293 lines
34 KiB
XML
293 lines
34 KiB
XML
<document id="467C8BB7747369D03D4E009C1E34C9C4" ID-CLB-Dataset="6279" ID-DOI="10.5852/ejt.2021.751.1381" ID-GBIF-Dataset="4e3af683-d8a6-40de-991b-c4584d66b8cc" ID-ISSN="2118-9773" ID-Zenodo-Dep="4906007" ID-ZooBank="1D27EA41-2396-463A-B1B2-0CCF79C9F3D2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623068023717" checkinUser="felipe" docAuthor="Zamani, Alireza, Chatzaki, Maria, Esyunin, Sergei L. & Marusik, Yuri M." docDate="2021" docId="03F98785035FEF401C617E7BFAB169AF" docLanguage="en" docName="ejt-751_68-114.pdf" docOrigin="European Journal of Taxonomy 751 (1)" docSource="http://dx.doi.org/10.5852/ejt.2021.751.1381" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Scotophaeus elburzensis Zamani & Chatzaki & Esyunin & Marusik 2021, sp. nov." docType="treatment" docUuid="CD564644-82B1-4E91-974E-9C6BEEB5CBA3" docUuidSource="ZooBank" docVersion="8" lastPageNumber="101" masterDocId="FFC0FFFD037FEF611E347F73FFC16D41" masterDocTitle="One new genus and nineteen new species of ground spiders (Araneae: Gnaphosidae) from Iran, with other taxonomic considerations" masterLastPageNumber="114" masterPageNumber="68" pageNumber="100" updateTime="1699000119853" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="DB2F6D40E742CBFB63BC32C45369B0E4">One new genus and nineteen new species of ground spiders (Araneae: Gnaphosidae) from Iran, with other taxonomic considerations</mods:title>
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<mods:namePart id="5A23BB1EEC650F21D24A2CC2EB0DAECC">Zamani, Alireza</mods:namePart>
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<mods:affiliation id="1D60A297CD4982ECE7835111585A3A9F">Zoological Museum, Biodiversity Unit, University of Turku, FI- 20014, Finland. Department of Molecular Biology and Genetics, Democritus University of Thrace, Dragana, 68100 Alexandroupolis, Greece. Perm State University, Bukireva Street 15, Perm, 614600, Russia. Institute for Biological Problems of the North RAS, Portovaya Str. 18, Magadan, 685000, Russia. Department of Zoology & Entomology, University of the Free State, Bloemfontein 9300, South Africa.</mods:affiliation>
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<mods:nameIdentifier id="55BB2720D49D688C8206173A8DBF121D" type="email">zamani.alireza5@gmail.com</mods:nameIdentifier>
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<mods:namePart id="7CC6217D1FFAFCC37DB452287D92347B">Chatzaki, Maria</mods:namePart>
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<mods:nameIdentifier id="EA2CEC362E372030AD2381BC34C8E634" type="email">maria.chatzaki@gmail.com</mods:nameIdentifier>
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<mods:namePart id="917D64C8B2E0152F7844BDF87D0A48EA">Esyunin, Sergei L.</mods:namePart>
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<mods:affiliation id="F53CCEDF171011E7E7A6A000FA25D1E2">Sergei.</mods:affiliation>
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<mods:nameIdentifier id="6F0A9DD986EEB0915AE91ABB048838D1" type="email">Sergei.Esyunin@psu.ru</mods:nameIdentifier>
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<mods:namePart id="8EE65DF37AAB10B2D8DB46DD49754664">Marusik, Yuri M.</mods:namePart>
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<mods:nameIdentifier id="74279E74335068411335DF8AF6487989" type="email">yurmar@mail.ru</mods:nameIdentifier>
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<mods:title id="FCE19A41B7F88B7E0239D7B6ADF7C01E">European Journal of Taxonomy</mods:title>
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<mods:part id="7816072B4FA7C219CF36C55D917AA393">
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<mods:date id="CC782DE61086FEB10E8E7DD7DE1F3089">2021</mods:date>
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<treatment id="03F98785035FEF401C617E7BFAB169AF" ID-DOI="http://doi.org/10.5281/zenodo.4906492" ID-GBIF-Taxon="182597853" ID-Zenodo-Dep="4906492" LSID="urn:lsid:zoobank.org:act:CD564644-82B1-4E91-974E-9C6BEEB5CBA3" httpUri="http://treatment.plazi.org/id/03F98785035FEF401C617E7BFAB169AF" lastPageId="33" lastPageNumber="101" pageId="32" pageNumber="100">
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<subSubSection id="C34A6518035FEF411C617E7BFC1E6C62" box="[597,991,264,291]" pageId="32" pageNumber="100" type="nomenclature">
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<paragraph id="8BEF3693035FEF411C617E7BFC1E6C62" blockId="32.[363,1225,264,362]" box="[597,991,264,291]" pageId="32" pageNumber="100">
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<heading id="D0A781FF035FEF411C617E7BFC1E6C62" box="[597,991,264,291]" centered="true" fontSize="11" level="2" pageId="32" pageNumber="100" reason="2">
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<taxonomicName id="4C504D10035FEF411C617E7BFCBC6C62" ID-CoL="98YVY" authority="Zamani & Chatzaki & Esyunin & Marusik, 2021" authorityName="Zamani & Chatzaki & Esyunin & Marusik" authorityYear="2021" box="[597,893,264,291]" class="Arachnida" family="Gnaphosidae" genus="Scotophaeus" kingdom="Animalia" order="Araneae" pageId="32" pageNumber="100" phylum="Arthropoda" rank="species" species="elburzensis" status="sp. nov.">
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<emphasis id="B924EA81035FEF411C617E7BFCBC6C62" bold="true" box="[597,893,264,291]" italics="true" pageId="32" pageNumber="100">Scotophaeus elburzensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A21757FA035FEF411DB07E7AFC1E6C62" box="[900,991,265,291]" pageId="32" pageNumber="100" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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<subSubSection id="C34A6518035FEF411F5F7E5FFC416C28" pageId="32" pageNumber="100" type="description">
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<paragraph id="8BEF3693035FEF411F5F7E5FFB086C06" blockId="32.[363,1225,264,362]" box="[363,1225,300,327]" pageId="32" pageNumber="100">
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urn:lsid:zoobank.org:act:
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</uri>
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</paragraph>
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<paragraph id="8BEF3693035FEF411C877E3CFC416C28" blockId="32.[363,1225,264,362]" box="[691,896,335,362]" pageId="32" pageNumber="100">
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<figureCitation id="136B2A16035FEF411C877E3CFC966C2B" box="[691,855,335,362]" captionStart="Fig" captionStartId="31.[189,232,1858,1884]" captionTargetBox="[189,1393,771,1817]" captionTargetId="figure-136@31.[189,1398,771,1819]" captionTargetPageId="31" captionText="Fig. 19. A–C, G–H. Scotophaeus anahita sp. nov., holotype, ♀ (ZMMU). A, H. Epigyne, ventral view. B–C. Epigyne, dorsal view. G. Chelicera, posterior view. – D–F, I. Scotophaeus elburzensis sp. nov., ♀ (D–E = specimen from Tehran Province, SMF; F, I = holotype, ZMMU). D, I. Epigyne, ventral view. E–F. Epigyne, dorsal view. Abbreviations: Bl = basal lobe; Ld = lateral duct; Lm = lateral margin; rLd = ridge of lateral duct; Tl = terminal lobe. Scale bars: A, C, F, H–I = 0.1 mm; D–E = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4906061" httpUri="https://zenodo.org/record/4906061/files/figure.png" pageId="32" pageNumber="100">Figs 19D–F, I</figureCitation>
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,
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<figureCitation id="136B2A16035FEF411D577E3CFC416C28" box="[867,896,335,361]" captionStart="Fig" captionStartId="32.[189,231,1962,1988]" captionTargetBox="[207,1398,1284,1919]" captionTargetId="figure-216@32.[189,1398,1284,1923]" captionTargetPageId="32" captionText="Fig. 20. Scotophaeus elburzensis sp. nov. (A, C = ♀ from Tehran Province, SMF; B = holotype, ♀, ZMMU). A–B. Habitus, dorsal view. C. Chelicerae, posterior view. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="http://doi.org/10.5281/zenodo.4906063" httpUri="https://zenodo.org/record/4906063/files/figure.png" pageId="32" pageNumber="100">20</figureCitation>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C34A6518035FEF411E897EE3FE5C6F90" pageId="32" pageNumber="100" type="diagnosis">
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||
<paragraph id="8BEF3693035FEF411E897EE3FEF76CEB" blockId="32.[189,310,400,426]" box="[189,310,400,426]" pageId="32" pageNumber="100">
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<heading id="D0A781FF035FEF411E897EE3FEF76CEB" bold="true" box="[189,310,400,426]" fontSize="11" level="3" pageId="32" pageNumber="100" reason="3">
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<emphasis id="B924EA81035FEF411E897EE3FEF76CEB" bold="true" box="[189,310,400,426]" pageId="32" pageNumber="100">Diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BEF3693035FEF411E897ECCFE5C6F90" blockId="32.[188,1400,447,721]" pageId="32" pageNumber="100">
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The new species is easily distinguished from its congeners by the atrium which is trapezoidal posteriorly and anteriorly an open dome, and by the peculiar placement of the two pairs of receptacles, one vertical to the other. As in the similar
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<taxonomicName id="4C504D10035FEF411C257D75FD486F61" authority="Zamani & Chatzaki & Esyunin & Marusik, 2021" authorityName="Zamani & Chatzaki & Esyunin & Marusik" authorityYear="2021" box="[529,649,518,544]" class="Arachnida" family="Gnaphosidae" genus="Scotophaeus" kingdom="Animalia" order="Araneae" pageId="32" pageNumber="100" phylum="Arthropoda" rank="species" species="anahita" status="sp. nov.">
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<emphasis id="B924EA81035FEF411C257D75FD486F61" box="[529,649,518,544]" italics="true" pageId="32" pageNumber="100">S. anahita</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A21757FA035FEF411CBB7D75FD2A6F61" box="[655,747,518,544]" pageId="32" pageNumber="100" rank="species">sp. nov.</taxonomicNameLabel>
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, the posterior part of the epigynal atrium is most similar to that of
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<taxonomicName id="4C504D10035FEF411F007D5AFE586F02" authorityName="Wunderlich" authorityYear="1995" box="[308,409,553,579]" class="Arachnida" family="Gnaphosidae" genus="Scotophaeus" kingdom="Animalia" order="Araneae" pageId="32" pageNumber="100" phylum="Arthropoda" rank="species" species="nanus">
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<emphasis id="B924EA81035FEF411F007D5AFE586F02" box="[308,409,553,579]" italics="true" pageId="32" pageNumber="100">S. nanus</emphasis>
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</taxonomicName>
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(see
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<bibRefCitation id="EFC14B62035FEF411FEC7D5AFD686F05" author="Wunderlich J." box="[472,681,553,580]" pageId="32" pageNumber="100" pagination="743 - 747" refId="ref18382" refString="Wunderlich J. 1995. Beschreibung einer bisher unbekannten Art der Gattung Scotophaeus Simon aus Osterreich (Arachnida: Araneae: Gnaphosidae). Beitrage zur Araneologie 4 (1994): 743 - 747. [publ. in Dec. 1995]" type="book chapter" year="1995">Wunderlich 1995</bibRefCitation>
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: figs 6–8), but the rest of the epigynal characters are unique enough to consider it a new species. This species can be further diagnosed from the related
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<taxonomicName id="4C504D10035FEF411ACB7D3FFAB66F27" authority="Zamani & Chatzaki & Esyunin & Marusik, 2021" authorityName="Zamani & Chatzaki & Esyunin & Marusik" authorityYear="2021" box="[1279,1399,588,615]" class="Arachnida" family="Gnaphosidae" genus="Scotophaeus" kingdom="Animalia" order="Araneae" pageId="32" pageNumber="100" phylum="Arthropoda" rank="species" species="anahita" status="sp. nov.">
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<emphasis id="B924EA81035FEF411ACB7D3FFACC6F27" box="[1279,1293,588,614]" italics="true" pageId="32" pageNumber="100">S</emphasis>
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.
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<emphasis id="B924EA81035FEF411B297D3FFAB66F27" box="[1309,1399,588,614]" italics="true" pageId="32" pageNumber="100">anahita</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A21757FA035FEF411E897D03FED86FCB" box="[189,281,624,650]" pageId="32" pageNumber="100" rank="species">sp. nov.</taxonomicNameLabel>
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by the lateral ducts being relatively straight and slightly curved outward (vs spiraled) and the terminal lobes being larger than the basal lobes and located posteriorly (vs smaller than basal lobes and located anteriorly).
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C34A6518035FEF411E897D84FE6D6E22" pageId="32" pageNumber="100" type="etymology">
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<paragraph id="8BEF3693035FEF411E897D84FE856E50" blockId="32.[189,324,759,785]" box="[189,324,759,785]" pageId="32" pageNumber="100">
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<heading id="D0A781FF035FEF411E897D84FE856E50" bold="true" box="[189,324,759,785]" fontSize="11" level="3" pageId="32" pageNumber="100" reason="3">
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<emphasis id="B924EA81035FEF411E897D84FE856E50" bold="true" box="[189,324,759,785]" pageId="32" pageNumber="100">Etymology</emphasis>
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</heading>
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||
</paragraph>
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<paragraph id="8BEF3693035FEF411E897C55FE6D6E22" blockId="32.[189,1398,806,867]" pageId="32" pageNumber="100">
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The specific epithet is an adjective and refers to the
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<collectingRegion id="4994F871035FEF411D2F7C55FCA86E01" box="[795,873,806,832]" country="Iran" name="Alborz" pageId="32" pageNumber="100">Elburz</collectingRegion>
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Mountain Range, from where the specimens have been collected.
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</paragraph>
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</subSubSection>
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<subSubSection id="C34A6518035FEF411E897CF9FE6F69C8" pageId="32" pageNumber="100" type="materials_examined">
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<paragraph id="8BEF3693035FEF411E897CF9FEAD6EE5" blockId="32.[189,364,906,932]" box="[189,364,906,932]" pageId="32" pageNumber="100">
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<heading id="D0A781FF035FEF411E897CF9FEAD6EE5" bold="true" box="[189,364,906,932]" fontSize="11" level="3" pageId="32" pageNumber="100" reason="3">
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<emphasis id="B924EA81035FEF411E897CF9FEAD6EE5" bold="true" box="[189,364,906,932]" pageId="32" pageNumber="100">Type material</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BEF3693035FEF411EDE7CCAFA826EB6" blockId="32.[189,1347,953,1015]" pageId="32" pageNumber="100">
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<materialsCitation id="3B383CCE035FEF411EDE7CCAFA826EB6" ID-GBIF-Occurrence="3127434307" collectingDate="2017-05-25" collectionCode="ZMMU" collectorName="L. Karami" country="Iran" latitude="36.1" location="Gheydar" longLatPrecision="1189" longitude="48.566666" pageId="32" pageNumber="100" specimenCount="1" specimenCount-female="1" stateProvince="Zanjan Province" typeStatus="holotype">
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<emphasis id="B924EA81035FEF411EDE7CCAFE9D6E92" bold="true" box="[234,348,953,979]" pageId="32" pageNumber="100">
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<typeStatus id="54EB8831035FEF411EDE7CCAFE9D6E92" box="[234,348,953,979]" pageId="32" pageNumber="100" type="holotype">Holotype</typeStatus>
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</emphasis>
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<collectingCountry id="F3477603035FEF411E897CAFFEC96EB7" box="[189,264,988,1014]" name="Iran" pageId="32" pageNumber="100">IRAN</collectingCountry>
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•
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<specimenCount id="9D56FD1A035FEF411F167CAEFEF86EB6" box="[290,313,989,1015]" pageId="32" pageNumber="100" type="female">♀</specimenCount>
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;
|
||
<collectingRegion id="4994F871035FEF411F737CAFFDCA6EB7" box="[327,523,988,1014]" country="Iran" name="Zanjan" pageId="32" pageNumber="100">Zanjan Province</collectingRegion>
|
||
,
|
||
<location id="8E8F6048035FEF411C237CAFFD416EB7" LSID="urn:lsid:plazi:treatment:03F98785035FEF401C617E7BFAB169AF:8E8F6048035FEF411C237CAFFD416EB7" box="[535,640,988,1014]" country="Iran" latitude="36.1" longLatPrecision="1189" longitude="48.566666" name="Gheydar" pageId="32" pageNumber="100" stateProvince="Zanjan Province">Gheydar</location>
|
||
;
|
||
<geoCoordinate id="EE645054035FEF411CB87CA9FD396EB6" box="[652,760,986,1015]" degrees="36" direction="north" minutes="06" orientation="latitude" pageId="32" pageNumber="100" precision="925" value="36.1">36°06′ N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE645054035FEF411D307CA9FCAC6EB6" box="[772,877,986,1015]" degrees="48" direction="east" minutes="34" orientation="longitude" pageId="32" pageNumber="100" precision="925" value="48.566666">48°34′ E</geoCoordinate>
|
||
;
|
||
<date id="FFEE1053035FEF411D4D7CAFFBD86EB7" box="[889,1049,988,1015]" pageId="32" pageNumber="100" value="2017-05-25">
|
||
<collectingDate id="EFAAE9BB035FEF411D4D7CAFFBD86EB7" box="[889,1049,988,1015]" pageId="32" pageNumber="100" value="2017-05-25">25 May 2017</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="26A55345035FEF411A117CAEFB5F6EB7" box="[1061,1182,988,1015]" pageId="32" pageNumber="100">L. Karami</collectorName>
|
||
leg.;
|
||
<collectionCode id="ED41AE56035FEF411AD47CAEFAFE6EB6" box="[1248,1343,989,1015]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="32" pageNumber="100" type="Museum">ZMMU</collectionCode>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BEF3693035FEF411E897B6EFE386976" blockId="32.[189,505,1053,1079]" box="[189,505,1053,1079]" pageId="32" pageNumber="100">
|
||
<heading id="D0A781FF035FEF411E897B6EFE386976" bold="true" box="[189,505,1053,1079]" fontSize="11" level="3" pageId="32" pageNumber="100" reason="3">
|
||
<emphasis id="B924EA81035FEF411E897B6EFE386976" bold="true" box="[189,505,1053,1079]" pageId="32" pageNumber="100">Other material examined</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BEF3693035FEF411E897B3FFE6F69C8" blockId="32.[189,1398,1100,1162]" pageId="32" pageNumber="100">
|
||
<materialsCitation id="3B383CCE035FEF411E897B3FFE6B69C8" ID-GBIF-Occurrence="3127434324" collectingDate="1978-05-25" collectionCode="SMF" collectorName="J. Martens & H. Pieper" country="Iran" elevation="2450" location="Koolak" pageId="32" pageNumber="100" specimenCount="1" specimenCount-female="1" stateProvince="Tehran Province">
|
||
<collectingCountry id="F3477603035FEF411E897B3FFEC96927" box="[189,264,1100,1126]" name="Iran" pageId="32" pageNumber="100">IRAN</collectingCountry>
|
||
•
|
||
<specimenCount id="9D56FD1A035FEF411F147B3FFE8D6926" box="[288,332,1100,1127]" pageId="32" pageNumber="100" type="female">1 ♀</specimenCount>
|
||
;
|
||
<collectingRegion id="4994F871035FEF411F6D7B3FFDDD6927" box="[345,540,1100,1126]" country="Iran" name="Tehran" pageId="32" pageNumber="100">Tehran Province</collectingRegion>
|
||
,
|
||
<location id="8E8F6048035FEF411C137B3FFDBC6927" LSID="urn:lsid:plazi:treatment:03F98785035FEF401C617E7BFAB169AF:8E8F6048035FEF411C137B3FFDBC6927" box="[551,637,1100,1126]" country="Iran" name="Koolak" pageId="32" pageNumber="100" stateProvince="Tehran Province">Koolak</location>
|
||
(= Kolakchal);
|
||
<elevation id="007DD1A0035FEF411D027B3FFBD76927" box="[822,1046,1100,1127]" metricMagnitude="3" metricUnit="m" metricValue="2.45" metricValueMax="2.5" metricValueMin="2.4" pageId="32" pageNumber="100" unit="m" value="2450.0" valueMax="2500.0" valueMin="2400.0">
|
||
<quantity id="4CA89B76035FEF411D027B3FFC166926" box="[822,983,1100,1127]" metricMagnitude="3" metricUnit="m" metricValue="2.45" metricValueMax="2.5" metricValueMin="2.4" pageId="32" pageNumber="100" unit="m" value="2450.0" valueMax="2500.0" valueMin="2400.0">2400–2500 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
;
|
||
<date id="FFEE1053035FEF411A177B3FFB006927" box="[1059,1217,1100,1127]" pageId="32" pageNumber="100" value="1978-05-25">
|
||
<collectingDate id="EFAAE9BB035FEF411A177B3FFB006927" box="[1059,1217,1100,1127]" pageId="32" pageNumber="100" value="1978-05-25">25 May 1978</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="26A55345035FEF411AF87B3EFA846926" box="[1228,1349,1101,1127]" pageId="32" pageNumber="100">J. Martens</collectorName>
|
||
and
|
||
<collectorName id="26A55345035FEF411E897B03FEED69C8" box="[189,300,1135,1162]" pageId="32" pageNumber="100">H. Pieper</collectorName>
|
||
leg.;
|
||
<collectionCode id="ED41AE56035FEF411F5A7B1CFE6B69C8" box="[366,426,1135,1161]" collectionName="Germany, Frankfurt-am-Main, Forschungsinstitut und Naturmuseum Senckenberg" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="32" pageNumber="100">SMF</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF2F661B035FEF411E8978D9FAAC6AA6" ID-DOI="http://doi.org/10.5281/zenodo.4906063" ID-Zenodo-Dep="4906063" httpUri="https://zenodo.org/record/4906063/files/figure.png" pageId="32" pageNumber="100" startId="32.[189,231,1962,1988]" targetBox="[207,1398,1284,1919]" targetPageId="32">
|
||
<paragraph id="8BEF3693035FEF411E8978D9FAAC6AA6" blockId="32.[189,1399,1962,2024]" pageId="32" pageNumber="100">
|
||
<emphasis id="B924EA81035FEF411E8978D9FED56A85" bold="true" box="[189,276,1962,1988]" pageId="32" pageNumber="100">Fig. 20.</emphasis>
|
||
<taxonomicName id="4C504D10035FEF411F2C78D9FDEC6A85" authority="Zamani & Chatzaki & Esyunin & Marusik, 2021" authorityName="Zamani & Chatzaki & Esyunin & Marusik" authorityYear="2021" box="[280,557,1962,1988]" class="Arachnida" family="Gnaphosidae" genus="Scotophaeus" kingdom="Animalia" order="Araneae" pageId="32" pageNumber="100" phylum="Arthropoda" rank="species" species="elburzensis" status="sp. nov.">
|
||
<emphasis id="B924EA81035FEF411F2C78D9FDEC6A85" box="[280,557,1962,1988]" italics="true" pageId="32" pageNumber="100">Scotophaeus elburzensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21757FA035FEF411C0778D8FD4A6A84" box="[563,651,1963,1989]" pageId="32" pageNumber="100" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(A, C = ♀ from Tehran Province, SMF; B = holotype, ♀, ZMMU).
|
||
<emphasis id="B924EA81035FEF411E8978BEFF356AA6" bold="true" box="[189,244,1997,2023]" pageId="32" pageNumber="100">A–B</emphasis>
|
||
. Habitus, dorsal view.
|
||
<emphasis id="B924EA81035FEF411C3678BEFDD66AA6" bold="true" box="[514,535,1997,2023]" pageId="32" pageNumber="100">C</emphasis>
|
||
. Chelicerae, posterior view. Scale bars: 0.2 mm, unless stated otherwise.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C34A6518035EEF401E897E7BFEF86E84" pageId="33" pageNumber="101" type="description">
|
||
<paragraph id="8BEF3693035EEF401E897E7BFE8F6C63" blockId="33.[189,334,264,290]" box="[189,334,264,290]" pageId="33" pageNumber="101">
|
||
<heading id="D0A781FF035EEF401E897E7BFE8F6C63" bold="true" box="[189,334,264,290]" fontSize="11" level="3" pageId="33" pageNumber="101" reason="3">
|
||
<emphasis id="B924EA81035EEF401E897E7BFE8F6C63" bold="true" box="[189,334,264,290]" pageId="33" pageNumber="101">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BEF3693035EEF401EDE7E44FE066C13" blockId="33.[188,1400,311,693]" box="[234,455,311,338]" pageId="33" pageNumber="101">
|
||
<emphasis id="B924EA81035EEF401EDE7E44FE846C10" bold="true" box="[234,325,311,337]" pageId="33" pageNumber="101">Female</emphasis>
|
||
(
|
||
<typeStatus id="54EB8831035EEF401F617E44FE7E6C13" box="[341,447,311,338]" pageId="33" pageNumber="101" type="holotype">holotype</typeStatus>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8BEF3693035EEF401E897E28FC116FF5" blockId="33.[188,1400,311,693]" pageId="33" pageNumber="101">
|
||
Habitus as in
|
||
<figureCitation id="136B2A16035EEF401F687E28FE2B6C34" box="[348,490,347,373]" captionStart="Fig" captionStartId="32.[189,231,1962,1988]" captionTargetBox="[207,1398,1284,1919]" captionTargetId="figure-216@32.[189,1398,1284,1923]" captionTargetPageId="32" captionText="Fig. 20. Scotophaeus elburzensis sp. nov. (A, C = ♀ from Tehran Province, SMF; B = holotype, ♀, ZMMU). A–B. Habitus, dorsal view. C. Chelicerae, posterior view. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="http://doi.org/10.5281/zenodo.4906063" httpUri="https://zenodo.org/record/4906063/files/figure.png" pageId="33" pageNumber="101">Fig. 20A–B</figureCitation>
|
||
. Total length 10.8. Carapace 5.25 long, 3.45 wide. Eye sizes and interdistance of PMEs: AME = 0.17, ALE = 0.15, PME = 0.14, PLE = 0.17, PME–PME = 0.13. Carapace yellowish brown, covered with dark, short setae. Chelicerae (
|
||
<figureCitation id="136B2A16035EEF401D267ED1FCBA6CFD" box="[786,891,418,444]" captionStart="Fig" captionStartId="32.[189,231,1962,1988]" captionTargetBox="[207,1398,1284,1919]" captionTargetId="figure-216@32.[189,1398,1284,1923]" captionTargetPageId="32" captionText="Fig. 20. Scotophaeus elburzensis sp. nov. (A, C = ♀ from Tehran Province, SMF; B = holotype, ♀, ZMMU). A–B. Habitus, dorsal view. C. Chelicerae, posterior view. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="http://doi.org/10.5281/zenodo.4906063" httpUri="https://zenodo.org/record/4906063/files/figure.png" pageId="33" pageNumber="101">Fig. 20C</figureCitation>
|
||
) brown, with 3 promarginal teeth. Sternum yellowish brown with dark borders, covered with black setae. Maxillae and labium yellow-brown with white tips. Legs yellowish brown with lighter femora. Abdomen gray, covered with white plumose hairs and dark setae dorsally, and grayish white with dark setae ventrally. Spinnerets yellowish brown and uniform in color. Measurements of legs: I: 12.20 (3.50, 1.95, 2.85, 2.30, 1.60), II: 12.00 (3.50, 1.95, 2.70, 2.30, 1.55), III: 11.50 (3.30, 1.70, 2.40, 2.55, 1.55), IV: 15.25 (4.00, 2.00, 3.45, 4.05, 1.75). Spination: I: Fe: 2d, 1pl; Ti: 4v; Mt: 2v. II: Fe: 2d, 2pl; Ti: 3v; Mt: 2v. III: Fe: 3d, 2pl, 2rl; Ti: 1d, 3pl, 3rl, 6v; Mt: 5pl, 5rl, 6v. IV: Fe: 3d, 2pl, 2rl; Ti: 2d, 3pl, 3rl, 6v; Mt: 5pl, 5rl, 6v.
|
||
</paragraph>
|
||
<paragraph id="8BEF3693035EEF401E897DAFFE486E21" blockId="33.[189,1399,732,864]" pageId="33" pageNumber="101">
|
||
Epigyne as in
|
||
<figureCitation id="136B2A16035EEF401F517DAFFDC46FB7" box="[357,517,732,758]" captionStart="Fig" captionStartId="31.[189,232,1858,1884]" captionTargetBox="[189,1393,771,1817]" captionTargetId="figure-136@31.[189,1398,771,1819]" captionTargetPageId="31" captionText="Fig. 19. A–C, G–H. Scotophaeus anahita sp. nov., holotype, ♀ (ZMMU). A, H. Epigyne, ventral view. B–C. Epigyne, dorsal view. G. Chelicera, posterior view. – D–F, I. Scotophaeus elburzensis sp. nov., ♀ (D–E = specimen from Tehran Province, SMF; F, I = holotype, ZMMU). D, I. Epigyne, ventral view. E–F. Epigyne, dorsal view. Abbreviations: Bl = basal lobe; Ld = lateral duct; Lm = lateral margin; rLd = ridge of lateral duct; Tl = terminal lobe. Scale bars: A, C, F, H–I = 0.1 mm; D–E = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4906061" httpUri="https://zenodo.org/record/4906061/files/figure.png" pageId="33" pageNumber="101">Fig. 19D–F, I</figureCitation>
|
||
. Atrium bordered by lateral margin (
|
||
<emphasis id="B924EA81035EEF401D837DAEFC1D6FB7" box="[951,988,733,758]" italics="true" pageId="33" pageNumber="101">Lm</emphasis>
|
||
); lateral ducts (
|
||
<emphasis id="B924EA81035EEF401AA27DAFFB746FB7" box="[1174,1205,732,758]" italics="true" pageId="33" pageNumber="101">Ld</emphasis>
|
||
) slightly curved outward, originating posteriorly to the atrium, forming a chitinized ʻridgeʼ to the prolateral side of the lateral margin (
|
||
<emphasis id="B924EA81035EEF401F4C7C57FE6B6E7C" box="[376,426,804,829]" italics="true" pageId="33" pageNumber="101">rLD</emphasis>
|
||
); each receptacle consists of two spherical lobes (
|
||
<emphasis id="B924EA81035EEF401A3A7C50FBE96E7C" box="[1038,1064,803,829]" italics="true" pageId="33" pageNumber="101">Bl</emphasis>
|
||
and
|
||
<emphasis id="B924EA81035EEF401A527C50FBBE6E7C" box="[1126,1151,803,829]" italics="true" pageId="33" pageNumber="101">Tl</emphasis>
|
||
) connected by large copulatory ducts.
|
||
</paragraph>
|
||
<paragraph id="8BEF3693035EEF401EDE7CFBFEEB6EE3" blockId="33.[189,313,904,965]" box="[234,298,904,930]" pageId="33" pageNumber="101">
|
||
<emphasis id="B924EA81035EEF401EDE7CFBFEEB6EE3" bold="true" box="[234,298,904,930]" pageId="33" pageNumber="101">Male</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BEF3693035EEF401E897CD8FEF86E84" blockId="33.[189,313,904,965]" box="[189,313,939,965]" pageId="33" pageNumber="101">Unknown.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34A6518035EEF401E897C9EFE10693C" pageId="33" pageNumber="101" type="discussion">
|
||
<paragraph id="8BEF3693035EEF401E897C9EFE856946" blockId="33.[189,324,1005,1031]" box="[189,324,1005,1031]" pageId="33" pageNumber="101">
|
||
<heading id="D0A781FF035EEF401E897C9EFE856946" bold="true" box="[189,324,1005,1031]" fontSize="11" level="3" pageId="33" pageNumber="101" reason="3">
|
||
<emphasis id="B924EA81035EEF401E897C9EFE856946" bold="true" box="[189,324,1005,1031]" pageId="33" pageNumber="101">Comments</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BEF3693035EEF401E897B6FFE10693C" blockId="33.[189,1399,1052,1149]" pageId="33" pageNumber="101">
|
||
There are differences in the size and position of the receptacles between the
|
||
<typeStatus id="54EB8831035EEF401A027B6EFBA86976" box="[1078,1129,1053,1079]" pageId="33" pageNumber="101">type</typeStatus>
|
||
specimen from
|
||
<collectingRegion id="4994F871035EEF401B117B6FFAB76977" box="[1317,1398,1052,1078]" country="Iran" name="Zanjan" pageId="33" pageNumber="101">Zanjan</collectingRegion>
|
||
(
|
||
<figureCitation id="136B2A16035EEF401EF27B4CFE86691B" box="[198,327,1087,1114]" captionStart="Fig" captionStartId="31.[189,232,1858,1884]" captionTargetBox="[189,1393,771,1817]" captionTargetId="figure-136@31.[189,1398,771,1819]" captionTargetPageId="31" captionText="Fig. 19. A–C, G–H. Scotophaeus anahita sp. nov., holotype, ♀ (ZMMU). A, H. Epigyne, ventral view. B–C. Epigyne, dorsal view. G. Chelicera, posterior view. – D–F, I. Scotophaeus elburzensis sp. nov., ♀ (D–E = specimen from Tehran Province, SMF; F, I = holotype, ZMMU). D, I. Epigyne, ventral view. E–F. Epigyne, dorsal view. Abbreviations: Bl = basal lobe; Ld = lateral duct; Lm = lateral margin; rLd = ridge of lateral duct; Tl = terminal lobe. Scale bars: A, C, F, H–I = 0.1 mm; D–E = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4906061" httpUri="https://zenodo.org/record/4906061/files/figure.png" pageId="33" pageNumber="101">Fig. 19F, I</figureCitation>
|
||
) and the additional material examined from
|
||
<collectingRegion id="4994F871035EEF401D5C7B4CFC7A6918" box="[872,955,1087,1113]" country="Iran" name="Tehran" pageId="33" pageNumber="101">Tehran</collectingRegion>
|
||
(
|
||
<figureCitation id="136B2A16035EEF401DF87B4CFB9F691B" box="[972,1118,1087,1114]" captionStart="Fig" captionStartId="31.[189,232,1858,1884]" captionTargetBox="[189,1393,771,1817]" captionTargetId="figure-136@31.[189,1398,771,1819]" captionTargetPageId="31" captionText="Fig. 19. A–C, G–H. Scotophaeus anahita sp. nov., holotype, ♀ (ZMMU). A, H. Epigyne, ventral view. B–C. Epigyne, dorsal view. G. Chelicera, posterior view. – D–F, I. Scotophaeus elburzensis sp. nov., ♀ (D–E = specimen from Tehran Province, SMF; F, I = holotype, ZMMU). D, I. Epigyne, ventral view. E–F. Epigyne, dorsal view. Abbreviations: Bl = basal lobe; Ld = lateral duct; Lm = lateral margin; rLd = ridge of lateral duct; Tl = terminal lobe. Scale bars: A, C, F, H–I = 0.1 mm; D–E = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4906061" httpUri="https://zenodo.org/record/4906061/files/figure.png" pageId="33" pageNumber="101">Fig. 19D–E</figureCitation>
|
||
), which are considered intraspecific variations.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34A6518035EEF401E897BD7FAB169AF" box="[189,1392,1188,1262]" pageId="33" pageNumber="101" type="distribution">
|
||
<paragraph id="8BEF3693035EEF401E897BD7FE9769FF" blockId="33.[189,342,1188,1214]" box="[189,342,1188,1214]" pageId="33" pageNumber="101">
|
||
<heading id="D0A781FF035EEF401E897BD7FE9769FF" bold="true" box="[189,342,1188,1214]" fontSize="11" level="3" pageId="33" pageNumber="101" reason="3">
|
||
<emphasis id="B924EA81035EEF401E897BD7FE9769FF" bold="true" box="[189,342,1188,1214]" pageId="33" pageNumber="101">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BEF3693035EEF401E897BA7FAB169AF" blockId="33.[189,1392,1235,1262]" box="[189,1392,1235,1262]" pageId="33" pageNumber="101">
|
||
Known only from the listed localities in
|
||
<collectingRegion id="4994F871035EEF401CAC7BA0FD2869AC" box="[664,745,1235,1261]" country="Iran" name="Zanjan" pageId="33" pageNumber="101">Zanjan</collectingRegion>
|
||
and
|
||
<collectingRegion id="4994F871035EEF401D167BA0FCB269AC" box="[802,883,1235,1261]" country="Iran" name="Tehran" pageId="33" pageNumber="101">Tehran</collectingRegion>
|
||
Provinces, northwestern and northern
|
||
<collectingCountry id="F3477603035EEF401B0D7BA7FAAD69AF" box="[1337,1388,1236,1262]" name="Iran" pageId="33" pageNumber="101">Iran</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |