169 lines
18 KiB
XML
169 lines
18 KiB
XML
<document id="3994423A0481AFB455E66BDF9D66156C" ID-DOI="10.5281/zenodo.5352660" ID-ISSN="1942-1354" ID-Zenodo-Dep="5352660" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630208527114" checkinUser="felipe" docAuthor="Dunford, James C." docDate="2009" docId="03FC87C67B11FF8EFF6CFD95FDACDD6D" docLanguage="en" docName="0090.pdf" docOrigin="Insecta Mundi 2009 (90)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Speyeria hesperis subsp. lurana dos Passos and Grey 1945" docType="treatment" docVersion="4" lastPageNumber="34" masterDocId="FFC5FFBE7B32FFADFFACFFD0FFFED816" masterDocTitle="Taxonomic overview of the greater fritillary genus Speyeria Scudder and the atlantis - hesperis species complexes, with species accounts, type images, and relevant literature (Lepidoptera: Nymphalidae)" masterLastPageNumber="74" masterPageNumber="1" pageNumber="34" updateTime="1699186102529" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:titleInfo id="8953C3124C04CD7E5E8673F66425610C">
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<mods:title id="A66EBB8BC3A9938F058C8791599E4698">Taxonomic overview of the greater fritillary genus Speyeria Scudder and the atlantis - hesperis species complexes, with species accounts, type images, and relevant literature (Lepidoptera: Nymphalidae)</mods:title>
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<mods:name id="B492CFD3CBFACDC8CAF153BCF4D1C504" type="personal">
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<mods:roleTerm id="9CEDC030BD8B00F6FF459366C9E6AF03">Author</mods:roleTerm>
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<mods:namePart id="508AEEC63A0045206EE86354554A1EF9">Dunford, James C.</mods:namePart>
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<mods:typeOfResource id="BABCA34084AB7044B3614755D22DBC88">text</mods:typeOfResource>
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<mods:title id="CAB4E67C94A70107939E130B197759A2">Insecta Mundi</mods:title>
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<mods:part id="E1DA0E60CC4ACA6B1B64BDD6D76E2D05">
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<mods:date id="EC9F56FDD74F987782CDCCB192F4A5ED">2009</mods:date>
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<mods:number id="60F0D2611CD6472C8C76242E205259D9">2009-09-26</mods:number>
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<mods:detail id="DA99C367C318524F92162A31315BADE1" type="volume">
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<mods:number id="F375C2BCB7733FF735DFC6BCDBD5DE50">2009</mods:number>
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<mods:detail id="117EA258D91D27EEF5D68BFAFE8B961D" type="issue">
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<mods:number id="7548159502E19863CF03176F5CE7FF73">90</mods:number>
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<mods:start id="C4C105455725C62FB02F414A70D9D163">1</mods:start>
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<mods:end id="02A17A7805335A7BC0032F1D2F9A9B5C">74</mods:end>
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<mods:identifier id="D405D045AF44BA2C02453473A7906358" type="DOI">10.5281/zenodo.5352660</mods:identifier>
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<mods:identifier id="843A8621412BBF224829BF3B34105D1B" type="ISSN">1942-1354</mods:identifier>
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<treatment id="03FC87C67B11FF8EFF6CFD95FDACDD6D" ID-DOI="http://doi.org/10.5281/zenodo.5450577" ID-Zenodo-Dep="5450577" LSID="urn:lsid:plazi:treatment:03FC87C67B11FF8EFF6CFD95FDACDD6D" httpUri="http://treatment.plazi.org/id/03FC87C67B11FF8EFF6CFD95FDACDD6D" lastPageNumber="34" pageId="35" pageNumber="34">
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<subSubSection id="C34F655B7B11FF8EFF6CFD95FC82DA4D" box="[192,892,579,604]" pageId="35" pageNumber="34" type="nomenclature">
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<paragraph id="8BEA36D07B11FF8EFF6CFD95FC82DA4D" blockId="35.[192,892,579,635]" box="[192,892,579,604]" pageId="35" pageNumber="34">
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<heading id="D0A281BC7B11FF8EFF6CFD95FC82DA4D" bold="true" box="[192,892,579,604]" fontSize="10" level="2" pageId="35" pageNumber="34" reason="3">
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<taxonomicName id="4C554D537B11FF8EFF6CFD95FC82DA4D" authority="dos Passos and Grey, 1945" authorityName="dos Passos and Grey" authorityYear="1945" box="[192,892,579,604]" class="Insecta" family="Nymphalidae" genus="Speyeria" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="34" phylum="Arthropoda" rank="subSpecies" species="hesperis" subSpecies="lurana">
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<emphasis id="B921EAC27B11FF8EFF6CFD95FC82DA4D" bold="true" box="[192,892,579,604]" pageId="35" pageNumber="34">
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<emphasis id="B921EAC27B11FF8EFF6CFD95FDEADA4A" bold="true" box="[192,532,581,604]" italics="true" pageId="35" pageNumber="34">Speyeria hesperis lurana</emphasis>
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dos Passos and Grey, 1945
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34F655B7B11FF8EFF6CFDB3FEBADA6D" box="[192,324,611,635]" pageId="35" pageNumber="34" type="description">
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<paragraph id="8BEA36D07B11FF8EFF6CFDB3FEBADA6D" blockId="35.[192,892,579,635]" box="[192,324,611,635]" pageId="35" pageNumber="34">
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(
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<figureCitation id="136E2A557B11FF8EFF64FDB3FEC2DA6D" box="[200,316,611,635]" captionStart="Figure 31-35" captionStartId="71.[192,276,1642,1664]" captionTargetBox="[192,1443,192,1614]" captionTargetId="figure-15@71.[192,1444,192,1614]" captionTargetPageId="71" captionText="Figure 31-35. Type images for Speyeria spp.; dorsal, ventral, and label images. 31) Speyeria hesperis greyi holotype, described as Speyeria atlantis greyi, male, AMNH. 32) Speyeria hesperis lurana holotype, described as Speyeria atlantis lurana, male, AMNH. 33) Speyeria hesperis irene lectotype, described as Argynnis irene, female, NMNH. 34) Speyeria hesperis hanseni holotype, described as Speyeria atlantis hanseni, male, LACM. 35) Speyeria hesperis dodgei holotype, described as Argynnis dodgei, male, AMNH. Images 31-32 and 35 by James C. Dunford; 33 by Robert Robbins; 34 by Weiping Xie." figureDoi="http://doi.org/10.5281/zenodo.5352682" httpUri="https://zenodo.org/record/5352682/files/figure.png" pageId="35" pageNumber="34">Figure 32</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C34F655B7B11FF8EFF6CFD74FC2EDACD" pageId="35" pageNumber="34" type="reference_group">
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<paragraph id="8BEA36D07B11FF8EFF6CFD74FC9ADAAD" blockId="35.[192,976,675,731]" box="[192,868,675,699]" pageId="35" pageNumber="34">
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<treatmentCitationGroup id="AB4511FE7B11FF8EFF6CFD74FC9ADAAD" box="[192,868,675,699]" pageId="35" pageNumber="34">
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<treatmentCitation id="0AF410C17B11FF8EFF6CFD74FCA1DAAD" author="dos Passos C. F. & L. P. Grey" box="[192,863,675,699]" page="8" pageId="35" pageNumber="34" year="1945">
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<taxonomicName id="4C554D537B11FF8EFF6CFD74FCA1DAAD" ID-CoL="5L8Q8" authority="dos Passos and Grey, 1945 b: 8" authorityName="dos Passos and Grey" authorityPageNumber="8" authorityYear="1945" box="[192,863,675,699]" class="Insecta" family="Nymphalidae" genus="Speyeria" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="34" phylum="Arthropoda" rank="subSpecies" species="atlantis" subSpecies="lurana">
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<emphasis id="B921EAC27B11FF8EFF6CFD74FE14DAAD" box="[192,490,675,699]" italics="true" pageId="35" pageNumber="34">Speyeria atlantis lurana</emphasis>
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<bibRefCitation id="EFC44B217B11FF8EFE5CFD73FCA1DAAD" author="dos Passos C. F. & L. P. Grey" box="[496,863,675,699]" pageId="35" pageNumber="34" pagination="1 - 17" refId="ref35862" refString="dos Passos C. F., and L. P. Grey. 1945 b. A new species and some new subspecies of Speyeria (Lepidoptera, Nymphalidae). American Museum Novitates 1297: 1 - 17." type="journal article" year="1945">dos Passos and Grey, 1945b: 8</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BEA36D07B11FF8EFF6CFD14FC2EDACD" blockId="35.[192,976,675,731]" box="[192,976,707,731]" pageId="35" pageNumber="34">
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<taxonomicName id="4C554D537B11FF8EFF6CFD14FD11DACD" authority="dos Passos and Grey" authorityName="dos Passos and Grey" authorityYear="1945" box="[192,751,707,731]" class="Insecta" family="Nymphalidae" genus="Speyeria" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="34" phylum="Arthropoda" rank="subSpecies" species="hesperis" subSpecies="lurana">
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<emphasis id="B921EAC27B11FF8EFF6CFD14FE11DACD" box="[192,495,707,731]" italics="true" pageId="35" pageNumber="34">Speyeria hesperis lurana</emphasis>
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dos Passos and Grey
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</taxonomicName>
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[
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<bibRefCitation id="EFC44B217B11FF8EFD57FD13FC3BDACD" author="Scott, J. A. & N. G. Kondla & S. M. Spomer" box="[763,965,707,731]" pageId="35" pageNumber="34" pagination="1 - 31" refId="ref45136" refString="Scott, J. A., N. G. Kondla, and S. M. Spomer. 1998. Speyeria hesperis and Speyeria atlantis are distinct species. Papilio 8: 1 - 31." type="journal article" year="1998">Scott et al. 1998</bibRefCitation>
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].
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</paragraph>
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</subSubSection>
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<subSubSection id="C34F655B7B11FF8EFF6CFCD3FD8EDB0D" box="[192,624,771,795]" pageId="35" pageNumber="34" type="description">
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<paragraph id="8BEA36D07B11FF8EFF6CFCD3FD8EDB0D" blockId="35.[192,624,771,795]" box="[192,624,771,795]" pageId="35" pageNumber="34">
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<emphasis id="B921EAC27B11FF8EFF6CFCD3FE68DB0D" bold="true" box="[192,406,771,795]" pageId="35" pageNumber="34">Common name.</emphasis>
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Lurana fritillary.
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</paragraph>
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</subSubSection>
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<subSubSection id="C34F655B7B11FF8EFF6CFC93FDACDD6D" pageId="35" pageNumber="34" type="materials_examined">
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||
<paragraph id="8BEA36D07B11FF8EFF6CFC93FB6ADB4D" blockId="35.[192,1172,835,859]" box="[192,1172,835,859]" pageId="35" pageNumber="34">
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<emphasis id="B921EAC27B11FF8EFF6CFC93FE67DB4D" bold="true" box="[192,409,835,859]" pageId="35" pageNumber="34">Type deposited.</emphasis>
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<materialsCitation id="3B3D3C8D7B11FF8EFE32FC93FB6ADB4D" box="[414,1172,835,859]" location="American Museum of Natural History" pageId="35" pageNumber="34" specimenCount="1" typeStatus="holotype">
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<typeStatus id="54EE88727B11FF8EFE32FC93FDF3DB4D" box="[414,525,835,859]" pageId="35" pageNumber="34" type="holotype">Holotype</typeStatus>
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at
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<location id="8E8A600B7B11FF8EFD9DFC93FBFEDB4D" LSID="urn:lsid:plazi:treatment:03FC87C67B11FF8EFF6CFD95FDACDD6D:8E8A600B7B11FF8EFD9DFC93FBFEDB4D" box="[561,1024,835,859]" name="American Museum of Natural History" pageId="35" pageNumber="34">American Museum of Natural History</location>
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||
(
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<figureCitation id="136E2A557B11FF8EFBA2FC93FB79DB4D" box="[1038,1159,835,859]" captionStart="Figure 31-35" captionStartId="71.[192,276,1642,1664]" captionTargetBox="[192,1443,192,1614]" captionTargetId="figure-15@71.[192,1444,192,1614]" captionTargetPageId="71" captionText="Figure 31-35. Type images for Speyeria spp.; dorsal, ventral, and label images. 31) Speyeria hesperis greyi holotype, described as Speyeria atlantis greyi, male, AMNH. 32) Speyeria hesperis lurana holotype, described as Speyeria atlantis lurana, male, AMNH. 33) Speyeria hesperis irene lectotype, described as Argynnis irene, female, NMNH. 34) Speyeria hesperis hanseni holotype, described as Speyeria atlantis hanseni, male, LACM. 35) Speyeria hesperis dodgei holotype, described as Argynnis dodgei, male, AMNH. Images 31-32 and 35 by James C. Dunford; 33 by Robert Robbins; 34 by Weiping Xie." figureDoi="http://doi.org/10.5281/zenodo.5352682" httpUri="https://zenodo.org/record/5352682/files/figure.png" pageId="35" pageNumber="34">Figure 32</figureCitation>
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).
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</materialsCitation>
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</paragraph>
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<paragraph id="8BEA36D07B11FF8EFF6CFC53FC84DB8D" blockId="35.[192,890,899,924]" box="[192,890,899,924]" pageId="35" pageNumber="34">
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||
<emphasis id="B921EAC27B11FF8EFF6CFC53FE85DB8D" bold="true" box="[192,379,899,923]" pageId="35" pageNumber="34">
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<typeStatus id="54EE88727B11FF8EFF6CFC53FEFBDB8D" box="[192,261,899,923]" pageId="35" pageNumber="34">Type</typeStatus>
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locality.
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</emphasis>
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Harney Peak, Black Hills,
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<collectingRegion id="4991F8327B11FF8EFD64FC53FC89DB8D" box="[712,887,899,923]" country="United States of America" name="South Dakota" pageId="35" pageNumber="34">South Dakota</collectingRegion>
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.
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</paragraph>
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<paragraph id="8BEA36D07B11FF8EFF6CFC13FBFADBED" blockId="35.[192,1440,963,1020]" pageId="35" pageNumber="34">
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<emphasis id="B921EAC27B11FF8EFF6CFC13FE51DBCD" bold="true" box="[192,431,963,987]" pageId="35" pageNumber="34">Type label data. “</emphasis>
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<materialsCitation id="3B3D3C8D7B11FF8EFE03FC13FBFEDBED" collectionCode="HARNEY, PEAK" collectorName="Cyril F. dos Passos & L. Paul Grey & Col. By A. C. & L. P. Grey" pageId="35" pageNumber="34" specimenCount="1" typeStatus="holotype">
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<typeStatus id="54EE88727B11FF8EFE03FC13FDE1DBCD" box="[431,543,963,987]" pageId="35" pageNumber="34" type="holotype">Holotype</typeStatus>
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<taxonomicName id="4C554D537B11FF8EFD89FC13FCADDBCD" box="[549,851,963,987]" class="Insecta" family="Nymphalidae" genus="Speyeria" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="34" phylum="Arthropoda" rank="subSpecies" species="atlantis" subSpecies="lurana">Speyeria atlantis lurana</taxonomicName>
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<collectorName id="26A053067B11FF8EFCF5FC13FBBCDBCD" box="[857,1090,963,987]" pageId="35" pageNumber="34">Cyril F. dos Passos</collectorName>
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and
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<collectorName id="26A053067B11FF8EFBD1FC14FAE2DBCD" box="[1149,1308,963,987]" pageId="35" pageNumber="34">L. Paul Grey</collectorName>
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;
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<collectionCode id="ED44AE157B11FF8EFA8BFC13FEEFDBED" pageId="35" pageNumber="34">HARNEY PEAK</collectionCode>
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, S. D. [male] 25 VI-39
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<collectorName id="26A053067B11FF8EFD8CFC33FD45DBED" box="[544,699,995,1020]" pageId="35" pageNumber="34">Col. By A. C.</collectorName>
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FREDERICK;
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<collectorName id="26A053067B11FF8EFCDAFC34FC0BDBED" box="[886,1013,995,1019]" pageId="35" pageNumber="34">L. P. Grey</collectorName>
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”
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="8BEA36D07B11FF8EFF6CFBF3FD08DCCD" blockId="35.[192,1441,1059,1244]" pageId="35" pageNumber="34">
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<emphasis id="B921EAC27B11FF8EFF6CFBF3FD16DC2D" bold="true" box="[192,744,1059,1083]" pageId="35" pageNumber="34">Identification, taxonomy, and variation.</emphasis>
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Both sexes are typically unsilvered, sympatrically occurring with silvered forms being those of
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<taxonomicName id="4C554D537B11FF8EFD3EFB94FB96DC4D" authority="(Scott et al. 1998)" baseAuthorityName="Scott" baseAuthorityYear="1998" box="[658,1128,1091,1115]" class="Insecta" family="Nymphalidae" genus="Speyeria" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="34" phylum="Arthropoda" rank="subSpecies" species="atlantis" subSpecies="pahasapa">
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<emphasis id="B921EAC27B11FF8EFD3EFB94FC73DC4D" box="[658,909,1091,1115]" italics="true" pageId="35" pageNumber="34">S. atlantis pahasapa</emphasis>
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||
(
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<bibRefCitation id="EFC44B217B11FF8EFC36FB93FBA1DC4D" author="Scott, J. A. & N. G. Kondla & S. M. Spomer" box="[922,1119,1091,1115]" pageId="35" pageNumber="34" pagination="1 - 31" refId="ref45136" refString="Scott, J. A., N. G. Kondla, and S. M. Spomer. 1998. Speyeria hesperis and Speyeria atlantis are distinct species. Papilio 8: 1 - 31." type="journal article" year="1998">Scott et al. 1998</bibRefCitation>
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||
)
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</taxonomicName>
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. Cliff Ferris (pers. comm.) notes that both ‘
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<emphasis id="B921EAC27B11FF8EFE2EFBB4FDECDC6D" box="[386,530,1123,1147]" italics="true" pageId="35" pageNumber="34">S. h. lurana</emphasis>
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’ and ‘
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<emphasis id="B921EAC27B11FF8EFDF9FBB4FCF6DC6D" box="[597,776,1123,1147]" italics="true" pageId="35" pageNumber="34">S. a. pahasapa</emphasis>
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’ fly together and interbreed (or at least copulate) in the Big Horn Mountains, Washakie Co.,
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<collectingRegion id="4991F8327B11FF8EFDD4FB53FD0EDC8D" box="[632,752,1155,1179]" country="United States of America" name="Wyoming" pageId="35" pageNumber="34">Wyoming</collectingRegion>
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. The status or distinctness of these two subspecies should be further evaluated.
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<bibRefCitation id="EFC44B217B11FF8EFE62FB73FD8FDCAD" author="Scott, J. A." box="[462,625,1187,1211]" pageId="35" pageNumber="34" pagination="1 - 129" refId="ref45004" refString="Scott, J. A. 2008 a. Corrections / reviews of 58 North American butterfly books. Papilio 19: 1 - 129." type="journal article" year="2008">Scott (2008a)</bibRefCitation>
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suggests that these Big Horn populations best be called ‘
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<emphasis id="B921EAC27B11FF8EFA9AFB73FA75DCAD" box="[1334,1419,1187,1211]" italics="true" pageId="35" pageNumber="34">lurana</emphasis>
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x
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<emphasis id="B921EAC27B11FF8EFF6CFB13FEE5DCCD" box="[192,283,1219,1243]" italics="true" pageId="35" pageNumber="34">dennisi</emphasis>
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’. Adult wingspan is on average
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<quantity id="4CAD9B357B11FF8EFD30FB13FD0CDCCD" box="[668,754,1219,1243]" metricMagnitude="-2" metricUnit="m" metricValue="5.5" pageId="35" pageNumber="34" unit="mm" value="55.0">55 mm</quantity>
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||
.
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</paragraph>
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<paragraph id="8BEA36D07B11FF8EFF6CFAD3FB70DD0D" blockId="35.[192,1166,1283,1307]" box="[192,1166,1283,1307]" pageId="35" pageNumber="34">
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||
<emphasis id="B921EAC27B11FF8EFF6CFAD3FEDEDD0D" bold="true" box="[192,288,1283,1307]" pageId="35" pageNumber="34">Range.</emphasis>
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Black Hills,
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<collectingRegion id="4991F8327B11FF8EFE16FAD3FD99DD0D" box="[442,615,1283,1307]" country="United States of America" name="South Dakota" pageId="35" pageNumber="34">South Dakota</collectingRegion>
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. Also recorded in
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<collectingRegion id="4991F8327B11FF8EFC97FAD3FC4EDD0D" box="[827,944,1283,1307]" country="United States of America" name="Wyoming" pageId="35" pageNumber="34">Wyoming</collectingRegion>
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||
(
|
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<bibRefCitation id="EFC44B217B11FF8EFC11FAD3FB7FDD0D" author="Grey, L. P. & A. H. Moeck & W. H. Evans" box="[957,1153,1283,1307]" pageId="35" pageNumber="34" pagination="129 - 147" refId="ref39693" refString="Grey, L. P., A. H. Moeck, and W. H. Evans. 1963. Notes on overlapping subspecies. II. Segregation in the Speyeria atlantis of the Black Hills (Nymphalidae). Journal of the Lepidopterists' Society 17: 129 - 147." type="journal article" year="1963">Grey et al. 1963</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BEA36D07B11FF8EFF6CFA93FDACDD6D" blockId="35.[192,1440,1347,1403]" pageId="35" pageNumber="34">
|
||
<emphasis id="B921EAC27B11FF8EFF6CFA93FE9BDD4D" bold="true" box="[192,357,1347,1371]" pageId="35" pageNumber="34">Life history.</emphasis>
|
||
I observed numerous individuals using creeks as flyways and feeding on various flowers in Spearfish Canyon,
|
||
<collectingRegion id="4991F8327B11FF8EFE08FAB3FDB1DD6D" box="[420,591,1379,1403]" country="United States of America" name="South Dakota" pageId="35" pageNumber="34">South Dakota</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |