treatments-xml/data/03/FC/87/03FC87E9FF8C0637FF4CCA2CFAC0C71C.xml

<document id="794D8C910880362945016DD8A6F7A702" ID-CLB-Dataset="27891" ID-DOI="10.11646/zootaxa.4526.2.4" ID-GBIF-Dataset="10aed39d-f494-4164-b41f-5c2e6bcb7be4" ID-ISSN="1175-5326" ID-Zenodo-Dep="2611530" ID-ZooBank="541D8D11-DCC0-4E5C-8B23-81D92D0F1BE9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553697344389" checkinUser="plazi" docAuthor="Sobral, Rafael, Grossi, Paschoal C. &amp; De Morais, José W." docDate="2018" docId="03FC87E9FF8C0637FF4CCA2CFAC0C71C" docLanguage="en" docName="zootaxa.4526.2.4.pdf" docOrigin="Zootaxa 4526 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Aegopsis diceratops Sobral &amp; Grossi &amp; De Morais 2018, new species" docType="treatment" docVersion="8" lastPageNumber="190" masterDocId="FFC5FF91FF840638FFDBC840FB0BC106" masterDocTitle="Two new species of Aegopsis Burmeister, 1847 (Coleoptera: Scarabaeidae: Dynastinae) from the central Brazilian Cerrado" masterLastPageNumber="194" masterPageNumber="175" pageNumber="183" updateTime="1698714249358" updateUser="ExternalLinkService">
<mods:mods id="A1119264B324E7A531FAC4BDD68E53C9" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="73D4D106B875829507584ACF3E17F802">
<mods:title id="1B7572D02429CEB14F878166E85BB992">Two new species of Aegopsis Burmeister, 1847 (Coleoptera: Scarabaeidae: Dynastinae) from the central Brazilian Cerrado</mods:title>
</mods:titleInfo>
<mods:name id="C4DD1120DAA2751DA882AF6455257CA0" type="personal">
<mods:role id="CAE01AC6DCF266D359CDB4E2B5CBFC7A">
<mods:roleTerm id="2D589311FDE701C27056343B04ABDE7A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="959AA46966522F564A011755BF7BB2CE">Sobral, Rafael</mods:namePart>
</mods:name>
<mods:name id="64BC3B8314099B5D29EE96886D8CBF24" type="personal">
<mods:role id="AB51A2AFB45E4E70C5C78C8C644C1F93">
<mods:roleTerm id="7587DEC5D7F86B63BB9011436DB24257">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="89C561A5835F8B6086DD0B5A0D5F0329">Grossi, Paschoal C.</mods:namePart>
</mods:name>
<mods:name id="934880668CA338A5DB59A6FA2295B353" type="personal">
<mods:role id="B559FD950BD6541741FF66238E622BBD">
<mods:roleTerm id="AF0BE22B6168C619758FCD4F568F9E9A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0AA9D211FA89B75E41F2858877363E7B">De Morais, José W.</mods:namePart>
</mods:name>
<mods:typeOfResource id="4F45036A0913B2A599B42B123610F8DD">text</mods:typeOfResource>
<mods:relatedItem id="CE0501E3D9B87C892BCDE63953E37285" type="host">
<mods:titleInfo id="702971D1506F22EF03425A07E9992C3A">
<mods:title id="DDE618E05852C1A2A1766AE317E636AE">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="71818C86C171BE309321DD4CF19324B3">
<mods:date id="AD5E089C1C4CC923B74CD34F31CBFAF6">2018</mods:date>
<mods:detail id="56E71AB29FF3C70F4ECDBD5E60F9B4F3" type="pubDate">
<mods:number id="EB6E010C0D03E02F720D699A042496D0">2018-11-29</mods:number>
</mods:detail>
<mods:detail id="DF84D66CCD015976BA357ACD53D1EC52" type="volume">
<mods:number id="FC5CE5FD42BA864EC1EBB696C0CF0525">4526</mods:number>
</mods:detail>
<mods:detail id="3A37CD0E5DC11676A14B488B3E82371B" type="issue">
<mods:number id="A9F4627C08243FAFAFD24E4FE38E6F23">2</mods:number>
</mods:detail>
<mods:extent id="5B35B5E7F2EBC72BA647BEE3F2F4621C" unit="page">
<mods:start id="AFC6DB1C9B09BAF0B1F105B1C8FBBEC8">175</mods:start>
<mods:end id="3820A0541DCEBFA229A17813788CDA74">194</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="0850C02298F833645C5F22571EC53233">journal article</mods:classification>
<mods:identifier id="6F899A405FF90AC460B9478813F7C2A8" type="CLB-Dataset">27891</mods:identifier>
<mods:identifier id="20230091451638E2E30C55772A4CA8C2" type="DOI">10.11646/zootaxa.4526.2.4</mods:identifier>
<mods:identifier id="20ED30019269A846B040FCC51BCDB5FB" type="GBIF-Dataset">10aed39d-f494-4164-b41f-5c2e6bcb7be4</mods:identifier>
<mods:identifier id="53F29F2013F6461721D692FA2288A2E3" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="FA26E6A0C151659F9F41E2E27702530E" type="Zenodo-Dep">2611530</mods:identifier>
<mods:identifier id="5EA1AA102A8ECE5B1FD915B31A482DBB" type="ZooBank">541D8D11-DCC0-4E5C-8B23-81D92D0F1BE9</mods:identifier>
</mods:mods>
<treatment id="03FC87E9FF8C0637FF4CCA2CFAC0C71C" ID-DOI="http://doi.org/10.5281/zenodo.5976893" ID-GBIF-Taxon="154507624" ID-Zenodo-Dep="5976893" LSID="urn:lsid:plazi:treatment:03FC87E9FF8C0637FF4CCA2CFAC0C71C" httpUri="http://treatment.plazi.org/id/03FC87E9FF8C0637FF4CCA2CFAC0C71C" lastPageId="15" lastPageNumber="190" pageId="8" pageNumber="183">
<subSubSection id="C34F6574FF8C0630FF4CCA2CF9FDC380" box="[151,758,620,646]" pageId="8" pageNumber="183" type="nomenclature">
<paragraph id="8BEA36FFFF8C0630FF4CCA2CF9FDC380" blockId="8.[151,758,620,679]" box="[151,758,620,646]" pageId="8" pageNumber="183">
<heading id="D0A28193FF8C0630FF4CCA2CF9FDC380" bold="true" box="[151,758,620,646]" fontSize="11" level="1" pageId="8" pageNumber="183" reason="1">
<emphasis id="B921EAEDFF8C0630FF4CCA2CF9FDC380" bold="true" box="[151,758,620,646]" pageId="8" pageNumber="183">
<taxonomicName id="4C554D7CFF8C0630FF4CCA2CF952C380" ID-CoL="64WV4" authority="Sobral &amp; Grossi" authorityName="Sobral &amp; Grossi &amp; De Morais" authorityYear="2018" box="[151,601,620,646]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="183" phylum="Arthropoda" rank="species" species="diceratops" status="sp. nov.">
<emphasis id="B921EAEDFF8C0630FF4CCA2CFA8DC380" bold="true" box="[151,390,620,646]" italics="true" pageId="8" pageNumber="183">Aegopsis diceratops</emphasis>
Sobral &amp; Grossi
</taxonomicName>
, 
<taxonomicNameLabel id="A2125796FF8C0630FDBDCA2CF9FDC380" box="[614,758,620,646]" pageId="8" pageNumber="183" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34F6574FF8C0630FF4CCACEFA3BC3A1" box="[151,304,654,679]" pageId="8" pageNumber="183" type="description">
<paragraph id="8BEA36FFFF8C0630FF4CCACEFA3BC3A1" blockId="8.[151,758,620,679]" box="[151,304,654,679]" pageId="8" pageNumber="183">
(
<figureCitation id="136E2A7AFF8C0630FF44CACEFA0BC3A1" box="[159,256,654,679]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStart-4="FIGURE 6" captionStart-5="FIGURE 7" captionStart-6="FIGURE 8" captionStart-7="FIGURE 9" captionStartId-0="4.[151,250,1882,1904]" captionStartId-1="5.[151,250,1876,1898]" captionStartId-2="6.[151,250,1693,1715]" captionStartId-3="7.[151,250,1292,1314]" captionStartId-4="9.[151,250,1775,1797]" captionStartId-5="10.[151,250,1892,1914]" captionStartId-6="11.[151,250,1322,1344]" captionStartId-7="12.[151,250,1976,1998]" captionTargetBox-0="[200,1377,193,1853]" captionTargetBox-1="[231,1354,193,1844]" captionTargetBox-2="[151,1415,201,1637]" captionTargetBox-3="[151,1436,193,1270]" captionTargetBox-4="[152,1421,205,1743]" captionTargetBox-5="[185,1389,202,1867]" captionTargetBox-6="[151,1436,193,1283]" captionTargetBox-7="[185,1375,193,1943]" captionTargetId-0="figure@4.[200,1377,193,1853]" captionTargetId-1="figure@5.[231,1354,193,1844]" captionTargetId-2="figure@6.[151,1436,193,1672]" captionTargetId-3="figure@7.[151,1436,193,1270]" captionTargetId-4="figure@9.[151,1436,193,1754]" captionTargetId-5="figure@10.[181,1389,193,1871]" captionTargetId-6="figure@11.[151,1436,193,1283]" captionTargetId-7="figure@12.[185,1375,193,1943]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionTargetPageId-4="9" captionTargetPageId-5="10" captionTargetPageId-6="11" captionTargetPageId-7="12" captionText-0="FIGURE 2. A, Head of Aegopsis bolboceridus male, in dorsal view; B, pronotum of Aegopsis bolboceridus male, in dorsal view; C, head of Aegopsis diceratops male, in dorsal view; D, pronotum of Aegopsis diceratops male, in dorsal view; E, head of Aegopsis vazdemelloi male, in dorsal view; F, pronotum of Aegopsis vazdemelloi male, in dorsal view. Black arrow indicating emarginate apex of clypeus in A, E; white arrow indicating micropunctures on pronotum in F. Scale bars: A, C, F, 2 mm; B, D, E, 3 mm." captionText-1="FIGURE 3. A, Canthus of Aegopsis bolboceridus male, in dorsal view; B, canthus of Aegopsis bolboceridus female, in dorsal view; C, canthus of Aegopsis diceratops male, in dorsal view; D, canthus of Aegopsis diceratops female, in dorsal view; E, canthus of Aegopsis vazdemelloi male, in dorsal view; F, canthus of Aegopsis vazdemelloi female, in dorsal view. White arrow indicating apex inconspicuous of canthi carina in A; black arrows indicating canthi carina split in C; black arrow indicating apex conspicuous of canthi carina in E; black arrows indicating external edge of canthi in B, D, F. Scale bars: 0.5 mm." captionText-2="FIGURE 4. A, Labrum of Aegopsis bolboceridus male, in dorsal view; B, maxillary palpomere IV of Aegopsis bolboceridus male, in dorsal view; C, maxillary palpomere IV of Aegopsis bolboceridus female, in dorsal view; D, labrum of Aegopsis diceratops male, in dorsal view; E, maxillary palpomere IV of Aegopsis diceratops male, in dorsal view; F, maxillary palpomere IV of Aegopsis diceratops female, in dorsal view; G, labrum of Aegopsis vazdemelloi male, in dorsal view; H, maxillary palpomere IV of Aegopsis vazdemelloi male, in dorsal view; I, maxillary palpomere IV of Aegopsis vazdemelloi female, in dorsal view. Arrows indicating the middle of the labrum in A, G; arrows indicating sensorial area in C, I. Scale bars: 0.5 mm." captionText-3="FIGURE 5. A, Mandible of Aegopsis bolboceridus male, in ventral view; B, mandible of Aegopsis diceratops male, in ventral view; C, mandible of Aegopsis vazdemelloi male, in ventral view; D, mentum of Aegopsis bolboceridus male, in ventral view; E, mentum of Aegopsis diceratops male, in ventral view; F, mentum of Aegopsis vazdemelloi male, in ventral view. Arrows indicating respectively apical protuberance and protuberance on internal lobe in A; Arrows indicating the internal lobe in B, C; arrow indicating the mentum carina in E. Scale bars: 0.5 mm." captionText-4="FIGURE 6. A, Aedeagus of Aegopsis bolboceridus, in lateral view; B, parameres of Aegopsis bolboceridus, in ventral view; C, parameres of Aegopsis bolboceridus, in dorsal view; D, aedeagus of Aegopsis diceratops in lateral view; E, parameres of Aegopsis diceratops in ventral view; F, parameres of Aegopsis diceratops in dorsal view; G, aedeagus of Aegopsis vazdemelloi in lateral view; H, parameres of Aegopsis vazdemelloi in ventral view; I, parameres of Aegopsis vazdemelloi in dorsal view. White arrows in the left indicating ventral carina of parameres in B, E, H; white arrows upward indicating inferior edge of parameres in B, E, H. Scale bars: A, D, G, 1 mm; B, C, E, F, H, I, 0.5 mm." captionText-5="FIGURE 7. A, Head of Aegopsis bolboceridus female in dorsal view; B, head of Aegopsis bolboceridus female in lateral view; C, head of Aegopsis diceratops female in dorsal view; D, head of Aegopsis diceratops female in lateral view; E, head of Aegopsis vazdemelloi female in dorsal view; F, head of Aegopsis vazdemelloi female in lateral view. Black arrow indicating head punctures in A; black arrow indicating clypeus trapezoidal in C; white arrow indicating vertex in C; black arrow indicating frontal tubercles in D; white arrow indicating head punctures in E. Scale bars: 1 mm." captionText-6="FIGURE 8. A, Mandible of Aegopsis bolboceridus female in ventral view; B, mandible of Aegopsis diceratops female in ventral view; C, mandible of Aegopsis vazdemelloi female in ventral view; D, mentum of Aegopsis bolboceridus female in ventral view; E, mentum of Aegopsis diceratops female in ventral view; F, mentum of Aegopsis vazdemelloi female in ventral view. Arrows indicating respectively apical protuberance and protuberance on internal lobe in A; arrow indicating middle part of mesal brushes in B; arrow indicating internal lobe in C. Scale bars: 0.5 mm." captionText-7="FIGURE 9. Habitus of Aegopsis diceratops. A, Major male in dorsal view; B, minor male in dorsal view; C, female in dorsal view. Scale bars: 3 mm." httpUri-0="https://zenodo.org/record/2611534/files/figure.png" httpUri-1="https://zenodo.org/record/2611536/files/figure.png" httpUri-2="https://zenodo.org/record/2611538/files/figure.png" httpUri-3="https://zenodo.org/record/2611541/files/figure.png" httpUri-4="https://zenodo.org/record/2611544/files/figure.png" httpUri-5="https://zenodo.org/record/2611546/files/figure.png" httpUri-6="https://zenodo.org/record/2611548/files/figure.png" httpUri-7="https://zenodo.org/record/2611550/files/figure.png" pageId="8" pageNumber="183">Fig. 2–9</figureCitation>
, 
<figureCitation id="136E2A7AFF8C0630FED7CACFFA2CC3A1" box="[268,295,655,679]" captionStart="FIGURE 11" captionStartId="14.[151,250,1333,1355]" captionTargetBox="[150,1437,193,1311]" captionTargetId="figure@14.[150,1437,193,1311]" captionTargetPageId="14" captionText="FIGURE 11. Distribution map of the species in the Aegopsis bolboceridus species group." httpUri="https://zenodo.org/record/2611555/files/figure.png" pageId="8" pageNumber="183">11</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34F6574FF8C0630FF4CCA96FA08C2A5" pageId="8" pageNumber="183" type="diagnosis">
<paragraph id="8BEA36FFFF8C0630FF4CCA96FA08C2A5" blockId="8.[151,1437,726,1975]" pageId="8" pageNumber="183">
<emphasis id="B921EAEDFF8C0630FF4CCA96FA1AC3E9" bold="true" box="[151,273,726,751]" pageId="8" pageNumber="183">Diagnosis.</emphasis>
Males: clypeus subrectangular, apex convex (
<figureCitation id="136E2A7AFF8C0630FCC6CA96F87CC3E9" box="[797,887,726,751]" captionStart="FIGURE 2" captionStartId="4.[151,250,1882,1904]" captionTargetBox="[200,1377,193,1853]" captionTargetId="figure@4.[200,1377,193,1853]" captionTargetPageId="4" captionText="FIGURE 2. A, Head of Aegopsis bolboceridus male, in dorsal view; B, pronotum of Aegopsis bolboceridus male, in dorsal view; C, head of Aegopsis diceratops male, in dorsal view; D, pronotum of Aegopsis diceratops male, in dorsal view; E, head of Aegopsis vazdemelloi male, in dorsal view; F, pronotum of Aegopsis vazdemelloi male, in dorsal view. Black arrow indicating emarginate apex of clypeus in A, E; white arrow indicating micropunctures on pronotum in F. Scale bars: A, C, F, 2 mm; B, D, E, 3 mm." httpUri="https://zenodo.org/record/2611534/files/figure.png" pageId="8" pageNumber="183">Fig. 2C</figureCitation>
); ocular canthi anteriorly protruding, posteriorly acute (
<figureCitation id="136E2A7AFF8C0630FF3ACABAFA33C215" box="[225,312,762,787]" captionStart="FIGURE 3" captionStartId="5.[151,250,1876,1898]" captionTargetBox="[231,1354,193,1844]" captionTargetId="figure@5.[231,1354,193,1844]" captionTargetPageId="5" captionText="FIGURE 3. A, Canthus of Aegopsis bolboceridus male, in dorsal view; B, canthus of Aegopsis bolboceridus female, in dorsal view; C, canthus of Aegopsis diceratops male, in dorsal view; D, canthus of Aegopsis diceratops female, in dorsal view; E, canthus of Aegopsis vazdemelloi male, in dorsal view; F, canthus of Aegopsis vazdemelloi female, in dorsal view. White arrow indicating apex inconspicuous of canthi carina in A; black arrows indicating canthi carina split in C; black arrow indicating apex conspicuous of canthi carina in E; black arrows indicating external edge of canthi in B, D, F. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/2611536/files/figure.png" pageId="8" pageNumber="183">Fig. 3C</figureCitation>
); internal lobe of mandibles partially covered by mesal brush (
<figureCitation id="136E2A7AFF8C0630FC29CABAFF41C215" box="[1010,1098,762,787]" captionStart="FIGURE 5" captionStartId="7.[151,250,1292,1314]" captionTargetBox="[151,1436,193,1270]" captionTargetId="figure@7.[151,1436,193,1270]" captionTargetPageId="7" captionText="FIGURE 5. A, Mandible of Aegopsis bolboceridus male, in ventral view; B, mandible of Aegopsis diceratops male, in ventral view; C, mandible of Aegopsis vazdemelloi male, in ventral view; D, mentum of Aegopsis bolboceridus male, in ventral view; E, mentum of Aegopsis diceratops male, in ventral view; F, mentum of Aegopsis vazdemelloi male, in ventral view. Arrows indicating respectively apical protuberance and protuberance on internal lobe in A; Arrows indicating the internal lobe in B, C; arrow indicating the mentum carina in E. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/2611541/files/figure.png" pageId="8" pageNumber="183">Fig. 5B</figureCitation>
); mentum subtrapezoidal with apical corners carinate (
<figureCitation id="136E2A7AFF8C0630FE79CB5EFAF2C231" box="[418,505,798,823]" captionStart="FIGURE 5" captionStartId="7.[151,250,1292,1314]" captionTargetBox="[151,1436,193,1270]" captionTargetId="figure@7.[151,1436,193,1270]" captionTargetPageId="7" captionText="FIGURE 5. A, Mandible of Aegopsis bolboceridus male, in ventral view; B, mandible of Aegopsis diceratops male, in ventral view; C, mandible of Aegopsis vazdemelloi male, in ventral view; D, mentum of Aegopsis bolboceridus male, in ventral view; E, mentum of Aegopsis diceratops male, in ventral view; F, mentum of Aegopsis vazdemelloi male, in ventral view. Arrows indicating respectively apical protuberance and protuberance on internal lobe in A; Arrows indicating the internal lobe in B, C; arrow indicating the mentum carina in E. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/2611541/files/figure.png" pageId="8" pageNumber="183">Fig. 5E</figureCitation>
); parameres, in ventral view, with apex acute, ventral carina basally round, inferior edge of parameres deeply concave (
<figureCitation id="136E2A7AFF8C0630FDEBCB02F982C25D" box="[560,649,834,859]" captionStart="FIGURE 6" captionStartId="9.[151,250,1775,1797]" captionTargetBox="[152,1421,205,1743]" captionTargetId="figure@9.[151,1436,193,1754]" captionTargetPageId="9" captionText="FIGURE 6. A, Aedeagus of Aegopsis bolboceridus, in lateral view; B, parameres of Aegopsis bolboceridus, in ventral view; C, parameres of Aegopsis bolboceridus, in dorsal view; D, aedeagus of Aegopsis diceratops in lateral view; E, parameres of Aegopsis diceratops in ventral view; F, parameres of Aegopsis diceratops in dorsal view; G, aedeagus of Aegopsis vazdemelloi in lateral view; H, parameres of Aegopsis vazdemelloi in ventral view; I, parameres of Aegopsis vazdemelloi in dorsal view. White arrows in the left indicating ventral carina of parameres in B, E, H; white arrows upward indicating inferior edge of parameres in B, E, H. Scale bars: A, D, G, 1 mm; B, C, E, F, H, I, 0.5 mm." httpUri="https://zenodo.org/record/2611544/files/figure.png" pageId="8" pageNumber="183">Fig. 6E</figureCitation>
). Females: clypeus trapezoidal, punctures absent on middle of vertex (
<figureCitation id="136E2A7AFF8C0630FF44CB26FBF2C279" box="[159,249,870,895]" captionStart="FIGURE 7" captionStartId="10.[151,250,1892,1914]" captionTargetBox="[185,1389,202,1867]" captionTargetId="figure@10.[181,1389,193,1871]" captionTargetPageId="10" captionText="FIGURE 7. A, Head of Aegopsis bolboceridus female in dorsal view; B, head of Aegopsis bolboceridus female in lateral view; C, head of Aegopsis diceratops female in dorsal view; D, head of Aegopsis diceratops female in lateral view; E, head of Aegopsis vazdemelloi female in dorsal view; F, head of Aegopsis vazdemelloi female in lateral view. Black arrow indicating head punctures in A; black arrow indicating clypeus trapezoidal in C; white arrow indicating vertex in C; black arrow indicating frontal tubercles in D; white arrow indicating head punctures in E. Scale bars: 1 mm." httpUri="https://zenodo.org/record/2611546/files/figure.png" pageId="8" pageNumber="183">Fig. 7C</figureCitation>
); frons with two distinct tubercles (
<figureCitation id="136E2A7AFF8C0630FD49CB26F9E5C279" box="[658,750,870,895]" captionStart="FIGURE 7" captionStartId="10.[151,250,1892,1914]" captionTargetBox="[185,1389,202,1867]" captionTargetId="figure@10.[181,1389,193,1871]" captionTargetPageId="10" captionText="FIGURE 7. A, Head of Aegopsis bolboceridus female in dorsal view; B, head of Aegopsis bolboceridus female in lateral view; C, head of Aegopsis diceratops female in dorsal view; D, head of Aegopsis diceratops female in lateral view; E, head of Aegopsis vazdemelloi female in dorsal view; F, head of Aegopsis vazdemelloi female in lateral view. Black arrow indicating head punctures in A; black arrow indicating clypeus trapezoidal in C; white arrow indicating vertex in C; black arrow indicating frontal tubercles in D; white arrow indicating head punctures in E. Scale bars: 1 mm." httpUri="https://zenodo.org/record/2611546/files/figure.png" pageId="8" pageNumber="183">Fig. 7D</figureCitation>
); mentum subtrapezoidal, apical corners with a weak carina (
<figureCitation id="136E2A7AFF8C0630FF44CBCAFBFDC2A5" box="[159,246,906,931]" captionStart="FIGURE 8" captionStartId="11.[151,250,1322,1344]" captionTargetBox="[151,1436,193,1283]" captionTargetId="figure@11.[151,1436,193,1283]" captionTargetPageId="11" captionText="FIGURE 8. A, Mandible of Aegopsis bolboceridus female in ventral view; B, mandible of Aegopsis diceratops female in ventral view; C, mandible of Aegopsis vazdemelloi female in ventral view; D, mentum of Aegopsis bolboceridus female in ventral view; E, mentum of Aegopsis diceratops female in ventral view; F, mentum of Aegopsis vazdemelloi female in ventral view. Arrows indicating respectively apical protuberance and protuberance on internal lobe in A; arrow indicating middle part of mesal brushes in B; arrow indicating internal lobe in C. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/2611548/files/figure.png" pageId="8" pageNumber="183">Fig. 8E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34F6574FF8C0630FF1CCBEEF959C605" pageId="8" pageNumber="183" type="materials_examined">
<paragraph id="8BEA36FFFF8C0630FF1CCBEEF959C605" blockId="8.[151,1437,726,1975]" pageId="8" pageNumber="183">
<emphasis id="B921EAEDFF8C0630FF1CCBEEF966C2C1" bold="true" box="[199,621,942,967]" pageId="8" pageNumber="183">
<typeStatus id="54EE885DFF8C0630FF1CCBEEFA3FC2C1" box="[199,308,942,967]" pageId="8" pageNumber="183" type="holotype">Holotype</typeStatus>
description, male. 
<emphasis id="B921EAEDFF8C0630FDF9CBEEF968C2C1" bold="true" box="[546,611,942,967]" italics="true" pageId="8" pageNumber="183">Color</emphasis>
:
</emphasis>
Surface lustrous; dorsum, venter and appendages, dark brown with a weak metallic shine (
<figureCitation id="136E2A7AFF8C0630FE5ECB92FAD4C2ED" box="[389,479,978,1003]" captionStart="FIGURE 9" captionStartId="12.[151,250,1976,1998]" captionTargetBox="[185,1375,193,1943]" captionTargetId="figure@12.[185,1375,193,1943]" captionTargetPageId="12" captionText="FIGURE 9. Habitus of Aegopsis diceratops. A, Major male in dorsal view; B, minor male in dorsal view; C, female in dorsal view. Scale bars: 3 mm." httpUri="https://zenodo.org/record/2611550/files/figure.png" pageId="8" pageNumber="183">Fig. 9A</figureCitation>
). 
<emphasis id="B921EAEDFF8C0630FE28CB92F931C2ED" bold="true" box="[499,570,978,1003]" pageId="8" pageNumber="183">
<emphasis id="B921EAEDFF8C0630FE28CB92F93AC2ED" bold="true" box="[499,561,978,1003]" italics="true" pageId="8" pageNumber="183">Head</emphasis>
:
</emphasis>
Frons moderately punctate; punctures moderate in size, becoming larger and mixed with smaller punctures posteriorly to clypeus. Cephalic horns moderately punctate, rugopunctate dorsally on outer side; punctures small, becoming larger near base. Clypeus densely punctate, punctures larger and coalescent basally, becoming smaller and fine apically; clypeus subrectangular, apex convex (
<figureCitation id="136E2A7AFF8C0630FB5ACC7EFFE9C551" box="[1153,1250,1086,1111]" captionStart="FIGURE 2" captionStartId="4.[151,250,1882,1904]" captionTargetBox="[200,1377,193,1853]" captionTargetId="figure@4.[200,1377,193,1853]" captionTargetPageId="4" captionText="FIGURE 2. A, Head of Aegopsis bolboceridus male, in dorsal view; B, pronotum of Aegopsis bolboceridus male, in dorsal view; C, head of Aegopsis diceratops male, in dorsal view; D, pronotum of Aegopsis diceratops male, in dorsal view; E, head of Aegopsis vazdemelloi male, in dorsal view; F, pronotum of Aegopsis vazdemelloi male, in dorsal view. Black arrow indicating emarginate apex of clypeus in A, E; white arrow indicating micropunctures on pronotum in F. Scale bars: A, C, F, 2 mm; B, D, E, 3 mm." httpUri="https://zenodo.org/record/2611534/files/figure.png" pageId="8" pageNumber="183">Fig. 2C</figureCitation>
). Ocular canthi anteriorly protruding, posteriorly acute, with a few setae posteriorly; anterior carina disrupted near cephalic horns base (
<figureCitation id="136E2A7AFF8C0630FF0CCCC6FA24C599" box="[215,303,1158,1183]" captionStart="FIGURE 3" captionStartId="5.[151,250,1876,1898]" captionTargetBox="[231,1354,193,1844]" captionTargetId="figure@5.[231,1354,193,1844]" captionTargetPageId="5" captionText="FIGURE 3. A, Canthus of Aegopsis bolboceridus male, in dorsal view; B, canthus of Aegopsis bolboceridus female, in dorsal view; C, canthus of Aegopsis diceratops male, in dorsal view; D, canthus of Aegopsis diceratops female, in dorsal view; E, canthus of Aegopsis vazdemelloi male, in dorsal view; F, canthus of Aegopsis vazdemelloi female, in dorsal view. White arrow indicating apex inconspicuous of canthi carina in A; black arrows indicating canthi carina split in C; black arrow indicating apex conspicuous of canthi carina in E; black arrows indicating external edge of canthi in B, D, F. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/2611536/files/figure.png" pageId="8" pageNumber="183">Fig. 3C</figureCitation>
). 
<emphasis id="B921EAEDFF8C0630FE98CCC6FAD8C599" bold="true" box="[323,467,1158,1183]" pageId="8" pageNumber="183">
<emphasis id="B921EAEDFF8C0630FE98CCC6FAC1C599" bold="true" box="[323,458,1158,1183]" italics="true" pageId="8" pageNumber="183">Mouthparts</emphasis>
:
</emphasis>
Labrum setose, almost semicircular, laterally acute (
<figureCitation id="136E2A7AFF8C0630FBC4CCC6FF73C599" box="[1055,1144,1158,1183]" captionStart="FIGURE 4" captionStartId="6.[151,250,1693,1715]" captionTargetBox="[151,1415,201,1637]" captionTargetId="figure@6.[151,1436,193,1672]" captionTargetPageId="6" captionText="FIGURE 4. A, Labrum of Aegopsis bolboceridus male, in dorsal view; B, maxillary palpomere IV of Aegopsis bolboceridus male, in dorsal view; C, maxillary palpomere IV of Aegopsis bolboceridus female, in dorsal view; D, labrum of Aegopsis diceratops male, in dorsal view; E, maxillary palpomere IV of Aegopsis diceratops male, in dorsal view; F, maxillary palpomere IV of Aegopsis diceratops female, in dorsal view; G, labrum of Aegopsis vazdemelloi male, in dorsal view; H, maxillary palpomere IV of Aegopsis vazdemelloi male, in dorsal view; I, maxillary palpomere IV of Aegopsis vazdemelloi female, in dorsal view. Arrows indicating the middle of the labrum in A, G; arrows indicating sensorial area in C, I. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/2611538/files/figure.png" pageId="8" pageNumber="183">Fig. 4D</figureCitation>
). Mandibles subtriangular with two prominent teeth apically and two ventral carina; in ventral view, apical protuberance absent, internal lobe partially covered by mesal brush (
<figureCitation id="136E2A7AFF8C0630FDC4CC8EF971C5E1" box="[543,634,1230,1255]" captionStart="FIGURE 5" captionStartId="7.[151,250,1292,1314]" captionTargetBox="[151,1436,193,1270]" captionTargetId="figure@7.[151,1436,193,1270]" captionTargetPageId="7" captionText="FIGURE 5. A, Mandible of Aegopsis bolboceridus male, in ventral view; B, mandible of Aegopsis diceratops male, in ventral view; C, mandible of Aegopsis vazdemelloi male, in ventral view; D, mentum of Aegopsis bolboceridus male, in ventral view; E, mentum of Aegopsis diceratops male, in ventral view; F, mentum of Aegopsis vazdemelloi male, in ventral view. Arrows indicating respectively apical protuberance and protuberance on internal lobe in A; Arrows indicating the internal lobe in B, C; arrow indicating the mentum carina in E. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/2611541/files/figure.png" pageId="8" pageNumber="183">Fig. 5B</figureCitation>
). Maxilla with galea subtriangular, teeth absent, external edge basally projecting, crossing external edge of stipes, anterior margin of external edge straight; maxillary palpomere IV four times longer than wide; sensorial area densely striate (
<figureCitation id="136E2A7AFF8C0630FD23CD56F85BC429" box="[760,848,1302,1327]" captionStart="FIGURE 4" captionStartId="6.[151,250,1693,1715]" captionTargetBox="[151,1415,201,1637]" captionTargetId="figure@6.[151,1436,193,1672]" captionTargetPageId="6" captionText="FIGURE 4. A, Labrum of Aegopsis bolboceridus male, in dorsal view; B, maxillary palpomere IV of Aegopsis bolboceridus male, in dorsal view; C, maxillary palpomere IV of Aegopsis bolboceridus female, in dorsal view; D, labrum of Aegopsis diceratops male, in dorsal view; E, maxillary palpomere IV of Aegopsis diceratops male, in dorsal view; F, maxillary palpomere IV of Aegopsis diceratops female, in dorsal view; G, labrum of Aegopsis vazdemelloi male, in dorsal view; H, maxillary palpomere IV of Aegopsis vazdemelloi male, in dorsal view; I, maxillary palpomere IV of Aegopsis vazdemelloi female, in dorsal view. Arrows indicating the middle of the labrum in A, G; arrows indicating sensorial area in C, I. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/2611538/files/figure.png" pageId="8" pageNumber="183">Fig. 4E</figureCitation>
). Mentum subtrapezoidal, apical corners with carina (
<figureCitation id="136E2A7AFF8C0630FF44CD7AFBF2C455" box="[159,249,1338,1363]" captionStart="FIGURE 5" captionStartId="7.[151,250,1292,1314]" captionTargetBox="[151,1436,193,1270]" captionTargetId="figure@7.[151,1436,193,1270]" captionTargetPageId="7" captionText="FIGURE 5. A, Mandible of Aegopsis bolboceridus male, in ventral view; B, mandible of Aegopsis diceratops male, in ventral view; C, mandible of Aegopsis vazdemelloi male, in ventral view; D, mentum of Aegopsis bolboceridus male, in ventral view; E, mentum of Aegopsis diceratops male, in ventral view; F, mentum of Aegopsis vazdemelloi male, in ventral view. Arrows indicating respectively apical protuberance and protuberance on internal lobe in A; Arrows indicating the internal lobe in B, C; arrow indicating the mentum carina in E. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/2611541/files/figure.png" pageId="8" pageNumber="183">Fig. 5E</figureCitation>
). 
<emphasis id="B921EAEDFF8C0630FED5CD7AFA86C455" bold="true" box="[270,397,1338,1363]" pageId="8" pageNumber="183">
<emphasis id="B921EAEDFF8C0630FED5CD7AFA8FC455" bold="true" box="[270,388,1338,1363]" italics="true" pageId="8" pageNumber="183">Prothorax</emphasis>
:
</emphasis>
Pronotum bordered, disc weakly convex and distinct punctate. Thoracic horn strong, apex slightly downturned. Punctures smaller from posterior margin of pronotum to apex of horn, becoming larger and denser on sides (
<figureCitation id="136E2A7AFF8C0630FEA9CDC2FADCC49D" box="[370,471,1410,1435]" captionStart="FIGURE 2" captionStartId="4.[151,250,1882,1904]" captionTargetBox="[200,1377,193,1853]" captionTargetId="figure@4.[200,1377,193,1853]" captionTargetPageId="4" captionText="FIGURE 2. A, Head of Aegopsis bolboceridus male, in dorsal view; B, pronotum of Aegopsis bolboceridus male, in dorsal view; C, head of Aegopsis diceratops male, in dorsal view; D, pronotum of Aegopsis diceratops male, in dorsal view; E, head of Aegopsis vazdemelloi male, in dorsal view; F, pronotum of Aegopsis vazdemelloi male, in dorsal view. Black arrow indicating emarginate apex of clypeus in A, E; white arrow indicating micropunctures on pronotum in F. Scale bars: A, C, F, 2 mm; B, D, E, 3 mm." httpUri="https://zenodo.org/record/2611534/files/figure.png" pageId="8" pageNumber="183">Fig. 2D</figureCitation>
). Anterior angles and lateral edges of horn completely wrinkled. 
<taxonomicName id="4C554D7CFF8C0630FAC5CDC2FA38C4B9" authority=": Subtriangular" authorityName="Subtriangular" class="Trilobita" family="Styginidae" genus="Scutellum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Corynexochida" pageId="8" pageNumber="183" phylum="Arthropoda" rank="genus">
<emphasis id="B921EAEDFF8C0630FAC5CDC2FE97C49D" bold="true" box="[1310,1436,1410,1435]" pageId="8" pageNumber="183">
<emphasis id="B921EAEDFF8C0630FAC5CDC2FE98C49D" bold="true" box="[1310,1427,1410,1435]" italics="true" pageId="8" pageNumber="183">Scutellum</emphasis>
:
</emphasis>
Subtriangular
</taxonomicName>
; punctures strong anteriorly and medially, setose anteriorly. 
<emphasis id="B921EAEDFF8C0630FC09CDE6FF29C4B9" bold="true" box="[978,1058,1446,1471]" pageId="8" pageNumber="183">
<emphasis id="B921EAEDFF8C0630FC09CDE6FF11C4B9" bold="true" box="[978,1050,1446,1471]" italics="true" pageId="8" pageNumber="183">Elytra</emphasis>
:
</emphasis>
Form 1.7 times longer than wide. Surface glabrous, densely covered by large ocellate punctures, microsculpture slightly wrinkled. Elytral punctures generally irregular, with distinct sutural striae; striae not impressed, punctures small. Humeral umbone weak, apical umbone distinct, round, not crossing elytral external edges. 
<emphasis id="B921EAEDFF8C0630FC4FCE53F8D8C72D" bold="true" box="[916,979,1554,1579]" pageId="8" pageNumber="183">
<emphasis id="B921EAEDFF8C0630FC4FCE53F8C1C72C" bold="true" box="[916,970,1555,1578]" italics="true" pageId="8" pageNumber="183">Legs</emphasis>
:
</emphasis>
Protibial surface rugopunctate dorsally, with two setose striae; protibia with three well-developed external teeth posteriorly decreasing in size, punctate on external edges of teeth. 
<emphasis id="B921EAEDFF8C0630FE6BCE1AF92CC775" bold="true" box="[432,551,1626,1651]" pageId="8" pageNumber="183">
<taxonomicName id="4C554D7CFF8C0630FE6BCE1AF914C775" box="[432,543,1626,1651]" class="Actinopterygii" family="Trichomycteridae" genus="Pygidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="183" phylum="Chordata" rank="genus">
<emphasis id="B921EAEDFF8C0630FE6BCE1AF914C775" bold="true" box="[432,543,1626,1651]" italics="true" pageId="8" pageNumber="183">Pygidium</emphasis>
</taxonomicName>
:
</emphasis>
Form 2.5 times wider than longer, strongly convex. Disc and anterior edge glabrous, hirsute laterally and on posterior edge; densely wrinkled on edges; larger and denser punctures medially. 
<emphasis id="B921EAEDFF8C0630FF4CCEE2FA00C7BD" bold="true" box="[151,267,1698,1723]" pageId="8" pageNumber="183">
<emphasis id="B921EAEDFF8C0630FF4CCEE2FA09C7BD" bold="true" box="[151,258,1698,1723]" italics="true" pageId="8" pageNumber="183">Genitalia</emphasis>
:
</emphasis>
Parameres, in dorsal view, as in 
<figureCitation id="136E2A7AFF8C0630FD5FCEE2F9D2C7BD" box="[644,729,1698,1723]" captionStart="FIGURE 6" captionStartId="9.[151,250,1775,1797]" captionTargetBox="[152,1421,205,1743]" captionTargetId="figure@9.[151,1436,193,1754]" captionTargetPageId="9" captionText="FIGURE 6. A, Aedeagus of Aegopsis bolboceridus, in lateral view; B, parameres of Aegopsis bolboceridus, in ventral view; C, parameres of Aegopsis bolboceridus, in dorsal view; D, aedeagus of Aegopsis diceratops in lateral view; E, parameres of Aegopsis diceratops in ventral view; F, parameres of Aegopsis diceratops in dorsal view; G, aedeagus of Aegopsis vazdemelloi in lateral view; H, parameres of Aegopsis vazdemelloi in ventral view; I, parameres of Aegopsis vazdemelloi in dorsal view. White arrows in the left indicating ventral carina of parameres in B, E, H; white arrows upward indicating inferior edge of parameres in B, E, H. Scale bars: A, D, G, 1 mm; B, C, E, F, H, I, 0.5 mm." httpUri="https://zenodo.org/record/2611544/files/figure.png" pageId="8" pageNumber="183">Fig. 6F</figureCitation>
; in lateral view, thicker than apex of phallobase, lateral carina strongly curved at base (
<figureCitation id="136E2A7AFF8C0630FE65CE86F916C7D9" box="[446,541,1734,1759]" captionStart="FIGURE 6" captionStartId="9.[151,250,1775,1797]" captionTargetBox="[152,1421,205,1743]" captionTargetId="figure@9.[151,1436,193,1754]" captionTargetPageId="9" captionText="FIGURE 6. A, Aedeagus of Aegopsis bolboceridus, in lateral view; B, parameres of Aegopsis bolboceridus, in ventral view; C, parameres of Aegopsis bolboceridus, in dorsal view; D, aedeagus of Aegopsis diceratops in lateral view; E, parameres of Aegopsis diceratops in ventral view; F, parameres of Aegopsis diceratops in dorsal view; G, aedeagus of Aegopsis vazdemelloi in lateral view; H, parameres of Aegopsis vazdemelloi in ventral view; I, parameres of Aegopsis vazdemelloi in dorsal view. White arrows in the left indicating ventral carina of parameres in B, E, H; white arrows upward indicating inferior edge of parameres in B, E, H. Scale bars: A, D, G, 1 mm; B, C, E, F, H, I, 0.5 mm." httpUri="https://zenodo.org/record/2611544/files/figure.png" pageId="8" pageNumber="183">Fig. 6D</figureCitation>
); in ventral view, apex acute, ventral carina basally round, inferior edge of parameres deeply emarginate (
<figureCitation id="136E2A7AFF8C0630FE35CEAAF94EC605" box="[494,581,1770,1795]" captionStart="FIGURE 6" captionStartId="9.[151,250,1775,1797]" captionTargetBox="[152,1421,205,1743]" captionTargetId="figure@9.[151,1436,193,1754]" captionTargetPageId="9" captionText="FIGURE 6. A, Aedeagus of Aegopsis bolboceridus, in lateral view; B, parameres of Aegopsis bolboceridus, in ventral view; C, parameres of Aegopsis bolboceridus, in dorsal view; D, aedeagus of Aegopsis diceratops in lateral view; E, parameres of Aegopsis diceratops in ventral view; F, parameres of Aegopsis diceratops in dorsal view; G, aedeagus of Aegopsis vazdemelloi in lateral view; H, parameres of Aegopsis vazdemelloi in ventral view; I, parameres of Aegopsis vazdemelloi in dorsal view. White arrows in the left indicating ventral carina of parameres in B, E, H; white arrows upward indicating inferior edge of parameres in B, E, H. Scale bars: A, D, G, 1 mm; B, C, E, F, H, I, 0.5 mm." httpUri="https://zenodo.org/record/2611544/files/figure.png" pageId="8" pageNumber="183">Fig. 6E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34F6574FF8C0636FF1CCF4EF8ECC4D5" lastPageId="14" lastPageNumber="189" pageId="8" pageNumber="183" type="description">
<paragraph id="8BEA36FFFF8C0630FF1CCF4EF8D7C64C" blockId="8.[151,1437,726,1975]" pageId="8" pageNumber="183">
<emphasis id="B921EAEDFF8C0630FF1CCF4EF901C621" bold="true" box="[199,522,1806,1831]" pageId="8" pageNumber="183">
Measurements of 
<typeStatus id="54EE885DFF8C0630FE47CF4EF90EC621" box="[412,517,1806,1831]" pageId="8" pageNumber="183" type="holotype">holotype</typeStatus>
.
</emphasis>
Body length: 
<quantity id="4CAD9B1AFF8C0630FD70CF4FF819C620" box="[683,786,1807,1831]" metricMagnitude="-2" metricUnit="m" metricValue="2.94" pageId="8" pageNumber="183" unit="mm" value="29.4">29.4 mm</quantity>
. Cephalic horns length: 
<quantity id="4CAD9B1AFF8C0630FBF8CF4EFF76C620" box="[1059,1149,1806,1831]" metricMagnitude="-3" metricUnit="m" metricValue="6.4" pageId="8" pageNumber="183" unit="mm" value="6.4">6.4 mm</quantity>
. Elytral length: 
<quantity id="4CAD9B1AFF8C0630FAEACF4EFE92C620" box="[1329,1433,1806,1831]" metricMagnitude="-2" metricUnit="m" metricValue="1.59" pageId="8" pageNumber="183" unit="mm" value="15.9">15.9 mm</quantity>
. Elytral width: 
<quantity id="4CAD9B1AFF8C0630FEE2CF73FA99C64C" box="[313,402,1843,1867]" metricMagnitude="-3" metricUnit="m" metricValue="8.8" pageId="8" pageNumber="183" unit="mm" value="8.8">8.8 mm</quantity>
. Pronotal width: 
<quantity id="4CAD9B1AFF8C0630FD8ACF73F9B3C64C" box="[593,696,1843,1867]" metricMagnitude="-2" metricUnit="m" metricValue="1.3199999999999998" pageId="8" pageNumber="183" unit="mm" value="13.2">13.2 mm</quantity>
. Protibial length: 
<quantity id="4CAD9B1AFF8C0630FCA4CF73F8D3C64C" box="[895,984,1843,1867]" metricMagnitude="-3" metricUnit="m" metricValue="7.9" pageId="8" pageNumber="183" unit="mm" value="7.9">7.9 mm</quantity>
.
</paragraph>
<paragraph id="8BEA36FFFF8C0630FF1CCF16FF40C6B1" blockId="8.[151,1437,726,1975]" pageId="8" pageNumber="183">
<emphasis id="B921EAEDFF8C0630FF1CCF16F900C669" bold="true" box="[199,523,1878,1903]" pageId="8" pageNumber="183">
<typeStatus id="54EE885DFF8C0630FF1CCF16FA34C669" box="[199,319,1878,1903]" pageId="8" pageNumber="183" type="paratype">Paratypes</typeStatus>
variation, males.
</emphasis>
Only minor males were found. Minor males (
<figureCitation id="136E2A7AFF8C0630FBD3CF16FF56C669" box="[1032,1117,1878,1903]" captionStart="FIGURE 9" captionStartId="12.[151,250,1976,1998]" captionTargetBox="[185,1375,193,1943]" captionTargetId="figure@12.[185,1375,193,1943]" captionTargetPageId="12" captionText="FIGURE 9. Habitus of Aegopsis diceratops. A, Major male in dorsal view; B, minor male in dorsal view; C, female in dorsal view. Scale bars: 3 mm." httpUri="https://zenodo.org/record/2611550/files/figure.png" pageId="8" pageNumber="183">Fig. 9B</figureCitation>
) with cephalic horns as long as vertex. Thoracic horn short, tuberculate, not crossing anterior margin of pronotum. Ocular canthi anteriorly rounder, but still protruding. Punctures of pronotum larger and denser than 
<typeStatus id="54EE885DFF8C0630FC3ACFDEFF4CC6B1" box="[993,1095,1950,1975]" pageId="8" pageNumber="183" type="holotype">holotype</typeStatus>
.
</paragraph>
<caption id="DF2A6677FF8D0631FF4CCEAFF899C698" httpUri="https://zenodo.org/record/2611544/files/figure.png" pageId="9" pageNumber="184" startId="9.[151,250,1775,1797]" targetBox="[152,1421,205,1743]" targetPageId="9">
<paragraph id="8BEA36FFFF8D0631FF4CCEAFF899C698" blockId="9.[151,1437,1775,1950]" pageId="9" pageNumber="184">
<emphasis id="B921EAEDFF8D0631FF4CCEAFFA21C602" bold="true" box="[151,298,1775,1797]" pageId="9" pageNumber="184">FIGURE 6. A</emphasis>
, Aedeagus of 
<taxonomicName id="4C554D7CFF8D0631FE60CEB0F9ABC603" authorityName=": Dechambre &amp; Grossi" authorityYear="1990" box="[443,672,1775,1797]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="184" phylum="Arthropoda" rank="species" species="bolboceridus">
<emphasis id="B921EAEDFF8D0631FE60CEB0F9ABC603" box="[443,672,1775,1797]" italics="true" pageId="9" pageNumber="184">Aegopsis bolboceridus</emphasis>
</taxonomicName>
, in lateral view; 
<emphasis id="B921EAEDFF8D0631FC92CEAFF851C602" bold="true" box="[841,858,1775,1796]" pageId="9" pageNumber="184">B</emphasis>
, parameres of 
<taxonomicName id="4C554D7CFF8D0631FC35CEB0FFD8C603" authorityName=": Dechambre &amp; Grossi" authorityYear="1990" box="[1006,1235,1775,1797]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="184" phylum="Arthropoda" rank="species" species="bolboceridus">
<emphasis id="B921EAEDFF8D0631FC35CEB0FFD8C603" box="[1006,1235,1775,1797]" italics="true" pageId="9" pageNumber="184">Aegopsis bolboceridus</emphasis>
</taxonomicName>
, in ventral view; 
<emphasis id="B921EAEDFF8D0631FA58CEAFFE9EC603" bold="true" box="[1411,1429,1775,1797]" pageId="9" pageNumber="184">C</emphasis>
, parameres of 
<taxonomicName id="4C554D7CFF8D0631FEF3CF4FF91AC622" authorityName=": Dechambre &amp; Grossi" authorityYear="1990" box="[296,529,1806,1828]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="184" phylum="Arthropoda" rank="species" species="bolboceridus">
<emphasis id="B921EAEDFF8D0631FEF3CF4FF91AC622" box="[296,529,1806,1828]" italics="true" pageId="9" pageNumber="184">Aegopsis bolboceridus</emphasis>
</taxonomicName>
, in dorsal view; 
<emphasis id="B921EAEDFF8D0631FD12CF4EF9D0C625" bold="true" box="[713,731,1806,1827]" pageId="9" pageNumber="184">D</emphasis>
, aedeagus of 
<taxonomicName id="4C554D7CFF8D0631FCAACF4FFF4AC622" authorityName="Sobral &amp; Grossi &amp; De Morais" authorityYear="2018" box="[881,1089,1806,1828]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="184" phylum="Arthropoda" rank="species" species="diceratops">
<emphasis id="B921EAEDFF8D0631FCAACF4FFF4AC622" box="[881,1089,1806,1828]" italics="true" pageId="9" pageNumber="184">Aegopsis diceratops</emphasis>
</taxonomicName>
in lateral view; 
<emphasis id="B921EAEDFF8D0631FB2FCF4EFE0EC625" bold="true" box="[1268,1285,1806,1827]" pageId="9" pageNumber="184">E</emphasis>
, parameres of 
<taxonomicName id="4C554D7CFF8D0631FF4CCF6DFA6FC644" authorityName="Sobral &amp; Grossi &amp; De Morais" authorityYear="2018" box="[151,356,1836,1858]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="184" phylum="Arthropoda" rank="species" species="diceratops">
<emphasis id="B921EAEDFF8D0631FF4CCF6DFA6FC644" box="[151,356,1836,1858]" italics="true" pageId="9" pageNumber="184">Aegopsis diceratops</emphasis>
</taxonomicName>
in ventral view; 
<emphasis id="B921EAEDFF8D0631FDCBCF6DF916C644" bold="true" box="[528,541,1837,1858]" pageId="9" pageNumber="184">F</emphasis>
, parameres of 
<taxonomicName id="4C554D7CFF8D0631FD6FCF6DF88AC644" authorityName="Sobral &amp; Grossi &amp; De Morais" authorityYear="2018" box="[692,897,1836,1858]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="184" phylum="Arthropoda" rank="species" species="diceratops">
<emphasis id="B921EAEDFF8D0631FD6FCF6DF88AC644" box="[692,897,1836,1858]" italics="true" pageId="9" pageNumber="184">Aegopsis diceratops</emphasis>
</taxonomicName>
in dorsal view; 
<emphasis id="B921EAEDFF8D0631FBFECF6DFF3DC645" bold="true" box="[1061,1078,1837,1859]" pageId="9" pageNumber="184">G</emphasis>
, aedeagus of 
<taxonomicName id="4C554D7CFF8D0631FB1ACF6DFE97C644" authorityName="Sobral &amp; Grossi &amp; De Morais" authorityYear="2018" box="[1217,1436,1836,1858]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="184" phylum="Arthropoda" rank="species" species="vazdemelloi">
<emphasis id="B921EAEDFF8D0631FB1ACF6DFE97C644" box="[1217,1436,1836,1858]" italics="true" pageId="9" pageNumber="184">Aegopsis vazdemelloi</emphasis>
</taxonomicName>
in lateral view; 
<emphasis id="B921EAEDFF8D0631FEE0CF0BFA44C666" bold="true" box="[315,335,1867,1888]" pageId="9" pageNumber="184">H</emphasis>
, parameres of 
<taxonomicName id="4C554D7CFF8D0631FE31CF0CF9CCC667" authorityName="Sobral &amp; Grossi &amp; De Morais" authorityYear="2018" box="[490,711,1867,1889]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="184" phylum="Arthropoda" rank="species" species="vazdemelloi">
<emphasis id="B921EAEDFF8D0631FE31CF0CF9CCC667" box="[490,711,1867,1889]" italics="true" pageId="9" pageNumber="184">Aegopsis vazdemelloi</emphasis>
</taxonomicName>
in ventral view; 
<emphasis id="B921EAEDFF8D0631FCA1CF0BF88FC666" bold="true" box="[890,900,1867,1888]" pageId="9" pageNumber="184">I</emphasis>
, parameres of 
<taxonomicName id="4C554D7CFF8D0631FBC4CF0CFFF7C667" authorityName="Sobral &amp; Grossi &amp; De Morais" authorityYear="2018" box="[1055,1276,1867,1889]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="184" phylum="Arthropoda" rank="species" species="vazdemelloi">
<emphasis id="B921EAEDFF8D0631FBC4CF0CFFF7C667" box="[1055,1276,1867,1889]" italics="true" pageId="9" pageNumber="184">Aegopsis vazdemelloi</emphasis>
</taxonomicName>
in dorsal view. White arrows in the left indicating ventral carina of parameres in 
<emphasis id="B921EAEDFF8D0631FC9DCF2AF85CC679" bold="true" box="[838,855,1898,1919]" pageId="9" pageNumber="184">B</emphasis>
, 
<emphasis id="B921EAEDFF8D0631FCBDCF2AF87CC679" bold="true" box="[870,887,1898,1919]" pageId="9" pageNumber="184">E</emphasis>
, 
<emphasis id="B921EAEDFF8D0631FC5DCF2AF891C679" bold="true" box="[902,922,1898,1919]" pageId="9" pageNumber="184">H</emphasis>
; white arrows upward indicating inferior edge of parameres in 
<emphasis id="B921EAEDFF8D0631FEC4CFC9FA3BC698" bold="true" box="[287,304,1929,1950]" pageId="9" pageNumber="184">B</emphasis>
, 
<emphasis id="B921EAEDFF8D0631FEE7CFC9FA46C698" bold="true" box="[316,333,1929,1950]" pageId="9" pageNumber="184">E</emphasis>
, 
<emphasis id="B921EAEDFF8D0631FE81CFC9FA65C698" bold="true" box="[346,366,1929,1950]" pageId="9" pageNumber="184">H</emphasis>
. Scale bars: 
<emphasis id="B921EAEDFF8D0631FE34CFC8F90AC69B" bold="true" box="[495,513,1928,1949]" pageId="9" pageNumber="184">A</emphasis>
, 
<emphasis id="B921EAEDFF8D0631FDD5CFC9F92BC698" bold="true" box="[526,544,1929,1950]" pageId="9" pageNumber="184">D</emphasis>
, 
<emphasis id="B921EAEDFF8D0631FDF6CFC8F935C698" bold="true" box="[557,574,1928,1950]" pageId="9" pageNumber="184">G</emphasis>
, 1 mm; 
<emphasis id="B921EAEDFF8D0631FD49CFC9F9A8C698" bold="true" box="[658,675,1929,1950]" pageId="9" pageNumber="184">B</emphasis>
, 
<emphasis id="B921EAEDFF8D0631FD6BCFC8F9C9C698" bold="true" box="[688,706,1928,1950]" pageId="9" pageNumber="184">C</emphasis>
, 
<emphasis id="B921EAEDFF8D0631FD15CFC9F9D4C698" bold="true" box="[718,735,1929,1950]" pageId="9" pageNumber="184">E</emphasis>
, 
<emphasis id="B921EAEDFF8D0631FD37CFC9F9F2C698" bold="true" box="[748,761,1929,1950]" pageId="9" pageNumber="184">F</emphasis>
, 
<emphasis id="B921EAEDFF8D0631FCDDCFC9F811C698" bold="true" box="[774,794,1929,1950]" pageId="9" pageNumber="184">H</emphasis>
, 
<emphasis id="B921EAEDFF8D0631FCFDCFC9F83BC698" bold="true" box="[806,816,1929,1950]" pageId="9" pageNumber="184">I</emphasis>
, 0.5 mm.
</paragraph>
</caption>
<caption id="DF2A6677FF8E0632FF4CCF24FF6BC6F2" httpUri="https://zenodo.org/record/2611546/files/figure.png" pageId="10" pageNumber="185" startId="10.[151,250,1892,1914]" targetBox="[185,1389,202,1867]" targetPageId="10">
<paragraph id="8BEA36FFFF8E0632FF4CCF24FF6BC6F2" blockId="10.[151,1436,1892,2036]" pageId="10" pageNumber="185">
<emphasis id="B921EAEDFF8E0632FF4CCF24FA21C67F" bold="true" box="[151,298,1892,1914]" pageId="10" pageNumber="185">FIGURE 7. A</emphasis>
, Head of 
<taxonomicName id="4C554D7CFF8E0632FE56CF24F978C67C" authorityName=": Dechambre &amp; Grossi" authorityYear="1990" box="[397,627,1892,1914]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="185" phylum="Arthropoda" rank="species" species="bolboceridus">
<emphasis id="B921EAEDFF8E0632FE56CF24F978C67C" box="[397,627,1892,1914]" italics="true" pageId="10" pageNumber="185">Aegopsis bolboceridus</emphasis>
</taxonomicName>
female in dorsal view; 
<emphasis id="B921EAEDFF8E0632FCBBCF24F87AC67F" bold="true" box="[864,881,1892,1913]" pageId="10" pageNumber="185">B</emphasis>
, head of 
<taxonomicName id="4C554D7CFF8E0632FC15CF24FFBFC67C" authorityName=": Dechambre &amp; Grossi" authorityYear="1990" box="[974,1204,1892,1914]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="185" phylum="Arthropoda" rank="species" species="bolboceridus">
<emphasis id="B921EAEDFF8E0632FC15CF24FFBFC67C" box="[974,1204,1892,1914]" italics="true" pageId="10" pageNumber="185">Aegopsis bolboceridus</emphasis>
</taxonomicName>
female in lateral view; 
<emphasis id="B921EAEDFF8E0632FF4CCFC2FBA2C69E" bold="true" box="[151,169,1922,1944]" pageId="10" pageNumber="185">C</emphasis>
, head of 
<taxonomicName id="4C554D7CFF8E0632FECACFC3FAEAC69E" authorityName="Sobral &amp; Grossi &amp; De Morais" authorityYear="2018" box="[273,481,1922,1944]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="185" phylum="Arthropoda" rank="species" species="diceratops">
<emphasis id="B921EAEDFF8E0632FECACFC3FAEAC69E" box="[273,481,1922,1944]" italics="true" pageId="10" pageNumber="185">Aegopsis diceratops</emphasis>
</taxonomicName>
female in dorsal view; 
<emphasis id="B921EAEDFF8E0632FD3ACFC3F9F8C69E" bold="true" box="[737,755,1923,1944]" pageId="10" pageNumber="185">D</emphasis>
, head of 
<taxonomicName id="4C554D7CFF8E0632FC80CFC3FF20C69E" authorityName="Sobral &amp; Grossi &amp; De Morais" authorityYear="2018" box="[859,1067,1922,1944]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="185" phylum="Arthropoda" rank="species" species="diceratops">
<emphasis id="B921EAEDFF8E0632FC80CFC3FF20C69E" box="[859,1067,1922,1944]" italics="true" pageId="10" pageNumber="185">Aegopsis diceratops</emphasis>
</taxonomicName>
female in lateral view; 
<emphasis id="B921EAEDFF8E0632FAF7CFC3FE36C69E" bold="true" box="[1324,1341,1923,1944]" pageId="10" pageNumber="185">E</emphasis>
, head of 
<taxonomicName id="4C554D7CFF8E0632FF4CCFE2FA7FC6B1" authorityName="Sobral &amp; Grossi &amp; De Morais" authorityYear="2018" box="[151,372,1953,1975]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="185" phylum="Arthropoda" rank="species" species="vazdemelloi">
<emphasis id="B921EAEDFF8E0632FF4CCFE2FA7FC6B1" box="[151,372,1953,1975]" italics="true" pageId="10" pageNumber="185">Aegopsis vazdemelloi</emphasis>
</taxonomicName>
female in dorsal view; 
<emphasis id="B921EAEDFF8E0632FDB0CFE1F973C6B0" bold="true" box="[619,632,1953,1974]" pageId="10" pageNumber="185">F</emphasis>
, head of 
<taxonomicName id="4C554D7CFF8E0632FD07CFE2F8B2C6B1" authorityName="Sobral &amp; Grossi &amp; De Morais" authorityYear="2018" box="[732,953,1953,1975]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="185" phylum="Arthropoda" rank="species" species="vazdemelloi">
<emphasis id="B921EAEDFF8E0632FD07CFE2F8B2C6B1" box="[732,953,1953,1975]" italics="true" pageId="10" pageNumber="185">Aegopsis vazdemelloi</emphasis>
</taxonomicName>
female in lateral view. Black arrow indicating head punctures in 
<emphasis id="B921EAEDFF8E0632FE84CF80FA7AC6D3" bold="true" box="[351,369,1984,2005]" pageId="10" pageNumber="185">A</emphasis>
; black arrow indicating clypeus trapezoidal in 
<emphasis id="B921EAEDFF8E0632FCAACF80F888C6D0" bold="true" box="[881,899,1984,2006]" pageId="10" pageNumber="185">C</emphasis>
; white arrow indicating vertex in 
<emphasis id="B921EAEDFF8E0632FB21CF80FE07C6D0" bold="true" box="[1274,1292,1984,2006]" pageId="10" pageNumber="185">C</emphasis>
; black arrow indicating frontal tubercles in 
<emphasis id="B921EAEDFF8E0632FE13CF9FFAD1C6F2" bold="true" box="[456,474,2015,2036]" pageId="10" pageNumber="185">D</emphasis>
; white arrow indicating head punctures in 
<emphasis id="B921EAEDFF8E0632FC57CF9FF896C6F2" bold="true" box="[908,925,2015,2036]" pageId="10" pageNumber="185">E</emphasis>
. Scale bars: 1 mm.
</paragraph>
</caption>
<caption id="DF2A6677FF8F0633FF4CCD6AF8BFC4BD" httpUri="https://zenodo.org/record/2611548/files/figure.png" pageId="11" pageNumber="186" startId="11.[151,250,1322,1344]" targetBox="[151,1436,193,1283]" targetPageId="11">
<paragraph id="8BEA36FFFF8F0633FF4CCD6AF8BFC4BD" blockId="11.[151,1436,1322,1467]" pageId="11" pageNumber="186">
<emphasis id="B921EAEDFF8F0633FF4CCD6AFA3AC439" bold="true" box="[151,305,1322,1344]" pageId="11" pageNumber="186">FIGURE 8. A</emphasis>
, Mandible of 
<taxonomicName id="4C554D7CFF8F0633FE11CD6BF9BFC446" authorityName=": Dechambre &amp; Grossi" authorityYear="1990" box="[458,692,1322,1344]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="186" phylum="Arthropoda" rank="species" species="bolboceridus">
<emphasis id="B921EAEDFF8F0633FE11CD6BF9BFC446" box="[458,692,1322,1344]" italics="true" pageId="11" pageNumber="186">Aegopsis bolboceridus</emphasis>
</taxonomicName>
female in ventral view; 
<emphasis id="B921EAEDFF8F0633FC61CD6AF8C0C439" bold="true" box="[954,971,1322,1343]" pageId="11" pageNumber="186">B</emphasis>
, mandible of 
<taxonomicName id="4C554D7CFF8F0633FBBACD6BFE3BC446" authorityName="Sobral &amp; Grossi &amp; De Morais" authorityYear="2018" box="[1121,1328,1322,1344]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="186" phylum="Arthropoda" rank="species" species="diceratops">
<emphasis id="B921EAEDFF8F0633FBBACD6BFE3BC446" box="[1121,1328,1322,1344]" italics="true" pageId="11" pageNumber="186">Aegopsis diceratops</emphasis>
</taxonomicName>
female in ventral view; 
<emphasis id="B921EAEDFF8F0633FEFCCD09FA32C459" bold="true" box="[295,313,1353,1375]" pageId="11" pageNumber="186">C</emphasis>
, mandible of 
<taxonomicName id="4C554D7CFF8F0633FE16CD09F9A0C459" authorityName="Sobral &amp; Grossi &amp; De Morais" authorityYear="2018" box="[461,683,1353,1375]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="186" phylum="Arthropoda" rank="species" species="vazdemelloi">
<emphasis id="B921EAEDFF8F0633FE16CD09F9A0C459" box="[461,683,1353,1375]" italics="true" pageId="11" pageNumber="186">Aegopsis vazdemelloi</emphasis>
</taxonomicName>
female in ventral view; 
<emphasis id="B921EAEDFF8F0633FC75CD09F8CBC458" bold="true" box="[942,960,1353,1374]" pageId="11" pageNumber="186">D</emphasis>
, mentum of 
<taxonomicName id="4C554D7CFF8F0633FB92CD09FE3AC459" authorityName=": Dechambre &amp; Grossi" authorityYear="1990" box="[1097,1329,1353,1375]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="186" phylum="Arthropoda" rank="species" species="bolboceridus">
<emphasis id="B921EAEDFF8F0633FB92CD09FE3AC459" box="[1097,1329,1353,1375]" italics="true" pageId="11" pageNumber="186">Aegopsis bolboceridus</emphasis>
</taxonomicName>
female in ventral view; 
<emphasis id="B921EAEDFF8F0633FEF8CD28FA3FC47B" bold="true" box="[291,308,1384,1405]" pageId="11" pageNumber="186">E</emphasis>
, mentum of 
<taxonomicName id="4C554D7CFF8F0633FE6CCD28F98FC47B" authorityName="Sobral &amp; Grossi &amp; De Morais" authorityYear="2018" box="[439,644,1383,1405]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="186" phylum="Arthropoda" rank="species" species="diceratops">
<emphasis id="B921EAEDFF8F0633FE6CCD28F98FC47B" box="[439,644,1383,1405]" italics="true" pageId="11" pageNumber="186">Aegopsis diceratops</emphasis>
</taxonomicName>
female in ventral view; 
<emphasis id="B921EAEDFF8F0633FCA6CD28F881C47B" bold="true" box="[893,906,1384,1405]" pageId="11" pageNumber="186">F</emphasis>
, mentum of 
<taxonomicName id="4C554D7CFF8F0633FBD6CD28FFE2C47B" authorityName="Sobral &amp; Grossi &amp; De Morais" authorityYear="2018" box="[1037,1257,1383,1405]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="186" phylum="Arthropoda" rank="species" species="vazdemelloi">
<emphasis id="B921EAEDFF8F0633FBD6CD28FFE2C47B" box="[1037,1257,1383,1405]" italics="true" pageId="11" pageNumber="186">Aegopsis vazdemelloi</emphasis>
</taxonomicName>
female in ventral view. Arrows indicating respectively apical protuberance and protuberance on internal lobe in 
<emphasis id="B921EAEDFF8F0633FB82CDC6FF60C49D" bold="true" box="[1113,1131,1414,1435]" pageId="11" pageNumber="186">A</emphasis>
; arrow indicating middle part of mesal brushes in 
<emphasis id="B921EAEDFF8F0633FEBACDE5FA79C4BC" bold="true" box="[353,370,1445,1466]" pageId="11" pageNumber="186">B</emphasis>
; arrow indicating internal lobe in 
<emphasis id="B921EAEDFF8F0633FD17CDE5F9D5C4BD" bold="true" box="[716,734,1445,1467]" pageId="11" pageNumber="186">C</emphasis>
. Scale bars: 0.5 mm.
</paragraph>
</caption>
<paragraph id="8BEA36FFFF8F0633FF1CCDA7FE7FC725" blockId="11.[151,1436,1511,2003]" pageId="11" pageNumber="186">
<emphasis id="B921EAEDFF8F0633FF1CCDA7F957C706" bold="true" box="[199,604,1511,1536]" pageId="11" pageNumber="186">
Measurements of male 
<typeStatus id="54EE885DFF8F0633FE04CDA7F95CC706" box="[479,599,1511,1536]" pageId="11" pageNumber="186" type="paratype">paratypes</typeStatus>
.
</emphasis>
Body length: 
<quantity id="4CAD9B1AFF8F0633FD24CDA8F8ADC4F9" box="[767,934,1512,1536]" metricMagnitude="-2" metricUnit="m" metricValue="2.045" metricValueMax="2.28" metricValueMin="1.81" pageId="11" pageNumber="186" unit="mm" value="20.45" valueMax="22.8" valueMin="18.1">18.1–22.8 mm</quantity>
. Cephalic horns length: 
<quantity id="4CAD9B1AFF8F0633FB62CDA7FE4FC4F9" box="[1209,1348,1511,1536]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" metricValueMax="1.6" metricValueMin="1.1" pageId="11" pageNumber="186" unit="mm" value="1.35" valueMax="1.6" valueMin="1.1">1.1–1.6 mm</quantity>
. Elytral length: 
<quantity id="4CAD9B1AFF8F0633FF37CE4BFA99C725" box="[236,402,1547,1572]" metricMagnitude="-2" metricUnit="m" metricValue="1.2449999999999999" metricValueMax="1.26" metricValueMin="1.23" pageId="11" pageNumber="186" unit="mm" value="12.45" valueMax="12.6" valueMin="12.3">12.3–12.6 mm</quantity>
. Elytral width: 
<quantity id="4CAD9B1AFF8F0633FDE5CE4BF9C3C725" box="[574,712,1547,1572]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" metricValueMax="7.1" metricValueMin="6.7" pageId="11" pageNumber="186" unit="mm" value="6.9" valueMax="7.1" valueMin="6.7">6.7–7.1 mm</quantity>
. Pronotal width: 
<quantity id="4CAD9B1AFF8F0633FC5CCE4BFF14C725" box="[903,1055,1547,1572]" metricMagnitude="-2" metricUnit="m" metricValue="1.02" metricValueMax="1.09" metricValueMin="0.95" pageId="11" pageNumber="186" unit="mm" value="10.2" valueMax="10.9" valueMin="9.5">9.5–10.9 mm</quantity>
. Protibial length: 
<quantity id="4CAD9B1AFF8F0633FB3DCE4BFE7BC725" box="[1254,1392,1547,1572]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="6.2" metricValueMin="5.8" pageId="11" pageNumber="186" unit="mm" value="6.0" valueMax="6.2" valueMin="5.8">5.8–6.2 mm</quantity>
.
</paragraph>
<paragraph id="8BEA36FFFF8F0633FF1CCE6FFA75C6D5" blockId="11.[151,1436,1511,2003]" pageId="11" pageNumber="186">
<emphasis id="B921EAEDFF8F0633FF1CCE6FF93DC74E" bold="true" box="[199,566,1583,1608]" pageId="11" pageNumber="186">
<typeStatus id="54EE885DFF8F0633FF1CCE6FFA3FC74E" box="[199,308,1583,1608]" pageId="11" pageNumber="186" type="paratype">Paratype</typeStatus>
description, females.
</emphasis>
Females as males in general aspect except the following (
<figureCitation id="136E2A7AFF8F0633FB39CE6FFE36C74E" box="[1250,1341,1583,1608]" captionStart="FIGURE 9" captionStartId="12.[151,250,1976,1998]" captionTargetBox="[185,1375,193,1943]" captionTargetId="figure@12.[185,1375,193,1943]" captionTargetPageId="12" captionText="FIGURE 9. Habitus of Aegopsis diceratops. A, Major male in dorsal view; B, minor male in dorsal view; C, female in dorsal view. Scale bars: 3 mm." httpUri="https://zenodo.org/record/2611550/files/figure.png" pageId="11" pageNumber="186">Fig. 9C</figureCitation>
). 
<emphasis id="B921EAEDFF8F0633FA8FCE6FFE97C74E" bold="true" box="[1364,1436,1583,1608]" pageId="11" pageNumber="186">
<emphasis id="B921EAEDFF8F0633FA8FCE6FFE99C74E" bold="true" box="[1364,1426,1583,1608]" italics="true" pageId="11" pageNumber="186">Head</emphasis>
:
</emphasis>
Cephalic horns absent. Frons with two strong tubercles (
<figureCitation id="136E2A7AFF8F0633FCE9CE13F89BC76A" box="[818,912,1619,1644]" captionStart="FIGURE 7" captionStartId="10.[151,250,1892,1914]" captionTargetBox="[185,1389,202,1867]" captionTargetId="figure@10.[181,1389,193,1871]" captionTargetPageId="10" captionText="FIGURE 7. A, Head of Aegopsis bolboceridus female in dorsal view; B, head of Aegopsis bolboceridus female in lateral view; C, head of Aegopsis diceratops female in dorsal view; D, head of Aegopsis diceratops female in lateral view; E, head of Aegopsis vazdemelloi female in dorsal view; F, head of Aegopsis vazdemelloi female in lateral view. Black arrow indicating head punctures in A; black arrow indicating clypeus trapezoidal in C; white arrow indicating vertex in C; black arrow indicating frontal tubercles in D; white arrow indicating head punctures in E. Scale bars: 1 mm." httpUri="https://zenodo.org/record/2611546/files/figure.png" pageId="11" pageNumber="186">Fig. 7D</figureCitation>
). Clypeus trapezoidal, slightly concave from frontoclypeal suture to apex (
<figureCitation id="136E2A7AFF8F0633FE04CE37F933C796" box="[479,568,1655,1680]" captionStart="FIGURE 7" captionStartId="10.[151,250,1892,1914]" captionTargetBox="[185,1389,202,1867]" captionTargetId="figure@10.[181,1389,193,1871]" captionTargetPageId="10" captionText="FIGURE 7. A, Head of Aegopsis bolboceridus female in dorsal view; B, head of Aegopsis bolboceridus female in lateral view; C, head of Aegopsis diceratops female in dorsal view; D, head of Aegopsis diceratops female in lateral view; E, head of Aegopsis vazdemelloi female in dorsal view; F, head of Aegopsis vazdemelloi female in lateral view. Black arrow indicating head punctures in A; black arrow indicating clypeus trapezoidal in C; white arrow indicating vertex in C; black arrow indicating frontal tubercles in D; white arrow indicating head punctures in E. Scale bars: 1 mm." httpUri="https://zenodo.org/record/2611546/files/figure.png" pageId="11" pageNumber="186">Fig. 7C</figureCitation>
). Surface with punctures larger and denser, especially on clypeus, wrinkles in front of tubercles; punctures absent on vertex near the pronotal margin. Ocular canthi strongly convex anteriorly, concave posteriorly (
<figureCitation id="136E2A7AFF8F0633FE51CEFFFAE3C7DE" box="[394,488,1727,1752]" captionStart="FIGURE 3" captionStartId="5.[151,250,1876,1898]" captionTargetBox="[231,1354,193,1844]" captionTargetId="figure@5.[231,1354,193,1844]" captionTargetPageId="5" captionText="FIGURE 3. A, Canthus of Aegopsis bolboceridus male, in dorsal view; B, canthus of Aegopsis bolboceridus female, in dorsal view; C, canthus of Aegopsis diceratops male, in dorsal view; D, canthus of Aegopsis diceratops female, in dorsal view; E, canthus of Aegopsis vazdemelloi male, in dorsal view; F, canthus of Aegopsis vazdemelloi female, in dorsal view. White arrow indicating apex inconspicuous of canthi carina in A; black arrows indicating canthi carina split in C; black arrow indicating apex conspicuous of canthi carina in E; black arrows indicating external edge of canthi in B, D, F. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/2611536/files/figure.png" pageId="11" pageNumber="186">Fig. 3D</figureCitation>
). 
<emphasis id="B921EAEDFF8F0633FE24CEFFF984C7DE" bold="true" box="[511,655,1727,1752]" pageId="11" pageNumber="186">
<emphasis id="B921EAEDFF8F0633FE24CEFFF98DC7DE" bold="true" box="[511,646,1727,1752]" italics="true" pageId="11" pageNumber="186">Mouthparts</emphasis>
:
</emphasis>
Labrum more convex anteriorly, sides round. Mandibles and teeth shorter, middle part of mesal brushes more concave (
<figureCitation id="136E2A7AFF8F0633FCCACEA3F865C7FA" box="[785,878,1763,1788]" captionStart="FIGURE 8" captionStartId="11.[151,250,1322,1344]" captionTargetBox="[151,1436,193,1283]" captionTargetId="figure@11.[151,1436,193,1283]" captionTargetPageId="11" captionText="FIGURE 8. A, Mandible of Aegopsis bolboceridus female in ventral view; B, mandible of Aegopsis diceratops female in ventral view; C, mandible of Aegopsis vazdemelloi female in ventral view; D, mentum of Aegopsis bolboceridus female in ventral view; E, mentum of Aegopsis diceratops female in ventral view; F, mentum of Aegopsis vazdemelloi female in ventral view. Arrows indicating respectively apical protuberance and protuberance on internal lobe in A; arrow indicating middle part of mesal brushes in B; arrow indicating internal lobe in C. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/2611548/files/figure.png" pageId="11" pageNumber="186">Fig. 8B</figureCitation>
). Maxilla shorter, galea and stipes less hirsute, maxillary palpomere IV 2.9 times longer than wide, sensorial area broader and shorter, striae sparser and almost indistinct (
<figureCitation id="136E2A7AFF8F0633FECFCF6BFA64C642" box="[276,367,1835,1860]" captionStart="FIGURE 4" captionStartId="6.[151,250,1693,1715]" captionTargetBox="[151,1415,201,1637]" captionTargetId="figure@6.[151,1436,193,1672]" captionTargetPageId="6" captionText="FIGURE 4. A, Labrum of Aegopsis bolboceridus male, in dorsal view; B, maxillary palpomere IV of Aegopsis bolboceridus male, in dorsal view; C, maxillary palpomere IV of Aegopsis bolboceridus female, in dorsal view; D, labrum of Aegopsis diceratops male, in dorsal view; E, maxillary palpomere IV of Aegopsis diceratops male, in dorsal view; F, maxillary palpomere IV of Aegopsis diceratops female, in dorsal view; G, labrum of Aegopsis vazdemelloi male, in dorsal view; H, maxillary palpomere IV of Aegopsis vazdemelloi male, in dorsal view; I, maxillary palpomere IV of Aegopsis vazdemelloi female, in dorsal view. Arrows indicating the middle of the labrum in A, G; arrows indicating sensorial area in C, I. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/2611538/files/figure.png" pageId="11" pageNumber="186">Fig. 4F</figureCitation>
). Mentum shorter, apex longer, less hirsute, carina not touching anterior angles (
<figureCitation id="136E2A7AFF8F0633FAE8CF6BFE84C642" box="[1331,1423,1835,1860]" captionStart="FIGURE 8" captionStartId="11.[151,250,1322,1344]" captionTargetBox="[151,1436,193,1283]" captionTargetId="figure@11.[151,1436,193,1283]" captionTargetPageId="11" captionText="FIGURE 8. A, Mandible of Aegopsis bolboceridus female in ventral view; B, mandible of Aegopsis diceratops female in ventral view; C, mandible of Aegopsis vazdemelloi female in ventral view; D, mentum of Aegopsis bolboceridus female in ventral view; E, mentum of Aegopsis diceratops female in ventral view; F, mentum of Aegopsis vazdemelloi female in ventral view. Arrows indicating respectively apical protuberance and protuberance on internal lobe in A; arrow indicating middle part of mesal brushes in B; arrow indicating internal lobe in C. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/2611548/files/figure.png" pageId="11" pageNumber="186">Fig. 8E</figureCitation>
). 
<emphasis id="B921EAEDFF8F0633FF4CCF10FA1EC66E" bold="true" box="[151,277,1871,1896]" pageId="11" pageNumber="186">
<emphasis id="B921EAEDFF8F0633FF4CCF10FA07C661" bold="true" box="[151,268,1872,1895]" italics="true" pageId="11" pageNumber="186">Pronotum</emphasis>
:
</emphasis>
Weakly convex; prothoracic horn absent; punctures large and dense, wrinkles absent. 
<taxonomicName id="4C554D7CFF8F0633FAC5CF0FFBE7C68A" authority=": Surface" authorityName="Surface" class="Trilobita" family="Styginidae" genus="Scutellum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Corynexochida" pageId="11" pageNumber="186" phylum="Arthropoda" rank="genus">
<emphasis id="B921EAEDFF8F0633FAC5CF0FFE97C66E" bold="true" box="[1310,1436,1871,1896]" pageId="11" pageNumber="186">
<emphasis id="B921EAEDFF8F0633FAC5CF0FFE98C66E" bold="true" box="[1310,1427,1871,1896]" italics="true" pageId="11" pageNumber="186">Scutellum</emphasis>
:
</emphasis>
Surface
</taxonomicName>
more punctate, punctures larger. 
<taxonomicName id="4C554D7CFF8F0633FDB1CF33F84EC68A" authority=": Convex" authorityName="Convex" box="[618,837,1907,1932]" class="Actinopterygii" family="Trichomycteridae" genus="Pygidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="11" pageNumber="186" phylum="Chordata" rank="genus">
<emphasis id="B921EAEDFF8F0633FDB1CF33F9E9C68A" bold="true" box="[618,738,1907,1932]" pageId="11" pageNumber="186">
<emphasis id="B921EAEDFF8F0633FDB1CF33F9D3C68A" bold="true" box="[618,728,1907,1932]" italics="true" pageId="11" pageNumber="186">Pygidium</emphasis>
:
</emphasis>
Convex
</taxonomicName>
, slightly concave near apex, apex compact. Strongly hirsute, laterally with long bristles, short bristles at middle. Coalescent punctures denser medially, annular punctures near apex.
</paragraph>
<caption id="DF2A6677FF880634FF4CCFF8FA82C6EA" httpUri="https://zenodo.org/record/2611550/files/figure.png" pageId="12" pageNumber="187" startId="12.[151,250,1976,1998]" targetBox="[185,1375,193,1943]" targetPageId="12">
<paragraph id="8BEA36FFFF880634FF4CCFF8FA82C6EA" blockId="12.[151,1436,1976,2028]" pageId="12" pageNumber="187">
<emphasis id="B921EAEDFF880634FF4CCFF8FA18C6CB" bold="true" box="[151,275,1976,1998]" pageId="12" pageNumber="187">FIGURE 9.</emphasis>
Habitus of 
<taxonomicName id="4C554D7CFF880634FE57CFF9F952C6C8" authorityName="Sobral &amp; Grossi &amp; De Morais" authorityYear="2018" box="[396,601,1976,1998]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="187" phylum="Arthropoda" rank="species" species="diceratops">
<emphasis id="B921EAEDFF880634FE57CFF9F952C6C8" box="[396,601,1976,1998]" italics="true" pageId="12" pageNumber="187">Aegopsis diceratops</emphasis>
</taxonomicName>
. 
<emphasis id="B921EAEDFF880634FDBECFF8F973C6CB" bold="true" box="[613,632,1976,1997]" pageId="12" pageNumber="187">A</emphasis>
, Major male in dorsal view; 
<emphasis id="B921EAEDFF880634FC7BCFF8F8BAC6CB" bold="true" box="[928,945,1976,1997]" pageId="12" pageNumber="187">B</emphasis>
, minor male in dorsal view; 
<emphasis id="B921EAEDFF880634FB03CFF8FFE1C6C8" bold="true" box="[1240,1258,1976,1998]" pageId="12" pageNumber="187">C</emphasis>
, female in dorsal view. Scale bars: 3 mm.
</paragraph>
</caption>
<caption id="DF2A6677FF890635FF4CCFF2FAC6C6E1" httpUri="https://zenodo.org/record/2611553/files/figure.png" pageId="13" pageNumber="188" startId="13.[151,250,1970,1992]" targetBox="[194,1312,193,1920]" targetPageId="13">
<paragraph id="8BEA36FFFF890635FF4CCFF2FAC6C6E1" blockId="13.[151,1436,1970,2023]" pageId="13" pageNumber="188">
<emphasis id="B921EAEDFF890635FF4CCFF2FA29C6C1" bold="true" box="[151,290,1970,1992]" pageId="13" pageNumber="188">FIGURE 10.</emphasis>
Habitus of 
<taxonomicName id="4C554D7CFF890635FE7ACFF3F975C6CE" authorityName="Sobral &amp; Grossi &amp; De Morais" authorityYear="2018" box="[417,638,1970,1992]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="188" phylum="Arthropoda" rank="species" species="vazdemelloi">
<emphasis id="B921EAEDFF890635FE7ACFF3F975C6CE" box="[417,638,1970,1992]" italics="true" pageId="13" pageNumber="188">Aegopsis vazdemelloi</emphasis>
</taxonomicName>
. 
<emphasis id="B921EAEDFF890635FD56CFF2F994C6C1" bold="true" box="[653,671,1970,1991]" pageId="13" pageNumber="188">A</emphasis>
, Major male in dorsal view; 
<emphasis id="B921EAEDFF890635FC0FCFF3F8EEC6CE" bold="true" box="[980,997,1971,1992]" pageId="13" pageNumber="188">B</emphasis>
, minor male in dorsal view; 
<emphasis id="B921EAEDFF890635FAC2CFF2FE20C6CE" bold="true" box="[1305,1323,1970,1992]" pageId="13" pageNumber="188">C</emphasis>
, female in dorsal view. Scale bars: 3 mm.
</paragraph>
</caption>
<caption id="DF2A6677FF8A0636FF4CCD75FF2FC44D" box="[151,1060,1333,1355]" httpUri="https://zenodo.org/record/2611555/files/figure.png" pageId="14" pageNumber="189" startId="14.[151,250,1333,1355]" targetBox="[150,1437,193,1311]" targetPageId="14">
<paragraph id="8BEA36FFFF8A0636FF4CCD75FF2FC44D" blockId="14.[151,1060,1333,1355]" box="[151,1060,1333,1355]" pageId="14" pageNumber="189">
<emphasis id="B921EAEDFF8A0636FF4CCD75FA15C44C" bold="true" box="[151,286,1333,1355]" pageId="14" pageNumber="189">FIGURE 11.</emphasis>
Distribution map of the species in the 
<taxonomicName id="4C554D7CFF8A0636FD7DCD76F886C44D" authorityName=": Dechambre &amp; Grossi" authorityYear="1990" box="[678,909,1333,1355]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="189" phylum="Arthropoda" rank="species" species="bolboceridus">
<emphasis id="B921EAEDFF8A0636FD7DCD76F886C44D" box="[678,909,1333,1355]" italics="true" pageId="14" pageNumber="189">Aegopsis bolboceridus</emphasis>
</taxonomicName>
species group.
</paragraph>
</caption>
<paragraph id="8BEA36FFFF8A0636FF1CCD32FE5EC4A8" blockId="14.[151,1437,1394,2031]" pageId="14" pageNumber="189">
<emphasis id="B921EAEDFF8A0636FF1CCD32F976C48D" bold="true" box="[199,637,1394,1419]" pageId="14" pageNumber="189">
Measurements of female 
<typeStatus id="54EE885DFF8A0636FDDBCD32F973C48D" box="[512,632,1394,1419]" pageId="14" pageNumber="189" type="paratype">paratypes</typeStatus>
.
</emphasis>
Body length: 
<quantity id="4CAD9B1AFF8A0636FCF0CD32F8DEC48C" box="[811,981,1394,1419]" metricMagnitude="-2" metricUnit="m" metricValue="2.315" metricValueMax="2.52" metricValueMin="2.11" pageId="14" pageNumber="189" unit="mm" value="23.15" valueMax="25.2" valueMin="21.1">21.1–25.2 mm</quantity>
. Elytral length: 
<quantity id="4CAD9B1AFF8A0636FB4DCD32FE4BC48C" box="[1174,1344,1394,1419]" metricMagnitude="-2" metricUnit="m" metricValue="1.4849999999999999" metricValueMax="1.55" metricValueMin="1.42" pageId="14" pageNumber="189" unit="mm" value="14.85" valueMax="15.5" valueMin="14.2">14.2–15.5 mm</quantity>
. Elytral width: 
<quantity id="4CAD9B1AFF8A0636FF3ECDD7FA64C4A8" box="[229,367,1431,1455]" metricMagnitude="-3" metricUnit="m" metricValue="7.75" metricValueMax="8.2" metricValueMin="7.3" pageId="14" pageNumber="189" unit="mm" value="7.75" valueMax="8.2" valueMin="7.3">7.3–8.2 mm</quantity>
. Head length: 
<quantity id="4CAD9B1AFF8A0636FDCACDD7F990C4A8" box="[529,667,1431,1455]" metricMagnitude="-3" metricUnit="m" metricValue="3.25" metricValueMax="3.3" metricValueMin="3.2" pageId="14" pageNumber="189" unit="mm" value="3.25" valueMax="3.3" valueMin="3.2">3.2–3.3 mm</quantity>
. Pronotal width: 
<quantity id="4CAD9B1AFF8A0636FC81CDD6FF0BC4A8" box="[858,1024,1430,1455]" metricMagnitude="-2" metricUnit="m" metricValue="1.045" metricValueMax="1.06" metricValueMin="1.03" pageId="14" pageNumber="189" unit="mm" value="10.45" valueMax="10.6" valueMin="10.3">10.3–10.6 mm</quantity>
. Protibial length: 
<quantity id="4CAD9B1AFF8A0636FB1CCDD6FE5AC4A8" box="[1223,1361,1430,1455]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="6.3" metricValueMin="5.7" pageId="14" pageNumber="189" unit="mm" value="6.0" valueMax="6.3" valueMin="5.7">5.7–6.3 mm</quantity>
.
</paragraph>
<paragraph id="8BEA36FFFF8A0636FF1CCDFAF8ECC4D5" blockId="14.[151,1437,1394,2031]" box="[199,999,1466,1491]" pageId="14" pageNumber="189">
<emphasis id="B921EAEDFF8A0636FF1CCDFAFAE4C4D5" bold="true" box="[199,495,1466,1491]" pageId="14" pageNumber="189">Geographic distribution.</emphasis>
<collectingCountry id="F342766FFF8A0636FE2ECDFAF935C4D5" box="[501,574,1466,1491]" name="Brazil" pageId="14" pageNumber="189">Brazil</collectingCountry>
: northwestern 
<collectingRegion id="4991F81DFF8A0636FD39CDFAF873C4D5" box="[738,888,1466,1491]" country="Brazil" name="Minas Gerais" pageId="14" pageNumber="189">Minas Gerais</collectingRegion>
(
<figureCitation id="136E2A7AFF8A0636FC5CCDFAF8D2C4D5" box="[903,985,1466,1491]" captionStart="FIGURE 11" captionStartId="14.[151,250,1333,1355]" captionTargetBox="[150,1437,193,1311]" captionTargetId="figure@14.[150,1437,193,1311]" captionTargetPageId="14" captionText="FIGURE 11. Distribution map of the species in the Aegopsis bolboceridus species group." httpUri="https://zenodo.org/record/2611555/files/figure.png" pageId="14" pageNumber="189">Fig. 11</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34F6574FF8A0636FF1CCD9EFAB7C781" pageId="14" pageNumber="189" type="materials_examined">
<paragraph id="8BEA36FFFF8A0636FF1CCD9EFAB7C781" blockId="14.[151,1437,1394,2031]" pageId="14" pageNumber="189">
<emphasis id="B921EAEDFF8A0636FF1CCD9EF92EC4F1" bold="true" box="[199,549,1502,1527]" pageId="14" pageNumber="189">
Material examined. 
<materialsCitation id="3B3D3CA2FF8A0636FE63CD9EF92EC4F1" ID-GBIF-Occurrence="2029160300" box="[440,549,1502,1527]" collectingDate="2013-10-18" collectorName="C. M. Oliveira" country="Brazil" determinerName="R. Sobral &amp; P. C. Grossi" elevation="978" latitude="-15.180667" location="Minas Gerais" pageId="14" pageNumber="189" specimenCount="978" stateProvince="Minas Gerais" typeStatus="holotype">
<typeStatus id="54EE885DFF8A0636FE63CD9EF92EC4F1" box="[440,549,1502,1527]" pageId="14" pageNumber="189" type="holotype">Holotype</typeStatus>
</materialsCitation>
</emphasis>
male dissected, labeled: a) “ 
<collectingCountry id="F342766FFF8A0636FCBCCD9FF8DAC4F1" box="[871,977,1503,1527]" name="Brazil" pageId="14" pageNumber="189">BRASIL</collectingCountry>
: 
<collectingRegion id="4991F81DFF8A0636FC07CD9EFF7DC4F1" box="[988,1142,1502,1527]" country="Brazil" name="Minas Gerais" pageId="14" pageNumber="189">Minas Gerais</collectingRegion>
. / Buritis. Faz. Querência. / 
<geoCoordinate id="EE615038FF8A0636FF76CE42FA37C71D" box="[173,316,1538,1563]" direction="south" orientation="latitude" pageId="14" pageNumber="189" precision="1" value="-15.180667">15°10'50.4''S</geoCoordinate>
;46°32'20.1'' / W. 
<specimenCount id="9D53FD76FF8A0636FDC2CE43F96AC71D" box="[537,609,1539,1563]" pageId="14" pageNumber="189" type="generic">
<quantity id="4CAD9B1AFF8A0636FDC2CE43F956C71D" box="[537,605,1539,1563]" metricMagnitude="2" metricUnit="m" metricValue="9.78" pageId="14" pageNumber="189" unit="m" value="978.0">
<elevation id="0078D1CCFF8A0636FDC2CE43F956C71D" box="[537,605,1539,1563]" metricMagnitude="2" metricUnit="m" metricValue="9.78" pageId="14" pageNumber="189" unit="m" value="978.0">978m</elevation>
</quantity>
.
</specimenCount>
<collectingDate id="EFAFE9D7FF8A0636FDB4CE43F9E5C71D" box="[623,750,1539,1563]" pageId="14" pageNumber="189" value="2013-10-18">
<date id="FFEB103FFF8A0636FDB4CE43F9E5C71D" box="[623,750,1539,1563]" pageId="14" pageNumber="189" value="2013-10-18">18-X-2013</date>
</collectingDate>
. / 
<collectorName id="26A05329FF8A0636FCCDCE42F8BBC71D" box="[790,944,1538,1563]" pageId="14" pageNumber="189">C.M.Oliveira</collectorName>
”; b) “ 
<taxonomicName id="4C554D7CFF8A0636FBDACE43FFE1C71C" authority="Sobral &amp; Grossi &amp; De Morais, 2018" authorityName="Sobral &amp; Grossi &amp; De Morais" authorityYear="2018" box="[1025,1258,1538,1562]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="189" phylum="Arthropoda" rank="species" species="diceratops" status="sp. nov.">
<emphasis id="B921EAEDFF8A0636FBDACE43FFE1C71C" box="[1025,1258,1538,1562]" italics="true" pageId="14" pageNumber="189">Aegopsis diceratops</emphasis>
</taxonomicName>
<emphasis id="B921EAEDFF8A0636FB22CE42FE4BC71D" bold="true" box="[1273,1344,1538,1563]" pageId="14" pageNumber="189">
<taxonomicNameLabel id="A2125796FF8A0636FB22CE42FE4BC71D" box="[1273,1344,1538,1563]" pageId="14" pageNumber="189" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
/ male 
<typeStatus id="54EE885DFF8A0636FF4CCE67FA36C739" box="[151,317,1575,1599]" pageId="14" pageNumber="189" type="holotype">HOLOTYPUS</typeStatus>
/ 
<determinerName id="7E36DA3BFF8A0636FE89CE67FAB0C739" box="[338,443,1574,1599]" pageId="14" pageNumber="189">R. Sobral</determinerName>
&amp; 
<determinerName id="7E36DA3BFF8A0636FE05CE67F950C739" box="[478,603,1574,1599]" pageId="14" pageNumber="189">P.C. Grossi</determinerName>
det. 2018”. 
<materialsCitation id="3B3D3CA2FF8A0636FD3FCE66FE92C739" ID-GBIF-Occurrence="2029160974" box="[740,1433,1574,1599]" collectingDate="2013-10-18" collectorName="C. M. Oliveira" country="Brazil" determinerName="R. Sobral &amp; P. C. Grossi" elevation="978" latitude="-15.180667" location="Minas Gerais" pageId="14" pageNumber="189" specimenCount="6" specimenCount-female="3" specimenCount-male="3" stateProvince="Minas Gerais" typeStatus="paratype">
<typeStatus id="54EE885DFF8A0636FD3FCE66F857C739" box="[740,860,1574,1599]" pageId="14" pageNumber="189" type="paratype">
<emphasis id="B921EAEDFF8A0636FD3FCE66F857C739" bold="true" box="[740,860,1574,1599]" pageId="14" pageNumber="189">Paratypes</emphasis>
</typeStatus>
: 
<specimenCount id="9D53FD76FF8A0636FCB1CE67F8CBC739" box="[874,960,1574,1599]" pageId="14" pageNumber="189" type="male">3 males</specimenCount>
and 
<specimenCount id="9D53FD76FF8A0636FC2ECE67FF6AC739" box="[1013,1121,1574,1599]" pageId="14" pageNumber="189" type="female">3 females</specimenCount>
with same data as holotype
</materialsCitation>
. 
<typeStatus id="54EE885DFF8A0636FF4CCE0AFBF4C765" box="[151,255,1610,1635]" pageId="14" pageNumber="189" type="holotype">Holotype</typeStatus>
and one female 
<typeStatus id="54EE885DFF8A0636FE1DCE0BF92DC765" box="[454,550,1611,1635]" pageId="14" pageNumber="189" type="paratype">paratype</typeStatus>
at CERPE, one male and one female 
<typeStatus id="54EE885DFF8A0636FC3CCE0BFF4CC765" box="[999,1095,1611,1635]" pageId="14" pageNumber="189" type="paratype">paratype</typeStatus>
at CEMT, one male and one female 
<typeStatus id="54EE885DFF8A0636FF31CE2FFA41C781" box="[234,330,1647,1671]" pageId="14" pageNumber="189" type="paratype">paratype</typeStatus>
at EPGC.
</paragraph>
</subSubSection>
<subSubSection id="C34F6574FF8A0636FF1CCED2FF76C7C9" pageId="14" pageNumber="189" type="etymology">
<paragraph id="8BEA36FFFF8A0636FF1CCED2FF76C7C9" blockId="14.[151,1437,1394,2031]" pageId="14" pageNumber="189">
<emphasis id="B921EAEDFF8A0636FF1CCED2FA45C7AD" bold="true" box="[199,334,1682,1707]" pageId="14" pageNumber="189">Etymology.</emphasis>
From the Greek, “ 
<emphasis id="B921EAEDFF8A0636FDF4CED3F940C7AC" box="[559,587,1683,1706]" italics="true" pageId="14" pageNumber="189">Di</emphasis>
” meaning “two”, “ 
<emphasis id="B921EAEDFF8A0636FCF5CED2F861C7AC" box="[814,874,1682,1706]" italics="true" pageId="14" pageNumber="189">keras</emphasis>
” meaning “horn” and “ 
<emphasis id="B921EAEDFF8A0636FB5FCED3FFA1C7AC" box="[1156,1194,1683,1706]" italics="true" pageId="14" pageNumber="189">ops</emphasis>
” meaning “face”, in reference to the two cephalic horns. This name is an adjective in the nominative singular.
</paragraph>
</subSubSection>
<subSubSection id="C34F6574FF8A0637FF1CCE9AFAC0C71C" lastPageId="15" lastPageNumber="190" pageId="14" pageNumber="189" type="discussion">
<paragraph id="8BEA36FFFF8A0636FF1CCE9AF876C6E9" blockId="14.[151,1437,1394,2031]" pageId="14" pageNumber="189">
<emphasis id="B921EAEDFF8A0636FF1CCE9AFA30C7F5" bold="true" box="[199,315,1754,1779]" pageId="14" pageNumber="189">Remarks.</emphasis>
<taxonomicName id="4C554D7CFF8A0636FE98CE9BF92DC7F4" authorityName="Sobral &amp; Grossi &amp; De Morais" authorityYear="2018" box="[323,550,1754,1778]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="189" phylum="Arthropoda" rank="species" species="diceratops">
<emphasis id="B921EAEDFF8A0636FE98CE9BF92DC7F4" box="[323,550,1754,1778]" italics="true" pageId="14" pageNumber="189">Aegopsis diceratops</emphasis>
</taxonomicName>
is known from Brazilian Cerrado areas, in a rupestrian field area, at 
<quantity id="4CAD9B1AFF8A0636FAEECE9BFE76C7F4" box="[1333,1405,1755,1779]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="14" pageNumber="189" unit="m" value="900.0">900 m</quantity>
of elevation, from the northwest of Minas Gerais state. This species is most similar to 
<taxonomicName id="4C554D7CFF8A0636FB7DCEBFFA55C63D" authority="Dechambre, 1999" authorityName="Dechambre" authorityYear="1999" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="189" phylum="Arthropoda" rank="species" species="chaminadei">
<emphasis id="B921EAEDFF8A0636FB7DCEBFFE97C610" box="[1190,1436,1790,1814]" italics="true" pageId="14" pageNumber="189">Aegopsis chaminadei</emphasis>
<bibRefCitation id="EFC44B0EFF8A0636FF4CCF62FA55C63D" author="Dechambre, R. - P." box="[151,350,1826,1851]" pageId="14" pageNumber="189" pagination="173 - 174" refId="ref8804" refString="Dechambre, R. - P. (1999) Une nouvelle espece d' Aegopsis Burmeister, 1847. Revue Francaise d'Entomologie, Nouvelle-Serie, 21, 173 - 174." type="journal article" year="1999">Dechambre, 1999</bibRefCitation>
</taxonomicName>
as both species have three teeth on protibia, the clypeal apex round, and the same shape of ocular canthus. However, 
<taxonomicName id="4C554D7CFF8A0636FEB4CF07F91AC658" authorityName="Dechambre" authorityYear="1999" box="[367,529,1862,1886]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="189" phylum="Arthropoda" rank="species" species="chaminadei">
<emphasis id="B921EAEDFF8A0636FEB4CF07F91AC658" box="[367,529,1862,1886]" italics="true" pageId="14" pageNumber="189">A. chaminadei</emphasis>
</taxonomicName>
occurs in an intersection area of Peruvian Yungas and Southwest Amazon moist forests, at 
<quantity id="4CAD9B1AFF8A0636FED1CF2AFA5AC684" box="[266,337,1898,1923]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="14" pageNumber="189" unit="m" value="450.0">450 m</quantity>
of elevation, from southeastern 
<collectingCountry id="F342766FFF8A0636FD62CF2BF9E7C684" box="[697,748,1899,1922]" name="Peru" pageId="14" pageNumber="189">Peru</collectingCountry>
(
<bibRefCitation id="EFC44B0EFF8A0636FD21CF2AF8B4C685" author="Dechambre, R. - P." box="[762,959,1898,1923]" pageId="14" pageNumber="189" pagination="173 - 174" refId="ref8804" refString="Dechambre, R. - P. (1999) Une nouvelle espece d' Aegopsis Burmeister, 1847. Revue Francaise d'Entomologie, Nouvelle-Serie, 21, 173 - 174." type="journal article" year="1999">Dechambre 1999</bibRefCitation>
; 
<bibRefCitation id="EFC44B0EFF8A0636FC11CF2AFFCAC685" author="Dinerstein, E. &amp; Olson, D. &amp; Joshi, A. &amp; Vynne, C. &amp; Burgess, N. D. &amp; Wikramanayake, E. &amp; Hahn, N. &amp; Palminteri, S. &amp; Hedao, P. &amp; Noss, R. &amp; Hansen, M. &amp; Locke, H. &amp; Ellis, E. C. &amp; Jones, B. &amp; Barber, C. V. &amp; Hayes, R. &amp; Kormos, C. &amp; Martin, V. &amp; Crist, E. &amp; Sechrest, W. &amp; Price, L. &amp; Baillie, J. E. M. &amp; Weeden, D. &amp; Suckling, K. &amp; Davis, C. &amp; Sizer, N. &amp; Moore, R. &amp; Thau, D. &amp; Birch, T. &amp; Potapov, P. &amp; Turubanova, S. &amp; Tyukavina, A. &amp; de Souza, N. &amp; Pintea, L. &amp; Brito, J. C. &amp; Llewellyn, O. A. &amp; Miller, A. G. &amp; Patzelt, A. &amp; Ghazanfar, S. A. &amp; Timberlake, J. &amp; Kloser, H. &amp; Shennan-Farpon, Y. &amp; Kindt, R. &amp; Lilleso, J. B. &amp; van Breugel, P. &amp; Graudal, L. &amp; Voge, M. &amp; Shammari, K. F. &amp; Saleem, M." box="[970,1217,1898,1923]" pageId="14" pageNumber="189" pagination="534 - 545" refId="ref8878" refString="Dinerstein, E., Olson, D., Joshi, A., Vynne, C., Burgess, N. D., Wikramanayake, E., Hahn, N., Palminteri, S., Hedao, P., Noss, R., Hansen, M., Locke, H., Ellis, E. C., Jones, B., Barber, C. V., Hayes, R., Kormos, C., Martin, V., Crist, E., Sechrest, W., Price, L., Baillie, J. E. M., Weeden, D., Suckling, K., Davis, C., Sizer, N., Moore, R., Thau, D., Birch, T., Potapov, P., Turubanova, S., Tyukavina, A., de Souza, N., Pintea, L., Brito, J. C., Llewellyn, O. A., Miller, A. G., Patzelt, A., Ghazanfar, S. A., Timberlake, J., Kloser, H., Shennan-Farpon, Y., Kindt, R., Lilleso, J. B., van Breugel, P., Graudal, L., Voge, M., Al- Shammari, K. F. &amp; Saleem, M. (2017) An ecoregion-based approach to protecting half the terrestrial realm. BioScience, 67, 534 - 545. https: // doi. org / 10.1093 / biosci / bix 014" type="journal article" year="2017">
Dinerstein 
<emphasis id="B921EAEDFF8A0636FB9ECF2BFF7DC684" box="[1093,1142,1898,1922]" italics="true" pageId="14" pageNumber="189">et al</emphasis>
. 2017
</bibRefCitation>
). Also, the shape of parameres and mentum is very distinct. Whereas 
<taxonomicName id="4C554D7CFF8A0636FD17CFCFF87BC6A0" authorityName="Dechambre" authorityYear="1999" box="[716,880,1934,1958]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="189" phylum="Arthropoda" rank="species" species="chaminadei">
<emphasis id="B921EAEDFF8A0636FD17CFCFF87BC6A0" box="[716,880,1934,1958]" italics="true" pageId="14" pageNumber="189">A. chaminadei</emphasis>
</taxonomicName>
has the apex of parameres almost as long as the base of parameres (
<figureCitation id="136E2A7AFF8A0636FEA8CFF2FAD2C6CD" box="[371,473,1970,1995]" captionStart="FIGURE 12" captionStartId="15.[151,250,901,923]" captionTargetBox="[172,1416,193,879]" captionTargetId="figure@15.[172,1416,193,879]" captionTargetPageId="15" captionText="FIGURE 12. A, Parameres of Aegopsis chaminadei in dorsal view; B, mentum of Aegopsis chaminadei in ventral view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/2611557/files/figure.png" pageId="14" pageNumber="189">Fig. 12A</figureCitation>
) and the mentum round (
<figureCitation id="136E2A7AFF8A0636FD22CFF2F854C6CD" box="[761,863,1970,1995]" captionStart="FIGURE 12" captionStartId="15.[151,250,901,923]" captionTargetBox="[172,1416,193,879]" captionTargetId="figure@15.[172,1416,193,879]" captionTargetPageId="15" captionText="FIGURE 12. A, Parameres of Aegopsis chaminadei in dorsal view; B, mentum of Aegopsis chaminadei in ventral view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/2611557/files/figure.png" pageId="14" pageNumber="189">Fig. 12B</figureCitation>
), 
<taxonomicName id="4C554D7CFF8A0636FCAFCFF3FF01C6CC" authorityName="Sobral &amp; Grossi &amp; De Morais" authorityYear="2018" box="[884,1034,1970,1994]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="189" phylum="Arthropoda" rank="species" species="diceratops">
<emphasis id="B921EAEDFF8A0636FCAFCFF3FF01C6CC" box="[884,1034,1970,1994]" italics="true" pageId="14" pageNumber="189">A. diceratops</emphasis>
</taxonomicName>
has the apex two times longer than base of parameres (
<figureCitation id="136E2A7AFF8A0636FEABCF96FAC8C6E9" box="[368,451,2006,2031]" captionStart="FIGURE 6" captionStartId="9.[151,250,1775,1797]" captionTargetBox="[152,1421,205,1743]" captionTargetId="figure@9.[151,1436,193,1754]" captionTargetPageId="9" captionText="FIGURE 6. A, Aedeagus of Aegopsis bolboceridus, in lateral view; B, parameres of Aegopsis bolboceridus, in ventral view; C, parameres of Aegopsis bolboceridus, in dorsal view; D, aedeagus of Aegopsis diceratops in lateral view; E, parameres of Aegopsis diceratops in ventral view; F, parameres of Aegopsis diceratops in dorsal view; G, aedeagus of Aegopsis vazdemelloi in lateral view; H, parameres of Aegopsis vazdemelloi in ventral view; I, parameres of Aegopsis vazdemelloi in dorsal view. White arrows in the left indicating ventral carina of parameres in B, E, H; white arrows upward indicating inferior edge of parameres in B, E, H. Scale bars: A, D, G, 1 mm; B, C, E, F, H, I, 0.5 mm." httpUri="https://zenodo.org/record/2611544/files/figure.png" pageId="14" pageNumber="189">Fig. 6F</figureCitation>
) and mentum subrectangular (
<figureCitation id="136E2A7AFF8A0636FCC2CF96F864C6E9" box="[793,879,2006,2031]" captionStart="FIGURE 5" captionStartId="7.[151,250,1292,1314]" captionTargetBox="[151,1436,193,1270]" captionTargetId="figure@7.[151,1436,193,1270]" captionTargetPageId="7" captionText="FIGURE 5. A, Mandible of Aegopsis bolboceridus male, in ventral view; B, mandible of Aegopsis diceratops male, in ventral view; C, mandible of Aegopsis vazdemelloi male, in ventral view; D, mentum of Aegopsis bolboceridus male, in ventral view; E, mentum of Aegopsis diceratops male, in ventral view; F, mentum of Aegopsis vazdemelloi male, in ventral view. Arrows indicating respectively apical protuberance and protuberance on internal lobe in A; Arrows indicating the internal lobe in B, C; arrow indicating the mentum carina in E. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/2611541/files/figure.png" pageId="14" pageNumber="189">Fig. 5E</figureCitation>
).
</paragraph>
<caption id="DF2A6677FF8B0637FF4CCBC5FA28C2BF" httpUri="https://zenodo.org/record/2611557/files/figure.png" pageId="15" pageNumber="190" startId="15.[151,250,901,923]" targetBox="[172,1416,193,879]" targetPageId="15">
<paragraph id="8BEA36FFFF8B0637FF4CCBC5FA28C2BF" blockId="15.[151,1436,900,953]" pageId="15" pageNumber="190">
<emphasis id="B921EAEDFF8B0637FF4CCBC5FA33C29C" bold="true" box="[151,312,901,923]" pageId="15" pageNumber="190">FIGURE 12. A</emphasis>
, Parameres of 
<taxonomicName id="4C554D7CFF8B0637FE09CBC5F9A1C29C" authorityName="Dechambre" authorityYear="1999" box="[466,682,900,922]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="190" phylum="Arthropoda" rank="species" species="chaminadei">
<emphasis id="B921EAEDFF8B0637FE09CBC5F9A1C29C" box="[466,682,900,922]" italics="true" pageId="15" pageNumber="190">Aegopsis chaminadei</emphasis>
</taxonomicName>
in dorsal view; 
<emphasis id="B921EAEDFF8B0637FC95CBC5F854C29C" bold="true" box="[846,863,901,922]" pageId="15" pageNumber="190">B</emphasis>
, mentum of 
<taxonomicName id="4C554D7CFF8B0637FC39CBC5FFB1C29C" authorityName="Dechambre" authorityYear="1999" box="[994,1210,900,922]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="190" phylum="Arthropoda" rank="species" species="chaminadei">
<emphasis id="B921EAEDFF8B0637FC39CBC5FFB1C29C" box="[994,1210,900,922]" italics="true" pageId="15" pageNumber="190">Aegopsis chaminadei</emphasis>
</taxonomicName>
in ventral view. Scale bars: 0.5 mm.
</paragraph>
</caption>
<paragraph id="8BEA36FFFF8B0637FF1CCBA5FAC0C71C" blockId="15.[151,1437,997,1563]" pageId="15" pageNumber="190">
Although the distribution of 
<taxonomicName id="4C554D7CFF8B0637FDD0CBA6F9AAC2F8" authorityName="Sobral &amp; Grossi &amp; De Morais" authorityYear="2018" box="[523,673,998,1022]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="190" phylum="Arthropoda" rank="species" species="diceratops">
<emphasis id="B921EAEDFF8B0637FDD0CBA6F9AAC2F8" box="[523,673,998,1022]" italics="true" pageId="15" pageNumber="190">A. diceratops</emphasis>
</taxonomicName>
is geographically close to Brasília municipality (one of the locality records of 
<taxonomicName id="4C554D7CFF8B0637FED5CC4AFAB4C524" authorityName=": Dechambre &amp; Grossi" authorityYear="1990" box="[270,447,1034,1058]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="190" phylum="Arthropoda" rank="species" species="bolboceridus">
<emphasis id="B921EAEDFF8B0637FED5CC4AFAB4C524" box="[270,447,1034,1058]" italics="true" pageId="15" pageNumber="190">A. bolboceridus</emphasis>
</taxonomicName>
), males of both species can be easily distinguished by the clypeal apex, which is convex (
<figureCitation id="136E2A7AFF8B0637FF44CC6DFBFDC540" box="[159,246,1069,1094]" captionStart="FIGURE 2" captionStartId="4.[151,250,1882,1904]" captionTargetBox="[200,1377,193,1853]" captionTargetId="figure@4.[200,1377,193,1853]" captionTargetPageId="4" captionText="FIGURE 2. A, Head of Aegopsis bolboceridus male, in dorsal view; B, pronotum of Aegopsis bolboceridus male, in dorsal view; C, head of Aegopsis diceratops male, in dorsal view; D, pronotum of Aegopsis diceratops male, in dorsal view; E, head of Aegopsis vazdemelloi male, in dorsal view; F, pronotum of Aegopsis vazdemelloi male, in dorsal view. Black arrow indicating emarginate apex of clypeus in A, E; white arrow indicating micropunctures on pronotum in F. Scale bars: A, C, F, 2 mm; B, D, E, 3 mm." httpUri="https://zenodo.org/record/2611534/files/figure.png" pageId="15" pageNumber="190">Fig. 2C</figureCitation>
) in the new species and truncate (
<figureCitation id="136E2A7AFF8B0637FDA4CC6DF9D2C540" box="[639,729,1069,1094]" captionStart="FIGURE 2" captionStartId="4.[151,250,1882,1904]" captionTargetBox="[200,1377,193,1853]" captionTargetId="figure@4.[200,1377,193,1853]" captionTargetPageId="4" captionText="FIGURE 2. A, Head of Aegopsis bolboceridus male, in dorsal view; B, pronotum of Aegopsis bolboceridus male, in dorsal view; C, head of Aegopsis diceratops male, in dorsal view; D, pronotum of Aegopsis diceratops male, in dorsal view; E, head of Aegopsis vazdemelloi male, in dorsal view; F, pronotum of Aegopsis vazdemelloi male, in dorsal view. Black arrow indicating emarginate apex of clypeus in A, E; white arrow indicating micropunctures on pronotum in F. Scale bars: A, C, F, 2 mm; B, D, E, 3 mm." httpUri="https://zenodo.org/record/2611534/files/figure.png" pageId="15" pageNumber="190">Fig. 2A</figureCitation>
) in 
<taxonomicName id="4C554D7CFF8B0637FCDCCC6EF8B1C540" authorityName=": Dechambre &amp; Grossi" authorityYear="1990" box="[775,954,1070,1094]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="190" phylum="Arthropoda" rank="species" species="bolboceridus">
<emphasis id="B921EAEDFF8B0637FCDCCC6EF8B1C540" box="[775,954,1070,1094]" italics="true" pageId="15" pageNumber="190">A. bolboceridus</emphasis>
</taxonomicName>
. Also, 
<taxonomicName id="4C554D7CFF8B0637FBD4CC6EFFADC540" authorityName="Sobral &amp; Grossi &amp; De Morais" authorityYear="2018" box="[1039,1190,1070,1094]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="190" phylum="Arthropoda" rank="species" species="diceratops">
<emphasis id="B921EAEDFF8B0637FBD4CC6EFFADC540" box="[1039,1190,1070,1094]" italics="true" pageId="15" pageNumber="190">A. diceratops</emphasis>
</taxonomicName>
differs by having the ocular canthi anteriorly protruding (
<figureCitation id="136E2A7AFF8B0637FDFCCC12F977C56C" box="[551,636,1106,1131]" captionStart="FIGURE 3" captionStartId="5.[151,250,1876,1898]" captionTargetBox="[231,1354,193,1844]" captionTargetId="figure@5.[231,1354,193,1844]" captionTargetPageId="5" captionText="FIGURE 3. A, Canthus of Aegopsis bolboceridus male, in dorsal view; B, canthus of Aegopsis bolboceridus female, in dorsal view; C, canthus of Aegopsis diceratops male, in dorsal view; D, canthus of Aegopsis diceratops female, in dorsal view; E, canthus of Aegopsis vazdemelloi male, in dorsal view; F, canthus of Aegopsis vazdemelloi female, in dorsal view. White arrow indicating apex inconspicuous of canthi carina in A; black arrows indicating canthi carina split in C; black arrow indicating apex conspicuous of canthi carina in E; black arrows indicating external edge of canthi in B, D, F. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/2611536/files/figure.png" pageId="15" pageNumber="190">Fig. 3C</figureCitation>
) and paired carinae on the apical angles of the mentum (
<figureCitation id="136E2A7AFF8B0637FB2FCC12FE41C56D" box="[1268,1354,1106,1131]" captionStart="FIGURE 5" captionStartId="7.[151,250,1292,1314]" captionTargetBox="[151,1436,193,1270]" captionTargetId="figure@7.[151,1436,193,1270]" captionTargetPageId="7" captionText="FIGURE 5. A, Mandible of Aegopsis bolboceridus male, in ventral view; B, mandible of Aegopsis diceratops male, in ventral view; C, mandible of Aegopsis vazdemelloi male, in ventral view; D, mentum of Aegopsis bolboceridus male, in ventral view; E, mentum of Aegopsis diceratops male, in ventral view; F, mentum of Aegopsis vazdemelloi male, in ventral view. Arrows indicating respectively apical protuberance and protuberance on internal lobe in A; Arrows indicating the internal lobe in B, C; arrow indicating the mentum carina in E. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/2611541/files/figure.png" pageId="15" pageNumber="190">Fig. 5E</figureCitation>
), while in 
<taxonomicName id="4C554D7CFF8B0637FF6FCC36FA6EC588" authorityName=": Dechambre &amp; Grossi" authorityYear="1990" box="[180,357,1142,1166]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="190" phylum="Arthropoda" rank="species" species="bolboceridus">
<emphasis id="B921EAEDFF8B0637FF6FCC36FA6EC588" box="[180,357,1142,1166]" italics="true" pageId="15" pageNumber="190">A. bolboceridus</emphasis>
</taxonomicName>
the ocular canthi are anteriorly straight (
<figureCitation id="136E2A7AFF8B0637FCEACC35F880C588" box="[817,907,1141,1166]" captionStart="FIGURE 3" captionStartId="5.[151,250,1876,1898]" captionTargetBox="[231,1354,193,1844]" captionTargetId="figure@5.[231,1354,193,1844]" captionTargetPageId="5" captionText="FIGURE 3. A, Canthus of Aegopsis bolboceridus male, in dorsal view; B, canthus of Aegopsis bolboceridus female, in dorsal view; C, canthus of Aegopsis diceratops male, in dorsal view; D, canthus of Aegopsis diceratops female, in dorsal view; E, canthus of Aegopsis vazdemelloi male, in dorsal view; F, canthus of Aegopsis vazdemelloi female, in dorsal view. White arrow indicating apex inconspicuous of canthi carina in A; black arrows indicating canthi carina split in C; black arrow indicating apex conspicuous of canthi carina in E; black arrows indicating external edge of canthi in B, D, F. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/2611536/files/figure.png" pageId="15" pageNumber="190">Fig. 3A</figureCitation>
), and the mentum carinae are absent (
<figureCitation id="136E2A7AFF8B0637FAEECC35FE84C588" box="[1333,1423,1141,1166]" captionStart="FIGURE 5" captionStartId="7.[151,250,1292,1314]" captionTargetBox="[151,1436,193,1270]" captionTargetId="figure@7.[151,1436,193,1270]" captionTargetPageId="7" captionText="FIGURE 5. A, Mandible of Aegopsis bolboceridus male, in ventral view; B, mandible of Aegopsis diceratops male, in ventral view; C, mandible of Aegopsis vazdemelloi male, in ventral view; D, mentum of Aegopsis bolboceridus male, in ventral view; E, mentum of Aegopsis diceratops male, in ventral view; F, mentum of Aegopsis vazdemelloi male, in ventral view. Arrows indicating respectively apical protuberance and protuberance on internal lobe in A; Arrows indicating the internal lobe in B, C; arrow indicating the mentum carina in E. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/2611541/files/figure.png" pageId="15" pageNumber="190">Fig. 5D</figureCitation>
). Females of 
<taxonomicName id="4C554D7CFF8B0637FEC2CCDAFAA5C5B4" authorityName="Sobral &amp; Grossi &amp; De Morais" authorityYear="2018" box="[281,430,1178,1202]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="190" phylum="Arthropoda" rank="species" species="diceratops">
<emphasis id="B921EAEDFF8B0637FEC2CCDAFAA5C5B4" box="[281,430,1178,1202]" italics="true" pageId="15" pageNumber="190">A. diceratops</emphasis>
</taxonomicName>
have a trapezoidal clypeus (
<figureCitation id="136E2A7AFF8B0637FD37CCDAF849C5B4" box="[748,834,1178,1203]" captionStart="FIGURE 7" captionStartId="10.[151,250,1892,1914]" captionTargetBox="[185,1389,202,1867]" captionTargetId="figure@10.[181,1389,193,1871]" captionTargetPageId="10" captionText="FIGURE 7. A, Head of Aegopsis bolboceridus female in dorsal view; B, head of Aegopsis bolboceridus female in lateral view; C, head of Aegopsis diceratops female in dorsal view; D, head of Aegopsis diceratops female in lateral view; E, head of Aegopsis vazdemelloi female in dorsal view; F, head of Aegopsis vazdemelloi female in lateral view. Black arrow indicating head punctures in A; black arrow indicating clypeus trapezoidal in C; white arrow indicating vertex in C; black arrow indicating frontal tubercles in D; white arrow indicating head punctures in E. Scale bars: 1 mm." httpUri="https://zenodo.org/record/2611546/files/figure.png" pageId="15" pageNumber="190">Fig. 7C</figureCitation>
) and strong frontal tubercles (
<figureCitation id="136E2A7AFF8B0637FB49CCDAFFE7C5B4" box="[1170,1260,1178,1203]" captionStart="FIGURE 7" captionStartId="10.[151,250,1892,1914]" captionTargetBox="[185,1389,202,1867]" captionTargetId="figure@10.[181,1389,193,1871]" captionTargetPageId="10" captionText="FIGURE 7. A, Head of Aegopsis bolboceridus female in dorsal view; B, head of Aegopsis bolboceridus female in lateral view; C, head of Aegopsis diceratops female in dorsal view; D, head of Aegopsis diceratops female in lateral view; E, head of Aegopsis vazdemelloi female in dorsal view; F, head of Aegopsis vazdemelloi female in lateral view. Black arrow indicating head punctures in A; black arrow indicating clypeus trapezoidal in C; white arrow indicating vertex in C; black arrow indicating frontal tubercles in D; white arrow indicating head punctures in E. Scale bars: 1 mm." httpUri="https://zenodo.org/record/2611546/files/figure.png" pageId="15" pageNumber="190">Fig. 7D</figureCitation>
), while females of 
<taxonomicName id="4C554D7CFF8B0637FF64CCFEFA71C5D0" authorityName=": Dechambre &amp; Grossi" authorityYear="1990" box="[191,378,1214,1238]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="190" phylum="Arthropoda" rank="species" species="bolboceridus">
<emphasis id="B921EAEDFF8B0637FF64CCFEFA71C5D0" box="[191,378,1214,1238]" italics="true" pageId="15" pageNumber="190">A. bolboceridus</emphasis>
</taxonomicName>
have a round clypeus (
<figureCitation id="136E2A7AFF8B0637FD6BCCFDF819C5D0" box="[688,786,1213,1238]" captionStart="FIGURE 7" captionStartId="10.[151,250,1892,1914]" captionTargetBox="[185,1389,202,1867]" captionTargetId="figure@10.[181,1389,193,1871]" captionTargetPageId="10" captionText="FIGURE 7. A, Head of Aegopsis bolboceridus female in dorsal view; B, head of Aegopsis bolboceridus female in lateral view; C, head of Aegopsis diceratops female in dorsal view; D, head of Aegopsis diceratops female in lateral view; E, head of Aegopsis vazdemelloi female in dorsal view; F, head of Aegopsis vazdemelloi female in lateral view. Black arrow indicating head punctures in A; black arrow indicating clypeus trapezoidal in C; white arrow indicating vertex in C; black arrow indicating frontal tubercles in D; white arrow indicating head punctures in E. Scale bars: 1 mm." httpUri="https://zenodo.org/record/2611546/files/figure.png" pageId="15" pageNumber="190">Fig. 7A</figureCitation>
) and inconspicuous frontal tubercles (
<figureCitation id="136E2A7AFF8B0637FB2BCCFDFE58C5D0" box="[1264,1363,1213,1238]" captionStart="FIGURE 7" captionStartId="10.[151,250,1892,1914]" captionTargetBox="[185,1389,202,1867]" captionTargetId="figure@10.[181,1389,193,1871]" captionTargetPageId="10" captionText="FIGURE 7. A, Head of Aegopsis bolboceridus female in dorsal view; B, head of Aegopsis bolboceridus female in lateral view; C, head of Aegopsis diceratops female in dorsal view; D, head of Aegopsis diceratops female in lateral view; E, head of Aegopsis vazdemelloi female in dorsal view; F, head of Aegopsis vazdemelloi female in lateral view. Black arrow indicating head punctures in A; black arrow indicating clypeus trapezoidal in C; white arrow indicating vertex in C; black arrow indicating frontal tubercles in D; white arrow indicating head punctures in E. Scale bars: 1 mm." httpUri="https://zenodo.org/record/2611546/files/figure.png" pageId="15" pageNumber="190">Fig. 7B</figureCitation>
). The morphological differences of these two species, as well as the fact that specimens of 
<taxonomicName id="4C554D7CFF8B0637FB8FCCA2FFE1C5FC" authorityName="Sobral &amp; Grossi &amp; De Morais" authorityYear="2018" box="[1108,1258,1250,1274]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="190" phylum="Arthropoda" rank="species" species="diceratops">
<emphasis id="B921EAEDFF8B0637FB8FCCA2FFE1C5FC" box="[1108,1258,1250,1274]" italics="true" pageId="15" pageNumber="190">A. diceratops</emphasis>
</taxonomicName>
were not found in the same localities as 
<taxonomicName id="4C554D7CFF8B0637FE75CD46F96BC418" authorityName=": Dechambre &amp; Grossi" authorityYear="1990" box="[430,608,1286,1310]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="190" phylum="Arthropoda" rank="species" species="bolboceridus">
<emphasis id="B921EAEDFF8B0637FE75CD46F96BC418" box="[430,608,1286,1310]" italics="true" pageId="15" pageNumber="190">A. bolboceridus</emphasis>
</taxonomicName>
, may indicate that these two species are geographically separated. In the Brazilian Cerrado, the rupestrian field is a phytophysiognomy that occurs above 
<quantity id="4CAD9B1AFF8B0637FBF6CD6AFF7DC444" box="[1069,1142,1322,1346]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="15" pageNumber="190" unit="m" value="900.0">900 m</quantity>
on ridges associated with quartzite, arenite and iron ore outcrops (
<bibRefCitation id="EFC44B0EFF8B0637FD80CD0DF821C460" author="Vasconcelos, M. F." box="[603,810,1357,1382]" pageId="15" pageNumber="190" pagination="241 - 246" refId="ref10029" refString="Vasconcelos, M. F. (2011) O que sao campos rupestres e campos de altitude nos topos de montanha do Leste do Brasil? Revista Brasileira de Botanica, 34, 241 - 246. https: // doi. org / 10.1590 / S 0100 - 84042011000200012" type="journal article" year="2011">Vasconcelos 2011</bibRefCitation>
). Rupestrian fields are similar to islands intercalated by valleys or different phytophysiognomies (
<emphasis id="B921EAEDFF8B0637FD58CD33F9AEC48C" box="[643,677,1395,1418]" italics="true" pageId="15" pageNumber="190">i.e.</emphasis>
, Cerrado fields or peat bogs), so we hypothesize that ancestral lineages of 
<taxonomicName id="4C554D7CFF8B0637FEFACDD6FADDC4A8" authorityName=": Dechambre &amp; Grossi" authorityYear="1990" box="[289,470,1430,1454]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="190" phylum="Arthropoda" rank="species" species="bolboceridus">
<emphasis id="B921EAEDFF8B0637FEFACDD6FADDC4A8" box="[289,470,1430,1454]" italics="true" pageId="15" pageNumber="190">A. bolboceridus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C554D7CFF8B0637FDCECDD6F9A4C4A8" authorityName="Sobral &amp; Grossi &amp; De Morais" authorityYear="2018" box="[533,687,1430,1454]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="190" phylum="Arthropoda" rank="species" species="diceratops">
<emphasis id="B921EAEDFF8B0637FDCECDD6F9A4C4A8" box="[533,687,1430,1454]" italics="true" pageId="15" pageNumber="190">A. diceratops</emphasis>
</taxonomicName>
may have been isolated by the Neogene uplift of the Brazilian Plateau and Quaternary climate oscillations, in which the ridges, lowlands, and phytophysiognomies were shaped. This has been observed in some species of lizards (
<bibRefCitation id="EFC44B0EFF8B0637FD0CCD9DF8C4C4F0" author="Domingos, F. M. C. B. &amp; Bosque, R. J. &amp; Cassimiro, J. &amp; Colli, G. R. &amp; Rodrigues, M. T. &amp; Santos, M. G. &amp; Beheregaray, L. B." box="[727,975,1501,1526]" pageId="15" pageNumber="190" pagination="113 - 124" refId="ref9181" refString="Domingos, F. M. C. B., Bosque, R. J., Cassimiro, J., Colli, G. R., Rodrigues, M. T., Santos, M. G. &amp; Beheregaray, L. B. (2014) Out of the deep: cryptic speciation in a Neotropical gecko (Squamata, Phyllodactylidae) revealed by species delimitation methods. Molecular Phylogenetics and Evolution, 80, 113 - 124. https: // doi. org / 10.1016 / j. ympev. 2014.07.022" type="journal article" year="2014">
Domingos 
<emphasis id="B921EAEDFF8B0637FC8FCD9FF886C4F0" box="[852,909,1502,1526]" italics="true" pageId="15" pageNumber="190">et al.</emphasis>
2014
</bibRefCitation>
) and presumed for some species of dung beetles (
<bibRefCitation id="EFC44B0EFF8B0637FF28CE42FAB4C71C" author="Nunes, R. V. &amp; Frizzas, M. R. &amp; Vaz-de-Mello, F. Z." box="[243,447,1538,1562]" pageId="15" pageNumber="190" pagination="125 - 129" refId="ref9547" refString="Nunes, R. V., Frizzas, M. R. &amp; Vaz-de-Mello, F. Z. (2012) Scarabaeinae (Coleoptera: Scarabaeidae) of a rupestrian field at Cafuringa, Distrito Federal, Brazil: commented list of species. Biota Neotropica, 12, 125 - 129. https: // doi. org / 10.1590 / S 1676 - 06032012000400013" type="journal article" year="2012">
Nunes 
<emphasis id="B921EAEDFF8B0637FE98CE43FA77C71C" box="[323,380,1538,1562]" italics="true" pageId="15" pageNumber="190">et al.</emphasis>
2012
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>