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<document id="F68007D7CE05DEB3AE023B6DF5D68956" ID-DOI="10.5281/zenodo.172764" ID-GBIF-Dataset="57f77567-9f74-4e9f-a79a-a19a41e960c8" ID-ISSN="1175­5326" ID-Zenodo-Dep="172764" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459761781846" checkinUser="plazi" docAuthor="Blahnik, Roger J. &amp; Holzenthal, Ralph W." docDate="2006" docId="03FC8793FFC86F71846AFDF3C2061EB5" docLanguage="en" docName="zt01233p052.pdf" docOrigin="Zootaxa 1233" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Culoptila hamata Blahnik &amp; Holzenthal, 2006, new species" docType="treatment" docVersion="10" lastPageNumber="27" masterDocId="FFC5FFEBFFD16F6B8562FFAFC6051D3B" masterDocTitle="Revision of the genus Culoptila (Trichoptera: Glossosomatidae)" masterLastPageNumber="52" masterPageNumber="1" pageNumber="26" updateTime="1698215326134" updateUser="plazi">
<mods:mods id="54ABBA444AF2D8206B4308590B11C239" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E0675A603FC0692D21305951C0438E45">
<mods:title id="6E1ED1F5CCBB588EDE06B9B39A59C76A">Revision of the genus Culoptila (Trichoptera: Glossosomatidae)</mods:title>
</mods:titleInfo>
<mods:name id="9E12D42702DE88BBC4574FAED77BA55E" type="personal">
<mods:role id="B3802B125BC2B4A8AF10AF7C5FD20EF5">
<mods:roleTerm id="01A6D092BCBAF14F76F6ACF47A213FFB">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FB248BB93441C4F7D5C6AC40FC038086">Blahnik, Roger J.</mods:namePart>
</mods:name>
<mods:name id="29A8ACD69C251F4B709096981CCA8910" type="personal">
<mods:role id="74F12A136E0FFFAF87E48436295494CD">
<mods:roleTerm id="2992CF066367216A29E26103165F120A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="20FD69B4ECAB979C87614D9356215693">Holzenthal, Ralph W.</mods:namePart>
</mods:name>
<mods:typeOfResource id="45C1DFF527ABDD003AA542876531BC59">text</mods:typeOfResource>
<mods:relatedItem id="7007E02FBF507DFFE35E885F3A0D10F2" type="host">
<mods:titleInfo id="3FB270CF0AE32DA278EEEA5421C25096">
<mods:title id="2ABAFA00D5EBC844192C26CE9B833BB4">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="6F95858075208137E71E0478743E7455">
<mods:date id="FCD0D8A6300F3AA5C2679F20BA6C76CF">2006</mods:date>
<mods:detail id="A833BCD8925ED1683D63F3D821B2EF92" type="volume">
<mods:number id="40FD4317EADBF8AA3D2470001519A609">1233</mods:number>
</mods:detail>
<mods:extent id="222A0F7319BF1B731BD34B3A05E772CF" unit="page">
<mods:start id="EABAC163D2D09C91FB2FC06B7C617DC5">1</mods:start>
<mods:end id="EB7C5E6EA346B7663216CE490F044EC5">52</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="AAD63CA95EA5369A65670419ABF811EA">journal article</mods:classification>
<mods:identifier id="98724AD610F52FBAE9A503A08FD3CEC8" type="DOI">10.5281/zenodo.172764</mods:identifier>
<mods:identifier id="ABEBBC4B37FFF5378022CD6ADBEC37F5" type="GBIF-Dataset">57f77567-9f74-4e9f-a79a-a19a41e960c8</mods:identifier>
<mods:identifier id="32D00EDCD0717A595FA1A0DEFDC626C7" type="ISSN">1175­5326</mods:identifier>
<mods:identifier id="83991967B72E771ABDB035D80069CD2D" type="Zenodo-Dep">172764</mods:identifier>
</mods:mods>
<treatment id="03FC8793FFC86F71846AFDF3C2061EB5" ID-DOI="http://doi.org/10.5281/zenodo.6256643" ID-GBIF-Taxon="119345079" ID-Zenodo-Dep="6256643" LSID="urn:lsid:plazi:treatment:03FC8793FFC86F71846AFDF3C2061EB5" httpUri="http://treatment.plazi.org/id/03FC8793FFC86F71846AFDF3C2061EB5" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26">
<subSubSection id="C34F650EFFC86F72846AFDF3C4711FA2" pageId="25" pageNumber="26" type="nomenclature">
<paragraph id="8BEA3685FFC86F72846AFDF3C47C1F4E" blockId="25.[264,633,603,665]" box="[264,633,603,630]" pageId="25" pageNumber="26">
<heading id="D0A281E9FFC86F72846AFDF3C47C1F4E" bold="true" box="[264,633,603,630]" fontSize="11" level="1" pageId="25" pageNumber="26" reason="1">
<emphasis id="B921EA97FFC86F72846AFDF3C47C1F4E" bold="true" box="[264,633,603,630]" pageId="25" pageNumber="26">
<taxonomicName id="4C554D06FFC86F72846AFDF3C7DF1F4D" box="[264,474,604,630]" class="Insecta" family="Glossosomatidae" genus="Culoptila" kingdom="Animalia" order="Trichoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="hamata" status="sp. nov.">
<emphasis id="B921EA97FFC86F72846AFDF3C7DF1F4D" bold="true" box="[264,474,604,630]" italics="true" pageId="25" pageNumber="26">Culoptila hamata</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A21257ECFFC86F72848BFDF4C47C1F4E" box="[489,633,603,629]" pageId="25" pageNumber="26" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BEA3685FFC86F72846AFDD0C4711FA2" blockId="25.[264,633,603,665]" box="[264,628,639,665]" pageId="25" pageNumber="26">
<figureCitation id="136E2A00FFC86F72846AFDD0C75D1FA2" box="[264,344,639,665]" captionStart="FIGURE 9" captionStartId="9.[264,368,1361,1385]" captionTargetBox="[279,1305,796,1311]" captionTargetId="figure@9.[279,1307,794,1320]" captionTargetPageId="9" captionText="FIGURE 9. Culoptila hamata, new species, wings. 9 A—forewing; 9 B—hind wing." httpUri="https://zenodo.org/record/172768/files/figure.png" pageId="25" pageNumber="26">Figs. 9</figureCitation>
A, B, 4, 22A, B, 23A, B
</paragraph>
</subSubSection>
<subSubSection id="C34F650EFFC86F72846AFD60C5EB1882" pageId="25" pageNumber="26" type="description">
<paragraph id="8BEA3685FFC86F72846AFD60C2571E8A" blockId="25.[264,1324,719,1465]" pageId="25" pageNumber="26">
This species is probably most closely related to
<taxonomicName id="4C554D06FFC86F728626FD60C20A1FD3" box="[836,1039,719,744]" class="Insecta" family="Glossosomatidae" genus="Culoptila" kingdom="Animalia" order="Trichoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="tapanti" status="sp. nov.">
<emphasis id="B921EA97FFC86F728626FD60C20A1FD3" box="[836,1039,719,744]" italics="true" pageId="25" pageNumber="26">Culoptila tapanti</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A21257ECFFC86F72817DFD60C25A1FD2" box="[1055,1119,719,745]" pageId="25" pageNumber="26" rank="species">n. sp.</taxonomicNameLabel>
Both species are similar in having an elongate curved tergum X, a phallobase that is relatively parallelsided, with an elongate, acute posterodorsal apex, and also in the possession of a single phallic spine, which is somewhat inflated mesally and tapers to a very fine needle­like apex.
<taxonomicName id="4C554D06FFC86F72842FFCC0C41E1EB3" box="[333,539,879,904]" class="Insecta" family="Glossosomatidae" genus="Culoptila" kingdom="Animalia" order="Trichoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="hamata">
<emphasis id="B921EA97FFC86F72842FFCC0C41E1EB3" box="[333,539,879,904]" italics="true" pageId="25" pageNumber="26">Culoptila hamata</emphasis>
</taxonomicName>
differs most notably C.
<taxonomicName id="4C554D06FFC86F728656FCC0C5821EB3" box="[820,903,879,904]" class="Insecta" family="Glossosomatidae" genus="Culoptila" kingdom="Animalia" order="Trichoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="tapanti">
<emphasis id="B921EA97FFC86F728656FCC0C5821EB3" box="[820,903,879,904]" italics="true" pageId="25" pageNumber="26">tapanti</emphasis>
</taxonomicName>
in the form of the apex of tergum X (narrowed and acute in
<taxonomicName id="4C554D06FFC86F72877FFC38C4921E8B" box="[541,663,919,944]" class="Insecta" family="Glossosomatidae" genus="Culoptila" kingdom="Animalia" order="Trichoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="hamata">
<emphasis id="B921EA97FFC86F72877FFC38C4921E8B" box="[541,663,919,944]" italics="true" pageId="25" pageNumber="26">C. hamata</emphasis>
</taxonomicName>
and deeply emarginate in
<taxonomicName id="4C554D06FFC86F7286AEFC38C2441E8B" box="[972,1089,919,944]" class="Insecta" family="Glossosomatidae" genus="Culoptila" kingdom="Animalia" order="Trichoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="tapanti">
<emphasis id="B921EA97FFC86F7286AEFC38C2441E8B" box="[972,1089,919,944]" italics="true" pageId="25" pageNumber="26">C. tapanti</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8BEA3685FFC86F72845AFC10C7C51942" blockId="25.[264,1324,719,1465]" pageId="25" pageNumber="26">
Adult. Length of forewing: male
<quantity id="4CAD9B60FFC86F7287B8FC10C5741EE2" box="[730,881,959,985]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" metricValueMax="3.2" metricValueMin="2.2" pageId="25" pageNumber="26" unit="mm" value="2.7" valueMax="3.2" valueMin="2.2">2.2–3.2 mm</quantity>
; female
<quantity id="4CAD9B60FFC86F728682FC10C2721EE2" box="[992,1143,959,985]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" metricValueMax="3.2" metricValueMin="2.4" pageId="25" pageNumber="26" unit="mm" value="2.8" valueMax="3.2" valueMin="2.4">2.4–3.2 mm</quantity>
. Color brown; forewing with small whitish mark at arculus on wing margin. Mesothorax of male slightly narrowed anteromesally; mesoscutal wart short, heart­shaped. Mesothoracic tegulae of male rounded, distinctly enlarged; tegular setae short; tegular gland short, pleated, concertina­like.
</paragraph>
<paragraph id="8BEA3685FFC86F72845AFB28C5EB1882" blockId="25.[264,1324,719,1465]" pageId="25" pageNumber="26">Male genitalia. Sternum VI process short, subtruncate. Tergum IX ventral margin subtruncately rounded, slightly produced posteroventrally. Inferior appendages moderately long, subacute apically. Tergum X very elongate, narrow, strongly curved, apex in ventral view acute; ventrolateral processes with apices incurved and posteriorly bent, bluntly rounded and distinctly sclerotized. Phallobase elongate, dorsal and ventral margins subparallel in lateral view, apicodorsal projection very elongate, straight, apex acute. Phallic apparatus with 1 phallic spine, narrow and needle­like, about 1/2 length of phallobase, with rather abrupt, bulbous expansion in basal half.</paragraph>
</subSubSection>
<subSubSection id="C34F650EFFC86F71846AFA40C32E1FD5" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26" type="materials_examined">
<paragraph id="8BEA3685FFC86F72846AFA40C7E31B33" blockId="25.[264,1323,1519,1905]" box="[264,486,1519,1544]" pageId="25" pageNumber="26">
<emphasis id="B921EA97FFC86F72846AFA40C7E31B33" box="[264,486,1519,1544]" italics="true" pageId="25" pageNumber="26">Material examined</emphasis>
</paragraph>
<paragraph id="8BEA3685FFC86F72845AF9B9C4641BBA" blockId="25.[264,1323,1519,1905]" pageId="25" pageNumber="26">
<emphasis id="B921EA97FFC86F72845AF9B9C5431B0B" bold="true" box="[312,838,1558,1584]" pageId="25" pageNumber="26">
<typeStatus id="54EE8827FFC86F72845AF9B9C7AF1B0B" box="[312,426,1558,1584]" pageId="25" pageNumber="26" type="holotype">Holotype</typeStatus>
male:
<collectingCountry id="F3427615FFC86F728767F9B9C4C71B0B" box="[517,706,1558,1584]" name="Costa Rica" pageId="25" pageNumber="26">COSTA RICA</collectingCountry>
: Alajuela:
</emphasis>
Río Toro, 3.0 km (road) SW Bajos del Toro,
<geoCoordinate id="EE615042FFC86F72842FF990C7DB1B62" box="[333,478,1599,1625]" direction="north" orientation="latitude" pageId="25" pageNumber="26" precision="15" value="10.203889">10°12’14&quot;N</geoCoordinate>
,
<geoCoordinate id="EE615042FFC86F728488F990C48B1B62" box="[490,654,1599,1625]" direction="west" orientation="longitude" pageId="25" pageNumber="26" precision="15" value="-84.31612">084°18’58&quot;W</geoCoordinate>
,
<quantity id="4CAD9B60FFC86F7287F8F990C4F31B62" box="[666,758,1599,1625]" metricMagnitude="3" metricUnit="m" metricValue="1.53" pageId="25" pageNumber="26" unit="m" value="1530.0">1530 m</quantity>
,
<date id="FFEB1045FFC86F728663F990C58B1B62" box="[769,910,1599,1625]" pageId="25" pageNumber="26" value="1990-09-03" valueMax="1990-09-04" valueMin="1990-09-03">3–4.ix.1990</date>
(Holzenthal, Blahnik, &amp; Huisman) (
<collectionCode id="ED44AE40FFC86F728472F9C8C7641BBA" box="[272,353,1639,1665]" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="25" pageNumber="26">UMSP</collectionCode>
000000386) (
<collectionCode id="ED44AE40FFC86F72849DF9C8C4501BBA" box="[511,597,1639,1665]" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="25" pageNumber="26">UMSP</collectionCode>
).
</paragraph>
<paragraph id="8BEA3685FFC86F71845AF921C32E1FD5" blockId="25.[264,1323,1519,1905]" lastBlockId="26.[264,1323,284,750]" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26">
<emphasis id="B921EA97FFC86F72845AF921C5051B93" bold="true" box="[312,768,1678,1704]" pageId="25" pageNumber="26">
<typeStatus id="54EE8827FFC86F72845AF921C7BF1B93" box="[312,442,1678,1704]" pageId="25" pageNumber="26" type="paratype">Paratypes</typeStatus>
:
<collectingCountry id="F3427615FFC86F7284A4F921C4851B93" box="[454,640,1678,1704]" name="Costa Rica" pageId="25" pageNumber="26">COSTA RICA</collectingCountry>
: Alajuela:
</emphasis>
same data as
<typeStatus id="54EE8827FFC86F7286C1F920C20F1B92" box="[931,1034,1679,1705]" pageId="25" pageNumber="26" type="holotype">holotype</typeStatus>
—
<specimenCount id="9D53FD0CFFC86F728154F920C2801B92" box="[1078,1157,1679,1705]" pageId="25" pageNumber="26" type="male">1 male</specimenCount>
(
<collectionCode id="ED44AE40FFC86F7281F7F920C2EE1B92" box="[1173,1259,1679,1705]" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="25" pageNumber="26">UMSP</collectionCode>
); Río Sarapiquí, ca.
<quantity id="4CAD9B60FFC86F7284A0F918C4031BEA" box="[450,518,1719,1745]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="25" pageNumber="26" unit="km" value="2.0">2 km</quantity>
SE Cariblanco,
<geoCoordinate id="EE615042FFC86F728780F918C5761BEA" box="[738,883,1719,1745]" direction="north" orientation="latitude" pageId="25" pageNumber="26" precision="15" value="10.298889">10°17’56&quot;N</geoCoordinate>
,
<geoCoordinate id="EE615042FFC86F7286EAF918C2291BEA" box="[904,1068,1719,1745]" direction="west" orientation="longitude" pageId="25" pageNumber="26" precision="15" value="-82.17194">082°10’19&quot;W</geoCoordinate>
,
<quantity id="4CAD9B60FFC86F728123F918C29D1BEA" box="[1089,1176,1719,1745]" metricMagnitude="2" metricUnit="m" metricValue="7.1" pageId="25" pageNumber="26" unit="m" value="710.0">710 m</quantity>
,
<date id="FFEB1045FFC86F7281CFF918C32F1BEA" box="[1197,1322,1719,1745]" pageId="25" pageNumber="26" value="1986-06-22">22.vi.1986</date>
(Holzenthal, Heyn, &amp; Armitage) —
<specimenCount id="9D53FD0CFFC86F7287D4F970C51A1BC2" box="[694,799,1759,1785]" pageId="25" pageNumber="26" type="male">15 males</specimenCount>
(
<collectionCode id="ED44AE40FFC86F728652F970C5831BC2" box="[816,902,1759,1785]" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="25" pageNumber="26">UMSP</collectionCode>
); Reserva Forestal San Ramón, Río San Lorencito &amp; tribs.,
<geoCoordinate id="EE615042FFC86F728746F8A8C4B11A1A" box="[548,692,1799,1825]" direction="north" orientation="latitude" pageId="25" pageNumber="26" precision="15" value="10.216111">10°12’58&quot;N</geoCoordinate>
,
<geoCoordinate id="EE615042FFC86F7287A0F8A8C5631A1A" box="[706,870,1799,1825]" direction="west" orientation="longitude" pageId="25" pageNumber="26" precision="15" value="-84.60694">084°36’25&quot;W</geoCoordinate>
,
<quantity id="4CAD9B60FFC86F728611F8A8C5C71A1A" box="[883,962,1799,1825]" metricMagnitude="2" metricUnit="m" metricValue="9.8" pageId="25" pageNumber="26" unit="m" value="980.0">980 m</quantity>
,
<date id="FFEB1045FFC86F7286B2F8A8C28A1A1A" box="[976,1167,1799,1825]" pageId="25" pageNumber="26" value="1987-03-30" valueMax="1987-04-01" valueMin="1987-03-30">30.iii.–1.iv.1987</date>
(Holzenthal, Hamilton, &amp; Heyn) —
<specimenCount id="9D53FD0CFFC86F728774F880C48A1A72" box="[534,655,1839,1865]" pageId="25" pageNumber="26" type="male">951 males</specimenCount>
, 1044 females (
<collectionCode id="ED44AE40FFC86F728624F880C5921A72" box="[838,919,1839,1865]" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="25" pageNumber="26">UMSP</collectionCode>
,
<collectionCode id="ED44AE40FFC86F7286C0F880C5FD1A72" box="[930,1016,1839,1865]" pageId="25" pageNumber="26">INBIO</collectionCode>
); same,
<date id="FFEB1045FFC86F728134F880C3051A72" box="[1110,1280,1839,1865]" pageId="25" pageNumber="26" value="1988-06-13" valueMax="1988-06-16" valueMin="1988-06-13">13–16.vi.1988</date>
(C. &amp; O. Flint, Holzenthal) —
<specimenCount id="9D53FD0CFFC86F728724F8F8C4B61A4A" box="[582,691,1879,1905]" pageId="25" pageNumber="26" type="male">92 males</specimenCount>
,
<specimenCount id="9D53FD0CFFC86F7287A2F8F8C5471A4A" box="[704,834,1879,1905]" pageId="25" pageNumber="26" type="female">59 females</specimenCount>
(
<collectionCode id="ED44AE40FFC86F728631F8F8C5B11A4A" box="[851,948,1879,1905]" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="25" pageNumber="26">NMNH</collectionCode>
); same,
<date id="FFEB1045FFC86F728174F8F8C29D1A4A" box="[1046,1176,1879,1905]" pageId="25" pageNumber="26" value="1990-05-01" valueMax="1990-05-04" valueMin="1990-05-01">1–4.v.1990</date>
(Holzenthal &amp; Blahnik) —
<specimenCount id="9D53FD0CFFCB6F7184AFFEB3C44A1C0D" box="[461,591,284,310]" pageId="26" pageNumber="27" type="male">176 males</specimenCount>
,
<specimenCount id="9D53FD0CFFCB6F718700FEB3C4F21C0D" box="[610,759,284,310]" pageId="26" pageNumber="27" type="female">138 females</specimenCount>
(
<collectionCode id="ED44AE40FFCB6F71866CFEB3C5611C0D" box="[782,868,284,310]" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="26" pageNumber="27">UMSP</collectionCode>
); same,
<date id="FFEB1045FFCB6F7186B3FEB3C28E1C0D" box="[977,1163,284,310]" pageId="26" pageNumber="27" value="1990-08-28" valueMax="1990-08-30" valueMin="1990-08-28">28–30.viii.1990</date>
(Holzenthal, Blahnik, &amp; Muñoz) —
<specimenCount id="9D53FD0CFFCB6F718721FEEBC4991C65" box="[579,668,324,350]" pageId="26" pageNumber="27" type="male">6 male</specimenCount>
(
<collectionCode id="ED44AE40FFCB6F7187D7FEEBC50E1C65" box="[693,779,324,350]" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="26" pageNumber="27">UMSP</collectionCode>
); same,
<date id="FFEB1045FFCB6F71861DFEEBC2191C65" box="[895,1052,324,350]" pageId="26" pageNumber="27" value="1991-03-06" valueMax="1991-03-10" valueMin="1991-03-06">6–10.iii.1991</date>
(Holzenthal, Muñoz, Huisman) —
<specimenCount id="9D53FD0CFFCB6F7184C8FEC3C4131CBD" box="[426,534,364,390]" pageId="26" pageNumber="27" type="male">59 males</specimenCount>
,
<specimenCount id="9D53FD0CFFCB6F718741FEC3C4A11CBD" box="[547,676,364,390]" pageId="26" pageNumber="27" type="female">51 females</specimenCount>
(
<collectionCode id="ED44AE40FFCB6F7187D7FEC3C50E1CBD" box="[693,779,364,390]" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="26" pageNumber="27">UMSP</collectionCode>
); Quebrada Latas,
<quantity id="4CAD9B60FFCB6F718688FEC3C2391CBD" box="[1002,1084,364,390]" metricMagnitude="3" metricUnit="m" metricValue="8.9" pageId="26" pageNumber="27" unit="km" value="8.9">8.9 km</quantity>
NE Bajos del Toro,
<geoCoordinate id="EE615042FFCB6F71846AFE3BC79C1C95" box="[264,409,404,430]" direction="north" orientation="latitude" pageId="26" pageNumber="27" precision="15" value="10.268888">10°16’08&quot;N</geoCoordinate>
,
<geoCoordinate id="EE615042FFCB6F7184C8FE3BC4481C95" box="[426,589,404,430]" direction="west" orientation="longitude" pageId="26" pageNumber="27" precision="15" value="-84.26">084°15’36&quot;W</geoCoordinate>
,
<quantity id="4CAD9B60FFCB6F71873DFE3BC4C51C95" box="[607,704,404,430]" metricMagnitude="3" metricUnit="m" metricValue="1.03" pageId="26" pageNumber="27" unit="m" value="1030.0">1030 m</quantity>
,
<date id="FFEB1045FFCB6F7187B3FE3BC53A1C95" box="[721,831,404,430]" pageId="26" pageNumber="27" value="1990-09-06">6.ix.1990</date>
(Holzenthal, Blahnik, &amp; Huisman) —
<specimenCount id="9D53FD0CFFCB6F71807CFE3BC74A1CED" pageId="26" pageNumber="27" type="male">3 males</specimenCount>
,
<specimenCount id="9D53FD0CFFCB6F718402FE13C7D01CED" box="[352,469,444,470]" pageId="26" pageNumber="27" type="female">3 females</specimenCount>
(
<collectionCode id="ED44AE40FFCB6F718488FE13C4451CED" box="[490,576,444,470]" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="26" pageNumber="27">UMSP</collectionCode>
); Reserva Bosque Nubosa Monte Verde, Río Peñas Blancas,
<geoCoordinate id="EE615042FFCB6F71846AFE4BC79C1CC5" box="[264,409,484,510]" direction="north" orientation="latitude" pageId="26" pageNumber="27" precision="15" value="10.3">10°18’00&quot;N</geoCoordinate>
,
<geoCoordinate id="EE615042FFCB6F7184C7FE4BC44C1CC5" box="[421,585,484,510]" direction="west" orientation="longitude" pageId="26" pageNumber="27" precision="15" value="-84.74">084°44’24&quot;W</geoCoordinate>
,
<quantity id="4CAD9B60FFCB6F718736FE4BC4A71CC5" box="[596,674,484,510]" metricMagnitude="2" metricUnit="m" metricValue="9.5" pageId="26" pageNumber="27" unit="m" value="950.0">950 m</quantity>
,
<date id="FFEB1045FFCB6F7187CFFE4BC5181CC5" box="[685,797,484,510]" pageId="26" pageNumber="27" value="1986-03-01">1.iii.1986</date>
(Holzenthal &amp; Fasth) —
<specimenCount id="9D53FD0CFFCB6F718121FE4BC29A1CC5" box="[1091,1183,484,510]" pageId="26" pageNumber="27" type="male">4 males</specimenCount>
,
<specimenCount id="9D53FD0CFFCB6F7181C9FE4BC32F1CC5" box="[1195,1322,484,510]" pageId="26" pageNumber="27" type="female">15 females</specimenCount>
(
<collectionCode id="ED44AE40FFCB6F718472FDA3C7631F1D" box="[272,358,524,550]" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="26" pageNumber="27">UMSP</collectionCode>
);
<emphasis id="B921EA97FFCB6F7184EEFDA4C42D1F1E" bold="true" box="[396,552,523,549]" pageId="26" pageNumber="27">Guanacaste:</emphasis>
Parque Nacional Guanacaste, Río Tempisquito, Maritza,
<geoCoordinate id="EE615042FFCB6F71846AFD9BC79C1F75" box="[264,409,564,590]" direction="north" orientation="latitude" pageId="26" pageNumber="27" precision="15" value="10.9580555">10°57’29&quot;N</geoCoordinate>
,
<geoCoordinate id="EE615042FFCB6F7184C7FD9BC44D1F75" box="[421,584,564,590]" direction="west" orientation="longitude" pageId="26" pageNumber="27" precision="15" value="-85.49694">085°29’49&quot;W</geoCoordinate>
,
<quantity id="4CAD9B60FFCB6F718736FD9BC4A71F75" box="[596,674,564,590]" metricMagnitude="2" metricUnit="m" metricValue="5.5" pageId="26" pageNumber="27" unit="m" value="550.0">550 m</quantity>
,
<date id="FFEB1045FFCB6F7187CFFD9BC5631F75" box="[685,870,564,590]" pageId="26" pageNumber="27" value="1990-08-30" valueMax="1990-08-31" valueMin="1990-08-30">30–31.viii.1990</date>
(Huisman, Blahnik, &amp; Quesada) —
<specimenCount id="9D53FD0CFFCB6F71806CFD9BC74A1F4D" pageId="26" pageNumber="27" type="male">60 males</specimenCount>
,
<specimenCount id="9D53FD0CFFCB6F71843CFDF3C7E51F4D" box="[350,480,604,630]" pageId="26" pageNumber="27" type="female">37 females</specimenCount>
(
<collectionCode id="ED44AE40FFCB6F718491FDF3C44C1F4D" box="[499,585,604,630]" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="26" pageNumber="27">UMSP</collectionCode>
); same,
<date id="FFEB1045FFCB6F7187CCFDF3C5651F4D" box="[686,864,604,630]" pageId="26" pageNumber="27" value="1992-07-13" valueMax="1992-07-16" valueMin="1992-07-13">13–16.vii.1992</date>
(F. Munoz) —
<specimenCount id="9D53FD0CFFCB6F718140FDF3C27B1F4D" box="[1058,1150,604,630]" pageId="26" pageNumber="27" type="male">2 males</specimenCount>
(
<collectionCode id="ED44AE40FFCB6F7181F3FDF3C2EC1F4D" box="[1169,1257,604,630]" pageId="26" pageNumber="27">INBIO</collectionCode>
); Río Tempisquito Sur, Maritza,
<geoCoordinate id="EE615042FFCB6F718729FD2BC4DE1FA5" box="[587,731,644,670]" direction="north" orientation="latitude" pageId="26" pageNumber="27" precision="15" value="10.95">10°57’00&quot;N</geoCoordinate>
,
<geoCoordinate id="EE615042FFCB6F718789FD2BC58A1FA5" box="[747,911,644,670]" direction="west" orientation="longitude" pageId="26" pageNumber="27" precision="15" value="-85.48">085°28’48&quot;W</geoCoordinate>
,
<quantity id="4CAD9B60FFCB6F7186FCFD2BC5EB1FA5" box="[926,1006,644,670]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="26" pageNumber="27" unit="m" value="600.0">600 m</quantity>
,
<date id="FFEB1045FFCB6F71869FFD2BC28E1FA5" box="[1021,1163,644,670]" pageId="26" pageNumber="27" value="1990-08-30">30.viii.1990</date>
(Huisman &amp; Quesada) —
<specimenCount id="9D53FD0CFFCB6F7184C9FD03C42C1FFD" box="[427,553,684,710]" pageId="26" pageNumber="27" type="male">172 males</specimenCount>
,
<specimenCount id="9D53FD0CFFCB6F71875BFD03C4B81FFD" box="[569,701,684,710]" pageId="26" pageNumber="27" type="female">34 females</specimenCount>
(
<collectionCode id="ED44AE40FFCB6F7187B3FD03C5221FFD" box="[721,807,684,710]" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="26" pageNumber="27">UMSP</collectionCode>
); ca.
<quantity id="4CAD9B60FFCB6F71860FFD03C5C61FFD" box="[877,963,684,710]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="26" pageNumber="27" unit="km" value="0.7">0.7 km</quantity>
N Est. Maritza,
<geoCoordinate id="EE615042FFCB6F7181F7FD03C3231FFD" box="[1173,1318,684,710]" direction="north" orientation="latitude" pageId="26" pageNumber="27" precision="15" value="10.96">10°57’36&quot;N</geoCoordinate>
,
<geoCoordinate id="EE615042FFCB6F71846AFD7BC7A11FD5" box="[264,420,724,750]" direction="east" orientation="longitude" pageId="26" pageNumber="27" precision="15" value="85.5">085°30’00&quot;E</geoCoordinate>
,
<quantity id="4CAD9B60FFCB6F7184D2FD7BC7F61FD5" box="[432,499,724,750]" metricMagnitude="2" metricUnit="m" metricValue="5.5" pageId="26" pageNumber="27" unit="m" value="550.0">550m</quantity>
,
<date id="FFEB1045FFCB6F71876BFD7BC4931FD5" box="[521,662,724,750]" pageId="26" pageNumber="27" value="1990-08-31">31.viii.1990</date>
(Huisman &amp; Quesada) —
<specimenCount id="9D53FD0CFFCB6F7186B2FD7BC2391FD5" box="[976,1084,724,750]" pageId="26" pageNumber="27" type="male">16 males</specimenCount>
,
<specimenCount id="9D53FD0CFFCB6F71812AFD7BC2BC1FD5" box="[1096,1209,724,750]" pageId="26" pageNumber="27" type="female">3 females</specimenCount>
(
<collectionCode id="ED44AE40FFCB6F7181AAFD7BC31B1FD5" box="[1224,1310,724,750]" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="26" pageNumber="27">UMSP</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C34F650EFFCB6F71846AFC8BC2061EB5" pageId="26" pageNumber="27" type="etymology">
<paragraph id="8BEA3685FFCB6F71846AFC8BC7801E06" blockId="26.[264,1323,804,910]" box="[264,389,804,829]" pageId="26" pageNumber="27">
<emphasis id="B921EA97FFCB6F71846AFC8BC7801E06" box="[264,389,804,829]" italics="true" pageId="26" pageNumber="27">Etymology</emphasis>
</paragraph>
<paragraph id="8BEA3685FFCB6F71845AFCE3C2061EB5" blockId="26.[264,1323,804,910]" pageId="26" pageNumber="27">
This species is named
<taxonomicName id="4C554D06FFCB6F71875DFCE3C4921E5E" box="[575,663,844,869]" class="Insecta" family="Glossosomatidae" genus="Culoptila" kingdom="Animalia" order="Trichoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="hamata">
<emphasis id="B921EA97FFCB6F71875DFCE3C4921E5E" box="[575,663,844,869]" italics="true" pageId="26" pageNumber="27">hamata</emphasis>
</taxonomicName>
, from the Latin
<emphasis id="B921EA97FFCB6F718637FCE3C5A41E5E" box="[853,929,844,869]" italics="true" pageId="26" pageNumber="27">hamus</emphasis>
, meaning hooked, in reference to the elongate, curved, or hooked apex of tergum X of this species.
</paragraph>
</subSubSection>
</treatment>
</document>
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