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<document id="46C02DA7CB89DBA44A71AB6CC82E5039" ID-DOI="http://dx.doi.org/10.11646/zootaxa.3884.3.7" ID-GBIF-Dataset="e29b2d13-73ec-4ca3-bf3b-80e19a58bcf9" ID-ISSN="1175-5326" ID-Zenodo-Dep="229641" ID-ZooBank="2061002B-A5F9-4BBE-91CB-4C21C954B879" checkinTime="1460968257452" checkinUser="plazi" docAuthor="Szederjesi, Tímea, Pavlíček, Tomás, Latif, Robabeh &amp; Csuzdi, Csaba" docDate="2014" docId="03F99722FFEFFFAAFF252252FC6BFD62" docLanguage="en" docName="zt03884p288.pdf" docOrigin="Zootaxa 3884 (3)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@598f6591" docStyleName="zootaxa.2013.journal_article" docTitle="Eisenia kontschani Szederjesi, Pavlicek &amp; Csuzdi, sp. nov." docType="treatment" docVersion="12" lastPageNumber="284" masterDocId="FFC0EF5AFFEEFFA8FFB2244FFFB1FFE6" masterDocTitle="Review of the Eisenia muganiensis (Michaelsen, 1910) species group with description of two new species (Oligochaeta: Lumbricidae)" masterLastPageNumber="288" masterPageNumber="282" pageNumber="283" updateTime="1698303584182" updateUser="ExternalLinkService">
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<mods:title id="C89795D8A3F488CDDBEC33DDDF7E4FA6">Review of the Eisenia muganiensis (Michaelsen, 1910) species group with description of two new species (Oligochaeta: Lumbricidae)</mods:title>
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<mods:namePart id="12C03B7FDCD921DEC27518CFCDF58097">Szederjesi, Tímea</mods:namePart>
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<mods:namePart id="BBC5B921C98576D44B76127F6AF90488">Pavlíček, Tomás</mods:namePart>
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<mods:namePart id="36D83EC548D8D3E23442642F05D13442">Latif, Robabeh</mods:namePart>
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<mods:namePart id="4F191C8A5A39A0C629CAD3969F682569">Csuzdi, Csaba</mods:namePart>
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<mods:date id="687EBC5D62FDF7F1E026F1320FB7D1A3">2014</mods:date>
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<treatment id="03F99722FFEFFFAAFF252252FC6BFD62" ID-DOI="http://doi.org/10.5281/zenodo.6144270" ID-GBIF-Taxon="119593474" ID-Zenodo-Dep="6144270" LSID="urn:lsid:plazi:treatment:03F99722FFEFFFAAFF252252FC6BFD62" httpUri="http://treatment.plazi.org/id/03F99722FFEFFFAAFF252252FC6BFD62" lastPageId="2" lastPageNumber="284" pageId="1" pageNumber="283">
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<emphasis id="B924FA26FFEFFFA9FF252252FCEEF9DE" bold="true" box="[151,863,1565,1592]" pageId="1" pageNumber="283">
<taxonomicName id="4C505DB7FFEFFFA9FF252252FD4BF9DE" authority="Szederjesi, Pavlicek &amp; Csuzdi" authorityName="Szederjesi, Pavlicek &amp; Csuzdi" box="[151,762,1565,1592]" class="Clitellata" family="Lumbricidae" genus="Eisenia" higherTaxonomySource="CoL" kingdom="Animalia" order="Crassiclitellata" pageId="1" pageNumber="283" phylum="Annelida" rank="species" species="kontschani" status="sp. nov.">
<emphasis id="B924FA26FFEFFFA9FF252252FECFF9D1" bold="true" box="[151,382,1565,1591]" italics="true" pageId="1" pageNumber="283">Eisenia kontschani</emphasis>
Szederjesi, Pavlíček &amp; Csuzdi
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<taxonomicNameLabel id="A217475DFFEFFFA9FCB02251FCEEF9DE" box="[770,863,1566,1592]" pageId="1" pageNumber="283" rank="species">sp. nov.</taxonomicNameLabel>
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<figureCitation id="136B3AB1FFEFFFA9FF25220FFF44F9BF" box="[151,245,1600,1625]" captionStart="FIGURE 2" captionStartId="2.[151,250,1043,1065]" captionTargetBox="[151,1432,697,1021]" captionTargetId="figure@2.[151,1436,697,1021]" captionTargetPageId="2" captionText="FIGURE 2. Eisenia kontschani sp. nov. A. Setal arrangement. B. Anterior body with clitellar region, lateral view. Mp = male pore, Tb = tubercle. Scale bar = 1 mm." httpUri="https://zenodo.org/record/229643/files/figure.png" pageId="1" pageNumber="283">
Figure
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A,B
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<subSubSection id="C34A75BFFFEFFFA9FF2522C4FBFBF909" pageId="1" pageNumber="283" type="materials_examined">
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<typeStatus id="54EB9896FFEFFFA9FF2522C4FEB5F942" box="[151,260,1675,1700]" pageId="1" pageNumber="283">Holotype</typeStatus>
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.
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/
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. Grazed Mediterranean garrigue near Hasankeyf, Batman Province,
<collectingCountry id="F34766A4FFEFFFA9FB6E22C4FA9CF942" box="[1244,1325,1675,1700]" name="Turkey" pageId="1" pageNumber="283">Turkey</collectingCountry>
,
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o
<number id="3D983413FFEFFFA9FAD222C3FACEF942" box="[1376,1407,1676,1700]" pageId="1" pageNumber="283" value="42.0">42</number>
'N
</geoCoordinate>
,
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<number id="3D983413FFEFFFA9FF2522FCFF02F92D" box="[151,179,1715,1739]" pageId="1" pageNumber="283" value="41.0">41</number>
o
<number id="3D983413FFEFFFA9FF0922FCFF6BF92D" box="[187,218,1715,1739]" pageId="1" pageNumber="283" value="24.0">24</number>
'E
</geoCoordinate>
. Leg. T. Pavlíček &amp; P. Cardet,
<date id="FFEE00F4FFEFFFA9FDFA22FCFD7BF92D" box="[584,714,1715,1739]" pageId="1" pageNumber="283" value="2009-04-23">23.04.2009</date>
.
</paragraph>
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<typeStatus id="54EB9896FFEFFFA9FF752299FE8EF909" box="[199,319,1750,1775]" pageId="1" pageNumber="283">Paratypes</typeStatus>
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.
<collectionCode id="ED41BEF1FFEFFFA9FEFE2299FE15F909" LSID="urn:lsid:biocol.org:col:34991" box="[332,420,1750,1775]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="1" pageNumber="283">HNHM</collectionCode>
/
<number id="3D983413FFEFFFA9FE1F2299FDBDF909" box="[429,524,1750,1775]" pageId="1" pageNumber="283" value="16954.5">16954, 5</number>
ex., locality and date same as that of the
<typeStatus id="54EB9896FFEFFFA9FC682299FBF7F909" box="[986,1094,1750,1775]" pageId="1" pageNumber="283">Holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C34A75BFFFEFFFA9FF7522B5FC2FF8D1" pageId="1" pageNumber="283" type="etymology">
<paragraph id="8BEF2634FFEFFFA9FF7522B5FC2FF8D1" blockId="1.[151,1437,1672,2027]" pageId="1" pageNumber="283">
<emphasis id="B924FA26FFEFFFA9FF7522B5FE2EF8F5" bold="true" box="[199,415,1786,1811]" pageId="1" pageNumber="283">Derivatio nominis</emphasis>
. The new species is named in honour of our college, the renowned mite taxonomist Dr. Jenő Kontschán for his continuous help in our earthworm researches.
</paragraph>
</subSubSection>
<subSubSection id="C34A75BFFFEFFFAAFF75230DFCF0FDDB" lastPageId="2" lastPageNumber="284" pageId="1" pageNumber="283" type="diagnosis">
<paragraph id="8BEF2634FFEFFFA9FF75230DFE30F80D" blockId="1.[151,1437,1672,2027]" pageId="1" pageNumber="283">
<emphasis id="B924FA26FFEFFFA9FF75230DFE8BF8BD" bold="true" box="[199,314,1858,1883]" pageId="1" pageNumber="283">Diagnosis</emphasis>
. Length
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, diameter 3.5
<quantity id="4CA88BD1FFEFFFA9FD0E230DFD4FF8BC" box="[700,766,1858,1883]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="283" unit="mm" value="4.0">4 mm</quantity>
, setae closely paired. Colour red-violet dorsally. First dorsal pore in
<number id="3D983413FFEFFFA9FF5C2329FF4CF899" box="[238,253,1894,1919]" pageId="1" pageNumber="283" value="4.0">4</number>
/
<number id="3D983413FFEFFFA9FEBA2329FEA7F899" box="[264,278,1894,1919]" pageId="1" pageNumber="283" value="5.0">5</number>
. Clitellum on
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2637,
<date id="FFEE00F4FFEFFFA9FDBE2328FDA7F899" box="[524,534,1895,1919]" pageId="1" pageNumber="283" value="1938-02-01">1</date>
</number>
/
<number id="3D983413FFEFFFA9FD922328FD81F899" box="[544,560,1895,1919]" pageId="1" pageNumber="283" value="2.0">2</number>
<number id="3D983413FFEFFFA9FD852328FDE4F899" box="[567,597,1895,1919]" pageId="1" pageNumber="283" value="38.0">38</number>
, tubercles on
<number id="3D983413FFEFFFA9FD462329FC8CF899" box="[756,829,1894,1919]" pageId="1" pageNumber="283" value="34.0" valueMax="36.0" valueMin="32.0">3236</number>
. Male pore on
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, surrounded by glandular crescents. Nephridial pores not visible. Four pairs of vesicles in
<number id="3D983413FFEFFFA9FCB623C4FC8EF845" box="[772,831,1931,1955]" pageId="1" pageNumber="283" value="10.5" valueMax="12.0" valueMin="9.0">912</number>
; spermathecae two pairs in
<date id="FFEE00F4FFEFFFA9FB3223C4FAB0F845" box="[1152,1281,1931,1955]" pageId="1" pageNumber="283" value="2011-09-10" valueMax="2011-09-10" valueMin="2011-09-10">
<number id="3D983413FFEFFFA9FB3223C4FB3FF845" box="[1152,1166,1931,1955]" pageId="1" pageNumber="283" value="9.0">9</number>
/
<number id="3D983413FFEFFFA9FB2523C4FB02F845" box="[1175,1203,1931,1955]" pageId="1" pageNumber="283" value="10.0">10</number>
<number id="3D983413FFEFFFA9FB7123C4FB6EF845" box="[1219,1247,1931,1955]" pageId="1" pageNumber="283" value="10.0">10</number>
/
<number id="3D983413FFEFFFA9FB5A23C4FAB0F845" box="[1256,1281,1931,1955]" pageId="1" pageNumber="283" value="11.0">11</number>
</date>
, open near
<emphasis id="B924FA26FFEFFFA9FA3523C5FA24F844" box="[1415,1429,1930,1954]" italics="true" pageId="1" pageNumber="283">d</emphasis>
. Calciferous glands in
<number id="3D983413FFEFFFA9FE2223E0FE69F821" box="[400,472,1967,1991]" pageId="1" pageNumber="283" value="12.0" valueMax="13.0" valueMin="11.0">1113</number>
. Hearts in segments
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. Nephridial bladders simple, sausage-shaped. Longitudinal musculature pinnate.
</paragraph>
<paragraph id="8BEF2634FFECFFAAFF7524D8FB77FE82" blockId="2.[151,1436,151,645]" pageId="2" pageNumber="284">
<emphasis id="B924FA26FFECFFAAFF7524D8FE04FF56" bold="true" box="[199,437,151,176]" pageId="2" pageNumber="284">External characters</emphasis>
.
<emphasis id="B924FA26FFECFFAAFE7124D7FD98FF56" box="[451,553,152,176]" italics="true" pageId="2" pageNumber="284">
<typeStatus id="54EB9896FFECFFAAFE7124D7FD98FF56" box="[451,553,152,176]" pageId="2" pageNumber="284">Holotype</typeStatus>
</emphasis>
<quantity id="4CA88BD1FFECFFAAFD8324D7FD30FF49" box="[561,641,152,176]" metricMagnitude="-2" metricUnit="m" metricValue="7.8" pageId="2" pageNumber="284" unit="mm" value="78.0">78 mm</quantity>
long and
<quantity id="4CA88BD1FFECFFAAFD4324D7FC82FF49" box="[753,819,152,176]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="2" pageNumber="284" unit="mm" value="4.0">4 mm</quantity>
wide. Number of segments
<number id="3D983413FFECFFAAFBC324D8FB2FFF56" box="[1137,1182,151,176]" pageId="2" pageNumber="284" value="169.0">169</number>
.
<emphasis id="B924FA26FFECFFAAFB1824D6FAADFF56" box="[1194,1308,153,176]" italics="true" pageId="2" pageNumber="284">
<typeStatus id="54EB9896FFECFFAAFB1824D6FAADFF56" box="[1194,1308,153,176]" pageId="2" pageNumber="284">Paratypes</typeStatus>
</emphasis>
<quantity id="4CA88BD1FFECFFAAFA9124D8FA2DFF49" box="[1315,1436,151,176]" metricMagnitude="-2" metricUnit="m" metricValue="6.75" metricValueMax="7.9" metricValueMin="5.6" pageId="2" pageNumber="284" unit="mm" value="67.5" valueMax="79.0" valueMin="56.0">5679 mm</quantity>
long and 3.5
<quantity id="4CA88BD1FFECFFAAFE8324F3FEC3FF32" box="[305,370,188,213]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="2" pageNumber="284" unit="mm" value="4.0">4 mm</quantity>
wide. Number of segments
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. Colour red-violet dorsally. Prostomium tanylobous. First dorsal pore at intersegmental furrow
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/
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. Setae closely paired. Setal arrangement behind clitellum:
<emphasis id="B924FA26FFECFFAAFB4624AEFAA1FF1E" box="[1268,1296,225,248]" italics="true" pageId="2" pageNumber="284">aa</emphasis>
:
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:
<emphasis id="B924FA26FFECFFAAFA8E24AFFAE7FF1E" box="[1340,1366,224,248]" italics="true" pageId="2" pageNumber="284">bc</emphasis>
:
<emphasis id="B924FA26FFECFFAAFAEC24AFFAC9FF1E" box="[1374,1400,224,248]" italics="true" pageId="2" pageNumber="284">cd</emphasis>
:
<emphasis id="B924FA26FFECFFAAFA3224AFFA2DFF1E" box="[1408,1436,224,248]" italics="true" pageId="2" pageNumber="284">dd</emphasis>
= 18.6:1.3:
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:
<number id="3D983413FFECFFAAFE87254BFE8EFEFB" box="[309,319,260,285]" pageId="2" pageNumber="284" value="1.0">1</number>
:38.6 (
<figureCitation id="136B3AB1FFECFFAAFE3C254BFE62FEFA" box="[398,467,260,285]" captionStart="FIGURE 2" captionStartId="2.[151,250,1043,1065]" captionTargetBox="[151,1432,697,1021]" captionTargetId="figure@2.[151,1436,697,1021]" captionTargetPageId="2" captionText="FIGURE 2. Eisenia kontschani sp. nov. A. Setal arrangement. B. Anterior body with clitellar region, lateral view. Mp = male pore, Tb = tubercle. Scale bar = 1 mm." httpUri="https://zenodo.org/record/229643/files/figure.png" pageId="2" pageNumber="284">
Fig.
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A). Male pores on segment
<number id="3D983413FFECFFAAFCA2254BFC9FFEFB" box="[784,814,260,285]" pageId="2" pageNumber="284" value="15.0">15</number>
, surrounded by glandular crescents protruding into the neighbouring segments. Nephridial pores not visible. Clitellum on segments
<number id="3D983413FFECFFAAFC5F2568FBFAFEA6" box="[1005,1099,295,320]" pageId="2" pageNumber="284" value="31.55" valueMax="37.1" valueMin="26.0">
2637,
<date id="FFEE00F4FFECFFAAFBF32567FBFAFEA6" box="[1089,1099,296,320]" pageId="2" pageNumber="284" value="1938-02-01">1</date>
</number>
/
<number id="3D983413FFECFFAAFBE72567FBD4FEA6" box="[1109,1125,296,320]" pageId="2" pageNumber="284" value="2.0">2</number>
<number id="3D983413FFECFFAAFBD92567FB3BFEA6" box="[1131,1162,296,320]" pageId="2" pageNumber="284" value="38.0">38</number>
. Tubercula pubertatis on segments
<number id="3D983413FFECFFAAFEB42503FEE1FE83" box="[262,336,332,357]" pageId="2" pageNumber="284" value="34.0" valueMax="36.0" valueMin="32.0">3236</number>
. Glandular tumescence on segments
<number id="3D983413FFECFFAAFD5F2503FCEEFE82" box="[749,863,332,356]" pageId="2" pageNumber="284" value="11.07" valueMax="12.14" valueMin="10.0">1012, 14</number>
,
<number id="3D983413FFECFFAAFCD92503FBB4FE82" box="[875,1029,332,357]" pageId="2" pageNumber="284" value="19.765" valueMax="23.32" valueMin="16.21">16, 2123, 32</number>
<number id="3D983413FFECFFAAFBA62503FB9EFE82" box="[1044,1071,332,356]" pageId="2" pageNumber="284" value="37.0">37</number>
<emphasis id="B924FA26FFECFFAAFB842503FBE3FE82" box="[1078,1106,332,356]" italics="true" pageId="2" pageNumber="284">ab</emphasis>
(
<figureCitation id="136B3AB1FFECFFAAFBD32503FB14FE82" box="[1121,1189,332,357]" captionStart="FIGURE 2" captionStartId="2.[151,250,1043,1065]" captionTargetBox="[151,1432,697,1021]" captionTargetId="figure@2.[151,1436,697,1021]" captionTargetPageId="2" captionText="FIGURE 2. Eisenia kontschani sp. nov. A. Setal arrangement. B. Anterior body with clitellar region, lateral view. Mp = male pore, Tb = tubercle. Scale bar = 1 mm." httpUri="https://zenodo.org/record/229643/files/figure.png" pageId="2" pageNumber="284">
Fig.
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B).
</paragraph>
<paragraph id="8BEF2634FFECFFAAFF752520FCF0FDDB" blockId="2.[151,1436,151,645]" pageId="2" pageNumber="284">
<emphasis id="B924FA26FFECFFAAFF752520FE02FE6E" bold="true" box="[199,435,367,392]" pageId="2" pageNumber="284">Internal characters</emphasis>
.
<collectingCountry id="F34766A4FFECFFAAFE75253FFDB4FE6E" box="[455,517,368,392]" name="Spain" pageId="2" pageNumber="284">Septa</collectingCountry>
<number id="3D983413FFECFFAAFDA02520FDAEFE6E" box="[530,543,367,392]" pageId="2" pageNumber="284" value="6.0">6</number>
/
<number id="3D983413FFECFFAAFD9B2520FDE0FE6E" box="[553,593,367,392]" pageId="2" pageNumber="284" value="7.5" valueMax="8.0" valueMin="7.0">78</number>
/
<number id="3D983413FFECFFAAFDE82520FDD6FE6E" box="[602,615,367,392]" pageId="2" pageNumber="284" value="9.0">9</number>
strongly thickened,
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<number id="3D983413FFECFFAAFCEC253FFCDDFE6E" box="[862,876,368,392]" pageId="2" pageNumber="284" value="9.0">9</number>
/
<number id="3D983413FFECFFAAFCC7253FFC21FE6E" box="[885,912,368,392]" pageId="2" pageNumber="284" value="10.0">10</number>
<number id="3D983413FFECFFAAFC12253FFC0AFE6E" box="[928,955,368,392]" pageId="2" pageNumber="284" value="10.0">10</number>
/
<number id="3D983413FFECFFAAFC76253FFC6CFE6E" box="[964,989,368,392]" pageId="2" pageNumber="284" value="11.0">11</number>
</date>
less so. Testes and funnels paired in segments
<number id="3D983413FFECFFAAFEBA25DBFEE1FE4A" box="[264,336,404,428]" pageId="2" pageNumber="284" value="10.5" valueMax="11.0" valueMin="10.0">1011</number>
, covered with peri-oesophageal testis sac. Four pairs of seminal vesicles in
<number id="3D983413FFECFFAAFB0525DBFB42FE4A" box="[1207,1267,404,428]" pageId="2" pageNumber="284" value="10.5" valueMax="12.0" valueMin="9.0">912</number>
. Spermathecae two pairs in
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/
<number id="3D983413FFECFFAAFE8825F7FECCFE36" box="[314,381,440,464]" pageId="2" pageNumber="284" value="10.1">10, 10</number>
/
<number id="3D983413FFECFFAAFE3525F7FE11FE36" box="[391,416,440,464]" pageId="2" pageNumber="284" value="11.0">11</number>
with external openings near setal line
<emphasis id="B924FA26FFECFFAAFCFD25F7FCECFE36" box="[847,861,440,464]" italics="true" pageId="2" pageNumber="284">d</emphasis>
. Calciferous glands in segment
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without well demarcated diverticula. Paired hearts appear in segments
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, with a pair of small extraoesophageal vessels in
<number id="3D983413FFECFFAAFACB2593FA29FE12" box="[1401,1432,476,500]" pageId="2" pageNumber="284" value="12.0">12</number>
. Nephridial bladders simple, sausage-shaped. Crop in segments
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, and gizzard in segments
<number id="3D983413FFECFFAAFB74264FFABEFDFE" box="[1222,1295,512,536]" pageId="2" pageNumber="284" value="17.5" valueMax="18.0" valueMin="17.0">1718</number>
. Typhlosolis T-shaped, large. Longitudinal musculature is of pinnate
<typeStatus id="54EB9896FFECFFAAFCBB266AFC8CFDDB" box="[777,829,549,573]" pageId="2" pageNumber="284">type</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C34A75BFFFECFFAAFF752608FC6BFD62" pageId="2" pageNumber="284" type="discussion">
<paragraph id="8BEF2634FFECFFAAFF752608FC6BFD62" blockId="2.[151,1436,151,645]" pageId="2" pageNumber="284">
<emphasis id="B924FA26FFECFFAAFF752608FE85FD86" bold="true" box="[199,308,583,608]" pageId="2" pageNumber="284">Remarks</emphasis>
. The new species is close to
<taxonomicName id="4C505DB7FFECFFAAFD222606FC2EFD86" authority="Perel, 1962" authorityName="Perel" authorityYear="1962" box="[656,927,583,608]" class="Clitellata" family="Lumbricidae" genus="Eisenia" higherTaxonomySource="CoL" kingdom="Animalia" order="Crassiclitellata" pageId="2" pageNumber="284" phylum="Annelida" rank="species" species="malevici">
<emphasis id="B924FA26FFECFFAAFD222606FCA3FD86" box="[656,786,584,608]" italics="true" pageId="2" pageNumber="284">E. malevici</emphasis>
<bibRefCitation id="EFC15BC5FFECFFAAFCAE2608FC2EFD86" author="Perel" box="[796,927,583,608]" pageId="2" pageNumber="284" refString="Perel, T. S. (1962) A new earthworm species from Western Siberia: Eisenia malevici sp. n. (Oligochaeta, Lumbricidae). Zoologicheski Zhurnal, 41, 454 - 456. [in Russian]" type="journal article" year="1962">
Perel,
<number id="3D983413FFECFFAAFCD52608FC2EFD86" box="[871,927,583,608]" pageId="2" pageNumber="284" value="1962.0">1962</number>
</bibRefCitation>
</taxonomicName>
but differs from it by the somewhat longer clitellum, the tanylobic head and the number of seminal vesicles (Table
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).
</paragraph>
</subSubSection>
</treatment>
</document>