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<document id="2D3FB6FBF76B18E67E3C96ABC1E2B059" ID-DOI="10.5281/zenodo.281652" ID-GBIF-Dataset="1d6d96e1-5b92-4633-b769-d9e7ab8fa254" ID-ISSN="1175-5326" ID-Zenodo-Dep="281652" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460495559621" checkinUser="plazi" docAuthor="Chan, Benny K. K. &amp; Hayashi, Ryota" docDate="2012" docId="03FC87D4FFD38A17C7D722E753F9F81F" docLanguage="en" docName="zt03367p048.pdf" docOrigin="Zootaxa 3367" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Oxynaspis ryukyuensis Chan &amp; Hayashi, 2012, n. sp." docType="treatment" docVersion="9" lastPageNumber="34" masterDocId="FFC5FFACFFDF8A1AC74020105772FFD1" masterDocTitle="Epibiotic barnacles (Crustacea: Cirripedia: Thoracica) collected by the Kumejima 2009 Expedition, with descriptions of two new species *" masterLastPageNumber="48" masterPageNumber="21" pageNumber="33" updateTime="1698310858039" updateUser="plazi">
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<mods:title id="E3AF83C398CEC4C321329BF363169DC3">Epibiotic barnacles (Crustacea: Cirripedia: Thoracica) collected by the Kumejima 2009 Expedition, with descriptions of two new species *</mods:title>
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<mods:namePart id="5CF71B43CE84908F11EC19788330C0D2">Chan, Benny K. K.</mods:namePart>
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<mods:namePart id="71F6C68BA21F165AF3BE25E453535509">Hayashi, Ryota</mods:namePart>
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<treatment id="03FC87D4FFD38A17C7D722E753F9F81F" ID-DOI="http://doi.org/10.5281/zenodo.6174491" ID-GBIF-Taxon="119520510" ID-Zenodo-Dep="6174491" LSID="urn:lsid:zoobank.org:act:4B6FE07F-1B1E-4461-9AEB-6A65F3EEF220" httpUri="http://treatment.plazi.org/id/03FC87D4FFD38A17C7D722E753F9F81F" lastPageId="13" lastPageNumber="34" pageId="12" pageNumber="33">
<subSubSection id="C34F6549FFD38A16C7D722E75635FCE2" pageId="12" pageNumber="33" type="nomenclature">
<paragraph id="8BEA36C2FFD38A16C7D722E75684FCC0" blockId="12.[151,502,759,819]" box="[151,502,759,785]" pageId="12" pageNumber="33">
<heading id="D0A281AEFFD38A16C7D722E75684FCC0" bold="true" box="[151,502,759,785]" fontSize="11" level="1" pageId="12" pageNumber="33" reason="1">
<emphasis id="B921EAD0FFD38A16C7D722E75684FCC0" bold="true" box="[151,502,759,785]" pageId="12" pageNumber="33">
<taxonomicName id="4C554D41FFD38A16C7D722E756DFFCC0" box="[151,429,759,785]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="12" pageNumber="33" phylum="Arthropoda" rank="species" species="ryukyuensis" status="sp. nov.">
<emphasis id="B921EAD0FFD38A16C7D722E756DFFCC0" bold="true" box="[151,429,759,785]" italics="true" pageId="12" pageNumber="33">Oxynaspis ryukyuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21257ABFFD38A16C6F322E75684FCC0" box="[435,502,759,785]" pageId="12" pageNumber="33" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BEA36C2FFD38A16C7D7230B5635FCE2" blockId="12.[151,502,759,819]" box="[151,327,795,819]" pageId="12" pageNumber="33">
(
<figureCitation id="136E2A47FFD38A16C7DF230B5794FCE2" box="[159,230,795,819]" captionStart="FIGURE 2" captionStartId="2.[151,244,1869,1890]" captionTargetBox="[153,1433,917,1847]" captionTargetId="figure@2.[153,1433,917,1848]" captionTargetPageId="2" captionText="FIGURE 2. Kumejima epibiotic barnacles. A, Calantica arcuata n. sp. (Left, holotype, RUMF-ZC- 1554, middle, paratype, RUMF- ZC- 1555, note the tip of carina was broken, right, paratype, NMNS- 6690 - 001); B, Euscalpellum c. f. squamoum Hiro, 1937 (left, RUMF-ZC- 1556, right ASIZCR- 000225); C, Oxynaspis celata Darwin, 1852 (RUMF-ZC- 1557); D, Oxynaspis ryukyuensis n. sp. (holotype, RUMF-ZC- 1558); E, Poecilasma obliqua (Hoek, 1907) (RUMF-ZC- 1560); F, Platylepas hexastylos (Fabricius, 1798) (RUMF-ZC- 1561). A-E, lateral view for stalked barnacles; F, anterior view for Platylepas hexastylos. Scale bars in mm to indicate the size of specimens." httpUri="https://zenodo.org/record/281654/files/figure.png" pageId="12" pageNumber="33">Figs 2</figureCitation>
D, 911)
</paragraph>
</subSubSection>
<subSubSection id="C34F6549FFD38A16C7D723725383FC13" pageId="12" pageNumber="33" type="materials_examined">
<paragraph id="8BEA36C2FFD38A16C7D723725383FC13" blockId="12.[151,1436,866,2034]" pageId="12" pageNumber="33">
<emphasis id="B921EAD0FFD38A16C7D7237256F3FCAA" bold="true" box="[151,385,866,891]" pageId="12" pageNumber="33">Material examined.</emphasis>
<typeStatus id="54EE8860FFD38A16C6C923725684FCAB" box="[393,502,866,890]" pageId="12" pageNumber="33" type="holotype">Holotype</typeStatus>
: RUMF-ZC-1558, one specimen (CL
<quantity id="4CAD9B27FFD38A16C4E62372548CFCAB" box="[934,1022,866,890]" metricMagnitude="-3" metricUnit="m" metricValue="5.4" pageId="12" pageNumber="33" unit="mm" value="5.4">5.4 mm</quantity>
) on an antipatharian coral; trawl 75,
<geoCoordinate id="EE615005FFD38A16C7D723965659FC4F" box="[151,299,902,926]" direction="north" orientation="latitude" pageId="12" pageNumber="33" precision="1" value="26.326433">26°19.586´N</geoCoordinate>
,
<geoCoordinate id="EE615005FFD38A16C679239656AAFC4F" box="[313,472,902,926]" direction="east" orientation="longitude" pageId="12" pageNumber="33" precision="1" value="126.70755">126°42.453´E</geoCoordinate>
,
<date id="FFEB1002FFD38A16C6A5239655F6FC4F" box="[485,644,902,926]" pageId="12" pageNumber="33" value="2009-11-19">19 Nov. 2009</date>
, depth
<quantity id="4CAD9B27FFD38A16C5982396542EFC4F" box="[728,860,902,926]" metricMagnitude="2" metricUnit="m" metricValue="1.465" metricValueMax="1.56" metricValueMin="1.3699999999999999" pageId="12" pageNumber="33" unit="m" value="146.5" valueMax="156.0" valueMin="137.0">
137
<specimenCount id="9D53FD4BFFD38A16C4502396542EFC4F" box="[784,860,902,926]" pageId="12" pageNumber="33" type="generic">156 m</specimenCount>
</quantity>
.
<typeStatus id="54EE8860FFD38A16C428239754ABFC4E" box="[872,985,903,927]" pageId="12" pageNumber="33" type="paratype">Paratypes</typeStatus>
: RUMF-ZC-1559, one specimen (CL
<quantity id="4CAD9B27FFD38A16C2CD239657B5FC13" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="12" pageNumber="33" unit="mm" value="5.0">5 mm</quantity>
), same data as
<typeStatus id="54EE8860FFD38A16C62E23BA56A6FC13" box="[366,468,938,962]" pageId="12" pageNumber="33" type="holotype">holotype</typeStatus>
. NHMS-6690-002, one specimen (CL
<quantity id="4CAD9B27FFD38A16C4C623BA5492FC13" box="[902,992,938,962]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="12" pageNumber="33" unit="mm" value="5.8">5.8 mm</quantity>
), same data as
<typeStatus id="54EE8860FFD38A16C3C823BA539FFC13" box="[1160,1261,938,962]" pageId="12" pageNumber="33" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C34F6549FFD38A16C78723DD5405FB80" pageId="12" pageNumber="33" type="diagnosis">
<paragraph id="8BEA36C2FFD38A16C78723DD5405FB80" blockId="12.[151,1436,866,2034]" pageId="12" pageNumber="33">
<emphasis id="B921EAD0FFD38A16C78723DD5633FC37" bold="true" box="[199,321,973,998]" pageId="12" pageNumber="33">Diagnosis.</emphasis>
Capitulum elongated, with 5 completely calcified, thin plates. Tergum triangular, apex acute, high. Scutum quadrangular, occludent margin almost parallel with carinal margin. Carina slightly concave, umbo of carina located very close to basal region of carina, length of distal arm of carina about 7.5 times length of basal arm. Two filamentary appendages on dorsal side of somatic body.
</paragraph>
</subSubSection>
<subSubSection id="C34F6549FFD38A16C787244C5210F905" pageId="12" pageNumber="33" type="description">
<paragraph id="8BEA36C2FFD38A16C787244C53B7FAF6" blockId="12.[151,1436,866,2034]" pageId="12" pageNumber="33">
<emphasis id="B921EAD0FFD38A16C787244C562BFBA4" bold="true" box="[199,345,1116,1141]" pageId="12" pageNumber="33">Description.</emphasis>
Hermaphrodite, capitulum narrow, surface covered by antipatharian; 5 fully calcified thin plates, covering whole capitulum (
<figureCitation id="136E2A47FFD38A16C68E24905565FB49" box="[462,535,1152,1176]" captionStart="FIGURE 2" captionStartId="2.[151,244,1869,1890]" captionTargetBox="[153,1433,917,1847]" captionTargetId="figure@2.[153,1433,917,1848]" captionTargetPageId="2" captionText="FIGURE 2. Kumejima epibiotic barnacles. A, Calantica arcuata n. sp. (Left, holotype, RUMF-ZC- 1554, middle, paratype, RUMF- ZC- 1555, note the tip of carina was broken, right, paratype, NMNS- 6690 - 001); B, Euscalpellum c. f. squamoum Hiro, 1937 (left, RUMF-ZC- 1556, right ASIZCR- 000225); C, Oxynaspis celata Darwin, 1852 (RUMF-ZC- 1557); D, Oxynaspis ryukyuensis n. sp. (holotype, RUMF-ZC- 1558); E, Poecilasma obliqua (Hoek, 1907) (RUMF-ZC- 1560); F, Platylepas hexastylos (Fabricius, 1798) (RUMF-ZC- 1561). A-E, lateral view for stalked barnacles; F, anterior view for Platylepas hexastylos. Scale bars in mm to indicate the size of specimens." httpUri="https://zenodo.org/record/281654/files/figure.png" pageId="12" pageNumber="33">Figs 2</figureCitation>
D, 9A). Carina slightly concave, apex reaching middle of carinal margin of tergum, umbo located very close to basal region of carinal margin, base non-forked (
<figureCitation id="136E2A47FFD38A16C36824B45305FB6D" box="[1064,1143,1188,1212]" captionStart="FIGURE 11" captionStartId="16.[151,244,1934,1955]" captionTargetBox="[156,1432,159,1933]" captionTargetId="figure@16.[135,1451,137,1948]" captionTargetPageId="16" captionText="FIGURE 11. Oxynaspis ryukyuensis n. sp. (holotype, RUMF-ZC- 1558). A, scutum, dorsal view, B, tergum, dorsal view, C, carina, lateral view, D, carina, basal view showing non-forked base, u—umbo, E, simple type setae on cirrus I; F, distal region of cirrus II; G, distal region of cirrus III; H. intermediate segment of posterior ramus of cirrus VI; I, somatic body, showing location of two filamentary appendage (Fa); J, penis. Scale bars in μm, except E in mm." httpUri="https://zenodo.org/record/281663/files/figure.png" pageId="12" pageNumber="33">Fig. 11</figureCitation>
C, D), length of distal arm 7.5 times length of basal arm (
<figureCitation id="136E2A47FFD38A16C6AA24D85543FB31" box="[490,561,1224,1248]" captionStart="FIGURE 2" captionStartId="2.[151,244,1869,1890]" captionTargetBox="[153,1433,917,1847]" captionTargetId="figure@2.[153,1433,917,1848]" captionTargetPageId="2" captionText="FIGURE 2. Kumejima epibiotic barnacles. A, Calantica arcuata n. sp. (Left, holotype, RUMF-ZC- 1554, middle, paratype, RUMF- ZC- 1555, note the tip of carina was broken, right, paratype, NMNS- 6690 - 001); B, Euscalpellum c. f. squamoum Hiro, 1937 (left, RUMF-ZC- 1556, right ASIZCR- 000225); C, Oxynaspis celata Darwin, 1852 (RUMF-ZC- 1557); D, Oxynaspis ryukyuensis n. sp. (holotype, RUMF-ZC- 1558); E, Poecilasma obliqua (Hoek, 1907) (RUMF-ZC- 1560); F, Platylepas hexastylos (Fabricius, 1798) (RUMF-ZC- 1561). A-E, lateral view for stalked barnacles; F, anterior view for Platylepas hexastylos. Scale bars in mm to indicate the size of specimens." httpUri="https://zenodo.org/record/281654/files/figure.png" pageId="12" pageNumber="33">Figs 2</figureCitation>
D, 9A). Tergum triangular, narrow, apex acute, umbo apical, occludent margin shortest, umbo apical (
<figureCitation id="136E2A47FFD38A16C6DA24FB5696FAD2" box="[410,484,1259,1283]" captionStart="FIGURE 2" captionStartId="2.[151,244,1869,1890]" captionTargetBox="[153,1433,917,1847]" captionTargetId="figure@2.[153,1433,917,1848]" captionTargetPageId="2" captionText="FIGURE 2. Kumejima epibiotic barnacles. A, Calantica arcuata n. sp. (Left, holotype, RUMF-ZC- 1554, middle, paratype, RUMF- ZC- 1555, note the tip of carina was broken, right, paratype, NMNS- 6690 - 001); B, Euscalpellum c. f. squamoum Hiro, 1937 (left, RUMF-ZC- 1556, right ASIZCR- 000225); C, Oxynaspis celata Darwin, 1852 (RUMF-ZC- 1557); D, Oxynaspis ryukyuensis n. sp. (holotype, RUMF-ZC- 1558); E, Poecilasma obliqua (Hoek, 1907) (RUMF-ZC- 1560); F, Platylepas hexastylos (Fabricius, 1798) (RUMF-ZC- 1561). A-E, lateral view for stalked barnacles; F, anterior view for Platylepas hexastylos. Scale bars in mm to indicate the size of specimens." httpUri="https://zenodo.org/record/281654/files/figure.png" pageId="12" pageNumber="33">Figs 2</figureCitation>
D, 9A, 11B). Scutum trapezoid, umbo at middle portion of occludent margin, basal margin straight, perpendicular to occludent margin, carinal margin shortest (
<figureCitation id="136E2A47FFD38A16C4AC251F5346FAF6" box="[1004,1076,1295,1319]" captionStart="FIGURE 2" captionStartId="2.[151,244,1869,1890]" captionTargetBox="[153,1433,917,1847]" captionTargetId="figure@2.[153,1433,917,1848]" captionTargetPageId="2" captionText="FIGURE 2. Kumejima epibiotic barnacles. A, Calantica arcuata n. sp. (Left, holotype, RUMF-ZC- 1554, middle, paratype, RUMF- ZC- 1555, note the tip of carina was broken, right, paratype, NMNS- 6690 - 001); B, Euscalpellum c. f. squamoum Hiro, 1937 (left, RUMF-ZC- 1556, right ASIZCR- 000225); C, Oxynaspis celata Darwin, 1852 (RUMF-ZC- 1557); D, Oxynaspis ryukyuensis n. sp. (holotype, RUMF-ZC- 1558); E, Poecilasma obliqua (Hoek, 1907) (RUMF-ZC- 1560); F, Platylepas hexastylos (Fabricius, 1798) (RUMF-ZC- 1561). A-E, lateral view for stalked barnacles; F, anterior view for Platylepas hexastylos. Scale bars in mm to indicate the size of specimens." httpUri="https://zenodo.org/record/281654/files/figure.png" pageId="12" pageNumber="33">Figs 2</figureCitation>
D, 9A, 11A).
</paragraph>
<paragraph id="8BEA36C2FFD38A16C787252355A0FA0B" blockId="12.[151,1436,866,2034]" pageId="12" pageNumber="33">
Maxilla rounded, simple setae on all margin (
<figureCitation id="136E2A47FFD38A16C59B2523545DFA9A" box="[731,815,1331,1355]" captionStart="FIGURE 10" captionStartId="15.[151,244,1943,1964]" captionTargetBox="[168,1406,162,1928]" captionTargetId="figure@15.[135,1451,127,1937]" captionTargetPageId="15" captionText="FIGURE 10. Oxynaspis ryukyuensis n. sp. (holotype, RUMF-ZC- 1558). A, maxilla; B, magnified view of maxilla, showing setae; C, maxillule; D, cutting margin of maxillule; E, mandible; F, lower margin and inferior angle of mandible; G, mandibular palp; H, setae at tip of mandibular palp. Scale bars in μm." httpUri="https://zenodo.org/record/281662/files/figure.png" pageId="12" pageNumber="33">Fig. 10</figureCitation>
A, B). Maxillule strongly notched (
<figureCitation id="136E2A47FFD38A16C38B2523526DFA9A" box="[1227,1311,1331,1355]" captionStart="FIGURE 10" captionStartId="15.[151,244,1943,1964]" captionTargetBox="[168,1406,162,1928]" captionTargetId="figure@15.[135,1451,127,1937]" captionTargetPageId="15" captionText="FIGURE 10. Oxynaspis ryukyuensis n. sp. (holotype, RUMF-ZC- 1558). A, maxilla; B, magnified view of maxilla, showing setae; C, maxillule; D, cutting margin of maxillule; E, mandible; F, lower margin and inferior angle of mandible; G, mandibular palp; H, setae at tip of mandibular palp. Scale bars in μm." httpUri="https://zenodo.org/record/281662/files/figure.png" pageId="12" pageNumber="33">Fig. 10</figureCitation>
C), 2 large spines above notch, region below notch protuberant, several short, simple setae in notch, approximately 8 setae below notch (
<figureCitation id="136E2A47FFD38A16C670256A56F3FA43" box="[304,385,1402,1426]" captionStart="FIGURE 10" captionStartId="15.[151,244,1943,1964]" captionTargetBox="[168,1406,162,1928]" captionTargetId="figure@15.[135,1451,127,1937]" captionTargetPageId="15" captionText="FIGURE 10. Oxynaspis ryukyuensis n. sp. (holotype, RUMF-ZC- 1558). A, maxilla; B, magnified view of maxilla, showing setae; C, maxillule; D, cutting margin of maxillule; E, mandible; F, lower margin and inferior angle of mandible; G, mandibular palp; H, setae at tip of mandibular palp. Scale bars in μm." httpUri="https://zenodo.org/record/281662/files/figure.png" pageId="12" pageNumber="33">Fig. 10</figureCitation>
D). Mandible with4 teeth, first separated from remainder, fourth smallest, lower margin short, with 3 short pectinations, inferior angle with 1 sharp pectination (
<figureCitation id="136E2A47FFD38A16C434258E54B7FA67" box="[884,965,1438,1462]" captionStart="FIGURE 10" captionStartId="15.[151,244,1943,1964]" captionTargetBox="[168,1406,162,1928]" captionTargetId="figure@15.[135,1451,127,1937]" captionTargetPageId="15" captionText="FIGURE 10. Oxynaspis ryukyuensis n. sp. (holotype, RUMF-ZC- 1558). A, maxilla; B, magnified view of maxilla, showing setae; C, maxillule; D, cutting margin of maxillule; E, mandible; F, lower margin and inferior angle of mandible; G, mandibular palp; H, setae at tip of mandibular palp. Scale bars in μm." httpUri="https://zenodo.org/record/281662/files/figure.png" pageId="12" pageNumber="33">Fig. 10</figureCitation>
E, F). Mandibular palps elongated, simple setae distally and on exterior margin (
<figureCitation id="136E2A47FFD38A16C57E25D255FFFA0B" box="[574,653,1474,1498]" captionStart="FIGURE 10" captionStartId="15.[151,244,1943,1964]" captionTargetBox="[168,1406,162,1928]" captionTargetId="figure@15.[135,1451,127,1937]" captionTargetPageId="15" captionText="FIGURE 10. Oxynaspis ryukyuensis n. sp. (holotype, RUMF-ZC- 1558). A, maxilla; B, magnified view of maxilla, showing setae; C, maxillule; D, cutting margin of maxillule; E, mandible; F, lower margin and inferior angle of mandible; G, mandibular palp; H, setae at tip of mandibular palp. Scale bars in μm." httpUri="https://zenodo.org/record/281662/files/figure.png" pageId="12" pageNumber="33">Fig. 10</figureCitation>
G, H).
</paragraph>
<paragraph id="8BEA36C2FFD38A16C78725F65210F905" blockId="12.[151,1436,866,2034]" pageId="12" pageNumber="33">
Cirrus I with anterior and posterior rami 6-segmented, anterior ramus slightly longer (
<figureCitation id="136E2A47FFD38A16C3D625F653A9FA2F" box="[1174,1243,1510,1534]" captionStart="FIGURE 9" captionStartId="14.[151,244,1967,1988]" captionTargetBox="[172,1383,203,1948]" captionTargetId="figure@14.[135,1451,163,1974]" captionTargetPageId="14" captionText="FIGURE 9. Oxynaspis ryukyuensis n. sp. (holotype, RUMF-ZC- 1558). A, right view of capitulum; B, cirrus I; C, cirrus II; D, cirrus III; E, cirrus IV; F, cirrus V; G, cirrus VI. Scale bars in μm, except A in mm." httpUri="https://zenodo.org/record/281661/files/figure.png" pageId="12" pageNumber="33">Fig. 9</figureCitation>
B); setae on both rami simple (
<figureCitation id="136E2A47FFD38A16C670261956F3F9F0" box="[304,385,1545,1569]" captionStart="FIGURE 11" captionStartId="16.[151,244,1934,1955]" captionTargetBox="[156,1432,159,1933]" captionTargetId="figure@16.[135,1451,137,1948]" captionTargetPageId="16" captionText="FIGURE 11. Oxynaspis ryukyuensis n. sp. (holotype, RUMF-ZC- 1558). A, scutum, dorsal view, B, tergum, dorsal view, C, carina, lateral view, D, carina, basal view showing non-forked base, u—umbo, E, simple type setae on cirrus I; F, distal region of cirrus II; G, distal region of cirrus III; H. intermediate segment of posterior ramus of cirrus VI; I, somatic body, showing location of two filamentary appendage (Fa); J, penis. Scale bars in μm, except E in mm." httpUri="https://zenodo.org/record/281663/files/figure.png" pageId="12" pageNumber="33">Fig. 11</figureCitation>
E). Cirrus II with anterior and posterior rami 13- and 12-segmented, respectively (
<figureCitation id="136E2A47FFD38A16C2742619520BF9F0" box="[1332,1401,1545,1569]" captionStart="FIGURE 9" captionStartId="14.[151,244,1967,1988]" captionTargetBox="[172,1383,203,1948]" captionTargetId="figure@14.[135,1451,163,1974]" captionTargetPageId="14" captionText="FIGURE 9. Oxynaspis ryukyuensis n. sp. (holotype, RUMF-ZC- 1558). A, right view of capitulum; B, cirrus I; C, cirrus II; D, cirrus III; E, cirrus IV; F, cirrus V; G, cirrus VI. Scale bars in μm, except A in mm." httpUri="https://zenodo.org/record/281661/files/figure.png" pageId="12" pageNumber="33">Fig. 9</figureCitation>
C); setae on both rami simple (
<figureCitation id="136E2A47FFD38A16C68C263D5569F994" box="[460,539,1581,1605]" captionStart="FIGURE 11" captionStartId="16.[151,244,1934,1955]" captionTargetBox="[156,1432,159,1933]" captionTargetId="figure@16.[135,1451,137,1948]" captionTargetPageId="16" captionText="FIGURE 11. Oxynaspis ryukyuensis n. sp. (holotype, RUMF-ZC- 1558). A, scutum, dorsal view, B, tergum, dorsal view, C, carina, lateral view, D, carina, basal view showing non-forked base, u—umbo, E, simple type setae on cirrus I; F, distal region of cirrus II; G, distal region of cirrus III; H. intermediate segment of posterior ramus of cirrus VI; I, somatic body, showing location of two filamentary appendage (Fa); J, penis. Scale bars in μm, except E in mm." httpUri="https://zenodo.org/record/281663/files/figure.png" pageId="12" pageNumber="33">Fig. 11</figureCitation>
F, G). Cirri III and IV with anterior and posterior rami 11-segmented (
<figureCitation id="136E2A47FFD38A16C27D263D52F3F994" box="[1341,1409,1581,1605]" captionStart="FIGURE 9" captionStartId="14.[151,244,1967,1988]" captionTargetBox="[172,1383,203,1948]" captionTargetId="figure@14.[135,1451,163,1974]" captionTargetPageId="14" captionText="FIGURE 9. Oxynaspis ryukyuensis n. sp. (holotype, RUMF-ZC- 1558). A, right view of capitulum; B, cirrus I; C, cirrus II; D, cirrus III; E, cirrus IV; F, cirrus V; G, cirrus VI. Scale bars in μm, except A in mm." httpUri="https://zenodo.org/record/281661/files/figure.png" pageId="12" pageNumber="33">Fig. 9</figureCitation>
D, E). Cirrus V with anterior and posterior rami 12- and 11-segmented (
<figureCitation id="136E2A47FFD38A16C4D8264154A9F9B8" box="[920,987,1617,1641]" captionStart="FIGURE 9" captionStartId="14.[151,244,1967,1988]" captionTargetBox="[172,1383,203,1948]" captionTargetId="figure@14.[135,1451,163,1974]" captionTargetPageId="14" captionText="FIGURE 9. Oxynaspis ryukyuensis n. sp. (holotype, RUMF-ZC- 1558). A, right view of capitulum; B, cirrus I; C, cirrus II; D, cirrus III; E, cirrus IV; F, cirrus V; G, cirrus VI. Scale bars in μm, except A in mm." httpUri="https://zenodo.org/record/281661/files/figure.png" pageId="12" pageNumber="33">Fig. 9</figureCitation>
F). Cirrus VI with anterior and posterior rami 15- and 14-segmented, respectively (
<figureCitation id="136E2A47FFD38A16C52D266455DDF95D" box="[621,687,1652,1676]" captionStart="FIGURE 9" captionStartId="14.[151,244,1967,1988]" captionTargetBox="[172,1383,203,1948]" captionTargetId="figure@14.[135,1451,163,1974]" captionTargetPageId="14" captionText="FIGURE 9. Oxynaspis ryukyuensis n. sp. (holotype, RUMF-ZC- 1558). A, right view of capitulum; B, cirrus I; C, cirrus II; D, cirrus III; E, cirrus IV; F, cirrus V; G, cirrus VI. Scale bars in μm, except A in mm." httpUri="https://zenodo.org/record/281661/files/figure.png" pageId="12" pageNumber="33">Fig. 9</figureCitation>
G). Intermediate segments of posterior rami of cirri III to VI with 3 pairs of long simple setae and 1 pair of short simple setae (
<figureCitation id="136E2A47FFD38A16C46D2688540FF961" box="[813,893,1688,1712]" captionStart="FIGURE 11" captionStartId="16.[151,244,1934,1955]" captionTargetBox="[156,1432,159,1933]" captionTargetId="figure@16.[135,1451,137,1948]" captionTargetPageId="16" captionText="FIGURE 11. Oxynaspis ryukyuensis n. sp. (holotype, RUMF-ZC- 1558). A, scutum, dorsal view, B, tergum, dorsal view, C, carina, lateral view, D, carina, basal view showing non-forked base, u—umbo, E, simple type setae on cirrus I; F, distal region of cirrus II; G, distal region of cirrus III; H. intermediate segment of posterior ramus of cirrus VI; I, somatic body, showing location of two filamentary appendage (Fa); J, penis. Scale bars in μm, except E in mm." httpUri="https://zenodo.org/record/281663/files/figure.png" pageId="12" pageNumber="33">Fig. 11</figureCitation>
H). Caudal appendages absent. Penis half length of cirrus VI (
<figureCitation id="136E2A47FFD38A16C66926AC560BF905" box="[297,377,1724,1748]" captionStart="FIGURE 11" captionStartId="16.[151,244,1934,1955]" captionTargetBox="[156,1432,159,1933]" captionTargetId="figure@16.[135,1451,137,1948]" captionTargetPageId="16" captionText="FIGURE 11. Oxynaspis ryukyuensis n. sp. (holotype, RUMF-ZC- 1558). A, scutum, dorsal view, B, tergum, dorsal view, C, carina, lateral view, D, carina, basal view showing non-forked base, u—umbo, E, simple type setae on cirrus I; F, distal region of cirrus II; G, distal region of cirrus III; H. intermediate segment of posterior ramus of cirrus VI; I, somatic body, showing location of two filamentary appendage (Fa); J, penis. Scale bars in μm, except E in mm." httpUri="https://zenodo.org/record/281663/files/figure.png" pageId="12" pageNumber="33">Fig. 11</figureCitation>
J). Dorsal margin of somatic body with 2 narrow, long filamentary appendages (
<figureCitation id="136E2A47FFD38A16C3B926AC523BF905" box="[1273,1353,1724,1748]" captionStart="FIGURE 11" captionStartId="16.[151,244,1934,1955]" captionTargetBox="[156,1432,159,1933]" captionTargetId="figure@16.[135,1451,137,1948]" captionTargetPageId="16" captionText="FIGURE 11. Oxynaspis ryukyuensis n. sp. (holotype, RUMF-ZC- 1558). A, scutum, dorsal view, B, tergum, dorsal view, C, carina, lateral view, D, carina, basal view showing non-forked base, u—umbo, E, simple type setae on cirrus I; F, distal region of cirrus II; G, distal region of cirrus III; H. intermediate segment of posterior ramus of cirrus VI; I, somatic body, showing location of two filamentary appendage (Fa); J, penis. Scale bars in μm, except E in mm." httpUri="https://zenodo.org/record/281663/files/figure.png" pageId="12" pageNumber="33">
Fig.
<date id="FFEB1002FFD38A16C26D26AC523BF905" box="[1325,1353,1724,1748]" pageId="12" pageNumber="33" value="2011-01">11</date>
</figureCitation>
I).
</paragraph>
</subSubSection>
<subSubSection id="C34F6549FFD38A16C78726CF535AF929" box="[199,1064,1759,1784]" pageId="12" pageNumber="33" type="distribution">
<paragraph id="8BEA36C2FFD38A16C78726CF535AF929" blockId="12.[151,1436,866,2034]" box="[199,1064,1759,1784]" pageId="12" pageNumber="33">
<emphasis id="B921EAD0FFD38A16C78726CF5612F929" bold="true" box="[199,352,1759,1784]" pageId="12" pageNumber="33">Distribution.</emphasis>
Currently only known from Kumejima Island, Ryukyu Islands.
</paragraph>
</subSubSection>
<subSubSection id="C34F6549FFD38A16C78727145320F8CA" box="[199,1106,1795,1820]" pageId="12" pageNumber="33" type="etymology">
<paragraph id="8BEA36C2FFD38A16C78727145320F8CA" blockId="12.[151,1436,866,2034]" box="[199,1106,1795,1820]" pageId="12" pageNumber="33">
<emphasis id="B921EAD0FFD38A16C7872714563CF8CD" bold="true" box="[199,334,1796,1820]" pageId="12" pageNumber="33">Etymology.</emphasis>
The name
<taxonomicName id="4C554D41FFD38A16C68B2714553DF8CD" box="[459,591,1796,1820]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="12" pageNumber="33" phylum="Arthropoda" rank="species" species="ryukyuensis">
<emphasis id="B921EAD0FFD38A16C68B2714553DF8CD" box="[459,591,1796,1820]" italics="true" pageId="12" pageNumber="33">ryukyuensis</emphasis>
</taxonomicName>
denotes the
<typeStatus id="54EE8860FFD38A16C59C2714547EF8CD" box="[732,780,1796,1820]" pageId="12" pageNumber="33">type</typeStatus>
locality, the Ryukyu Islands.
</paragraph>
</subSubSection>
<subSubSection id="C34F6549FFD38A17C7872738536FF904" lastPageId="13" lastPageNumber="34" pageId="12" pageNumber="33" type="discussion">
<paragraph id="8BEA36C2FFD38A17C7872738557CFE1F" blockId="12.[151,1436,866,2034]" lastBlockId="13.[151,1437,151,1999]" lastPageId="13" lastPageNumber="34" pageId="12" pageNumber="33">
<emphasis id="B921EAD0FFD38A16C78727385649F891" bold="true" box="[199,315,1832,1856]" pageId="12" pageNumber="33">Remarks.</emphasis>
Previous studies on the diversity of
<taxonomicName id="4C554D41FFD38A16C5942738543BF8EE" box="[724,841,1832,1855]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="12" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="B921EAD0FFD38A16C5942738543BF8EE" box="[724,841,1832,1855]" italics="true" pageId="12" pageNumber="33">Oxynaspis</emphasis>
</taxonomicName>
have received very scant attention. A recent revision of the
<taxonomicName id="4C554D41FFD38A16C79E275B559CF8B2" authority="Van Syoc &amp; Dekelboum 2011" authorityName="Van Syoc &amp; Dekelboum" authorityYear="2011" box="[222,750,1867,1891]" class="Maxillopoda" family="Oxynaspididae" kingdom="Animalia" order="Pedunculata" pageId="12" pageNumber="33" phylum="Arthropoda" rank="family">
Oxynaspididae (
<bibRefCitation id="EFC44B33FFD38A16C6D4275B5595F8B2" author="Van" box="[404,743,1867,1891]" pageId="12" pageNumber="33" refString="Van Syoc, R. &amp; Dekelboum, A. (2011) Oxynaspididae (Crustacea, Cirripedia): phylogenetic and evolutionary ecology, with description of three new genera and six new species. Zootaxa, 3103, 1 - 32." type="journal article" year="2011">Van Syoc &amp; Dekelboum 2011</bibRefCitation>
)
</taxonomicName>
erected two new genera,
<taxonomicName id="4C554D41FFD38A16C34C275C57BCF856" authority="Van Syoc &amp; Dekelboum, 2011" authorityName="Van Syoc &amp; Dekelboum" authorityYear="2011" class="Maxillopoda" family="Oxynaspididae" genus="Pycnaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pedunculata" pageId="12" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="B921EAD0FFD38A16C34C275C530CF8B2" box="[1036,1150,1868,1891]" italics="true" pageId="12" pageNumber="33">Pycnaspis</emphasis>
<bibRefCitation id="EFC44B33FFD38A16C3C4275C57BCF856" author="Van" pageId="12" pageNumber="33" refString="Van Syoc, R. &amp; Dekelboum, A. (2011) Oxynaspididae (Crustacea, Cirripedia): phylogenetic and evolutionary ecology, with description of three new genera and six new species. Zootaxa, 3103, 1 - 32." type="journal article" year="2011">Van Syoc &amp; Dekelboum, 2011</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C554D41FFD38A16C643276055A1F856" authority="Van Syoc &amp; Dekelboum, 2011" authorityName="Van Syoc &amp; Dekelboum" authorityYear="2011" box="[259,723,1903,1927]" class="Maxillopoda" family="Oxynaspididae" genus="Minyaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pedunculata" pageId="12" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="B921EAD0FFD38A16C64327605604F856" box="[259,374,1904,1927]" italics="true" pageId="12" pageNumber="33">Minyaspis</emphasis>
<bibRefCitation id="EFC44B33FFD38A16C63C276055A1F856" author="Van" box="[380,723,1903,1927]" pageId="12" pageNumber="33" refString="Van Syoc, R. &amp; Dekelboum, A. (2011) Oxynaspididae (Crustacea, Cirripedia): phylogenetic and evolutionary ecology, with description of three new genera and six new species. Zootaxa, 3103, 1 - 32." type="journal article" year="2011">Van Syoc &amp; Dekelboum, 2011</bibRefCitation>
</taxonomicName>
, in addition to
<taxonomicName id="4C554D41FFD38A16C4372760549EF856" box="[887,1004,1904,1927]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="12" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="B921EAD0FFD38A16C4372760549EF856" box="[887,1004,1904,1927]" italics="true" pageId="12" pageNumber="33">Oxynaspis</emphasis>
</taxonomicName>
.
<taxonomicName id="4C554D41FFD38A16C4BA2760531EF856" box="[1018,1132,1904,1927]" class="Maxillopoda" family="Oxynaspididae" genus="Pycnaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pedunculata" pageId="12" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="B921EAD0FFD38A16C4BA2760531EF856" box="[1018,1132,1904,1927]" italics="true" pageId="12" pageNumber="33">Pycnaspis</emphasis>
</taxonomicName>
is a monotypic genus, containing the thick and strong plated
<taxonomicName id="4C554D41FFD38A16C560278455CBF87A" box="[544,697,1940,1963]" class="Maxillopoda" family="Oxynaspididae" genus="Pycnaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pedunculata" pageId="12" pageNumber="33" phylum="Arthropoda" rank="species" species="connectens">
<emphasis id="B921EAD0FFD38A16C560278455CBF87A" box="[544,697,1940,1963]" italics="true" pageId="12" pageNumber="33">P. connectens</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFC44B33FFD38A16C588278253E4F87A" author="Broch" box="[712,1174,1938,1963]" pageId="12" pageNumber="33" refString="Broch, H. (1931) Papers from Dr. Th. Mortensen's Pacific Expedition 1914 ~ 1916, LVI. Indomalayan Cirripedia. Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening i Kobenhavn, 91, 1 - 146." type="journal article" year="1931">
Broch, 1931, = ex
<taxonomicName id="4C554D41FFD38A16C4DD278353E4F87A" box="[925,1174,1939,1963]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="12" pageNumber="33" phylum="Arthropoda" rank="species" species="connectens">
<emphasis id="B921EAD0FFD38A16C4DD278353E4F87A" box="[925,1174,1939,1963]" italics="true" pageId="12" pageNumber="33">Oxynaspis connectens</emphasis>
</taxonomicName>
</bibRefCitation>
).
<emphasis id="B921EAD0FFD38A16C3EE27845261F87A" box="[1198,1299,1940,1963]" italics="true" pageId="12" pageNumber="33">Miyaspis</emphasis>
contains 15 species, all of which have a capitulum partially covered by plates and a basally forked carina (
<bibRefCitation id="EFC44B33FFD38A16C3FE27A65668F823" author="Van" pageId="12" pageNumber="33" refString="Van Syoc, R. &amp; Dekelboum, A. (2011) Oxynaspididae (Crustacea, Cirripedia): phylogenetic and evolutionary ecology, with description of three new genera and six new species. Zootaxa, 3103, 1 - 32." type="journal article" year="2011">Van Syoc &amp; Dekelboum 2011</bibRefCitation>
).
<taxonomicName id="4C554D41FFD38A16C66D27CB56D0F823" box="[301,418,2011,2034]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="12" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="B921EAD0FFD38A16C66D27CB56D0F823" box="[301,418,2011,2034]" italics="true" pageId="12" pageNumber="33">Oxynaspis</emphasis>
</taxonomicName>
presently contains nine species (
<taxonomicName id="4C554D41FFD38A16C45527CB530BF823" authority="Darwin, 1852" authorityName="Darwin" authorityYear="1852" box="[789,1145,2010,2034]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="12" pageNumber="33" phylum="Arthropoda" rank="species" species="celata">
<emphasis id="B921EAD0FFD38A16C45527CB54A7F823" box="[789,981,2010,2034]" italics="true" pageId="12" pageNumber="33">Oxynaspis celata</emphasis>
<bibRefCitation id="EFC44B33FFD38A16C49D27CA530BF823" author="Darwin" box="[989,1145,2010,2034]" pageId="12" pageNumber="33" refString="Darwin, C. (1852) A monograph on the sub-class Cirripedia with figures of all species. The Lepadidae: or pedunculated barnacles. Ray Society, London. 400 pp. (1851, published 1852)" type="book" year="1852">Darwin, 1852</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C554D41FFD38A16C3C627CB52EAF823" authority="Totton, 1940" authorityName="Totton" authorityYear="1940" box="[1158,1432,2010,2034]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="12" pageNumber="33" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="B921EAD0FFD38A16C3C627CB538CF823" box="[1158,1278,2010,2034]" italics="true" pageId="12" pageNumber="33">O. gracilis</emphasis>
<bibRefCitation id="EFC44B33FFD38A16C24627CB52EAF823" author="Totton" box="[1286,1432,2010,2034]" pageId="12" pageNumber="33" refString="Totton, A. K. (1940) New species of the cirripede genus Oxynaspis commensal with Antipatharia. The Annals and Magazine of Natural History, Series, 11, 6, 465 - 487." type="journal article" year="1940">Totton, 1940</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C554D41FFD28A17C7D7208856E5FF7E" authority="Totton, 1940" authorityName="Totton" authorityYear="1940" box="[151,407,151,176]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="alatae">
<emphasis id="B921EAD0FFD28A17C7D72088578DFF61" box="[151,255,152,176]" italics="true" pageId="13" pageNumber="34">O. alatae</emphasis>
<bibRefCitation id="EFC44B33FFD28A17C646208856E5FF7E" author="Totton" box="[262,407,151,175]" pageId="13" pageNumber="34" refString="Totton, A. K. (1940) New species of the cirripede genus Oxynaspis commensal with Antipatharia. The Annals and Magazine of Natural History, Series, 11, 6, 465 - 487." type="journal article" year="1940">Totton, 1940</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C554D41FFD28A17C6E220885445FF7E" authority="Totton, 1940" authorityName="Totton" authorityYear="1940" box="[418,823,151,176]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="cancellatae">
<emphasis id="B921EAD0FFD28A17C6E2208855EDFF61" box="[418,671,152,176]" italics="true" pageId="13" pageNumber="34">Oxynaspis cancellatae</emphasis>
<bibRefCitation id="EFC44B33FFD28A17C5E620885445FF7E" author="Totton" box="[678,823,151,175]" pageId="13" pageNumber="34" refString="Totton, A. K. (1940) New species of the cirripede genus Oxynaspis commensal with Antipatharia. The Annals and Magazine of Natural History, Series, 11, 6, 465 - 487." type="journal article" year="1940">Totton, 1940</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C554D41FFD28A17C402208853E8FF7E" authority="Hiro, 1931" authorityName="Hiro" authorityYear="1931" box="[834,1178,151,176]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="pacifica">
<emphasis id="B921EAD0FFD28A17C4022088536AFF61" box="[834,1048,152,176]" italics="true" pageId="13" pageNumber="34">Oxynaspis pacifica</emphasis>
<bibRefCitation id="EFC44B33FFD28A17C35F208753E8FF7E" author="Hiro" box="[1055,1178,151,175]" pageId="13" pageNumber="34" refString="Hiro, F. (1931) Notes on some new Cirripedia from Japan. Memoirs of the College of Science, Koyoto Imperial University, Series B., 7 (3), 143 - 158." type="journal article" year="1931">Hiro, 1931</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C554D41FFD28A17C3E620885663FF02" authority="Newman, 1972" authorityName="Newman" authorityYear="1972" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="rossi">
<emphasis id="B921EAD0FFD28A17C3E620885225FF61" box="[1190,1367,152,176]" italics="true" pageId="13" pageNumber="34">Oxynaspis rossi</emphasis>
<bibRefCitation id="EFC44B33FFD28A17C21E20885663FF02" author="Newman" pageId="13" pageNumber="34" refString="Newman, W. A. (1972) An Oxynaspid (Cirripedia, Thoracica) from the Eastern Pacific. Crustaceana, 23, 202 - 208" type="journal article" year="1972">Newman, 1972</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C554D41FFD28A17C65C20AC5431FF02" authority="Van Syoc &amp; Dekelboum, 2011" authorityName="Van Syoc &amp; Dekelboum" authorityYear="2011" box="[284,835,187,212]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="perekrestenkoi">
<emphasis id="B921EAD0FFD28A17C65C20AC5690FF05" box="[284,482,188,212]" italics="true" pageId="13" pageNumber="34">O. perekrestenkoi</emphasis>
<bibRefCitation id="EFC44B33FFD28A17C6A920AC5431FF02" author="Van" box="[489,835,187,211]" pageId="13" pageNumber="34" refString="Van Syoc, R. &amp; Dekelboum, A. (2011) Oxynaspididae (Crustacea, Cirripedia): phylogenetic and evolutionary ecology, with description of three new genera and six new species. Zootaxa, 3103, 1 - 32." type="journal article" year="2011">Van Syoc &amp; Dekelboum, 2011</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C554D41FFD28A17C40F20AC521EFF02" authority="Van Syoc &amp; Dekelboum, 2011" authorityName="Van Syoc &amp; Dekelboum" authorityYear="2011" box="[847,1388,187,212]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="joankovennae">
<emphasis id="B921EAD0FFD28A17C40F20AC537EFF05" box="[847,1036,188,212]" italics="true" pageId="13" pageNumber="34">O. joankovennae</emphasis>
<bibRefCitation id="EFC44B33FFD28A17C35420AC521EFF02" author="Van" box="[1044,1388,187,211]" pageId="13" pageNumber="34" refString="Van Syoc, R. &amp; Dekelboum, A. (2011) Oxynaspididae (Crustacea, Cirripedia): phylogenetic and evolutionary ecology, with description of three new genera and six new species. Zootaxa, 3103, 1 - 32." type="journal article" year="2011">Van Syoc &amp; Dekelboum, 2011</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C554D41FFD28A17C7D720F055D1FF26" authority="Van Syoc &amp; Dekelboum, 2011" authorityName="Van Syoc &amp; Dekelboum" authorityYear="2011" box="[151,675,223,247]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="joandianeae">
<emphasis id="B921EAD0FFD28A17C7D720F05631FF26" box="[151,323,223,247]" italics="true" pageId="13" pageNumber="34">O. joandianeae</emphasis>
<bibRefCitation id="EFC44B33FFD28A17C60B20F055D1FF26" author="Van" box="[331,675,223,247]" pageId="13" pageNumber="34" refString="Van Syoc, R. &amp; Dekelboum, A. (2011) Oxynaspididae (Crustacea, Cirripedia): phylogenetic and evolutionary ecology, with description of three new genera and six new species. Zootaxa, 3103, 1 - 32." type="journal article" year="2011">Van Syoc &amp; Dekelboum, 2011</bibRefCitation>
</taxonomicName>
) and four sub-species of
<taxonomicName id="4C554D41FFD28A17C4FD20F05351FF26" box="[957,1059,223,247]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="celata">
<emphasis id="B921EAD0FFD28A17C4FD20F05351FF26" box="[957,1059,223,247]" italics="true" pageId="13" pageNumber="34">O. celata</emphasis>
</taxonomicName>
(see remarks on
<taxonomicName id="4C554D41FFD28A17C3A420F05238FF26" box="[1252,1354,223,247]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="celata">
<emphasis id="B921EAD0FFD28A17C3A420F05238FF26" box="[1252,1354,223,247]" italics="true" pageId="13" pageNumber="34">O. celata</emphasis>
</taxonomicName>
above;
<bibRefCitation id="EFC44B33FFD28A17C7D72114569FFECA" author="Van" box="[151,493,259,283]" pageId="13" pageNumber="34" refString="Van Syoc, R. &amp; Dekelboum, A. (2011) Oxynaspididae (Crustacea, Cirripedia): phylogenetic and evolutionary ecology, with description of three new genera and six new species. Zootaxa, 3103, 1 - 32." type="journal article" year="2011">Van Syoc &amp; Dekelboum 2011</bibRefCitation>
). In Zevinas (1982) revision of the poecilasmatid barnacles,
<taxonomicName id="4C554D41FFD28A17C3DF21145655FEEE" authority="Totton, 1940" authorityName="Totton" authorityYear="1940" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="cancellatae">
<emphasis id="B921EAD0FFD28A17C3DF211452EEFECA" box="[1183,1436,259,283]" italics="true" pageId="13" pageNumber="34">Oxynaspis cancellatae</emphasis>
<bibRefCitation id="EFC44B33FFD28A17C7D721385655FEEE" author="Totton" box="[151,295,295,319]" pageId="13" pageNumber="34" refString="Totton, A. K. (1940) New species of the cirripede genus Oxynaspis commensal with Antipatharia. The Annals and Magazine of Natural History, Series, 11, 6, 465 - 487." type="journal article" year="1940">Totton, 1940</bibRefCitation>
</taxonomicName>
was considered a junior synonym to
<taxonomicName id="4C554D41FFD28A17C5952138537EFEEE" authority="Broch, 1931" authorityName="Broch" authorityYear="1931" box="[725,1036,295,319]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="connectens">
<emphasis id="B921EAD0FFD28A17C59521385404FEEE" box="[725,886,296,319]" italics="true" pageId="13" pageNumber="34">O. connectens</emphasis>
<bibRefCitation id="EFC44B33FFD28A17C43E2137537EFEEE" author="Broch" box="[894,1036,295,319]" pageId="13" pageNumber="34" refString="Broch, H. (1931) Papers from Dr. Th. Mortensen's Pacific Expedition 1914 ~ 1916, LVI. Indomalayan Cirripedia. Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening i Kobenhavn, 91, 1 - 146." type="journal article" year="1931">Broch, 1931</bibRefCitation>
</taxonomicName>
, while
<bibRefCitation id="EFC44B33FFD28A17C32121385268FEEE" author="Newman" box="[1121,1306,295,319]" pageId="13" pageNumber="34" refString="Newman, W. A. (1972) An Oxynaspid (Cirripedia, Thoracica) from the Eastern Pacific. Crustaceana, 23, 202 - 208" type="journal article" year="1972">Newman (1972)</bibRefCitation>
considered them two separate species. Even
<bibRefCitation id="EFC44B33FFD28A17C54E215B5406FEB3" box="[526,884,330,354]" pageId="13" pageNumber="34" refString="Van Syoc, R. &amp; Dekelboum, A. (2011) Oxynaspididae (Crustacea, Cirripedia): phylogenetic and evolutionary ecology, with description of three new genera and six new species. Zootaxa, 3103, 1 - 32." type="journal article">Van Syoc &amp; Dekelboum (2011)</bibRefCitation>
erected a new genus
<taxonomicName id="4C554D41FFD28A17C329215C53A9FEB2" box="[1129,1243,332,355]" class="Maxillopoda" family="Oxynaspididae" genus="Pycnaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pedunculata" pageId="13" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="B921EAD0FFD28A17C329215C53A9FEB2" box="[1129,1243,332,355]" italics="true" pageId="13" pageNumber="34">Pycnaspis</emphasis>
</taxonomicName>
to accommodate
<taxonomicName id="4C554D41FFD28A17C7D7217F5646FE57" box="[151,308,367,390]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="connectens">
<emphasis id="B921EAD0FFD28A17C7D7217F5646FE57" box="[151,308,367,390]" italics="true" pageId="13" pageNumber="34">O. connectens</emphasis>
</taxonomicName>
, due to its distinctly strong and thick capitular plates. In the present study, we adopt the classification of
<bibRefCitation id="EFC44B33FFD28A17C7F621835552FE7B" box="[182,544,402,426]" pageId="13" pageNumber="34" refString="Van Syoc, R. &amp; Dekelboum, A. (2011) Oxynaspididae (Crustacea, Cirripedia): phylogenetic and evolutionary ecology, with description of three new genera and six new species. Zootaxa, 3103, 1 - 32." type="journal article">Van Syoc &amp; Dekelboum (2011)</bibRefCitation>
. No other
<taxonomicName id="4C554D41FFD28A17C5D82183547FFE7B" box="[664,781,403,426]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="13" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="B921EAD0FFD28A17C5D82183547FFE7B" box="[664,781,403,426]" italics="true" pageId="13" pageNumber="34">Oxynaspis</emphasis>
</taxonomicName>
species are reported to be junior synonyms of
<taxonomicName id="4C554D41FFD28A17C263218352E9FE7B" box="[1315,1435,402,426]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="B921EAD0FFD28A17C263218352E9FE7B" box="[1315,1435,402,426]" italics="true" pageId="13" pageNumber="34">O. gracilis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C554D41FFD28A17C78621A7565CFE1F" box="[198,302,438,462]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="alatae">
<emphasis id="B921EAD0FFD28A17C78621A7565CFE1F" box="[198,302,438,462]" italics="true" pageId="13" pageNumber="34">O. alatae</emphasis>
</taxonomicName>
(see
<bibRefCitation id="EFC44B33FFD28A17C62A21A65573FE1F" author="Zevina" box="[362,513,438,462]" pageId="13" pageNumber="34" refString="Zevina, G. B. (1982) Barnacles of the suborder Lepadomorpha (Cirripedia, Thoracica) of the world oceans. II. Opredeliteli Faune SSSR, 133, 1 - 221. (in Russian)." type="journal article" year="1982">Zevina, 1982</bibRefCitation>
).
</paragraph>
<paragraph id="8BEA36C2FFD28A17C78721CA5686FDE8" blockId="13.[151,1437,151,1999]" pageId="13" pageNumber="34">
<taxonomicName id="4C554D41FFD28A17C78721CA56E1FE20" box="[199,403,473,497]" class="Maxillopoda" family="Oxynaspididae" genus="Oxnaspis" kingdom="Animalia" order="Pedunculata" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="pacifica">
<emphasis id="B921EAD0FFD28A17C78721CA56E1FE20" box="[199,403,473,497]" italics="true" pageId="13" pageNumber="34">Oxnaspis pacifica</emphasis>
</taxonomicName>
,
<taxonomicName id="4C554D41FFD28A17C6E421CA568CFE20" box="[420,510,474,497]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="rossi">
<emphasis id="B921EAD0FFD28A17C6E421CA568CFE20" box="[420,510,474,497]" italics="true" pageId="13" pageNumber="34">O. rossi</emphasis>
</taxonomicName>
,
<taxonomicName id="4C554D41FFD28A17C55021CA55AAFE20" box="[528,728,473,497]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="perekrestenkoi">
<emphasis id="B921EAD0FFD28A17C55021CA55AAFE20" box="[528,728,473,497]" italics="true" pageId="13" pageNumber="34">O. perekrestenkoi</emphasis>
</taxonomicName>
,
<taxonomicName id="4C554D41FFD28A17C5A921CA54DBFE20" box="[745,937,473,497]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="joankovennae">
<emphasis id="B921EAD0FFD28A17C5A921CA54DBFE20" box="[745,937,473,497]" italics="true" pageId="13" pageNumber="34">O. joankovennae</emphasis>
</taxonomicName>
and
<taxonomicName id="4C554D41FFD28A17C4A621CA53E7FE20" box="[998,1173,473,497]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="joandianeae">
<emphasis id="B921EAD0FFD28A17C4A621CA53E7FE20" box="[998,1173,473,497]" italics="true" pageId="13" pageNumber="34">O. joandianeae</emphasis>
</taxonomicName>
have large uncalcified spaces between the scutum and carina, differing from
<taxonomicName id="4C554D41FFD28A17C44021EE54DAFDC4" box="[768,936,509,533]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="ryukyuensis" status="sp. nov.">
<emphasis id="B921EAD0FFD28A17C44021EE54DAFDC4" box="[768,936,509,533]" italics="true" pageId="13" pageNumber="34">O. ryukyuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21257ABFFD28A17C4F121ED5480FDC4" box="[945,1010,509,533]" pageId="13" pageNumber="34" rank="species">
<emphasis id="B921EAD0FFD28A17C4F121ED5480FDC4" bold="true" box="[945,1010,509,533]" pageId="13" pageNumber="34">sp. n.</emphasis>
</taxonomicNameLabel>
, which has the capitulum completely covered by the capitular plates.
</paragraph>
<paragraph id="8BEA36C2FFD28A17C78722545450FB80" blockId="13.[151,1437,151,1999]" pageId="13" pageNumber="34">
The location of the umbo on an opercular plate is an important taxonomic character in identifying stalked barnacles (see
<bibRefCitation id="EFC44B33FFD28A17C658227856C2FD51" author="Zevina" box="[280,432,616,640]" pageId="13" pageNumber="34" refString="Zevina, G. B. (1982) Barnacles of the suborder Lepadomorpha (Cirripedia, Thoracica) of the world oceans. II. Opredeliteli Faune SSSR, 133, 1 - 221. (in Russian)." type="journal article" year="1982">Zevina, 1982</bibRefCitation>
). In
<taxonomicName id="4C554D41FFD28A17C6A122795524FD51" box="[481,598,617,640]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="13" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="B921EAD0FFD28A17C6A122795524FD51" box="[481,598,617,640]" italics="true" pageId="13" pageNumber="34">Oxynaspis</emphasis>
</taxonomicName>
, the umbo of the scutum is located at the occludent margin; the location of the umbo on the carina, located at the middle or below the middle portion of the carinal margin, is a morphological characteristic used to identify
<taxonomicName id="4C554D41FFD28A17C6B822A0551FFD16" box="[504,621,688,711]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="13" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="B921EAD0FFD28A17C6B822A0551FFD16" box="[504,621,688,711]" italics="true" pageId="13" pageNumber="34">Oxynaspis</emphasis>
</taxonomicName>
species (
<bibRefCitation id="EFC44B33FFD28A17C5A622A054E8FD19" author="Newman" box="[742,922,688,712]" pageId="13" pageNumber="34" refString="Newman, W. A. (1972) An Oxynaspid (Cirripedia, Thoracica) from the Eastern Pacific. Crustaceana, 23, 202 - 208" type="journal article" year="1972">Newman, 1972</bibRefCitation>
;
<bibRefCitation id="EFC44B33FFD28A17C4E922A1527EFD19" author="Van" box="[937,1292,688,712]" pageId="13" pageNumber="34" refString="Van Syoc, R. &amp; Dekelboum, A. (2011) Oxynaspididae (Crustacea, Cirripedia): phylogenetic and evolutionary ecology, with description of three new genera and six new species. Zootaxa, 3103, 1 - 32." type="journal article" year="2011">Van Syoc &amp; Dekelboum 2011</bibRefCitation>
).
<bibRefCitation id="EFC44B33FFD28A17C26322A05793FD3A" author="Annandale" pageId="13" pageNumber="34" refString="Annandale, N. (1909) An account of the Indian Cirripedia Pedunculata Part 1. Family Lepadidae (sensu stricto). Records of the Indian Museum, 2 (2), 61 - 137." type="journal article" year="1909">Annandale (1909)</bibRefCitation>
and
<bibRefCitation id="EFC44B33FFD28A17C65722C456C3FD3A" author="Totton" box="[279,433,723,747]" pageId="13" pageNumber="34" refString="Totton, A. K. (1940) New species of the cirripede genus Oxynaspis commensal with Antipatharia. The Annals and Magazine of Natural History, Series, 11, 6, 465 - 487." type="journal article" year="1940">Totton (1940)</bibRefCitation>
measured the length ratio of the distal arm (carinal length above the umbo) and basal arm (carinal length below the umbo) of the carina of
<taxonomicName id="4C554D41FFD28A17C5FA22E8545DFCDE" box="[698,815,760,783]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="13" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="B921EAD0FFD28A17C5FA22E8545DFCDE" box="[698,815,760,783]" italics="true" pageId="13" pageNumber="34">Oxynaspis</emphasis>
</taxonomicName>
for species comparisons. Comparing the present specimen with
<taxonomicName id="4C554D41FFD28A17C647230C56B4FCE2" box="[263,454,795,819]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="celata">
<emphasis id="B921EAD0FFD28A17C647230C56B4FCE2" box="[263,454,795,819]" italics="true" pageId="13" pageNumber="34">Oxynaspis celata</emphasis>
</taxonomicName>
collected in the present study (see
<taxonomicName id="4C554D41FFD28A17C40F230C54C4FCE2" box="[847,950,795,819]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="celata">
<emphasis id="B921EAD0FFD28A17C40F230C54C4FCE2" box="[847,950,795,819]" italics="true" pageId="13" pageNumber="34">O. celata</emphasis>
</taxonomicName>
in the previous section), the location of the umbo on the carina exhibited great differences between
<taxonomicName id="4C554D41FFD28A17C4492350541DFC86" box="[777,879,831,855]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="celata">
<emphasis id="B921EAD0FFD28A17C4492350541DFC86" box="[777,879,831,855]" italics="true" pageId="13" pageNumber="34">O. celata</emphasis>
</taxonomicName>
and
<taxonomicName id="4C554D41FFD28A17C4E523505339FC86" box="[933,1099,831,855]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="ryukyuensis" status="sp. nov.">
<emphasis id="B921EAD0FFD28A17C4E523505339FC86" box="[933,1099,831,855]" italics="true" pageId="13" pageNumber="34">O. ryukyuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21257ABFFD28A17C312232F53E3FC86" box="[1106,1169,831,855]" pageId="13" pageNumber="34" rank="species">
<emphasis id="B921EAD0FFD28A17C312232F53E3FC86" bold="true" box="[1106,1169,831,855]" pageId="13" pageNumber="34">n. sp.</emphasis>
</taxonomicNameLabel>
The umbo of the carina of
<taxonomicName id="4C554D41FFD28A17C7F523735669FCAA" box="[181,283,867,891]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="celata">
<emphasis id="B921EAD0FFD28A17C7F523735669FCAA" box="[181,283,867,891]" italics="true" pageId="13" pageNumber="34">O. celata</emphasis>
</taxonomicName>
was located in the proximal one third of the carina and the length of the distal arm was about twice the length of the basal arm. In
<taxonomicName id="4C554D41FFD28A17C69123975508FC4F" box="[465,634,902,926]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="ryukyuensis" status="sp. nov.">
<emphasis id="B921EAD0FFD28A17C69123975508FC4F" box="[465,634,902,926]" italics="true" pageId="13" pageNumber="34">O. ryukyuensis</emphasis>
</taxonomicName>
<emphasis id="B921EAD0FFD28A17C5C3239655B9FC4F" bold="true" box="[643,715,902,926]" pageId="13" pageNumber="34">
<taxonomicNameLabel id="A21257ABFFD28A17C5C3239655B7FC4F" box="[643,709,902,926]" pageId="13" pageNumber="34" rank="species">n. sp.</taxonomicNameLabel>
,
</emphasis>
the location of the carinal umbo was located very close to the basal region, with the length of the distal arm 7.5 times greater than the basal arm (Table 1). Both specimens of
<taxonomicName id="4C554D41FFD28A17C2C123BB57A9FC37" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="celata">
<emphasis id="B921EAD0FFD28A17C2C123BB57A9FC37" italics="true" pageId="13" pageNumber="34">O. celata</emphasis>
</taxonomicName>
and
<taxonomicName id="4C554D41FFD28A17C65323DF56C8FC37" box="[275,442,974,998]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="ryukyunesis" status="sp. nov.">
<emphasis id="B921EAD0FFD28A17C65323DF56C8FC37" box="[275,442,974,998]" italics="true" pageId="13" pageNumber="34">O. ryukyunesis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21257ABFFD28A17C68323DE5570FC37" box="[451,514,974,998]" pageId="13" pageNumber="34" rank="species">
<emphasis id="B921EAD0FFD28A17C68323DE5570FC37" bold="true" box="[451,514,974,998]" pageId="13" pageNumber="34">n. sp.</emphasis>
</taxonomicNameLabel>
were adults with mature egg masses and thus such differences in the locations of the umbones should not be due to age variation.. In addition, the dorsal side of the somatic body of
<taxonomicName id="4C554D41FFD28A17C3B523E252EEFBD8" box="[1269,1436,1009,1033]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="ryukyuensis" status="sp. nov.">
<emphasis id="B921EAD0FFD28A17C3B523E252EEFBD8" box="[1269,1436,1009,1033]" italics="true" pageId="13" pageNumber="34">O. ryukyuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21257ABFFD28A17C7D7240557AAFBFC" box="[151,216,1045,1069]" pageId="13" pageNumber="34" rank="species">
<emphasis id="B921EAD0FFD28A17C7D7240557AAFBFC" bold="true" box="[151,216,1045,1069]" pageId="13" pageNumber="34">n. sp.</emphasis>
</taxonomicNameLabel>
has two filamentary appendages, whereas filamentary appendages were absent from
<taxonomicName id="4C554D41FFD28A17C3EB24065261FBFC" box="[1195,1299,1045,1069]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="celata">
<emphasis id="B921EAD0FFD28A17C3EB24065261FBFC" box="[1195,1299,1045,1069]" italics="true" pageId="13" pageNumber="34">O. celata</emphasis>
</taxonomicName>
, suggesting that
<taxonomicName id="4C554D41FFD28A17C788242A561CFB80" box="[200,366,1081,1105]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="ryukyuensis" status="sp. nov.">
<emphasis id="B921EAD0FFD28A17C788242A561CFB80" box="[200,366,1081,1105]" italics="true" pageId="13" pageNumber="34">O. ryukyuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21257ABFFD28A17C635242956C6FB80" box="[373,436,1081,1105]" pageId="13" pageNumber="34" rank="species">
<emphasis id="B921EAD0FFD28A17C635242956C6FB80" bold="true" box="[373,436,1081,1105]" pageId="13" pageNumber="34">n. sp.</emphasis>
</taxonomicNameLabel>
differs from
<taxonomicName id="4C554D41FFD28A17C508242A55DCFB80" box="[584,686,1081,1105]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="celata">
<emphasis id="B921EAD0FFD28A17C508242A55DCFB80" box="[584,686,1081,1105]" italics="true" pageId="13" pageNumber="34">O. celata</emphasis>
</taxonomicName>
(Table 1).
</paragraph>
<paragraph id="8BEA36C2FFD28A17C787244C536FF904" blockId="13.[151,1437,151,1999]" pageId="13" pageNumber="34">
The location of the carinal umbo has been described and clearly illustrated for
<taxonomicName id="4C554D41FFD28A17C31F244D5201FBA5" authority="Darwin, 1852" authorityName="Darwin" authorityYear="1852" box="[1119,1395,1116,1141]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="celata">
<emphasis id="B921EAD0FFD28A17C31F244D53BAFBA4" box="[1119,1224,1117,1141]" italics="true" pageId="13" pageNumber="34">O. celata</emphasis>
<bibRefCitation id="EFC44B33FFD28A17C393244C5201FBA5" author="Darwin" box="[1235,1395,1116,1140]" pageId="13" pageNumber="34" refString="Darwin, C. (1852) A monograph on the sub-class Cirripedia with figures of all species. The Lepadidae: or pedunculated barnacles. Ray Society, London. 400 pp. (1851, published 1852)" type="book" year="1852">Darwin, 1852</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C554D41FFD28A17C2C1244D56ACFB49" authority="Broch, 1922" authorityName="Broch" authorityYear="1922" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="13" pageNumber="34" phylum="Arthropoda" rank="subSpecies" species="celata" subSpecies="japonica">
<emphasis id="B921EAD0FFD28A17C2C1244D5634FB49" italics="true" pageId="13" pageNumber="34">O. celata japonica</emphasis>
<bibRefCitation id="EFC44B33FFD28A17C60E249056ACFB49" author="Broch" box="[334,478,1152,1176]" pageId="13" pageNumber="34" refString="Broch, H. (1922) Papers from Dr. Th. Mortensen's Pacific Expedition 1914 - 1916, X. Studies on Pacific Cirripedes. Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening I Kobenhavn, 73, 215 - 358." type="journal article" year="1922">Broch, 1922</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C554D41FFD28A17C6AB24915448FB49" authority="Totton, 1940" authorityName="Totton" authorityYear="1940" box="[491,826,1152,1176]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="13" pageNumber="34" phylum="Arthropoda" rank="subSpecies" species="celata" subSpecies="hirtae">
<emphasis id="B921EAD0FFD28A17C6AB249155EDFB49" box="[491,671,1152,1176]" italics="true" pageId="13" pageNumber="34">O. celata hirtae</emphasis>
<bibRefCitation id="EFC44B33FFD28A17C5E724915448FB49" author="Totton" box="[679,826,1152,1176]" pageId="13" pageNumber="34" refString="Totton, A. K. (1940) New species of the cirripede genus Oxynaspis commensal with Antipatharia. The Annals and Magazine of Natural History, Series, 11, 6, 465 - 487." type="journal article" year="1940">Totton, 1940</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C554D41FFD28A17C408249153BBFB49" authority="Annandale, 1902" authorityName="Annandale" authorityYear="1902" box="[840,1225,1152,1176]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="13" pageNumber="34" phylum="Arthropoda" rank="subSpecies" species="celata" subSpecies="indica">
<emphasis id="B921EAD0FFD28A17C4082491548DFB49" box="[840,1023,1152,1176]" italics="true" pageId="13" pageNumber="34">O. celata indica</emphasis>
Annandale, 1902
</taxonomicName>
(illustrated in figs
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<bibRefCitation id="EFC44B33FFD28A17C64424B556E3FB6D" author="Totton" box="[260,401,1188,1212]" pageId="13" pageNumber="34" refString="Totton, A. K. (1940) New species of the cirripede genus Oxynaspis commensal with Antipatharia. The Annals and Magazine of Natural History, Series, 11, 6, 465 - 487." type="journal article" year="1940">Totton 1940</bibRefCitation>
; redrawn in
<figureCitation id="136E2A47FFD28A17C56324B45504FB6D" box="[547,630,1188,1212]" captionStart="FIGURE 16" captionStartId="24.[151,244,1109,1130]" captionTargetBox="[156,1422,164,1070]" captionTargetId="figure@24.[95,1444,108,1088]" captionTargetPageId="24" captionText="FIGURE 16. Comparisons between Calantica arcuata n. sp. (I) and allied species (Group I sensu Newman &amp; Jones, 2011), and between Oxynaspis ryukyuensis n. sp. (N) and allied congeners. A, C. darwini Jones &amp; Hoise, 2009 (redrawn from Jones &amp; Hoise 2009); B, C. pusilla Utinomi, 1970 a (redrawn from Utinomi 1970 a); C, C. graphica Rosell, 1991 (redrawn from Rosell 1991); D, C. kruegeri Hiro, 1932 (redrawn from Hiro 1932); E, C. studeri (Weltner, 1922) (redrawn from Zevina 1978); F, C. trispinosa Hoek, 1883 (redrawn from Hoek 1883); G, C. eos (Pilsbry, 1907) (redrawn from Pilsbry 1907); H, C. affinis (Broch, 1922) (redrawn from Broch 1922); I, C. arcuata n. sp. (present study, RUMF-ZC- 1554); J, Oxynaspis alatae Totton, 1940 (redrawn from Totton 1940); K, Oxynaspis gracilis Totton, 1940 (redrawn from Totton 1940); L, O. celata Darwin, 1852 (redrawn from Darwin 1852); M. O. celata japonica Broch, 1922 (redrawn from Broch 1922); N, O. c. hirtae Totton, 1940 (redrawn from Totton, 1940); O, O. c. indica Annandale, 1909 (redrawn from Totton 1940); P, O. cancellatae Totton, 1940 (redrawn from Totton 1940); Q, O. ryukyuensis n. sp. (present study, (RUMF-ZC- 1558). Arrow in I indicates the diagnostic carina in C. arcuata n. sp., u indicates the location of the umbo of the carina in Oxynaspis species. The distal arm indicates the length of carinal margin above the umbo, basal arm indicate length of carinal margin below the umbo (see J). The length ratio of the distal arm to basal arm is a diagnostic character to identify Oxynaspis species." httpUri="https://zenodo.org/record/281668/files/figure.png" pageId="13" pageNumber="34">Fig. 16</figureCitation>
),
<taxonomicName id="4C554D41FFD28A17C5CD24B554A2FB6D" authority="Totton, 1940" authorityName="Totton" authorityYear="1940" box="[653,976,1188,1212]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="cancellatae">
<emphasis id="B921EAD0FFD28A17C5CD24B55440FB6D" box="[653,818,1188,1212]" italics="true" pageId="13" pageNumber="34">O. cancellatae</emphasis>
<bibRefCitation id="EFC44B33FFD28A17C47C24B554A2FB6D" author="Totton" box="[828,976,1188,1212]" pageId="13" pageNumber="34" refString="Totton, A. K. (1940) New species of the cirripede genus Oxynaspis commensal with Antipatharia. The Annals and Magazine of Natural History, Series, 11, 6, 465 - 487." type="journal article" year="1940">Totton, 1940</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C554D41FFD28A17C49C24B55381FB6D" authority="Totton, 1940" authorityName="Totton" authorityYear="1940" box="[988,1267,1188,1212]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="B921EAD0FFD28A17C49C24B55325FB6D" box="[988,1111,1188,1212]" italics="true" pageId="13" pageNumber="34">O. gracilis</emphasis>
<bibRefCitation id="EFC44B33FFD28A17C31F24B55381FB6D" author="Totton" box="[1119,1267,1188,1212]" pageId="13" pageNumber="34" refString="Totton, A. K. (1940) New species of the cirripede genus Oxynaspis commensal with Antipatharia. The Annals and Magazine of Natural History, Series, 11, 6, 465 - 487." type="journal article" year="1940">Totton, 1940</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4C554D41FFD28A17C27224B55659FB0E" authority="Totton, 1940" authorityName="Totton" authorityYear="1940" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="alatae">
<emphasis id="B921EAD0FFD28A17C27224B552EEFB6D" box="[1330,1436,1188,1212]" italics="true" pageId="13" pageNumber="34">O. alatae</emphasis>
<bibRefCitation id="EFC44B33FFD28A17C7D724D95659FB0E" author="Totton" box="[151,299,1223,1248]" pageId="13" pageNumber="34" refString="Totton, A. K. (1940) New species of the cirripede genus Oxynaspis commensal with Antipatharia. The Annals and Magazine of Natural History, Series, 11, 6, 465 - 487." type="journal article" year="1940">Totton, 1940</bibRefCitation>
</taxonomicName>
. The umbo of the carina in
<taxonomicName id="4C554D41FFD28A17C52524D855BFFB31" box="[613,717,1224,1248]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="celata">
<emphasis id="B921EAD0FFD28A17C52524D855BFFB31" box="[613,717,1224,1248]" italics="true" pageId="13" pageNumber="34">O. celata</emphasis>
</taxonomicName>
(see pl.
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<bibRefCitation id="EFC44B33FFD28A17C42424D75489FB0E" author="Darwin" box="[868,1019,1223,1247]" pageId="13" pageNumber="34" refString="Darwin, C. (1852) A monograph on the sub-class Cirripedia with figures of all species. The Lepadidae: or pedunculated barnacles. Ray Society, London. 400 pp. (1851, published 1852)" type="book" year="1852">Darwin 1852</bibRefCitation>
;
<figureCitation id="136E2A47FFD28A17C34824D7532BFB0E" box="[1032,1113,1223,1247]" captionStart="FIGURE 16" captionStartId="24.[151,244,1109,1130]" captionTargetBox="[156,1422,164,1070]" captionTargetId="figure@24.[95,1444,108,1088]" captionTargetPageId="24" captionText="FIGURE 16. Comparisons between Calantica arcuata n. sp. (I) and allied species (Group I sensu Newman &amp; Jones, 2011), and between Oxynaspis ryukyuensis n. sp. (N) and allied congeners. A, C. darwini Jones &amp; Hoise, 2009 (redrawn from Jones &amp; Hoise 2009); B, C. pusilla Utinomi, 1970 a (redrawn from Utinomi 1970 a); C, C. graphica Rosell, 1991 (redrawn from Rosell 1991); D, C. kruegeri Hiro, 1932 (redrawn from Hiro 1932); E, C. studeri (Weltner, 1922) (redrawn from Zevina 1978); F, C. trispinosa Hoek, 1883 (redrawn from Hoek 1883); G, C. eos (Pilsbry, 1907) (redrawn from Pilsbry 1907); H, C. affinis (Broch, 1922) (redrawn from Broch 1922); I, C. arcuata n. sp. (present study, RUMF-ZC- 1554); J, Oxynaspis alatae Totton, 1940 (redrawn from Totton 1940); K, Oxynaspis gracilis Totton, 1940 (redrawn from Totton 1940); L, O. celata Darwin, 1852 (redrawn from Darwin 1852); M. O. celata japonica Broch, 1922 (redrawn from Broch 1922); N, O. c. hirtae Totton, 1940 (redrawn from Totton, 1940); O, O. c. indica Annandale, 1909 (redrawn from Totton 1940); P, O. cancellatae Totton, 1940 (redrawn from Totton 1940); Q, O. ryukyuensis n. sp. (present study, (RUMF-ZC- 1558). Arrow in I indicates the diagnostic carina in C. arcuata n. sp., u indicates the location of the umbo of the carina in Oxynaspis species. The distal arm indicates the length of carinal margin above the umbo, basal arm indicate length of carinal margin below the umbo (see J). The length ratio of the distal arm to basal arm is a diagnostic character to identify Oxynaspis species." httpUri="https://zenodo.org/record/281668/files/figure.png" pageId="13" pageNumber="34">Fig. 16</figureCitation>
L),
<taxonomicName id="4C554D41FFD28A17C3C224D85247FB31" box="[1154,1333,1224,1248]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="13" pageNumber="34" phylum="Arthropoda" rank="subSpecies" species="celata" subSpecies="hirtae">
<emphasis id="B921EAD0FFD28A17C3C224D85247FB31" box="[1154,1333,1224,1248]" italics="true" pageId="13" pageNumber="34">O. celata hirtae</emphasis>
</taxonomicName>
(see figs 10, 11 in
<bibRefCitation id="EFC44B33FFD28A17C64324FC56FDFAD2" author="Totton" box="[259,399,1259,1283]" pageId="13" pageNumber="34" refString="Totton, A. K. (1940) New species of the cirripede genus Oxynaspis commensal with Antipatharia. The Annals and Magazine of Natural History, Series, 11, 6, 465 - 487." type="journal article" year="1940">Totton 1940</bibRefCitation>
;
<figureCitation id="136E2A47FFD28A17C6DC24FB569CFAD2" box="[412,494,1259,1283]" captionStart="FIGURE 16" captionStartId="24.[151,244,1109,1130]" captionTargetBox="[156,1422,164,1070]" captionTargetId="figure@24.[95,1444,108,1088]" captionTargetPageId="24" captionText="FIGURE 16. Comparisons between Calantica arcuata n. sp. (I) and allied species (Group I sensu Newman &amp; Jones, 2011), and between Oxynaspis ryukyuensis n. sp. (N) and allied congeners. A, C. darwini Jones &amp; Hoise, 2009 (redrawn from Jones &amp; Hoise 2009); B, C. pusilla Utinomi, 1970 a (redrawn from Utinomi 1970 a); C, C. graphica Rosell, 1991 (redrawn from Rosell 1991); D, C. kruegeri Hiro, 1932 (redrawn from Hiro 1932); E, C. studeri (Weltner, 1922) (redrawn from Zevina 1978); F, C. trispinosa Hoek, 1883 (redrawn from Hoek 1883); G, C. eos (Pilsbry, 1907) (redrawn from Pilsbry 1907); H, C. affinis (Broch, 1922) (redrawn from Broch 1922); I, C. arcuata n. sp. (present study, RUMF-ZC- 1554); J, Oxynaspis alatae Totton, 1940 (redrawn from Totton 1940); K, Oxynaspis gracilis Totton, 1940 (redrawn from Totton 1940); L, O. celata Darwin, 1852 (redrawn from Darwin 1852); M. O. celata japonica Broch, 1922 (redrawn from Broch 1922); N, O. c. hirtae Totton, 1940 (redrawn from Totton, 1940); O, O. c. indica Annandale, 1909 (redrawn from Totton 1940); P, O. cancellatae Totton, 1940 (redrawn from Totton 1940); Q, O. ryukyuensis n. sp. (present study, (RUMF-ZC- 1558). Arrow in I indicates the diagnostic carina in C. arcuata n. sp., u indicates the location of the umbo of the carina in Oxynaspis species. The distal arm indicates the length of carinal margin above the umbo, basal arm indicate length of carinal margin below the umbo (see J). The length ratio of the distal arm to basal arm is a diagnostic character to identify Oxynaspis species." httpUri="https://zenodo.org/record/281668/files/figure.png" pageId="13" pageNumber="34">Fig. 16</figureCitation>
N),
<taxonomicName id="4C554D41FFD28A17C55A24FC559EFAD5" box="[538,748,1260,1284]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="13" pageNumber="34" phylum="Arthropoda" rank="subSpecies" species="celata" subSpecies="japonica">
<emphasis id="B921EAD0FFD28A17C55A24FC559EFAD5" box="[538,748,1260,1284]" italics="true" pageId="13" pageNumber="34">O. celata japonica</emphasis>
</taxonomicName>
(see fig.
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<bibRefCitation id="EFC44B33FFD28A17C4DA24FB5350FAD2" author="Broch" box="[922,1058,1259,1283]" pageId="13" pageNumber="34" refString="Broch, H. (1922) Papers from Dr. Th. Mortensen's Pacific Expedition 1914 - 1916, X. Studies on Pacific Cirripedes. Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening I Kobenhavn, 73, 215 - 358." type="journal article" year="1922">Broch 1922</bibRefCitation>
;
<figureCitation id="136E2A47FFD28A17C36F24FB530DFAD2" box="[1071,1151,1259,1283]" captionStart="FIGURE 16" captionStartId="24.[151,244,1109,1130]" captionTargetBox="[156,1422,164,1070]" captionTargetId="figure@24.[95,1444,108,1088]" captionTargetPageId="24" captionText="FIGURE 16. Comparisons between Calantica arcuata n. sp. (I) and allied species (Group I sensu Newman &amp; Jones, 2011), and between Oxynaspis ryukyuensis n. sp. (N) and allied congeners. A, C. darwini Jones &amp; Hoise, 2009 (redrawn from Jones &amp; Hoise 2009); B, C. pusilla Utinomi, 1970 a (redrawn from Utinomi 1970 a); C, C. graphica Rosell, 1991 (redrawn from Rosell 1991); D, C. kruegeri Hiro, 1932 (redrawn from Hiro 1932); E, C. studeri (Weltner, 1922) (redrawn from Zevina 1978); F, C. trispinosa Hoek, 1883 (redrawn from Hoek 1883); G, C. eos (Pilsbry, 1907) (redrawn from Pilsbry 1907); H, C. affinis (Broch, 1922) (redrawn from Broch 1922); I, C. arcuata n. sp. (present study, RUMF-ZC- 1554); J, Oxynaspis alatae Totton, 1940 (redrawn from Totton 1940); K, Oxynaspis gracilis Totton, 1940 (redrawn from Totton 1940); L, O. celata Darwin, 1852 (redrawn from Darwin 1852); M. O. celata japonica Broch, 1922 (redrawn from Broch 1922); N, O. c. hirtae Totton, 1940 (redrawn from Totton, 1940); O, O. c. indica Annandale, 1909 (redrawn from Totton 1940); P, O. cancellatae Totton, 1940 (redrawn from Totton 1940); Q, O. ryukyuensis n. sp. (present study, (RUMF-ZC- 1558). Arrow in I indicates the diagnostic carina in C. arcuata n. sp., u indicates the location of the umbo of the carina in Oxynaspis species. The distal arm indicates the length of carinal margin above the umbo, basal arm indicate length of carinal margin below the umbo (see J). The length ratio of the distal arm to basal arm is a diagnostic character to identify Oxynaspis species." httpUri="https://zenodo.org/record/281668/files/figure.png" pageId="13" pageNumber="34">Fig. 16</figureCitation>
M),
<taxonomicName id="4C554D41FFD28A17C3F024FC5215FAD5" box="[1200,1383,1260,1284]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="13" pageNumber="34" phylum="Arthropoda" rank="subSpecies" species="celata" subSpecies="indica">
<emphasis id="B921EAD0FFD28A17C3F024FC5215FAD5" box="[1200,1383,1260,1284]" italics="true" pageId="13" pageNumber="34">O. celata indica</emphasis>
</taxonomicName>
(see figs
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<bibRefCitation id="EFC44B33FFD28A17C674250056CDFAF6" author="Totton" box="[308,447,1295,1319]" pageId="13" pageNumber="34" refString="Totton, A. K. (1940) New species of the cirripede genus Oxynaspis commensal with Antipatharia. The Annals and Magazine of Natural History, Series, 11, 6, 465 - 487." type="journal article" year="1940">Totton 1940</bibRefCitation>
;
<figureCitation id="136E2A47FFD28A17C68C251F556FFAF6" box="[460,541,1295,1319]" captionStart="FIGURE 16" captionStartId="24.[151,244,1109,1130]" captionTargetBox="[156,1422,164,1070]" captionTargetId="figure@24.[95,1444,108,1088]" captionTargetPageId="24" captionText="FIGURE 16. Comparisons between Calantica arcuata n. sp. (I) and allied species (Group I sensu Newman &amp; Jones, 2011), and between Oxynaspis ryukyuensis n. sp. (N) and allied congeners. A, C. darwini Jones &amp; Hoise, 2009 (redrawn from Jones &amp; Hoise 2009); B, C. pusilla Utinomi, 1970 a (redrawn from Utinomi 1970 a); C, C. graphica Rosell, 1991 (redrawn from Rosell 1991); D, C. kruegeri Hiro, 1932 (redrawn from Hiro 1932); E, C. studeri (Weltner, 1922) (redrawn from Zevina 1978); F, C. trispinosa Hoek, 1883 (redrawn from Hoek 1883); G, C. eos (Pilsbry, 1907) (redrawn from Pilsbry 1907); H, C. affinis (Broch, 1922) (redrawn from Broch 1922); I, C. arcuata n. sp. (present study, RUMF-ZC- 1554); J, Oxynaspis alatae Totton, 1940 (redrawn from Totton 1940); K, Oxynaspis gracilis Totton, 1940 (redrawn from Totton 1940); L, O. celata Darwin, 1852 (redrawn from Darwin 1852); M. O. celata japonica Broch, 1922 (redrawn from Broch 1922); N, O. c. hirtae Totton, 1940 (redrawn from Totton, 1940); O, O. c. indica Annandale, 1909 (redrawn from Totton 1940); P, O. cancellatae Totton, 1940 (redrawn from Totton 1940); Q, O. ryukyuensis n. sp. (present study, (RUMF-ZC- 1558). Arrow in I indicates the diagnostic carina in C. arcuata n. sp., u indicates the location of the umbo of the carina in Oxynaspis species. The distal arm indicates the length of carinal margin above the umbo, basal arm indicate length of carinal margin below the umbo (see J). The length ratio of the distal arm to basal arm is a diagnostic character to identify Oxynaspis species." httpUri="https://zenodo.org/record/281668/files/figure.png" pageId="13" pageNumber="34">Fig. 16</figureCitation>
O),
<taxonomicName id="4C554D41FFD28A17C508250055B3FAF6" box="[584,705,1295,1319]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="B921EAD0FFD28A17C508250055B3FAF6" box="[584,705,1295,1319]" italics="true" pageId="13" pageNumber="34">O. gracilis</emphasis>
</taxonomicName>
(see fig.
<quantity id="4CAD9B27FFD28A17C46C251F5425FAF6" box="[812,855,1295,1319]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="13" pageNumber="34" unit="in" value="8.0">8 in</quantity>
<bibRefCitation id="EFC44B33FFD28A17C41F25005499FAF6" author="Totton" box="[863,1003,1295,1319]" pageId="13" pageNumber="34" refString="Totton, A. K. (1940) New species of the cirripede genus Oxynaspis commensal with Antipatharia. The Annals and Magazine of Natural History, Series, 11, 6, 465 - 487." type="journal article" year="1940">Totton 1940</bibRefCitation>
;
<figureCitation id="136E2A47FFD28A17C4B7251F5334FAF6" box="[1015,1094,1295,1319]" captionStart="FIGURE 16" captionStartId="24.[151,244,1109,1130]" captionTargetBox="[156,1422,164,1070]" captionTargetId="figure@24.[95,1444,108,1088]" captionTargetPageId="24" captionText="FIGURE 16. Comparisons between Calantica arcuata n. sp. (I) and allied species (Group I sensu Newman &amp; Jones, 2011), and between Oxynaspis ryukyuensis n. sp. (N) and allied congeners. A, C. darwini Jones &amp; Hoise, 2009 (redrawn from Jones &amp; Hoise 2009); B, C. pusilla Utinomi, 1970 a (redrawn from Utinomi 1970 a); C, C. graphica Rosell, 1991 (redrawn from Rosell 1991); D, C. kruegeri Hiro, 1932 (redrawn from Hiro 1932); E, C. studeri (Weltner, 1922) (redrawn from Zevina 1978); F, C. trispinosa Hoek, 1883 (redrawn from Hoek 1883); G, C. eos (Pilsbry, 1907) (redrawn from Pilsbry 1907); H, C. affinis (Broch, 1922) (redrawn from Broch 1922); I, C. arcuata n. sp. (present study, RUMF-ZC- 1554); J, Oxynaspis alatae Totton, 1940 (redrawn from Totton 1940); K, Oxynaspis gracilis Totton, 1940 (redrawn from Totton 1940); L, O. celata Darwin, 1852 (redrawn from Darwin 1852); M. O. celata japonica Broch, 1922 (redrawn from Broch 1922); N, O. c. hirtae Totton, 1940 (redrawn from Totton, 1940); O, O. c. indica Annandale, 1909 (redrawn from Totton 1940); P, O. cancellatae Totton, 1940 (redrawn from Totton 1940); Q, O. ryukyuensis n. sp. (present study, (RUMF-ZC- 1558). Arrow in I indicates the diagnostic carina in C. arcuata n. sp., u indicates the location of the umbo of the carina in Oxynaspis species. The distal arm indicates the length of carinal margin above the umbo, basal arm indicate length of carinal margin below the umbo (see J). The length ratio of the distal arm to basal arm is a diagnostic character to identify Oxynaspis species." httpUri="https://zenodo.org/record/281668/files/figure.png" pageId="13" pageNumber="34">Fig. 16</figureCitation>
K) and
<taxonomicName id="4C554D41FFD28A17C3DD25005274FAF6" box="[1181,1286,1295,1319]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="alatae">
<emphasis id="B921EAD0FFD28A17C3DD25005274FAF6" box="[1181,1286,1295,1319]" italics="true" pageId="13" pageNumber="34">O. alatae</emphasis>
</taxonomicName>
(see fig.
<quantity id="4CAD9B27FFD28A17C230251F52EEFAF6" box="[1392,1436,1295,1319]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="13" pageNumber="34" unit="in" value="3.0">3 in</quantity>
<bibRefCitation id="EFC44B33FFD28A17C7D725245653FA9A" author="Totton" box="[151,289,1331,1355]" pageId="13" pageNumber="34" refString="Totton, A. K. (1940) New species of the cirripede genus Oxynaspis commensal with Antipatharia. The Annals and Magazine of Natural History, Series, 11, 6, 465 - 487." type="journal article" year="1940">Totton 1940</bibRefCitation>
;
<figureCitation id="136E2A47FFD28A17C66C25235609FA9A" box="[300,379,1331,1355]" captionStart="FIGURE 16" captionStartId="24.[151,244,1109,1130]" captionTargetBox="[156,1422,164,1070]" captionTargetId="figure@24.[95,1444,108,1088]" captionTargetPageId="24" captionText="FIGURE 16. Comparisons between Calantica arcuata n. sp. (I) and allied species (Group I sensu Newman &amp; Jones, 2011), and between Oxynaspis ryukyuensis n. sp. (N) and allied congeners. A, C. darwini Jones &amp; Hoise, 2009 (redrawn from Jones &amp; Hoise 2009); B, C. pusilla Utinomi, 1970 a (redrawn from Utinomi 1970 a); C, C. graphica Rosell, 1991 (redrawn from Rosell 1991); D, C. kruegeri Hiro, 1932 (redrawn from Hiro 1932); E, C. studeri (Weltner, 1922) (redrawn from Zevina 1978); F, C. trispinosa Hoek, 1883 (redrawn from Hoek 1883); G, C. eos (Pilsbry, 1907) (redrawn from Pilsbry 1907); H, C. affinis (Broch, 1922) (redrawn from Broch 1922); I, C. arcuata n. sp. (present study, RUMF-ZC- 1554); J, Oxynaspis alatae Totton, 1940 (redrawn from Totton 1940); K, Oxynaspis gracilis Totton, 1940 (redrawn from Totton 1940); L, O. celata Darwin, 1852 (redrawn from Darwin 1852); M. O. celata japonica Broch, 1922 (redrawn from Broch 1922); N, O. c. hirtae Totton, 1940 (redrawn from Totton, 1940); O, O. c. indica Annandale, 1909 (redrawn from Totton 1940); P, O. cancellatae Totton, 1940 (redrawn from Totton 1940); Q, O. ryukyuensis n. sp. (present study, (RUMF-ZC- 1558). Arrow in I indicates the diagnostic carina in C. arcuata n. sp., u indicates the location of the umbo of the carina in Oxynaspis species. The distal arm indicates the length of carinal margin above the umbo, basal arm indicate length of carinal margin below the umbo (see J). The length ratio of the distal arm to basal arm is a diagnostic character to identify Oxynaspis species." httpUri="https://zenodo.org/record/281668/files/figure.png" pageId="13" pageNumber="34">Fig. 16</figureCitation>
J) are expanded, forming a distinct angle and are located at the middle or proximal one third of the carina, differing from the specimens of
<taxonomicName id="4C554D41FFD28A17C5C72548545DFABE" box="[647,815,1367,1391]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="ryukyuensis" status="sp. nov.">
<emphasis id="B921EAD0FFD28A17C5C72548545DFABE" box="[647,815,1367,1391]" italics="true" pageId="13" pageNumber="34">O. ryukyuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21257ABFFD28A17C4772547540AFABE" box="[823,888,1367,1391]" pageId="13" pageNumber="34" rank="species">
<emphasis id="B921EAD0FFD28A17C4772547540AFABE" bold="true" box="[823,888,1367,1391]" pageId="13" pageNumber="34">n. sp.</emphasis>
</taxonomicNameLabel>
, which have the umbo located very close to the basal region of the carina (
<figureCitation id="136E2A47FFD28A17C692256A5554FA43" box="[466,550,1402,1426]" captionStart="FIGURE 16" captionStartId="24.[151,244,1109,1130]" captionTargetBox="[156,1422,164,1070]" captionTargetId="figure@24.[95,1444,108,1088]" captionTargetPageId="24" captionText="FIGURE 16. Comparisons between Calantica arcuata n. sp. (I) and allied species (Group I sensu Newman &amp; Jones, 2011), and between Oxynaspis ryukyuensis n. sp. (N) and allied congeners. A, C. darwini Jones &amp; Hoise, 2009 (redrawn from Jones &amp; Hoise 2009); B, C. pusilla Utinomi, 1970 a (redrawn from Utinomi 1970 a); C, C. graphica Rosell, 1991 (redrawn from Rosell 1991); D, C. kruegeri Hiro, 1932 (redrawn from Hiro 1932); E, C. studeri (Weltner, 1922) (redrawn from Zevina 1978); F, C. trispinosa Hoek, 1883 (redrawn from Hoek 1883); G, C. eos (Pilsbry, 1907) (redrawn from Pilsbry 1907); H, C. affinis (Broch, 1922) (redrawn from Broch 1922); I, C. arcuata n. sp. (present study, RUMF-ZC- 1554); J, Oxynaspis alatae Totton, 1940 (redrawn from Totton 1940); K, Oxynaspis gracilis Totton, 1940 (redrawn from Totton 1940); L, O. celata Darwin, 1852 (redrawn from Darwin 1852); M. O. celata japonica Broch, 1922 (redrawn from Broch 1922); N, O. c. hirtae Totton, 1940 (redrawn from Totton, 1940); O, O. c. indica Annandale, 1909 (redrawn from Totton 1940); P, O. cancellatae Totton, 1940 (redrawn from Totton 1940); Q, O. ryukyuensis n. sp. (present study, (RUMF-ZC- 1558). Arrow in I indicates the diagnostic carina in C. arcuata n. sp., u indicates the location of the umbo of the carina in Oxynaspis species. The distal arm indicates the length of carinal margin above the umbo, basal arm indicate length of carinal margin below the umbo (see J). The length ratio of the distal arm to basal arm is a diagnostic character to identify Oxynaspis species." httpUri="https://zenodo.org/record/281668/files/figure.png" pageId="13" pageNumber="34">Fig. 16</figureCitation>
Q; Table 1). Measuring the
<typeStatus id="54EE8860FFD28A17C42B256B54E9FA42" box="[875,923,1403,1427]" pageId="13" pageNumber="34">type</typeStatus>
illustration of
<emphasis id="B921EAD0FFD28A17C30C256B53B3FA43" box="[1100,1217,1403,1426]" italics="true" pageId="13" pageNumber="34">Oxynapsis</emphasis>
species mentioned above (except
<taxonomicName id="4C554D41FFD28A17C67E258F56DBFA67" box="[318,425,1438,1462]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="indica">
<emphasis id="B921EAD0FFD28A17C67E258F56DBFA67" box="[318,425,1438,1462]" italics="true" pageId="13" pageNumber="34">O. indica</emphasis>
</taxonomicName>
illustrated in
<bibRefCitation id="EFC44B33FFD28A17C50A258F55AAFA67" author="Totton" box="[586,728,1438,1462]" pageId="13" pageNumber="34" refString="Totton, A. K. (1940) New species of the cirripede genus Oxynaspis commensal with Antipatharia. The Annals and Magazine of Natural History, Series, 11, 6, 465 - 487." type="journal article" year="1940">Totton 1940</bibRefCitation>
), the length ratio of the distal arm to basal arm was
<quantity id="4CAD9B27FFD28A17C276258E520AFA67" box="[1334,1400,1438,1462]" metricMagnitude="-2" metricUnit="m" metricValue="4.826" pageId="13" pageNumber="34" unit="in" value="1.9">1.9 in</quantity>
<taxonomicName id="4C554D41FFD28A17C2C1258F57AFFA0B" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="alatae">
<emphasis id="B921EAD0FFD28A17C2C1258F57AFFA0B" italics="true" pageId="13" pageNumber="34">O. alatae</emphasis>
</taxonomicName>
,
<quantity id="4CAD9B27FFD28A17C7AF25D25641FA0B" box="[239,307,1474,1498]" metricMagnitude="-2" metricUnit="m" metricValue="3.81" pageId="13" pageNumber="34" unit="in" value="1.5">1.5 in</quantity>
<taxonomicName id="4C554D41FFD28A17C67E25D356CBFA0B" box="[318,441,1474,1498]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="B921EAD0FFD28A17C67E25D356CBFA0B" box="[318,441,1474,1498]" italics="true" pageId="13" pageNumber="34">O. gracilis</emphasis>
</taxonomicName>
,
<quantity id="4CAD9B27FFD28A17C68C25D25562FA0B" box="[460,528,1474,1498]" metricMagnitude="-2" metricUnit="m" metricValue="4.826" pageId="13" pageNumber="34" unit="in" value="1.9">1.9 in</quantity>
<taxonomicName id="4C554D41FFD28A17C55B25D355F4FA0B" box="[539,646,1474,1498]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="celata">
<emphasis id="B921EAD0FFD28A17C55B25D355F4FA0B" box="[539,646,1474,1498]" italics="true" pageId="13" pageNumber="34">O. celata</emphasis>
</taxonomicName>
,
<quantity id="4CAD9B27FFD28A17C5D825D255AEFA0B" box="[664,732,1474,1498]" metricMagnitude="-2" metricUnit="m" metricValue="5.334" pageId="13" pageNumber="34" unit="in" value="2.1">2.1 in</quantity>
<taxonomicName id="4C554D41FFD28A17C5A725D354B2FA0B" box="[743,960,1474,1498]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="13" pageNumber="34" phylum="Arthropoda" rank="subSpecies" species="celata" subSpecies="japonica">
<emphasis id="B921EAD0FFD28A17C5A725D354B2FA0B" box="[743,960,1474,1498]" italics="true" pageId="13" pageNumber="34">O. celata japonica</emphasis>
</taxonomicName>
,
<quantity id="4CAD9B27FFD28A17C49225D25364FA0B" box="[978,1046,1474,1498]" metricMagnitude="-2" metricUnit="m" metricValue="6.096" pageId="13" pageNumber="34" unit="in" value="2.4">2.4 in</quantity>
<taxonomicName id="4C554D41FFD28A17C36125D353DBFA0B" box="[1057,1193,1474,1498]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="13" pageNumber="34" phylum="Arthropoda" rank="subSpecies" species="celata" subSpecies="hirtae">
<emphasis id="B921EAD0FFD28A17C36125D353DBFA0B" box="[1057,1193,1474,1498]" italics="true" pageId="13" pageNumber="34">O. c. hirtae</emphasis>
</taxonomicName>
,
<quantity id="4CAD9B27FFD28A17C3FB25D2538DFA0B" box="[1211,1279,1474,1498]" metricMagnitude="-2" metricUnit="m" metricValue="4.318" pageId="13" pageNumber="34" unit="in" value="1.7">1.7 in</quantity>
<taxonomicName id="4C554D41FFD28A17C24A25D352E7FA0B" box="[1290,1429,1474,1498]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="13" pageNumber="34" phylum="Arthropoda" rank="subSpecies" species="celata" subSpecies="indica">
<emphasis id="B921EAD0FFD28A17C24A25D352E7FA0B" box="[1290,1429,1474,1498]" italics="true" pageId="13" pageNumber="34">O. c. indica</emphasis>
</taxonomicName>
, obviously differing from
<quantity id="4CAD9B27FFD28A17C6FE25F65570FA2F" box="[446,514,1510,1534]" metricMagnitude="-1" metricUnit="m" metricValue="1.905" pageId="13" pageNumber="34" unit="in" value="7.5">7.5 in</quantity>
<taxonomicName id="4C554D41FFD28A17C54D25F755CAFA2F" box="[525,696,1510,1534]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="ryukyuensis" status="sp. nov.">
<emphasis id="B921EAD0FFD28A17C54D25F755CAFA2F" box="[525,696,1510,1534]" italics="true" pageId="13" pageNumber="34">O. ryukyuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21257ABFFD28A17C58325F65477FA2F" box="[707,773,1510,1534]" pageId="13" pageNumber="34" rank="species">
<emphasis id="B921EAD0FFD28A17C58325F65477FA2F" bold="true" box="[707,773,1510,1534]" pageId="13" pageNumber="34">n. sp.</emphasis>
</taxonomicNameLabel>
(Table 1). The basal location of the carinal umbo of
<taxonomicName id="4C554D41FFD28A17C2C125F75669F9F0" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="ryukyuensis" status="sp. nov.">
<emphasis id="B921EAD0FFD28A17C2C125F75669F9F0" italics="true" pageId="13" pageNumber="34">O. ryukyuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21257ABFFD28A17C662261A5612F9F3" box="[290,352,1546,1570]" pageId="13" pageNumber="34" rank="species">
<emphasis id="B921EAD0FFD28A17C662261A5612F9F3" bold="true" box="[290,352,1546,1570]" pageId="13" pageNumber="34">n. sp.</emphasis>
</taxonomicNameLabel>
is consistent in the
<typeStatus id="54EE8860FFD28A17C57D261955EDF9F0" box="[573,671,1545,1569]" pageId="13" pageNumber="34" type="holotype">holotype</typeStatus>
and the two
<typeStatus id="54EE8860FFD28A17C46E261A54EFF9F3" box="[814,925,1546,1570]" pageId="13" pageNumber="34" type="paratype">paratypes</typeStatus>
. The length ratio of the distal arm to the basal arm is 7.9 and 8.0 for the two
<typeStatus id="54EE8860FFD28A17C6B4263E5511F997" box="[500,611,1582,1606]" pageId="13" pageNumber="34" type="paratype">paratypes</typeStatus>
. There is no detailed description of
<emphasis id="B921EAD0FFD28A17C4BD263E53ACF994" box="[1021,1246,1581,1605]" italics="true" pageId="13" pageNumber="34">O. c. novazelandica</emphasis>
in
<bibRefCitation id="EFC44B33FFD28A17C244263D52EEF994" author="Broch" box="[1284,1436,1581,1605]" pageId="13" pageNumber="34" refString="Broch, H. (1922) Papers from Dr. Th. Mortensen's Pacific Expedition 1914 - 1916, X. Studies on Pacific Cirripedes. Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening I Kobenhavn, 73, 215 - 358." type="journal article" year="1922">Broch (1922)</bibRefCitation>
but
<emphasis id="B921EAD0FFD28A17C786264256DFF9B8" box="[198,429,1617,1641]" italics="true" pageId="13" pageNumber="34">O. c. novazelandica</emphasis>
was described as morphologically very close to
<taxonomicName id="4C554D41FFD28A17C4AE2642530BF9B8" box="[1006,1145,1617,1641]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="13" pageNumber="34" phylum="Arthropoda" rank="subSpecies" species="celata" subSpecies="indica">
<emphasis id="B921EAD0FFD28A17C4AE2642530BF9B8" box="[1006,1145,1617,1641]" italics="true" pageId="13" pageNumber="34">O. c. indica</emphasis>
</taxonomicName>
, with only the maxillule differing between the two species. In the present study, we believe the external morphology of
<emphasis id="B921EAD0FFD28A17C3FD266552E9F95C" box="[1213,1435,1653,1677]" italics="true" pageId="13" pageNumber="34">O. c. novazelandica</emphasis>
is close to that of
<taxonomicName id="4C554D41FFD28A17C6262689569EF961" box="[358,492,1688,1712]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="13" pageNumber="34" phylum="Arthropoda" rank="subSpecies" species="celata" subSpecies="indica">
<emphasis id="B921EAD0FFD28A17C6262689569EF961" box="[358,492,1688,1712]" italics="true" pageId="13" pageNumber="34">O. c. indica</emphasis>
</taxonomicName>
and thus morphologically differs from
<taxonomicName id="4C554D41FFD28A17C4F02689532AF961" box="[944,1112,1688,1712]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="ryukyuensis" status="sp. nov.">
<emphasis id="B921EAD0FFD28A17C4F02689532AF961" box="[944,1112,1688,1712]" italics="true" pageId="13" pageNumber="34">O. ryukyuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21257ABFFD28A17C321268953D0F960" box="[1121,1186,1689,1713]" pageId="13" pageNumber="34" rank="species">
<emphasis id="B921EAD0FFD28A17C321268953D0F960" bold="true" box="[1121,1186,1689,1713]" pageId="13" pageNumber="34">n. sp.</emphasis>
</taxonomicNameLabel>
, therefore confirming that
<taxonomicName id="4C554D41FFD28A17C78826AD561CF905" box="[200,366,1724,1748]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="ryukyuensis" status="sp. nov.">
<emphasis id="B921EAD0FFD28A17C78826AD561CF905" box="[200,366,1724,1748]" italics="true" pageId="13" pageNumber="34">O. ryukyuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21257ABFFD28A17C63526AD56C6F904" box="[373,436,1725,1749]" pageId="13" pageNumber="34" rank="species">
<emphasis id="B921EAD0FFD28A17C63526AD56C6F904" bold="true" box="[373,436,1725,1749]" pageId="13" pageNumber="34">n. sp.</emphasis>
</taxonomicNameLabel>
does not belong to the
<taxonomicName id="4C554D41FFD28A17C5F926AD5452F905" box="[697,800,1724,1748]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="celata">
<emphasis id="B921EAD0FFD28A17C5F926AD5452F905" box="[697,800,1724,1748]" italics="true" pageId="13" pageNumber="34">O. celata</emphasis>
</taxonomicName>
cryptic species group.
</paragraph>
</subSubSection>
<subSubSection id="C34F6549FFD28A17C78726F053F9F81F" pageId="13" pageNumber="34" type="reference_group">
<paragraph id="8BEA36C2FFD28A17C78726F053F9F81F" blockId="13.[151,1437,151,1999]" pageId="13" pageNumber="34">
<taxonomicName id="4C554D41FFD28A17C78726F056B7F929" box="[199,453,1760,1784]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="cancellatae">
<emphasis id="B921EAD0FFD28A17C78726F056B7F929" box="[199,453,1760,1784]" italics="true" pageId="13" pageNumber="34">Oxynaspis cancellatae</emphasis>
</taxonomicName>
(see fig.
<quantity id="4CAD9B27FFD28A17C57226F0552DF929" box="[562,607,1760,1784]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="13" pageNumber="34" unit="in" value="1.0">1 in</quantity>
<bibRefCitation id="EFC44B33FFD28A17C52726F15587F929" author="Totton" box="[615,757,1760,1784]" pageId="13" pageNumber="34" refString="Totton, A. K. (1940) New species of the cirripede genus Oxynaspis commensal with Antipatharia. The Annals and Magazine of Natural History, Series, 11, 6, 465 - 487." type="journal article" year="1940">Totton 1940</bibRefCitation>
;
<figureCitation id="136E2A47FFD28A17C44126F05420F929" box="[769,850,1760,1784]" captionStart="FIGURE 16" captionStartId="24.[151,244,1109,1130]" captionTargetBox="[156,1422,164,1070]" captionTargetId="figure@24.[95,1444,108,1088]" captionTargetPageId="24" captionText="FIGURE 16. Comparisons between Calantica arcuata n. sp. (I) and allied species (Group I sensu Newman &amp; Jones, 2011), and between Oxynaspis ryukyuensis n. sp. (N) and allied congeners. A, C. darwini Jones &amp; Hoise, 2009 (redrawn from Jones &amp; Hoise 2009); B, C. pusilla Utinomi, 1970 a (redrawn from Utinomi 1970 a); C, C. graphica Rosell, 1991 (redrawn from Rosell 1991); D, C. kruegeri Hiro, 1932 (redrawn from Hiro 1932); E, C. studeri (Weltner, 1922) (redrawn from Zevina 1978); F, C. trispinosa Hoek, 1883 (redrawn from Hoek 1883); G, C. eos (Pilsbry, 1907) (redrawn from Pilsbry 1907); H, C. affinis (Broch, 1922) (redrawn from Broch 1922); I, C. arcuata n. sp. (present study, RUMF-ZC- 1554); J, Oxynaspis alatae Totton, 1940 (redrawn from Totton 1940); K, Oxynaspis gracilis Totton, 1940 (redrawn from Totton 1940); L, O. celata Darwin, 1852 (redrawn from Darwin 1852); M. O. celata japonica Broch, 1922 (redrawn from Broch 1922); N, O. c. hirtae Totton, 1940 (redrawn from Totton, 1940); O, O. c. indica Annandale, 1909 (redrawn from Totton 1940); P, O. cancellatae Totton, 1940 (redrawn from Totton 1940); Q, O. ryukyuensis n. sp. (present study, (RUMF-ZC- 1558). Arrow in I indicates the diagnostic carina in C. arcuata n. sp., u indicates the location of the umbo of the carina in Oxynaspis species. The distal arm indicates the length of carinal margin above the umbo, basal arm indicate length of carinal margin below the umbo (see J). The length ratio of the distal arm to basal arm is a diagnostic character to identify Oxynaspis species." httpUri="https://zenodo.org/record/281668/files/figure.png" pageId="13" pageNumber="34">Fig. 16</figureCitation>
P) and
<taxonomicName id="4C554D41FFD28A17C4E526F0533FF929" box="[933,1101,1760,1784]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="ryukyuensis" status="sp. nov.">
<emphasis id="B921EAD0FFD28A17C4E526F0533FF929" box="[933,1101,1760,1784]" italics="true" pageId="13" pageNumber="34">O. ryukyuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21257ABFFD28A17C31526F053E4F929" box="[1109,1174,1760,1784]" pageId="13" pageNumber="34" rank="species">
<emphasis id="B921EAD0FFD28A17C31526F053E4F929" bold="true" box="[1109,1174,1760,1784]" pageId="13" pageNumber="34">n. sp.</emphasis>
</taxonomicNameLabel>
have the carinal umbo located close to the basal region (see
<figureCitation id="136E2A47FFD28A17C505271355E5F8CA" box="[581,663,1795,1819]" captionStart="FIGURE 16" captionStartId="24.[151,244,1109,1130]" captionTargetBox="[156,1422,164,1070]" captionTargetId="figure@24.[95,1444,108,1088]" captionTargetPageId="24" captionText="FIGURE 16. Comparisons between Calantica arcuata n. sp. (I) and allied species (Group I sensu Newman &amp; Jones, 2011), and between Oxynaspis ryukyuensis n. sp. (N) and allied congeners. A, C. darwini Jones &amp; Hoise, 2009 (redrawn from Jones &amp; Hoise 2009); B, C. pusilla Utinomi, 1970 a (redrawn from Utinomi 1970 a); C, C. graphica Rosell, 1991 (redrawn from Rosell 1991); D, C. kruegeri Hiro, 1932 (redrawn from Hiro 1932); E, C. studeri (Weltner, 1922) (redrawn from Zevina 1978); F, C. trispinosa Hoek, 1883 (redrawn from Hoek 1883); G, C. eos (Pilsbry, 1907) (redrawn from Pilsbry 1907); H, C. affinis (Broch, 1922) (redrawn from Broch 1922); I, C. arcuata n. sp. (present study, RUMF-ZC- 1554); J, Oxynaspis alatae Totton, 1940 (redrawn from Totton 1940); K, Oxynaspis gracilis Totton, 1940 (redrawn from Totton 1940); L, O. celata Darwin, 1852 (redrawn from Darwin 1852); M. O. celata japonica Broch, 1922 (redrawn from Broch 1922); N, O. c. hirtae Totton, 1940 (redrawn from Totton, 1940); O, O. c. indica Annandale, 1909 (redrawn from Totton 1940); P, O. cancellatae Totton, 1940 (redrawn from Totton 1940); Q, O. ryukyuensis n. sp. (present study, (RUMF-ZC- 1558). Arrow in I indicates the diagnostic carina in C. arcuata n. sp., u indicates the location of the umbo of the carina in Oxynaspis species. The distal arm indicates the length of carinal margin above the umbo, basal arm indicate length of carinal margin below the umbo (see J). The length ratio of the distal arm to basal arm is a diagnostic character to identify Oxynaspis species." httpUri="https://zenodo.org/record/281668/files/figure.png" pageId="13" pageNumber="34">Fig. 16</figureCitation>
Q). The carinal umbo of
<taxonomicName id="4C554D41FFD28A17C4F527145328F8CD" box="[949,1114,1796,1820]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="cancellatae">
<emphasis id="B921EAD0FFD28A17C4F527145328F8CD" box="[949,1114,1796,1820]" italics="true" pageId="13" pageNumber="34">O. cancellatae</emphasis>
</taxonomicName>
is located higher than in
<taxonomicName id="4C554D41FFD28A17C2C127145669F8EE" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="ryukyuensis" status="sp. nov.">
<emphasis id="B921EAD0FFD28A17C2C127145669F8EE" italics="true" pageId="13" pageNumber="34">O. ryukyuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21257ABFFD28A17C66327385611F891" box="[291,355,1832,1856]" pageId="13" pageNumber="34" rank="species">
<emphasis id="B921EAD0FFD28A17C66327385611F891" bold="true" box="[291,355,1832,1856]" pageId="13" pageNumber="34">n. sp.</emphasis>
</taxonomicNameLabel>
, resulting in the length ratio of the distal arm to the basal arm in
<taxonomicName id="4C554D41FFD28A17C30F27385386F8EE" box="[1103,1268,1831,1855]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="cancellatae">
<emphasis id="B921EAD0FFD28A17C30F27385386F8EE" box="[1103,1268,1831,1855]" italics="true" pageId="13" pageNumber="34">O. cancellatae</emphasis>
</taxonomicName>
as 5.7, shorter than that of
<taxonomicName id="4C554D41FFD28A17C665275C56BCF8B2" box="[293,462,1867,1891]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="ryukyuensis" status="sp. nov.">
<emphasis id="B921EAD0FFD28A17C665275C56BCF8B2" box="[293,462,1867,1891]" italics="true" pageId="13" pageNumber="34">O. ryukyuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21257ABFFD28A17C698275B556BF8B2" box="[472,537,1867,1891]" pageId="13" pageNumber="34" rank="species">
<emphasis id="B921EAD0FFD28A17C698275B556BF8B2" bold="true" box="[472,537,1867,1891]" pageId="13" pageNumber="34">n. sp.</emphasis>
</taxonomicNameLabel>
, which is 7.5. The base of cirrus I of
<taxonomicName id="4C554D41FFD28A17C499275C530DF8B2" box="[985,1151,1867,1891]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="cancellatae">
<emphasis id="B921EAD0FFD28A17C499275C530DF8B2" box="[985,1151,1867,1891]" italics="true" pageId="13" pageNumber="34">O. cancellatae</emphasis>
</taxonomicName>
has a single filamentary appendage (see fig. 13b in
<bibRefCitation id="EFC44B33FFD28A17C696277F552CF856" author="Broch" box="[470,606,1903,1927]" pageId="13" pageNumber="34" refString="Broch, H. (1931) Papers from Dr. Th. Mortensen's Pacific Expedition 1914 ~ 1916, LVI. Indomalayan Cirripedia. Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening i Kobenhavn, 91, 1 - 146." type="journal article" year="1931">Broch 1931</bibRefCitation>
) but a filamentary appendage is absent from the base of cirrus I of
<taxonomicName id="4C554D41FFD28A17C2C127605669F87A" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="ryukyuensis" status="sp. nov.">
<emphasis id="B921EAD0FFD28A17C2C127605669F87A" italics="true" pageId="13" pageNumber="34">O. ryukyuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21257ABFFD28A17C66327835610F87A" box="[291,354,1939,1963]" pageId="13" pageNumber="34" rank="species">
<emphasis id="B921EAD0FFD28A17C66327835610F87A" bold="true" box="[291,354,1939,1963]" pageId="13" pageNumber="34">n. sp.</emphasis>
</taxonomicNameLabel>
(
<figureCitation id="136E2A47FFD28A17C633278256C5F87B" box="[371,439,1938,1962]" captionStart="FIGURE 9" captionStartId="14.[151,244,1967,1988]" captionTargetBox="[172,1383,203,1948]" captionTargetId="figure@14.[135,1451,163,1974]" captionTargetPageId="14" captionText="FIGURE 9. Oxynaspis ryukyuensis n. sp. (holotype, RUMF-ZC- 1558). A, right view of capitulum; B, cirrus I; C, cirrus II; D, cirrus III; E, cirrus IV; F, cirrus V; G, cirrus VI. Scale bars in μm, except A in mm." httpUri="https://zenodo.org/record/281661/files/figure.png" pageId="13" pageNumber="34">Fig. 9</figureCitation>
B). The capitulum with completely calcified plates, the location of the carinal umbo and the absence of a filamentary appendage at the base of cirrus I define
<taxonomicName id="4C554D41FFD28A17C4D827A7534CF81F" box="[920,1086,1974,1998]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="ryukyuensis" status="sp. nov.">
<emphasis id="B921EAD0FFD28A17C4D827A7534CF81F" box="[920,1086,1974,1998]" italics="true" pageId="13" pageNumber="34">O. ryukyuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21257ABFFD28A17C30527A753F6F81E" box="[1093,1156,1975,1999]" pageId="13" pageNumber="34" rank="species">
<emphasis id="B921EAD0FFD28A17C30527A753F6F81E" bold="true" box="[1093,1156,1975,1999]" pageId="13" pageNumber="34">n. sp.</emphasis>
</taxonomicNameLabel>
.
</paragraph>
</subSubSection>
</treatment>
</document>