7DFE51" blockId="5.[136,1452,158,509]" pageId="5" pageNumber="286">
|
||
<bibRefCitation id="EFC44B0BFFCDFFE11CB2FE5CFE6CFE18" author="Thomas, W. W. & Hensold, N. & Foster, R. & Ree, R. H. & Soares Neto, R. L." box="[189,424,410,436]" pageId="5" pageNumber="286" pagination="1239 - 1247" refId="ref9525" refString="Thomas, W. W., Hensold, N., Foster, R., Ree, R. H. & Soares Neto, R. L. (2021) Aenigmanu, a new genus of Picramniaceae from western Amazonia. Taxon 70: 1239 - 1247. https: // doi. org / 10.1002 / tax. 12588" type="journal article" year="2021">
|
||
Thomas
|
||
<emphasis id="B921EAE8FFCDFFE11D10FE5DFE93FE18" box="[287,343,410,436]" italics="true" pageId="5" pageNumber="286">et al.</emphasis>
|
||
(2021)
|
||
</bibRefCitation>
|
||
described the stipules of
|
||
<taxonomicName id="4C554D79FFCDFFE11ECAFE5DFC04FE18" box="[709,960,410,437]" class="Magnoliopsida" family="Picramniaceae" genus="Aenigmanu" kingdom="Plantae" order="Picramniales" pageId="5" pageNumber="286" phylum="Tracheophyta" rank="species" species="alvareziae">
|
||
<emphasis id="B921EAE8FFCDFFE11ECAFE5DFC04FE18" box="[709,960,410,437]" italics="true" pageId="5" pageNumber="286">Aenigmanu alvareziae</emphasis>
|
||
</taxonomicName>
|
||
as “
|
||
<emphasis id="B921EAE8FFCDFFE11FFEFE5DFBD7FE18" box="[1009,1043,411,436]" italics="true" pageId="5" pageNumber="286">ca.</emphasis>
|
||
6.5 ×
|
||
<quantity id="4CAD9B1FFFCDFFE11855FE5CFB70FE18" box="[1114,1204,410,437]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="5" pageNumber="286" unit="mm" value="7.5">7.5 mm</quantity>
|
||
”, which seems to be a typographical error. Based in the available specimens and the original illustrations of
|
||
<taxonomicName id="4C554D79FFCDFFE11849FE79FB1DFE74" box="[1094,1241,446,473]" class="Magnoliopsida" family="Picramniaceae" genus="Aenigmanu" kingdom="Plantae" order="Picramniales" pageId="5" pageNumber="286" phylum="Tracheophyta" rank="species" species="alvareizae">
|
||
<emphasis id="B921EAE8FFCDFFE11849FE79FB1DFE74" box="[1094,1241,446,473]" italics="true" pageId="5" pageNumber="286">A. alvareizae</emphasis>
|
||
</taxonomicName>
|
||
, this should be
|
||
<emphasis id="B921EAE8FFCDFFE11985FE79FA68FE74" box="[1418,1452,447,472]" italics="true" pageId="5" pageNumber="286">ca.</emphasis>
|
||
6.5 ×
|
||
<quantity id="4CAD9B1FFFCDFFE11CC6FE24FEE5FE51" box="[201,289,482,509]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="5" pageNumber="286" unit="mm" value="1.5">1.5 mm</quantity>
|
||
, as in the diagnosis of
|
||
<taxonomicName id="4C554D79FFCDFFE11E2FFE25FD76FE50" authorityName="Londono-Ech. & A. M. Trujillo" authorityYear="2023" box="[544,690,482,509]" class="Magnoliopsida" family="Picramniaceae" genus="Aenigmanu" kingdom="Plantae" order="Picramniales" pageId="5" pageNumber="286" phylum="Tracheophyta" rank="species" species="chococola">
|
||
<emphasis id="B921EAE8FFCDFFE11E2FFE25FD76FE50" box="[544,690,482,509]" italics="true" pageId="5" pageNumber="286">A. chococola</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |